GEOLOGIJA 49/1, 7–44, Ljubljana 2006
Late Eocene benthic foraminiferal fauna from clastic sequence
of the Socka - Dobrna area and its chronostratigraphic
importance (Slovenia)
Poznoeocenska benti~na foraminiferna favna iz klasti~nega zaporedja med Socko in Dobrno in njen kronostratigrafski pomen
Franc CIMERMAN1, Bogomir JELEN2 & Dragomir SKABERNE2
1Paleontolo{ki in{titut Ivana Rakovca ZRC-SAZU, Novi trg 2, SI -1000 Ljubljana
cimerman@zrc-sazu.si
2Geolo{ki zavod Slovenije, Dimi~eva 14, SI -1109 Ljubljana
bogomir.jelen@geo-zs.si;   dragomir.skaberne@geo-zs.si;
Key words: Benthic foraminifera, Upper Eocene, Priabonian, Socka beds, Socka, Dobrna, Slovenia
Klju~ne besede: benti~ne foraminifere, zgornji eocen, priabonij, sote{ke plasti, Socka, Dobrna, Slovenija
Abstract
We surveyed, sampled and described Late Eocene (Priabonian) benthic foraminiferal fauna from two sections in the type locality of the Socka beds, between Socka and Dobrna. We present a short history of research of these beds emphasising their dating, from the middle of 19th century on. We describe the lithological characteristics and lithostratigraphy of the sections. In total 118 taxa of benthic foraminifers were identified, and the Priabonian age of the Soca beds established.
Kratka vsebina
V ~lanku obravnavamo pozno eocensko (priabonijsko) benti~no foraminiferno favno iz dveh profilov, ki smo ju posneli in vzor~evali na tipi~nem ozemlju sote{kih plasti med Socko in Dobrno. V uvodu je predstavljena kratka zgodovina raziskovanj sote{kih plasti, predvsem problem njihove starosti, od sredine 19. stoletja do danes. Opisane so litolo{ke zna~ilnosti in litostratigrafija obeh profilov. Skupno je bilo dolo~enih 118 taksonov benti~nih foraminifer in ugotovljena priabonijska starost sote{kih plasti.
Introduction                                    Southern Karavanke Tertiary, the type locality of the Socka beds between Socka and This paper contains only a portion of a         Dobrna (Fig. 1), and to compare it with the broader Geological Survey of Slovenia         stratigraphy in other Slovenian Tertiary project, Tertiary Tectofacies of the Southern         tectonostratigraphic units. The history sec-Karavanke Mountains. Its goal was to de-         tion was written by Bogomir Jelen, the litho-scribe and evaluate the stratigraphy of the         logic and lithostratigraphic description of
8
Franc Cimerman, Bogomir Jelen & Dragomir Skaberne
Fig. 1. Location map showing Socka-Dobrna area and detailed position of studied sections
sections by Dragomir Skaberne, and benthic foraminifers were described by Franc Cimerman.
Historical development of the problem
The problems associated with the stratigraphy of the Southern Karavanke Tertiary were recognized in the middle of the 19th century because of interest in the fast-growing coal industry. Coal exposures are not rare in basal Tertiary beds in central and eastern Slovenia, at that time a part of the Austrian-Hungarian Empire. These were targeted by geologists from the just established State Geological Survey in Vienna. Initially they distinguished between higher grade Glanzkohle (German term for bituminous and sub-bituminous coal in English terminology), which occurred in the northern coal belt between the Drava and Savinja rivers, and the
lower grade brown coal from the southern belt between the Savinja and Sava rivers.
Morlot (1849a) was the first to assign an Eocene age to the basal coal-bearing strata at Socka and Dobrna in the northern coal belt, and in 1850 the Miocene age of the southern, brown coal-bearing strata (note that the Oligocene Formation was established later, in 1854). M orlot’s (1849a) statement was based on Unger’s opinion (in Morlot, 1849b) of an Eocene age of the Socka macroflora. Influenced by the pale-obotanist Buch (1851), Morlot in 1853 changed his mind, and accepted a Miocene age for the Socka flora. Thus he started the painful flip-flopping leaps from one age of the Socka beds to another and back, which lasted for a century and a half. Ettings-hausen (1853) noted that the fossil macro-flora from the Zagorje locality in the southern coal belt was like the Eocene flora with many Miocene species.
Late Eocene benthic foraminiferal fauna from clastic sequence of the Socka - Dobrna area...             9
Rolle (1857, 1858) first tackled “this hard problem” in his own words, after the introduction of the Oligocene as a new epoch between the Eocene and the Miocene Formations. He was not very successful, but he was probably the first researcher who used the term Socka beds (in German, Sotzka-Schichten; Rolle, 1858) for the basal Tertiary fresh water coal-bearing strata of the middle and northern coal belts. Namely, he introduced in the former north coal belt the middle coal belt, based on lithological differences and fossil content, but without evidence for age differentiation (Rolle, 1857). He (Rolle , 1858) assumed the Socka beds were Eocene and/or Oli-gocene in age. Ettingshausen (1858) published an article which revised Unger’s flora and proposed a Late Eocene age of the Socka beds. Zollikofer (1859) stated that the coal-bearing strata of the southern coal belt are Oligocene. In contrast, in his 1861/ 1862 work in which Zollikofer dealt with the area east of the Savinja river, he dismissed the Oligocene age and subdivided the coal-bearing strata in the southern coal belt between the Eocene to Oligocene and the earliest Neogene age.
Ettingshausen (1872, 1877, 1885) dealt with the fossil leaf assemblages from the southern coal belt in detail. The assemblage from the coal footwall indicated a tropical, or at least subtropical climate, whereas the assemblage from the hangingwall indicated it formed in the temperate one. It followed that the tropical/subtropical flora corresponded to the Eocene, and the temperate flora to the Aquitanian. Stur (1864, 1871) advanced the idea that Socka and Eibiswald (in Slovene: Ivnik) beds from the southern rim of the Graz basin are of the same age, i.e. of Early Neogene age. He (S t u r , 1871) was the first to extend the term of Socka beds to the southern coal belt under the name Eibiswald and Socka beds. Bittner (1884) stated concisely that it was previously known that the Tertiary succession of the southern belt consists of two main rock units: the lower coal-bearing Socka beds unit, which is most probably of Oli-gocene age, and the upper unit which corresponds to the Neogene development of the Vienna basin in Austria. Hoernes (1876, 1892) temporarily solved the question of the age of the coal-bearing strata at the base of
the Tertiary system. In 1892 he formulated the following conclusions (paraphrased): 1) Socka beds, fresh water coal-bearing strata of the northern and middle coal belt belong to a much older stage than the coal-bearing strata of the southern belt. In the northern coal belt they partly belong to the Cretaceous Formation, which is more or less proved, while the age of the other part of northern and middle coal belt is still problematic. With regard to the underlying Cretaceous coal-bearing strata of the northern coal belt, they are not considerably younger, but Eocene in age. 2) The Oligocene coal-bearing strata in the southern coal belt have less in common with the Socka beds than they do with the Lower Miocene Eibiswald coal-bearing strata. Teller (1889,1896, 1898, 1907), expressed the opinion that Eocene deposits are completely missing. Tertiary deposits are divided into the Oligocene and the Miocene “lithostratigraphic” units. The coal-bearing strata comprise the youngest unit of the Oligocene succession in the northern, middle and southern coal belts.
Petrascheck (1926/1929) summarized existing stratigraphic ideas. He considered all coal-bearing strata at the base of the Tertiary System to be Socka beds and accepted their Oligocene age. Papp (1954, 1955) inferred the Chattian (Upper Oli-gocene) age from the Miogypsina (Miogy-psinoides) cf. formosensis Jabe & Hanzawa and Lepidocyclina (Nephrolepidina) tourn-oueri (Lemoine & Douvillé) found in the conformable marine marly clay, overlying the Socka beds of the southern coal-belt. Ku{~er (1955) proposed the idea that the Chattian rocks were missing in the southern coal belt. Szöts (1956) reasoned that the Chattian stage was the equivalent of the Aquitanian stage, therefore, according to him, the Socka beds were of the Aquitanian, i.e. of Early Miocene age, with Oligocene layers completely missing. Rijavec (1958) followed Papp’s view. Majzon (1958) argued for a Lattorfian (Lower Oligocene) age of the Socka beds. Thenius (1959) was convinced that Anthracotherium illyricum Teller and Miogypsina (Miogypsinoides) cf. formosensis Jabe & Hanzawa indicated the (late) Stampian (late Chattian) age of the Socka beds in the southern coal belt.
Ku{~er’s (1967) stratigraphic positional relationship scheme was based on the Rupe-
10
Franc Cimerman, Bogomir Jelen & Dragomir Skaberne
lian (Middle Oligocene) age of the Socka beds and the conformable overlying marine marly clay in the southern and middle coal belts. Their age was determined by the characteristic benthic microforaminiferal taxa accompanying Tritaxia szaboi (Hantken) from the marine marly clay which exhibited a close similarity with the Middle Oligocene (at that time Rupelian) microforaminiferal fauna of Hungary and Transylvania (NW Rumania). Then, in 1975, Papp came forth with new taxonomic results: his Miogypsina (Miogypsinoides) formosensis Jabe & Hanzawa and Miogypsina (Miogypsinoides) bantamensis Tan Sin Hok from the marine marly clay overlying Socka beds in the southern coal belt indicated a late Early Egerian (= late Chattian, Late Oligocene) age. From the map of M i o ~ & @ n i d a r ~ i ~ (1977), one can conclude that at the Socka site, the coal-bearing strata are Helvetian s.l. in age – equivalent to Ottnangian/Kar-patian. Cimerman (1979) kept the Rupelian age for the “Socka” beds in the southern coal belt. He (ibid.) stated that the use of term Socka beds in the southern coal belt is not justified and proposed to write Socka in quotation marks – “Socka” beds. The same year saw revival of the idea of Eocene age deposits in central Slovenia. Miku` (1979) advanced the thesis that the Socka beds of the western Southern Karavanke Mts. (middle coal belt) were not Rupelian, but Middle Eocene in age.
Drobne et al. (1979) published a Late Eocene age of a very small limestone patch in the eastern Southern Karavanke Mts. Mio~ (1978) distinguished between the Middle Oligocene and the Helvetian s. l. Socka beds in the Socka-Dobrna type area. Buser (1979) and Premru (1983) retained the Middle Oligocene (Rupelian) age of the southern coal belt Socka beds, while Buser allowed their Eocene age in the middle coal belt, though not naming them as such. Ani~i} & Juri{a (1985a) avoided the term Socka beds. In the southern coal belt Ani~i} & Juri{a (1985a, b) recognized both, the Middle Oligocene (Rupelian) and the Late Oligocene (Chattian) coal-bearing strata. In the middle coal belt, Ani~i} & Juri{a (1985a, b) assigned the Helvetian s.l. age to the basal coal-bearing strata, remarking that an older age was possible. B aldi (1984) correlated the Socka beds with the   nano-
plakton zone NP 23, (Rupelian). Drobne et al. (1985) correlated the Socka beds of the western Southern Karavanke Mts. (middle coal belt) with the late Middle Eocene, and the Socka beds of the southern coal belt with the Early Oligocene NP 21/22.
Hamrla (1987) presented a detailed stratigraphy of the Zre~e area in the northern coal belt. He distinguished between the Late Cretaceous, older Paleogene and Helvetian s.l. coal-bearing beds. However, Brezigar (in Brezigar et al. 1988) was the first to speculate about the Eocene age of the Socka beds in the whole Southern Karavanke geotectonic unit, and thought that the Tertiary stratigraphy might be different within the Southern Karavanke and Savinja Alps geotectonic units. R o y d e n & Baldi (1988) once again correlated the Socka beds of the middle and southern coal belt with the NP 23 zone. In the same year Mihajlovi} & Jungwirth (1988) also placed the Socka beds of the southern coal belt into the NP 23 zone, as did Pav{i~ and J e l e n (in J e l e n et al., 1990). C s o n t o s et al. (1992) considered the lithothamnium limestone at the base of the Tertiary succession in the southern coal belt to be Late Eocene, followed by a gap between it and the overlying “Socka” beds of the NP 23 zone. In 1992, our working group, dedicated to solving the problem of this high strati-graphic inconsistency, published the first note on its work (Jelen et al., 1992).
The above summary shows how confusing and instable the position of the basal Tertiary coal-bearing strata was. Since each change in their age attribution resulted into changes in age upsection, and accordingly a revision of their stratigraphical relationships, we can understand how critical the drastic age inconsistency of the basal coal-bearing strata was.
The problem was to find a solution to the issue or, how to get the key to go to the root of the problem that had been baffling so many past workers.
The first palynological inspection of the coal-bearing strata at the base of the Tertiary successions showed great differences in the composition of palynological assemblages from the northern, the middle, and the southern coal belts (Jelen, unpublished). After palynological analysis we applied a structural-stratigraphic modelling to
Late Eocene benthic foraminiferal fauna from clastic sequence of the Socka - Dobrna area...           11
establish an as much as possible flawless lithochronology of the middle coal belt, in which the Socka-Dobrna type area is situated. Luckily, in the type area the litho-chronology could be satisfactorily established. Sampling was first performed in successions where the lithochronology was considered to be well established. Foramin-iferal samples were collected from one metre long trenches in 1-metre intervals, if outcrop conditions were favourable, or at each metre, if considered necessary. In addition, we developed a new laboratory preparation method that allowed us to recover highly concentrated foraminifers (Jelen, unpublished).
Lithology and lithostratigraphy in the Vra~ek section area
The composite Vra~ek section (Vr/I and Vr/II) is situated north of Dobrna, south of Vra~kov vrh (708m), along the way from the @ebljar farm north of the Klanec village toward the Trojn{ek farm (Figs. 1, 2a). This area is more structuraly complex than was expected at the beginning of field work.
Tectonically disturbed limestones and pyroclastic rocks of Triassic age and older Eocene limestones are overlain by Upper Priabonian (Jelen et al., 2000), mostly clastic sedimentary rocks. The contact is partly tectonized. Eocene beds were folded together with basement rocks. All units are uncon-formably overlain by Miocene coarse-grained rocks with remains of ostrean valves of the Klanec beds and sandy marly rocks of the Dobrna marl.
Eocene beds are slightly bent into a syn-cline. The area is cut by several Dinaric (NW – SE), subvertical, sinistral faults into narrow, 20 to 60 m wide bands or wedges. Within the bands the Eocene marly beds are compressed, crushed and fissile. Fissility is steep and chiefly fault-parallel. In non-fissil parts, the rocks are predominantly massive, in places faintly laminated. The section comprises somewhat more than 200 m of the Eocene beds.
In the first, shorter segment of section, which is separated from the second main segment by a fault, Eocene rocks occur on tectonically disturbed green volcaniclastic rocks (tuffs, tuffites with thin intercalations
of dark grey biomicritic limestones) of presumably Middle Triassic age.
The Eocene succession starts with a one meter thick, prevailingly light to medium grey fine-crystalline dolomite (unit 1 on Fig. 2a). The rock contains lenses, laminae and thin layers of xylitic coal, 2 mm to 5 cm thick in its lower half. In the upper part traces of plant root systems can be seen as well as limonitized, originally pyritic concretions.
The dolomitic bed is followed by a 1.7 m thick clayey coal interval (unit 2 on Fig. 2a). It consists of 5 to 30 cm thick layers of carbargillitic, detritic and xylitic coal and two 2 to 5 cm thick predominantly clayey beds with a lower content of organic, coaly components.
Above the coaly interval follows a dark yellowish brown, somewhat weathered, marly silty shale of weakly expressed parallel lamination (unit 3 on Fig. 2a) The rock can be compared to weathered yellowish grey marly silty shale at the beginning of the second, main part of the section. That rock is 14 m thick, it overlies the tectonized limestone with numerous calcitic veins, and is probably Triassic in age.
In the second part of the section, the yellow grey marly silty shale passes into brownish grey to dark brownish massive silty marl-stone (unit 4 on Fig. 2a) approximately 20 m thick. It contains numerous ostracods and occasional bivalve and gastropod shells.
The dark brownish massive silty marl-stone passes into yellowish grey, somewhat weathered marly silty shale (unit 5 on Fig. 2a) comprising a thickness of 7 m, up to the contact with a limestone lens.
The limestone lens (unit 6 on Fig. 2a) is 5.2 m thick and can be genetically divided in two parts. The lower part, 4.2 m thick, bordering along a sharp contact on the underlying yellowish grey marly silty shale, was macroscopically subdivided into three sequences. They start with up to 15 cm thick layers of calcirudite with angular clasts of intramicrite and biointramicrite up to 4 cm in size. On the lower side of these layers and within overlying macroscopically homogeneous, 0.7 to 2.4 m thick intervals of biomicritic limestone stylolites parallel to bedding are visible. In the upper part the lens is terminated with a 1.2 m thick normally graded carbonate turbiditic sequence.
12
Franc Cimerman, Bogomir Jelen & Dragomir Skaberne
Fig. 2. Stratigraphic columns of:
a) Vra~ek section (Vr I and Vr II) b) Zavrh section (Zv II). Karpatian part was also determined by foraminifers but this is not a topic
of this article
Late Eocene benthic foraminiferal fauna from clastic sequence of the Socka - Dobrna area...           13
Above the limestone lens lies a medium to light grey, partly weathered yellowish grey, tectonically disturbed marly silty shale (unit 7 on Fig. 2a). Macrofossils are rare. In places rare plant remains can be found. In the section above the limestone lens more than 150 m of real thickness of these rocks are comprised.
The lithostratigraphically highest part of marly rocks (of lithostratigraphic unit 7) in the Vra~ek section is exposed in the Zavrh area, where it was covered with the Zavrh (Zv) section.
Lithology and lithostratigraphy in the Zavrh section area
The Zavrh (Zv) section is situated between Zavrh and ^re{kova in the area of the Vratca settlement (Figs. 1, 2b). The lower part of Tertiary beds to pre-Tertiary rocks is covered. Along the cartway leading to ^re{kova, however, they are abundantly exposed. Approximately parallel to the section a fault passes in the NW-SE direction. The horizontal component of the dextral displacement along the fault amounts to about 27 m.
The contact between the pre-Tertiary basement and Tertiary rocks is tectonical. In the basement a crystalline dolomite (dolo-sparite) occurs, on which erosional remains of red, probably weathered muddy rocks occur higher in the slope. The Tertiary sequence starts with a light grey silty clay. The thickness of the covered part of Tertiary rocks is an estimated 30 m. In rare outcrops, a grey massive to slightly laminated silty marlstone with rare plant remains occurs. The rock is brownish weathered and exhibits spheroidal parting. In the lower part occur rare, up to 0.1m thick layers of biodolomicrite with terrigenous admixture.
In the area of the carefully logged Zavrh (Zv/II) section there is a weathered light yelow grey silty marlstone with poorly expressed lamination in its lower part (unit 1 on Fig. 2b). Additionally, there are plant remains among the fossils, and in the middle part ostracods and rare echinoderm fragments. We found no fossils in the upper portion of Zv/II.
Following an angular discordance (250), a 6.5 m thick lenticular body of blocky conglomerate (unit 2 on Fig. 2b) overlies the
silty marlstone. The blocky conglomerate is cut off in the SW by the aforementioned fault that passes parallel to the section. Toward the NE the conglomerate thins and becomes finer grained. It contains numerous fossil fragments: of red algae (lithothamnia), bivalves, bryozoans, echinoderms and of other organisms. The blocky conglomerate consists of approximately 75% blocks and pebbles and 25% of muddy matrix. Blocks and pebbles range in size from 1 to 100 cm, on an average 20 cm, and are very poorly sorted. The size distribution of pebbles and blocks is bimodal. The first mode is in the range from 1 to 20 cm, and the second one in the 30 to 100 cm range. Nineteen blocks, 20 cm or more in size, and 4 smaller pebbles were examined microscopically. In these we recognized 11 lithofacies, represented in 1 to 4 blocks or pebbles: dolomitic breccia with limonitized dolomicritic cement, dolomicro-sparite with brecciated structure, crystalline dolosparite, dolomitized biomicrite, altered crystallolithic (andesitic) volcaniclastic rock, calcareous dolomite breccia, calcitic micaceous quartzy very fine-grained sandstone, partly recrystallized volcanic glass with microliths, recrystallized intrabio-micrite, intrabiomicrite with admixture of terrigenous components, and biopeldolo-micrite. Blocks and pebbles are of Triassic to inclusively Eocene age, and are derived in part directly from the underlying rocks.
The blocky conglomerate is followed at the end of the section by monotonous parallelly laminated, slightly micaceous marly silty shale (unit 3 on Fig. 2b) with rare up to 1mm thick lenses of very fine-grained sandstone. Along with rare plant remains, visibile with naked eye, there is also a fora-miniferal fauna.
Benthic foraminiferal fauna from the Socka beds
Eocene benthic foraminifers from Vra~ek II (Vr/II-) and Zavrh II (Zv/II-) sections were studied. In the two sections a total of 118 taxa were determined, belonging to 85 genera and 53 families.
Vra~ek II
From the VRA^EK II section (Fig. 2a) 12 samples (Vr/II-001, 002, 003, 004, 005, 005A,
14
Franc Cimerman, Bogomir Jelen & Dragomir Skaberne
009, 010, 011, 011A, 012, 014) were analyzed. In the section occur 107 taxa, out of them 49 only here. The highest number of taxa, 53, were found in sample Vr-005A, 39 in sample Vr-005, and in the remaining samples from 2 to 29 taxa.
Zavrh II
From the ZAVRH II section (Fig. 2b) 8 samples were analyzed, (Zv/II-011, 010, 009, 008, 007, 006, 005, 004, samples in sections are numbered in stratigraphic order, from the bottom upwards). In the section occur 75 taxa, of them 12 only here. The maximum taxa number was found in Zv/II-6, 37, and the smallest in the uppermost collected sample Zv/II-004, only 6; otherwise the number of taxa varies in a sample between 21 and 37.
Ecological interpretation of foraminiferal fauna
In samples Vr/II-001, 002 no foraminifers are present, only ostracods. The first fora-minifers appear in sample Vr/II-003 as individual hauerinids: Quinqueloculina sp. 3 and Pyrgo subsphaerica (d’Orbigny). This part of the Vra~ek II section represents the passage from fresh water or brackish environment into a marine setting. Foraminifers become more numerous in sample Vr/II-004, which is entirely marine. The Zv II section is wholly marine. In samples Vr/II-004, 005, 005A the following large foraminiferal genera occur: Heterostegina sp., Operculina cf. gomezi (two juvenile forms), Discocyclina trabayensis vicenzensis (this species was determined by Vlasta ]osovi} ), Astero-cyclina sp., Eoannularia eocenica, and the large miliolid Borelis vonderschmitti. The species Calcarina lecalvezae was established in both sections (Vr-014, Zv-009). These are shallow marine species, and are, according to their ecological characteristics, in contradiction with the remaining foraminiferal fauna in the same samples. Therefore we presume that the fauna was transported, washed out from shallower areas into the deeper sea. In the Vra~ek II section appears in marl a limestone lens with discocyclinas (the foraminiferal fauna has not yet been determined and interpreted). In Fig. 2a the lens is marked with number 6. Also at the
Povlak farm, about 1km NW of Dobrna, an Eocene limestone with nummulites, discocyclinas, Borelis, and corals is exposed. The limestone at Povlak and the limestone lens at the Vra~ek II section may represent remains of sediments of the shallow sea from which the large foraminifers were redepos-ited. For some more frequent genera, Murray (1991, 288-290 and 323-326) reports ecological data from recent environments. Among them appear 20 genera that are identical to genera from our sections. For each of these genera the habitat extends from the shelf (either internal or external) to the bathyal. The shelf is indicated also by the miliolids (Quin-queloculina). The upper slope (200-2000 m) is indicated by genera Ammobaculites, Bathy-siphon, Karreriella, Repmanina, Textularia, Bulimina, Globocassidulina, Hoeglundina, Pullenia, Reussella, Trifarina and Uvigerina. The genus Laticarinina that was found in both sections is an indicator of even greater bathyal depths. However, the observation of a relatively small number of planktonic forms is consistent with deposition of the sediments in the bathyal domain, but rather on the exernal shelf with a passage to the bathyal. According to the bathymetric interpretation of sediments at Biarritz (Gruenig & H e rb , 1984) they were deposited in the realm extending from neritic to bathyal.
Comparison with Late Eocene faunas in Europe
By comparisons of the occurrence of the species in Vra~ek II and Zavrh II sections with those in other European localities of the Eocene (Possagno in north Italy, Buda marl in Hungary and Upper Eocene beds with Tritaxia szaboi in Hungary and Biarritz in France) we want to demonstrate the degree of similarity of our fauna with them.
In north Italy, B r a g a et al. (1975) established in Paleocene, and Lower and Middle Eocene at Possagno 99 benthic species, of which 10 (=8%) are also our species. From the Upper Eocene 126 benthic species were determined. Among these 13 (=11%) are found in our sections.
In the Buda marl Sztràkos (1982) established 471 forms of foraminifers, out of these 416 were determined to the species or subspecies level. 42 (=35%) of our species
Late Eocene benthic foraminiferal fauna from clastic sequence of the Socka - Dobrna area...           15
are identical to those. From the Bartonian and Priabonian beds with Tritaxia szaboi, Sztràkos (1987) determined 429 species. Of these, 45 (=38%) are also our species.
The fauna from the Biarritz locality was investigated by H alkyard (1919), G r u e -n i g & H e r b (1984), Gruenig (1985) and Sztràkos (In: Mathelin & Sztràkos , 1993). Halkyard (1919) established the presence of 345 foraminiferal species. Gruenig (1985) determined 87 species. By far the most complete review was performed by Sztràkos (In: Mathelin & Sztràkos, 1993) who recorded 557 species. With the species from Biarritz 77 (=65%) species are identical, or two thirds of species from the Vra~ek II and Zavrh II sections are found in Biarritz. Outstanding among them is Marginulina globosa Halkyard. It was previously known only from Biarritz, and Halkyard’s specimen was the only one of this species until Sztràkos found additional three individuals in the Aquitanian basin in beds of the zone with Turborotalia cerroazulensis s. l. (Middle Priabonian) (Sztràkos, 1993, 44. In: M a t h e l i n & S z t r à k o s ). The species is obviously very rare. Our find is from the third and easternmost locality. The species is interesting and important as an indicator of the connection of our Late Eocene sea toward the west.
Dating of the Socka beds
For dating the species of benthic fora-minifers that are restricted to Eocene and that do not survive the Eocene-Oligocene boundary are critical. There are 22 such species (Heterostegina sp. does not belong to them) in our two sections:
Pavonitina biarritzensis Dorothia fallax Karrerotextularia olianensis Sigmoilopsis bartoniensis Borelis vonderschmitti Reticulopalmula arborescens Marginulina globosa Spirolingulina acutimargo Ceratocancris crassa Lamarckina cristellaroides Sporobulimina eocaena Sagrinopsis aspera Tubulogenerina tubulifera
Uvigerina chirana
Schlosserina asterites
Laticarinina altocamerata
Halkyardia minima
Melonis halkyardi
Eoannularia eocenica
Calcarina lecalvezae
Operculina cf. gomezi
Discocyclina trabayensis vicenzensis
Serra-Kiel et al. (1998) proposed the biozoning of the Paleocene and Eocene on the basis of shallow marine benthic fora-minifers in 20 SB (shallow benthic) biozones. The last two biozones, SB 19 and SB 20, comprise the Priabonian (upper part of NP 18 to lowermost part of NP 21 or upper part of P 15 to P 17. After Serra-Kiel et al. 1998) Several species extend vertically through biozones, in our sections. These are:
- Borelis vonderschmitti which comprises the top part of SB 18, the whole SB 19 and lower part of SB 20
- Operculina gomezi covers the upper part of SB 18, entire SB 19 and entire SB20
- Halkyardia minima comprises the zones SB 15 to the end of SB 20
-   Discocyclina trabajensis vicenzensis comprises SB 20
- Calcarina lecalvezae comprises the zones SB 17 and SB 18.
The authors of Calcarina lecalvezae, Bou-langer & Poignant (1971, 96), report the for Upper Eocene for its range, but they also found it in the Upper Lutetian. Hete-rostegina is also stratigraphically important and is represented by a few individuals in the Vra~ek II section; it does not appear before the Priabonian (H o t t i nger , 1977, 142; Loeblich & Tappan , 1987, 684).
The most data for detailed dating could be obtained from planktic foraminifers. However, they are rare relative to benthic foraminifers. From samples Vr/II-005, 010, 011, 011A, 014 and Zv/II-009 Monique Toumarkine determined 15 planktic species belonging to four genera and three families. In a written communication she qualified them as species without great stratigraphic value, as for example Catapsidrax dissimilis (Cushman & Bermudez), Globorotalia suteri Bolli, the range of which extends from the Eocene into the Miocene. The other ones are globigerinas from the Eocene-Oligocene boundary. Only speciesTurborotalia cerro-
16
Franc Cimerman, Bogomir Jelen & Dragomir Skaberne
azulensis cerroazulensis (Cole) indicates a Late Eocene age. Foraminifers are not very well preserved, they are often deformed and their apertures filled with sediment.
The benthic and planktic foraminifers permit us to conclude that the beds in the Vra~ek II and Zavrh II sections are Priabonian in age, which also confirms the Priabonian age of the Socka beds.
Systematics
Order FORAMINIFERIDA Eichwald, 1830 Family: Bathysiphonidae Avnimelech, 1952
Bathysiphon taurinensis Sacco, 1893
Pl. 1, Fig. 1
1893 Bathysiphon taurinensis - Sacco, p. 168, pl. 11, fig. 2 (Fide Ellis & Messina 1940)
1975 Bathysiphon taurinense Sacco - Popescu, p.
32, pl. 1, fig. 2 Locality: Vr-005, Vr-005A, Zv-008, Zv-009, Zv-011
Family: Ammodiscidae Reuss, 1862
Repmanina charoides (Jones & Parker), 1860
1860 Trochammina squamata Jones & Parker var. charoides - Jones & Parker, p. 304 (Fide Ellis & Messina 1940)
1982 Glomospira charoides (Jones & Parker) -Foraminiferi padani, pl. 1, fig. 3
1975 Glomospira charoides (Jones & Parker) -
Braga et. al., p. 102 Locality: Zv-010
Family: Lituolidae de Blaiville, 1827
Ammobaculites agglutinans (d’Orbigny), 1846
Pl. 1, Figs. 2-3
1846 Spirolina agglutinans - d’Orbigny, p. 137, pl. 7, figs. 10-12
1985 Ammobaculites agglutinans (d’Orbigny) -Papp & Schmid, p. 196, pl. 45, figs. 6-9
1985 Haplophragmoides - Ammobaculites sp. -Grünig, p. 255, pl. 1, figs. 10-12
1993 Ammobaculites agglutinans (d’Orbigny) -Sztràkos (In: Mathelin & Sztràkos), p. 73, pl. 1, fig. 15
Locality: Vr-012
Family: Spiroplectamminidae Cushman, 1927
Spiroplectinella dalmatina (de Witt Puyt), 1941
Pl. 1, Fig. 4
1941 Textularia dalmatina - de Witt Puyt, p. 45, pl. 1, figs. 4-5
1975 Spiroplectammina dalmatina (de Witt Puyt)
-  Braga et al., p. 102
1985 Spiroplectammina dalmatina (de Witt Puyt)
-  Grünig, p. 256, pl. 2, figs. 1-4 Locality: Vr-010, Zv-008, Zv-009
Vulvulina haeringensis (Gümbel), 1868
Pl. 1, Fig. 6
1868 Venilina haeringensis - Gümbel, p. 71, pl. 2, fig. 84
1975 Vulvulina haeringensis (Gümbel) - Braga et al., p 91, pl. 1, figs. 4, 23; p. 102
1985 Vulvulina haeringensis (Gümbel) - Grünig, p. 256, pl. 2, figs. 10-12
1993 Vulvulina haeringensis (Gümbel) - Sztràkos (In: Mathelin & Sztràkos), p. 73, pl. 2, fig. 12 Locality: Vr-004, Vr-005, Vr-005A, Vr-009, Vr-011, Vr-012, Zv-006, Zv-008
Family: Pavonitinidae Loeblich & Tappan, 1961
Pavonitina biarritzensis Sztràkos, 1987
Pl. 1, Figs. 7-8
1987b Pavonitina biarritzensis - Sztràkos, p. 129, pl. 1, figs. 1-3, pl. 3, figs. 1-3, textfig. 1
1993 Pavonitina biarritzensis Sztràkos - Sztràkos (In: Mathelin & Sztràkos), p. 73, pl. 24, fig. 5 Locality: Vr-005A, Zv-005, Zv-006, Zv-008, Zv-010, Zv-011
Family: Prolixoplectidae Loeblich & Tappan, 1985
Plectina dalmatina (Schubert), 1811
Pl. 1, Figs. 9-10
1911 Gaudryina dalmatina Schubert - Liebus, p. 75, pl. 3, fig. 5
1919 Clavulina gaudryinoises Halkyard non Fornasini - Halkyard, p. 46, pl. 3, figs. 1-3
1937aPlectina dalmatina (Schubert) - Cushman, p. 107, pl. 12, fig. 8
1985  Plectina dalmatina (Schubert) - Grünig, p.
158, pl. 3, figs. 1-6 Locality: Vr-004, Vr-005, Vr-009, Vr-010, Vr-012, Zv-006, Zv-007, Zv-009, Zv-010, Zv-011
Late Eocene benthic foraminiferal fauna from clastic sequence of the Socka - Dobrna area...           17
PLATE 1
1 Bathisyphon taurinensis Sacco, Vr-009, x28. 2-3 Ammobaculites agglutinans (d’Orbigny). 2 Vr-012, x40. 3 Vr-012, x40. 4 Spiroplectinella dalmatina (de Witt Puyt), Vr-009, x60. 5 Spiroplectinella pectinata (Hantken), Vr-010, x40. 6 Vulvulina haeringensis (Gümbel), Vr-009, x40. 7-8 Pavonitina biarritzensis Sztràkos. 7 Zv/II-006, x40. 8 Zv/II-006, x40. 9-10 Plectina dalmatina (Schubert). 9 Vr-009, x40, 10 Vr-009, x40. 11-12 Tritaxia kruhelensis (Woicik). 11 Vr-012, x40. 12 Vr-012, x40. 13 Tritaxia szaboi (Hantken), Vr-009, x40.
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Franc Cimerman, Bogomir Jelen & Dragomir Skaberne
Family: Tritaxiidae Plotnikova, 1979
Tritaxia kruhelensis (Woicik), 1903
Pl. 1, Figs. 11-12
1903 Clavulina szaboi var. kruhelensis - Woicik, p. 498, pl. 6, fig. 21 (Fide Ellis & Messina 1940)
1956 Clavulinoides kruhelensis (Woicik) - Hagn, p. 117, pl. 10, figs. 2-4, 8-9
1993 Clavulinoides kruhelensis (Woicik) -Sztràkos (In: Mathelin & Sztràkos), p. 73, pl. 3, fig. 19
Locality: Vr-012, Zv-004
Tritaxia szaboi (Hantken), 1868
Pl. 1, Fig. 13
1868 Clavulina Szaboi - Hantken, p. 83, pl. 1, figs. 4, 6, 7
1868 Rhabdogonium Szaboi - Hantken, p. 90, pl. 1, fig. 18
1956 Clavulinoides Szaboi (Hantken) - Hagn, p. 116, pl. 10, fig. 1
1975 Tritaxia szaboi (Hantken) - Braga et al. p. 103, pl. 4, figs. 1-2
1993  Clavulinoides szaboi (Hantken) - Sztràkos (In: Mathelin & Sztràkos), p. 73, pl. 4, fig. 1; pl. 25, fig. 5 Locality: Vr-004, Vr-005A, Zv-005, Zv-006, Zv-008, Zv-010, Zv-011
Family: Eggerellidae Cushman, 1937
Dorothia fallax Hagn, 1956
Pl. 2, Figs. 1-2
1956 Dorothia fallax - Hagn, p. 119, pl. 9, figs. 18-19; pl. 18, fig. 10
1975 Dorothia fallax Hagn - Braga et al., p. 92, pl. 1, fig. 25
1993  Dorothia    fallax    Hagn    -    Sztràkos    (In: Mathelin & Sztràkos), p. 39, pl. 24, fig. 8 Locality: Zv-010, Zv-011
Karreriella halkyardi Cushman, 1936
Pl. 2, figs. 3-4
1936 Karreriella halkyardi - Cushman, p. 36, pl. 5, fig. 16
1985 Karreriella halkyardi Cushman - Grünig, p. 258, pl. 3, figs. 21-22
1993 Karreriella halkyardi Cushman - Sztràkos (In: Mathelin & Sztràkos), p. 39, pl. 24, fig. 9 Locality: Vr-004, Vr-005, Vr-005A, Vr-010, Zv-004, Zv-006, Zv-008, Zv-009, Zv-010
Martinottiella sp.
Pl. 2, Fig. 5
Locality: Vr-004
Family: Textulariidae Ehrenberg, 1838
Semivulvulina pectinata (Hantken), 1875
Pl. 1, Fig. 5
1875 Vulvulina pectinata - Hantken, p. 68, pl. 7, fig. 10
1975 Vulvulina aff. pectinata Hantken - Braga et al. p. 102. pl. 4, fig. 3
1985 Spiroplectammina aff. pectinata (Hantken) - Grünig, p. 256, pl. 2, figs. 5-8
1993 “Semivulvulina”   cf.   pectinata   (Reuss)   -Sztràkos (In: Mathelin & Sztràkos), p. 40, pl. 2, fig. 11 Locality: Vr-005A, Vr-009, Vr-010, Vr-011, Vr-012, Vr-014, Zv-005, Zv-006, Zv-008, Zv-010
Textularia adalta Cushman, 1926
Pl. 2, Fig. 6
1926 Textularia adalta - Cushman, p. 29, pl. 4, fig. 2 (Fide Ellis & Messina 1940)
1950 Textularia adalta Cushman - Ruiz de Gaona & Colom, p. 416, figs. 5/22-26
1985 Textularia adalta Cushman - Grünig, p. 257, pl. 3, figs. 16-17
1993  Textularia adalta Cushman - Sztràkos (In:
Mathelin & Sztràkos), p. 73, pl. 3, fig. 9 Locality: Vr-005, Vr-005A, Vr-009, Zv-010
Karrerotextularia olianensis (Ruiz de Gaona & Colom), 1950
Pl. 2, Figs. 7-8
1950 Siphotextularia olianensis - Ruiz de Gaona & Colom, p. 413, figs. 16/1-9
1985 Siphotextularia concava (Karrer) - Grünig, p. 257, pl. 2, figs. 18-19
1991 Siphotextularia olianensis Ruiz de Gaona & Colom - Barbin & Keller-Grünig, p. 241, pl. 1, figs. 3-4
1993 Karrerotextularia olianensis (Ruiz de Gaona & Colom) - Sztràkos (In: Mathelin & Sztràkos), p. 40, pl. 3, figs. 17-18; pl. 25, figs. 3-4
Locality: Vr-009
Family: Pseudogaudryinidae Loeblich & Tappan, 1985
Pseudogaudryina textilaroides (Hantken), 1875
Pl. 2, Fig. 9
1875 Gaudryina textilaroides - Hantken, p. 15, pl. 1, fig. 6
Late Eocene benthic foraminiferal fauna from clastic sequence of the Socka - Dobrna area...           19
PLATE 2
1-2 Dorothia fallax Hagn. 1 Zv/II-011, x40. 2 Zv/II-011, x40. 3-4 Karreriella halkyardi Cushman. 3 Vr-010, x80. 4 Zv/II-006, x60. 5 Martinottiella sp., Vr-004, x80. 6 Textularia adalta Cushman, Vr-009, x40. 7-8 Karrerotextularia olianensis (Ruiz de Gaona & Colom). 7 Vr-009, x120. 8 Vr-009, x120. 9 Pseudogaudryina textilaroides (Hantken), Vr-009, x28. 10-11 Cylindroclavulina colomi Hagn. 10 Vr-009, x40. 11 Vr-012, x40. 12-13 Cylindroclavulina rudislosta (Hantken). 12 Vr-009, x28. 13 Vr-009, x28. 14 Tritaxilina pupa (Gümbel), Vr-005A, x60.
20
Franc Cimerman, Bogomir Jelen & Dragomir Skaberne
1993 Pseudogaudryina ? textilaroides Hantken -Sztràkos (In: Mathelin & Sztràkos), p. 41, pl. 4, fig. 5
Locality: Vr-009
Family: Valvulinidae Berthelin, 1880
Cylindroclavulina colomi Hagn, 1956
Pl. 2, Figs. 10-11
1956 Cylindroclavulina colomi - Hagn, p. 123, pl. 10, figs. 6-7; pl. 18, figs. 8. 9
1975 Cylindroclavulina colomi Hagn - Braga & al. p. 103, pl. 4, fig. 6
1985 Cylindroclavulina colomi Hagn - Grünig, p. 258, pl. 4, fig. 5
1993 Cylindroclavulina colomi Hagn - Sztràkos (In: Mathelin & Sztràkos), p. 73, pl. 4, fig. 2 Locality: Vr-004, Vr-005A, Vr-009, Vr-012, Vr-014, Zv-006, Zv-007
Cylindroclavulina rudislosta (Hantken), 1889
Pl. 2, Figs. 12-13
1889 Clavulina rudislosta - Hantken, p. 383
1875 Clavulina cylindrica Hantken (non d’Orbigny 1825) - Hantken, p. 18, pl. 1, fig. 8
1956 Cylindroclavulina rudislosta (Hantken) -Hagn, p. 122, pl. 10, fig. 5
1991 Cylindroclavulina rudislosta (Hantken) -Barbin & Keller-Grünig, p. 240
1993 Cylindroclavulina   rudislosta   (Hantken)  -Sztràkos (In: Mathelin & Sztràkos), p. 73, pl. 4, fig. 4 Locality: Vr-004, Vr-005, Vr-005A, Vr-009, Vr-012, Vr-014, Zv-004, Zv-005, Zv-007
Tritaxilina pupa (Gümbel), 1868
Pl. 2, Fig. 14
1868 Gaudryina pupa - Gümbel, p. 602, pl. 1, fig. 3
1950 Tritaxilina pupa (Gümbel) - Ruiz de Gaona & Colom, p. 421, fig. 14/23
1975 Tritaxilina pupa (Gümbel) - Braga et al., p. 93, pl. 1, figs. 24, 27
1987aTritaxilina pupa (Gümbel) - Sztràkos, p. 35,
pl. 3, figs. 11-12 Locality: Vr-005A, Vr-011, Vr-014, Zv-008, Zv-009, Zv-010, Zv-011
Family: CornusPiridae Schultze, 1854
Cornuspira angigyra (Reuss), 1850
Pl. 3, Fig. 1
1850 Operculina angigyra - Reuss, p. 370, pl. 46, fig. 19
1993  Cornuspira angigyra (Reuss) - Sztràkos (In: Mathelin & Sztràkos), p. 74, pl. 4, fig. 13 Locality: Zv-006
Family: Spiroloculinidae Wiesner, 1920
Spiroloculina dorsata Reuss, 1866
1866 Spiroloculina dorsata - Reuss, p. 123
1855  Spiroloculina limbata - Bornemann, p. 348, pl. 19, fig. 1
1961 Spiroloculina    canaliculata    d’Orbigny    -
Kaasschieter, p. 154, pl. 3, figs. 20-23 Locality: Vr-004, Vr-005A
Spiroloculina sp. 1
Pl. 3, Figs. 2-3
Locality: Vr-004, Vr-005, Vr-005A, Zv-005, Zv-010, Zv-011
Spiroloculina sp. 2
Pl. 3, Fig. 4
Locality: Vr-005, Vr-011, Zv-006, Zv-007,
Zv-008, Zv-009, Zv-010, Zv-011
Family: Hauerinidae Schwager, 1876
Quinqueloculina depressa d’Orbigny, 1826
Pl. 3, Fig. 5
1826 Quinqueloculina depressa - d’Orbigny, p. 302, no. 38
1878  Triloculina depressa d’Orbigny - Terquem, pl. 8, figs. 1-11
1904 Quinqueloculina    depressa    d’Orbigny    -Fornasini, p. 68, pl. 4, fig. 8
1993 Quinqueloculina    depressa    d’Orbigny    -Sztràkos (In: Mathelin & Sztràkos), p. 74, pl. 5, fig. 4 Locality: Vr-005A, Zv-006, Zv-009, Zv-010, Zv-011
Quinqueloculina impressa Reuss, 1851
Pl. 3, Fig. 6
1851 Quinqueloculina impressa - Reuss, p. 87, pl. 7, fig. 59
1993 Quinqueloculina impressa Reuss - Sztràkos (In: Mathelin & Sztràkos), p. 74, pl. 5, fig. 6 Locality: Vr-004, Vr-005, Vr-005A, Vr-009, Vr-010, Vr-011, Vr-011A, Vr-014, Zv-004, Zv-005, Zv-006, Zv-007, Zv-008, Zv-009, Zv-010, Zv-011
Quinqueloculina sp. 1
Pl. 3, Figs. 7-8
Locality: Vr-004, Vr-011, Vr-011A, Zv-006, Zv-007, Zv-009, Zv-010, Zv-011
Late Eocene benthic foraminiferal fauna from clastic sequence of the Socka - Dobrna area...           21
PLATE 3
1 Cornuspira angigyra (Reuss), Zv/II-006, x40. 2-3 Spiroloculina sp. 1. 2 Zv/II-011, x40. 3 Zv/II-011, x40. 4 Spiroloculina sp. 2. Vr-010, x60 5 Quinqueloculina depressa d’Orbigny, Zv/II-010, x60. 6 Quinqueloculina impressa Reuss, Vr-009, x60. 7-8 Quinqueloculina sp. 1. 7 Zv/II-009, x60. 8 Zv/II-009, x60. 9 Pyrgo simplex (d’Orbigny), Zv/II-006, x80. 10-11 Pyrgo sp. 1. 10 Zv/II-005, x80. 11 Zv/II-006, x80. 12-13 Pyrgo sp. 2. 12 Zv/II-009, x80. 13 Pyrgo sp. 2, Zv/II-009, x80. 14-16 Nov. gen. sp. 1. 14 Zv/II-005, x120. 15 Zv/ II-010, x60. 16 Zv/II-007, x80.
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Franc Cimerman, Bogomir Jelen & Dragomir Skaberne
Quinqueloculina sp. 2
Locality: Vr-011A, Zv-006, Zv-007, Zv-009, Zv-010, Zv-011
Quinqueloculina sp. 3
Pl. 4, Fig. 5 Locality: Vr-003
Pyrgo simplex (d’Orbigny), 1846
Pl. 3, Fig. 9
1846 Biloculina simplex - d’Orbigny, p. 264, pl. 15, figs. 25-27
1975 Pyrgo simplex (d’Orbigny) - Braga et al., p. 104
1985 Pyrgo simplex (d’Orbigny) - Grünig, p. 260 fig. 9
1993  Pyrgo simplex (d’Orbigny) – Sztràkos (In:
Mathelin & Sztràkos), p. 74, pl. 5, fig. 9 Locality: Vr-012, Zv-005, Zv-006, Zv-008
Pyrgo subsphaerica (d’Orbigny), 1839
Pl. 4, Fig.6
1839 Biloculina subsphaerica - d’Orbigny, p. 162, pl. 8, figs. 25-27
1949 Pyrgo subsphaerica (d’Orbigny) - Cuvillier & Szakall, p. 45, pl. 20, fig. 2
1985  Pyrgo subsphaerica (d’Orbigny) - Grünig, p.
260 Locality: Vr-003
Pyrgo sp. 1
Pl. 3, Figs. 10-11
Locality: Zv-005, Zv-006
Pyrgo sp. 2
Pl. 3, Figs. 12-13
Locality: Zv-009, Zv-010, Zv-011
Nov. gen. sp. 1
Pl. 3, Figs. 14-16
Locality: Vr-004, Vr-005A, Zv-005, Zv-006, Zv-007, Zv-008, Zv-010, Zv-011
Sigmoilopsis bartoniensis (Ruiz de Gaona & Colom), 1950
Pl. 4, Figs. 1-2
1951 Sigmoilina bartoniensis - Ruiz de Gaona & Colom, p. 410, figs. 17/1-16
1985 Sigmoilopsis sp. - Grünig, p. 260, pl. 4, figs. 14-16
1993 Sigmoilopsis bartoniensis Ruiz de Gaona & Colom - Sztràkos (In: Mathelin & Sztràkos), p. 74, pl. 5, fig. 12
Locality: Vr-011, Vr-011A, Zv-005, Zv-
006, Zv-007, Zv-008, Zv-009, Zv-010, Zv-011
Family: Alveolinidae Ehrenberg, 1839
Borelis vonderschmitti (Schweighauser), 1951
Pl. 4, Figs. 3-4
1951 Neoalveolina             vonderschmitti             -
Schweighauser, p. 468, figs. 1-4
1979 Borelis aff. vonderschmitti (Schweighauser) - Drobne K. et al., p. 160, pl. 4, figs. 1-2
1990   Borelis vonderschmitti (Schweighauser) -Pavlovec et al., p. 442, pl.1, figs. 1-3
Locality: Vr-005, Vr-005A
Family: Nodosariidae Ehrenberg, 1838
Nodosaria latejugata Gümbel, 1868
Pl. 4, Fig. 11
1868 Nodosaria latejugata - Gümbel, p. 41, pl. 1, fig. 32
1956 Nodosaria latejugata Gümbel - Hagn, p. 137, pl. 12, fig. 11; pl. 13, fig. 1
1987aNodosaria latejugata Gümbel - Sztràkos, p 33
1991   Nodosaria latejugata Gümbel - Barbin & Keller-Grünig, p. 240
Locality: Vr-004, Vr-005 Vr-010, Vr-011,
Vr-012, Zv-008
Pyramidulina bactridium (Reuss), 1866
Pl. 4, Figs. 8-9
1866 Nodosaria bactridium - Reuss, p. 130, pl. 1, figs. 24-25
1993 Pyramidulina bactridium (Reuss) - Sztràkos (In: Mathelin & Sztràkos). p. 75, pl. 7, figs. 15-16
Locality: Vr-005A
Family: Vaginulinidae Reuss, 1860
Lenticulina arcuatostriata (Hantken), 1868
Pl. 4, Fig. 13
1868 Cristellaria (Robulina) arcuato striata -Hantken, p. 93, pl. 2, fig. 30
1950 Robulus arcuatostriatus (Hantken) - Ruiz de Gaona & Colom, p. 402, figs. 6/1-9
1956 Robulus arcuato-striatus (Hantken) - Hagn, p. 127, pl. 11, fig. 4
1975 Lenticulina arcuatostriata (Hantken) -Braga et al., p. 104
1985 Lenticulina   arcuato-striata   (Hantken)   -
Grünig, p. 261, pl. 4, fig. 19 Locality: Vr-005A, Vr-010, Vr-011, Zv-005, Zv-010
Late Eocene benthic foraminiferal fauna from clastic sequence of the Socka - Dobrna area...           23
PLATE 4
1-2 Sigmoilopsis bartoniensis (Ruiz de Gaona & Colom). 1 Zv/II-010, x40. 2 Vr-011A, x40. 3-4 Borelis vonderschmitti (Schweighauser). 3 Vr-005A, x160. 4 Vr-005A, x120. 5 Quinqueloculina sp. 3, Vr/II-003, x40 6 Pyrgo subsphaerica (d’Orbigny), Vr/II-003, x80. 7-8 Rectobolivina zsigmondyi (Hantken). 7 Zv/II-009, x60. 8 Zv/II-009, x40. 9-10 Pyramidulina bactridium (Reuss). 9 Vr-005A, x28. 10 Vr-005A, x40. 11 Nodosaria latejugata Gümbel, Zv/II-006, x28. 12 Lenticulina budensis (Hantken), Vr-012, x40. 13 Lenticulina arcuatostriata (Hantken), Vr-011, x28. 14 Lenticulina limbosa (Reuss), Zv/II-008, x28.
24
Franc Cimerman, Bogomir Jelen & Dragomir Skaberne
Lenticulina budensis (Hantken), 1875
Pl. 4, Fig. 12
1875 Robulina budensis - Hantken, p. 58, pl. 7, fig. 1
1950  Robulus   budensis   (Hantken)   -   Ruiz   de
Gaona & Colom, p. 403, figs. 7/7-8 Locality: Vr-010, Vr-012
Lenticulina limbosa (Reuss), 1863
Pl. 4, Fig. 14
1863 Robulina limbosa - Reuss, p. 55, pl. 6, fig. 69a-b
1875 Robulina limbosa Reuss - Hantken, p. 57, pl. 6, fig. 11
1950 Rubulus limbosus (Reuss) - Ruiz de Gaona & Colom, p. 405, fig. 5/1-14
1956 Robulus limbosus (Reuss) - Hagn, p. 127, pl. 11, fig. 2
1993  Lenticulina limbosa (Reuss) - Sztràkos (In:
Mathelin & Sztràkos), p. 75, pl. 8, fig. 6 Locality: Zv-008
Marginulinopsis behmi (Reuss), 1866
Pl. 5, Fig. 1
1866 Cristellaria behmi - Reuss, p. 22, pl. 2, fig. 37
1868 Cristellaria behmi Reuss - Gümbel, s. 55, pl. 1, fig. 61
1875 Marginulina behmi (Reuss) - Hantken, p. 48, pl. 5, figs. 1-2
1950 Marginulina behmi (Reuss) - Ruiz de Gaona & Colom, p. 384, fig. 11/18-26
1956 Marginulina behmi (Reuss) - Hagn, p. 131, pl. 11, fig. 11
1975 Marginulina behmi (Reuss) - Braga et al., p. 105, pl. 4, fig. 13
1985 Marginulina behmi (Reuss) - Grünig, p. 262, pl. 5, figs. 17-19
1993 Marginulina behmi (Reuss) - Sztràkos (In: Mathelin & Sztràkos), p. 43, pl. 26, figs. 14-16
Locality: Vr-004, Vr-005, Vr-009, Vr-010,
Vr-011, Vr-012, Vr-014, Zv-006, Zv-008,
Zv-009, Zv-010, Zv-011
Marginulinopsis porvaensis (Hantken), 1875
Pl. 5, Figs. 2-3
1875 Cristellaria porvaensis - Hantken, p. 50, pl. 14, fig. 1
1950 Marginulina porvaensis (Hatken) - Ruiz de Gaona & Colom, p. 387, fig. 11/1-14
1987aMarginulinopsis porvaensis (Hantken) -Sztràkos, p. 32, pl. 5, fig. 20
1993 Astacolus porvaensis (Hantken) - Sztràkos (In: Mathelin & Sztràkos), p. 76, pl. 26, fig. 13
Locality: Vr-010, Vr-011, Zv-005, Zv-007, Zv-008, Zv-009, Zv-010
Percultazonaria fragaria (Gümbel), 1868
Pl. 5, figs. 4-5
1868 Marginulina fragaria - Gümbel, p. 57, pl. 1, figs. 58a-c
1950 Marginulina fragaria - Ruiz de Gaona & Colom, p. 386, figs. 11/15-17
1956 Marginulinopsis fragaria (Gümbel) - Hagn, p. 134, pl. 12, figs. 3, 8, 10
1975 Marginulinopsis fragaria (Gümbel) - Braga et al., p. 105
1985 Marginulinopsis fragaria (Gümbel) - Grünig, p. 161, pl. 5, figs. 24-28
1991 Marginulinopsis fragaria (Gümbel) - Barbin & Keller-Grünig, p. 240, pl. 2, figs. 6-12
1993 Percultazonaria fragaria (Gümbel) -Sztràkos (In: Mathelin & Sztràkos), p. 76, pl. 9, figs. 1-2
Locality: Vr-014, Zv-004, Zv-006, Zv-007,
Zv-009
Saracenaria hantkeni Cushman, 1933
Pl. 5, Fig. 6
1933 Saracenaria arcuata d’Orbigny var. hantkeni - Cushman, p. 4, pl. 1, figs. 11-12 (Fide Ellis & Messina 1940)
1956 Saracenaria hantkeni Cushman - Hagn, p. 138, pl. 13, fig. 4
1993  Saracenaria hantkeni Cushman - Sztràkos (In: Mathelin & Sztràkos), p. 76, pl. 9, fig. 6 Locality: Vr-004, Vr-009, Vr-010, Zv-005, Zv-006, Zv-007, Zv-010, Zv-011
Palmula budensis (Hantken), 1875
Pl. 5, Fig. 7
1875 Flabellina budensis - Hantken, p. 44, pl. 4, fig. 17
1984   Frondicularia budensis (Hantken) - Grünig & Herb, p. 263
1985  Frondicularia budensis (Hantken) - Grünig, p. 261, pl. 4, fig. 21
1993  Palmula budensis (Hantken) - Sztràkos (In: Mathelin & Sztràkos), p. 76, pl. 9, fig. 10 Locality: Vr-011, Zv-005, Zv-006, Zv-007, Zv-008
Palmula tenuissima (Hantken), 1875
Pl. 5, Figs. 8-9
1875 Frondicularia tenuissima - Hantken, p. 43, pl. 13, fig. 11
1950 Frondicularia tenuissima Hantken - Ruiz de Gaona & Colom, p. 377
Late Eocene benthic foraminiferal fauna from clastic sequence of the Socka - Dobrna area...           25
PLATE 5
1 Marginulinopsis behmi (Reuss), Zv/II-006, x60. 2-3 Marginulinopsis porvaensis (Hantken). 2 Zv/II-006, x28. 3 Zv/II-010, x40. 4-5 Percultazonaria fragaria (Gümbel). 4 Zv/II-006, x40. 5 Vr-012, x28. 6 Saracenaria hantkeni (Cushman), Vr-010, x40. 7 Palmula budensis (Hantken), Zv/II-006, x80. 8-9 Palmula tenuissima (Hantken). 8 Zv/II-008, x80. 9 Zv/II-005, x60. 10 Marginulina globosa (Halkyard), Zv/II-006, x120. 11-12 Reticulopalmula arborescens (Halkyard). 11 Vr-005, x40. 12 Vr-005, x40. 13-14 Vaginulinopsis cumulicostata (Gümbel). 13 Vr-004, x28. 14 Vr-004, x28.
26
Franc Cimerman, Bogomir Jelen & Dragomir Skaberne
1993 Frondovaginulina tenuissima (Hantken) -Sztràkos (In: Mathelin & Sztràkos), p. 76, pl. 9, fig. 8
Locality: Vr-005A, Zv-008
Reticulopalmula arborescens (Halkyard), 1919
Pl. 5, Figs. 11-12
1919 Frondicularia arborescens - Halkyard, p. 88, pl. 5, figs. 8-9
1981 Coleites arborescens (Halkyard) - Hagn et al., p. 91
1985 Coleites arborescens (Halkyard) - Grünig, p. 275, pl. 6, figs. 14-15
1993 Reticulopalmula arborescens (Halkyard) -Sztràkos (In: Mathelin & Sztràkos), p. 43, pl. 26, figs. 8-9
Locality: Vr-005, Vr-009, Vr-010, Vr-011,
Vr-012, Zv-005, Zv-006, Zv-008 Zv-010,
Zv-011
Marginulina globosa Halkyard, 1919
Pl. 5, Fig. 10
1919 Marginulina pyramidale Karrer var. globosa - Halkyard, p. 85, pl. 5, fig. 2
1993 Marginulina globosa Halkyard - Sztràkos (In: Mathelin & Sztràkos), p. 44, pl. 23, fig. 15; pl. 48, figs. 20-22
Locality: Vr-011, Zv-006
Vagunulinopsis cumulicostata (Gümbel), 1868
Pl. 5, Figs. 13-14
1868 Cristellaria cumulicostata - Gümbel, p. 60, pl. 1, fig. 67
1956 Vaginulinopsis cumulicostata (Gümbel) -Hagn, p. 135, pl. 13, fig. 3
1975 Vaginulinopsis cumulicostata (Gümbel) -Braga et al. p. 105, pl. 4, fig. 8
1985 Vaginulinopsis cumulicostata (Gümbel) -Grünig, p. 263, pl. 6, fig. 6
1993 Vaginulinopsis cumulicostatus (Gümbel) -Sztràkos (In: Mathelin & Sztràkos), p. 76, pl. 10, fig. 12
Locality: Vr-004, Vr-005, Vr-010, Vr-012,
Zv-005
Spirolingulina acutimargo (Halkyard), 1919
Pl. 6, Fig. 1
1919 Lingulinopsis acutimargo - Halkyard, p. 81, pl. 5, fig. 1
1949   Lingulina acutimargo (Halkyard) - Cuvillier & Szakall, p. 81, pl. 29, fig. 11
1950  Lingulina acutimargo (Halkyard) - Ruiz de Gaona & Colom, p. 384, fig. 11/31-3
1993 Spirolingulina acutimargo (Halkyard) -Sztràkos (In: Mathelin & Sztràkos), p. 44, pl. 27, fig. 2
Locality: Vr-004, Zv-009
Family: Ceratobuliminidae Cushman, 1927
Ceratocancris crassa (Halkyard), 1919
Pl. 6, Figs. 2-4
1919 Pulvinulina haueri d’Orbigny var. crassa -Halkyard, p. 125, pl. 7, fig. 6
1957 Ceretobulimina haueri d’Orbigny var. crassa (Halkyard) - Sacal & Debourle, p. 46, pl. 20, fig. 1
1993 Ceratocancris crassa (Halkyard) - Sztràkos (In: Mathelin & Sztràkos), p. 45, pl. 11, fig. 17
Locality: Vr-014
Lamarckina cristellaroides (Terquem), 1882
Pl. 6, Fig. 7
1882 Rotalina cristellaroides - Terquem, p. 57, pl. 3, fig. 15a-c
1961 Lamarckina cristellaroides (Terquem) -Kaasschieter, p. 231, pl. 15, figs. 3-4
1970 Lamarckina cristellaroides (Terquem) - Le Calvez, p. 203, pl. 37, fig. 4
1993 Lamarckina cristellaroides (Terquem) -Sztràkos (In: Mathelin & Sztràkos), p. 77, pl. 11, fig. 19; pl. 29, fig. 6
Locality: Vr-005A
Family: Epistominidae Wedekind, 1937
Hoeglundina eocenica (Cushman & Hanna), 1927
Pl. 6, Figs. 5-6
1927 Epistomina eocenica - Cushman & Hanna, p. 53, pl. 5, figs. 4-5
1975 Hoeglundina eocenica (Cushman & Hanna)
-  Braga et al., p. 110
1981 Hoeglundina eocenica (Cushman & Hanna)
-  Lindenberg et al., p. 148
1985 Hoeglundina eocenica (Cushman & Hanna)
-  Grünig, p. 277, pl. 11, figs. 22-24
1987aHoeglundina eocenica (Cushman & Hanna)
-  Sztràkos, p. 31, pl. 15, fig. 9 Locality: Vr-005, Vr-010, Vr-011A, Vr-012, Zv-010
Family: Bolivinidae Glaesner, 1937
Bolivina nobilis Hantken, 1875
Pl. 6, Figs. 9-10
1875 Bolivina nobilis - Hantken, p. 65, pl. 15, fig. 4a-b
Late Eocene benthic foraminiferal fauna from clastic sequence of the Socka - Dobrna area...           27
PLATE 6
1 Spirolingulina acutimargo (Halkyard), Vr-004, x40. 2-4 Ceratocancris crassa (Halkyard). 2 spiral view, Vr-014, x40. 3 edge view, Vr-014, x40. 4 umbilical view, Vr-014, x40. 5-6 Hoeglundina eocenica (Cushman & Hanna). 5 spiral view, Vr-010, x28. 6 umbilical view, Vr-010, x28. 7 Lamarckina cristellaroides (Terquem), Vr-005A. 8-9 Bolivina nobilis Hantken. 8 Vr-010, x80. 9 Vr-010, x80. 10 Bolivina semistriata Hantken, Vr-005A, x40. 11 Aragonia janoscheki Gohrbandt, Vr-010, x120. 12-13 Globocassidulina globosa (Hantken). 12 Vr.011, x80. 13 Vr-009, x80. 14-17 Sporobulimina eocaena Bykova. 14 side view, Vr-008, x120. 15 apertural view, Vr-005A, x120. 16 apical view, Zv/II-008, x120. 17 apertural view, Zv/II-008, x120.
28
Franc Cimerman, Bogomir Jelen & Dragomir Skaberne
1950  Bolivina nobilis Hantken - Ruiz de Gaona & Colom, p. 364
1956  Bolivina nobilis Hantken - Hagn, p. 147
1975  Bolivina nobilis Hantken - Braga et al., p. 106, pl. 5, figs. 1-2
1985  Bolivina nobilis Hantken - Grünig, p. 265, pl. 5, figs. 12-14
1991  Bolivina nobilis Hantken - Barbin & Keller-Grünig, p. 240
1993  Bolivina  nobilis  Hantken  -  Sztràkos  (In: Mathelin & Sztràkos), p. 78, pl. 32, fig. 9 Locality: Vr-004, Vr-005, Vr-009, Zv-007, Zv-008, Zv-009
Bolivina semistriata Hantken, 1875
Pl. 6, Fig. 10
1875  Bolivina semistriata - Hantken, p. 65, pl. 7, fig. 13
1956  Bolivina semistriata Hantken - Hagn, p. 147
1993  Bolivina semistriata Hantken - Sztràkos (In: Mathelin & Sztràkos), p. 78, pl. 32, fig. 10 Locality: Vr-011, Vr-012, Zv-005, Zv-006, Zv-010
Bolivina pl. sp.
Locality: Vr-005, Vr-009, Vr-011, Vr-012, Zv-005, Zv-007, Zv-008, Zv-009, Zv-010, Zv-011
Family: Loxostomatidae Loeblich & Tap-pan, 1962
Aragonia janoscheki Gohrbandt, 1962
Pl. 6, Fig. 11
1962  Aragonia janoscheki - Gohrbandt, p. 81, pl. 3, figs. 5a-b
1975  Aragonia janoscheki Gohrbandt - Braga et
al., p. 106, pl. 5, fig. 7 Locality: Vr-010, Vr-012
Family: Cassidulinidae d’Orbigny, 1839
Globocassidulina globosa (Hantken), 1875
Pl. 6, Figs. 12-13
1875  Cassidulina globosa - Hantken, p. 64, pl. 16, figs. 2a-b
1956  Cassidulina globosa Hantken - Hagn, p. 167, pl. 14, figs. 9-10
1975 Globocassidulina globosa (Hantken) - Braga et al., p. 108, pl. 6, fig. 4
1985 Globocassidulina     globosa     (Hantken)    -Grünig, p. 273, pl. 10, fig. 3
1987aGlobocassidulina     globosa      Hantken     -Sztràkos, p. 31, pl. 11, fig. 22
1993 Globocassidulina     globosa     (Hantken)    -Sztràkos (In: Mathelin & Sztràkos), p. 79
Locality: Vr-004, Vr-005, Vr-005A, Vr-009, Vr-011, Vr-011A
Family: Turrilinidae Cushman, 1927
Sporobulimina eocaena Bykova, 1959
Pl. 6, Figs. 14-17
1959 Sporobulimina eocaena - Bykova, p. 76, pl. 12, fig. 6a-b (Fide Ellis & Messina 1940)
1993 Sporobulimina eocaena Bykova - Sztràkos (In: Mathelin & Sztràkos), p. 46, pl. 33, figs. 11-12
Locality: Vr-005A, Zv/II-008
Family: Siphogenerinoididae Saidova, 1981
Rectobolivina zsigmondyi (Hantken), 1868
Pl. 4, Figs. 7-8
1868 Nodosaria (Dentalina) zsigmondyi -Hantken, p. 87, pl. 1, fig. 12
1982 Rectobolivina zsigmondyi (Hantken) -Sztràkos, pl. 15, fig. 14
1987 Rectobolivina    zsigmondyi    (Hantken)    -
Reiser, p. 91, pl. 9, figs. 13, 19 Locality: Zv-005, Zv-008, Zv-009, Zv-010
Sagrinopsis aspera (Terquem), 1882
Pl. 7, Figs. 1-3
1882 Textilaria aspera - Terquem, p. 147, pl. 15, fig. 14 (Fide Ellis & Messina 1940)
1919 Siphogenerina hexagona Halkyard -Halkyard, p. 41, pl. 6, fig. 5a-b
1975 Bifarina aff. selseyensis (Heron-Allen & Earland) - Braga et al., p. 108, pl. 5, fig. 9
1985 Rectobolivina sp. - Grünig, p. 266, pl. 8, fig. 22
1993  Sagrinopsis aspera (Terquem) - Sztràkos (In: Mathelin & Sztràkos), p. 47, pl. 33, fig. 15 Locality: Vr-005, Vr-005A
Tubulogenerina tubulifera (Parker & Jones), 1863
Pl. 7, Figs. 4-5
1863 Textularia tubulifera - Parker & Jones, p. 94, fig. 2 (Fide Ellis & Messina 1940)
1837 Tubulogenerina tubulifera (Parker & Jones)
-  Cushman, p. 215, pl. 24, figs. 14-16
1970 Tubulogenerina tubulifera (Parker & Jones)
-  Le Calvez, p. 122, pl. 26, fig. 7
1987aTubulogenerina tubulifera (Parker & Jones)
- Sztràkos, p. 35, pl. 7, fig. 20; pl. 18, fig. 2
1991 Tubulogenerina tubulifera (Parker & Jones)
-  Gibson et al., p. 307, pl. 1, figs. 1-11; pl. 3, figs. 6-8
Locality: Vr-005, Vr-005A
Late Eocene benthic foraminiferal fauna from clastic sequence of the Socka - Dobrna area...           29
PLATE 7
1-3 Sagrinopsis aspera (Terquem). 1 Vr-005, x80. 2 Vr-005, x120. 3 Vr-005A, x120. 4-5 Tubulogenerina cf. tubulifera (Parker & Jones). 4 Vr-005A, x120. 5 Vr-005, x120. 6 Bulimina ovata d’Orbigny, Vr-010, x60. 7 Bulimina subtruncana Hagn, Vr-010, x160. 8 Bulimina truncana Gümbel, Vr-005, x80. 9-10 Uvigerina gracilis Reuss. 9 Vr-005, x160. 10 Vr-005, x160. 11 Uvigerina chirana Cushman & Stone, Vr-005, x120. 12 Uvigerina eocaena Gümbel, Vr-010, x80. 13 Angulogerina muralis (Terquem), Zv/II-008, x160. 14 Angulogerina pulchella Cushman & Edwards, Vr-005, x160. 15 Angulogerina globosa (Stoltz), Vr-014, x120. 16 Koleshnikovella elongata (Halkyard), Zv/II-010, x120.
30
Franc Cimerman, Bogomir Jelen & Dragomir Skaberne
Family: Buliminidae Jones, 1875
Bulimina ovata d’Orbigny, 1846
Pl. 7, Fig. 6
1846  Bulimina ovata - d’Orbigny, p. 185, pl. 11, figs. 13-14
1961  Bulimina ovata d’Orbigny - Kaasschieter, p. 191, pl. 9, fig. 6
1985  Bulimina ovata d’Orbigny - Grünig, p. 266,
pl. 6, fig. 22 Locality: Vr-010
Bulimina subtruncana Hagn, 1956
Pl. 7, Fig. 7
1954  Bulimina subtruncana - Hagn, p. 17, pl. 3, fig. 19; pl. 4, fig. 9
1985 Bulimina subtruncana Hagn - Grünig, p.
266, pl. 6, fig. 23 Locality: Vr-010
Bulimina truncana Gümbel, 1868
Pl. 7, Fig. 8
1868  Bulimina truncana - Gümbel, p. 66, pl. 2, fig. 77a-b
1975  Bulimina truncana Gümbel - Hantken, p. 61, pl. 7, fig. 5
1975  Bulimina truncana Gümbel - Braga et al., p. 106, pl. 4, figs. 15-16
1985  Bulimina truncana Gümbel - Grünig, p. 267,
pl. 7, figs. 26-27 Locality: Vr-005, Vr-005A, Vr-010, Vr-011A, Zv-009
Family: Uvigerinidae Haeckel, 1894
Uvigerina chirana Cushman & Stone, 1947
Pl. 7, Fig. 11
1947  Uvigerina chirana - Cushman & Stone, p. 17, pl. 2, fig. 25 (Fide Ellis & Messina 1940)
1977  Uvigerina chirana Cushman & Stone - Braga et al., p. 95, pl. 5, fig. 10
1985  Uvigerina   chirana   Cushman   &   Stone   -
Grünig, p. 267, pl. 7, figs. 3-5 Locality: Vr-005, Zv-004, Zv-005
Uvigerina eocaena Gümbel, 1868
Pl. 7, Fig. 12
1868  Uvigerina eocaena - Gümbel, p. 645, pl. 2, fig. 78
1975  Uvigerina eocaena Gümbel - Braga et al., p. 107, pl. 5, fig. 17
1985  Uvigerina eocaena Gümbel - Grünig, p. 267, pl. 7, figs. 8-10
1993  Uvigerina eocaena Gümbel - Sztràkos (In: Mathelin & Sztràkos), p. 79, pl. 34, figs. 9-10 Locality: Vr-004, Vr-010, Vr-011 Zv-005, Zv-006, Zv-008, Zv-010, Zv-011
Uvigerina gracilis Reuss, 1851
Pl. 7, Figs. 9-10
1851 Uvigerina gracilis - Reuss, p. 77, pl. 5, fig. 39
1993  Uvigerina   gracilis   Reuss   -   Sztràkos   (In: Mathelin & Sztràkos), p. 79, pl. 34, fig. 13 Locality: Vr-005
Angulogerina globosa (Stoltz), 1925
Pl. 7, Fig. 15
1925 Uvigerina tenuistriata var. globosa – Stoltz, p. 130 (Fide Ellis & Messina 1940)
1993  Angulogerina globosa (Stoltz) - Sztràkos (In: Mathelin & Sztràkos), p. 48, pl. 34, fig. 20 Locality: Vr-014
Angulogerina muralis (Terquem), 1882
Pl. 7, Fig. 13
1882 Uvigerina muralis - Terquem, p. 119, pl. 12, figs. 26-29 (Fide Ellis & Messina 1940)
1919 Tritaxia dehiscens - Halkyard, p. 44, pl. 3, fig. 8
1985 Trifarina muralis (Terquem) - Grünig, p. 268, pl. 7, fig. 26
1993 Angulogerina muralis (Terquem) - Sztràkos (In: Mathelin & Sztràkos), p. 79, pl. 43, fig. 18
Locality: Zv/II-008
Angulogerina pulchella Cushman & Edwards, 1937
Pl. 7, Fig. 14
1937 Angulogerina pulchella - Cushman & Edwards, p. 61, pl. 8, fig. 19 (Fide Ellis & Messina 1940)
1993 Angulogerina pulchella Cushman & Edwards - Sztràkos (In: Mathelin & Sztràkos), p. 48, pl. 34, fig. 17
Locality: Vr-005
Trifarina sp. 1
Locality: Vr-012
Koleshnikovella elongata (Halkyard), 1919
Pl. 7, Fig. 16
1919 Tritaxia elongata - Halkyard, p. 45, pl. 3, fig. 9
1987aUvigerinella elongata (Halkyard) - Sztràkos, p. 36, pl. 18, fig. 5
1993 Koleshnikovella elongata (Halkyard) -Sztràkos (In: Mathelin & Sztràkos). p. 48, pl. 34, fig. 16
Late Eocene benthic foraminiferal fauna from clastic sequence of the Socka - Dobrna area...           31
PLATE 8
1-2 Reussella oberburgensis (Reuss). 1 Vr-005A, x120. 2 Vr-005A, x120. 3-4 Siphonodosaria crassisepta (Halkyard). 4 Vr-010, x40. 4 Vr-010, x40. 5-6 Nodogenerina atlantisae Cushman. 5 Zv/II-006, x80. 6 Zv/ II-008, x80. 7-8 Fursenkoina halkyardi (Cushman). 7 Vr-012, x60. 8 Zv/II-008, x60. 9-10 Sigmavirgulina tortuosa (Brady). 9 Vr-005, x80. 10 Vr-010, x120. 11-12 Neoeponides schreibersii (d’Orbigny). 11 Zv/II-005, x40. 12, Zv/II-005, x40. 13-14 Eponides ouachitaensis Howe & Wallace. 13 Vr-011A, x60. 14 Vr-011A, x60. 15 Schlosserina asterites (Gümbel), Vr-014, x28. 16-17 Neoconorbina sp. 16 spiral view, Vr-005A, x80. 17 umbilical view Vr-005A, x80.
32
Franc Cimerman, Bogomir Jelen & Dragomir Skaberne
Locality: Zv-010
Family: Reussellidae Cushman, 1933
Reussella oberburgensis (Reuss), 1864
Pl. 8, Figs. 1-2
1864 Verneuilina oberburgensis Frey. in litt. -Reuss, p. 6, pl. 1, fig. 2
1962 Reussella oberburgensis (Freyer) -Gohrbandt, p. 82, pl. 3, fig. 6a-c
1985 Reussella oberburgensis (Freyer) - Grünig, p. 267
1993 Reussella oberburgensis (Freyer) - Sztràkos (In: Mathelin & Sztràkos), p. 79, pl. 34, fig. 26
Locality: Vr-004, Vr-005, Vr-005A, Vr-011
Family: Fursenkoinidae Loeblich & TaPpan, 1961
Fursenkoina halkyardi (Cushman), 1936
Pl. 8, Figs. 7-8
1936 Virgulina halkyardi - Cushman, p. 47, pl. 7, fig. 5
1937bVirgulina halkyardi Cushman - Cushman, p. 11, pl. 1, figs. 26-27
1985 Fursenkoina halkyardi (Cushman) - Grünig, p. 273, pl. 9, figs. 25-27
1993 Fursenkoina halkyardi (Cushman) -Sztràkos (In: Mathelin & Sztràkos), p. 49, pl. 35, fig. 6
Locality: Vr-005, Vr-012, Zv-007, Zv-009,
Zv-010, Zv-011
Sigmavirgulina tortuosa (Brady), 1881
Pl. 8, Figs. 9-10
1881 Bolivina tortuosa - Brady, p. 57 (Fide Ellis & Messina 1940)
1960 Sigmavirgulina tortuosa (Brady) - Barker,
pl. 52, figs. 31-32 Locality: Vr-005, Vr-005A, Vr-010, Vr-011, Vr-011A, Vr-012, Zv-005
Family: Stilostomellidae Finlay, 1947
Siphonodosaria crassisepta (Halkyard), 1919
Pl. 8, Figs. 3-4
1919 Nodosaria pauperata d’Orbigny var. crassisepta - Halkyard, p. 72, pl. 4, figs. 12-13
1941 Dentalina annulata (Reuss) - de Witt Puyt, p. 51, pl. 1, figs. 23-25
1957 Nodogenerina crassisepta (Halkyard) -Sacal & Debourle, p. 16, pl. 4, fig. 8
1993 Siphonodosaria   crassisepta   (Halkyard)   -Sztràkos (In: Mathelin & Sztràkos), p. 51, pl. 14, figs. 3-11 Locality: Vr-004, Vr-005A, Vr-009, Vr-010, Vr-014, Zv-005, Zv-006, Zv-007, Zv-008, Zv-009, Zv-010
Family: Eponididae Hofker, 1951
Eponides ouachitaensis Howe & Wallace, 1932
Pl. 8, Figs. 13-14
1932 Eponides ouachitaensis - Howe & Wallace, p. 69, pl. 13, fig. 8 (Fide Ellis & Messina 1940)
1950 Eponides ouachitaensis Howe & Wallace -Ruiz de Gaona & Colom, p. 378, fig. 13/36-44
Locality: Vr-004
“Eponides” dalmatinus (de Witt Puyt), 1941
Pl. 11, Figs. 14-15
1941 Eponides carolinensis Cushman var. dalmatina - de Witt Puyt, p. 66, pl. 1, figs. 52, 57; pl. 2, fig. 2
1950 Eponides carolinensis Cushman var. navarraensis - Ruiz de Gaona & Colom, p. 377, fig. 14/5-10
1956 Rotalia dalmatina (de Witt Puyt) - Hagn, p. 165, pl. 15, figs. 2-3
1993 Neoeponides navaransis (Ruiz de Gaona & Colom) - Sztràkos (In: Mathelin & Sztràkos), p. 52, figs. 6-7 Locality: Vr-005A, Vr-010, Vr-011, Zv/II-006, Zv/II-010
Family: Mississippinidae Saidova, 1981
Schlosserina asterites (Gümbel), 1868
Pl. 8, Fig. 15
1868 Rosalina asterites - Gümbel, p. 658, pl. 2, fig. 101a-c
1981 Schlosserina asterites (Gümbel) -Lindenberg et al., p. 148
1991 Schlosserina asterites (Gümbel) - Barbin & Keller-Grünig, p. 241, pl. 1, figs. 13-15
1981 Schlosserina asterites (Gümbel) - Hagn et al., p. 91
1983 Schlosserina asterites (Gümbel) - Setiawan, p. 134, pl. 15, fig. 1
1993 Schlosserina asterites Hagn - Sztràkos (In: Mathelin & Sztràkos), p. 80, pl. 15, fig. 2 Locality: Vr-014
Family: Discorbidae Ehrenberg, 1838
Neoeponides schreibersii (d’Orbigny), 1846
Late Eocene benthic foraminiferal fauna from clastic sequence of the Socka - Dobrna area...           33
PLATE 9
1-2 Cibicidoides ungerianus (d’Orbigny). 1 spiral view, Vr-010, x60. 2 spiral view, Zv/II-008, x60. 3 Laticarinina altocamerata (Heron-Allen & Earland), umbilical view, Zv/II-008, x80. 4 Planulina compressa (Hantken), Vr-005A, x80. 5-6 Planulina costata (Hantken). 5 spiral view, Vr-012, x40. 6 umbilical view, Vr-010, x40. 7-8 Lobatula lobatula (Walker & Jacob). 7 umbilical view, Vr-012, x40. 8 umbilical view, Vr-005, x28. 9-10 Cycloloculina eocenica (Terquem). 9 Vr-005A, x80. 10 Vr-005A, x64. 11 Planorbulina difformis Roemer, Vr-005A, x60. 12 Halkyardia minima (Liebus), Vr-011, x80. 13 Halkyardia sp., Vr-011A, x120. 14-15 Korobkovella grosserugosa (Gümbel). 14 Vr-010, x60. 15 Vr-010, x60. 16 Sphaerogypsina globula (Reuss), Vr-005A, x120.
34
Franc Cimerman, Bogomir Jelen & Dragomir Skaberne
Pl. 8, Figs. 11-12
1846 Rotalina schreibersii - d’Orbigny, p. 154, pl. 8, figs. 4-6
1993 Neoeponides schreibersii (d’Orbigny) -Sztràkos (In: Mathelin & Sztràkos), p. 81, pl. 15, fig. 8
Locality: Zv-005, Zv-006, Zv-009, Zv-010
Family: Rosalinidae Reiss, 1963
Neoconorbina sp. 1
Pl. 8, Figs. 16-17 Locality: Vr-005, Vr-005A
Family: Parrelloididae Hofker, 1956
Cibicidoides ungerianus (d’Orbigny), 1846
Pl. 9, Figs. 1-2
1846 Rotalina ungeriana - d’Orbigny, p. 157, pl. 8, figs. 16-18
1950 Cibicides granosus (Reuss) - Ruiz de Gaona & Colom, p. 368, fig. 13/21-27
1956 Cibicides ungerianus (d’Orbigny) - Hagn, p. 181, pl. 17, figs. 10-11
1975 Heterolepa ungeriana (d’Orbigny) - Braga et al., p. 98
1991 Cibicidoides ungerianus (d’Orbigny) -Barbin & Keller-Grünig, p. 24
1993 Cibicidoides ungerianus (d’Orbigny) -Sztràkos (In: Mathelin & Sztràkos), p. 81, pl. 16, fig. 11
Locality: Vr-004, Vr-010, Vr-011, Zv-004,
Zv-006, Zv-008
Family: Discorbinellidae Sigal, 1952
Laticarinina altocamerata (Heron-Allen & Earland), 1922
Pl. 9, Fig. 3
1922 Truncatulina tenuimargo Brady var. altocamerata - Heron-Allen & Earland, p. 209, pl. 7, figs. 24-27 (Fide Ellis & Messina 1940)
1964 Laticarinina altocamerata (Heron-Allen & Earland) - Loeblich & Tappan, p. C580, fig. 457, 4
1983 Laticarinina altocamerata (Heron-Allen & Earland) - Setiawan, p. 116. pl. 5, fig. 4
1991 Laticarinina altocamerata (Heron-Allen & Earland) - Barbin & Keller-Grünig, p. 242 Locality: Vr-005A, Zv-008 Family: Planulinidae Bermudez, 1952
Planulina compressa (Hantken), 1875
Pl. 9, Fig. 4
1875 Truncatulina compressa - Hantken, p. 72, pl. 8, fig. 8a-b
1956 Planulina compressa (Hantken) - Hagn, p. 179, pl. 17, fig. 5a-b
1975 Planulina compressa (Hantken) - Braga et al., p. 95, pl. 2, fig. 13a-c
1993  Planulina compressa (Hantken) - Sztràkos (In:
Mathelin & Sztràkos), p. 81, pl. 17, fig. 1 Locality: Vr-005A
Planulina costata (Hantken), 1875
Pl. 9, Figs. 5-6
1875 Truncatulina costata - Hantken, p. 73, pl. 9, fig. 2
1956   Planulina costata Hantken – Hagn, p. 178, pl. 17, figs. 3a-b, 4a-b
1984 Planulina costata (Hantken) - Grünig & Herb, p. 262
1993  Planulina costata (Hantken) - Sztràkos (In: Mathelin & Sztràkos), p. 81, pl. 16, fig. 13; pl. 38, fig. 9 Locality: Vr-004, Vr-005, Vr-005A, Vr-010, Vr-011, Vr-012, Zv-005, Zv-006, Zv-007, Zv-008, Zv-009, Zv-010
Family: Cibicididae Cushman, 1927
Lobatula lobatula (Walker & Jacob), 1798
Pl. 9, Figs. 7-8
1798 Nautilus lobatulus - Walker & Jacob (in Kanmacher), p. 642, pl. 14, fig. 36 (Fide Ellis & Messina 1940))
1957   Cibicides lobatulus (Walker & Jacob) - Sacal & Debourle, p. 68, pl. 33, fig. 1
1962 Cibicides lobatulus (Walker & Jacob) -Gohrbandt, p. 113
1991 Cibicides lobatulus (Walker & Jacob) -Barbin & Keller-Grünig, p. 240, pl. 1, figs. 17-18
1993  Lobatula   lobatula   (Walker   &   Jacob)   -Sztràkos (In: Mathelin & Sztràkos), p. 81, pl. 17, fig. 9 Locality: Vr-005, Vr-005A, Zv-005, Zv-010
Cycloloculina eocenica (Terquem), 1882
Pl. 9, Figs. 9-10
1882 Planorbulina eocenica - Terquem, p. 90, pl. 9, fig. 15a-b
1970 Cycloloculina eocenica (Terquem) - Le Calvez, p. 186, pl. 35, fig. 4
1993 Cycloloculina eocenica (Terquem) -Sztràkos (In: Mathelin & Sztràkos), p. 81, pl. 19, fig. 1; pl. 43, fig. 6
Locality: Vr-004, Vr-005A
Family: Planorbulinidae Schwager, 1877
Planorbulina difformis Roemer, 1838
Pl. 9, Fig. 11
Late Eocene benthic foraminiferal fauna from clastic sequence of the Socka - Dobrna area...           35
PLATE 10
1-2 Asterigerina brencei Haque. 1 umbilical view, Vr-005A, x120. 2 umbilical view, Vr-005A, x120. 3-4 Asterigerina sp. 3 edge view, Vr-005, x80. 4 umbilical view, Vr-004, x80. 5 Asterigerina rotula (Kaufmann), edge view, Vr-009, x40. 6 Nonion scaphum (Fichtel & Moll), side view, Vr-005, x80. 7-8 Melonis halkyardi (Cushman). 7 side view, Zv/II-008, x80. 8 apertural view, Vr-005, x80. 9 Pullenia bulloides (d’Orbigny), apertural view, Zv/II-011, x80. 10 Pullenia quinqueloba (Reuss), side view, Vr-010, x80. 11-12 Queraltina epistominoides Marie. 11 Vr-005A, x60. 12 Marie, Vr-005A, x60. 13 Chilostomella ovoidea Reuss, Zv/II-011, x60. 14-15 Charltonina budensis (Hantken). 14 spiral view, Vr-005A, x60. 15 umbilical view, Vr-005A, x60. 16 Anomalinoides affinis (Hantken), Vr-005, x80.
36
Franc Cimerman, Bogomir Jelen & Dragomir Skaberne
1993 Planorbulina difformis Roemer - Sztràkos (In: Mathelin & Sztràkos), p. 81, pl. 38, figs. 5-6
Locality: Vr-005A
Family: Cymbaloporidae Cushman, 1927
Halkyardia minima (Liebus), 1911
Pl. 9, Fig. 12
1911 Cymbalopora radiata Hagenow var. minima - Liebus, p. 952, pl. 3, fig. 7a-c
1969   Halkyardia minima (Liebus) - Cimerman, p. 298, pl. 58, figs. 1-6
1976 Halkyardia minima (Liebus) - [iki} et al., p. 195, pl. 6, fig. 1
1979 Halkyardia minima (Liebus) - Drobne et al., p. 158
1985 Halkyardia minima (Liebus) - Grünig, p. 272, pl. 9, figs. 16-17
1991 Halkyardia minima (Liebus) - Barbin & Keller-Grünig, p. 241
1993  Halkyardia minima (Liebus) - Sztràkos (In: Mathelin & Sztràkos), p. 54, pl. 38, fig. 12 Locality: Vr-005A, Vr-011, Vr-011A, Vr-014
Halkyardia sp.
Pl. 9, Fig. 13 Locality: Vr-011A
Family: Stilostomellidae Finlay, 1947
Nodogenerina atlantisae (Cushman), 1939
Pl. 8, Figs. 5-6
1939 Ellipsonodosaria atlantisae - Cushman, p. 70, pl. 12, figs. 3-4 (Fide Ellis & Messina 1940)
1993 Nodogenerina atlantisae (Cushman) -Sztràkos (In: Mathelin & Sztràkos), p. 80, pl. 13, figs. 16-18; pl. 34, figs. 18-19
Locality: Zv-006, Zv-008
Family: Victoriellidae Chapman & Crespin, 1930
Korobkovella grosserugosa (Gümbel), 1868
Pl. 9, Figs. 14-15
1868 Truncatulina grosserugosa - Gümbel, p. 82, pl. 2, fig. 104
1950 Anomalina grosserugosa (Gümbel) - Ruiz de Gaona & Colom, p. 362
1957 Anomalina grosserugosa (Gümbel) - Sacal & Debourle, p. 66, pl. 33, fig. 6
1970   Anomalina grosserugosa (Gümbel) - Le Calvez, p. 198, pl. 40, fig. 2
1983 Anomalina grosserugosa (Gümbel) -Setiawan, p. 132, pl. 14, fig. 1
1993 Korobkovella    grosserugosa    (Gümbel)    -Sztràkos (In: Mathelin & Sztràkos), p. 82, pl. 19. Fig. 2 Locality: Vr-004, Vr-005A, Vr-009, Vr-010, Vr-012, Vr-014, Zv-006, Zv-007
Family: Acervulinidae Schultze, 1854
Sphaerogypsina globula (Reuss), 1848
Pl. 9, Fig. 16
1848 Ceriopora globulus - Reuss, p. 33, pl. 5, fig. 7
1950 Gypsina globula (Reuss) - Ruiz de Gaona & Colom, p. 380
1979 Sphaerogypsina globula (Reuss) - Drobne et al., p. 158, pl. 3, fig. 1; pl. 4, figs. 3-4
1985 Sphaerogypsina globula (Reuss) - Grünig, p. 272, pl. 9, figs. 14-15
1993 Sphaerogypsina globula (Reuss) - Sztràkos (In: Mathelin & Sztràkos), p. 82, pl. 38, fig. 13
Locality: Vr-004, Vr-005, Vr-005A
Family: Asterigerinidae d’Orbigny, 1839
Asterigerina brencei Haque, 1960
Pl. 10, Figs. 1-2
1960  Asterigerina brencei Haque - Setiawan, p.
118, pl. 9, fig. 3 Locality: Vr-005, Vr-009, Vr-010
Asterigerina rotula (Kaufmann), 1867
Pl. 10, Fig. 5
1867 Hemistegina rotula - Kaufmann, p. 150, pl. 8, fig. 19 (Fide Ellis & Messina 1940)
1886 Pulvinulina rotula (Kaufmann) - Uhlig, p. 193, pl. 3, figs. 5-6; pl. 5, figs. 6-7
1979 Asterigerina rotula (Kaufmann) - Drobne et al., p. 158, pl. 3, fig. 2; pl. 4, fig. 5
1993 Asterigerina rotula (Kaufmann) - Sztràkos (In: Mathelin & Sztràkos), p. 82, pl. 19, fig. 7
Locality: Vr-010, Vr-011
Asterigerina sp. 1
Pl. 10, Figs. 3-4 Locality: Vr-005, Vr-005A
Family: Nonionidae Schultze, 1854
Nonion scaphum (Fichtel et Moll), 1798
Pl. 10, Fig. 6
1798 Nautilus scapha - Fichtel & Moll, p. 105, pl. 19, figs. d-f (Fide Ellis & Messina 1940)
1846 Nonionina communis d’Orbigny -d’Orbigny, p. 106, pl. 5, fig. 7
Late Eocene benthic foraminiferal fauna from clastic sequence of the Socka - Dobrna area...           37
PLATE 11
1-2 Gyroidinoides girardanus (Reuss). 1 spiral view, Zv/II-006, x40. 2 umbilical view, Zv/II-006, x40. 3-4 Heterolepa eocaena (Gümbel). 3 spiral view, Zv/II-006, x28. 4 umbilical view, Zv/II-011, x35. 5 Heterolepa reussi (Silvestri), spiral view, Vr-012, x50. 6-8 Gavelinella acuta (Plummer). 6 spiral view, Vr-009, x40. 7 edge view, Vr-009, x400. 8 spiral view, Vr-009, x40. 9 Pararotalia armata (d’Orbigny), spiral view, Vr-011, x60. 10-11 Pararotalia audouini (d’Orbigny). 10 spiral view, Vr-011, x80. 11 spiral view, Vr-011, x80. 12-13 Calcarina lecalvezae Boulanger & Poignant. 12 Vr-014, x28. 13 Zv/II-009, x28. 14-15 “Eponides” dalmatinus (de Witt Puyt). 14 spiral view, Vr-009, x28, 15 umbilical view, Zv/II-010, x28.
38
Franc Cimerman, Bogomir Jelen & Dragomir Skaberne
1961 Nonion scaphum (Fichtel & Moll) -Kaasschieter, p. 204, pl. 10, fig. 5
1993  Nonion commune (d’Orbigny) - Sztràkos (In: Mathelin & Sztràkos), p. 82, pl. 19, fig. 11 Locality: Vr-005
Melonis halkyardi (Cushman), 1936
Pl. 10, Figs. 7-8
1936 Nonion halkyardi - Cushman, p. 63, pl. 12, fig. 1 (Fide Ellis & Messina 1940)
1987aMelonis halkyardi (Cushman) - Sztràkos, p.
32, pl. 15, fig. 6 Locality: Vr-005, Vr-012, Zv/II-008, Zv/II-011
Pullenia bulloides (d’Orbigny), 1846
Pl. 10, Fig. 9
1846 Nonionina bulloides - d’Orbigny, p. 107, pl. 5, fig. 9-10
1875 Pullenia bulloides (d’Orbigny) - Hantken, p. 59, pl. 10, fig. 9
1987aPullenia bulloides (d’Orbigny) - Sztràkos,
p. 34 Locality: Vr-005A, Vr-011, Vr-011A, Zv-006, Zv-010, Zv-011
Pullenia quinqueloba (Reuss), 1851
Pl. 10, Fig. 10
1851 Nonionina quinqueloba - Reuss, p. 71, pl. 5, fig. 31
1987 Pullenia quinqueloba (Reuss) - Wenger, p. 299, pl. 13, figs. 16, 20
1991 Pullenia quinqueloba (Reuss) - Barbin & Keller-Grünig, p. 240
1993  Pullenia quinqueloba (Reuss) - Sztràkos (In: Mathelin & Sztràkos), p. 82, pl. 19, fig. 17 Locality: Vr-005A, Zv-005, Zv-006
Family: Almaenidae Myatlyuk, 1959
Queraltina epistominoides Marie, 1950
Pl. 10, Figs. 11-12
1950 Queraltina epistominoides - Marie, p. 74, figs. 1-3, 8-9 (Fide Ellis & Messina 1940)
1983 Queraltina epistominoides Marie -Setiawan, p. 133, pl. 14, figs. 2-3
1985 Almaena epistominoides (Marie) - Grünig, p. 276
1987aQueraltina epistominoides Marie - Sztràkos, p. 34
1991 Queraltina epistominoides Marie - Barbin & Keller-Grünig, p. 241, pl. 2, fig. 16
1993 Queraltina epistominoides Marie - Sztràkos (In: Mathelin & Sztràkos), p. 56, pl. 39, fig. 20
Locality: Vr-004, Vr-005A, Vr-012, Vr-014
Family: Chilostomellidae Brady, 1881
Chilostomella ovoidea Reuss, 1850
Pl. 10, Fig. 13
1850 Chilostomella ovoidea - Reuss, p. 380, pl. 48, fig. 12
1993 Chilostomella cf. ovoidea Reuss - Sztràkos (In: Mathelin & Sztràkos), p. 82, pl. 20. fig. 1
Locality: Zv-005, Zv-006, Zv-009, Zv-010,
Zv-011
Family: Osangulariidae Loeblich & Tappan, 1964
Charltonina budensis (Hantken), 1875
Pl. 10, Figs. 14-15
1875 Truncatulina budensis - Hantken, p. 75, pl. 8, fig. 6
1919 Pulvinulina acutimargo - Halkyard, p. 120, pl. 7, fig. 4
1985 Alabamina acutimargo (Halkyard) - Grünig, p. 274, pl. 10. fig. 9-11
1987aAlabamina budensis (Hantken) - Sztràkos, p. 32, pl. 12, fig. 8
1993  Charltonina budensis (Hantken) - Sztràkos (In: Mathelin & Sztràkos), p. 57, pl. 20, fig. 4 Locality: Vr-005A, Vr-009, Vr-010, Vr-011, Vr-012
Family: Heterolepidae Gonzales-Donoso, 1969
Anomalinoides affinis (Hantken), 1875
Pl. 10, Fig. 16
1875 Pulvinulina affinis - Hantken, p. 78, pl. 10, fig. 6
1941 Anomalina dalmatina nov. sp. - van Bellen, p. 1001, fig. 26a-c
1956 Anomalinoides dalmatinus (van Bellen) -Hagn, p. 177, pl. 17, fig. 1a-b
1993  Anomalinoides affinis (Hantken) - Sztràkos (In: Mathelin & Sztràkos), p. 82, pl. 20, fig. 8 Locality: Vr-004, Vr-005A, Zv-006, Zv-007, Zv-009, Zv-010, Zv-011
Heterolepa eocaena (Gümbel), 1868
Pl. 11, Fig. 3-4
1868 Rotalia eocaena - Gümbel, p. 72, pl. 2, fig. 87a-b
1956 Cibicides eocaenus (Gümbel) - Hagn, p. 180, pl. 17, figs. 8-9
1985  Heterolepa eocaena (Gümbel) - Grünig, p.
275, pl. 11, figs. 1-3 Locality: Vr-004, Vr-005, Vr-005A, Vr-
Late Eocene benthic foraminiferal fauna from clastic sequence of the Socka - Dobrna area...           39
PLATE 12
1-2 Operculina cf. gomezi Colom & Bauza. 1 Vr-005A, x32. 2 Vr-005A, x32. 3 Eoannularia eoceniva Cole & Bermudez, Vr-005A, x64. 4 Asterocyclina sp. Vr-005A, x80. 5 Discocyclina trabayensis vicenzensis Less, Vr-005A, x64. 6 Discocyclina s. l., Vr-005A, x64. 7-9Heterostegoina sp. 1. 7 Vr-005A, x64 . 8 Vr-005A, x64. 9 Vr-005A, x64.
40
Franc Cimerman, Bogomir Jelen & Dragomir Skaberne
012, Zv-005, Zv-011
Heterolepa reussi (Silvestri), 1906
Pl. 11, Fig. 5
1906 Truncatulina dutemplei (d’Orbigny) var. reussi - Silvestri, p. 33 (Fide Ellis & Messina 1940)
1956 Cibicides dalmatinus van Bellen - Hagn, p. 180, pl. 18, fig. 5a-b
1985  Heterolepa reussi (Silvestri) - Grünig, p. 76,
pl. 11, figs. 7-9 Locality: Vr-011, Vr-012
Family: Gavelinellidae Hofker, 1956
Gyroidinoides girardanus (Reuss), 1851
Pl. 11, Figs. 1-2
1851 Rotalina girardana - Reuss, p. 73, pl. 5, fig. 34
1993 Gyroidinoides girardanus (Reuss) - Sztràkos (In: Mathelin & Sztràkos), p. 83, pl. 21, fig. 3 Locality: Vr-004, Vr-005A, Vr-010, Vr-012, Zv-006, Zv-007, Zv-009, Zv-011
Gavelinella acuta (Plummer), 1926
Pl. 11, Figs. 6-8
1926 Anomalina ammonoides (Reuss) var. acuta n. var. - Plummer, p. 149, pl. 10, fig. 2a-c
1983 Anomalina acuta Plummer - Setiawan, p. 131, pl. 14, fig. 3
1987aGavelinella acuta (Plummer) - Sztràkos, p. 31, pl. 13, figs. 12, 13
Locality: Vr-009
Family: Linderinidae Loeblich & Tappan, 1984
Eoannularia eocenica Cole & Bermudez, 1944
Pl. 12, Fig. 3
1944 Eoannularia eocenica - Cole & Bermudez, p. 12, pl. 1, figs. 11-16 (Fide Ellis & Messina 1940)
1976 Eoannularia eocenica Cole & Bermudez – Sirel, p. 80, pl. 1. figs. 1-3; pl. 2, figs. 1, 2, 4, 5, 8, 9
Locality: Vr-005A, Vr-011
Family: Rotaliidae Ehrenberg, 1839
Pararotalia armata (d’Orbigny), 1826
Pl. 11, Fig. 9
1826 Rotalia armata - d’Orbigny, p. 273, pl. 8, fig. no. 22
1970 Pararotalia armata (d’Orbigny) - Le Calvez, p. 161, pl. 39, fig. 1
1985 Pararotalia armata (d’Orbigny) - Grünig, p. 270, pl. 8, fig. 9
1993 Pararotalia armata (d’Orbigny) - Sztràkos (In: Mathelin & Sztràkos), p. 83, pl. 42, fig. 11
Locality: Vr-011
Pararotalia audouini (d’Orbigny), 1826
Pl. 11, Figs. 10-11
1826  Rotalia audouini - d’Orbigny, p. 273, no. 19
1970   Pararotalia audouini (d’Orbigny) - Le Calvez, p. 162, pl. 34, fig. 8
1975 Pararotalia audouini (d’Orbigny) - Braga et al., p. 107, pl. 6, figs. 8-9
1983 Pararotalia audouini (d’Orbigny) -Setiawan, p. 120
1993 Pararotalia audouini (d’Orbigny) - Sztràkos (In: Mathelin & Sztràkos), p. 83, pl. 42, fig. 12
Locality: Vr-011
Family: Calcarinidae Schwager, 1876
Calcarina lecalvezae Boulanger & Poignant, 1971
Pl. 11, Figs. 12-13
1971   Calcarina le calvezae - Boulanger & Poignant, p. 96, pl. 1, figs. 1-10
1991  Calcarina lecalvezae Boulanger & Poignant
- Barbin & Keller-Grünig, p. 242, pl. 1, fig. 22
1993  Calcarina lecalvezae Boulanger & Poignant
- Sztràkos (In: Mathelin & Sztràkos), p. 83, pl. 22, figs. 7-9
Locality: Vr-014, Zv-009
Family: Nummulitidae de Blainville, 1827
Heterostegina sp. 1
Pl. 12, Figs. 7-9 Locality: Vr-005, Vr-005A
Operculina cf. gomezi Colom & Bauza, 1950
Pl. 12, Figs. 1-2
1950 Operculina canalifera gomezi - Colom & Bauza, p. 217, pl. 16, figs. 1-3; texte, figs. 1-2. (Fide Ellis & Messina 1940)
1971 Operculina canalifera gomezi Colom & Bauza - Ferrer, p. 35, fig. 19
1977 Operculina gomezi Colom & Bauza -Hottinger, p. 98, figs. 38 A-F
1993 Operculina gomezi Colom & Bauza -Mathelin (In: Mathelin & Sztràkos), p. 62, pl. 44, fig. 4
Late Eocene benthic foraminiferal fauna from clastic sequence of the Socka - Dobrna area...           41
Locality: Vr-005A
Family: Discocyclinidae Galloway, 1928
Discocyclina trabayensis vicenzensis Less, 1987
Pl. 12, Fig. 5
1987 Discocyclina trabayensis Neumann, 1955 vicenzensis n. ssp. - Less, p. 172, pl. 18, figs. 3-5, textfig. 27y
Locality: Vr-005A
Discocyclina s. l.
Pl. 12, Fig. 6
Locality: Vr-004, Vr-005, Vr-005A, VR-009
Asterocyclina sp. 1
Pl. 12, Fig. 4
Locality: Vr-004, Vr-005, Vr-005A
Acknowledgemets
We thank the Slovenian Research Agency for financing this project. To the late Monique Toumarkine, who in the meantime passed away, go our thanks for determinations of planktic species, to Vlasta ]osovi} for determination of Lepidocyclina, and to Jo‘e ^ar, Ladislav Placer and Laszlo Fodor for discussions on tectonic deformations in profiles. Thanks also to Tom Popit and Alenka Eva ^rne who technically prepared the plates, and to Simon Pirc for translation of the text into English, and to Glenn S. Jacks for English corrections of the final version.
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