folia biologica et geologica 58/1, 5–30, ljubljana 2017
TWO NEW SCREE PLANT COMMUNITIES IN THE TRIGLAV 
MOUNTAINS (JULIAN ALPS, SLOVENIA)
DVE NOVI RASTLINSKI ZDRUŽBI MELIŠČ V TRIGLAVSKEM 
POGORJU (JULIJSKE ALPE, SLOVENIJA)
Igor DAKSKOBLER* & Branko ZUPAN**
http://dx.doi.org/10.3986/fbg0018
abStRact
two new scree plant communities in the triglav Moun-
tains (Julian alps, Slovenia)
In the Triglav Mountains, mainly on the slopes and pla-
teaus to the west of Mt. Triglav (Glava v Zaplanji, Vrh Zele-
nic) we found new localities of three rare species of Slove-
nian flora, Crepis terglouensis, Cerastium uniflorum and 
Geum reptans, and made a phytosociological inventory of 
their sites. Based on comparisons with similar communities 
within which they occur elsewhere in the Eastern and South-
eastern Alps we described two new associations, Crepido ter-
glouensis-Potentilletum nitidae (alliance Thlaspion rotundi-
folii) and Saxifrago carniolicae-Cerastietum uniflorae (alli-
ance Arabidion caeruleae).  
Key words: Alpine flora, phytosociology, synsystemat-
ics, Crepis terglouensis, Cerastium uniflorum, Geum reptans, 
Triglav National Park, Slovenia
iZVleČeK
Dve novi rastlinski združbi melišč v triglavskem pogorju 
(Julijske alpe, Slovenija)
V Triglavskem pogorju, predvsem na pobočjih in plano-
tah zahodno od Triglava (Glava v Zaplanji, Vrh Zelenic) smo 
našli nova nahajališča treh redkih vrst v flori Sloveniji, Cre-
pis terglouensis, Cerastium uniflorum in Geum reptans in 
fitocenološko popisali njihova rastišča. Na podlagi primer-
jav s podobnimi združbami, v katerih uspevajo omenjene 
vrste drugod v Vzhodnih in Jugovzhodnih Alpah, smo opis-
ali dve novi asociaciji Crepido terglouensis-Potentilletum ni-
tidae (zveza Thlaspion rotundifolii) in Saxifrago carniolicae-
-Cerastietum uniflorae (zveza Arabidion caeruleae).  
Ključne besede: alpska flora, fitocenologija, sinsistema-
tika, Crepis terglouensis, Cerastium uniflorum, Geum rep-
tans, Triglavski narodni park, Slovenija 
 * Institute of Biology, Scientific Research Centre of the Slovenian Academy of Sciences and Arts, Regional unit Tolmin, Brunov 
drevored 13, SI-5220 Tolmin and Biotechnical Faculty of the University in Ljubljana, Department of Forestry and Renewable 
Forest Resources, Večna pot 83, SI-1000 Ljubljana, Igor.Dakskobler@zrc-sazu.si
 ** Savica 6, SI-4264 Bohinjska Bistrica 
Igor DAKSKoBLEr & BrAnKo ZUPAn: TWo nEW SCrEE PLAnT CoMMUnITIES In THE TrIgLAV MoUnTAInS
6 folia biologica et geologica 58/1 – 2017
When mapping the flora on the slopes and plateaus 
west of Mt. Triglav in 2015 and 2016 we found and re-
corded fascinating scree communities with three rare 
species of Slovenian flora – Crepis terglouensis, Cerasti-
um uniflorum and Geum reptans. We compared them 
to similar communities that had been studied in the 
Triglav Mountains and in the Julian Alps by T. Wra-
ber (1972) and with similar communities elsewhere in 
the Eastern Alps, and classified them into a syntaxo-
nomic system.
1 INTRODUCTION
2 METHODS
Alpine communities under Mt. Triglav were studied 
applying the Braun-Blanquet method (Braun-
-Blanquet 1964). A total of 29 relevés (of which five 
had already been published by T. Wraber, 1972 and 
one by the authors of this paper – Dakskobler & Su-
rina, 2017) were entered into the FloVegSi database 
(Fauna, Flora, Vegetation and Paleovegetation of Slo-
venia) of the Jovan Hadži Institute of Biology at SRC 
SASA (T. Seliškar, Vreš et A. Seliškar 2003). They 
were arranged into a working table based on hierarchi-
cal classification. We transformed the combined cov-
er-abundance values with numerical values (1–9) ac-
cording to van der Maarel (1979). Numerical com-
parisons were performed with the SyN-TAX 2000 
program package (Podani 2001). The relevés were 
compared by means of (unweighted) average linkage 
method – UPGMA, using Wishart’s similarity ratio.
In the first step we used numerical analyses as the 
basis on which we formed floristically homogeneous 
groups of relevés that were subsequently compared, 
using the same methodology, with similar communi-
ties in the Eastern Alps, also using hierarchical classi-
fication and the same method as when we compared 
individual relevés.  
The nomenclature sources for the names of vascu-
lar plants are the Mala flora Slovenije (Martinčič et 
al. 2007) and Flora alpina (Aeschimann et al. 2004a,b). 
Prof. Andrej Martinčič determined the collected 
mosses. For the names of syntaxa we follow Englisch 
et al. (1993), Theurillat (2004), Šilc & Čarni (2012), 
E. Pignatti & S. Pignatti (2014) and Mucina et al. 
(2016). In the classification of species into phytosocio-
logical groups (groups of diagnostic species) we main-
ly refer to the Flora alpina (Aeschimann et al. 2004a,b). 
The geographic coordinates of relevés are determined 
according to the Slovenian geographic coordinate sys-
tem D 48 (5th zone) on the Bessel ellipsoid and with 
Gauss-Krüger projection.
The relevés discussed in this article were made in 
the Triglav range of the Julian Alps, mostly on the pla-
teuas west and southwest of Triglav. The geological 
bedrock in the study area is mainly calcareous, lime-
stone and dolomite limestone (Buser 2009). The stud-
ied communities occur on initial soils (lithosols) – Lo-
vrenčak (1998), Vidic et al. (2015). The climate is cold 
and humid, with mean annual precipitation of 2,500 to 
3,000 mm (Zupančič 1998) and mean annual air tem-
perature of -2 ºC to 0 ºC 
3. RESULTS AND DISCUSSION
3.1 Review of the studied syntaxa, with types of 
newly described communities 
Thlaspietea rotundifolii Br.-Bl. 1948
Thlaspietalia rotundifolii Br.-Bl. in Br.-Bl. et Jenny 
1926
Thlaspion rotundifolii Jenny-Lips 1930
Papaveri julici-Thlaspietum rotundifolii T. Wraber 1970
Crepidetum terglouensis Seibert 1977
Crepido terglouensis-Potentilletum nitidae ass. nov. 
hoc. loco, the nomenclature type, holotypus, is rele-
vé 12 in Table 1.
-typicum, subass. nov., the nomenclature type is the 
same as the nomenclature type of   the association
-caricetosum firmae, the nomenclature type, holo-
typus, is relevé 5 in Table 1.
Saxifrago sedoidis-Geumetum reptantis nom. prov.
Arabidetalia caeruleae Rübel ex Nordhagen 1937
Arabidion caeruleae Br.-Bl. In Br.-Bl. et Jenny 1926
Saxifrago carniolicae-Cerastietum uniflorae ass. 
nov. hoc loco, the nomenclature type, holotypus, is 
relevé 18 in Table 1.
Asplenietea trichomanis (Br.-Bl. in Meier et Br.-Bl. 
1934) Oberd. 1977
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Potentilletalia caulescentis Br.-Bl. in Br.-Bl. et Jenny 
1926
Physoplexido comosae-Saxifragion petraeae Muci-
na et Theurillat 2015 (syn. Androsaco-Drabion to-
Figure 1: Localities of recorded stands in the Triglav mountains; yellow circle – stands of the association Crepido terglouensis-
Potentilletum nitidae, red circle – stands of the association Saxifrago-Cerastietum uniflorae, blue circle – stand of the associa-
tion Saxifrago-Geumetum reptantis
Slika 1: Nahajališča preučenih sestojev v Triglavskem pogorju: rumen krog – sestoji asociacije Crepido terglouensis-Potentille-
tum nitidae, rdeč krog – sestoji asociacije  Saxifrago-Cerastietum uniflorae, moder krog – sestoj asociacije Saxifrago-Geume-
tum reptantis
mentosae T. Wraber 1970,  Phyteumato-Saxifragion 
petraeae Mucina in Šilc et Čarni 2012)
Potentilletum nitidae Wikus 1959
Igor DAKSKoBLEr & BrAnKo ZUPAn: TWo nEW SCrEE PLAnT CoMMUnITIES In THE TrIgLAV MoUnTAInS
8 folia biologica et geologica 58/1 – 2017
Figure 2: Distribution of Crepis terglouensis in Slovenia
Slika 2: Razširjenost vrste Crepis terglouensis v Sloveniji
Figure 3: Dendrogram of recorded stands in the Triglav Mountains, UPGMA, 1– similarity ratio 
PN Potentilletum nitidae; CTPN Crepido terglouensis-Potentilletum nitidae,Ss Salicetum serpyllifoliae nom. prov., SP Saxifra-
getum paniculatae nom. prov., SpCF Saxifrago paniculatae-Caricetum fuliginosae, PT Papaveri julici-Thlaspietum rotundifolii, 
CU Saxifrago-Cerastietum uniflorae
Slika 3: Dendrogram popisov preučenih sestojev v Triglavskem pogorju, UPGMA, komplement Wishartovega koeficienta 
podobnosti
PN
C
TP
N
C
TP
N
C
TP
N
C
TP
N
C
TP
N
C
TP
N
C
TP
N
C
TP
N
C
TP
N
C
TP
N
C
TP
N
C
TP
N
C
TP
N
C
TP
N
C
TP
N Ss SP SP
Sp
C
F
PT PT PT C
U
C
U
C
U
C
U
C
U
C
U
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3.2 Crepis terglouensis and its communities in 
Slovenia
According to Flora alpina (Aeschimann et al. 2004b: 
668) Crepis terglouensis is an eastern-Alpine species, a 
character species of the alliance Thlaspion rotundifolii 
that comprises vegetation of subalpine-alpine calcare-
ous screes. The species also gave its name to the alpine 
community Crepidetum terglouensis Seibert 1977 from 
the Eastern Alps, which was presented with a phytoso-
ciological table by Eggensberger (1994: 64–66, 76–
77). Certain similarities with this community can be 
observed also in the relevé published by Sutter (1969: 
353) that he had made together with T. Wraber under 
the Planika Lodge at Mt. Triglav. T. Wraber (1972, 
1990) recorded Crepis terglouensis mainly in the stands 
of a special form of the association Papaveri julici-
-Thlaspietum rotundifolii. He found it on two screes at 
Kredarica and on Grlo pass between Oltar and Dovški 
Križ. E. & S. Pignatti (2014, 2016) mention it as a rare 
species in the stands of two other alpine scree commu-
nities, Leontodontetum montani and Papaveretum rha-
etici, and in the special form (Seleginella-Homogyne) of 
the association Seslerio-Caricetum sempervirentis. A 
phytosociological inventory of its two new localities in 
the Slovenian Alps (on talus under Prestreljenik in the 
Kanin Mountains – Praprotnik, 1997, 2002, and on 
the scree under Mt. Stol in the Karavanke Mts. – 
Novak 2015) has not been made until now. Novak 
(ibid.) mentions companion species Sesleria caerulea 
and Campanula cochleariifolia for the locality under 
Mt. Stol. In our research we made most of the relevés 
with Crepis terglouensis on fine talus on Glava nad Za-
planjo under Mt. Triglav and on the neighbouring 
slopes towards Dolič (9648/2), but we also found a new 
locality on a similar site on Vrata pass between Zel-
narica and Zadnji Vogel (9648/4) at the elevation of 
2,192 m (leg. & det. I. Dakskobler, B. Anderle and B. 
Zupan, 23. 8. 2016, herbarium LJS), which is a new lo-
cality of this species in the new quadrant (Figures 1, 2). 
In the comparison of our relevés with Crepis terglouen-
sis and (or) Cerastium uniflorum with similar relevés 
made by T. Wraber (1972), the relevés with co-domi-
nating Potentilla nitida and Crepis terglouensis grouped 
Figure 4: Dendrogram of syntaxa Potentilletum nitidae (PN), Crepidetum terglouensis (CT) and Crepido terglouensis-Potentil-
letum nitidae (CTPN), UPGMA, 1– similarity ratio 
Slika 4: Dendrogram sintaksonov Potentilletum nitidae (PN), Crepidetum terglouensis (CT) and Crepido terglouensis-Potentil-
letum nitidae (CTPN), UPGMA, komplement Wishartovega koeficienta podobnosti 
Igor DAKSKoBLEr & BrAnKo ZUPAn: TWo nEW SCrEE PLAnT CoMMUnITIES In THE TrIgLAV MoUnTAInS
10 folia biologica et geologica 58/1 – 2017
separately (Figure 3). We obviously inventoried a form 
of alpine vegetation that is transitional between scree 
communities, chasmophytic communities and stony 
grasslands, which had already been demonstrated by 
Sutter’s relevé (ibid.) and mentioned also by Eggens-
berger (ibid.). We therefore made a synthetic table 
(Appendix 1) in which we compared 15 relevés of the 
studied stands with 33 relevés of the association Poten-
tilletum nitidae (T. Wraber 1972, Table 3) and with 25 
relevés of the association Crepidetum terglouensis 
(Eggensberger 1994, Table 6, columns 85–111). The 
results (Figure 4) indicate that our relevés are more 
similar to the stands of the association Potentilletum 
nitidae. They could therefore be classified into the new 
subassociation Potentilletum nitidae crepidetosum ter-
glouensis, but the analysis of diagnostic species (Table 
2, column 1) indicates the predominance of scree spe-
cies from the order Thlaspietelia rotundifolii (37.43%) 
over the diagnostic chasmophytic species from the 
order Potentilletalia caulescentis (23.8%). Most of the 
relevés were made on fine talus that is almost consis-
tently mixed with well-decomposed humus (mull), on 
levelled terrain with a relatively persistent snow cover. 
As a rule, Potentilla nitida is the dominant species in 
inventoried stands and its joint occurrence with the 
characteristic scree species Crepis terglouensis on rela-
tively small but ecologically unique areas between 
rocks (fine talus) can indicate a stage in development 
(succession) that could be partly associated with the 
ongoing climate change, reduced precipitation vol-
umes and shorter periods of snow cover on the pla-
teaus to the west of Mt. Triglav, as demonstrated by the 
measurements of the Triglav Glacier at a similar eleva-
tion (Gabrovec et al. 2014). This stage in succession 
can be treated also at the rank of the new association 
Crepido terglouensis-Potentilletum nitidae, which is 
classified into the alliance Thlaspion rotundifolii. Di-
agnostic species of the new association are Potentilla 
nitida, Crepis terglouensis, Alyssum ovirense, Eritrichi-
um nanum and Minuartia cherlerioides. We distin-
guish between two subassociations, typical (-typicum) 
and -caricetosum firmae. The differential species of the 
latter are Carex firma, Silene acaulis and Salix retusa, 
which might indicate a transition towards stony alpine 
grasslands from the association Gentiano terglouensis-
-Caricetum firmae.
Figure 5: Distribution of Cerastium uniflorum in Slovenia
Slika 5: Razširjenost vrste Cerastium uniflorum v Sloveniji
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3.3 Cerastium uniflorum and its communities in 
Slovenia
Cerastium uniflorum is an Alpine-Carpathian species, 
a character species of silicate subalpine and alpine 
screes (alliance Androsacion alpinae ) – Aeschimann 
et al. (2004a: 298). Its distribution in Slovenia is shown 
in Figure 5. Its reported localities are only on Mt. Tri-
glav and its vicinity, on Mt. Rjavina and on Mt. Man-
gart. T. Wraber (1990: 130, 1972) reported its occur-
rence in the stands of associations Potentilletum niti-
dae and Papaveri julici-Thlaspietum rotundifolii.  E. & 
S. Pignatti (2014, 2016) recorded this species in the 
stands of associations Drabetum hoppeanae, Saxifrage-
tum sedoidis, Leontodontetum montani, Papaveretum 
rhaetici, Festucetum pulchellae, Vitaliano-Eritrichetum 
nani, Arabidetum caeruleae, Saxifragetum bryoidis and 
several others. In our relevés it occurs with individual 
specimens within associations Saxifrago paniculatae-
-Caricetum fuliginosae (Dakskobler & Surina 2017) 
and Crepido-Potentilletum nitidae (this article), but it is 
also one of the dominating species in six relevés on 
mainly fine talus and on sites with a persistent snow 
cover (Figure 1, relevés 17–21 in Table 1). These relevés 
grouped separately from other relevés of alpine screes 
and alpine swards on rock ledges (Figure 3). The most 
frequent species in addition to Cerastium uniflorum is 
Saxifrage sedoides, so we compared them with the as-
sociation Saxifragetum sedoidis from the Dolomites (E. 
& S. Pignatti 2016: 204: Association table 9.2, 406– 
407) – Appendix 2. Surina (2005) prepared a phytoso-
ciological table for the Krn Mts. in which he presented 
a similar syntaxon Saxifragetum stellaro-sedoidis var. 
geogr. Ranunculus traunfellneri, but in his stands he 
did not record Cerastium uniflorum, which is absent 
also from the original description of this association 
(Englisch 1999). Floristic similarity of our relevés 
with the relevés of the stands of the association Saxi-
fragetum sedoidis is only 41% (Sørensen 1948), which 
does not allow for its classification into this associa-
tion. These stands can also be explained as a succes-
sional stage, because in the extreme conditions of the 
alpine belt Cerastium uniflorum can grow on various 
sites and overgrows even areas where it may not have 
occurred 50 or 100 years ago. For this reason and based 
on the composition of diagnostic species (Table 2, col-
umn 2) we classify them into the new association Saxi-
frago carniolicae-Cericetum uniflorae and into the alli-
ance Arabidion caeruleae. Its diagnostic species are 
Cerastium uniflorum, Saxifraga sedoides and Saxifraga 
exarata subsp. carniolica. The latter is endemic to the 
Southeastern Alps and a character species of alpine 
chasmophytic communities (Potentilletum nitidae, Po-
tentillo clusianae-Campanuletum zoysii) – T. Wraber 
(2006: 70) and of stony grasslands from alliances Cari-
cion firmae and Seslerion variae (Aeschimann et al. 
2004: 2010). The first two listed species characterise an 
alpine community on fine talus in areas with long-
lasting snow cover and the third characterises the new 
association mainly in terms of phytogeography and 
partly indicates similar development of communities 
on fine gravel with communities on stony alpine 
swards and rock crevices. Abundant Sagina saginoides 
in one of the stands indicates locally improved soil 
conditions with an abundance of fine weathered mate-
rial (mull).
3.4 Geum reptans in the triglav Mountains
Aeschimann et al. (2004a: 758) classify the South-
European montane species Geum reptans as a charac-
ter species of the alliance Androsacion alpinae. The 
only reported localities in the Julian Alps so far have 
been those on Mts. Mangart and Kanjavec (T. Wra-
ber 2006, Zupan & Dakskobler 2007, Figure 6). On 
8 August 2016 we found a new locality of this scree 
species on the western slopes of Triglav, in the cirque 
under Vrh Zelenic, on the elevation of 2,060 m 
(Dakskobler 2017, Figure 1). The site is a well-over-
grown, consolidated scree with rocks of various sizes. 
Its species composition is shown in Column 2 of Table 
3. Column 1 in the same table presents the relevé of the 
community on the locality at Teme between Kanjavec 
and Poprovec (Zupan & Dakskobler 2007), which 
was identified as a transition between the community 
of Potentilletum nitidae and the stand of the associa-
tion Papaveri julici-Thlaspietum rotundifolii. The stand 
under Vrh Zelenic cannot be classified into either of 
the mentioned associations. It characterises a scree 
community with many species of stony Alpine swards 
(from the association Gentiano terglouensis-Caricetum 
firmae). It is provisionally classified into the associa-
tion Saxifrago sedoidis-Geumetum reptantis nom. prov. 
and into the alliance Thlaspion rotundifolii. In addi-
tion to Geum reptans the species that characterise this 
community are mainly those from the genus Saxifraga 
(S. sedoides, S. paniculata, S. aizoides, S. crustata, S. 
squarrosa) and two typical scree species, Festuca nitida 
and Poa minor.
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The result of floristic mapping in the western part of 
Mt. Triglav (Glava v Zaplanji, Vrh Zelenic) and on 
Vrata pass between Velika Zelnarica and Zadnji Vogal 
are new localities of three alpine scree and (or) chas-
mophytic species that are rare in Slovenia and listed on 
the Red List (Anon. 2002): Cerastium uniflorum (new 
localities in quadrant 9648/2), Crepis terglouensis (new 
localities in quadrants 9648/2 and 9648/4) and Geum 
reptans (new locality in the existing quadrant, 9648/2, 
new to the flora of Triglav). We made a phytosocio-
logical inventory of the communities in which they 
grow and determined untypical stands on fine talus 
with fine weathered material (mull) that are transi-
tional between alpine chasmophytic, scree and snow 
bed communities and could be a successional stage in 
areas that decades ago were covered by snow for much 
longer periods than they are today. Based on their 
comparison with similar communities from the East-
ern and Southeastern Alps we described two new as-
sociations, Crepido terglouensis-Potentilletum nitidae 
and Saxifrago carniolicae-Cerastietum uniflorae. Even 
though their stands are located in the vicinity of quite 
popular mountain trails to Triglav they are not yet en-
dangered by increasing numbers of tourists in the 
summer as there are no other interventions into this 
area. In addition, neither Cerastium uniflorum nor 
Crepis terglouensis are prominent species and are quite 
inconspicuous outside their flowering season. The new 
locality of Geum reptans, which occurs also in the non-
typical scree community (Saxifrago sedoidis-Geume-
tum reptantis nom. prov.), is in a remote pathless area 
in the cirque under Vrh Zelenic and therefore still out-
side any direct influence of man.
Figure 6: Distribution of Geum reptans in Slovenia
Slika 6: Razširjenost vrste Geum reptans  v Sloveniji
4 CONCLUSIONS
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5.1 Uvod
Pri kartiranju flore na pobočjih in planotah zahodno 
od Triglava smo v letih 2015 in 2016 našli in popisali 
zanimive meliščne združbe, v katerih uspevajo tri 
redke vrste slovenske flore: Crepis terglouensis, Cera-
stium uniflorum in Geum reptans. Primerjali smo jih s 
podobnimi združbami, ki jih je v Triglavskem pogorju 
in v Julijskih Alpah raziskal T. Wraber (1972), in s 
podobnimi združbami drugod v Vzhodnih Alpah in 
jih uvrstili v sintaksonomski sistem. 
5.2 Metode
Alpinske združbe pod Triglavom smo preučevali po 
ustaljeni srednjeevropski metodi (Braun-Blanquet 
1964).  Skupno 29 popisov (od tega jih je pet predho-
dno objavil že T. Wraber, 1972, enega pa mi – 
Dakskobler & Surina, 2017) smo vnesli v podatkov-
no bazo FloVegSi (T. Seliškar, Vreš & A. Seliškar 
2003). Fitocenološke popise smo v delovno tabelo ure-
dili na podlagi hierarhične klasifikacije. Kombinirane 
ocene zastiranja in pogostnosti smo pretvorili v šte-
vilčne vrednosti (1–9) – van der Maarel 1979). Popise 
smo primerjali z metodo kopičenja na podlagi povezo-
vanja (netehtanih) srednjih razdalj – (Unweighted) 
average linkage method – UPGMA. Uporabljali smo 
programski paket SyN-TAX 2000 (Podani 2001) in 
kot mero različnosti komplement koeficienta »simila-
rity ratio«. V prvem koraku smo na podlagi numerič-
nih analiz oblikovali floristično homogene skupine 
popisov, ki smo jih nato z enakim metodološkim pri-
stopom primerjali s podobnimi združbami v Vzho-
dnih Alpah, pri tem pa prav tako uporabili hierarhično 
klasifikacijo in isto metodo kot pri primerjavi posa-
mičnih popisov.  
Nomenkalturna vira za imena prarotnic in semenk 
sta Mala flora Slovenije (Martinčič et al. 2007) in 
Flora alpina (Aeschimann et al. 2004a,b). Mahove, ki v 
preučenih združbah nimajo večjega zastiranja, je dolo-
čil prof. Andrej Martinčič. Nomenklaturni viri za 
imena sintaksonov so Englisch et al. (1993), Theuril-
lat (2004), Šilc & Čarni (2012), E. Pignatti & S. Pi-
gnatti (2014) in Mucina et al. (2016). Pri uvrščanju 
vrst v fitocenološke skupine smo v glavnem upoštevali 
delo Flora alpina (Aeschimann et al., ibid.). Geograf-
ske koordinate popisov smo določili po slovenskem ge-
ografskem koordinatnem sistemu D 48 (cona 5) na Bes-
selovem elipsoidu in z  Gauss-Krügerjevo projekcijo. 
Popise smo naredili v Triglavskem pogorju v Julij-
skih Alpah, večino na planotah zahodno in jugozaho-
dno od vrha Triglava. Kot geološka podlaga se poja-
vljata na raziskovanem območju apnenec in dolomit 
(Buser 2009). Preučene združbe uspevajo na kamni-
ščih (Lovrenčak 1998, Vidic et al. 2015). Podnebje je 
hladno in vlažno, s povprečno letno višino padavin 
med 2500 mm in 3000 mm (Zupančič 1998) in sre-
dnjo letno temperature zraka med -2 ºC in 0 ºC (Ce-
gnar 1998). 
5.3. Rezultati in razprava
5.3.1 Pregled preučenih sintaksonov s tipi na novo opi-
sanih združb 
Thlaspietea rotundifolii Br.-Bl. 1948
Thlaspietalia rotundifolii Br.-Bl. in Br.-Bl. et Jenny 
1926
Thlaspion rotundifolii Jenny-Lips 1930
Papaveri julici-Thlaspietum rotundifolii T. Wraber 
1970
Crepidetum terglouensis Seibert 1977
Crepido terglouensis-Potentilletum nitidae ass. nov. 
hoc. loco, nomenklaturni tip,  holotypus, je popis 12 
v preglednici 1.
-typicum, subass. nov., nomenklaturni tip je isti 
kot nomenklaturni tip asociacije
-caricetosum firmae, nomenklaturni tip, holotypus, 
je popis 5 v preglednici 1.
Saxifrago sedoidis-Geumetum reptantis nom. prov.
Arabidetalia caeruleae Rübel ex Nordhagen 1937
Arabidion caeruleae Br.-Bl. In Br.-Bl. et Jenny 1926
Saxifrago carniolicae-Cerastietum uniflorae ass. 
nov. hoc loco, nomenklaturni tip, holotypus, je 
popis 18 v preglednici 1.
Asplenietea trichomanis (Br.-Bl. in Meier et Br.-Bl. 
1934) Oberd. 1977
Potentilletalia caulescentis Br.-Bl. in Br.-Bl. et Jenny 
1926
Physoplexido comosae-Saxifragion petraeae Mucina 
et Theurillat 2015 (sin. Androsaco-Drabion tomen-
tosae T. Wraber 1970, Phyteumato-Saxifragion pe-
traeae Mucina in Šilc et Čarni 2012)
Potentilletum nitidae Wikus 1959
5.3.2 Vrsta Crepis terglouensis in združbe, v katerih 
uspeva v Sloveniji
Po delu Flora alpina (Aeschimann et al. 2004b: 668) je 
Crepis terglouensis vzhodnoalpska vrsta, značilnica 
zveze Thlaspion rotundifoliae, ki združuje rastje subal-
5 POVZETEK
Igor DAKSKoBLEr & BrAnKo ZUPAn: TWo nEW SCrEE PLAnT CoMMUnITIES In THE TrIgLAV MoUnTAInS
14 folia biologica et geologica 58/1 – 2017
pinsko-alpinskih melišč na karbonatni podlagi. Po njej 
se imenuje vzhodnoalpska alpinska združba Crepide-
tum terglouensis Seibert 1977, ki jo je s fitocenološko 
tabelo podrobno predstavil Eggensberger (1994: 64–
66, 76–77). Nekaj podobnosti s to združbo naj bi imel 
tudi popis, ki ga je objavil Sutter (1969: 353) in sta ga 
naredila skupaj s T. Wraberjem pod planinskim 
domom Planika pod Triglavom. T. Wraber (1972, 
1990) triglavski dimek omenja predvsem v posebni 
obliki asociacije Papaveri julici-Thlaspietum rotundifo-
lii. Našel ga je na dveh meliščih na Kredarici in na pre-
lazu Grlo med Oltarjem in Dovškim križem. E. & S. 
Pignatti (2014, 2016) to vrsto kot redko navajata v 
sestojih dveh drugih alpinskih meliščnih združb Leon-
todontetum montani in Papaveretum rhaetici in v po-
sebni obliki (Seleginella-Homogyne) asociacije Seslerio-
-Caricetum sempervirentis. Dve novi nahajališči te 
vrste v slovenskih Alpah (na grušču pod Prestreljeni-
kom v Kaninskem pogorju – Praprotnik, 1997, 2002 
in na melišču pod Stolom v Karavnkah – Novak 2015) 
fitocenološko do zdaj nista popisani. Na nahajališču 
pod Stolom, Novak (ibid.) omenja spremljevalni vrsti 
Sesleria caerulea in Campanula cochleariifolia. Pri 
naših raziskavah smo večino popisov triglavskega 
dimka naredili na drobnem grušču na Glavi nad Za-
planjo pod Triglavom in na sosednjih vzpetinah v 
smeri proti Doliču (9648/2) – slika 1, novo nahajališče 
pa smo našli tudi na enakem rastišču na prelazu Vrata 
med Zelnarico in Zadnjim Voglom (9648/4), na nad-
morski višini 2192 m (leg. & det. I. Dakskobler, B. An-
derle in B. Zupan, 23. 8. 2016, herbarij LJS), kar je novo 
nahajališče te vrste v novem kvadrantu (slika 2). Po 
primerjavi naših popisov z vrstama Crepis terglouensis 
in/ali Cerastium uniflorum s podobnimi popisi T. 
Wraberja (1972), so se posebej združevali popisi, v 
katerih skupaj prevladujeta vrsti Potentilla nitida in 
Crepis terglouensis (slika 3). Očitno smo popisali pre-
hodno obliko alpinskega rastja med meliščnimi združ-
bami, združbami skalnih razpok in kamnitih travišč, 
ki jo je z enim popisom ponazoril že Sutter (ibid.) in 
jo omenja tudi Eggensberger (ibid.). Zato smo izdela-
li sintezno tabelo (Dodatek 1), v kateri smo 15 popisov 
preučenih sestojev primerjali s 33 popisi asociacije Po-
tentilletum nitidae (T. Wraber 1972, preglednica 3) in 
s 25 popisi asociacije Crepidetum terglouensis (Eggens-
berger 1994, preglednica 6, stolpci 85–111). Rezultati 
(slika 4) kažejo na večjo podobnost naših popisov s 
sestoji asociacije Potentilletum nitidae. Mogoče bi jih 
bilo torej uvrstiti v novo subasociacijo, Potentilletum 
nitidae crepidetosum terglouensis, toda analiza diagno-
stičnih vrst (preglednica 2, stolpec 1) kaže na prevlado 
meliščnih vrst iz reda Thlaspietelia rotundifolii (37,43 
%) nad diagnostičnimi vrstami skalnih razpok iz reda 
Potentilletalia caulescentis (23,8 %). Večino popisov 
smo naredili na drobnem grušču, med katerim je sko-
raj vedno nekaj dobro razkrojenega humusa (sprsteni-
ne), na uravnavah, kjer se razmeroma dolgo zadržuje 
sneg. Dominantna vrsta popisanih sestojev je navadno 
Potentilla nitida, njeno družno uspevanje s tipično me-
liščno vrsto Crepis terglouensis na razmeroma majh-
nih, a ekološko svojskih površinah med skalovjem 
(droben grušč) lahko označuje določeno razvojno 
(sukcesijsko) stopnjo, ki je morda deloma povezana 
tudi z zdajšnjimi podnebnimi spremembami, očitno 
manjšo količino snežnih padavin in krajšimi obdobji s 
snežno odejo na planotah zahodno od Triglava, kar 
dokazujejo meritve bližnjega Triglavskega ledenika na 
podobni nadmorski višini (Gabrovec et al. 2014). To 
sukcesijsko stopnjo lahko obravnavamo tudi v rangu 
nove asociacije Crepido terglouensis-Potentilletum niti-
dae, ki jo uvrščamo v zvezo Thlaspion rotundifolii. Di-
agnostične vrste nove asociacije so Potentilla nitida, 
Crepis terglouensis, Eritrichium nanum in Minuartia 
cherlerioides. Razlikujemo dve subasociaciji, tipično 
(-typicum) in -caricetosum firmae. Razlikovalnice sle-
dnje so vrste Carex firma, Silene acaulis in Salix retusa, 
ki morda nakazujejo prehod proti kamnitim alpin-
skim traviščem iz asociacije Gentiano terglouensis-Ca-
ricetum firmae. 
5.3.3 Vrsta Cerastium uniflorum in združbe, v katerih 
uspeva v Sloveniji
Cerastium uniflorum je alpsko-karpatska vrsta, značil-
nica zveze silikatnih melišč subalpinskega in alpinske-
ga pasu Androsacion alpinae (Aeschimann et al. 
2004a: 298). Njeno razširjenost v Sloveniji prikazuje 
slika 5. Znana nahajališča so le na Triglavu in v njego-
va soseščini, na Rjavini in Mangartu. T. Wraber 
(1990: 130, 1972) navaja njeno pojavljanje v sestojih 
asociacij Potentilletum nitidae in Papaveri julici-Thla-
spietum rotundifolii. E. & S. Pignatti (2014, 2016) sta 
to vrsto popisala v sestojih asociacij Drabetum hoppea-
nae, Saxifragetum sedoidis, Leontodontetum montani, 
Papaveretum rhaetici, Festucetum pulchellae, Vitalia-
no-Eritrichetum nani, Arabidetum caeruleae, Saxifra-
getum bryoidis in v še nekaterih drugih. V naših popi-
sih jo imamo kot posamično primes v sestojih asociacij 
Saxifrago paniculatae-Caricetum fuliginosae (Daksko-
bler & Surina 2017) in Crepido-Potentilletum nitidae 
(ta članek), je pa ena od prevladujočih vrst v šestih po-
pisih na večinoma drobnem grušču in na krajih, kjer se 
dolgo zadržuje sneg (slika 6, popisi 17–21 v preglednici 
1). Ti popisi so se združevali ločeno od drugih popisov 
alpinskih melišč in alpskih trat na skalnatih policah 
(slika 2). Poleg vrste Cerastium uniflorum je na njih 
Igor DAKSKoBLEr & BrAnKo ZUPAn: TWo nEW SCrEE PLAnT CoMMUnITIES In THE TrIgLAV MoUnTAInS
15folia biologica et geologica 58/1 – 2017
najbolj pogosta vrsta Saxifrage sedoides, zato smo jih 
primerjali s popisi asociacije Saxifragetum sedoidis iz 
Dolomitov (E. & S. Pignatti 2016: 204: asociacijska 
preglednica 9.2, 406–407) – Dodatek 2. V Krnskem 
pogorju je Surina (2005) s fitocenološko tabelo pred-
stavil nekoliko podoben sintakson Saxifragetum stella-
ro-sedoidis var. geogr. Ranunculus traunfellneri, vendar 
v njegovih sestojih ni popisal enocvetne smiljke in tudi 
v izvornem opisu te asociacije (Englisch 1999) je ni. 
Floristična podobnost naših popisov s popisi sestojev 
asociacije Saxifragetum sedoidis je precej manj kot po-
lovična, le 41-odstotna (Sørensen 1948), kar ne dopu-
šča uvrstitve v to asociacijo. Tudi te sestoje lahko razlo-
žimo kot razvojno (sukcesijsko) stopnjo, saj vrsta Cera-
stium uniflorum v skrajnih razmerah alpinskega pasu 
lahko uspeva na različnih rastiščih in porašča tudi 
površine, ki jih morda pred 50 ali 100 leti še ni pora-
ščala. Zato in na podlagi sestave diagnostičnih vrst (ta-
bela 2, stolpec 2) jih uvrščamo v novo asociacijo Saxi-
frago carniolicae-Cericetum uniflorae in v zvezo Arabi-
dion caeruleae. Njene diagnostične vrste so Cerastium 
uniflorum, Saxifraga sedoides in Saxifraga exarata 
subsp. carniolica. Slednja je endemit Jugovzhodnih Alp 
in značilnica alpinskih združb skalnih razpok (Poten-
tilletum nitidae, Potentillo clusianae-Campanuletum 
zoysii)  – T. Wraber (2006: 70) in kamnitih travišč iz 
zvez Caricion firmae in Seslerion variae (Aeschimann 
et al. 2004: 2010). Prvi dve našteti vrsti označujeta al-
pinsko združbo drobnega grušča v območjih z dolgo-
trajno snežno odejo, tretja pa novo asociacijo označuje 
predvsem fitogeografsko in deloma kaže na razvojno 
povezanost združb drobnega grušča z združbami ka-
mnitih alpskih trat in skalnih razpok. V enem sestoju 
je v veliki količini prisotna vrsta Sagina saginoides, ki 
kaže na krajevno boljše talne razmere, z obilico drobne 
preperine (sprstenine). 
5.3.4 Vrsta Geum reptans v Triglavskem pogorju
Tudi južnoevropsko montansko vrsto Geum reptans 
Aeschimann et al. (2004a: 758) uvrščajo med značil-
nice zveze Androsacion alpinae. V Julijskih Alpah so 
bila do zdaj znana le nahajališča na Mangartu in Ka-
njavcu (T. Wraber 2006, Zupan & Dakskobler 2007, 
slika 7). 8. 8. 2016 smo našli novo nahajališče te melišč-
ne vrste na zahodnih pobočjih Triglava, v krnici pod 
Vrhom Zelenic, na nadmorski višini 2060 m (Daksko-
bler 2017). Rastišče je ustaljeno melišče z različno ve-
likimi kosi kamenja in precej poraslo. Njegova vrstna 
sestava je v stolpcu 2, v preglednici 3. V stolpcu 1 te 
tabele je popis združbe na nahajališču na Temenu med 
Kanjavcem in Poprovcem (Zupan & Dakskobler 
2007), ki smo ga označili kot prehod med združbo tri-
glavske rože (Potentilletum nitidae) in sestojem asocia-
cije Papaveri julici-Thlaspietum rotundifolii. Sestoja 
pod Vrhom Zelenic ne moremo uvrstiti v nobeno od 
obeh navedenih asociacij. Označuje združbo melišč s 
precej vrstami kamnitih alpskih trat (iz asociacije Gen-
tiano terglouensis-Caricetum firmae). Začasno jo uvr-
ščamo v provizorno asociacijo Saxifrago sedoidis-Geu-
metum reptantis in v zvezo Thlaspion rotundifolii. 
Združbo poleg prevladujoče vrste Geum reptans ozna-
čujejo predvsem vrste iz rodu Saxifraga (S. sedoides, S. 
paniculata, S. aizoides, S. crustata, S. squarrosa) ter dve 
tipični meliščni vrsti: Festuca nitida in Poa minor. 
5.4 Sklepne misli
Razultat florističnega kartiranja v zahodnem delu Tri-
glava (Glava v Zaplanji, Vrh Zelenic)  in na prevalu 
Vrata med Veliko Zelnarico in Zadnjim Voglom so 
nova nahajališča treh v Sloveniji redkih vrst alpinskih 
melišč in/ali skalnih razpok, ki so uvrščene na rdeči 
seznam (Anon. 2002): Cerastium uniflorum (nova na-
hajališča v kvadrantu 9648/2), Crepis terglouensis 
(nova nahajališča v kvadrantih 9648/2 in 9648/4) ter 
Geum reptans (novo nahajališče v že znanem kvadran-
tu, 9648/2, novost v flori Triglava). Fitocenološko smo 
popisali združbe, v katerih uspevajo navedene vrste in 
ugotovili njihove netipične sestoje na drobnem grušču 
z drobno preperino (sprstenino), ki so prehodni med 
alpinskimi združbami skalnih razpok, melišč in sne-
žnih dolinic ter so lahko sukcesijska stopnja na površi-
nah, ki so bile pred desetletji precej dlje pokrita s sne-
gom. Na podlagi primerjave s podobnimi združbami 
iz Vzhodnih in Jugovzhodnih Alp smo opisali dve novi 
asociaciji Crepido terglouensis-Potentilletum nitidae in 
Saxifrago carniolicae-Cerastietum uniflorae. Čeprav so 
njuni sestoji v bližini precej obiskanih planinskih poti, 
ki vodijo proti Triglavu, jih vsako leto večji poletni tu-
ristični obisk za zdaj ne ogroža, saj drugih posegov v ta 
prostor tu ni in sta vrsti Cerastium uniflorum in Crepis 
terglouensis precej neopazni, še posebej v času, ko ne 
cvetita. Novo nahajališče vrste Geum reptans, ki tudi 
raste v netipični meliščni združbi (Saxifrago sedoidis-
-Geumetum reptantis nom. prov.), je v odmaknjenem 
brezpotju  v krnici pod Vrhom Zelenic in nanj za zdaj 
človek nima nobenega neposrednega vpliva.
Igor DAKSKoBLEr & BrAnKo ZUPAn: TWo nEW SCrEE PLAnT CoMMUnITIES In THE TrIgLAV MoUnTAInS
16 folia biologica et geologica 58/1 – 2017
We are extremely grateful to Prof. Dr. Andrej Martinčič 
for his determination of mosses. Iztok Sajko prepared 
Figure 1 for print. Academician Dr. Mitja Zupančič 
and Prof. Nanika Holz helped us with valuable im-
provements and corrections. The authors acknowledge 
the financial support from the Slovenian Research 
Agency (research core funding No. P1-0236). English 
translation by Andreja Šalamon Verbič.
ACKNOWLEDGEMENTS
REFERENCES – LITERATURA
Aeschimann, D., K.  Lauber, D. M. Moser & J.-P. Theurillat, 2004a: Flora alpina. Bd. 1: Lycopodiaceae-Apia-
ceae. Haupt Verlag, Bern, Stuttgart, Wien.
Aeschimann, D., K.  Lauber, D. M. Moser & J.-P.Theurillat, 2004b: Flora alpina. Bd. 2: Gentianaceae−Orchi-
daceae. Haupt Verlag, Bern, Stuttgart, Wien.
Aeschimann, D., K.  Lauber, D. M. Moser & J.-P.Theurillat, 2004c: Flora alpina. Bd. 3: Register. Haupt Verlag, 
Bern, Stuttgart, Wien. 
Anonymous, 2002: Pravilnik o uvrstitvi ogroženih rastlinskih in živalskih vrst v rdeči seznam. Uradni list RS 
82/2002.
Braun-Blanquet, J., 1964: Pflanzensoziologie. Grundzüge der Vegetationskunde. 3. Auflage. Springer, Wien – 
New york.
Buser, S., 2009: Geološka karta Slovenije 1: 250.000. Geological map of Slovenia 1: 250,000. Geološki zavod Slo-
venije, Ljubljana.
Cegnar, T., 1998: Temperatura zraka. In: J. Fridl, D. Kladnik, M. Orožen Adamič & D. Perko (eds.): Geografski 
atlas Slovenije. Država v prostoru in času. Državna založba Slovenije, Ljubljana, pp. 100–101.
Dakskobler, I., 2017: Dve botanični opombi pod črto: plazeča sretena (Geum reptans) pod Triglavom in venerini 
laski (Adiantum capillus-veneris) v grapi Volarje pri Seliščih. Proteus (Ljubljana) 79 (5): 216–223.
Dakskobler, I. & B. Surina, 2017: Phytosociological analysis of alpine swards and heathlands (pioneer patches) on 
ridges and peaks in the Julian Alps (NW Slovenia). Hacquetia (Ljubljana) 16 (1): 49–171. https://doi.org/10.1515/
hacq-2016-0022 
Eggensberger, P., 1994: Die Pflanzengesellschaften der subalpinen umd alpinen Stufe der Ammergauer Alpen und 
ihre Stellung in den Ostalpen. Ber. Bayer. Bot. Ges., Beihefte 8: 3–239.
Englisch, T., 1999: Multivariate Analysen zur Synsystematik und Standortsökologie der Schneebodenvegetation 
(Arabidetalia caerulae) in den Nördlichen Kalkalpen. Stapfia (Linz) 59: 1– 211 + Tables.
Englisch, T., M. Valachovič, L. Mucina, G. Grabherr, G.& T. Ellamuer, 1993: Thlaspietea rotundifolii. In: G. 
Grabherr & L. Mucina  (eds.): Die Pflanzengesellschaften Österreichs. Teil II: Natürliche waldfreie Vegetation, 
Gustav Fischer Verlag, Jena - Stuttgart - New york, pp. 276–342.
Gabrovec, M., M. Hrvatin, B. Komac, J. Ortar, M. Pavšek, M. Topole, M. Triglav Čekada & M. Zorn, 2014: 
Triglavski ledenik. Založba ZRC, Ljubljana.
Lovrenčak, F., 1998: Prsti. In: J. Fridl, D. Kladnik, M. Orožen Adamič & D. Perko (eds.): Geografski atlas Sloveni-
je. Država v prostoru in času. Državna založba Slovenije, Ljubljana, pp. 114–115.
Maarel van der, E., 1979: Transformation of cover-abundance values in phytosociology and its effects on communi-
ty similarity. Vegetatio 39 (2): 97–114. https://doi.org/10.1007/BF00052021 
Martinčič, A., T. Wraber, N. Jogan, A. Podobnik, B. Turk, B. Vreš, V. Ravnik, B. Frajman, S. Strgulc 
Krajšek, B. Trčak, T. Bačič, M. A. Fischer, K. Eler & B. Surina,  2007: Mala flora Slovenije. Ključ za dolo-
čanje praprotnic in semenk. Četrta, dopolnjena in spremenjena izdaja. Tehniška založba Slovenije, Ljubljana. 
Mucina, L., H. Bültmann, K. Dierssen, J-P. Theurillat, T. Raus, A. Čarni, K. Šumberová, W. Willner, J. 
Dengler, R. Gavilán García, M. Chytrý, M. Hájek, R. Di Pietro, D. Iakushenko, J. Pallas, F. J. A. 
Daniëls, E., Bergmeier, A. Santos Guerra, N. Ermakov, M. Valachovič, J. H. J.  Schaminée, T. Lysen-
ko, y. P. Didukh, S. Pignatti, J. S. Rodwell, J. Capelo, H. E. Weber, A. Solomeshch, P. Dimopoulos, C. 
Aguiar, S. M.  Hennekens & L. Tichý, 2016: Vegetation of Europe: hierarchical floristic classification system 
of vascular plant, bryophyte, lichen, and algal communities. Applied Vegetation Science 19, Suplement 1: 3–264. 
http://dx.doi.org/10.1111/avsc.12257 
Igor DAKSKoBLEr & BrAnKo ZUPAn: TWo nEW SCrEE PLAnT CoMMUnITIES In THE TrIgLAV MoUnTAInS
17folia biologica et geologica 58/1 – 2017
Novak, Š., 2015: Crepis terglouensis (Hacquet) Kerner. Notulae ad floram Sloveniae. Hladnikia (Ljubljana) 36: 
47–50.
Pignatti, E. & S. Pignatti, 2014: Plant Life of the Dolomites. Vegetation Structure and Ecology. Publication of the 
Museum of Nature South Tyrol Nr. 8, Naturmuseum  Südtirol, Bozen, Springer Verlag, Heidelberg. https://doi.
org/10.1007/978-3-642-31043-0
Pignatti, E. & S. Pignatti, 2016: Plant Life of the Dolomites.Vegetation Tables. Publication of the Museum of Na-
ture South Tyrol Nr. 11, Bozen, Springer Verlag, Heidelberg. https://doi.org/10.1007/978-3-662-48032-8
Podani, J., 2001: SyN-TAX 2000. Computer Programs for Data Analysis in Ecology and Systematics. User’s Manual, 
Budapest.
Praprotnik,  N., 1997: Crepis terglouensis (Hacq.) A. Kerner. In: Jogan N.: Nova nahajališča. Hladnikia (Ljubljana) 
8/9: 59.
Praprotnik, N., 2002: Triglavski dimek. Gea (Ljubljana) 12 (5): 65–66.
Seliškar, T., B. Vreš & A. Seliškar, 2003: FloVegSi 2.0. Fauna, Flora, Vegetation and Paleovegetation of Slovenia. 
Computer programme for arranging and analysis of biological data. Biološki inštitut ZRC SAZU, Ljubljana. 
Sørensen, Th., 1948: A method of establishing groups of equal amplitude in plant sociology based on similarity of 
species content. Det Kongelige Danske Videnskaberns Selskab, Biologiske Skrifter (København) 5 (4): 1–34.
Surina, B., 2005: Subalpinska in alpinska vegetacija Krnskega pogorja v Julijskih Alpah. Scopolia (Ljubljana) 57: 
1–122.
Sutter, R., 1969: Ein Beitrag zur Kenntnis der soziologischen Bindung süd-südostalpiner Reliktendemismen. Acta 
Bot. Croatica (Zagreb) 28: 349–365.
Šilc, U. & A. Čarni, 2012: Conspectus of vegetation syntaxa in Slovenia. Hacquetia (Ljubljana) 11 (1): 113–164. 
https://doi.org/10.2478/v10028-012-0006-1 
Theurillat, J.-P., 2004: Pflanzensoziologisches System. In: Aeschimann, D., K. Lauber, D. M. Moser & J.-P. Theuri-
llat: Flora alpina 3: Register. Haupt Verlag, Bern, Stuttgart, Wien, pp. 301–313.
Vidic, N. J., T. Prus, H. Grčman, M. Zupan, A. Lisec, T. Kralj, B, Vrščaj, J. Rupreht, M. Šporar, M. Suha-
dolc, R. Mihelič, R. & F. Lobnik, 2015: Tla Slovenije s pedološko karto v merilu 1: 250 000. Soils of Slovenia 
with soil map 1: 250 000. European Union & University of Ljubljana, Luxemburg, Ljubljana. 
Wraber, T., 1972: Contributo alla conoscenza della vegetazione pioniere (Asplenietea rupestria e Thlaspeetea rotun-
difolii) delle Alpi Giulie. Tesi di laurea. Università degli Studi di Trieste, Facoltà di Scienze, Trieste (Doktorska 
naloga, 81 pp.)
Wraber, T., 1990: Sto znamenitih rastlin na Slovenskem. Prešernova družba, Ljubljana.
Wraber, T., 2006: 2 x Sto alpskih rastlin na Slovenskem. Prešernova družba, Ljubljana.
Zupan, B. & I. Dakskobler,  2007: Geum reptans L. Notulae ad floram Sloveniae 83. Hladnikia (Ljubljana) 20: 
36–38.
Zupančič, B., 1998: Padavine. In: J. Fridl, D., Kladnik, M. Orožen Adamič & D. Perko (eds.): Geografski atlas Slo-
venije. Država v prostoru in času. Državna založba Slovenije, Ljubljana, pp. 98–99.
Figures 7–14:  Photo/Foto: I. Dakskobler
Igor DAKSKoBLEr & BrAnKo ZUPAn: TWo nEW SCrEE PLAnT CoMMUnITIES In THE TrIgLAV MoUnTAInS
18 folia biologica et geologica 58/1 – 2017
Figure 7: Crepis terglouensis
Slika 7: Triglavski dimek (Crepis terglouensis)
Figure 8: Stand of the association Crepido terglouensis-Potentilletum nitidae
Slika 8: Sestoj asociacije Crepido terglouensis-Potentilletum nitidae
Igor DAKSKoBLEr & BrAnKo ZUPAn: TWo nEW SCrEE PLAnT CoMMUnITIES In THE TrIgLAV MoUnTAInS
19folia biologica et geologica 58/1 – 2017
Figure 9: Glava nad Zaplanjo under Mt. Triglav, the area where we have recorded stands of the association Crepido terglouen-
sis-Potentilletum nitidae 
Figure 9: Glava nad Zaplanjo, območje, kjer smo popisovali sestoje asociacije Crepido terglouensis-Potentilletum nitidae
Figure 10: Cerastium uniflorum
Slika 10: Enocvetna smiljka (Cerastium uniflorum)
Igor DAKSKoBLEr & BrAnKo ZUPAn: TWo nEW SCrEE PLAnT CoMMUnITIES In THE TrIgLAV MoUnTAInS
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Figure 11: Stand of the association Saxifrago carniolicae-Cerastietum uniflorae
Slika 11: Sestoj asociacije Saxifrago carniolicae-Cerastietum uniflorae
Figure 12: Plateau west from Mt. Triglav, area, where we have recorded stands of the association Saxifrago carniolicae-
Cerastietum uniflorae
Slika 12: Planota zahodno od Triglava, kjer smo popisovali sestoje asociacije Saxifrago carniolicae-Cerastietum uniflo-
rae
Igor DAKSKoBLEr & BrAnKo ZUPAn: TWo nEW SCrEE PLAnT CoMMUnITIES In THE TrIgLAV MoUnTAInS
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Figure 14: Stand of the association Saxifrago sedoidis-Geumetum raptantis
Slika 14: Sestoj asociacije Saxifrago sedoidis-Geumetum raptantis
Figure 13: Geum reptans under Mt. Triglav
Slika 13: Plazeča sretena (Geum reptans) pod Triglavom
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table 1 : Crepido terglouensis-Potentilletum nitidae and Saxifrago carniolicae-Cerastietum uniflorae
Preglednica 1: Crepido terglouensis-Potentilletum nitidae in Saxifrago carniolicae-Cerastietum uniflorae
Number of relevé
(Zaporedna številka popisa) 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21
Database number of relevé 
(Delovna številka popisa)    20
01
47
25
80
65
25
76
26
26
32
98
26
34
03
26
34
04
25
76
13
25
76
23
26
25
76
26
34
06
25
76
64
26
34
08
26
34
09
25
76
14
25
76
17
25
80
67
26
25
77
25
80
68
25
80
73
25
80
71
26
25
78
Author of the relevé
(Avtor popisa) T
W ID ID
BZ ID ID ID ID
BZ
ID
BZ ID ID ID
BZ ID ID ID
BZ
ID
BZ ID ID ID ID ID ID
Elevation in m (Nadmorska 
višina v m)     25
20
25
35
23
20
21
92
25
40
25
41
24
24
25
09
24
60
25
40
23
20
25
40
25
45
24
50
25
30
25
10
25
20
25
00
25
15
25
05
25
20
Aspect (Lega)                                  SE N
W N
E
SW N
W
SW SW S
E W SS
W
N
E
SE SE S SS
E
SE N
W N
N
W
W
SE N
W
Slope in degree
(Nagib v stopinjah) 10 3 5 0-3 10 5 5 2 30 5 3 10 5 15 15 1 20 25 35 5 20
Parent material
(Matična podlaga) Gr Gr Gr Gr Gr Gr Gr Gr Gr Gr Gr Gr Gr Gr Gr Gr Gr Gr Gr Gr Gr
Soil (Tla) Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Re Li Li Li Li Li
Stoniness in % (Kamnitost v %) 70 80 70 90 100 100 100 60 20 100 70 100 100 70 70 5 20 70 70 20 90
Cover of herb layer in %
(Zastiranje zeliščne plasti v %)    E1 30 30 30 30 40 40 30 40 80 30 30 30 40 30 30 90 90 30 30 80 30
Cover of moss layer in %
(Zastiranje mahovne plasti v %)     E0 . . . . . . . . . . . . . . . 10 60 . . . .
Number of species (Število vrst)    18 14 10 12 19 15 7 3 5 6 3 12 18 15 11 13 6 7 7 12 6
Relevé area
(Velikost popisne ploskve)   m
2 10 10 4 5 10 10 3 4 2 10 3 10 10 10 5 2 10 2 2 10 10
Date of taking relevé
(Datum popisa)          
7/
23
/1
96
3
9/
1/
20
15
7/
31
/2
01
5
8/
23
/2
01
6
8/
24
/2
01
6
8/
24
/2
01
6
8/
1/
20
15
8/
1/
20
15
8/
24
/2
01
6
8/
24
/2
01
6
7/
31
/2
01
5
8/
24
/2
01
6
8/
24
/2
01
6
8/
1/
20
15
8/
1/
20
15
9/
1/
20
15
8/
24
/2
01
6
9/
1/
20
15
9/
1/
20
15
9/
1/
20
15
8/
24
/2
01
6
Locality (Nahajališče)
K
re
da
ri
ca
G
la
va
 v
 Z
ap
la
nj
i 
St
an
ič
ev
 d
om
 
V
ra
ta
-Z
el
na
ri
ca
 
G
la
va
 v
 Z
ap
la
nj
i 
G
la
va
 v
 Z
ap
la
nj
i 
G
la
va
 v
 Z
ap
la
nj
i 
K
re
da
ri
ca
G
la
va
 v
 Z
ap
la
nj
i 
G
la
va
 v
 Z
ap
la
nj
i 
D
ov
šk
a 
vr
at
ca
 
G
la
va
 v
 Z
ap
la
nj
i 
G
la
va
 v
 Z
ap
la
nj
i 
G
la
va
 v
 Z
ap
la
nj
i 
G
la
va
 v
 Z
ap
la
nj
i 
Z
ap
la
nj
a-
M
or
be
gn
a
G
la
va
 v
 Z
ap
la
nj
i 
Z
ap
la
nj
a-
M
or
be
gn
a 
Z
ap
la
nj
a-
Pl
em
en
ic
e
Z
ap
la
nj
a-
Pl
am
en
ic
e 
G
la
va
 v
 Z
ap
la
nj
i 
Quadrant (Kvadrant)
96
49
/1
96
48
/2
96
49
/1
96
48
/4
96
48
/2
96
48
/2
96
48
/2
96
49
/1
96
48
/2
96
48
/2
96
49
/1
96
48
/2
96
48
/2
96
48
/2
96
48
/2
96
49
/1
96
48
/2
96
49
/1
96
49
/1
96
49
/1
96
48
/2
Coordinate GK y (D-48) m
41
18
86
41
00
68
41
27
23
40
75
39
41
00
65
41
00
57
41
00
88
41
18
65
41
00
15
41
00
82
41
32
36
41
01
73
41
01
78
41
00
48
41
02
13
41
04
48
41
01
03
41
03
69
41
06
02
41
04
48
41
01
19
Coordinate GK X (D-48) m
51
37
96
4
51
37
60
4
51
38
74
8
51
33
84
9
51
37
60
4
51
37
59
7
51
37
33
4
51
37
88
7
51
37
49
9
51
37
59
7
51
38
75
7
51
37
64
4
51
37
72
6
51
37
45
4
51
37
62
8
51
37
75
5
51
37
66
7
51
37
84
2
51
38
03
3
51
37
75
4
51
37
67
6
Diagnostic species of the syntaxa (Diagnostične vrste sintaksonov)
Pr
. 1
-1
5
Fr
- 1
-1
5
Pr
. 1
6-
21
Fr
. 1
6-
21
PS Potentilla nitida E1 2 2 1 1 1 2 3 3 4 3 2 3 3 2 2 . . . . . . 15 100 0 0
TR1 Crepis terglouensis E1 1 1 2 3 1 + 2 1 1 + 1 2 1 1 1 . . . . . . 15 100 0 0
PC Eritrichium nanum E1 1 + . . + + + . . + . + 1 . . . . . . . . 8 53 0 0
TR1 Alyssum ovirense E1 1 + . . + + . . . 1 . + + . . . . . . . . 7 47 0 0
PC Minuartia cherlerioides E1 . 1 . . 1 1 . . . + . + + . . . . . . . . 6 40 0 0
AA Cerastium uniflorum E1 + . . . + . . . . . . . . . . 1 2 3 2 4 1 2 13 6 100
TR1 Saxifraga sedoides E1 . . . . . . . . . . . . . . . 1 + + + + 2 0 0 6 100
Cfir Saxifraga exarata subsp. carniolica E1 . . . . . . . . . . . . . . . + . + + 1 . 0 0 4 67
TR1 Thlaspion rotundifolii
Papaver julicum E1 + . . . . . . . . . . + + . . . . 1 + . 1 3 20 3 50
Thlaspi cepeaefolium (T. rotun-
difolium, Noccaea rotundifolia) E1 . . . + . . . . . . . . . . + . . . . . . 2 13 0 0
TR2 Thlaspieetalia rotundifolii
Poa minor E1 . . . . . . . . + . . + + 1 + . . . . . . 5 33 0 0
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Achillea atrata E1 + . . . . . . . . . . . . 1 + + . . . . . 3 20 1 17
Cerastium carinthiacum subsp. 
carinthiacum E1 . . + 1 . . . . 1 . . . . 1 . . . . . . . 4 27 0 0
Moehringia ciliata E1 . . . . . . . . . . . . . + . . . . + . . 1 7 1 17
Armeria alpina E1 . . + . . . . . . . . . . . . . . . . . . 1 7 0 0
AC Arabidetalia caeruleae
Salix retusa E1 + . . + + + . . . . . . . . . . . . . . . 4 27 0 0
Salix serpyllifolia E1 . + . . . . . . . . . . . + . + + . . . . 2 13 2 33
CD Carex capillaris E1 . . . . + . . . . . . . + . + . . . . . . 3 20 0 0
Carex ornithopodoides E1 . . . . + + . . . . . . . . . . . . . . . 2 13 0 0
Carex parviflora E1 . . . . . . . . . . . . . . . + . . . . . 0 0 1 17
SH Sagina saginoides E1 . . . . . . . . . . . . . . . 4 . . . 1 . 0 0 2 33
 Saxifraga androsacea E1 . . . . . . . . . . . . . . . 1 . . . + . 0 0 2 33
DH Sesleria ovata E1 + . . . . . . . . . . . . . . . . . . . . 1 7 0 0
TR3 Thlaspietea rotundifolii
Taraxacum alpinum E1 . r . . + . . . . . . . + + . . . . . . . 4 27 0 0
Saxifraga oppositifolia s.str. E1 + . . . + . . . . . . . . . . . . . 1 . . 2 13 1 17
Linaria alpina E1 . . . . . . . . . . . + . . . . . . . . . 1 7 0 0
Festuca nitida E1 . . . . . . . . . . . . . + . . . . . . . 1 7 0 0
Arabis alpina E1 . . . . . . . . . . . . . . . . . . . . + 0 0 1 17
PC Potentilletalia caulescentis
Valeriana elongata E1 + + . . + . . . . . . . . . . . . . . . . 3 20 0 0
Festuca alpina E1 . . . . . + . . + . . . . . . . . . . 1 . 2 13 1 17
Campanula cochleariifolia E1 . . . . . + . . . . . . . + . . . . . . . 2 13 0 0
Saxifraga paniculata E1 . . . . . . . . . . . . 1 . + . . . . . . 2 13 0 0
Petrocallis pyrenaica E1 . . . . . . . . . . . . . + + . . . . + . 2 13 1 17
Cfir Caricion firmae
Minuartia verna E1 1 . + + . . + 1 . . . + + . . . + . . + . 7 47 2 33
Silene acaulis E1 + 2 + 1 + . . . . . . . . . . . . + . + . 5 33 2 33
Carex firma E1 1 + 1 1 1 1 . . . . 1 . . . . . . . . . . 7 47 0 0
Minuartia sedoides E1 1 2 + 1 . . + . . . . . + . . . . 1 . . . 6 40 1 17
Phyteuma sieberi E1 . + . + + . . . . . . + + + + . . . . . . 7 47 0 0
Sesleria sphaerocephala E1 . . . . + + + . . + . . + . . . . . . . . 5 33 0 0
Gentiana terglouensis E1 . 1 . . + + . . . . . . . + . . . . . . . 4 27 0 0
Festuca quadriflora E1 . . + . . . . . . . . . . . . + . . . + . 1 7 2 33
Saussurea pygmaea E1 . . . . . . . . . . . . . . + . . . . . . 1 7 0 0
Veronica aphylla E1 . . . . . . . . . . . . . . . . . . . + . 0 0 1 17
OE Oxytropido-Elynion
Arenaria ciliata E1 . . . . + + . . . . . . . . . . . . . . . 2 13 0 0
Erigeron uniflorus E1 . . . . . . . . . . . . + . . + . . . . . 1 7 1 17
Lloydia serotina E1 . . . . . . . . . . . . + . . . . . . . . 1 7 0 0
ES Elyno-Seslerietea
PAT Poa alpina E1 + + + + . + . . . . . + . + . 1 4 1 + 1 1 7 47 6 100
Myosotis alpestris E1 + . . . . . . . . . . + + + + . . . . . 1 5 33 1 17
Polygonum viviparum E1 + . . 1 + + + . . . . . + . . . . . . . . 6 40 0 0
ML Mosses (Mahovi)
Syntrichia norvegica E0 . . . . . . . . . . . . . . . 1 4 . . . . 0 0 2 33
Bryum sp. E0 . . . . . . . . . . . . . . .  + . . . . . 0 0 1 17
legend - legenda
ID Igor Dakskobler
BZ Branko Zupan
TW Tone Wraber
Gr Gravel - grušč
Li Lithosol - kamnišče
Pr. Presence  (number of relevés in which the species is presented) - število popisov, v katerih se pojavlja vrsta 
Fr. Frequency in % - frekvenca v % 
Re Rendzina - rendzina
PS Physoplexido-Saxifragion petraeae
AA Androsacion alpinae 
CD Caricetalia davallianae
SH Salicetea herbaceae
DH Drabion hoppeanae
PAT Poo alpinae-Trisetetalia
Number of relevé
(Zaporedna številka popisa) 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 Pr
. 1
-1
5
Fr
- 1
-1
5
Pr
. 1
6-
21
Fr
. 1
6-
21
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24 folia biologica et geologica 58/1 – 2017
table 2: groups of diagnostic species in the stands of syntaxa Crepido terglouensis-Potentilletum nitidae (ctPn) and Saxifrago 
carniolicae-Cerastietum uniflorae (Sccu), relative frequencies
Preglednica 2: Skupine diagnostičnih vrst v sestojih sintaksonov Crepido terglouensis-Potentilletum nitidae (ctPn) and Saxifrago 
carniolicae-Cerastietum uniflorae (Sccu), relativne frekvence
Successive number (Zaporedna številka) 1 2
Sign for syntaxa (Oznaka sintaksona) CtPn ScCu
Number of relevés (Število popisov) 15 6
Arabidetalia caeruleae 8,304 26,01
Thlaspion rotundifolii 16,07 18,01
Thlaspieetalia rotundifolii 8,333 4,002
Thlaspietea rotundifolii 4,762 4,002
Potentilletalia caulescentis 23,81 4,002
Caricion firmae 25,6 24,01
Oxytropido-Elynion 2,381 2,001
Elyno-Seslerietea 10,71 14,01
Moses (Mahovi) 0 4,002
Total (Skupaj) 100 100
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table 3: communities with Geum reptans in the triglav Mts.
Preglednica 3: Združbi z vrsto Geum reptans v triglavskem pogorju
Number of relevé (Zaporedna številka popisa)          1 2
Database number of relevé (Delovna številka popisa)    217413 262400  
Author of the relevé (Avtor popisa) IDBZ ID
Elevation in m (Nadmorska višina v m)     2470 2060
Aspect (Lega)                                  SSE SW
Slope in degrees (Nagib v stopinjah)           5 25
Parent material (Matična podlaga) Gr Gr
Soil (Tla) Li Li
Stoniness in % (Kamnitost v %) 20 80
Cover of herb layer in % (Zastiranje zeliščne plasti v %): E1 80 35
Number of species (Število vrst)               13 31
Relevé area (Velikost popisne ploskve)   m2 5 100
Date of taking relevé (Datum popisa)          7/26/2007 8/8/2016
Locality (Nahajališče) Poprovec-Teme Vrh Zelenic-Skok 
Quadrant (Kvadrant) 9648/2 9648/2
Coordinate GK y (D-48) m 408402 409533
Coordinate GK X (D-48) m 5135506 5138047
AA Androsacion alpinae Pr.
Geum reptans E1 4 3 2
AC Arabidetalia caeruleae
Doronicum glaciale E1 . + 1
Galium noricum E1 . + 1
Salix retusa E1 . + 1
TR1 Thlaspion rotundifolii
Thlaspi cepeaefolium (T. rotundifolium, Noccaea rotundifolia) E1 + + 2
Alyssum ovirense E1 + . 1
Papaver julicum E1 + . 1
Saxifraga sedoides E1 . 1 1
TR2 Thlaspieetalia rotundifolii
Achillea atrata E1 + + 2
Moehringia ciliata E1 + + 2
Festuca nitida E1 . 1 1
Poa minor E1 . 1 1
TR3 Thlaspietea rotundifolii
Saxifraga oppositifolia s.str. E1 + . 1
Taraxacum alpinum agg. E1 + . 1
Arabis alpina E1 . + 1
PS Physoplexido-Saxifragion petraeae
Potentilla nitida E1 1 . 1
Paederota lutea E1 . + 1
Saxifraga crustata E1 . + 1
Saxifraga squarrosa E1 . + 1
PC Potentilletalia caulescentis
Saxifraga paniculata E1 + 1 2
Festuca alpina E1 + . 1
Campanula cochleariifolia E1 . + 1
CA Caricion austroalpinae
 Arabis vochinensis E1 . + 1
Cfir Caricion ferrugineae
 Gentiana pumila E1 . + 1
OE Oxytropido-Elynion
 Arenaria ciliata E1 . + 1
Cfir Caricion firmae
Minuartia verna E1 + + 2
Carex firma E1 . + 1
Dryas octopetala E1 . + 1
Pedicularis rostratocapitata E1 . + 1
Phyteuma sieberi E1 . + 1
Silene acaulis E1 . + 1
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26 folia biologica et geologica 58/1 – 2017
ES Elyno-Seslerietea
Polygonum viviparum E1 . 1 1
Aster bellidiastrum E1 . + 1
Pedicularis verticillata E1 . + 1
Achillea clavenae E1 . + 1
MC Montio-Cardaminetea
Saxifraga aizoides E1 . 1 1
PAT Poo alpinae-Trisetetalia
Poa alpina E1 + 1 2
legend - legenda
ID Igor Dakskobler
BZ Branko Zupan
Gr Gravel - grušč
Li Lithosol - kamnišče
Number of relevé (Zaporedna številka popisa)          1 2 Pr.
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27folia biologica et geologica 58/1 – 2017
appendix 1: Synthetic table of communities with Potentilla nitida and/or Crepis terglouensis in eastern and Southeastern alps
Dodatek 1: Sintezna tabela združb z vrstama Potentilla nitida in/ali Crepis terglouensis v Vzhodnih in Jugovzhodnih alpah
Successive number (Zaporedna številka)  1 2 3 4
Number of relevés (Število popisov) 25 15 33 1
Sign for syntaxa (Oznaka sintaksonov) CT CTPN PN CTPN
TR1 Thlaspion rotundifolii
Crepis terglouensis E1 100 100 . 1
Galium megalospermum E1 76 . . .
Arabis bellidifolia subsp. bellidifolia E1 52 . . .
Pritzelago alpina subsp. alpina (Hutchinsia alpina) E1 44 . 3 .
Saxifraga aphyla E1 12 . . .
Thlaspi cepeaefolium (T. rotundifolium, Noccaea rotundifolia) E1 8 13 . .
Alyssum ovirense E1 . 47 15 1
Papaver julicum E1 . 20 12 .
Saxifraga sedoides E1 . . 15 .
Festuca rupicaprina E1 . . 8 .
TR2 Thlaspietalia rotundifolii
Poa cenisia E1 24 . . .
Poa minor E1 16 33 . .
Linaria alpina E1 8 7 6  +
Doronicum grandiflorum E1 4 . . .
Cerastium carinthiacum E1 . 27 3  +
Achillea atrata E1 . 20 3 .
Armeria alpina E1 . 7 .  +
Moehringia ciliata E1 . 7 . .
Sedum atratum E1 . . 3 .
AC Arabidetalia caeruleae
 Salix serpyllifolia E1 16 13 15 1
MC Saxifraga aizoides E1 12 . 3 .
  Ranunculus alpestris E1 4 . . .
 Salix retusa E1 . 27 24 .
CD Carex capillaris E1 . 20 9 .
AA Cerastium uniflorum E1 . 13 3 .
 Carex ornithopodoides E1 . 13 3 .
DH Sesleria ovata E1 . 7 . .
JT Carex fuliginosa E1 . . 30 .
Ranunculus traunfellneri E1 . . 12 .
Doronicum glaciale E1 . . 6 .
Salix reticulata E1 . . 3 .
Trifolium pallescens E1 . . 3 .
TR3 Thlaspietea rotundifolii
Athamantha cretensis E1 36 . . .
Trisetum distichophyllum E1 12 . . .
Gypsophila repens E1 8 . . .
Silene vulgaris subsp. glareosa E1 8 . . .
Taraxacum alpinum E1 . 27 . .
Saxifraga oppositifolia s.str. E1 . 13 70 .
Festuca nitida E1 . 7 . .
Pritzelago alpina subsp. brevicaulis E1 . . 3 .
PS Physoplexido-Saxifragion petraeae
Potentilla nitida E1 . 100 97 2
Saxifraga squarrosa E1 . . 73 .
Saxifraga crustata E1 . . 27 .
Campanula zoysii E1 . . 24 .
Paederota lutea E1 . . 3 .
PC Potentilletalia caulescentis
Campanula cochleariifolia E1 56 13 42  +
Eritrichium nanum E1 . 53 52  +
Minuartia cherlerioides E1 . 40 55 .
Valeriana elongata E1 . 20 3 .
Festuca alpina E1 . 13 79 .
Petrocallis pyrenaica E1 . 13 42  +
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Saxifraga paniculata E1 . 13 61 .
Draba tomentosa E1 . . 45 .
Saxifraga exarata subsp. carniolica E1 . . 24 .
Potentilla clusiana E1 . . 21 1
Arabis bellidifolia subsp. stellulata E1  . 18 .
Saxifraga burseriana E1 . . 12 .
Androsace helvetica E1 . . 6 .
Primula auricula E1 . . 3 .
Dianthus sylvestris E1 . . 3 .
CF Caricion ferrugineae
 Viola biflora E1 40 . 3 .
Heliosperma pusillum E1 12 . . .
Carex ferruginea E1 4 . . .
Cfir Caricion firmae
Festuca quadriflora E1 76 7 6 .
Carex firma E1 56 47 45  +
Dryas octopetala E1 32 . 3 .
Minuartia verna E1 24 47 61  +
Silene acaulis E1 24 33 30  +
Minuartia sedoides E1 12 40 82  +
Crepis kerneri E1 8 . . .
Saxifraga caesia E1 8 . 3 .
Phyteuma sieberi E1 . 47 . .
Sesleria sphaerocephala E1 . 33 70 .
Gentiana terglouensis E1 . 27 52  +
Saussurea pygmaea E1 . 7 6 .
Helianthemum alpestre E1 . . 18 .
Gentiana orbicularis E1 . . 12 .
Salix alpina E1 . . 9  +
Oxytropis neglecta E1 . . 6 .
Pedicularis rostratocapitata E1 . . 6 .
Carex rupestris E1 . . 3 .
Pedicularis rosea E1 . . 3 .
Veronica aphylla E1 . . 3 .
OE Oxytropido-Elynion
Arenaria ciliata E1  13 33 .
Lloydia serotina E1 . 7 15 .
Erigeron uniflorus E1 . 7 3 .
ES Elyno-Seslerietea
Polygonum viviparum E1 40 40 39 .
Sesleria caerulea E1 28 . 9 .
Euphrasia salisburgensis E1 12 . 3 .
Carex sempervirens E1 4 . 3 .
Pedicularis oederi E1 4 . . .
Galium anisophyllon E1 4 . . .
PaT Poa alpina E1 . 47 55 .
Myosotis alpestris E1 . 33 3 .
Draba aizoides E1 . . 33 .
Erigeron glabratus E1 . . 18 .
Achillea clavenae E1 . . 15 .
Leontopodium alpinum E1 . . 6 .
Agrostis alpina E1 . . 3 .
Aster bellidiastrum E1 . . 3 .
EP Erico-Pinetea
 Asperula aristata E1 8 . . .
 Rhodothamnus chamaecistus E1 . . 3 .
O other species (Druge vrste)
 Saxifraga sp. E1 . . 3 .
 Thymus sp. E1 . . 3 .
ML Mosses and lichens (Mahovi in lišaji)
Tortella tortuosa E0 8 . . .
Distichum capillaceum E0 8 . . .
Successive number (Zaporedna številka)  1 2 3 4
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Ctenidium molluscum E0 4 . . .
Orthotecium rufescens E0 4 . . .
Cladonia sp. E0 4 . . .
Pohlia cruda E0 4 . . .
Encalypta streptocarpa E0 4 . . .
Hypnum bambergeri E0 4 . . .
Ctenidium procerrimum E0 4 . . .
legend - legenda
CT Crepidetum terglouensis (Eggensbeger 1994, Table 6, Columns 85–111)
CTPN Crepido-Potentilletum nitidae, this article, Table 1, Coliumns 1-15
PN Potentilletum nitidae, T. Wraber 1972, Table  3
CTPN Crepido-Potentilletum nitidae, Sutter (1969: 353) 
MC Montio-Cradaminetea
CD Caricetalia davallianae
AA Androsacion alpinae 
DH Drabion hoppeanae
PAT Poo alpinae-Trisetetalia
JT Juncetea trifidi
Successive number (Zaporedna številka)  1 2 3 4
appendix 2: communities with Cerastium uniflorum and Saxifraga sedoides in Se alps
Preglednica 2: Združbi z vrstama Cerastium uniflorum in Saxifraga sedoides v JV alpah
Successive number (Zaporedna številka) 1 2
Number of relevés (Število popisov) 15 6
Sign for syntaxa (Oznaka sintaksonov) Ss ScSu
AC Arabidetalia caeruleae
AA Cerastium uniflorum E1 47 100
MC Saxifraga stellaris subsp. alpigena E1 13  
 Soldanella minima E1 13 .
AA Doronicum clusii E1 7 .
 Ranunculus alpestris E1 7 .
DH Draba hoppeana E1 7 .
 Salix serpyllifolia E1 . 33
 Saxifraga androsacea E1 . 33
SH Sagina saginoides E1 . 33
 Carex parviflora E1 . 17
TR1 Thlaspion rotundifolii
Saxifraga sedoides E1 100 100
Pritzelago alpina subsp. alpina E1 60 .
Thlaspi cepeaefolium (T. rotundifolium, Noccaea rotundifolia) E1 53 .
Papaver rhaeticum E1 27 .
Arabis bellidifolia subsp. bellidifolia E1 20 .
Papaver julicum E1 . 50
TR2 Thlaspietalia rotundifolii
Poa minor E1 67 .
Arabis alpina E1 60 17
Moehringia ciliata E1 47 17
Cerastium carinthiacum E1 20 .
Achillea oxyloba E1 13 .
Achillea atrata E1 . 17
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Successive number (Zaporedna številka) 1 2
TR3 Thlaspietea rotundifolii
Saxifraga oppositifolia s.str. E1 13 17
Pritzelago alpina subsp. brevicaulis E1 7 .
Taraxacum alpinum E1 7 .
PC Potentilletalia caulescentis
Festuca alpina E1 20 17
Campanula cochleariifolia E1 13 .
Androsace hausmanii E1 7 .
Phyteuma sieberi E1 7 .
Valeriana elongata E1 7 .
Cystopteris regia E1 7 .
Petrocallis pyrenaica E1 . 17
Saxifraga exarata subsp. carniolica E1 . 67
Cfir Caricion firmae
Minuartia sedoides E1 20 17
Sesleria sphaerocephala E1 13 .
Minuartia verna E1 7 33
Silene acaulis E1 7 33
OE Erigeron uniflorus E1 7 17
Festuca quadriflora E1 . 33
Veronica aphylla E1 . 17
ES Elyno-Seslerietea
PAT Poa alpina E1 27 100
 Polygonum viviparum E1 13 .
CF Heliosperma pusillum E1 7 .
 Myosotis alpestris E1 . 17
M Mosses and lichens (Mahovi in lišaji)
Musci spp. E0 13 .
Grimmia sp. E0 7 .
Marchantia sp. E0 7 .
Syntrichia norvegica E0 . 33
Bryum sp. E0 . 17
legend - legenda
Ss Saxifragetum sedoidis, E. & S. Pignatti, 2016, Association Table 9.2
ScCu  Saxifrago carniolicae-Cerastietum uniflorae, this article
MC Montio-Cradaminetea
CD Caricetalia davallianae
AA Androsacion alpinae 
DH Drabion hoppeanae
PAT Poo alpinae-Trisetetalia
SH Salicetea herbaceae
CF Caricion ferrugineae