175
Key words: Checklist, Red list, 
hornworts, liverworts, threatened 
species, Slovenia.
 
Ključne besede: Seznam, 
Rdeči seznam, rogovnjaki 
(Anthocerotophyta), jetrenjaki, 
(Marchantiophyta), ogrožene vrste, 
Slovenija.
Corresponding author:
Andrej Martinčič
E-mail:  
andrej.martincic@siol.net
Received: 20. 3. 2024
Accepted: 24. 5. 2024
New checklist and the Red list of the 
hornworts (Anthocerotophyta) and 
liverworts (Marchantiophyta) of Slovenia
Abstract
The new Checklist of hornworts (Anthocerotophyta) and liverworts 
(Marchantiophyta) of Slovenia within current political boundaries comprises 181 
species, additional 4 subspecies and 2 varieties. Additional 5 species are denoted 
by a question mark since reports are considered questionable. An alphabetically 
arranged tabular presentation of genera, species and lower taxa shows their presence 
in the phytogeographical units of Slovenia. The records from the periods before 
and after 1958 are marked with different symbols. Each species also has a new 
Red List status in Slovenia. The checklist includes the data from the literature and 
Herbarium LJU. Annotations to selected species give a critical evaluation of the 
literature and herbarium data. A list of taxa that should be excluded from the flora 
of Slovenia, the synonyms and the used literature with floristic data not cited in 
this article are also added.
Izvleček
Novi Seznam rogovnjakov (Anthocerotophyta) in jetrenjakov (Marchantiopyta) 
Slovenije v današnjih političnih mejah obsega 181 vrst, dodatne 4 podvrste in 2 
varieteti. Nadaljnih 5 vrst je označenih z vprašajem, ker gre za navedbe, ki so iz 
kakršnega koli vzroka dvomljive. Po abecedi urejen tabelarni prikaz rodov in vrst 
ter nižjih taksonov prikazuje prisotnost v posameznih fitogeografskih enotah. Z 
ustreznim znakom so prikazani podatki pred letom 1958 in po njem. Vsaka vrsta 
ima oznako statusa iz Rdečega seznama za Slovenijo. Seznam vključuje podatke 
iz literature in herbarija LJU. Pripombe kritično vrednotijo določene literaturne 
in herbarijske podatke. Dodan je spisek taksonov, ki jih je treba črtati iz flore 
Slovenije, sinonimi in uporabljena literatura s florističnimi podatki za Slovenijo, ki 
ni bila citirana v tekstu.
Andrej Martinčič1
1 Ljubljana, Slovenija
DOI: 10.2478/hacq-2023-001323/2 • 2024, 175–197
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To present the distribution of particular taxa, we 
used the phytogeographical division of Slovenia accord-
ing to M. Wraber (1969), which was slightly modified 
(Martinčič, 2003). The central Soča Valley was therefore 
excluded from the sub-Mediterranean region – or Adriatic 
province sensu Zupančič & Žagar (1995) as proposed by 
Dakskobler (1996) – and included in the pre-Alpine re-
gion. Similarly, we divided the so far uniform subregion of 
the Alpine region of Pohorje into three parts: Pohorje in 
the strict sense, Dravski Kozjak – the region north of the 
Drava river, and Mežiško-Mislinjska dolina with Strojna 
– in the west of Pohorje. The phytogeographical units are 
presented in Figure 1 with the following abbreviations:
AJ – Alpine region: Julijske Alpe
AK – Alpine region: Karavanke
AS – Alpine region: Kamniško-Savinjske Alpe
AP – Alpine region: Pohorje
AZ – Alpine region: Dravski Kozjak
AM – Alpine region: Mežiško-Mislinjska dolina-Strojna
DN – Dinaric region
PA – pre-Alpine region
PD – pre-Dinaric region
SM – Submediterranean region
SP – sub-Pannonian region
Introduction
The work by Scopoli entitled »Flora carniolica« (1st edi-
tion 1760, 2nd edition 1772) marks the beginning of re-
search of the bryophyte flora of Slovenia. It was followed 
by numerous works which, however, gave information, 
also in the form of florae, only for particular regions of 
Slovenia within the present-day political boundaries. 
Thirteen years after the publication of the Checklist of 
hornworts and liverworts of Slovenia (Martinčič, 2011; 
in Slovenian), it is necessary to provide a new, updated 
checklist. Over the last period, a substantial increase in 
floristic data has occured, mainly due to intensive field 
work and the processing of herbarium material in the 
Herbarium LJU. Another reasons for the updated Check-
list are recent taxonomical and nomenclatural changes in 
the European bryophytes flora (Ros et al., 2007; Hodg-
etts & Lockhart, 2020; Hodgetts et al., 2020).
Methods
The new checklist is based on data from literature sour-
ces published before 2024, the collection of Herbarium 
LJU, and the author‘s previously unpublished data. It 
considers all floristic works and a part of published phy-
tosociological relevés in which we assume liverworts were 
correctly identified. We critically evaluated data from ol-
der floristic literature related to the border areas between 
Slovenia and Italy. After the collapse of the Austro-Hun-
garian Monarchy, a part of these areas was added to Ita-
ly, and the data from these parts (mainly the vicinity of 
Gorizia and Trieste) was reported both by Pavletić (1955) 
and Düll et al. (1999). In the absence of appropriate her-
baria specimens, it was impossible to review the accuracy 
of all data reported in the literature. The only exception 
was the herbarium material kept in LJU. In the last de-
cades of the 19th century, it was obtained mainly from S. 
Robič, J. Šafer, J. Breidler and A. Paulin. The material 
contributed by the first two authors was revised or reclas-
sified by J. Breidler and J. Głowacki. F. Dolšak collected 
liverwort samples in the early 20th century. 
The reports that are in any way questionable but could 
not be revised because there is no herbarium material 
available were marked with a question mark. Some of the 
floristic data obtained from literature dates back to the 
period before 1915 and a part comes from the period be-
tween 1958 and 2023. Most of the herbarium material 
in LJU was collected after 1960, predominantly by the 
author of this paper.
The nomenclature and taxonomy used in the checklist 
follow Hodgetts et al. (2020).
Figure 1: Phytogeographic division of Slovenia (according to 
M. Wraber (1969), modified).
Slika 1: Fitogeografska razdelitev Slovenije (po M. Wraberju (1969), 
spremenjeno).
The accompanying Red List is added to the checklist 
(Table 1), although the categories are mostly based on the 
previously published Red List (Martinčič, 2016a). The 
species were classified into the Red List categories using 
the IUCN criteria 3.1 (IUCN 2001). Their application to 
bryophytes follows Hallingbäck et al. (1998). However, 
the only realistic threat assessment criterion is criterion B, 
which is expressed with the number of currently known 
localities: 1 (CR), 2–5 (EN), 6–10 (VU). The year of the 
last record was added for the taxa under DD category.
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New checklist of the hornworts and liverworts of Slovenia
SLO RED LIST – Status of taxon in Red List of Slovenia
Status taksona v Rdečem seznamu za Slovenijo
RE – Regionally extinct – izumrla v Sloveniji
CR – Critically endangered – skrajno ogrožena vrsta
EN – Endangered – prizadeta vrsta
VU – Vulnerable – ranljiva vrsta
NT – Near threatened – potencialno ogrožena vrsta
DD – Data deficient – premalo znana vrsta, stari podatki
DD-n – Data deficient, new – premalo znana vrsta, 
recentni podatki
LC – Least concern – neogrožena vrsta
List of taxa with distribution in phytogeographic 
regions of Slovenia
The following symbols were used:
● – Literature or herbarium report after 1958
Literaturni ali herbarijski podatek po letu 1958
○ – Literature or herbarium report before 1958
Literaturni ali herbarijski podatek pred letom 1958
? – Questionable report – dvomljiv podatek
SLO RED LIST AJ AK AS AP AZ AM DN PA PD SM SP
Anastrophyllum (Spruce) Steph. (Anastrophyllaceae)
michauxii (F. Weber) H. Buch LC ○ ● ●
Aneura Dumort. (Aneuraceae)
pinguis (L.) Dumort. LC ● ● ● ● ● ● ● ● ● ● ○
Anthelia (Dumort.) Dumort. (Antheliaceae)
juratzkana (Limpr.) Trevis. LC ● ●
Anthoceros L. (Anthocerotaceae)
agrestis Paton LC ● ●
punctatus L. (1) DD-1913 ○ ○
Apopellia (Grolle) Nebel & D. Quandt. (Pelliaceae)
endiviifolia (Dicks.) Nebel & D. Quandt. LC ● ● ● ● ● ● ● ● ● ● ●
Arnellia Lindb. (Arnelliaceae)
fennica (Gottsche) Lindb. DD-1905 ○
Asterella P. Beauv. (Aytoniaceae)
lindenbergiana (Corda ex Nees) Lindb. ex Arnell (2) VU ●
Barbilophozia Loeske (Anastrophyllaceae)
barbata (Schmidel ex Schreb.) Loeske LC ● ● ● ○ ○ ● ● ●
hatcheri (A. Evans) Loeske LC ● ● ●
lycopodioides (Wallr.) Loeske LC ● ● ●
sudetica (Nees ex Huebener) L. Söderstr. EN ● ●
Bazzania Gray (Lepidoziaceae)
flaccida (Dumort.) Grolle (3) DD-n-1999 ●
tricrenata /Wahlenb.) Lindb. LC ● ○ ● ○ ○ ○
trilobata (L.) Gray var. trilobata LC ● ● ● ● ● ● ● ● ● ● ●
var. depauperata (Müll. Frib.) Grolle (4) DD-n 2010 ●
Blasia L. (Blasiaceae)
pusilla L. LC ○ ● ○ ● ● ● ○ ●
Blepharostoma (Dumort.) Dumort. (Blepharostomataceae)
trichophyllum (L.) Dumort. subsp. trichophyllum LC ● ● ● ● ● ● ● ● ● ●
subsp. brevirete (Bryhn. & Kaal.) R. M. Schust. LC ● ●
Calypogeia Raddi (Calypogeiaceae)
arguta Nees & Mont. LC ● ● ● ● ● ○
azurea Stotler & Crotz LC ● ● ● ● ● ● ● ● ● ● ●
Table 1: List of taxa with their distributions in phytogeographic regions of Slovenia.
Tabela 1: Seznam taksonov s prikazom razširjenosti v fitogeografskih enotah Slovenije.
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SLO RED LIST AJ AK AS AP AZ AM DN PA PD SM SP
fissa (L.) Raddi LC ● ● ● ● ● ● ● ● ○ ●
integristipula Steph. LC ● ● ● ● ● ● ● ● ●
muelleriana (Schiffn.) Müll. Frib. LC ● ● ● ● ● ● ○ ●
neesiana (C. Massal. & Carestia) Müll. Frib. LC ● ● ● ● ● ●
suecica (Arnell & J. Perss.) Müll. Frib. LC ● ● ● ●
Cephalozia (Dumort.) Dumort. (Cephaloziaceae)
ambigua C. Massal. VU ● ●
bicuspidata (L.) Dumort. subsp. bicuspidata LC ● ● ● ● ● ● ● ● ● ○ ○
subsp. lammersiana (Huebener) R. M. Schust. VU ●
lacinulata (J. B. Jack ex Gottsche & Rabenh.) Spruce LC ● ● ● ○
Cephaloziella (Spruce) Schiffn. (Cephaloziellaceae)
baumgartneri Schiffn. (5) DD-n 1999 ●
divaricata (Sm.) Schiffn. DD-1913 ○ ○ ○ ○
elachista (J. B. Jack ex Gottsche & Rabenh.) Schiffn. DD-1908 ○
hampeana (Nees) Schiffn. ex Loeske (6) EN ●
integerrima (Lindb.) Warnst. (7) DD-1912 ○
rubella (Nees) Warnst. LC ● ○ ● ○ ● ○
spinigera (Lindb.) Jørg. (8) DD-n 2021 ●
Chiloscyphus Corda (Lophocoleaceae)
pallescens (Ehrh.) Dumort. var. pallescens LC ● ● ● ● ● ● ● ● ● ● ●
polyanthos (L.) Corda var. polyanthos LC ● ● ● ● ● ● ● ● ● ● ●
var. rivularis (Schrad.) Lindb. & Arnell LC ● ○ ● ● ○ ○ ○
Clevea Lindb. (Cleveaceae)
hyalina (Sommerf.) Lindb. LC ● ● ●
Cololejeunea (Spruce) Steph. (Lejeuneaceae)
calcarea (Lib.) Steph. LC ● ● ● ● ○ ● ● ● ● ○
rossetiana (C. Massal.) Schiffn. DD-1912 ○ ○
Conocephalum Hill (Conocephalaceae)
conicum (L.) Dumort. LC ● ● ● ● ● ● ● ● ● ● ●
salebrosum Szweyk., Buczk. & Odrzyk. LC ● ● ● ●
Corsinia Raddi (Corsiniaceae)
coriandrina (Spreng.) Lindb. (9) DD-1841 ?
Crossocalyx Meyl. (Anastrophyllaceae)
hellerianus (Nees ex Lindenb.) Meyl. DD-1913 ○ ○ ○
Diplophyllum (Dumort.) Dumort. (Scapaniaceae) 
albicans (L.) Dumort. LC ● ● ● ● ● ● ● ● ● ●
obtusifolium (Hook.) Dumort. LC ○ ● ● ○ ● ● ○ ○ ●
taxifolium (Wahlenb.) Dumort. VU ○ ● ●
Fossombronia Raddi (Fossombroniaceae)
foveolata Lindb. (10) DD-1917 ○
pusilla (L.) Nees DD-1912 ○ ○ ○ ○
wondraczekii (Corda) Dumort. ex Lindb. VU ○ ● ○ ●
Frullania Raddi (Frullaniaceae)
cleistostoma Schiffn. & W. Wollny (11) DD-n 2007 ●
dilatata (L.) Dumort. LC ● ● ● ● ● ● ● ● ● ● ●
fragilifolia (Taylor) Gottsche, Lindenb. & Nees LC ● ○ ● ● ● ● ○
jackii Gottsche VU ● ○ ○
riparia Hampe VU ○ ● ● ●
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tamarisci (L.) Dumort. LC ● ● ● ● ● ● ● ● ●
teneriffae (F. Weber) Nees (12) ?
Fuscocephaloziopsis Fulford (Cephaloziaceae)
catenulata (Huebener) Vaňa & L. Söderstr. LC ● ● ● ● ○ ● ● ● ●
connivens (Dicks.) Vaňa & L. Söderstr. LC ● ○ ●
leucantha (Spruce) Vaňa & L. Söderstr. LC ● ● ● ● ●
loitlesbergeri (Schiffn.) Vaňa & L. Söderstr. EN ● ●
lunulifolia (Dumort.) Vaňa & L. Söderstr. LC ● ● ● ● ● ● ● ●
pleniceps (Austin) Vaňa & L. Söderstr. VU ● ● ○
Geocalyx Nees (Geocalycaceae)
graveolens (Schrad.) Nees (13) EN ●
Gymnocolea (Dumort.) Dumort. (Anastrophyllaceae)
inflata (Huds.) Dumort. LC ● ○ ● ○
Gymnomitrion Corda (Gymnomitriaceae)
concinnatum (Lightf.) Corda (14) DD-1894 ○
corallioides Nees (15) DD-1894 ○
Harpanthus Nees (Harpanthaceae)
flotovianus (Nees) Nees EN ● ○
scutatus (F. Weber & D. Mohr) Spruce DD-1912 ○ ○
Isopaches H. Buch (Anastrophyllaceae)
bicrenatus (Schmidel ex Hoffm.) H. Buch NT ○ ● ○ ○
Jungermannia L. (Jungermanniaceae)
atrovirens Dumort. LC ● ● ● ● ● ● ● ● ● ● ●
polaris Lindb. VU ●
pumila With. LC ● ● ● ●
Kurzia G. Martens (Lepidoziaceae)
pauciflora (Dicks.) Grolle LC ○ ○ ● ● ○ ●
Lejeunea Lib. (Lejeuneaceae)
cavifolia (Ehrh.) Lindb. LC ● ● ● ● ● ● ● ● ● ● ●
patens Lindb. (16) ?
Lepidozia (Dumort.) Dumort. (Lepidoziaceae)
reptans (L.) Dumort. LC ● ● ● ● ● ● ● ● ● ●
Liochlaena Nees (Jungermanniaceae)
lanceolata Nees LC ● ● ● ○ ● ● ● ● ● ○
subulata (A. Evans) Schljakov LC ● ● ● ● ● ●
Lophocolea (Dumort.) Dumort. (Lophocoleaceae)
bidentata (L.) Dumort LC ● ● ● ● ● ● ● ● ● ● ●
coadunata (Sw.) Mont. LC ● ● ● ● ● ● ● ● ● ●
heterophylla (Schrad.) Dumort. LC ● ● ● ● ● ● ● ● ● ● ●
minor Nees LC ○ ● ● ● ● ● ● ○ ● ○
Lophozia (Dumort.) Dumort. (Lophoziaceae)
ascendens (Warnst.) R. M. Schust. (17) EN ●
guttulata (Lindb. & Arnell) A. Evans LC ○ ○ ● ○
longiflora (Nees) Schiffn. (18) EN ●
ventricosa (Dicks.) Dumort. LC ● ● ● ● ● ○ ○
Lophoziopsis Konstant. & Vilnet (Lophoziaceae)
excisa (Dicks.) Konstant. & Vilnet (19) VU ? ● ●
polaris (R. M. Schust.) Konstant. & Vilnet. (20) ?
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Lunularia Adans. (Lunulariaceae)
cruciata (L.) Dumort. ex Lindb. LC ● ● ● ● ●
Mannia Corda (Aytoniaceae)
fragrans (Balb.) Frye. & L. Clark DD-1901 ○ ○ ○
gracilis (F. Weber) D. B. Schill & D. G. Long VU ● ● ○
triandra (Scop.) Grolle (21) LC ● ○ ○ ○ ○ ○
Marchantia L. (Marchantiaceae)
polymorpha L. subsp. polymorpha LC ● ● ● ● ● ●
subsp. montivagans Bischl. & Boissel.-Dub. LC ● ● ○ ●
subsp. ruderalis Bischl. & Boissel.-Dub. LC ● ● ○ ● ● ● ● ● ● ●
quadrata Scop. (22) LC ● ● ● ● ● ● ● ● ● ●
Marsupella Dumort. (Gymnomitriaceae)
apiculata Schiffn. (23) DD-n 2006
emarginata (Ehrh.) Dumort. LC ● ● ○ ○
funckii (F. Weber & D. Mohr) Dumort. LC ● ○ ○ ○ ○ ○ ○ ○
sprucei (Limpr.) Bernet DD-1908 ○
Mesoptychia (Lindb.) A. Evans (Jungermanniaceae)
badensis (Gottsche ex Rabenh.) L. Söderstr. & Váňa VU ○ ● ○ ○
bantriensis (Hook.) L. Söderstr. & Váňa VU ● ● ●
collaris (Nees) L. Söderstr. & Váňa LC ● ● ● ○ ● ● ● ● ● ●
heterocolpos (Thed. ex Hartm.) L. Söderstr. & Váňa LC ● ● ● ○ ● ● ●
turbinata (Raddi) L. Söderstr. & Váňa DD-1912 ○ ○ ○ ○ ○
Metzgeria Raddi (Metzgeriaceae)
conjugata Lindb. LC ● ● ● ● ● ● ● ● ● ● ●
furcata (L.) Corda LC ● ● ● ● ● ● ● ● ● ● ●
pubescens (Schrank) Raddi LC ● ● ● ● ● ● ● ●
simplex Lorb. ex Müll. Frib. EN ● ● ●
violacea (Ach.) Dumort. (24) DD-n 2006
Microlejeunea (Spruce) Steph. (Lejeuneaceae)
ulicina (Taylor) Steph. LC ● ● ● ●
Moerckia Gottsche (Moerckiaceae)
flotowiana (Nees) Schiffn. (25) DD-1905 ○
hibernica (Hook.) Gottsche DD-1912 ○ ○
Mylia Gray (Myliaceae)
anomala (Hook.) Gray LC ● ● ●
taylorii (Hook.) Gray LC ● ● ● ● ● ○ ●
Nardia Gray (Gymnomitriaceae)
geoscyphus (De Not.) Lindb. DD-1908 ○ ○
scalaris Gray LC ● ● ● ● ● ● ●
Neoorthocaulis L. Söderstr. (Anastrophyllaceae)
attenuatus (Mart.) L. Söderstr., De Roo & Hedd. LC ● ● ● ●
floerkei (F. Weber & D. Mohr) L. Söderstr., De Roo & Hedd. LC ● ● ● ○ ●
Nowellia Mitt. (Cephaloziaceae)
curvifolia (Dicks.) Mitt. LC ● ● ● ● ● ● ● ● ● ● ○
Odontoschisma (Dumort.) Dumort. (Cephaloziaceae)
denudatum (Mart.) Dumort. LC ● ● ● ○ ○ ● ● ● ●
fluitans (Nees) L. Söderstr. & Váňa EN ● ●
sphagni (Dicks.) Dumort. DD-1915 ○ ○ ○
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Pedinophyllum Lindb. ex Nordst. (Plagiochilaceae)
interruptum (Nees) Kaal. LC ● ● ● ● ● ● ● ● ● ● ●
Pellia Raddi (Pelliaceae)
epiphylla (L.) Corda LC ○ ● ● ● ● ● ● ● ● ●
neesiana (Gottsche) Limpr. LC ○ ● ● ● ● ● ● ●
Peltolepis Lindb. (Cleveaceae)
quadrata (Saut.) Müll. Frib. VU ● ●
Phaeoceros Prosk. (Notothyladaceae)
carolinianus (Michx.) Prosk. (26) DD-1884 ○
laevis (L.) Prosk. (27) DD-1908 ○ ○ ○ ○
Plagiochila (Dumort.) Dumort. (Plagiochilaceae)
asplenioides (L.) Dumort. LC ● ● ● ● ● ● ● ● ● ● ●
porelloides (Torr. ex Nees) Lindenb. LC ● ● ● ● ● ● ● ● ● ● ●
Porella L. (Porellaceae)
arboris-vitae (Vith.) Grolle LC ● ○ ○ ● ● ● ● ● ●
baueri (Schiff.) C. E. O. Jensen VU ● ● ●
cordaeana (Huebener) Moore LC ● ● ● ○ ● ● ● ●
obtusata (Taylor) Trevis. LC ● ● ●
platyphylla (L.) Pfeiff. LC ● ● ● ● ● ● ● ● ● ●
Protolophozia (R. M. Schust.) Schljakov (Cephaloziellaceae)
elongata (Steph.) Schljakov VU ● ● ○
Ptilidium Nees (Ptilidiaceae)
ciliare (L.) Hampe LC ● ● ○ ● ● ○ ● ○
pulcherrimum (Weber) Vain. LC ● ● ● ● ○ ● ● ● ○
Radula Dumort. (Radulaceae)
complanata (L.) Dumort. LC ● ● ● ● ● ● ● ● ● ● ●
lindenbergiana Gottsche ex C. Hartm. LC ● ● ● ○ ●
Reboulia Raddi (Aytoniaceae)
hemisphaerica (L.) Raddi LC ● ● ● ○ ● ● ○ ● ● ○
Riccardia Gray (Aneuraceae)
chamaedryfolia (With.) Grolle LC ● ● ● ● ● ● ● ●
latifrons (Lindb.) Lindb. LC ○ ● ○ ○ ○ ● ○ ● ●
multifida (L.) Gray LC ● ● ○ ● ● ● ● ● ○ ○
palmata (Hedw.) Carruth. LC ● ● ● ● ● ● ● ● ● ●
Riccia L. (Ricciaceae)
bifurca Hoffm. VU ● ● ●
canaliculata Hoffm. (28) EN ● ●
cavernosa Hoffm. (29) EN ●
ciliata Hoffm. (30) DD-1894 ○
fluitans L. NT ● ●
glauca L. LC ○ ○ ○ ○ ● ○ ○ ●
huebeneriana Lindenb. (31) EN ●
sorocarpa Bisch. VU ○ ○ ○ ● ● ●
Ricciocarpos Corda (Ricciaceae)
natans (L.) Corda (32) EN ●
Saccobasis H. Buch (Scapaniaceae)
polita (Nees) H. Buch (33) DD-1908 ○
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New checklist of the hornworts and liverworts of Slovenia
SLO RED LIST AJ AK AS AP AZ AM DN PA PD SM SP
Sauteria Nees (Cleveaceae)
alpina (Nees) Nees (34) LC ● ○ ○ ? ?
Scapania (Dumort.) Dumort. (Scapaniaceae)
aequiloba (Schwagr.) Dumort. LC ● ● ● ○ ○ ● ● ○ ● ○
apiculata Spruce (35) ?
aspera M. Bernet & Bernet LC ● ● ○ ○ ● ○ ● ○
calcicola (Arnell & J. Perss.) Ingham VU ● ● ●
compacta (Roth) Dumort. (36) DD-n 2007
crassiretis Bryhn (37) DD-1921 ○
curta (Mart.) Dumort. LC ● ● ● ○ ● ○ ○
cuspiduligera (Nees) Müll. Frib. VU ●
helvetica Gottsche LC ● ● ○ ○ ○
irrigua (Nees ( Nees LC ● ● ● ● ○ ●
nemorea (L.) Grolle LC ● ● ● ● ● ● ● ● ● ● ●
paludicola Loeske & Müll. Frib. (38) CR ●
subalpina (Nees ex Lindenb.) Dumort. LC ● ● ● ● ●
umbrosa (Schrad.) Dumort. LC ● ● ● ● ○ ●
undulata (L.) Dumort. LC ● ● ● ● ● ○ ○
verrucosa Heeg VU ● ● ○
Schistochilopsis (N. Kitag.) Konstant. (Scapaniaceae)
incisa (Schrad.) Konstant. LC ● ● ● ● ● ○ ●
Schljakovianthus Konstant. & Vilnet (Anastrophyllaceae)
quadrilobus (Lindb.) Konstant. & Vilnet VU ● ○ ● ●
Solenostoma Mitt. (Solenostomataceae)
confertissimum (Nees) Schljakov LC ● ● ○ ○
gracillimum (Sm.) R. M. Schust. LC ○ ● ● ● ● ● ● ● ●
hyalinum (Lyell) Mitt. LC ○ ● ○ ● ● ● ● ○
obovatum (Nees) C. Massal. NT ● ● ● ●
sphaerocarpum (Hook.) Steph. LC ● ● ● ○ ● ○ ● ●
subellipticum (Lindb. ex Heeg) R. M. Schust. DD-1908 ○
Southbya Spruce (Southbyaceae)
tophacea (Spruce) Spruce (39) EN ●
Sphenolobus (Anastrophyllaceae)
minutus (Schreb. ex D. Crantz) Berggr. LC ● ● ● ● ● ● ○
Syzygiella Spruce (Adelanthaceae)
 autumnalis (DC.) D. Feldberg, Váňa, Hentschel & Heinrichs LC ○ ● ○ ? ○
Trichocolea Dumort. (Trichocoleaceae)
tomentella (Ehrh.) Dumort. LC ● ● ● ● ● ● ● ● ●
Trilophozia (R. M. Schust.) Bakalin (Lophoziaceae)
quinquedentata (Huds.) Bakalin LC ● ● ● ● ○ ● ○ ●
Tritomaria Schiffn. ex Loeske (Lophoziaceae)
exsecta (Schmidel) Schiffn. ex Loeske LC ● ● ● ● ● ● ○
exsectiformis (Breidl.) Schiffn. ex Loeske EN ○ ● ○
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Annotations
1 Düll et al. (1999) suspect that we are actually dealing 
with species Anthoceros agrestis Paton. For the time be-
ing, however, this supposition cannot be verified as the-
re is no herbarium material available.
Düll et al. (1999) domnevajo, da gre v resnici za vrsto 
Anthoceros agrestis Paton. Domneve za sedaj ni mogoče 
preveriti, ker ni na voljo herbarijskega materiala.
2 The only recent locality is in AJ-Na Jezeru pod Rokavi 
(Martinčič, 2016b).
Edino recentno nahajališče je AJ-Na Jezeru pod Ro-
kavi (Martinčič, 2016b).
3 Only reported by Düll (1999) for the AJ-Možnica valley. 
Edina navedba je AJ-dolina Možnice (Düll, 1999).
4 Only reported for AM-valley of Strojnska reka river un-
der Zeleni breg (Martinčič, 2012).
Edina navedba je AM-dolina Strojnske reke pod Ze-
lenim bregom (Martinčič, 2012).
5 The only reliable locality is at DN-Rakov Škocjan: Zelške 
jame near Rakek (Sguazin & Polli, 1999). The locality 
reported by Schiffner (1906, leg. Loitlesberger) and later 
Pavletić (1955) for the banks of the river Soča near the 
town Gorica is currently situated within Italian territory.
Edino zanesljivo nahajališče je DN-Rakov Škocjan 
– Zelške jame pri Rakeku (Sguazin & Polli, 1999). 
Lokaliteta »ob Soči pri Gorici« (Schiffner, 1906, leg. 
Loitlesberger; Pavletić, 1955) je sedaj v Italiji.
6 The only recent locality is in PD-Kočevski Rog: Rajhe-
navski virgin forest (Ódor & van Dort, 2002).
Edino recentno nahajališče je PD-Kočevski Rog: Raj-
henavski pragozd (Ódor & van Dort, 2002).
7 The only site reported by Głowacki (1912) is at PA-
-Žovnek near the town of Braslovče. 
Edino nahajališče je v PA-Žovnek pri Braslovčah 
(Głowacki, 1912). 
8 The only site reported by Lobnik Cimerman (2022) is 
at AP-south of Lovrenška jezera lakes.
Edino nahajališče je v AP-južno od Lovrenških jezer 
(Lobnik Cimerman, 2022).
9 The species is reported only for Morež in the Julian Alps 
(Pokorny, 1860, leg. Sendtner 1841). This record was la-
ter considered also by Pavletić (1955), Düll et al. (1999), 
Söderström et al. (2002), Sabovljević & Natcheva (2006) 
and Ros et al. (2007). In Europe, this species is a Medi-
terranean element that only occasionally reaches further 
into Submediterranean regions, so the above mentioned 
locality in the subalpine or alpine belt is highly doubtful.
Vrsta je navedena samo za Morež v Julijskih Alpah 
(Pokorny, 1860, leg. Sendtner 1841). Podatek so upo-
števali še Pavletić (1955), Düll et al. (1999), Söder-
ström et al. (2002), Sabovljević & Natcheva (2006) in 
Ros et al. (2007). Ker je vrsta v Evropi mediteranski 
element, ki le tu in tam sega v submediteranske predele, 
je navedeno nahajališče v subalpinskem ali alpinskem 
pasu, močno dvomljivo. 
10 The species is reported only for the PA-marsh at the 
foothills of Šišenski hrib hill in Ljubljana. The locality 
was refered in collection »Cryptogamae exsiccatae« - 
editae a Museo Hist. Nat. Vindobonensi, No 3779 
(legit K. Loitlesberger, 1917).
Vrsta je navedena samo za močvirje na vznožju 
Šišenskega hriba v Ljubljani. Nahajališče je bilo ob-
javljeno v zbirki »Cryptogamae exsiccatae« - editae a 
Museo Hist. Nat. Vindobonensi, No. 3779 (legit. K. 
Loitlesberger, 1917).
11 The only locality is SM-Steske at village Dornberk 
(Martinčič, 2007). The species is reported for Slovenia 
also by Sabovljević & Natcheva (2006) and Ros et al. 
(2007) – without stated locality or source.
Edino nahajališče so SM-Steske pri Dornberku 
(Martinčič, 2007). Vrsto navajajo za Slovenijo tudi 
Sabovljević & Natcheva (2006) in Ros et al. (2007); v 
obeh primerih brez vira ali lokalitete.
12 The species is reported from Slovenia by Sabovljević 
& Natcheva (2006) – without locality or source. As 
this species is a West-Mediterranean-Atlantic element 
the above mentioned report is doubtfull.
Vrsto navajata za Slovenijo Sabovljević & Natcheva 
(2006), vendar gre le za splošno navedbo brez vira ali 
lokalitete. Ker je vrsta zahodnomediteransko-atlantski 
element, je navedba dvomljiva.
13 Only reported by Martinčič (2017) for DN-Kobilji 
curek at settlement Rob in vicinity of Velike Lašče.
Navedena samo za DN-Kobilji curek pri Robu pri 
Velikih Laščah (Martinčič, 2017).
14 The only locality is in the mountain Pohorje – at the 
stream Plešiščica above settlement Lovrenc na Pohor-
ju (Breidler, 1894).
Edino nahajališče vrste je na Pohorju – graben potoka 
Plešiščica nad Lovrencem na Pohorju (Breidler, 1894).
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New checklist of the hornworts and liverworts of Slovenia
15 The only locality is in the mountain Pohorje – at the 
stream Plešiščica above settlement Lovrenc na Pohor-
ju (Breidler, 1894).
Edino nahajališče vrste je na Pohorju – graben po-
toka Plešiščica nad Lovrencem na Pohorju (Breidler, 
1894).
16 In his manuscript list of mosses collected in the vici-
nity of Bohinj and Bled, Wallace (1980) refers also to 
L. patens. However, as it is an Atlantic-Mediterranean 
element, its occurence in the phytogeographical re-
gion Julian Alps is doubtful.
Wallace (1980) navaja v rokopisnem seznamu ma-
hov, ki jih je nabiral v okolici Bohinja in Bleda, tudi 
vrsto L. patens. Ker je vrsta atlantsko-mediteranski 
element, je njeno nastopanje v fitogeografskem ob-
močju Julijske Alpe zelo dvomljivo. Verjetno gre za 
vrsto L. cavifolia.
17 The only locality is on the AK-mountain Olševa, north 
slope (leg. R. Čas, det. A. Martinčič, LJU, 2003) and 
reported for AK-Olševa, Zadnji travnik by Ž. Lobnik 
Cimerman (In Strgulc Krajšek S. (ed.) et al., 2023).
Edino nahajališče vrste je v Karavankah, na Olševi, 
severno pobočje (Martinčič, LJU, 2003), navedena 
tudi za AK-Olševa, pri Zadnjem travniku (Lobnik 
Cimerman Ž. v Strgulc Krajšek S. (ed.) et al., 2023).
18 The only locality is on the AS-mountain Veliki Trav-
nik (Martinčič, LJU, 1996).
Edino nahajališče vrste je v Kamniško-Savinjskih 
Alpah: VK. Travnik v Smrekovškem pogorju (Martin-
čič, LJU, 1996).
 19 Loitlesberger (1905) reported for Mangart (2100 m 
s. m.) Lophozia excisa as »eine ihr nahestehende Pflan-
ze«, but after Głowacki (1910) this record is doubtful.
Loitlesberger (1905) navaja vrsto za Mangart s 
pripombo »eine ihr nahestehende Pflanze«, vendar 
Głowacki (1910) dvomi v pravilnost podatka.
20 The species is reported from Slovenia by Sabovljević 
& Natcheva (2006) and Ros et al. (2007) – without 
locality or source. As this species is an Arctic element, 
not found in Europe outside the Arctic, above men-
tioned report is highly improbable.
Vrsto navajajo za Slovenijo Sabovljević & Natcheva 
(2006) in Ros et al. (2007), vendar gre le za splošno 
navedbo, brez vira ali lokalitete. Ker je vrsta arktični 
element, ki v Evropi ni bila najdena južno od ark-
tičnih predelov, je navedba povsem neverjetna.
21 The species was described by Scopoli: in the first edi-
tion of the renowned Flora carniolica (1760) with a 
descriptive name »Marchantia pileo hemisphaerica 
integro triloculari«, and in the second edition (1772) 
already with the binary name of »Marchantia trian-
dra«. Its locus classicus is situated in Slovenia between 
towns Idrija and Spodnja Idrija.
Vrsto je opisal Scopoli: v 1. izdaji znamenite Flora 
Carniolica (1760) še z opisnim imenom »Marchantia 
pileo hemisphaerica integro triloculari«, v drugi izdaji 
(1772) pa že z binarnim imenom »Marchantia trian-
dra«. Locus classicus je med Idrijo in Spodnjo Idrijo.
22 The species was described by Scopoli: in the first edi-
tion of the renowned Flora carniolica (1760) with the 
descriptive name »Marchantia pileo hemisphaerico, 
semiquadrifido, quadriloculari«, in the second edi-
tion (1772) already with the binary name »Marchan-
tia quadrata«.
Vrsto je opisal Scopoli: v 1. izdaji znamenite Flora 
carniolica (1760) še z opisnim imenom »Marchantia 
pileo hemisphaerico, semiquadrifido, quadriloculari«, 
v drugi izdaji (1772) pa že z binarnim imenom »Mar-
chantia quadrata«.
23 The species is reported from Slovenia by Sabovljević 
& Natcheva (2006) and Ros et al. (2007) – without 
locality or source.
Vrsto navajajo za Slovenijo Sabovljević & Natcheva 
(2006) in Ros et al. (2007), v obeh primerih pa gre le 
za splošno navedbo.
24 The species is reported from Slovenia by Sabovljević 
& Natcheva (2006) and Ros et al. (2007) – without 
locality or source.
Vrsto navajajo za Slovenijo Sabovljević & Natcheva 
(2006) in Ros et al. (2007), v obeh primerih gre za 
splošno navedbo, brez vira ali lokalitete.
25 Only reported by Loitlesberger ( 1905) for DN-Tr-
novski gozd: Paradana (Paradanastrasse)
Navedena samo za DN-Trnovski gozd: ob cesti v 
Paradano (Loitlesberger, 1905).
26 The only site is in PD-Debeli hrib at Lavrica – vicini-
ty of town Ljubljana (LJU, A. Paulin sub Anthoceros 
laevis, 1884; det. Martinčič: Phaeoceros laevis subsp. 
carolinianus; Kerner: Schedae ad floram exsiccatam 
austro-hungaricam No.1933, 1888, sub Anthoceros 
laevis, leg. A. Paulin).
Edino nahajališče je v PD-Debeli hrib nad Lavrico 
blizu Ljubljane (LJU, A. Paulin kot Anthoceros laevis, 
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New checklist of the hornworts and liverworts of Slovenia
1884; det. Martinčič – Phaeoceros laevis subsp. caroli-
nianus; Kerner: Schedae ad floram exsiccatam austro-
-hungaricam No. 1933, 1888, kot Anthoceros laevis, 
leg. A. Paulin).
27 On the base of literature data only (Robič, 1893; 
Breidler, 1894; Głowacki, 1908; Paulin mscr.), wit-
hout herbarium specimens, it is not possible to de-
cide, if this records belong to Phaeoceros laevis or to 
P. carolinianus?
Samo na podlagi navedb v literaturi (Robič, 1893; 
Breidler, 1894; Głowacki, 1908; Paulin mscr.), brez 
herbarijskega materiala, ni mogoče opredeliti, ali ti 
stari podatki pripadajo vrsti Phaeoceros laevis ali P. ca-
rolinianus?
28 Recently only reported for PA-Šmartno pod Šmarno 
goro in the vicinity of town Ljubljana at river Gračeni-
ca (Strgulc Krajšek & Lobnik Cimerman, 2021) and 
for SP-Radvanje in the vicinity of town Maribor (Lob-
nik Cimerman in Strgulc Krajšek (ed.) et al., 2023).
Edini recentni navedbi sta za PA-Šmartno pod 
Šmarno Goro, ob Gračenici (Strgulc Krajšek & Lob-
nik Cimerman, 2021) in za SP-Radvanje pri Maribo-
ru, na vznožju Pohorja (Lobnik Cimerman v Strgulc 
Krajšek (ed.) et al., 2023).
29 Only reported by Lobnik Cimerman & Strgulc Kraj-
šek (2022) for SP-Zrkovci at town Maribor and by 
Lobnik Cimerman et al. (2023) for SP-Prekmurje: 
Beltinci, S from the settlement Dokležovje.
Edini navedbi sta za SP-Zrkovci pri Mariboru 
(Lobnik Cimerman & Strgulc Krajšek, (2022) in za 
SP-Beltinci, južno od Dokležovja (Lobnik Cimerman 
et al., 2023).
30 Only reported by Breidler (1894) for AZ-Kozjak: 
Srednje above settlement Brestrnica.
Edina navedba je za AZ-Kozjak: Srednje nad Brestr-
nico (Breidler, 1894).
31 The only locality is on SP-right bank of river Mura, 
north of settlement Mota (Trčak, herb., 2013).
Edino nahajališče je v SP-desni breg reke Mure, se-
verno od naselja Mota (Trčak, herb., 2013).
32 Only reported by Babij & Jogan (2001) for SP-Pre-
kmurje: near the village Mali Bakovci and by Lobnik 
& Šabeder (in Martinčič (ed.) et al., 2020) for SP-
-Prekmurje: west of village Petišovci.
Vrsta je navedena samo za SP-Prekmurje: zahodno 
od vasi Mali Bakovci, v opuščeni gramoznici (Babij & 
Jogan, 2001) in zahodno od vasi Petišovci, v mrtvici 
reke Mure (Lobnik Cimerman & Šabeder v Martinčič 
(ed.) et al., 2020).
33 Only reported by Głowacki (1908) for Pohorje Mts.: 
Rakovec above settlement Vitanje
Vrsta je navedena samo za Pohorje: Rakovec nad 
Vitanjem (Głowacki, 1908).
34 Both data outside the Alpine phytogeographical re-
gion are questionable. The first (PA: Šklendovec un-
der Kum – Paulin mscr.) oving to the very low altitude 
of the locality at ca. 500 m s. m. – all other localities 
are in the subalpine and alpine belt. The second data 
(DN: the Trnovski gozd plateau, Smrekova draga, 
1100 m – Grom, 1969) is questionable due to the 
uncertainty of author in determination and absence 
of herbarium specimens.
Oba podatka zunaj alpskega fitogeografskega ob-
močja sta dvomljiva. Prvi (PA: Šklendrovec pod 
Kumom – Paulin mscr.) zaradi zelo nizke nadmorske 
višine nahajališča, ok. 500 m) – vsa druga nahajališča 
so v subalpinskem ali alpinskem pasu. Drugi poda-
tek (DN: Trnovski gozd, Smrekova draga, 1100 m – 
Grom, 1969) je dvomljiv zaradi nezanesljivosti avtorja 
pri določanju, manjkajo pa tudi herbarijski primerki.
35 Reported only for DN: Trnovski gozd – Mrzla Draga 
near the village of Lokve by Grom (1969). The report 
is to be considered uncertain since it is not supported 
by herbarium specimens.
Navedena samo za DN: Trnovski gozd, Mrzla Dra-
ga pri Lokvah (Grom, 1969). Podatek je dvomljiv, ker 
ni herbarijskega materiala.
36 The species is reported from Slovenia by Söderström 
et al. (2002), Sabovljević & Natcheva (2006) and Ros 
et al. (2007) – without locality or source.
Vrsto navajajo za Slovenijo Söderström et al. 
(2002), Sabovljević & Natcheva (2006) in Ros et al. 
(2007) – v vseh primerih gre za splošno navedbo, brez 
vira ali lokalitete.
37 The only locality is in the Julian Alps: The Kot valley 
(LJU – leg. F. Dolšak, 1921, det. A. Martinčič). The 
reports by Grom (1963) for AJ-Pokljuka and DN-Tr-
novski gozd were excluded since they were erroneo-
usly identified.
Edino nahajališče je v Julijskih Alpah: dolina Kot 
(LJU – leg. F. Dolšak 1921, det. A. Martinčič). Na-
vedbe Groma (1963) za Pokljuko in Trnovski gozd 
temeljijo na napačni determinaciji.
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38 Only reported by Martinčič (2017) for Pohorje Mts.-
Stegnetovo močvirje.
Navedena samo za Pohorje-Stegnetovo močvirje 
(Martinčič, 2017).
39 In this year the species was collected the first time in 
Slovenia, on the Littoral (Simonov zaliv near Izola – 
pers. comm. by Aljaž Jakob).
Letos je bila vrsta prvič najdena v Sloveniji, na Oba-
li (Simonov zaliv pri Izoli - osebno sporočilo Aljaža 
Jakoba).
Excluded Taxa
Mannia pilosa (Hornem.) Frye & Clark
Pavletić (1955) reports that the species thrives in Slove-
nia as well, i.e. in the Julian Alps. This reference is based 
on F. Kern (1910) and was later taken up by Düll et al. 
(1999), Söderström et al. (2002), Sabovljević & Natcheva 
(2006) and Ros et al. (2007). Today, however, the locali-
ty – Mt. Viš (Iof Fuart) in the Western Julian Alps – is 
situated in Italy.
Pavletić (1955) navaja, da uspeva vrsta tudi v Sloveniji, 
v Julijskih Alpah. Navedba temelji na podatku F. Kerna 
(1910). Temu so sledili kasneje tudi Düll et al. (1999), 
Söderström et al. (2002), Sabovljević & Natcheva (2006) 
ter Ros et al. (2007). Vendar je nahajališče – gora Viš (Iof 
Fuart) v Zahodnih Julijskih Alpah, sedaj v Italiji.
Marchantia paleacea Bertol.
Based on Loitlesberger‘s reference (1905), Pavletić re-
ports the species also for Slovenia. This was later taken 
up by Düll et al. (1999), Söderström et al. (2002), Sa-
bovljević & Natcheva (2006) and Ros et al. (2007). Ne-
vertheless, its only locality – Stračice in Gorizia – is today 
situated in Italy (comp. Martinčič, 2007).
Na podlagi navedbe Loitlesbergerja (1905) navaja Pav-
letić (1955) vrsto tudi za Slovenijo. Temu so sledili kasne-
je tudi Düll et al. (1999), Söderström et al. (2002), Sabo-
vljević & Natcheva (2006) in Ros et al. (2007). Vendar je 
edino nahajališče, Stračice v Gorici, sedaj v Italiji (prim. 
Martinčič, 2007).
Fuscocephaloziopsis albescens (Hook.) Váňa & Söderstr.
The only report by Grom (1963) for DN-Trnovski 
gozd, Mrzla Draga near the village of Lokve is based on 
erroneous determination.
Edini podatek daje Grom (1963) za DN-Trnovski 
gozd, Mrzla Draga pri Lokvah, vendar temelji na napačni 
določitvi.
Riccia bicarinata Lindb.
Based on reference in Bischler & Jovet-Ast (1973), Sö-
derström et al. (2002) reported the species from Slovenia 
as well. This was later taken up by Sabovljević & Natche-
va (2006) and Ros et al. (2007). Nevertheless, the locality 
i.e. Valica in Istria, is situated in Croatia.
Na podlagi navedbe v Bischler & Jovet-Ast (1973) 
navajajo Söderström et al. (2002) vrsto tudi za Sloveni-
jo. Temu so sledili kasneje tudi Sabovljević & Natcheva 
(2006) in Ros et al. (2007). Vendar je nahajališče, Istra: 
Valica v Hrvaški. 
Southbya nigrella (De Not.) Henriq.
Based on Loitlesberger,s reference (1905), Pavletić 
(1955) reports species from Slovenia as well. This was 
later taken up by Düll et al. (1999), Söderström et al. 
(2002), Sabovljević & Natcheva (2006) and Ros et al. 
(2007). However, its only locality – »along the Soča River 
near the town Gorizia« is today situated in Italy.
Na podlagi navedbe Loitlesbergerja (1905) navaja Pav-
letić (1955) vrsto tudi za Slovenijo. Temu so sledili kas-
neje Düll et al. (1999), Söderström et al. (2002), Sabo-
vljević & Natcheva (2006) in Ros et al. (2007). Vendar 
je edino nahajališče »vzdolž reke Soče pri Gorici« sedaj 
v Italiji.
Targionia hypophylla L. 
Pavletić (1955) reports the species for Slovenia, altho-
ugh without any specific locality, but only with a general 
description »along the Austrian border«. This description 
is based on a misinterpretation in »Die Lebermoose Ste-
iermarks« (Breidler, 1894). In this work, Breidler does 
not refer to the species as belonging to the territory of 
the former Styria, but merely notes that among other 
species, Targionia hypophylla could also thrive in these 
parts. This incorrect reference was later taken up by Sö-
derström et al. (2002), Sabovljević & Natcheva (2006) 
and Ros et al. (2007).
Za Slovenijo navaja vrsto Pavletić (1955), vendar brez 
točnega nahajališča, le s splošno oznako »vzdolž avstrij-
ske meje«. Navedba temelji na napačnem razumevanju 
pripombe v delu »Die Lebermoose Steiermarks« (Breid-
ler, 1894). Breidler v tem delu ne navaja vrste za ozeml-
je nekdanje Štajerske, temveč le pripominja, da bi med 
drugimi vrstami tudi Targionia hypophylla lahko uspevala 
v teh predelih. Tej napačni navedbi so sledili kasneje Sö-
derström et al. (2002), Sabovljević & Natcheva (2006) in 
Ros et al. (2007).
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Discussion
The New Checklist
Based on the current taxonomy and nomenclature 
(Hodgetts et al., 2020), the New Checklist of hornworts 
(Anthocerotophyta) and liverworts (Marchantiophyta) of 
Slovenia comprises 181 species, 4 additional subspecies 
and 2 varieties, which is an increase compared to the pre-
vious List (Martinčič, 2011). A significant increase is also 
evident in the number of species in individual geograph-
ic-phytogeographic units (Table 2).
Table 2: Number of taxa by geographic-phytogeographic units 
of Slovenia
Tabela 2: Število taksonov v posameznih geografsko-fitogeo-
grafskih enotah Slovenije
SLO AJ AK AS AP AZ AM DN PA PD SM SP
spp 181 130 101 111 114 68 57 108 111 83 56 65
subsp 4 3 3 3 1 1 1 1 1 1 1 1
var 2 1 1 1 2 1 1 1
? 5
The differences in the number of species between ge-
ographic-phytogeographic units are relatively large. This 
is partly associated with differences in the intensity of 
floristic research in particular areas and, even more im-
portantly, with the ecological diversity of habitats and el-
evation ranges, which are the most notable in the Alpine 
region. Compared with other European countries, the 
total number of taxa indicates that the bryophyte flora 
of Slovenia is relatively rich despite its small territory 
(20,273 km2). On the other hand, there are 19 (10.5%) 
species with a single known locality, half found in the 
period before 1915. This year marks the end of the first 
intensive research of the bryophyte flora of Slovenia, fol-
lowed only by a few short papers between 1920 and 1945. 
Not earlier than after 1958, the research of the bryophyte 
flora has started up again.
Red List 
Under different threat categories, 38 (20.99%) spe-
cies and subspecies are listed: 1 (0.5%) of them CR, 14 
(7.7%) as EN and 23 (12.7%) as VU. Furthermore, 3 
(1.6%) species are listed as near threatened (NT), while 
33 (18.2%) species are data deficient (DD and DD-n). 
There are currently 111 (61.3%) species and subspecies in 
the Least Concern (LC) category. The number and per-
centage of species in the various threat categories in the 
Slovenian Red List are not entirely comparable with some 
other countries, although the IUCN 3.1 (2001) and Hal-
lingbäck et al. (1998) serve as the basis for categorisation. 
The main reason is that authors tend to adapt these crite-
ria, to some extent, to the specific situation in their coun-
tries, especially regarding the number of recently reported 
localities. Floristic research available for a particular coun-
try also has an impact on the species categorisation. 
Povzetek
Uvod
Scopolijevo delo z naslovom »Flora carniolica« (1. izda-
ja 1760, 2. izdaja 1772) označuje začetek raziskovanja 
mahovne flore v Sloveniji. Sledila so številna dela za po-
samezne predele, pa tudi za celotno ozemlje Slovenije. 
Po trinajstih letih od zadnjega dela »Seznam jetrenjakov 
(Marchantiophyta) in rogovnjakov (Athocerotophyta) 
Slovenije (Martinčič, 2011)« je napočil čas za nov, po-
sodobljen Seznam, saj je v tem obdobju močno naraslo 
število florističnih podatkov. Drugi vzrok za Novi poso-
dobljen Seznam pa so velike taksonomske in nomenkla-
turne spremembe v evropski mahovni flori (Hodgetts & 
Lockhart, 2020; Hodgetts et al., 2020).
Metode
Novi Seznam temelji na podatkih iz literature, objavljene 
do leta 2024, herbarijske zbirke LJU ter avtorjevih neob-
javljenih podatkov. Upoštevali smo vsa floristična dela 
in tisti del objavljenih fitocenoloških popisov, za katere 
smo menili, da so bili v njih mahovi pravilno določeni. 
V starejši floristični literaturi smo kritično ovrednotili 
tiste podatke, ki se nanašajo na mejne predele med Slo-
venijo in Italijo. Po razpadu Avstroogrske monarhije je 
del omenjenega ozemlja pripadel Italiji, podatki iz teh 
predelov (predvsem okolica Gorice in Trsta) pa so obja-
vljeni tako pri Pavletiču (1955) kakor tudi v Düll et al. 
(1999). Revizija točnosti vseh navedb v literaturi sicer ni 
bila mogoča, ker nam niso bili dostopni ustrezni herbariji. 
Izjema je material v Herbariju LJU. V zadnjih desetletjih 
19. stoletja so ga prispevali v glavnem S. Robič, J. Šafer, 
J. Breidler in A. Paulin. Material prvih dveh sta revidi-
rala oz. nanovo določila J. Breidler in J. Głowacki. Na 
začetku 20. stoletja je nabiral mahovni material F. Dolšak. 
Herbarijski material v LJU nam je omogočil tudi, da smo 
opravili revizijo enega dela literaturnih podatkov amater-
skega briologa S. Groma, ki je deloval v letih 1958–1969. 
Navedbe, ki so iz kakršnega koli vzroka dvomljive, ven-
dar ni herbarijskega materiala za revizijo, smo v pregledu 
označili z vprašajem. En del florističnih podatkov v lit-
eraturi izvira iz obdobja pred letom 1915, drugi del pa 
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Andrej Martinčič
New checklist of the hornworts and liverworts of Slovenia
iz obdobja 1958–2023. Herbarijski material v LJU je bil 
nabran zlasti po letu 1960, največji del je prispeval avtor 
tega prispevka.
Nomenklatura in taksonomija, uporabljeni v Novem 
Seznamu, sledita delu Hodgetts et al. (2020). 
Za prikaz razširjenosti posameznih taksonov smo up-
orabili fitogeografsko razdelitev Slovenije po M. Wraberju 
(1969), a smo jo nekoliko spremenili (Martinčič, 2003). 
Tako smo srednje Posočje izločili iz submediteranskega 
območja, kot je predlagal že Dakskobler (1996) in ga 
priključili predalpskemu območju. Prav tako pa smo raz-
delili enotno podobmočje alpskega območja – Pohorje, 
v tri dele: Pohorje v ožjem pomenu, Dravski Kozjak – 
ozemlje severno od reke Drave in Mežiško-Mislinjska 
dolina s Strojno – zahodno od Pohorja. Razpored enot je 
razviden na sliki 1, kratice pa so naslednje:
AJ – alpsko območje: Julijske Alpe
AK – alpsko območje: Karavanke
AS – alpsko območje: Kamniško-Savinjske Alpe
AP – alpsko območje: Pohorje
AM – alpsko območje: Mežiško-Mislinjska dolina-Strojna 
AZ – alpsko območje: Dravski Kozjak
DN – dinarsko območje
PA – predalpsko območje
PD – preddinarsko območje
SM – submediteransko območje
SP – subpanonsko območje
Vsakemu taksonu je dodana še oznaka ogroženosti, 
čeprav temelji v največji meri na Rdečem seznamu, ki je že 
bil objavljen (Martinčič, 2016a). Za opredelitev pripad-
nosti taksonov posameznim kategorijam smo upoštevali 
IUCN kriterije 3.1 (IUCN 2001). Njihovo uporabo za 
mahove smo povzeli po Hallingbäck et al. (1998). Vendar 
smo mnenja, da je v našem primeru edini realni kriterij za 
oceno ogroženosti – kriterij B, ki se izraža s številom tre-
nutno poznanih recentnih nahajališč: 1 (CR), 2-5 (EN), 
6-10 (VU). Pri taksonih iz kategorije DD je dodana tudi 
letnica zadnjega podatka.
V Rdečem seznamu smo uporabili naslednje kategorije: 
Regionalno izumrla vrsta (RE)
Skrajno ogrožena vrsta (CR)
Prizadeta vrsta (EN)
Ranljiva vrsta (VU)
Potencialno ogrožena vrsta (NT)
Premalo znana vrsta, stari podatki (DD)
Premalo znana vrsta, recentni podatki (DD-n)
Neogrožena vrsta (LC)
Rezultati
Novi Seznam
Novi Seznam rogovnjakov (Anthocerotophyta) in jetren-
jakov (Marchantiophyta) Slovenije obsega na podlagi 
trenutne taksonomije in nomenklature (Hodgetts et al., 
2020) 181 vrst, nadalje 4 podvrste in 2 varieteti. Dvom-
ljivih vrst, ki so bile navedene v literaturi in podatkov ni 
mogoče revidirati, ker ni herbarijskega materiala, je 5. 
Izločenih vrst je 6. Razpored po posameznih fitogeograf-
skih enotah je razviden v tabeli 1.
Razlike v številu vrst med posameznimi fitogeografs-
kimi enotami so razmeroma velike (Tabela 1). Deloma je 
to povezano z razlikami v intenzivnosti florističnih razis-
kovanj posameznih predelov. Predvsem pa so razlike, tako 
kot pri listnatih mahovih, pogojene z ekološko pestrostjo 
habitatov, vključno z razponom nadmorskih višin, kar je 
najbolj razvidno v alpskem območju. Primerjava števila 
taksonov z nekaterimi sosednjimi državami kaže, da 
je mahovna flora Slovenije kljub površinski majhnosti 
(20.273 km2) relativno bogata.
Rdeči seznam
V kategorije, ki kažejo na ogroženost taksonov, je uvrščeno 
37 (20,1%) vrst in podvrst: 1 (0,5%) vrsta je v CR kat-
egoriji, 13 (7,1%) vrst je v kategoriji EN in 23 (12,5%) 
v kategoriji VU. 3 (1,6%) vrste so v kategoriji NT, 33 
(17,9%) vrst je v kategorijah DD in DD-n. V kategori-
ji LC-neogrožene vrste je 111 (60,3%) vrst in podvrst. 
Število vrst in procentna razmerja v kategorijah Rdečega 
seznama Slovenije niso enostavno primerljive z različnimi 
drugimi državami – čeprav je tudi pri njih podlaga za kat-
egorizacijo IUCN 3.1 in delo Hallingbäck et al. (1998). 
Razlog za to je, da avtorji prilagajajo kriterije do določene 
mere specifičnim razmerah v svojih državah. Vpliv pa 
imata tudi stopnja floristične raziskanosti in ohranjenost 
biotopov.
Synonyms 
Alicularia minor (Nees) Limpr = Nardia geoscyphus
Alicularia scalaris (Schrad.) Corda = Nardia scalaris
Anastrophyllum cavifolium (H. Buch et S. W. Arnell) La-
mes = Orthocaulis cavifolium
Anastrophyllum hellerianum (Nees ex Lindenb.) R. M. 
Schust. = Crossocalyx hellerianus
Anastrophyllum minutum (Schreb.) R. M. Schust. = Sphe-
nolobus minutus
Anastrophyllum minutum var. weberi (Mart.) Kartt. = 
Sphenolobus minutus
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New checklist of the hornworts and liverworts of Slovenia
Aneura latifrons Lindb. = Riccardia latifrons
Aneura multifida (L.) Dumort. = Riccardia multifida
Aneura palmata (Hedw.) Dumort. = Riccardia palmata
Anthoceros laevis L. = Phaeoceros laevis
Aplozia amplexicaulis (Dumort.) Dumort. = Solenostoma 
sphaerocarpum
Aplozia autumnalis (DC.) Heeg = Syzygiella autumnalis
Aplozia crenulata (Mitt.) Lindb. = Solenostoma gracillimum
Aplozia lanceolata auct. = Liochlaena lanceolata
Aplozia lurida (Dumort.) Dumort. = Solenostoma sphae-
rocarpum
Aplozia nana (Nees) Breidl. var. confertissima (Nees) Heeg 
= Solenostoma confertissimum
Aplozia pumila (With.) Dumort. = Jungermannia pumila
Aplozia riparia (Tayl.) Dumort. = Jungermannia atrovirens
Aplozia riparia var. potamophila Müller Arg. ex Bernet = 
Jungermannia atrovirens
Aplozia sphaerocarpa (Hook.) Dumort. = Solenostoma 
sphaerocarpum
Apometzgeria pubescens (Schrank.) Kuwah. = Metzgeria 
pubescens
Asterella gracilis (F. Weber) Underw. = Mannia gracilis
Asterella hemisphaerica (L.) P. Beauv. = Reboulia hemispha-
erica
Athalamia hyalina (Sommerf.) S. Hatt. = Clevea hyalina
Athalamia hyalina var. suecica (Lindb.) S. Hatt. = Clevea 
hyalina
Barbilophozia attenuata (Mart.) Loeske = Neoorthocaulis 
attenuatus
Barbilophozia gracilis (Schleich. ex Steph.) Müll. Frib. = 
Neoorthocaulis attenuatus
Barbilophozia floerkei (F. Weber & D. Mohr) Loeske = 
Neoorthocaulis floerkei
Barbilophozia quadriloba (Lindb.) Loeske = Schljako-
vianthus quadrilobus
Bazzania triangularis (Schleich. ex Steudel) Loeske = 
Bazzania tricrenata
Bellincinia laevigata O. Kuntze = Porella arboris-vitae
Bellincinia platyphylla O. Kuntze = Porella platyphylla
Bellincinia rivularis O. Kuntze = Porella cordaeana
Blepharozia ciliaris (L.) Dumort. = Ptilidium ciliare
Blepharozia pulcherrima (G. Weber) Lindb. = Ptilidium 
pulcherrimum
Cephalozia bicuspidata var. alpicola (C. Massal.) C. Mas-
sal. & Carestia = Fuscocephaloziopsis pleniceps
Cephalozia catenulata (Huebener) Lindb. = Fuscocephalo-
ziopsis catenulata
Cephalozia connivens (Dicks.) Lindb. = Fuscocephaloziop-
sis connivens
Cephalozia divaricata (Sm.) Spruce = Cephaloziella diva-
ricata
Cephalozia integerrima Lindb. = Cephaloziella integerrima
Cephalozia jackii Limpr. = Cephaloziella rubella
Cephalozia leucantha Spruce = Fuscocephaloziopsis leucan-
tha
Cephalozia loitlesbergeri Schiffn. = Fuscocephaloziopsis loit-
lesbergeri
Cephalozia lunulifolia (Dumort.) Dumort. = Fuscocepha-
loziopsis lunulifolia
Cephalozia media Lindb. = Fuscocephaloziopsis lunulifolia
Cephalozia myriantha Lindb. = Cephaloziella rubella
Cephalozia pleniceps (Austin) Lindb. = Fuscocephaloziopsis 
pleniceps
Cephalozia reclusa (Tayl.) Dumort. = Fuscocephaloziopsis 
catenulata
Cephalozia starkei (Nees) Breidl. = Cephaloziella divari-
cata
Cephalozia symbolica (Gottsche) Breidl. = Fuscocephalo-
ziopsis lunulifolia
Cephaloziella bifida (Lindb.) Schiffn. = Cephaloziella ru-
bella
Cephaloziella byssacea (Roth) Warnst. = Cephaloziella 
hampeana
Cephaloziella leucantha Spruce = Fuscocephaloziopsis leu-
cantha
Cephaloziella myriantha (Lindb.) Schiffn. = Cephaloziella 
rubella
Cephaloziella raddiana (C. Massal.) Schiffn. = Cephalo-
ziella rubella
Chiloscyphus coadunatus (Sw.) J. J. Engel & R. M. Schust. 
= Lophocolea coadunata
Chiloscyphus coadunatus var. rivularis Loeske = Lophocolea 
bidentata
Chiloscyphus minor (Nees) J. J. Engel & R. M. Schust. = 
Lophocolea minor
Chiloscyphus polyanthos var. pallescens (Ehrh. ex Hoffm.) 
C. Hartm. = Chiloscyphus pallescens
Chiloscyphus profundus (Nees) J. J. Engel & R. M. Schust. 
= Lophocolea heterophylla
Cladopodiella fluitans (Nees) H. Buch = Odontoschisma 
fluitans
Corsinia marchantioides Raddi = Corsinia coriandrina
Diplophylleia albicans (L.) Trev. = Diplophyllum albicans
Diplophylleia obtusifolia (Hook.) Trev. = Diplophyllum 
obtusifolium
Diplophylleia taxifolia (Wahlenb.) Trev. = Diplophyllum 
taxifolium
Duvalia rupestris Nees = Mannia triandra
Eucalyx hyalinus (Lyell) Carrington = Solenostoma hyalinum
Eucalyx subellipticus (Lindb. ex Kaal.) Breidl. = Solenosto-
ma subellipticum
Eucephalozia media Schiffn. = Fuscocephaloziopsis lunuli-
folia
Fimbriaria lindenbergiana Corda = Asterella lindenbergiana
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New checklist of the hornworts and liverworts of Slovenia
Fossombronia cristata Lindb. = Fossombronia wondraczekii
Fossombronia dumortieri Hueben. et Genth. ex Lindb. = 
Fossombronia foveolata
Frullania cesatiana De Not. = Frullania riparia
Frullania inflata Gottsche = Frullania cleistostoma
Grimaldia barbifrons Bischl. = Mannia fragrans
Grimaldia triandra (Scop.) Lindb. = Mannia triandra
Haplozia crenulata (Sm.) Dumort. = Solenostoma gracil-
limum
Haplozia lanceolata (Schrad.) Dumort. = Liochlaena lan-
ceolata
Haplozia pumila (With.) Dumort. = Jungermannia pumila
Haplozia schiffneri (Loitl.) K.M. = Jungermannia polaris
Haplozia sphaerocarpa (Hook.) Dumort. = Solenostoma 
sphaerocarpum
Haplozia sphaerocarpa var. nana (Nees) K. Müll. = Soleno-
stoma sphaerocarpum
Hypenantron lindenbergianum (Corda) Głow. = Asterella 
lindenbergiana
Hypenantron pilosum O. Kuntze = Mannia gracilis
Jamesoniella autumnalis (DC.) Steph. = Syzygiella autum-
nalis
Jungermannia acuta Lindenb. = Mesoptychia collaris
Jungermannia alpestris Schleich. ex F. Weber = Lophozia 
collaris
Jungermannia badensis Gottsche = Mesoptychia badensis
Jungermannia bantriensis Hook. = Mesoptychia bantriensis
Jungermannia barbata Schmidel ex Schreb. = Barbilopho-
zia barbata
Jungermannia bicrenata Schmidel ex Hoffm. = Isopaches 
bicrenatus
Jungermannia bicuspidata L. = Cephalozia bicuspidata
Jungermannia catenulata Hueben. = Fuscocephaloziopsis 
catenulata
Jungermannia confertissima Nees = Solenostoma confertis-
simum
Jungermannia connivens Dicks. = Fuscocephaloziopsis con-
nivens
Jungermannia crenulata Sm. non Schmidel = Solenostoma 
gracillimum
Jungermannia excisa Dicks. = Lophoziopsis excisa
Jungermannia curvifolia Dicks. = Nowellia curvifolia
Jungermannia execta Schrad. = Tritomaria execta
Jungermannia exectiformis Breidl. = Tritomaria exectiformis 
subsp. exectiformis
Jungermannia floerkei F. Weber & D. Mohr = Neoortho-
caulis floerkei
Jungermannia gracilis Schleich. ex Steph. = Neoorthocaulis 
attenuatus
Jungermannia gracillima Sm. = Solenostoma gracillimum
Jungermannia helleriana Nees ex Lindenb. = Crossocalyx 
hellerianus
Jungermannia heterocolpos Thed. ex Hartm. = Mesoptychia 
heterocolpos
Jungermannia hyalina Lyell = Solenostoma hyalinum
Jungermannia incisa Schrad. = Schistochilopsis incisa
Jungermannia inflata Huds. = Gymnocolea inflata
Jungermannia lanceolata L. = Jungermannia atrovirens
Jungermannia leiantha Grolle = Liochlaena lanceolata
Jungermannia lycopodioides Wallr. = Barbilophozia lycopo-
dioides
Jungermannia michauxii F. Weber = Anastrophyllum mi-
chauxii
Jungermannia minuta Schreb. = Sphenolobus minutus
Jungermannia mülleri Nees ex Lindenb. = Mesoptychia 
collaris
Jungermannia obovata Nees = Solenostoma obovatum
Jungermannia quinquedentata Huds. = Trilophozia quin-
quedentata
Jungermannia riparia Taylor = Jungermannia atrovirens
Jungermannia sphaerocarpa Hook. = Solenostoma sphaero-
carpum
Jungermannia starkii (Funck) Nees = Cephaloziella divaricata
Jungermannia subulata A. Evans = Liochlaena subulata
Jungermannia tamarisci L. = Frullania tamarisci
Jungermannia taylori Hook. = Mylia taylori
Jungermannia trichophylla L. = Blepharostoma trichophyl-
lum subsp. trichophyllum
Jungermannia turbinata Raddi = Mesoptychia turbinata
Jungermannia ventricosa Dicks. = Lophozia ventricosa
Kantia arguta (Nees & Mont.) Lindb. = Calypogeia arguta
Kantia calypogeia Lindb. = Calypogeia fissa
Kantia suecica Arnell & J. Perss. = Calypogeia suecica
Kantia trichomanis auct. = Calypogeia azurea
Kantia trichomanis var. neesiana Massal. & Carest. = 
Calypogeia neesiana
Kantia trichomanis var. repanda Nees = Calypogeia fissa 
Leiocolea alpestris (Schleicher ex F. Weber) Isov. = Mesop-
tychia collaris
Leiocolea badensis (Gottsche) Jörg. = Mesoptychia badensis
Leiocolea bantriensis (Hook.) Jörg. = Mesoptychia bantri-
ensis
Leiocolea collaris (Nees) Schljakov = Mesoptychia collaris
Leiocolea heterocolpos (Thed. ex Hartm.) Buch = Mesopty-
chia heterocolpos
Leiocolea muelleri (Nees ex Lindenb.) Jörg. = Mesoptychia 
collaris
Leiocolea turbinata (Raddi) Buch = Mesoptychia turbinata
Lejeunea calcarea Lib. = Cololejeunea calcarea
Lejeunea cavifolia var. planiuscula Lindb. apud Breidler, 
Głowacki = Lejeunea cavifolia
Lejeunea echinata (Hook.) Tayl. = Cololejeunea calcarea
Lejeunea rossettiana C. Massal. = Cololejeunea rossettiana
Lejeunea serpyllifolia Lib. = Lejeunea cavifolia
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New checklist of the hornworts and liverworts of Slovenia
Lejeunea ulicina (Tayl.) Gottsche et al. = Microlejeunea 
ulicina
Lepidozia setacea auct. = Kurzia pauciflora
Leptoscyphus taylori (Hook.) Mitt. = Mylia taylori
Lophocolea cuspidata Limpr. = Lophocolea coadunata
Lophozia alpestris (Schleich. ex F. Weber) A. Evans = Me-
soptychia collaris
Lophozia badensis (Gottsche) Schiffn. = Mesoptychia ba-
densis
Lophozia bantriensis (Hook.) Steph. = Mesoptychia ban-
triensis
Lophozia barbata (Schmidt) Dumort. = Barbilophozia 
barbata
Lophozia bicrenata (Schmidel ex Hoffm.) Dumort. = Iso-
paches bicrenatus
Lophozia collaris (Nees) Schljakov = Mesoptychia collaris
Lophozia debiliformis R. M. Schust. et Damsh. = Barbi-
lophozia sudetica
Lophozia elongata (Lindb.) Steph. = Protolophozia elongata
Lophozia excisa (Dicks.) Dumort. = Lophoziopsis excisa
Lophozia floerkei Dumort. = Neoorthocaulis floerkei
Lophozia gracilis (Schleich. ex Steph.) Steph. = Neoortho-
caulis attenuatus
Lophozia heterocolpos (Thed. ex C. Hartm.) M. Howe = 
Mesoptychia heterocolpos
Lophozia hornschuchiana (Nees) Macoum = Mesoptychia 
bantriensis
Lophozia incisa (Schrad.) Dumort. = Schistochilopsis incisa
Lophozia lycopodioides (Wallr.) Cog. = Barbilophozia lyco-
podioides
Lophozia michauxii (Web.) Steph. = Anastrophyllum mi-
chauxii
Lophozia polaris (R. M. Schust.) R. M. Schust. = Lopho-
ziopsis polaris
Lophozia polita (Nees) = Saccobasis polita
Lophozia porphyroleuca (Nees) Schiffn. = Lophozia guttu-
lata
Lophozia quadriloba (Lindb.) A. Evans = Schljakovianthus 
quadrilobus
Lophozia quinquedentata (Huds.) Cogn. = Trilophozia 
quinquedentata
Lophozia sudetica (Nees ex Hueben.) Grolle = Barbilopho-
zia sudetica
Lophozia turbinata (Raddi) Steph. = Mesoptychia turbinata
Lophozia ventricosa var. porphyroleuca (Nees) Hartm. = 
Lophozia guttulata
Madotheca baueri Schiffn. = Porella baueri
Madotheca cordaeana (Hueben.) Dumort. = Porella corda-
eana
Madotheca jackii Schiffn. = Porella platyphylla
Madotheca laevigata (Schrad.) Dumort. = Porella arboris-
vitae
Madotheca platyphylla (L.) Dumort. = Porella platyphylla
Madotheca rivularis Nees = Porella cordaeana
Marchantia alpestris (Nees) Burgeff = Marchantia 
polymorpha subsp. montivagans
Marchantia triandra Scop. = Mannia triandra
Marsupella ehrharti Corda = Marsupella emarginata
Marsupella neglecta (Limpr.) Steph. = Marsupella sprucei
Mastigobryum deflexum (Mart.) Nees = Bazzania tricre-
nata
Mastigobryum triangularis (Schleich. ex Steude) Steph. = 
Bazzania tricrenata
Metzgeria conjugata var. simplex (Lorb.) Aleffi & Schuma-
cker = Metzgeria simplex
Metzgeria fruticulosa (Dicks.) A. Evans = Metzgeria vio-
lacea
Nardia crenulata (Mitt.) Lindb. = Solenostoma gracillimum
Nardia haematosticta Lindb. = Nardia geoscyphus
Nardia hyalina Lindb. = Solenostoma hyalinus
Neesiella rupestris (Nees) Schiffn. = Mannia triandra
Pallavicinia hibernica (Hook.) Lindb. = Moerckia hiber-
nica
Pedinophyllum pyrenaicum Lindb. = Pedinophyllum inter-
ruptum
Pellia calycina (Tayl.) Nees = Apopellia endiviifolia
Pellia endiviifolia (Dicks.) Dumort. = Apopellia endivii-
folia
Peltolepis grandis (Lindb.) Lindb. = Peltolepis quadrata
Plagiochila interrupta (Nees) Dumort. = Pedinophyllum 
interruptum
Plectocolea obovata (Nees) Lindb. = Solenostoma obovatum
Porella rivularis (Nees) Lindb. = Porella cordaeana
Preissia quadrata (Scop.) Nees = Marchantia quadrata
Riccardia pinguis Gray = Aneura pinguis
Riccardia sinuata (Dicks. ex Hook.) Trevis = Riccardia 
chamaedryfolia
Riccia fluitans var. canaliculata (Hoffm.) A. Roth = Riccia 
canaliculata
Riccia intumescens (Bischl.) Heeg = Riccia ciliata
Riccia minima L. = Riccia sorocarpa
Sarcoscyphus ehrhardtii Corda = Marsupella emarginata
Sarcoscyphus emarginatus (Ehrh.) C. Hartm. = Marsupella 
emarginata
Sarcoscyphus funckii (F. Weber & D. Mohr) Nees = Mar-
supella funckii
Sarcoscyphus neglectus Limpr. = Marsupella sprucei
Scapania bartlingii (Hampe) Nees = Scapania cuspiduligera
Scapania convexa Pearson = Scapania umbrosa
Scapania dentata (Dumort.) Dumort. = Scapania undulata
Scapania nemorosa Dumort. = Scapania nemorea
Scapania rosacea (Corda) Nees = Scapania curta
Solenostoma amplexicaule (Dumort.) Steph. = Solenostoma 
sphaerocarpum
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New checklist of the hornworts and liverworts of Slovenia
Solenostoma crenulata (Sm.) R.H. Schust. = Solenostoma 
gracillimum
Solenostoma schiffneri (Loitl.) Müll. Frib. = Jungermannia 
polaris
Sphagoecetis communis (Dicks.) Nees = Odontoschisma 
sphagni
Sphagnoecetis communis var. macrior Nees = Odontoschi-
sma denudatum
Sphenolobus exectiformis (Breidl.) Steph. = Tritomaria exec-
tiformis
Sphenolobus exectus (Schmid.) Steph. = Tritomaria execta
Sphenolobus hellerianus (Nees ex Lindenb.) Steph. = Cros-
socalyx hellerianus
Sphenolobus michauxii (F. Weber) Steph. = Anastrophyl-
lum michauxii
Stephanina complanata O. Kuntze = Radula complanata
Stephanina lindenbergiana (Gottsche) O. Kuntze = Radula 
lindenbergiana
Telaranea setacea auct. = Kurzia pauciflora
Tritomaria polita (Nees) Jörg. = Saccobasis polita
Tritomaria quinquedentata (Huds.) H. Buch = Trilophozia 
quinquedentata
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