CONTRIBUTION TO THE KNOWLEDGE OF LEPIDOPTERA FAUNA 
OF THE KARST
Radovan ŠTANTA1, Matjaž ZADRGAL2
1 Miren 163, SI-5291 Miren, Slovenija, e-mail: radovan.stanta@gmail.com
2 Pod Lazami 53, Vrtojba, SI-5290 Šempeter pri Gorici, Slovenija, 
e-mail: matjaz.zadrgal@si.mahle.com
Abstract - The Macrolepidoptera fauna of Komenski and Sežanski Kras region has
been under our examination during the years 1985-2015, first by random checks and
later with planned, daily and night time inventory. We recorded 755 species of
Macrolepidoptera during 422 inventories and records of 36 additional species were
found in other sources, altogether 791 species or 52 % of Slovenian Macrolepidoptera
fauna are present in the region. The families Hepialidae, Psychidae, Limacodidae,
Heterogynidae, Zygaenidae, Sesiidae, Cossiidae, Thyrididae and families from Bombi-
coidea to Noctuidea are classified among Macrolepidoptera according to the old clas-
sification for older literature data comparability. 
We have devided the investigated area into five zones – sub-areas, in accordance
with the various ecological characteristics. We are stating the presence of species,
their ecological characteristics, frequency rating and interesting informations from
our own observations for each sub-area in the list of species. In rare and interesting
species of the area we mention the exact data of the finds. Some of more interesting
species are presented with comments and photos. We compare our results with other
known data on the Karst fauna, as well as those for Gorizia and Trieste karst fauna of
the neighboring Friuli - Venezia Giulia region. In addition we also present the differ-
ences between the sub-areas, especially between the warmer north-west and the colder
south-east part of the area. Furthermore we highlight the phenomenon of the Karst
plateau diversity, where in a small area xerophilous, mesophilous and hygrophilous
species are met due to Karst's heterogeneous structure of microhabitats, where hy-
grophilous species retain in sinkholes, mesophilous on their edges, and xerophilous
species on the slopes. The importance of nature protection of the area is also empha-
sized. The finding of the geometrid Eupithecia limbata Staudinger, 1879 (Geometridae)
in the Slovenian fauna is being mentioned for the first time. 
69
ACTA ENTOMOLOGICA SLOVENICA
LJUBLJANA, DECEMBER 2016       Vol. 24, øt. 2: 69–150
KEY WORDS: Lepidoptera, Macrolepidoptera, Fauna, List of Species, Slovenia, Karst,
Komenski kras, Sežanski kras
Izvleček – PRISPEVEK K POZNAVANJU FAVNE METULJEV (LEPIDOPTERA)
KRASA
Predstavljamo favno makro metuljev (Macrolepidoptera) Komenskega in Sežan-
skega krasa, ki smo jo proučevali v obdobju 1985-2015, najprej z naključnimi, kasneje
pa z načrtnimi, dnevnimi in nočnimi popisi metuljev. Na skupno 422 popisih smo za-
beležili 755 vrst makro metuljev, dodatnih 36 vrst pa predstavljajo še podatki iz
drugih virov, kar skupaj znese 791 vrst ali 52 % slovenske favne makro metuljev. Za-
radi primerljivosti podatkov iz starejše literature so v prispevku po stari razdelitvi
med makro metulje uvrščene tudi družine Hepialidae, Psychidae, Limacodidae, He-
terogynidae, Zygaenidae, Sesiidae, Cossiidae, Thyrididae ter družine od Bombicoidea
do Noctuidea. 
V prispevku smo obravnavano območje, zaradi različnih ekoloških značilnosti,
razdelili na pet con - podobmočij. Pri seznamu vrst navajamo prisotnost vrst v posa-
mezni coni, ekološke značilnosti vrste, ocenjeno pogostost ter zanimive informacije
iz lastnih opazovanj. Pri redkih in zanimivejših vrstah v območju navajamo dejanske
podatke o najdbah. Nekaj zanimivejših vrst je predstavljenih s komentarji in fotogra-
fijami. V prispevku primerjamo rezultate opazovanj z drugimi znanimi podatki o
favni Krasa, kot tudi s favno Goriškega in Tržaškega krasa sosednje Furlanije -
Julijske krajine. Prav tako predstavljamo razlike v favni med opredeljenimi conami,
predvsem med toplejšim severozahodnim in hladnejšim jugovzhodnim delom območja.
Izpostavljamo tudi fenomen raznolikosti kraške planote, kjer na majhnem območju
srečamo kserotermne, mezofilne in higrofilne vrste, saj Kras s kraškimi pojavi nudi
zelo heterogeno strukturo mikrohabitatov, kjer se v vrtačah zadržujejo higrofilne, na
njihovem robu mezofilne, na pobočjih pa kserotermne vrste metuljev. Poudarjen je
tudi naravovarstveni pomen območja. Prvič za favno Slovenije omenjamo najdbo
pedica Eupithecia limbata Staudinger, 1879 (Geometridae).
KLJUČNE BESEDE: Lepidoptera, Macrolepidoptera, favna, seznam vrst, Slovenija, Kras,
Komenski kras, Sežanski kras
Introduction
Parent Kras (Karst) has always been interesting for researchers of different
disciplines, also for entomologists, among whom were quite some of lepidopterologists.
Trieste as a major economic and intellectual center of this area offered in the past a
research starting point for the Karst hinterland and wider surroundings, because here
it was the second largest natural history museum at the time of the Austro-Hungarian
Empire. This is also reflected in Lepidoptera literature as the area of Kras is cited in
many publications.
According to the current political division the majority of these data belongs to
the Italian side of the Kras. Data from the Slovenian part of the Kras are less
Acta entomologica slovenica, 24 (2), 2016
70
comprehensive and there are few recent data. With an aim to supplement the knowledge
of fauna of Macrolepidoptera of the Komenski and Sežanski kras, we herein present
a more complete review, which is the outcome of our 30-year-old survey of
Lepidoptera in this area.
Kras with a capital letter or parent Kras, which is named also Komenski and
Sežanski kras, is a limestone plateau the size of around 550 km2. It lies in the
southwest of Slovenia and in neighboring Italy, in Friuli - Venezia Giulia, which
possesses about a quarter of the total area. During the turbulent political history of
Kras in the last century, in the beginning it belonged to Austro-Hungarian Monarchy,
then to Italy, after the Second World War was shared by Italy and Yugoslavia and is
now divided between Italy and Slovenia. The border runs along the ridge Žekenc so
that the north-western part and a narrow strip of hinterland of Gulf of Trieste is on
the Italian side, while the rest, just over three quarters of the surface, is the Slovenian
part.
Natural boundaries are Gulf of Trieste to the southwest, Friulian plain in the
northwest and the Vipava Valley in the north and northeast, where border continues
through the valleys of Branica and Raša. On the southeast border is more blurred as
Kras passes in neighboring regions with similar characteristics, such as hills Vremščica,
Brkini, Matarsko podolje and Podgorski Kras. The only distinctive dividing line is
the Glinščica valley, which connects the southern boundary with the Gulf of Trieste.
Inside the plateau, except in the extreme north and south are gently rounded hills,
surrounded by valleys, sinkholes and larger planes. On the southwestern and
northeastern edge the plateau descends with steep slopes down to the lowlands. The
average altitude is between 300 and 400 m, with regard that the peaks V. Gradišče
and Ograda in the far southwest exceed 700 m, at the northern edge Trstelj and Stol
also rise over 600 m, in the northwest the plateau gradually reduces even at 50 m.
Kras’s geological basis is Cretaceous and Tertiary lime stone, just to the northwest
is occurring dolomite. Because of this permeable bedrock Karst does not have surface
waters. Intermittent lake Doberdob - Lago di Doberdo and Prelosno Lake - Lago di
Pietrarossa near Tržič - Monfalcone in Italy are two exceptions. Soil cover on the
plateau is shallow, particularly in the northwest and in places where the wind and
rain swept it away. Thicker soil is located only in numerous sinkholes and in valleys.
In the middle part of the Karst occurs famous red soil, jerovica or terra rossa.
The climate of the Kras is characterized by a mixture of sub-Mediterranean and
continental influences. Sub-Mediterranean influence is reflected in warm and relatively
dry summer and relatively high rainfall in spring and autumn. Continental influence,
however, is indicated in sporadic incursions of cold air in the form of the famous
burja (bora) wind and the distribution of average temperatures (Gogala, 2003).
The average annual temperature across the plateau varies between 10-12 °C,
except in the extreme north-west and in Brestoviški dol, where is 2 °C higher. In the
far southwest, which is on average higher than 500 m is 2 °C lower. The average
annual rainfall in the west, in the areas nearest the sea is around 1300 mm and is
gradually increasing towards the northeast, where it reaches 1800 mm
(www.arso.gov.si).
Radovan Øtanta, Matjaæ Zadrgal: Contribution to the knowledge of Lepidoptera fauna of the Karst
71
The dynamic karst terrain conditions large macroclimate differences. Valleys and
sinkholes are filled at night with cold air, which brings a lot of dew, therefore even
marsh plant species can be found there. The exact opposite is the dry slopes and
peaks of the hills, where dew may be completely absent. On a sunny day grasslands
and sparse woods get very hot during the day and cool at night strongly, due to the
radiation during clear nights. Minor fluctuations in temperature between day and
night are in denser stands.
Because of intensive logging and grazing of sheep, goats and cattle the plateau of
Kras was in the past almost naked. In the middle of the 19th century, the Austro-
Hungarian authorities began the systematic forestation of the Karst with the black
pine. It has proved to be very successful tree species to grow on the deserted plateau,
and therefore, mainly due to the abandonment of livestock, the Kras in recent years
turned into a landscape with a large proportion of forest or land in abandonment. You
can still admire the entire plateau interlacing and sharing its distinctive habitats, such
as dry grasslands, scrub and forest.
Typical plant associations of the Karst are divided into forest, scrub and the asso-
ciations of dry grasslands. 
Typical forest associations in the area are:
• The association of hop hornbeam and autumn moor grass (Seslerio autumnalis -
Ostryetum carpinifoliae) is a transitional form from grasslands to potential natural
association of downy oak and hop hornbeam.
• The association of sessile oak and autumn moor grass (Seslerio - Quercetum petreae)
appears on deeper grounds.
• The association of downy oak and hop hornbeam (Ostryo  carpinifoliae - Quercetum
pubescentis) is a climax forest association, we find it in various positions and it is
relatively frequent.
• Black pine grooves - almost two hundred years after forestation of the Karst with
black pine on poorer and rocky grounds a large amount of aging grooves are in a
poor state of health. Based on the development of succession, the oldest pines
already decay in the bottom layer of the stands and indigenous xerophilic tree
species are emerging.
Typical scrubland associations of the area are:
• The association of hazel and spring bell (Galantho nivalis - Coryletum avellanae)
occurs in the deeper, richer soils.
• Association of common dogwood and manna ash (Fraxino orni - Cornetum
hungaricae) can be found on various habitats.
• The association mahaleb cherry and rocky buckthorn (Frangulo rupestris - Prunetum
mahaleb) is a typical association of rocky habitats.
• The association wild privet elmleaf blackberry (Rubo ulmifolii - Ligustretum) is a
typical association of nutrient - rich habitats.
Acta entomologica slovenica, 24 (2), 2016
72
• Association with blackthorn Prunus spinosa can be found in eutrophic habitats.
• The association smoke tree and rock buckthorn (Frangulo rupestris - Cotinetum
coggygriae) is found between grasslands and forest edges.
Typical associations of dry meadows are:
• The most widespread and extremely rich in flora is the association of low sedge and
rock centaury (Carici humilis - Centaureetum rupestris), which grows from lowlands
to mountain belt.
• The association Genisto sericeae - Seslerietum juncifoliae is typical for rocky slopes
and ridges exposed to the wind.
• The association Danthonio-Scorzoneretum villosae is linked to the deeper soil and
the flora is very diverse (www.razvojkrasa.si/si/narava).
A review of literature on Lepidoptera from Karst
Compared to other “Carniola” countries, where Scopoli (1763) with his
Entomologia Carniolica laid the foundation and reached to the former top of
entomological science, butterflies of Karst are mentioned for the first time over 100
years later. The first records of 4 species we traced in Curò (1874, 1878), locations
are not exact, only Karst.
In the next period which lasts until the collapse of the Austro-Hungarian Empire
at the end of World War I, or after, we trace much more publications of various
researchers. Galvagni (1909) in his work covers the period between 1900 and 1909,
he states data for the landscapes around the northern Adriatic and Dalmatia. For
Trieste Karst lists 43 species, of which 3 from current Slovenian side. Ivan Hafner
(1909-1912) mentions two species in general for Kras and 2 specifically for Sežana
and Divača. Among the 587 species that Hafner (1910) states in his work
Macrolepidoptera from Gorica - Gorizia and the surrounding area, there are also 29
species of Kras. The data cover the period between 1892 and 1909 and are from two
locations, Sežana and Kobdilj. Five articles follow with one species: Naufock (1913),
Stauder (1913, 1914, 1921), Rebel (1919). 
Loebel (1920, 1921) recorded 145 species, mostly from Sežana and its
surroundings, others from Divača. In the Stauder's articles (1919/20, 1920/21, 1922,
1923, 1925, 1926, 1927) 76 species were listed, most of the current Slovenian side of
the Kras, almost half of the species records are citations of other authors.
The most complete review of this period can be found in Carrara (1926), in which
the "Territory of Trieste" (Territorio di Trieste) lists 584 species of Macrolepidoptera.
For certain species only exceptionally indicates the exact locality and date. He worked
between Križ - S. Croce and Žavlje - Zaule or Aquilinia in the Gulf of Trieste and
between Prosek - Prosecco, Opčine - Opicina or Villa Opicina and Bazovica -
Basovizza on the Karst plateau between 1907 and 1924. It can be concluded that
almost all species quoted for Trieste Karst belong to locations that are now on the
Italian side of the border.
Radovan Øtanta, Matjaæ Zadrgal: Contribution to the knowledge of Lepidoptera fauna of the Karst
73
During both world wars, when the entire Kras was under the Italian political
power, the entomological activities decreased. We find only two publications: Loebel
(1941) and Verity (1928) with one species.
After connecting Primorska to parent country in September 1947 and after the
normalization of the post-war situation, the exploration of butterflies had increased
again. Domestic entomologists have concentrated their activities mainly in the Triglav
National Park and reconnected areas, among which was also Primorska region with
Kras. But the parent Kras was quite neglected in comparison with the coastal areas
and the Karst edge, the Vipava Valley with southern slopes of the Trnovski gozd and
western parts of the Julian Alps.
The first publications of this period, signed by Italian entomologist Verity (1947,
1950, 1953) with four species for Slovenian Karst and two for the Italian side,
followed by Proust and Wehrli (1934-1954) with two species. Carnelutti and Michieli
(1955) indicate three new species of the Slovenian Karst among the novelties for
Slovenia. Two articles followed with one species of the Trieste Karst (Melis, 1957;
Storace, 1963), Pinker (1965) with eleven species on the Slovenian side and Bartol et
al. (1965) with one. Carnelutti (1971) critically cites only findings by then, those of
three species of Kras. Pleterski (1972) gives 160 species of owlet moths (Noctuidae)
without naming for Sežana, provided only to the level of genera and 158 species of
other families, also without naming. In the article by Michieli (1978) 67 more rare or
more interesting species of Lepidoptera from "Vipava Valley and neighboring karst
areas" are presented, where Karst is not precisely defined. Considering the known
prevalence of certain species, we can conclude that the author had in mind the entire
south-western Dinaric karst area to the Karst edge and beyond. Many of these species
can be considered as representatives of the parent Kras. Especially we can mention
Chelis maculosa and Omia cymbalariae for Divača, since this is the only information
from the Kras for these two species.
With the political independence of Slovenia in 1991 comes a new fruitful period
of entomological research. The following year Slovenia gets the first comprehensive
review of Macrolepidoptera (Carnelutti, 1992a, 1992b). Kras is included in the
zoogeographical region of Primorska. Some articles with publications of new species
records for Slovenia include new records for Kras. Štanta (1995, 1996, 2008) and
Lasan (1997, 2000) recorded each of three and Predovnik (2001, 2002) two species.
During this period, Čelik and Rebeušek (1996) collect and publish the first detailed
data on endangered butterfly species of Slovenia, of which 13 are also found in the
Kras. Here are some research works or reports, which deal mainly with butterflies of
Kras: Verovnik, 2000, Čelik, 2003, Čelik et al., 2005, Polak, 2009, Bedjanič et al.
2009.
In the publication by Flamigni et al. (2007) recent data for 26 species of Ennominae
are presented for the Italian part of the Karst, mentioned are also 3 interesting findings
in the Komen Karst: Gnopharmia stevenaria, Ennomos autumnaria and Nychiodes
dalmatina. Cited after Lasan (2000)  is the discovery of Isturgia murinaria for the
"Carso Triestino SLO" and marked on map at Šmarje pri Sežani. It is a
misinterpretation as Šmarje near Koper is the site where the species was found.
Acta entomologica slovenica, 24 (2), 2016
74
The most extensive recent work in this field (Deutsch, 2008) deals with the
province of Gorizia and Trieste in Friuli Venezia Giulia - Italy between 1973 and
2005. In it 905 species are listed, because it also includes Microlepidoptera. Among
the 12 present locations, there are 7 situated in the Gorizia and Trieste Karst. If we
consider only Macrolepidoptera with these seven karst sites, we get 359 species.
In The Atlas of Slovenian butterflies (Verovnik & al., 2012) there are confirmed
117 species of butterflies for the parent Kras. With the recent work by Koren & al.
(2013) the prevalence of sister species Pyrgus malvae / malvoides are presented at
their intersection with several new locations including the Kras. Some new butterflly
sites for the Slovenian part of the Karst is also found in Čelik (2014).
In a recent work (Jež et al. 2015) R. Štanta recorded 149 species of nocturnal
Macrolepidoptera found on three illuminated churches in the central Kras.
From the posts from the beginning of activity until the 60s of the last century, we
see that entomologists operated primarily in the area of  Trieste and at the railway
lines, which run through the Kras, Vienna - Trieste - Venice and Gorizia - Trieste -
Rijeka. Other parts with few exceptions have been unexplored due to the difficult
access. Greater coverage area and greater efficiency of the inventory has recently
been enabled with the development of other means of transport and new ways of
night hunt, especially of the portable UV lamps.
Much of the data from the listed literature is not precise enough to be accurately
placed in time and space. Nevertheless, with a high degree of accuracy we can
estimate that 670 species of Macrolepidoptera are recorded for the Trieste and Gorizia
Karst, which now belongs to Italy. For the Slovenian part of the Karst approximately
390 species are published.
Listed among Macrolepidoptera after that scheme also were: Hepialidae, Psychidae,
Limacodidae, Heterogynidae, Zygaenidae, Sesiidae, Cossiidae and Thyrididae. The
old systematics in the listed literature has led us to decide that we maintain the same
classification in this paper in order to facilitate comparisons.
Material and methods
Inventories of Lepidoptera were conducted with day and night observations. In
the case of daytime flying species we have carried out the locations overview and
classical catching with butterfly net, as well as searching for different developmental
stages on their food plants.
In the case of observations at night between 1985 and 1990, we mainly inspected
lit areas under public lighting, which was renovated a little earlier and partly newly
built. In it mainly 125W mercury vapor lamps were applied, which were in terms of
attracting moths highly effective. The advantage of this method is data collecting in
the same day from several sites. We used this method also later occasionally, but the
aging of lamps and moreover with their replacement with more ecologic sodium-
vapor lamps became completely ineffective. In the next period, most of the data were
obtained by means of attracting moths on a white upright placed canvas, illuminated
with 300W mercury-vapour lamp with a high proportion of UV light, powered by
Radovan Øtanta, Matjaæ Zadrgal: Contribution to the knowledge of Lepidoptera fauna of the Karst
75
220V electric generator. From 2000 onwards, we also used special light tents
illuminated with super actinic UV tube lamps Blacklight and Blacklight blue 15W
powered by 12V batteries covered by a translucent white synthetic fabric.
In the early spring and autumn months we used to attract moths with wine bait
smell which has proved to be a very effective method.  A mixture of wine, vinegar
and sugar was used with which we sprinkled tree trunks just before dark. Owlet
moths - Noctuidae and Erebidae prefer the bait.
We identified the species mainly on the spot already. Exceptionally, we took a
small number of specimens away from the nature for subsequent determination. In
some difficult determinable species we have used also the inspection of the genital
armatures. Collected specimens of Lepidoptera are deposited in the collections of the
authors.
Localities and dates of inventories of species
The border of the studied Komenski and Sežanski kras area is presented in Figure
1.
Addressed area was divided into 5 approximately equal zones. They are labeled
with numbers from 1 to 5, and they follow as shown in Fig. 1, from northwest to
southeast. This division allows us a more accurate analysis and show the prevalence
and easier detection of species which have more local distribution. 
In the Table 1 we present an overview of 70 locations, which provide data and
dates of inventories.
Acta entomologica slovenica, 24 (2), 2016
76
Fig. 1: Map of Komenski and
Sežanski kras region divided into 5
zones
Locations are named after the nearest town, hamlet, peak or after the provincial
name. The average altitudes of locations are also listed and accurate or raw location
coordinates in case if we were not able to identify them precisely, due to the size of
the observed areas or due to more surveys in the vicinity of the same location. The
names of locations are cited after Atlas Slovenije (1996).
Dates of observations or excursions are arranged chronologically for each location
and gathered in groups of authors observations. Day and night observations are not
specifically separated, so you can find two or more locations at the same date. More
locations can also be found when we visited more long distant locations during one
observation.
Authors of observations are marked only with initials of their names explained
below in order to save space. Considered are only those authors who have joined us
on observations with their equipment.
(RŠ) = Radovan Štanta, Miren, (MZ) = Matjaž Zadrgal, Vrtojba, (BZ) = Bojan
Zadravec, Nova Gorica, (CM) = dr. Carlo Morandini, Campoformido (Italy), (SG) =
Stanislav Gomboc, Kranj, (ML) = Mojmir Lasan, Ljubljana, (TL) = Tone Lesar(†),
Maribor (JD) = Jozef Debets, Solčava
Table 1: Overview of localities of observed species and associated zones, as iden-
tified on the map and dates of the observations. 
Radovan Øtanta, Matjaæ Zadrgal: Contribution to the knowledge of Lepidoptera fauna of the Karst
77
Zone Name, altitude and coordinates of
localities after WGS84
Dates and authors of observations
1 Brestovica, 66 m, 45.81258, 13.62173 13.2.1997, 28.6.1995, 31.5.1998, 22.6.2013 all (RŠ); 6.9.1991,
9.5.1992, 16.5.1994, 9.5.2013 all (RŠ&MZ); 24.9.1994,
26.9.1995, 16.5.2000 (MZ)
1 Gornja Brestovica, 70 m, 45.81236,
13.64460
16.4.1995 (RŠ); 20.7.1991, 19.3.1995, 10.9.1995, 3.4.2009,
13.11.2009 all (RŠ&MZ); 10.7.1997 (RŠ&ML); 26.3.1999
(RŠ&SG)
1 Hudi Log, 200 m, 45.83984, 13.61076 29.5.1997, 3.9.2012, 24. 8.2013 all (RŠ)
1 Klariči, 30 m, 45.81217, 13.60492 25.6.1988, 30.6.1988, 9.5.1992, 14.7.1992, 28.8.1993,
11.10.1993, 29.12.1994, 4.6.1995, 28.6.1995, 1.7.1995,
26.9.1995, 2.10.1995, 9.10.1995, 25.3.1996, 24.6.1996,
16.1.1998, 15.3.1998, 31.5.1998, 7.7.2001, 29.9.2011, 13.9.2015
all (RŠ); 19.7.1988, 9.11.1991, 7.9.1993, 18.3.1994, 25.3.1994,
24.4.1994, 13.11.1994, 22.1.1995, 26.1.1995, 18.3.1995,
19.3.1995, 26.3.1995, 7.9.1995, 29.10.1995, 13.11.1996,
15.3.2002, 13.11.2009, 21.4.2010, 27.9.2013, 7.10.2014 all
(RŠ&MZ); 4.11.1993, 5.7.1995, 15.9.1995, 6.5.2000,
22.4.2006, 11.6.2010, 6.6.2012, 9.6.2013, 15.3.2014 all (MZ)
1 Klariči – Gnojne, 120 m, 45.80362,
13.61014
18.6.2014 (RŠ); 24.9.2014 (RŠ&MZ)
1 Korita na Krasu, 226 m, 45.83243,
13.62280
27.6.2004 (RŠ)
Acta entomologica slovenica, 24 (2), 2016
78
1 Kostanjevica na Krasu, 277 m,
45.84405, 13.64199
22.4.1981, 25.7.1981, 31.3.1987, 9.8.1987, 23.4.1988,
16.4.1989, 16.8.1989, 19.3.1990, 1.5.1991, 19.5.1991,
10.10.1991, 18.6.1993, 24.9.1993, 4.10.1993, 11.7.1994,
19.8.1994, 27.11.1994, 20.1.1995, 22.4.1995, 19.6.1995,
2.10.1995, 25.10.1995, 20.4.1996, 4.7.1996, 4.11.1996,
16.3.1997, 29.5.1997, 22.6.1997, 20.4.1998, 10.7.1998,
16.7.1998, 26.3.2003, 24.5.2003, 17.4.2006, 11.4.2007 all
(RŠ);10.10.1986, 2.5.1987, 23.5.1987, 6.6.1987, 11.7.1987,
15.7.1987, 25.7.1987, 23.8.1987, 15.8.1987, 19.9.1987,
27.5.1988, 11.6.1988, 19.7.1988, 24.9.1988, 7.4.1989,
14.4.1989, 23.4.1989, 6.5.1989, 10.6.1989, 1.7.1989, 26.7.1989,
25.8.1989, 1.10.1990, 13.3.1991, 7.4.1993, 22.5.1993,
22.9.1993, 25.3.1994, 31.3.1994, 16.5.1994, 17.2.1995,
1.3.1995, 2.4.1995, 12.7.1995, 26.2.1997, 9.7.2002, 30.10.2002
all (RŠ&MZ); 21.7.2001 (RŠ&BZ)
1 Lokvica, 215 m, 45.86114, 13.60280 23.11.1994, 28.6.1996, 4.11.1996, 19.7.2007, 26.8.2008,
8.10.2009, 21.11.2009, 22.6.2010, 5.9.2012 all (RŠ); 25.5.2002
(RŠ&BZ&SG)
1 Lokvica – Devetaki, 168 m, 45.86710,
13.58514
23.6.2006, 11.8.2006 all (RŠ)
1 Lokvica – Medvejšče, 300 m,
45.86370, 13.62460
27.6.2004, 30.3.2006, 2.9.2006, 30.3.2007, 6.4.2007, 25.2.2008,
9.4.2011, 1.8.2012, 14.9.2014, 17.9.2014 all (RŠ); 9.4.2007
(RŠ&MZ)
1 Lokvica – Segeti, 203 m, 45.85439,
13.61112
23.5.1988, 4.6.1989, 28.9.2003, 23.4.2006, 2.9.2011 all (RŠ);
11.4.2007 (RŠ&MZ&BZ&SG)
1 Miren – Gmajna, 215 m, 45.87602,
13.59923
19.11.1987, 2.9.1991, 27.10.1992, 19.3.1993, 29.10.1993,
11.11.1993, 5.11.1994, 24.3.1995, 15.4.2012 all (RŠ);
13.11.1994, 1.3.1995, 7.9.2008 (RŠ&MZ)
1 Nova vas, 200 m, 45.84173, 13.58372 14.10.2012, 14.9.2014 all (RŠ)
1 Novelo, 333 m, 45.84675, 13.66125   20.10.1989, 12.7.1996, 14.5.2011 all (RŠ); 14.7.1994,
15.8.1994, 26.10.1996, 9.7.2002 all (RŠ&MZ)
1 Opatje selo, 170 m, 45.85230,
13.58193
19.8.1978, 14.5.1989, 8.7.1989, 16.8.1989, 10.10.1991,
15.3.1992, 8.9.1992, 13.6.1993, 24.9.1993, 11.10.1993,
1.11.1993, 17.6.1994, 19.4.1995, 28.6.1996, 23.4.2006,
4.6.2006, 23.6.2006, 7.1.2007, 21.9.2011 all (RŠ); 2.5.1987,
6.6.1987, 11.7.1987, 2.8.1987, 15.8.1987, 23.8.1987, 19.9.1987,
6.5.1988, 27.5.1988, 11.6.1988, 19.7.1988, 13.8.1988,
24.9.1988, 7.4.1989, 14.4.1989, 6.5.1989, 12.5.1989, 10.6.1989,
1.7.1989, 26.7.1989, 25.8.1989, 27.10.1989, 5.5.1990,
21.9.1990, 1.6.1991, 7.9.1993, 22.1.1995, 26.1.1995 all
(RŠ&MZ); 9.5.1987, 12.7.1987, 17.7.1987, 11.9.1987,
7.5.1995, 18.6.2004, 20.3.2007 all (MZ)
1 Sela na Krasu, 230 m, 45.82128,
13.61729
8.7.1989, 7.1.2007, 13.11.2010, 9.11.2011, 2.11.2013 all (RŠ);
10.11.2014 (MZ)
Radovan Øtanta, Matjaæ Zadrgal: Contribution to the knowledge of Lepidoptera fauna of the Karst
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1 Temnica, 402 m, 45.84578, 13.67785 4.10.1993, 25.10.1995, 11.8.2008, 21.7.2009, 1.8.2009,
9.4.2011, 14.5.2011 all (RŠ)
1 Trstelj, 622 m, 45.85787, 13.70660 19.5.1991, 18.6.1993, 15.7.2000, 1.8.2009, 6.8.2009, 16.7.2010
all (RŠ); 14.6.1991, 2.9.2009 all (RŠ&MZ)
1 Trstelj – Vrtovka, 406 m, 45.86455,
13.66558
23.7.2009 (RŠ&MZ&TL); 25.7.2010 (MZ)
1 Vale, 120 m, 45.81299, 13.67324 23.6.1999 (RŠ)
1 Vojščica, 309 m, 45.82857, 13.67339 19.3.1988, 4.10.1991, 4.10.1993, 11.10.1993, 11.7.1994,
30.10.1994, 9.10.1995, 15.11.1995, 25.3.1996, 4.7.1996,
11.10.1996, 4.11.1996, 10.7.1998, 18.8.2002, 2.5.2004,
27.12.2012 all (RŠ); 10.10.1986, 22.9.1993, 31.3.1994,
24.4.1994, 16.5.1994, 14.7.1994, 17.2.1995, 2.4.1995,
12.7.1995, 7.9.1995, 26.10.1996, 26.2.1997, 10.6.1999, 4.6.2003
all (RŠ&MZ); 1.7.1987, 17.7.1993, 16.10.1993, 7.5.1994,
6.6.2011, 29.7.2012 all (MZ)
2 Brje pri Komnu, 178 m, 45.78370,
13.71933
18.10.1995, 4.11.1996 all (RŠ); 22.5.1993, 2.4.1995,
29.10.1995, 2.5.1999, 30.4.2003, 25.9.2009, 25.4.2014 all
(RŠ&MZ); 15.6.2001, 21.7.2013, 26.7.2014 (MZ)
2 Gorjansko, 200 m, 45.80232,
13.71403
15.11.1995, 4.11.1996, 13.2.1997, 21.5.2000, 2.5.2004,
3.7.2005, 22.5.2011, 9.6.2013, 22.6.2013 all (RŠ); 24.4.1994,
16.5.1994, 29.10.1995, 20.10.1996, 13.11.1996, 26.2.1997,
22.5.2009 all (RŠ&MZ); 11.9.1996, 5.11.1996, 29.4.2004,
15.5.2007, 18.6.2007, 20.5.2009, 5.7.2009 all (MZ)
2 Gorjansko – Krmanjek, 150 m,
45.79801, 13.70130
29.9.2011, 2.10.2011 all (RŠ)
2 Gorjansko – Podklanec, 200 m,
45.80630, 13.70645
3.7.2009 (RŠ&MZ)
2 Ivanji Grad, 297 m, 45.81891,
13.70680
1.4.2001, 13.11.2010, 9.11.2012 all (RŠ)
2 Klanec pri Komnu, 180 m, 45.80956,
13.69820
3.7.2005 (RŠ); 21.4.2010 (RŠ&MZ)
2 Kobjeglava – Zavodi, 270 m,
45.81415, 13.80066
29.10.1995, 19.5.1997, 8.5.2001 all (RŠ&MZ)
2 Komen, 290 m, 45.81728, 13.74532 12.4.2007, 29.9.2010; 17.7.2015 all (RŠ); 10.10.1986, 2.5.1987,
23.5.1987, 22.9.1993, 25.3.1994, 19.5.2011, 24.9.2013 all
(RŠ&MZ); 23.4.1996 (MZ&TL); 28.5.2012, 13.6.2013,
5.8.2013 all (MZ)
2 Komen – Devinščina, 240 m,
45.80893, 13.72971
13.11.2010, 29.5.2011 all (RŠ)
2 Komen – Jažmerca, 210 m, 45.80122,
13.76386
5.6.2010, 18.9.2012 all (RŠ&MZ&BZ)
2 Lipa na Krasu, 366 m, 45.84386,
13.70732
30.6.1990, 18.8.1994, 27.11.1994, 20.4.1996, 12.7.1996,
29.5.1997, 16.5.1998, 10.7.1998, 13.6.1999, 26.4.2000,
18.6.2001, 9.6.2002, 29.7.2002, 25.2.2008, 1.8.2009, 19.6.2010,
9.4.2011, 14.5.2011 all (RŠ); 15.8.1994, 5.9.1994, 18.7.1995,
10.9.1995, 12.5.2006, 19.5.2010 all (RŠ&MZ); 21.7.2001
(RŠ&BZ); 14.5.2000, 9.6.2000, 14.5.2010, 16.6.2010 all (MZ)
Acta entomologica slovenica, 24 (2), 2016
80
2 Mali Dol, 270 m, 45.83355, 13.76515 2.5.2004, 20.5.2007, 4.10.2007 all (RŠ)
2 Preserje pri Komnu, 264 m, 45.81811,
13.72852
29.10.1995 (RŠ)
2 Škrbina, 360 m, 45.84590, 13.72486 18.8.1994, 27.11.1994, 14.5.2011, 6.5.2014 all (RŠ); 24.4.1994,
15.8.1994, 5.9.1994, 18.7.1995 all (RŠ&MZ)
2 Tublje pri Komnu, 200 m, 45.77696,
13.73963
29.10.1995 (RŠ&MZ)
2 Volčji Grad, 255 m, 45.80259,
13.74697
8.6.2004 (RŠ)
2 Zagrajec, 280 m, 45.81530, 13.69937 2.10.1995, 13.2.1997, 1.6.2003, 13.10.2008, 30.9.2010,
30.10.2010, 2.10.2011 all (RŠ); 15.11.1992, 26.4.1993,
1.3.1995, 26.7.2008, 22.5.2009, 10.9.2010, 25.10.2013 all
(RŠ&MZ); 6.11.2004 (MZ)
3 Brje pri Koprivi, 280 m, 45.77852,
13.84864
16.10.1998, 25.4.2014 all (RŠ&MZ); 25.10.2009 (RŠ&BZ)
3 Dobravlje, 310 m, 45.76291, 13.88159 28.9.1996, 20.7.2013 all (RŠ)
3 Dutovlje, 315 m, 45.75632, 13.83525 21.6.1991, 9.8.1994, 18.10.1995, 15.11.1995, 11.10.1996,
4.11.1996, 14.10.1998, 17.8.2013 all (RŠ); 25.3.1994,
31.3.1994, 24.4.1994, 16.5.1994, 2.4.1995, 7.9.1995, 29.10.1995
all (RŠ&MZ); 1.6.1994, 15.9.1995, 23.3.1996 all (MZ)
3 Kobjeglava, 320 m, 45.81688,
13.80965
2.10.1995, 18.10.1995, 15.11.1995, 4.11.1996 all (RŠ);
19.5.1987, 15.11.1992, 31.3.1994, 7.9.1995, 26.4.1997,
27.9.1997 all (RŠ&MZ); 18.5.1996 (MZ)
3 Kobjeglava – Jelenca, 285 m,
45.80901, 13.80898
5.8.1988, 15.6.1990, 22.5.1997, 22.5.1999, 23.5.1999,
21.5.2000, 26.5.2000, 20.5.2007, 8.10.2008, 22.5.2011, 4.6.2012
all (RŠ); 25.4.1993, 19.5.1997, 9.5.2012, 25.10.2013, 6.5.2014
all (RŠ&MZ); 17.5.2004 (RŠ&MZ&BZ); 13.5.1997
(RŠ&MZ&SG); 16.5.1996 (RŠ&ML)
3 Kopriva, 280 m, 45.78177, 13.83379 22.5.2011, 29.5.2011, 30.5.2011, 10.6.2011, 26.6.2011,
20.8.2011, 3.9.2011, 15.6.2012, 27.6.2012, 18.7.2012,
13.8.2012, 10.7.2013, 4.8.2013, 17.8.2013, 7.9.2013 all (RŠ);
2.4.1995, 15.5.2013 all (RŠ&MZ); 5.7.2013 (MZ); 29.6.2013
(RŠ&MZ&CM); 6.7.2014 (RŠ&CM)
3 Krajna vas, 250 m, 45.76314,13.80289 
14.9.2010, 21.5.2014 all (RŠ); 21.5.2014, 18.7.2014 all
(RŠ&MZ)
3 Kreplje, 280 m, 45.74515, 13.83082 8.6.2004 (RŠ)
3 Pliskovica, 280 m, 45.77545,
13.78746  
23.6.1999, 10.8.2010, 29.5.2011, 10.8.2011, 15.8.2011,
17.8.2013 all (RŠ); 22.9.1993 (RŠ&MZ)
3 Ponikve, 325 m, 45.78986, 13.85696 12.6.2013, 17.6.2013 all (RŠ); 30.3.2014 (RŠ&MZ); 29.6.2013
(RŠ&MZ&CM)
3 Skopo, 290 m, 45.77385, 13.82180 29.5.2011, 30.5.2011, 10.6.2011, 26.6.2011, 28.7.2011,
20.8.2011, 3.9.2011, 12.10.2011, 4.8.2013, 17.8.2013, 7.9.2013
all (RŠ); 28.3.2012 (RŠ&MZ)
3 Štanjel, 300 m, 45.82069, 13.83588 2.5.1997 (RŠ); 1.6.1996 (MZ)
Radovan Øtanta, Matjaæ Zadrgal: Contribution to the knowledge of Lepidoptera fauna of the Karst
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3 Tomaj, 360 m, 45.75698, 13.85754  12.10.2014 (RŠ)
3 Veliki Dol, 220 m, 45.77083,
13.75269
15.6.2012, 27.6.2012, 28.7.2012, 13.8.2012, 15.5.2013,
13.8.2014 all (RŠ); 29.10.1995 (RŠ&MZ)
3 Vrhovlje, 350 m, 45.72525, 13.83003 8.6.2004, 17.8.2013, 6.6.2014 all (RŠ); 8.8.2014 (RŠ&MZ)
4 Grahovo Brdo, 340 m, 45.74383,
13.90570  
21.3.2014 (RŠ)
4 Kazlje, 340 m, 45.75553, 13.90355 30.7.2005; 13.5.2015 all (RŠ); 17.9.2011, 31.10.2011,
28.3.2012, 14.11.2014; 19.5.2015 all (RŠ&MZ)
4 Podbreže, 380 m, 45.72658, 13.92076 6.6.2014 (RŠ); 1.5.2013, 28.5.2013, 12.7.2013 all (RŠ&MZ);
25.4.2013 (MZ)
4 Povir, 420 m, 45.69437, 13.93855 22.5.2005 (RŠ); 28.4.2012, 11.10.2013; 29.6.2015 all
(RŠ&MZ); 22.8.2005 (MZ)
4 Sežana, 350 m, 45.70722, 13.84790 15.9.1999 (RŠ&MZ); 2.10.1997 (MZ)
4 Sežana – Dol Leskovec, 345 m,
45.69850, 13.86794
14.6.1986, 23.5.1999, 10.7.2005, 17.8.2013 all (RŠ)
4 Šepulje, 350 m, 45.75282, 13.87099 16.10.1998 (RŠ&MZ)
4 Šmarje pri Sežani, 296 m, 45.72203,
13.86662 
26.6.2011, 28.7.2011, 15.5.2013, 4.8.2013 all (RŠ)
4 Štorje, 355 m, 45.75193, 13.92337 28.4.1996, 28.9.1996, 10.7.2005 all (RŠ); 6.5.2015 (RŠ&MZ)
5 Divača, 435 m, 45.66786, 13.94995 8.7.2002, 22.5.2005 all (RŠ); 25.10.1992, 20.10.1996,
15.9.1999, 15.9.2000 all (RŠ&MZ); 25.9.1999 (MZ)
5 Jirmanec, 665 m, 45.64414, 13.90096 17.6.2011, 8.7.2011 all (RŠ); 4.6.2011, 19.8.2011, 28.7.2012,
14.6.2013, 5.9.2013, 19.10.2013 all (RŠ&MZ); 9.6.2013 (MZ);
7.9.2014 (RŠ&MZ&BZ); 8.5.2015 (RŠ&MZ&BZ&SG);
3.9.2015 (RŠ&MZ&CM)
5 Kozina, 500 m, 45.61421, 13.94821  30.6.1992, 2.6.1994 all (RŠ)
5 Krvavi Potok, 520 m, 45.62601,
13.91472 
30.6.1992 (RŠ); 18.4.2013 (RŠ&BZ&SG)
5 Lokev, 455 m, 45.65583, 13.90865 23.5.1999, 8.7.2002, 6.6.2014, 25.8.2014 all (RŠ); 30.9.2011,
16.3.2012 all (RŠ&MZ); 4.9.2010 (RŠ&MZ&BZ); 25.6.2010
(RŠ&MZ&BZ&CM); 19.10.2012 (RŠ&MZ&BZ); 26.5.2012,
9.6.2013 all (MZ)
5 Rodik, 500 m, 45.62878, 13.96455 15.9.1999 (RŠ&MZ)
5 Vrhpolje, 550 m, 45.62760, 13.91480 8.3.2014 (RŠ&MZ)
5 Vrhpolje – Sv. Tomaž, 600 m,
45.63210, 13.92089
17.6.2011, 8.7.2011, 16.6.2012 all (RŠ)
Explanation of terms to the table showing all species observed in the area -
Table 2
We present the data in a table with an overview of all of the species noticed in the
area. Data present species according to the content that represent family and name of
the species, the presence of the species in each zone (presence is marked with an x),
time of occurrence of an adult butterfly (flying period) with an indication of the
months when it was spotted, the frequency of species throughout the area and notes
where peculiarities are presented and also interesting facts or accurate data in the
case of rare species.
The classification and nomenclature of the species is listed according to Nieukerken
et al. in Zhang [ed.] (2011). All dates, which are stated in the text are provided in the
format day.month.year.
Lepidoptera flying period is provided with the consecutive number of the month
(1 = January, 2 = February, ...). It indicates when the adult imago was observed, with
a few exceptions when it is an observation of caterpillars, as explained in a footnote.
Frequency of species is estimated on the basis of the average number of observed
specimens of each species in the entire 30 year period inventory. If the species is
present everywhere, the rating applies to the entire Karst if not this is applied only to
segments where the presence of the species is confirmed or for micro-location in
them, if the species is distinctly local (abandoned ponds, distinctly dry and warm
slopes, the tops of hills ...). The rating applies to the time interval when specimens of
the species fly except with a few exceptions, where the frequency is estimated on the
basis of observations of caterpillars (Cucullia, Thaumetopoea, Psychidae ...)
Frequency of species in the table is divided into 5 categories: very rare, rare, less
common, common and very common. Very rare means that there has been observed
less than one specimen per year, a rare - observed one to 5 specimens per year, less
common - observed over 5 specimens per year to three specimens in one observation,
frequent - observed from 4 to 20 specimens in one observation, very common - seen
more than 20 specimens in one observation.
Specimens of rarer, more interesting and more difficult determinable species
which are mentioned by exact data, are deposited in the collections of the authors as
evidence.
Results
During the thirty years of observations (1985-2015) Macrolepidoptera fauna of
Komenski and Sežanski kras region, we recorded 755 species of Macrolepidoptera at
the total of 422 inventories. 
Including some data before 1985, we collected 15,400 records of the species finds.
A complete species checklist is shown in Table 2 and represents data for the 25
families of Macrolepidoptera, as day or night flying species. Families with the greatest
number of species identified in the area are Noctuidae (230 species), followed by
Geometridae (212 species) and Erebidae (83 species).
Acta entomologica slovenica, 24 (2), 2016
82
By classifying and analyzing the data by zones we tried to determine the potential
differences between them. Most of the records are from zone 1, 2 and 3, which repre-
sent 83% of the total. Among the most notable is zone no. 1 with 38% of the performed
examinations and the largest number of locations, which leads to a greater number of
found species (636). Zones 4 and 5 were less often visited, due to greater distances
from the place of residence, so the total number of data and collected species is
smaller (Fig. 2). 
In zone 1, we find the majority of the species (56) which were found in only one
of the zones. The zone 5 follows with 33 species, while in the central zones the
number of such species is lower (Fig. 3).
Overview of interesting species of the area
This chapter presents some interesting, more scarce or locally distributed species
with comments and specific data of findings.
Radovan Øtanta, Matjaæ Zadrgal: Contribution to the knowledge of Lepidoptera fauna of the Karst
83
Fig. 2: Number of species found
in each zone
Fig. 3: Number of species in
this zone only
Zygaenidae
Jordanita budensis Ad. & Au. Spey.
A rare species which with isolated populations occupies area from Spain through
southern Europe, Asia Minor and the Caucasus to Mongolia. In neighborhood it has
been found in the Trieste Karst Villa Opicina - Opčine (Deutsch, 2008). In Slovenia,
it is known from Nanos (Štanta, Zadrgal personal observation), Vipava Valley and
Kozina (Deutsch, 2008), southern slope of Trnovski gozd (Kovk, Kucelj) and Kačiči
near Divača (S. Gomboc personal comment) and Brkini (S. Gomboc and M. Lasan
personal comment). For the determination the analysis of genital structures, especially
in females is necessary due to the visual similarity with related species Jordanita
globulariae and Jordanita notata. The species is active during the day, at night it
might come to artificial light, as came the specimen of 5.6.2014. Three males were
found in Komenski Kras: Kobjeglava – Jelenca, 23.5.1999 1♂, 21.5.2000 1♂; Kob-
jeglava – Jelenca, 6.5.2014 (RŠ&MZ) gen. prep. & coll. (RŠ); several specimens
have been observed in Kazlje, 13.5.2015 3♂1♀ leg., gen. prep. & coll. (RŠ).
Zygaena punctum Ochs.
Mediterranean – Eastern European xerophilous species that inhabits karst meadows
where its food plants of the genus Eryngium grow. As a rare species, it is regularly
found in Goriški and Komenski kras. Occasionally more numerous populations occur
in the area around Brestovica and Komen.
Klariči, 4.6.1995, 1.7.1995, 24.6.1996; Opatje selo, 28.6.1996; Kostanjevica,
22.6.1997; Komen - Mali Dol, 20.5.2007; Kobjeglava – Jelenca, 20.5.2007; Kopriva,
29.5.2011; Vrhovlje, 6.6.2014 all leg. & coll. (RŠ); Klariči, 9.6.2013; Komen,
13.6.2013 both leg. & coll. (MZ).
Hesperiidae
Pyrgus malvae L. and Pyrgus malvoides Elw. & Edw.
In the Atlas of butterflies of Slovenija (Verovnik et al., 2012) a doubt is expressed
about the accuracy of the data collected on sister species P. malvae / malvoides in
Primorska, where the species mix, so we conducted an analysis of the genital structures
of all 24 specimens from the Karst, which were available to us. 16 specimens with a
total of 8 locations belong to P. malvoides. The remaining 8 specimens belong to P.
malvae, all from a single location - south-east from Krajna vas. Although all the spe-
cimens are only from the zones 1, 2 and 3, they adequately complement the image of
prevalence of P. malvoides at its eastern border, according to Koren et al., 2013.
P. malvoides:
Kostanjevica na Krasu, 22.4.1995 1♂1♀; Kostanjevica na Krasu, 20.4.1998 1♂2♀;
Komen, 22.5.2010 1♂; Lipa na Krasu, 19.5.2010 2♂; Lipa na Krasu, 14.5.2011 1♀;
Novelo, 14.5.2011 1♀;  Kobjeglava – Jelenca, 23.5.2010 2♂; Škrbina, 6.5.2014 1♂
all leg., gen. prep. & coll. (RŠ); Opatje selo, 7.5.1995 1♂; Lipa na Krasu, 14.5.2000
1♂; Brestovica, 16.5.2000 1♂ all leg., gen. prep. & coll. (MZ).
P. malvae:
Krajna vas, 21.5.2014 7♂1♀ leg., gen. prep. & coll. (RŠ).
Acta entomologica slovenica, 24 (2), 2016
84
Lycaenidae
Polyommatus escheri Hübn.
Escher's blue is in Slovenia only known from flysch slopes in the coastal part of
Primorska and in a small area in the west of Ilirska Bistrica (Verovnik et al., 2012).
By locating one male in Klariči near Brestovica is confirmed the data of 1992 that
species is present also in Komenski Kras (UL97) (Čelik, Rebeušek, 1996): Klariči,
6.6.2012 leg. & coll. (MZ).
Geometridae
Rhoptria asperaria Hübn.
It is distributed from the Canary Islands, via Morocco, Algeria and South Europe
to Asia Minor. It is not rare in the coastal maquis of the Mediterranean countries, but
is mostly local. It flies in at least two generations from March to November. In our
vicinity it has been found in Friuli and Istria (Flamigni, et al., 2007). The only
published data for Slovenia is one from the beginning of May 1854 from the vicinity
of Gradišče at Vipava (Mann, 1854), so Carnelutti included it in the red list (Carnelutti,
1992a) as presumed extinct (EX?). It was found in one specimen in Kras: Komen,
24.9.2013 (RŠ&MZ) coll. (MZ).
Ennomos quercaria Hübn.
Widespread in southern and central Europe to Iran. It flies between May and Oc-
tober in two generations. Inhabits oak forests because the larva feeds on various oaks
- Quercus spp. In the south of the Apennine and the Balkan peninsulas it is quite
Radovan Øtanta, Matjaæ Zadrgal: Contribution to the knowledge of Lepidoptera fauna of the Karst
85
Fig. 4: Rhoptria asperaria female, Komen, 24.9.2013, leg. and coll. M. Zadrgal
common, but in the north it is rarer and more local. It is very rare in Slovenia. Long
after the discovery in Vipava (Mann, 1854) it was no longer observed, therefore the
red list (Carnelutti, 1992a) classified it as extinct (EX) for the Dinaric karst region
and possibly extinct (EX?) for the Primorska region. It was later found in Podgorski
kras (Flamigni et al., 2007), 23.9.2011 leg. (CM&RŠ) coll. (CM), near Koper (M.
Sukič personal comment) and in Kozjansko (S. Gomboc personal comment). We got
it only once in Kras: Ponikve, 29.6.2013 (RŠ&MZ&CM) coll. (RŠ).
Lycia graecarius Staud.
The species can be found in southern and western Balkans, in Italy extends back
to the western border of Friuli, which is also the most eastern border of a similar
species Lycia florentina (Flamigni et al., 2007). It was noted for the Slovenian Pri-
morska (Carnelutti, 1992a) due to some wrongly determinated specimens. From this
sister species in Kras only Lycia graecarius is present. It flies from the end of February
to April, depending on the height and it is relatively rare.
Vojščica, 19.3.1988 (RŠ); Klariči, 18.3.1994 (RŠ&MZ); Kostanjevica na Krasu,
25.3.1994 (RŠ&MZ); Gorjansko, 26.2.1997 (RŠ&MZ); Jirmanec, 4.6.2011 (RŠ&MZ)
– larva.
Paraboarmia viertlii Boh.
It is widespread in South East Europe and extends back to the Caucasus and the
Middle East. It occurs in late June and early July. In Slovenia was first detected in
Podgorski Kras (Habeler, 2011), which is also the only so far known location. We
got it in three locations in Kras, in the woods and at the edge of the forest with plenty
of oak, its food plant.
Kopriva, 26.6.2011, 27.6.2012, 10.7.2013 all leg. & coll. (RŠ); Ponikve, 29.6.2013
(RŠ&MZ&CM); Podbreže, 12.7.2013 (RŠ&MZ); Kopriva, 6.7.2014 (RŠ&CM).
Acta entomologica slovenica, 24 (2), 2016
86
Fig. 5: A female of Paraboarmia viertlii attracted by artificial ultra-violet light,
Ponikve, 29.6.2013 
Peribatodes correptaria Zell.
Karst is situated at the western border of distribution of this geometrid moth of
South-Eastern Europe. Its westernmost find, which is also the first finding in Italy
(Deutsch, 2008), derives from the surroundings of Tržič – Monfalcone. In Carnelutti,
1992b, it is listed still as P. perversaria and mentioned in our country from Primorska.
As it turned out later, this species is widespread only in Western Europe, so all the
data in our region are attributed to the species P. correptaria.
We found P. correptaria in three locations in Kras: Kobjeglava – Zavodi,
29.10.1995 leg. & coll. (RŠ); Jirmanec, 4.6.2011 and 14.6.2013 (RŠ&MZ); Komen,
24.9.2013 (RŠ&MZ).
Eumannia lepraria Reb.
Its prevalence area extends from Romania via Hungary and east Austria to the
north of Italy. In Slovenia, it is known only from Primorska (Carnelutti, 1992b),
listed as Tephronia sepiaria. Later species were separated, therefore the old data for
T. sepiaria in our area are attributed to the species of E. lepraria. It flies in July and
August and inhabits both warm, dry and as well highly humid biotopes. Larvae live
on various lichens on wood and stone.
Gorenja Brestovica, 20.7.1991 (RŠ&MZ); Temnica, 14.7.1994 leg. & coll. (RŠ);
Lipa na Krasu, 18.7.1995 (RŠ&MZ); Zagrajec, 26.7.2008 leg. & coll. (RŠ); Trstelj –
Vrtovka, 23.7.2009 (RŠ&MZ&TL); Temnica, 11.8.2011 leg. & coll. (RŠ); Jirmanec,
19.8.2011 (RŠ&MZ); Kopriva, 5.7.2013 leg. & coll. (MZ).
Colostygia sericeata Schwin.
It occurs in the Apennine and the Balkan Peninsulas. In Slovenia, it is known only
from Primorska (Carnelutti, 1992), classified still as Colostygia multistrigaria. Since
C. sericeata was only recently separated from the western C. multistrigaria, old data
Radovan Øtanta, Matjaæ Zadrgal: Contribution to the knowledge of Lepidoptera fauna of the Karst
87
Fig. 6: Eumannia lepraria male, Trstelj – Vrtovka, 23.7.2009, leg. and coll. R.
Štanta
of this species in our country can be attributed to C. sericeata. We found it in Bresto-
viška dolina in Kras, where is most common in warm, dry, sunny slope, where thrives
Asperula cynanchica L., one of its food plants. Other nutritional plant Galium verum
L.  can be found at the bottom of the valley, where geometrid moth however is much
more rare. Klariči, 4.11.1993; Sela na Krasu, 10.11.2014 vse leg. & coll. MZ; Sela na
Krasu, 14.11.2010, 9.11.2011, 25.11.2011, 6.11.2012; Gorjansko, 13.11.2010 all leg.
& coll. (RŠ).
Eupithecia schiefereri Boh.
From the similar and more common E. venosata differs by a gray base color, in
addition unambiguous determination requires analysis of the genital structures. Car-
nelutti (1989) cited it from Krvavec. At Kras we found it on two locations of Komenski
kras. Larvae could be found in June on Silene latifolia Poire. ssp. alba Mill., where
they mainly feed with flowers and seeds. Komen – Jažmerca, 5.6.2010 1♂1♀ leg.,
gen. prep. & coll. (RŠ); Komen – Jažmerca, 19.5.2011 2♂2♀ (RŠ&MZ); Komen –
Jažmerca, 10.5.2012 (larva 5.6.2011) leg. & coll. (RŠ); Kobjeglava – Jelenca, 9.5.2012
leg. & coll. (MZ).
Eupithecia limbata Staud.
Published records of the species in the Slovenian fauna can not be found. Heinz
Habeler has found it at Podgorski Kras (S. Gomboc personal comment). In our
vicinity it has been found in the Trieste Karst in Basovizza - Bazovica (www.lepifo-
rum.de, 2013). The species is locally common in Spain and Italy, in the Balkans and
in the Middle East. Tied to the karst terrain, where grows food plant genus Eryngium.
Acta entomologica slovenica, 24 (2), 2016
88
Fig. 7: A male of Colostygia sericeata on a warm, dry karst slope near Sela na
Krasu, 14.11.2010
Radovan Øtanta, Matjaæ Zadrgal: Contribution to the knowledge of Lepidoptera fauna of the Karst
89
Fig. 8: Eupithecia schiefereri
- full grown larva on its host
plant Silene latifolia alba,
Komen – Jažmerca, 10.6.2011
Fig. 9: Eupithecia schiefereri - three females and a male (above right), coll. R.
Štanta
Acta entomologica slovenica, 24 (2), 2016
90
Fig. 11: Larva of Eupithecia limbata feeding on Eryngium amethystinum inflo-
rescence, Hudi Log, 3.9.2012
Fig. 10: Male of Eupithecia limbata sitting on a limestone rock, ex. larva Lokvica
– Medvejšče, 10.7.2015, leg. and coll. R. Štanta
Five larvae were found on flowers of Eryngium amethystinum L. on three locations
of Goriški kras: Lokvica-Segeti, 2.9.2011; Hudi Log, 3.9.2012; Hudi Log, 24.8.2013
all larvae leg. & det. (RŠ); Lokvica – Medvejšče, 10.7.2015, 12.7.2015 (larvae
14.9.2014) leg. & coll. (RŠ).
Notodontidae
Paradrymonia vittata Staud.
This species of South European distribution reaches at our region the northeast
limit of distribution. It is present throughout Kras and is not rare. It is tied to the
forests and thickets, where thrives Acer campestre L., its plant nutrient. It flies from
the beginning of May until mid-July. Here are only some data: Kostanjevica na
Krasu, 11.7.1994 leg. & coll. (RŠ); Kostanjevica na Krasu, 1.7.1989 (RŠ&MZ);
Gorenja Brestovica, 10.7.1997 (RŠ&ML); Jirmanec, 4.6.2011 (RŠ&MZ); Brestovica,
9.5.2013 (RŠ&MZ); Ponikve, 29.6.2013 (RŠ&MZ); Podbreže, 12.7.2013 (RŠ&MZ);
Klariči, 18.6.2014 leg. & coll. (RŠ); Kopriva, 6.7.2014 (RŠ&CM).
Erebidae
Nudaria mundana L.
European species that prefers rocky mountain areas above 700 m altitude, was
found in the Kras at three lowland locations: Kopriva (280 m), Ponikve (325 m) and
Lokev (455 m), dominated by dry meadows with hedges and forest edge. 
Lokev, 25.6.2010 (RŠ&MZ&BZ); Kopriva, 27.6.2012 leg. & coll. (RŠ); Ponikve,
29.6.2013 (RŠ&MZ&CM).
Eilema pseudocomplana Dan.
This species is locally distributed in central and southern Europe, from Spain to
Iran and is not common. It has one generation which flies in July and August. In Slo-
venia we have little published data, probably because of the similarity with E. com-
plana, but is not uncommon on the Karst plateau, especially in August, when is the
peak of its occurrence.
Lokev, 4.9.2010 (RŠ&MZ&BZ); Skopo, 28.7.2011 leg. & coll. (RŠ); Temnica,
11.8.2012 (RŠ&MZ); Jirmanec, 28.7.2012 (RŠ&MZ); Komen, 5.8.2013 leg. & coll.
(MZ).
Amata marjana Staud.
Some authors designate it as Amata kruegeri (Ragusa, 1904). Xerophilous species
that inhabits dry, open karst meadows. In Goriški kras - around Brestovica and
Lokvica two populations can be found, they occur approximately two weeks before
the sister species of Amata phegea, common throughout all Karst. 
Lokvica, 18.6.1993, 18.6.2013; Lipa na Krasu, 13.6.1999; Klariči, 31.5.1998,
1.6.2001, 8.6.2013 all leg. & coll. (RŠ); Opatje selo, 18.6 2004 leg. & coll. (MZ).
Radovan Øtanta, Matjaæ Zadrgal: Contribution to the knowledge of Lepidoptera fauna of the Karst
91
Spiris slovenica Dan.
Until now known as subspecies of Coscinia (Spiris) striata, has been now identified
as a valid species (Huemer, 2012) which replaces the previously asserted C. striata in
Slovenia, Italy, southern Austria and Croatia. It is active during the day, at night can
be attracted with artificial light. Its habitat are dry meadows. At Kras is a common
and widespread everywhere: 
Sežana - Dol Leskovec, 14.6.1986; Opatje selo, 8.7.1989; Dutovlje, 21.6.1991;
Lipa na Krasu, 18.6.2001; Komen, 22.6.2013; Brestovica, 22.6.2013; Vrhovlje,
6.6.2014 all leg. & coll. (RŠ); Lokev, 9.6.2013 leg. & coll. (MZ).
Hypena lividalis Hübn.
This interesting South European erebid moth we got only once in the Komenski
kras. It probably came to us as a migrant. 
Zagrajec, 30.9.2010 leg. & coll. (RŠ).
Noctuidae
Euchalcia modestoides Poole
This Eurasian species is not common, it flies in June and July. One male was
found southwest of Ponikve: 
Ponikve, 12.06.2013 leg. & coll. (RŠ).
Cucullia gozmanyi G. & L. Ron.
In Slovenia, it is known only in the Komenski kras (Lasan, 2000). Here are listed
some of the findings, which expand the known distribution of the species to the
Goriški kras:
Kobjeglava – Jelenca, 15.4.1998 ex larva on Scrophularia phenicaeum L. (larvae
22.5.1997); Kostanjevica na Krasu, 3. - 15.4.1998 (larvae 29.5.1997); Hudi log,
20.4.1998 (larvae 29.5.1997); 
Lipa na Krasu, 15. - 18.4.2009 ex ovum on S. phenicaeum (ovum 9.5.2010, larvae
12. - 14.5.2010); 
Kobjeglava – Jelenca, 12. - 26.4.2001 (larvae 23.5.2000); Gorjansko, 2. - 8.6.2001
(larvae 21.5.2000) all leg. & coll. (RŠ).
Schinia cardui Hübn.
This owlet moth has been found so far in three locations in Slovenia: Laško (Car-
nelutti, 1971), Košaki at Maribor (Lesar, Jež, 2006) and Sečovlje (S. Gomboc personal
comment). From our proximity is known in Austria, Croatian Istria and Triest Karst,
Fernetti (Fernetiči) (L. Morin personal comment). This xerophilous species we found
in dry, open areas in Karst, particularly along the ways on Picris hieracioides L.
where it feeds on flowers and seeds.
Sežana, 4.9.2010 3x larvae (RŠ&MZ&BZ); Pliskovica, 9. - 12.8.2011 2♂1♀
(larvae 14.9.2010); Krajna vas, 14.8.2011 1♂ (larva 14.9.2010); Pliskovica, 24. -
28.7.2012 5♂6♀ (larvae 15.8.2011); Veliki Dol, 28.7.2012 1♂ (larva 15.8.2011);
Acta entomologica slovenica, 24 (2), 2016
92
Pliskovica, 15.8.2011 1♀, Lokvica – Medvejšče, 1.8.2012 1♀ all leg. & coll. (RŠ);
Vojščica, 29.7.2012; Brje pri Komnu, 21.7.2013, 26.7.2014 all leg. & coll. (MZ).
Oligia dubia Heyd.
It occurs around the Alps from Switzerland to the Pannonian plains in the east. It
is more common on the southern slopes of the Alps and the eastern provinces of the
northern Adriatic. At Kras it is common from the middle of May until the end of
June. In the middle of this period, when is its summit occurrence O. dubia is almost
as frequent as O. latruncula. Typical pattern of the front wings differentiates O.
dubia from O. latruncula due to the lighter colored field between the post-median
fascia and sub-terminal fascia. In a typical O. dubia the lower part of the post-median
fascia is light, almost white, and it is in the lower edge of the wings slightly curved
outwards, while in a typical O. latruncula it is on average darker, almost straight and
reaches the lower edge of the wing at right angle. There are also some exceptions, so
an analysis of genital armatures is required.
Kostanjevica na Krasu, 27.5.1988 (RŠ&MZ); Kobjeglava, 19.5.1997 (RŠ&MZ);
Lokvica, 25.5.2002 (RŠ&BZ&SG); Lipa na Krasu, 12.5.2006 (RŠ&MZ); Zagrajec,
22.5.2009 (RŠ&MZ); Komen – Jažmerca, 5.6.2010 (RŠ&MZ&BZ); Komen,
19.5.2011 (RŠ&MZ); Kopriva, 30.5.2011 (RŠ); Skopo, 30.5.2011 and 10.6.2011
(RŠ); Jirmanec, 4.6.2011 (RŠ&MZ); Vrhpolje - Sv. Tomaž, 16.6.2012 (RŠ); Ponikve,
12.6.2013 (RŠ); Ponikve, 29.6.2013 (RŠ&MZ&CM); Brje pri Komnu, 25.4.2014
(RŠ&MZ) all gen. prep. & coll. (RŠ).   
Radovan Øtanta, Matjaæ Zadrgal: Contribution to the knowledge of Lepidoptera fauna of the Karst
93
Fig. 12: Two caterpillars of Schinia cardui feeding on flowers of Picris hieracioi-
des, Pliskovica, 14.9.2010
Perigrapha rorida Friv.
Mediterranean-Asian owlet moth occurs in March and April. Larvae are fed exc-
lusively with Paliurus spina-christi Mill. Regularly occurring in west of Goriški and
Komenski kras. 
Klariči, 25.3.1994, 26.3.1995, 15.3.2002 (RŠ & MZ); Vojščica, 25.3.1996 leg. &
coll. (RŠ); Klariči, 19. - 25.3.1996 ex larva leg. & coll. (RŠ); Klariči, 15.03.2014 leg.
& coll. (MZ).
Euxoa cos Hübn.
It is a Mediterranean – West Asian species, Carnelutti (1992) is mentioning it for
Vipava - old data. It has recently been found in Iamiano - Jamlje in Gorizia karst in
Italy (Deutsch, 2008). One specimen was found also on the Slovenian side: 
Temnica, 29.8.2008 leg. & coll. (MZ).
Discussion
755 species were found in the area of the Karst, compared with approximately
390 previously mentioned literature data, quite a significant increase in knowledge of
Macrolepidoptera for Komenski and Sežanski kras. Comparison with approximately
670 recorded species on the more explored Italian side indicates that the vast majority
of the registered fauna on the Italian side was found with our observations. Larger
number of species is mostly due to greater surface of Kras on the Slovenian side and
because there is a greater distance from the Gulf of Trieste, having on average, cooler
climate. Typical species of these cooler areas, which are not confirmed for the Italian
side, are populated in the southeastern and eastern part of the plateau. These are eg.:
Acta entomologica slovenica, 24 (2), 2016
94
Fig. 13: Euxoa cos male, Temnica, 29.8.2008, leg. and coll. M. Zadrgal
Euphyia biangulata, Nudaria mundana, Brachionycha nubeculosa, Mniotype adusta,
Staurophora celsia, Agrotis cinerea and others.
Fauna on the Italian side leads in the number of hygrophilous species, some of
them were not found on our side: Lycaena dispar Gastropacha populifolia, Perizoma
flavofasciata, Cerura vinula, Herminia tenuialis, Deltote bankiana, Deltote uncula,
Apterogenum ypsillon, Mythimna pallens, Mythimna impure and others. The literature
does not always show the exact locations of these species; mainly they are surroundings
of karst lakes, Doberdob lake - Lago di Doberdo and Prelosno lake - Lago di Pietrarossa
and for some wetland areas of nearby Tržič - Monfalcone or Milje - Muggia.
We found some hygrophilous species on the Slovenian side in the surroundings of
ponds and their residues as the only surface water. The most suitable due to multiannual
abandonment and large edge vegetation zone is a pond in Komen, where we found
the majority of hygrophilous species: Athetis gluteosa, Rhizedra lutosa, Globia spar-
ganii and others.
There are fewer differences regarding xerophilous species among the Italian and
the Slovenian part of the Karst. Thanks mainly to the warmer northwestern part on
our side we found there almost all present species: Polygonia egea, Gnopharmia
stevenaria, Rhoptria asperaria, Colostygia sericeata, Dysauxes famula, Metachrostis
velox, Zebeeba falsalis, Meganephria bimaculosa, Dryobotodes monochroma, Aporo-
phyla canescens, Perigrapha rorida and others.
In the studied area of the Karst the most notable regarding diversity are zones 1
and 5. Among the 56 species, which were found only in the zone 1 stand out xerop-
hilous species. The main factor is the impact of a warmer local climate, especially in
Brestoviški dol and neighboring lower parts of Goriški and Komenski kras, where
the influence of the sea could be felt. Therefore some xerophilous species we found
only here: Acanthopsyche ecksteini, Pieris ergane, Polyommatus escheri, Polygonia
egea, Nychiodes dalmatina, Amata marjana, Catocala conjuncta, Odice suava, Dry-
obotodes carbonis, Perigrapha rorida, Euxoa cos and others.
Another group typical only for zone 1 are random finds species, which are more
common in the near Vipava Valley: Lycaena hippothoe, Clostera anastomosis, Glup-
hisia crenata, Furcula furcula, Furcula bifida, Spilosoma lutea, Euproctis chrysorr-
hoea, Sphrageidus similis, Acronicta auricoma, Apamea crenata.
The discovery of Erebid moth Pelosia muscerda in Klariči is also interesting, the
latter is probably accidental guest from the surroundings of karst lakes on the Italian
side. Similarly, Dryobota labecula and Dryobotodes tenebrosa probably flew from
the stands of holly oaks at the coast of the Gulf of Trieste. 
Zone 5 is the second according to the number of unique species. We found 33
species here which in other zones can not be found. A key role in this has the set of
hills with altitude about 700 m from Jirmanec, more commonly known as Kokoš to
V. Gradišče and Ograda. Colder climate on the top generates the presence of certain
species, which were found only here: Peribatodes secundaria, Charissa obscurata,
Scotopteryx moeniata, Catarhoe rubidata, Euphyia biangulata, Thera vetustata, Hy-
drelia flammeolaria, Eupithecia trisignaria, Eupithecia absinthiata, Xanthia icteritia,
Eugnorisma depuncta and others.
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95
The warm southern slopes are different, there are some interesting xerophilous
and mesophilous species: Phaiogramma etruscaria, Euphyia adumbraria, Horisme
calligraphata, Perigrapha i-cinctum slovenica.
For the entire Kras, especially for the central zones 2,3 and 4 which are characte-
rized by the coexistence of species with different ecological requirements. This is
enabled by a heterogeneous structure of microhabitats, conditioned by karst pheno-
mena. Thus we find in small areas xerophilous, mesophilous and hygrophilous species.
Hygrophilous species retain in sinkholes, some of the rarer are: Coenonympha oedip-
pus, Acosmetia caliginosa, Photedes morrisii sohnretheli, Mythimna riparia, Orthosia
gracilis. At the edge of sinkholes mesophilous species are found, among which we
highlight Ennomos autumnaria, Odontopera bidentata, Polia nebulosa, Moma alpium;
on the slopes xerophilous species of Lepidoptera, among the rarer eg.: Ennomos
quercaria, Zekelita antiqualis, Praestilbia armeniaca, Aporophyla canescens.
Among central zones slightly stands out zone 2 with 23 species which are not
found elsewhere and consist of mainly xerophilous species: Gnopharmia stevenaria,
Rhoptria asperaria, Eublemma parva, Epimecia ustula, because the lower southwest
part of the zone falls within the warmer climate area.
Due to the small number of inventories in zones 4 and 5, the number of species
found there is the smallest. We expect that with additional research the differences in
correlation with other zones will diminish, as in these two zones over 110 ubiquitous
species are missing but are being represented in all other zones.
A brief mention also deserve the most common species of Karst. In selection of
the latter we took into account the average abundance throughout the period of 30
years. We canceled out seasonal fluctuations, which were quite large. Kras is being
largely dominated by the tree species of Lepidoptera due to its overgrow with shrubs
and trees. Among these species we can find: Colotois pennaria, Operophtera brumata,
Eupithecia ericeata, Thaumetopoea pityocampa, Diloba caeruleocephala, Craniop-
hora ligustri, Amphipyra pyramidea, Allophyes oxyacanthae, Conistra vaccinii, Co-
nistra erythrocephala, Orthosia cerasi, Orthosia cruda.
The butterflies were somewhat neglected in our observations, so we were unable
to confirm all known species. According to the Atlas of butterflies of Slovenia
(Verovnik & al., 2012) 9 additional species are also present: Leptotes pirithous,
Satyrium acaciae, Satyrium w-album, Apatura ilia, Aphantopus hyperantus, Araschnia
levana, Brenthis ino, Satyrus ferula and Colias hyale, as well as 15 species, which
were found during the inventory of the Kras by Stanislav Gomboc: Hyles gallii,
Laothoe populi, Peribatodes umbraria, Idaea muricata, Scopula confinaria, Nebula
achromaria, Epirrita autumnata, Aplocera praeformata, Atolmis rubricollis, Hy-
phoraia aulica, Lygephila procax, Eublemma ostrina, Bryophila domestica, Hoplod-
rina octogenarian and Noctua orbona.
A few more species known from the literature are expected to be found, but we
have not managed to find them yet: Acanthopsyche atra, Apterona helicoidella (parth.
form), Paranthrene tabaniformis, Synanthedon stomoxiformis, Daphnis nerii, Erannis
ankeraria, Charissa variegata, Cerura vinula, Chelis maculosa, Catocala diversa,
Omia cymbalariae, Dichonia aeruginea.
Acta entomologica slovenica, 24 (2), 2016
96
Along with these, the number of species found in the Karst region rose to 791,
which represents more than 52 % of known Macrolepidoptera species in Slovenia
where from Carnelutti (1992a, 1992b), Gomboc & Lasan (2006) and Lesar & Govedič
(2010) is to conclude that the slovenian fauna of Macrolepidoptera counts around
1500 species.
For slovenian Kras is in older literature also mentioned Pyrgus onopordi, which
now applies to extinct species in Slovenia (Verovnik & al., 2012) and some dubious
species: Rebelia sapho, Asthena anseraria, Autophila anaphanes and Diarsia mendica,
some of them had already been critically discussed: Pyrgus cirsii, Coscinia cribraria,
Malacosoma franconica (Carnelutti, 1971).
Species Conservation overview
Karst is one of the hotspots of biodiversity, but we have witnessed accelerated
change of the Kras area in recent decades. The main reason is the abandonment of
traditional agricultural use of the landscape, resulting in overgrowth, thereby the re-
ducement of the typical dry grasslands. The increase of urban environments, indu-
strialization and the introduction of intensive farming also have a negative impact,
but in the addressed area represent more marginal role. With the disappearance of the
typical karst landscape we are losing the diversity of flora and fauna. The public is
recently aware of this natural wealth, since almost the entire area of the central Kras
has been placed in a special protection area Natura 2000 (Kras SI5000023), exempted
are only Sežana, Divača and Kozina with their neighborhoods. 
In the attached checklist of species there are 67 species, which have the status of
the endangered species in Slovenia (UL RS, 82/2002), from which 43 are in the cat-
egory of endangered species - E  and 24 of them are vulnerable species - V.
There are 7 prioirity species living in Kras according to the Habitats Directive
(92/43/EC), of those are 4 from Attachment II.: Callimorpha (Euplagia) quadripunc-
taria, Coenonympha oedippus, Eriogaster catax in Euphydryas aurinia and 3 from
Attachment IV.: Lopinga achine, Parnassius mnemosyne and Zerynthia polyxena.
Protected species on the state level are laid down with Regulation of the conser-
vation of natural habitats and of wild fauna and flora (Ur. l. RS, 2004a). 23 species
are listed in both attachments.
In Attachment 1, which protects the species and populations, there are 30 species
from Kras (following in alphabetical order, as in the Regulation): Callimorpha (Eu-
plagia) quadripunctaria, Carcharodus flocciferus (floccifera),  Chazara briseis (Ex?),
Chesias rufata, Coenonympha oedippus, Diaphora luctuosa, Eriogaster catax, Eu-
blemma purpurina, Euphydryas aurinia, Eupithecia ochridata, Gnopharmia steve-
naria, Gortyna puengeleri, Hecatera cappa, Hemaris fuciformis, Hemaris tityus,
Lopinga achine, Nudaria mundana, Paradrymonia vittata, Parnassius mnemosyne,
Perigrapha i-cinctum, Perisomena (Saturnia) caecigena, Phragmatobia luctifera,
Plebicula (Polyommatus) escheri, Polygonia egea, Praestilbia armeniaca, Saturnia
pyri, Schinia cardui, Zerynthia polyxena, Zygaena ephialtes, Zygaena punctum.
Radovan Øtanta, Matjaæ Zadrgal: Contribution to the knowledge of Lepidoptera fauna of the Karst
97
42 species is from Attachment 2, which protects habitats: Acanthopsyche zelleri,
Apamea aquila, Archanara sparganii, Atethmia centrago, Athetis gluteosa, Callimor-
pha (Euplagia) quadripunctaria, Carcharodus flocciferus (floccifera), Chazara briseis
(Ex?), Chesias rufata, Chortodes (Photedes) morrisii sohnretheli (in Regulation as
Chortodes morrisii and Chortodes sohnretheli), Coenonympha oedippus, Colostygia
sericeata, Coscinia slovenica (in Regulation as Coscinia striata), Crassagrotis crassa
(Agrotis bigramma), Cucullia gozmanyi, Diaphora luctuosa, Eilema pseudocomplana,
Eriogaster catax, Eublemma purpurina, Euphydryas aurinia, Eupithecia ochridata,
Euxoa cos, Gnopharmia stevenaria, Hemaris fuciformis, Hemaris tityus, Metachrostis
dardouini, Metachrostis velox, Nudaria mundana, Odice suava, Orbona fragariae,
Parnassius mnemosyne, Perigrapha i-cinctum, Phragmatobia luctifera, Plebicula
(Polyommatus) escheri, Plebicula (Polyommatus) thersites, Polygonia egea, Praestil-
bia armeniaca, Rhizedra lutosa, Rhyparia purpurata, Zerynthia polyxena, Zygaena
ephialtes, Zygaena punctum.
Data on the recorded species of Macrolepidoptera confirm that the importance of
the nature protection area is enormous and the collected data could represent the
basis for the monitoring and supervision of the endangered, protected and other
species.
Acknowledgements
We are very grateful to all of our dear friends and colleagues who have accompanied
us on excursions and shared with us data: Bojan Zadravec, dr. Carlo Morandini,
Stanislav Gomboc, Mojmir Lasan, Tone Lesar (†), Jozef Debets and all the other
occasional companions. Special thanks to Stanislav Gomboc for his data of Slovenian
Kras and furthermore to Lucio Morin and dr. Carlo Morandini for the information
from the Italian side.
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103
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ta
 g
er
m
an
ic
a
(R
Š)
; L
ok
vi
ca
,
9.
4.
20
07
 (R
Š&
M
Z)
Ac
an
th
op
sy
ch
e 
ze
lle
ri
(M
an
n,
 1
85
5)
x
x
x
4
co
m
m
on
Lo
kv
ic
a,
 2
3.
4.
20
06
 (R
Š)
; O
pa
tje
 se
lo
, 7
.4
.2
00
7
(R
Š)
; K
os
ta
nj
ev
ic
a 
na
 K
ra
su
, 1
1.
4.
20
07
 (R
Š)
Acta entomologica slovenica, 24 (2), 2016
104
C
an
ep
ho
ra
 h
ir
su
ta
(P
od
a,
 1
76
1)
x
x
6
ra
re
K
os
ta
nj
ev
ic
a 
na
 K
ra
su
, 1
1.
6.
19
88
 (R
Š&
M
Z)
;
Za
gr
aj
ec
, 4
.6
.2
01
4 
e.
 p
. 2
5.
10
.2
01
3 
(R
Š&
M
Z)
Pa
ch
yt
he
lia
 v
ill
os
el
la
(O
ch
se
nh
ei
m
er
, 1
81
0)
x
x
5,
6
ra
re
Li
pa
 n
a 
K
ra
su
, 9
.6
.2
00
0 
(M
Z)
; V
oj
šč
ic
a,
 6
.6
.2
01
1
(M
Z)
; K
om
en
, 2
8.
5.
20
12
 (M
Z)
Pt
ilo
ce
ph
al
a 
pl
um
ife
ra
 m
ed
ite
rr
an
ea
(L
ed
er
er
 1
85
2)
x
x
3,
4
ra
re
Li
pa
 n
a 
K
ra
su
, 2
0.
4.
19
96
 (R
Š&
M
Z)
; K
os
ta
nj
ev
ic
a
na
 K
ra
su
, 1
6.
3.
19
97
 (R
Š)
; B
rje
 p
ri 
K
om
nu
,
4.
4.
19
99
 (M
Z)
; K
os
ta
nj
ev
ic
a 
na
 K
ra
su
, 1
7.
4.
20
06
(R
Š)
M
eg
al
op
ha
ne
s v
ic
ie
lla
(D
en
is
 &
 S
ch
iff
er
m
ül
le
r, 
17
75
)
x
x
x
x
x
5,
6
co
m
m
on
Ph
al
ac
ro
pt
er
ix
 p
ra
ec
el
le
ns
(S
ta
ud
in
ge
r, 
18
70
)
x
x
x
x
x
2,
3,
4
ve
ry
 c
om
m
on
C
O
SS
ID
A
E 
(4
)
C
os
su
s c
os
su
s(
Li
nn
ae
us
, 1
75
8)
x
x
x
5,
6,
7
ra
re
K
la
rič
i, 
16
.5
.1
99
4 
(R
Š&
M
Z)
; S
ko
po
, 2
6.
6.
20
11
(R
Š)
; P
on
ik
ve
, 2
9.
6.
20
13
 (R
Š&
M
Z&
C
M
); 
K
ra
jn
a
va
s, 
18
.7
.2
01
4 
(R
Š&
M
Z)
Pa
ra
hy
po
pt
a 
ca
es
tr
um
(H
üb
ne
r, 
18
03
-1
80
8)
x
x
x
6,
7
ra
re
G
or
en
ja
 B
re
st
ov
ic
a,
 2
0.
7.
19
91
 (R
Š&
M
Z)
; Š
kr
bi
na
,
18
.7
.1
99
5 
(R
Š&
M
Z)
; Z
ag
ra
je
c,
 2
6.
7.
20
08
(R
Š&
M
Z)
; V
el
ik
i D
ol
, 2
7.
6.
20
12
 (R
Š)
  
D
ys
pe
ss
a 
ul
ul
a
(B
or
kh
au
se
n,
 1
79
0)
x
x
x
x
x
4,
5,
6,
7
no
t c
om
m
on
Ze
uz
er
a 
py
ri
na
(L
in
na
eu
s, 
17
61
)
x
x
x
x
6,
7,
8,
10
no
t c
om
m
on
K
os
ta
nj
ev
ic
a 
na
 K
ra
su
, 1
.7
.1
98
9 
(R
Š&
M
Z)
;
K
ob
je
gl
av
a,
 2
.1
0.
19
95
 (R
Š)
; P
on
ik
ve
, 2
9.
6.
20
13
(R
Š&
M
Z&
C
M
); 
K
op
riv
a,
 6
.7
.2
01
4 
(R
Š&
C
M
)
SE
SI
ID
A
E 
(4
)
Sy
na
nt
he
do
n 
an
dr
en
ae
fo
rm
is
(L
as
pe
yr
es
, 1
80
1)
x
/
ve
ry
 ra
re
O
pa
tje
 se
lo
, 2
3.
4.
20
06
 - 
la
rv
ae
 o
n 
Vi
bu
rn
um
la
nt
an
a
(R
Š)
Sy
na
nt
he
do
n 
m
yo
pa
ef
or
m
is
(B
or
kh
au
se
n,
 1
78
9)
x
/
no
t c
om
m
on
K
om
en
, 1
2.
4.
20
07
 - 
la
rv
ae
 o
n 
M
al
us
 d
om
es
tic
a
(R
Š)
Sy
na
nt
he
do
n 
tip
ul
ifo
rm
is
(C
le
rc
k,
 1
75
9)
x
/
no
t c
om
m
on
K
os
ta
nj
ev
ic
a 
na
 K
ra
su
, 2
4.
5.
20
03
 - 
la
rv
ae
 o
n 
Ri
be
s
sp
p.
 (R
Š)
C
ha
m
ae
sp
he
ci
a 
ae
ri
fr
on
s(
Ze
lle
r, 
18
47
)
x
/
no
t c
om
m
on
V
oj
šč
ic
a,
 2
7.
12
.2
01
2 
(R
Š)
; S
el
o 
na
 K
ra
su
,
2.
11
.2
01
3 
(R
Š)
 - 
ex
uv
ia
e 
an
d 
la
rv
ae
 in
 th
e 
ro
ot
s o
f
Sa
tu
re
ja
 m
on
ta
na
Radovan Øtanta, Matjaæ Zadrgal: Contribution to the knowledge of Lepidoptera fauna of the Karst
105
H
ET
ER
O
G
Y
N
ID
A
E 
(1
)
H
et
er
og
yn
is
 p
en
el
la
(H
üb
ne
r, 
18
19
)
x
x
x
5,
6
ve
ry
 c
om
m
on
Ji
rm
an
ec
, 4
.6
.2
01
1 
pu
pa
 (R
Š&
M
Z)
; P
od
br
ež
e,
9.
6.
20
13
 e
. p
. 1
.5
.2
01
3 
(R
Š&
M
Z)
; V
rh
ov
lje
,
6.
6.
20
14
 e
xu
vi
ae
 (R
Š)
; K
az
lje
, 1
3.
5.
20
15
 (R
Š)
LI
M
A
C
O
D
ID
A
E 
(1
)
Ap
od
a 
lim
ac
od
es
(H
uf
na
ge
l, 
17
66
)
x
x
x
x
x
5,
6,
7,
8
co
m
m
on
ZY
G
A
EN
ID
A
E 
(1
6)
Jo
rd
an
ita
 b
ud
en
si
s(
A
d.
 &
 A
u.
 S
pe
ye
r, 
18
58
)
x
x
5
ve
ry
 ra
re
D
at
a 
in
 th
e 
C
ha
pt
er
 re
ga
rd
in
g 
in
te
re
st
in
g 
sp
ec
ie
s
Jo
rd
an
ita
 n
ot
at
a
(Z
el
le
r, 
18
47
)
x
x
6
ra
re
Li
pa
 n
a 
K
ra
su
, 9
.6
.2
00
2 
(R
Š)
; K
op
riv
a,
 1
0.
6.
20
11
(R
Š)
 - 
in
sp
ec
tio
n 
of
 th
e 
ge
ni
ta
l s
tru
ct
ur
es
 (R
Š)
Jo
rd
an
ita
 c
hl
or
os
(H
üb
ne
r, 
18
13
)
x
6,
7
ra
re
Ji
rm
an
ec
, 1
6.
7.
20
10
 (M
Z)
; J
irm
an
ec
, 8
.7
.2
01
1 
(R
Š)
Jo
rd
an
ita
 g
lo
bu
la
ri
ae
(H
üb
ne
r, 
17
93
)
x
x
x
x
x
5,
6
co
m
m
on
Ad
sc
ita
 m
an
ni
i(
Le
de
re
r, 
18
53
)
x
x
x
x
x
5,
6,
7
co
m
m
on
Ad
sc
ita
 st
at
ic
es
(L
in
na
eu
s, 
17
58
)
x
5
ve
ry
 ra
re
K
ob
je
gl
av
a 
- J
el
en
ca
, 2
6.
5.
20
00
 (R
Š)
Zy
ga
en
a 
pu
nc
tu
m
O
ch
se
nh
ei
m
er
, 1
80
8
x
x
x
5,
6
ra
re
D
at
a 
in
 th
e 
C
ha
pt
er
 re
ga
rd
in
g 
in
te
re
st
in
g 
sp
ec
ie
s
Zy
ga
en
a 
pu
rp
ur
al
is
(B
rü
nn
ic
h,
 1
76
3)
x
x
x
x
x
4,
5,
6
ve
ry
 c
om
m
on
Zy
ga
en
a 
ca
rn
io
lic
a
(S
co
po
li,
 1
76
3)
x
x
x
x
x
6,
7
co
m
m
on
Zy
ga
en
a 
lo
ti
(D
en
is
 &
 S
ch
iff
er
m
ül
le
r, 
17
75
)
x
x
x
x
x
5,
6
ve
ry
 c
om
m
on
Zy
ga
en
a 
os
te
ro
de
ns
is
R
ei
ss
, 1
92
1
x
x
5
ve
ry
 ra
re
O
pa
tje
 S
el
o,
 9
.6
.1
99
5 
(M
Z)
; K
la
rič
i, 
31
.5
.1
99
8
(R
Š)
; K
ob
je
gl
av
a 
- J
el
en
ca
, 2
6.
5.
20
00
 (R
Š)
Zy
ga
en
a 
vi
ci
ae
(D
en
is
 &
 S
ch
iff
er
m
ül
le
r, 
17
75
)
x
x
6
ra
re
Li
pa
 n
a 
K
ra
su
, 9
.6
.2
00
0 
(M
Z)
; L
ip
a 
na
 K
ra
su
,
9.
6.
20
02
 (R
Š)
; J
irm
an
ec
, 1
7.
6.
20
11
 (R
Š)
Zy
ga
en
a 
ep
hi
al
te
s(
Li
nn
ae
us
, 1
76
7)
x
6
ve
ry
 ra
re
K
la
rič
i, 
25
.6
.1
98
8 
(R
Š)
; K
la
rič
i, 
5.
7.
19
95
 (M
Z)
Zy
ga
en
a 
tr
an
sa
lp
in
a
(E
sp
er
, 1
78
0)
x
x
x
x
x
5,
6
no
t c
om
m
on
N
ov
a 
va
s, 
14
.9
.2
01
4 
- u
nu
su
al
 d
at
e,
 o
ne
 m
al
e
ob
se
rv
ed
Zy
ga
en
a 
fil
ip
en
du
la
e
(L
in
na
eu
s, 
17
58
)
x
x
x
x
x
5,
6
ve
ry
 c
om
m
on
Zy
ga
en
a 
lo
ni
ce
ra
e
(S
ch
ev
en
, 1
77
7)
x
x
x
x
5,
6
co
m
m
on
Acta entomologica slovenica, 24 (2), 2016
106
TH
Y
R
ID
ID
A
E 
(1
)
Th
yr
is
 fe
ne
st
re
lla
(S
co
po
li,
 1
76
3)
x
6,
7
ve
ry
 ra
re
O
pa
tje
 se
lo
, 8
.7
.1
98
9 
(R
Š)
; K
la
rič
i, 
24
.6
.1
99
6 
(R
Š)
PA
PI
LI
O
N
ID
A
E 
(4
)
Ze
ry
nt
hi
a 
po
ly
xe
na
(D
en
is
 &
 S
ch
iff
er
m
ül
le
r, 
17
75
)
x
x
x
x
x
3,
4,
5
co
m
m
on
K
os
ta
nj
ev
ic
a 
na
 K
ra
su
, 1
6.
4.
19
89
 (R
Š)
; L
ip
a 
na
K
ra
su
, 2
6.
4.
20
00
 (R
Š)
; D
ut
ov
lje
, 1
8.
4.
20
09
 (R
Š)
Pa
rn
as
si
us
 m
ne
m
os
yn
e
(L
in
na
eu
s, 
17
58
)
x
x
x
x
5,
6
no
t c
om
m
on
K
ob
je
gl
av
a,
 1
5.
6.
19
90
 (R
Š)
; T
rs
te
lj,
 1
9.
5.
19
91
(R
Š)
; L
ip
a 
na
 K
ra
su
, 1
4.
5.
20
10
 (M
Z)
; K
az
lje
,
13
.5
.2
01
5 
(R
Š)
Ip
hi
cl
id
es
 p
od
al
ir
iu
s(
Li
nn
ae
us
, 1
75
8)
x
x
x
x
x
3,
4,
5,
6,
7,
8
co
m
m
on
Pa
pi
lio
 m
ac
ha
on
Li
nn
ae
us
, 1
75
8
x
x
x
x
x
4,
5,
6,
7,
8,
9
no
t c
om
m
on
H
ES
PE
R
II
D
A
E 
(1
5)
Er
yn
ni
s t
ag
es
(L
in
na
eu
s, 
17
58
)
x
x
x
x
x
4,
5,
6,
7,
8,
9
co
m
m
on
C
ar
ch
ar
od
us
 a
lc
ea
e
(E
sp
er
, 1
78
0)
x
6
ve
ry
 ra
re
B
re
st
ov
ic
a,
 2
8.
6.
19
95
 (R
Š)
C
ar
ch
ar
od
us
 fl
oc
ci
fe
ra
(Z
el
le
r, 
18
47
)
x
x
x
x
5,
6,
7
ra
re
K
la
rič
i, 
31
.5
.1
99
8 
(R
Š)
; L
ok
ev
, 8
.7
.2
00
2 
(R
Š)
;
Li
pa
 n
a 
K
ra
su
, 2
2.
5.
20
10
 (R
Š)
; K
ob
je
gl
av
a,
22
.5
.2
01
1 
(R
Š)
; K
ra
jn
a 
va
s, 
21
.5
.2
01
4 
(R
Š)
;
V
rh
ov
lje
, 6
.6
.2
01
4 
(R
Š)
Sp
ia
lia
 se
rt
or
iu
s(
H
of
fm
an
ns
eg
g,
 1
80
4)
x
x
x
x
x
4,
5,
6,
7,
8
co
m
m
on
Py
rg
us
 c
ar
th
am
i(
H
üb
ne
r, 
18
13
)
x
x
x
x
x
5,
6
no
t c
om
m
on
B
re
st
ov
ic
a,
 1
6.
6.
20
02
 (R
Š)
; L
ip
a 
na
 K
ra
su
,
30
.6
.1
99
0 
(R
Š)
; G
or
ja
ns
ko
, 2
2.
5.
20
11
 (R
Š)
;
K
op
riv
a,
 2
9.
5.
20
11
 (R
Š)
; P
od
br
ež
e,
 6
.6
.2
01
4 
(R
Š)
Py
rg
us
 m
al
va
e
(L
in
na
eu
s, 
17
58
)
x
5
ra
re
D
at
a 
in
 th
e 
C
ha
pt
er
 re
ga
rd
in
g 
in
te
re
st
in
g 
sp
ec
ie
s
Py
rg
us
 m
al
vo
id
es
(E
lw
es
 &
 E
dw
ar
ds
, 1
89
7)
x
x
4,
5,
6
no
t c
om
m
on
D
at
a 
in
 th
e 
C
ha
pt
er
 re
ga
rd
in
g 
in
te
re
st
in
g 
sp
ec
ie
s
Py
rg
us
 a
rm
or
ic
an
us
(O
be
rth
ür
, 1
91
0)
x
x
x
5,
6
ra
re
K
la
rič
i, 
6.
5.
20
00
 (M
Z)
; K
ob
je
gl
av
a,
 2
2.
5.
20
11
(R
Š)
; K
om
en
, 2
9.
5.
20
11
 (R
Š)
; G
or
ja
ns
ko
,
22
.6
.2
01
3 
(R
Š)
; V
rh
ov
lje
, 6
.6
.2
01
4 
(R
Š)
; L
ok
ev
,
6.
6.
20
14
 (R
Š)
Py
rg
us
 a
lv
eu
s t
re
be
vi
ce
ns
is
(W
ar
re
n,
 1
92
6)
x
6
ve
ry
 ra
re
Lo
ke
v,
 6
.6
.2
01
4 
(R
Š)
Radovan Øtanta, Matjaæ Zadrgal: Contribution to the knowledge of Lepidoptera fauna of the Karst
107
H
et
er
op
te
ru
s m
or
ph
eu
s(
Pa
lla
s, 
17
71
)
x
x
x
6,
7
ra
re
O
pa
tje
 se
lo
, 2
3.
6.
20
06
 (R
Š)
; K
ob
je
gl
av
a,
 1
5.
6.
19
90
(R
Š)
C
ar
te
ro
ce
ph
al
us
 p
al
ae
m
on
(P
al
la
s, 
17
71
)
x
5
ve
ry
 ra
re
Šk
rb
in
a,
 6
.5
.2
01
4 
(R
Š)
Th
ym
el
ic
us
 li
ne
ol
a
(O
ch
se
nh
ei
m
er
, 1
80
8)
x
x
x
x
x
5,
6,
7
ve
ry
 c
om
m
on
Th
ym
el
ic
us
 sy
lv
es
tr
is
(P
od
a,
 1
76
1)
x
x
6,
7
ra
re
Li
pa
 n
a 
K
ra
su
, 1
9.
6.
20
10
 (R
Š)
; D
ob
ra
vl
je
,
20
.7
.2
01
3 
(R
Š)
H
es
pe
ri
a 
co
m
m
a
(L
in
na
eu
s, 
17
58
)
x
x
x
x
x
6,
7,
8,
9
no
t c
om
m
on
O
ch
lo
de
s s
yl
va
nu
s(
Es
pe
r, 
17
77
)
x
x
x
x
x
5,
6,
7,
8,
9
co
m
m
on
PI
ER
ID
A
E 
(1
2)
Le
pt
id
ea
 si
na
pi
s(
Li
nn
ae
us
, 1
75
8)
x
x
x
x
x
3,
4,
5,
6,
7,
8,
9
ve
ry
 c
om
m
on
An
th
oc
ha
ri
s c
ar
da
m
in
es
(L
in
na
eu
s, 
17
58
)
x
x
x
x
x
3,
4,
5,
6
co
m
m
on
Ap
or
ia
 c
ra
ta
eg
i(
Li
nn
ae
us
, 1
75
8)
x
x
x
x
x
4,
5,
6,
7
ve
ry
 c
om
m
on
K
os
ta
nj
ev
ic
a 
na
 K
ra
su
, 2
2.
4.
19
81
 (R
Š)
 - 
th
e
ea
rli
es
t d
at
e
Pi
er
is
 b
ra
ss
ic
ae
(L
in
na
eu
s, 
17
58
)
x
x
x
x
x
5,
6,
7,
8,
9,
10
no
t c
om
m
on
Pi
er
is
 m
an
ni
i(
M
ay
er
, 1
85
1)
x
x
x
x
x
3,
4,
5,
6,
7,
8,
9
co
m
m
on
Pi
er
is
 ra
pa
e
(L
in
na
eu
s, 
17
58
)
x
x
x
x
x
3,
4,
5,
6,
7,
8,
9,
10
co
m
m
on
Pi
er
is
 e
rg
an
e
(G
ey
er
, 1
82
8)
x
6,
8
ve
ry
 ra
re
V
al
e,
 2
3.
6.
19
99
 (R
Š)
; V
oj
šč
ic
a,
 1
8.
8.
20
02
 (R
Š)
Pi
er
is
 n
ap
i(
Li
nn
ae
us
, 1
75
8)
x
x
x
x
x
3,
4,
5,
6,
7,
8,
9,
10
ve
ry
 c
om
m
on
Po
nt
ia
 e
du
sa
(F
ab
ric
iu
s, 
17
77
)
x
x
x
4,
5,
6,
8,
9
no
t c
om
m
on
C
ol
ia
s c
ro
ce
us
(G
eo
ff
ro
y 
&
 F
ou
rc
ro
y,
 1
78
5)
x
x
x
x
x
4,
5,
6,
7,
8,
9,
10
,
11
co
m
m
on
C
ol
ia
s a
lfa
ca
ri
en
si
sR
ib
be
, 1
90
5
x
x
x
x
x
4,
5,
6,
7,
8,
9,
10
co
m
m
on
G
on
ep
te
ry
x 
rh
am
ni
(L
in
na
eu
s, 
17
58
)
x
x
x
x
x
1 
- 1
2
co
m
m
on
Acta entomologica slovenica, 24 (2), 2016
108
LY
C
A
EN
ID
A
E 
(3
0)
H
am
ea
ri
s l
uc
in
a
(L
in
na
eu
s, 
17
58
)
x
x
x
x
x
4,
5,
7,
8
co
m
m
on
K
op
riv
a,
 2
9.
5.
20
11
 (R
Š)
Ly
ca
en
a 
ph
la
ea
s(
Li
nn
ae
us
, 1
76
1)
x
x
x
x
x
3,
4,
5,
6,
7,
8,
9,
1
0,
11
co
m
m
on
K
op
riv
a 
an
d 
V
rh
ov
lje
, 1
7.
8.
20
13
 (R
Š)
Ly
ca
en
a 
tit
yr
us
(P
od
a,
 1
76
1)
x
x
x
x
x
4,
5,
6,
7,
8
no
t c
om
m
on
Ly
ca
en
a 
al
ci
ph
ro
n
(R
ot
te
m
bu
rg
, 1
77
5)
x
5
ve
ry
 ra
re
Lo
ke
v,
 2
6.
5.
20
12
 (M
Z)
Ly
ca
en
a 
hi
pp
ot
ho
e
(L
in
na
eu
s, 
17
61
)
x
6,
7
ve
ry
 ra
re
O
pa
tje
 se
lo
, 1
9.
7.
19
89
 (R
Š)
; K
os
ta
nj
ev
ic
a 
na
K
ra
su
, 3
0.
6.
19
90
 (R
Š)
Fa
vo
ni
us
 q
ue
rc
us
(L
in
na
eu
s, 
17
58
)
x
x
6,
7,
8,
9
ra
re
C
al
lo
ph
ry
s r
ub
i(
Li
nn
ae
us
, 1
75
8)
x
x
x
x
x
3,
4,
5,
6
co
m
m
on
Sa
ty
ri
um
 sp
in
i(
D
en
is
 &
 S
ch
iff
er
m
ül
le
r, 
17
75
)
x
x
x
x
x
6,
7
co
m
m
on
Sa
ty
ri
um
 il
ic
is
(E
sp
er
, 1
77
9)
x
x
x
x
x
6,
7
ve
ry
 c
om
m
on
C
ac
yr
eu
s m
ar
sh
al
li
B
ut
le
r, 
18
98
x
9,
10
no
t c
om
m
on
O
pa
tje
 se
lo
, 2
1.
9.
20
11
 (R
Š)
; N
ov
a 
va
s, 
14
.1
0.
20
12
(R
Š)
C
up
id
o 
m
in
im
us
(F
ue
ss
ly
, 1
77
5)
x
x
x
x
x
4,
5,
6,
7
co
m
m
on
K
op
riv
a,
 2
9.
5.
20
11
 (R
Š)
C
up
id
o 
ar
gi
ad
es
(P
al
la
s, 
17
71
)
x
x
x
4,
5,
6,
7,
8,
9
no
t c
om
m
on
C
up
id
o 
al
ce
ta
s(
H
of
fm
an
ns
eg
g,
 1
80
4)
x
x
x
x
4,
5,
6,
7,
8,
9
no
t c
om
m
on
C
el
as
tr
in
a 
ar
gi
ol
us
(L
in
na
eu
s, 
17
58
)
x
x
x
x
x
4,
5,
6,
7,
8,
9
co
m
m
on
K
op
riv
a,
 2
9.
5.
20
11
 (R
Š)
; S
ko
po
, 1
7.
8.
20
13
 (R
Š)
Ps
eu
do
ph
ilo
te
s v
ic
ra
m
a
(M
oo
re
, 1
86
5)
x
x
x
3,
4,
5,
6,
7,
8
ra
re
G
or
en
ja
 B
re
st
ov
ic
a,
 1
6.
4.
19
95
 (R
Š)
; O
pa
tje
 se
lo
,
19
.4
.1
99
5 
(R
Š)
; L
ip
a 
na
 K
ra
su
, 2
6.
4.
20
00
 (R
Š)
Sc
ol
ita
nt
id
es
 o
ri
on
(P
al
la
s, 
17
71
)
x
x
x
x
4,
5,
6,
7
no
t c
om
m
on
G
la
uc
op
sy
ch
e 
al
ex
is
(P
od
a,
 1
76
1)
x
x
x
4,
5
ra
re
Pl
eb
ei
us
 a
rg
us
(L
in
na
eu
s, 
17
58
)
x
x
x
x
x
5,
6,
7,
8
ve
ry
 c
om
m
on
Pl
eb
ei
us
 id
as
(L
in
na
eu
s, 
17
61
)
x
x
x
x
x
5,
6,
7,
8,
9
co
m
m
on
Radovan Øtanta, Matjaæ Zadrgal: Contribution to the knowledge of Lepidoptera fauna of the Karst
109
Pl
eb
ei
us
 a
rg
yr
og
no
m
on
(B
er
gs
trä
ss
er
, 1
77
9)
x
x
x
5,
6,
7,
9
no
t c
om
m
on
Ar
ic
ia
 a
ge
st
is
(D
en
is
 &
 S
ch
iff
er
m
ül
le
r, 
17
75
)
x
x
x
x
x
4,
5,
6,
7,
8,
9,
10
co
m
m
on
Po
ly
om
m
at
us
 se
m
ia
rg
us
(R
ot
te
m
bu
rg
, 1
77
5)
x
x
x
x
x
5,
6,
7
ve
ry
 c
om
m
on
Po
ly
om
m
at
us
 e
sc
he
ri
(H
üb
ne
r, 
18
23
)
x
6
ve
ry
 ra
re
D
at
a 
in
 th
e 
C
ha
pt
er
 re
ga
rd
in
g 
in
te
re
st
in
g 
sp
ec
ie
s
Po
ly
om
m
at
us
 d
or
yl
as
(D
en
is
 &
 S
ch
iff
er
m
ül
le
r, 
17
75
)
x
x
4,
5,
6,
7
ra
re
K
os
ta
nj
ev
ic
a 
na
 K
ra
su
, 2
3.
4.
19
88
 (R
Š)
;
K
os
ta
nj
ev
ic
a 
na
 K
ra
su
, 1
8.
6.
19
93
 (R
Š)
; L
ip
a 
na
K
ra
su
, 1
4.
5.
20
11
 (R
Š)
Po
ly
om
m
at
us
 a
m
an
du
s(
Sc
hn
ei
de
r, 
17
92
)
x
5,
6
no
t c
om
m
on
K
rv
av
i P
ot
ok
, 3
0.
6.
19
92
 (R
Š)
; V
rh
po
lje
 - 
Sv
.
To
m
až
, 1
6.
6.
20
12
 (R
Š)
; L
ok
ev
, 9
.6
.2
01
3 
(M
Z)
 
Po
ly
om
m
at
us
 th
er
si
te
s(
C
an
te
ne
r, 
18
35
)
x
x
5,
6
ve
ry
 ra
re
Lo
ke
v,
 2
6.
5.
20
12
 (M
Z)
; K
la
rič
i, 
11
.6
.2
01
0 
(M
Z)
Po
ly
om
m
at
us
 ic
ar
us
(R
ot
te
m
bu
rg
, 1
77
5)
x
x
x
x
x
5,
6,
7,
8,
9
ve
ry
 c
om
m
on
Po
ly
om
m
at
us
 d
ap
hn
is
(D
en
is
 &
 S
ch
iff
er
m
ül
le
r, 
17
75
)
x
x
6,
7,
8
ra
re
Po
ly
om
m
at
us
 b
el
la
rg
us
(R
ot
te
m
bu
rg
, 1
77
5)
x
x
x
x
x
5,
6,
7,
8,
9,
10
ve
ry
 c
om
m
on
Po
ly
om
m
at
us
 c
or
id
on
(P
od
a,
 1
76
1)
x
x
x
x
x
7,
8
ve
ry
 c
om
m
on
Pl
is
ko
vi
ca
, K
op
riv
a,
 S
ko
po
, D
ut
ov
lje
, V
rh
ov
lje
,
Se
ža
na
 - 
D
ol
 L
es
ko
ve
c 
al
l 1
7.
8.
20
13
 (R
Š)
N
Y
M
PH
A
LI
D
A
E 
(4
9)
Li
by
th
ea
 c
el
tis
(L
ai
ch
ar
tin
g,
 1
78
2)
x
x
x
2,
3,
4,
6,
7
co
m
m
on
Pa
ra
rg
e 
ae
ge
ri
a
(L
in
na
eu
s, 
17
58
)
x
x
x
x
x
2,
3,
4,
5,
6,
7,
8,
9,
10
,1
1
co
m
m
on
La
si
om
m
at
a 
m
eg
er
a
(L
in
na
eu
s, 
17
58
)
x
x
x
x
x
3,
4,
5,
6,
7,
8,
9,
10
co
m
m
on
La
si
om
m
at
a 
m
ae
ra
(L
in
na
eu
s, 
17
58
)
x
x
x
x
x
4,
5,
6,
7,
8
co
m
m
on
Lo
pi
ng
a 
ac
hi
ne
(S
co
po
li,
 1
76
3)
x
x
6
ve
ry
 ra
re
Se
ža
na
 - 
D
ol
 L
es
ko
ve
c,
 1
4.
6.
19
86
 (R
Š)
; T
rs
te
lj,
18
.6
.1
99
3 
(R
Š)
C
oe
no
ny
m
ph
a 
oe
di
pp
us
(F
ab
ric
iu
s, 
17
87
)
x
x
x
6,
7
no
t c
om
m
on
O
pa
tje
 se
lo
, 8
.7
.1
98
9 
(R
Š)
; L
ok
vi
ca
, 2
0.
6.
19
98
(R
Š)
; B
rje
 p
ri 
K
om
nu
, 1
2.
6.
19
99
 (M
Z)
; P
lis
ko
vi
ca
,
23
.6
.1
99
9 
(R
Š)
; V
al
e,
 2
3.
6.
19
99
 (R
Š)
; L
ip
a 
na
K
ra
su
, 1
8.
6.
20
01
 (R
Š)
Acta entomologica slovenica, 24 (2), 2016
110
C
oe
no
ny
m
ph
a 
ar
ca
ni
a
(L
in
na
eu
s, 
17
61
)
x
x
x
x
x
5,
6,
7
ve
ry
 c
om
m
on
C
oe
no
ny
m
ph
a 
gl
yc
er
io
n
(B
or
kh
au
se
n,
 1
78
8)
x
6,
7
co
m
m
on
Lo
ke
v,
 8
.7
.2
00
2 
(R
Š)
; J
irm
an
ec
,  
17
.6
.2
01
1 
(R
Š)
C
oe
no
ny
m
ph
a 
pa
m
ph
ilu
s(
Li
nn
ae
us
, 1
75
8)
x
x
x
x
x
4,
5,
6,
7,
8,
9,
10
ve
ry
 c
om
m
on
Er
eb
ia
 a
et
hi
op
s(
Es
pe
r, 
17
77
)
x
x
x
7,
8
no
t c
om
m
on
K
ob
je
gl
av
a,
 5
.8
.1
98
8 
(R
Š)
; O
pa
tje
 se
lo
, 1
1.
8.
20
06
(R
Š)
; J
irm
an
ec
, 8
.7
.2
01
1 
(R
Š)
Er
eb
ia
 m
ed
us
a
(D
en
is
 &
 S
ch
iff
er
m
ül
le
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17
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 (R
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17
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(R
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(L
in
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eu
s, 
17
58
)
x
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 c
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17
58
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 (R
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14
 (R
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17
75
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 (R
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 &
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17
75
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 (R
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in
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17
64
)
x
8
EX
?
M
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it 
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17
58
)
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17
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17
58
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17
58
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17
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on
Radovan Øtanta, Matjaæ Zadrgal: Contribution to the knowledge of Lepidoptera fauna of the Karst
111
Ar
gy
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(L
in
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17
58
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17
58
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17
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17
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Šk
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 2
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 (R
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17
67
)
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17
64
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17
63
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in
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17
58
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17
58
)
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17
58
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17
58
)
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17
58
)
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17
58
)
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17
58
)
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17
58
)
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Acta entomologica slovenica, 24 (2), 2016
112
M
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17
75
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Š)
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R
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in
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17
58
)
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17
66
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17
75
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94
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17
75
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17
75
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17
75
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Radovan Øtanta, Matjaæ Zadrgal: Contribution to the knowledge of Lepidoptera fauna of the Karst
113
LA
SI
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17
58
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10
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58
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(L
in
na
eu
s, 
17
58
)
x
x
6,
7
ra
re
Li
pa
 n
a 
K
ra
su
, 1
0.
7.
19
98
 (R
Š)
; P
on
ik
ve
, 2
9.
6.
20
13
(R
Š&
M
Z&
C
M
)
M
al
ac
os
om
a 
ca
st
re
ns
is
(L
in
na
eu
s, 
17
58
)
x
x
x
x
x
6,
7
no
t c
om
m
on
La
si
oc
am
pa
 tr
ifo
lii
(D
en
is
 &
 S
ch
iff
er
m
ül
le
r, 
17
75
)
x
x
x
x
x
8,
9,
10
ve
ry
 c
om
m
on
La
si
oc
am
pa
 q
ue
rc
us
(L
in
na
eu
s, 
17
58
)
x
x
x
x
7,
8
ra
re
M
ac
ro
th
yl
ac
ia
 ru
bi
(L
in
na
eu
s, 
17
58
)
x
x
x
x
4,
5
no
t c
om
m
on
D
en
dr
ol
im
us
 p
in
i(
Li
nn
ae
us
, 1
75
8)
x
x
x
x
x
5,
6,
7,
8,
9
no
t c
om
m
on
Ji
rm
an
ec
, 5
.9
.2
01
3 
(R
Š&
M
Z)
 - 
th
e 
la
te
st
 d
at
e
Ph
yl
lo
de
sm
a 
tr
em
ul
ifo
lia
(H
üb
ne
r, 
18
10
)
x
x
x
x
x
3,
4,
5,
6
co
m
m
on
G
as
tr
op
ac
ha
 q
ue
rc
ifo
lia
(L
in
na
eu
s, 
17
58
)
x
x
x
6,
7,
8,
9
ra
re
K
os
ta
nj
ev
ic
a 
na
 K
ra
su
, 1
9.
7.
19
88
 (R
Š&
M
Z)
;
O
pa
tje
 se
lo
, 1
7.
7.
19
93
 (M
Z)
; T
rs
te
lj,
 1
6.
7.
20
10
(R
Š)
; K
op
riv
a,
 1
8.
7.
20
12
 (R
Š)
O
do
ne
st
is
 p
ru
ni
(L
in
na
eu
s, 
17
58
)
x
x
x
x
5,
6,
7,
8,
9
no
t c
om
m
on
O
pa
tje
 se
lo
, 1
.7
.1
98
9 
(R
Š&
M
Z)
; T
rs
te
lj,
 1
4.
6.
19
91
(R
Š&
M
Z)
; J
irm
an
ec
, 5
.9
.2
01
3 
(R
Š&
M
Z)
; K
op
riv
a,
6.
7.
20
14
 (R
Š&
C
M
)
B
R
A
H
M
A
EI
D
A
E 
(1
)
Le
m
on
ia
 ta
ra
xa
ci
(D
en
is
 &
 S
ch
iff
er
m
ül
le
r, 
17
75
)
x
x
x
x
9,
10
no
t c
om
m
on
K
os
ta
nj
ev
ic
a 
na
 K
ra
su
, 2
4.
9.
19
93
 (R
Š)
; V
oj
šč
ic
a,
11
.1
0.
19
96
 (R
Š)
; K
az
lje
, 1
7.
9.
20
11
 (R
Š&
M
Z)
;
K
op
riv
a,
 1
8.
9.
20
14
 (R
Š&
M
Z&
C
M
)
Acta entomologica slovenica, 24 (2), 2016
114
EN
D
R
O
M
ID
A
E 
(1
)
En
dr
om
is
 v
er
si
co
lo
ra
(L
in
na
eu
s, 
17
58
)
x
x
x
x
x
3,
4
no
t c
om
m
on
SA
TU
R
N
II
D
A
E 
(5
)
Ag
lia
 ta
u
(L
in
na
eu
s, 
17
58
)
x
x
x
x
4,
5
ra
re
K
ob
je
gl
av
a,
 1
9.
5.
19
87
 (R
Š&
M
Z)
; B
rje
 p
ri 
K
om
nu
,
30
.4
.2
00
3 
(R
Š&
M
Z)
; P
ov
ir,
 2
8.
4.
20
12
 (R
Š&
M
Z)
;
Ji
rm
an
ec
, 8
.5
.2
01
5 
(R
Š&
M
Z&
B
Z&
SG
)
Sa
tu
rn
ia
 p
yr
i(
D
en
is
 &
 S
ch
iff
er
m
ül
le
r, 
17
75
)
x
x
x
x
x
4,
5,
6
no
t c
om
m
on
Sa
tu
rn
ia
 p
av
on
ie
lla
(S
co
po
li,
 1
76
3)
x
x
x
x
x
3,
4
no
t c
om
m
on
Sa
tu
rn
ia
 c
ae
ci
ge
na
K
up
id
o,
 1
82
5
x
x
x
x
x
9,
10
no
t c
om
m
on
K
az
lje
, 1
7.
9.
20
11
 (R
Š&
M
Z)
 - 
th
e 
ea
rli
es
t d
at
e,
la
rv
ae
 o
n 
Q
ue
rc
us
 p
et
ra
ea
An
th
er
ae
a 
ya
m
am
ai
(G
ué
rin
-M
én
ev
ill
e,
 1
86
1)
x
x
x
x
x
7,
8
no
t c
om
m
on
SP
H
IN
G
ID
A
E 
(1
4)
M
ar
um
ba
 q
ue
rc
us
(D
en
is
 &
 S
ch
iff
er
m
ül
le
r, 
17
75
)
x
x
x
x
x
6,
7,
8
co
m
m
on
N
ov
el
o,
 9
.7
.2
00
2 
(R
Š&
M
Z)
; G
or
ja
ns
ko
 -
Po
dk
la
ne
c,
 3
.7
.2
00
9 
(R
Š&
M
Z)
; J
irm
an
ec
, 4
.6
.2
01
1
(R
Š&
M
Z)
; K
op
riv
a,
 2
7.
6.
20
12
 (R
Š)
M
im
as
 ti
lia
e
(L
in
na
eu
s, 
17
58
)
x
x
x
x
4,
5,
6,
7,
8
no
t c
om
m
on
Sm
er
in
th
us
 o
ce
lla
ta
(L
in
na
eu
s, 
17
58
)
x
5
ve
ry
 ra
re
O
pa
tje
 se
lo
, 6
.6
.1
98
7 
(R
Š&
M
Z)
Ag
ri
us
 c
on
vo
lv
ul
i(
Li
nn
ae
us
, 1
75
8)
x
x
x
x
8,
9,
10
no
t c
om
m
on
Ac
he
ro
nt
ia
 a
tr
op
os
(L
in
na
eu
s, 
17
58
)
x
x
8,
9,
10
ve
ry
 ra
re
V
oj
šč
ic
a,
 9
.1
0.
19
95
 (R
Š)
; M
al
i D
ol
, 2
6.
8.
20
14
 e
. l
.
22
.7
.2
01
4 
(M
Z)
Sp
hi
nx
 li
gu
st
ri
Li
nn
ae
us
, 1
75
8
x
x
7,
8
ve
ry
 ra
re
O
pa
tje
 se
lo
, 2
6.
7.
19
89
 (R
Š&
M
Z)
; L
ip
a 
na
 K
ra
su
,
15
.8
.1
99
4 
(R
Š&
M
Z)
Sp
hi
nx
 p
in
as
tr
iL
in
na
eu
s, 
17
58
x
x
x
x
x
5,
6,
7
co
m
m
on
H
em
ar
is
 ti
ty
us
(L
in
na
eu
s, 
17
58
)
x
x
5
ve
ry
 ra
re
Lo
ke
v,
 2
6.
5.
20
12
 (M
Z)
; P
od
br
ež
e,
 6
.5
.2
01
5
(R
Š&
M
Z)
Radovan Øtanta, Matjaæ Zadrgal: Contribution to the knowledge of Lepidoptera fauna of the Karst
115
H
em
ar
is
 fu
ci
fo
rm
is
(L
in
na
eu
s, 
17
58
)
x
x
x
5
ve
ry
 ra
re
K
os
ta
nj
ev
ic
a 
na
 K
ra
su
, 1
9.
5.
19
91
 (R
Š)
; L
ip
a 
na
K
ra
su
, 1
4.
5.
20
11
 (R
Š)
; K
ra
jn
a 
va
s, 
21
.5
.2
01
4 
(R
Š)
 
M
ac
ro
gl
os
su
m
 st
el
la
ta
ru
m
(L
in
na
eu
s, 
17
58
)
x
x
x
x
3,
4,
5,
6,
7,
8,
9,
10
,1
1
no
t c
om
m
on
H
yl
es
 e
up
ho
rb
ia
e
(L
in
na
eu
s, 
17
58
)
x
x
6
ra
re
Tr
st
el
j, 
14
.6
.1
99
1 
(R
Š&
M
Z)
; L
ok
vi
ca
, 2
6.
8.
20
08
(R
Š)
 la
rv
ae
; M
ire
n 
- G
m
aj
na
, 7
.9
.2
00
8 
(R
Š)
 la
rv
ae
 
H
yl
es
 li
vo
rn
ic
a
(E
sp
er
, 1
78
0)
x
x
5
ve
ry
 ra
re
O
pa
tje
 se
lo
, 6
.5
.1
98
8 
(R
Š&
M
Z)
;  
Ji
rm
an
ec
,
8.
5.
20
15
 (R
Š&
M
Z&
B
Z&
SG
)
D
ei
le
ph
ila
 e
lp
en
or
(L
in
na
eu
s, 
17
58
)
x
x
x
5,
6
ve
ry
 ra
re
O
pa
tje
 se
lo
, 1
0.
6.
19
89
 (R
Š&
M
Z)
; B
rje
 p
ri 
K
om
nu
,
22
.5
.1
99
3 
(R
Š&
M
Z)
; V
oj
šč
ic
a,
 1
2.
7.
19
95
(R
Š&
M
Z)
; L
ok
ev
, 2
5.
6.
20
10
 (R
Š&
M
Z)
D
ei
le
ph
ila
 p
or
ce
llu
s(
Li
nn
ae
us
, 1
75
8)
x
x
x
x
x
4,
5,
6,
7,
8
co
m
m
on
G
EO
M
ET
R
ID
A
E 
(2
12
)
Ab
ra
xa
s g
ro
ss
ul
ar
ia
ta
(L
in
na
eu
s, 
17
58
)
x
6,
7
ve
ry
 ra
re
V
oj
šč
ic
a,
 1
.7
.1
98
7 
(M
Z)
Li
gd
ia
 a
du
st
at
a
(D
en
is
 &
 S
ch
iff
er
m
ül
le
r, 
17
75
)
x
x
x
x
x
3,
4,
5,
6,
7,
8,
9
co
m
m
on
St
eg
an
ia
 tr
im
ac
ul
at
a
(d
e 
V
ill
er
s, 
17
89
)
x
x
x
5,
6,
7
ve
ry
 ra
re
V
rh
po
lje
 - 
Sv
. T
om
až
, 1
6.
6.
20
12
 (R
Š)
; J
irm
an
ec
,
14
.6
.2
01
3 
(R
Š&
M
Z)
;  
K
om
en
, 1
7.
7.
20
15
 (R
Š)
H
el
io
m
at
a 
gl
ar
ea
ri
a
(D
en
is
 &
 S
ch
iff
er
m
ül
le
r, 
17
75
)
x
x
x
x
x
4,
5,
6,
7,
8
co
m
m
on
M
ac
ar
ia
 a
lte
rn
at
a
(D
en
is
 &
 S
ch
iff
er
m
ül
le
r, 
17
75
)
x
x
x
x
x
4,
5,
6,
7,
8,
9
co
m
m
on
M
ac
ar
ia
 li
tu
ra
ta
(C
le
rc
k,
 1
75
9)
x
x
x
x
x
4,
5,
6,
7,
8,
9
no
t c
om
m
on
C
hi
as
m
ia
 c
la
th
ra
ta
(L
in
na
eu
s, 
17
58
)
x
x
x
x
x
4,
5,
6,
8,
9
ve
ry
 c
om
m
on
Is
tu
rg
ia
 a
re
na
ce
ar
ia
(D
en
is
 &
 S
ch
iff
er
m
ül
le
r, 
17
75
)
x
7,
8
ve
ry
 ra
re
K
os
ta
nj
ev
ic
a 
na
 K
ra
su
, 2
5.
7.
19
87
 (R
Š&
M
Z)
;
O
pa
tje
 se
lo
, 1
3.
8.
19
88
 (R
Š&
M
Z)
; G
or
en
ja
B
re
st
ov
ic
a,
 2
0.
7.
19
91
 (R
Š&
M
Z)
G
no
ph
ar
m
ia
 st
ev
en
ar
ia
(B
oi
sd
uv
al
, 1
84
0)
x
5
ve
ry
 ra
re
B
rje
 p
ri 
K
om
nu
, 2
.5
.1
99
9 
 (R
Š&
M
Z)
Rh
op
tr
ia
 a
sp
er
ar
ia
(H
üb
ne
r, 
18
17
)
x
9
ve
ry
 ra
re
D
at
a 
in
 th
e 
C
ha
pt
er
 re
ga
rd
in
g 
in
te
re
st
in
g 
sp
ec
ie
s
Pe
tr
op
ho
ra
 c
hl
or
os
at
a
(S
co
po
li,
 1
76
3)
x
5
ve
ry
 ra
re
K
ob
je
gl
av
a 
- J
el
en
ca
, 1
6.
5.
19
96
 (R
Š&
M
L)
Acta entomologica slovenica, 24 (2), 2016
116
Pl
ag
od
is
 p
ul
ve
ra
ri
a
(L
in
na
eu
s, 
17
58
)
x
x
x
x
3,
4,
7,
8
no
t c
om
m
on
Po
ni
kv
e,
 3
0.
3.
20
14
 (R
Š&
M
Z)
Pl
ag
od
is
 d
ol
ab
ra
ri
a
(L
in
na
eu
s, 
17
67
)
x
x
6,
7,
8,
9
ra
re
K
os
ta
nj
ev
ic
a 
na
 K
ra
su
, 9
.7
.2
00
2 
(R
Š&
M
Z)
;
Te
m
ni
ca
, 1
1.
8.
20
08
 (R
Š)
; T
rs
te
lj,
 2
.9
.2
00
9
(R
Š&
M
Z)
; T
rs
te
lj,
 1
6.
7.
20
10
 (R
Š)
O
pi
st
ho
gr
ap
tis
 lu
te
ol
at
a
(L
in
na
eu
s, 
17
58
)
x
x
x
x
x
4,
5,
6,
7,
8,
9
co
m
m
on
Ps
eu
do
pa
nt
he
ra
 m
ac
ul
ar
ia
(L
in
na
eu
s, 
17
58
)
x
x
x
x
x
4,
5
ve
ry
 c
om
m
on
Ap
ei
ra
 sy
ri
ng
ar
ia
(L
in
na
eu
s, 
17
58
)
x
x
x
6,
7,
9
ve
ry
 ra
re
O
pa
tje
 se
lo
, 2
1.
9.
19
90
 (R
Š&
M
Z)
; K
om
en
 -
Ja
žm
er
ca
, 5
.6
.2
01
0 
(R
Š&
M
Z&
B
Z)
; K
op
riv
a,
18
.9
.2
01
4 
(R
Š&
M
Z&
C
M
)
En
no
m
os
 a
ut
um
na
ri
a
(W
er
ne
bu
rg
, 1
85
9)
x
x
9
ve
ry
 ra
re
K
ob
je
gl
av
a,
 7
.9
.1
99
5 
(R
Š&
M
Z)
; J
irm
an
ec
,
5.
9.
20
13
 (R
Š&
M
Z)
En
no
m
os
 q
ue
rc
in
ar
ia
(H
uf
na
ge
l, 
17
67
)
x
6
ve
ry
 ra
re
Lo
ke
v,
 2
5.
6.
20
10
 (R
Š&
M
Z&
B
Z)
En
no
m
os
 e
ro
sa
ri
a
(D
en
is
 &
 S
ch
iff
er
m
ül
le
r, 
17
75
)
x
x
9,
10
ra
re
K
az
lje
, 1
7.
9.
20
11
 (R
Š&
M
Z)
; L
ok
ev
, 3
0.
9.
20
11
(R
Š&
M
Z)
En
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os
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ia
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üb
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13
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 2
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13
 (R
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C
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)
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ia
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ab
ric
iu
s, 
17
75
)
x
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ra
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 se
lo
, 1
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89
 (R
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or
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üb
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ia
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uf
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ge
l, 
17
67
)
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 2
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11
 (R
Š)
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 1
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5
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 1
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13
 (R
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or
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 1
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ia
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in
na
eu
s, 
17
58
)
x
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co
m
m
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pa
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a 
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ra
su
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.2
00
9 
(R
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 - 
th
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rli
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t d
at
e
Radovan Øtanta, Matjaæ Zadrgal: Contribution to the knowledge of Lepidoptera fauna of the Karst
117
O
ur
ap
te
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in
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17
58
)
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6
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oj
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 1
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 (R
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 2
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 (R
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in
na
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s, 
17
61
)
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10
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17
58
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ia
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in
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17
58
)
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17
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x
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no
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om
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K
os
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 K
ra
su
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.1
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5 
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Š&
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G
or
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6.
2.
19
97
 (R
Š&
M
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la
rič
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.1
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99
8 
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Š)
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ri
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aj
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en
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 &
 S
ch
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m
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le
r, 
17
75
)
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10
,1
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üb
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.2
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4 
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Š&
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ab
ric
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76
)
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7.
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95
 (M
Z)
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en
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hu
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 1
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2)
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3,
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no
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5,
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no
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pa
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 se
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1 
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Š&
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av
a,
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9.
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95
 (R
Š&
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ag
ra
je
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 2
2.
5.
20
09
(R
Š&
M
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op
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 7
.9
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3 
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Š)
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 v
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lii
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oh
at
sc
h,
 1
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3)
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x
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7
no
t c
om
m
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K
op
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 2
6.
6.
20
11
 (R
Š)
; K
op
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 2
7.
6.
20
12
 (R
Š)
;
Po
ni
kv
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 2
9.
6.
20
13
 (R
Š&
M
Z&
C
M
); 
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13
 (M
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od
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13
 (R
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Acta entomologica slovenica, 24 (2), 2016
118
Pe
rib
at
od
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om
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ar
ia
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at
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75
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no
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ci
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ta
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in
na
eu
s, 
17
58
)
x
x
x
5,
6
ra
re
O
pa
tje
 se
lo
, 1
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99
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ko
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0.
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11
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75
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no
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 K
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5.
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94
 (R
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 3
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20
11
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Š)
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 7
.9
.2
01
3 
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Š)
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4 
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4,
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 1
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99
6 
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Š)
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gi
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uf
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ge
l, 
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67
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4,
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9
no
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Po
vi
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2 
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3
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5.
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14
 (R
Š&
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ch
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r, 
17
75
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8
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75
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7
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8.
3.
20
12
 (R
Š&
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Z)
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at
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ar
ia
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in
na
eu
s, 
17
58
)
x
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x
x
4,
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6,
7,
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9
ve
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om
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8
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at
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in
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17
58
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6
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om
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in
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17
58
)
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75
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Š&
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rs
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ko
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75
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on
Radovan Øtanta, Matjaæ Zadrgal: Contribution to the knowledge of Lepidoptera fauna of the Karst
119
C
am
pa
ea
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ar
ga
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ta
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a
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in
na
eu
s, 
17
61
)
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5,
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 S
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r, 
17
75
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6,
9
ve
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13
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8.
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51
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75
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75
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94
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Š)
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99
5
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Š&
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Z)
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pa
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00
7 
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Š)
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uf
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ge
l, 
17
67
)
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8,
9,
10
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eo
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 p
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ia
Li
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us
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75
8
x
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6,
7
ve
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 ra
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Te
m
ni
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5.
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19
87
 (R
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M
Z)
; K
op
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 1
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20
11
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Š)
C
om
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ae
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aj
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ia
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en
is
 &
 S
ch
iff
er
m
ül
le
r, 
17
75
)
x
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x
x
5,
6
ra
re
K
om
en
 - 
Ja
žm
er
ca
, 5
.6
.2
01
0 
(R
Š&
M
Z&
B
Z)
;
K
op
riv
a,
 2
2.
5.
20
11
 (R
Š)
; J
irm
an
ec
, 1
4.
6.
20
13
(R
Š&
M
Z)
 
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et
id
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 sm
ar
ag
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a
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ab
ric
iu
s, 
17
87
)
x
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x
x
6,
7,
8
ra
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 K
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19
87
 (R
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M
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;
O
pa
tje
 se
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3.
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19
87
 (R
Š&
M
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; T
em
ni
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,
11
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8 
(R
Š)
H
em
ith
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tiv
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ia
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üb
ne
r, 
17
89
)
x
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6
ra
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a 
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 K
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1.
6.
19
88
 (R
Š&
M
Z)
;
K
op
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 1
5.
6.
20
12
 (R
Š)
Ph
ai
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ra
m
m
a 
et
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ar
ia
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el
le
r, 
18
49
)
x
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6
ve
ry
 ra
re
V
rh
po
lje
 - 
Sv
. T
om
až
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6.
6.
20
12
 (R
Š)
; K
op
riv
a,
29
.6
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01
3 
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Š&
M
Z&
C
M
)
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al
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ia
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co
po
li,
 1
76
3)
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x
x
6,
7
ra
re
G
or
ja
ns
ko
 - 
Po
dk
la
ne
c,
 3
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.2
00
9 
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Š&
M
Z)
;
Po
ni
kv
e,
 2
9.
6.
20
13
 (R
Š&
M
Z&
C
M
)
Acta entomologica slovenica, 24 (2), 2016
120
H
em
is
to
la
 c
hr
ys
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ra
sa
ri
a
(E
sp
er
, 1
79
5)
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 2
0.
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91
 (R
Š&
M
Z)
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ko
po
,
20
.8
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01
1 
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; K
op
riv
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 6
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.2
01
4 
(R
Š&
C
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)
Em
m
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ae
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ia
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üb
ne
r, 
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oj
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 4
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10
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58
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 1
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58
)
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08
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Š)
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 K
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4.
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11
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 1
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7.
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12
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Š)
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ct
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67
)
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9
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K
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11
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99
4 
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ob
je
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av
a 
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16
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6 
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Š&
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L)
; L
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 K
ra
su
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5.
20
06
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Š&
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Z)
Radovan Øtanta, Matjaæ Zadrgal: Contribution to the knowledge of Lepidoptera fauna of the Karst
121
Sc
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ul
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vi
rg
ul
at
a
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 &
 S
ch
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le
r, 
17
75
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6,
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9
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co
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li,
 1
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3)
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x
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 &
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 (R
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x
4,
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 8
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0.
20
09
 (R
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Sc
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in
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s, 
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58
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x
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re
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99
4 
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os
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su
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19
.6
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99
5 
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Š)
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irm
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.9
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Š&
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99
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9,
10
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10
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99
5 
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Š)
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 1
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9.
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10
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 1
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11
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Š)
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in
na
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s, 
17
58
)
x
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9
ve
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re
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or
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9.
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96
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 2
9.
6.
20
15
(R
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a
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 1
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9)
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6,
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8
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om
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6)
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7
no
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ae
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31
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10
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4.
20
14
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Š&
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gl
av
a
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5.
10
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3 
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Š&
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ab
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iu
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17
75
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9
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O
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6.
7.
19
89
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Š&
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ob
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99
7 
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9.
5.
20
11
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 2
6.
6.
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11
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Š)
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9
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 1
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9.
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5)
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x
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9
ra
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pa
tje
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.5
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99
0 
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ob
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99
7 
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 2
8.
4.
20
12
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Š&
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;
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rje
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5.
4.
20
14
 (R
Š&
M
Z)
Acta entomologica slovenica, 24 (2), 2016
122
C
yc
lo
ph
or
a 
po
ra
ta
(L
in
na
eu
s, 
17
67
)
x
x
x
x
x
4,
5,
7,
8,
9
ra
re
V
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šč
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a,
 2
4.
4.
19
94
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Š&
M
Z)
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rs
te
lj,
 6
.8
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9
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 1
0.
9.
20
10
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8.
9.
20
12
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Z)
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as
te
lb
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ge
r, 
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97
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9
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8.
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09
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 1
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9.
20
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in
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eu
s, 
17
58
)
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x
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9
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Tr
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9.
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09
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0
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Š&
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3 
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4 
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47
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8
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Po
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5 
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99
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7.
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98
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9 
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13
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in
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eu
s, 
17
67
)
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x
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10
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 4
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96
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10
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11
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67
)
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8,
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no
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iv
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 1
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 (R
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11
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om
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9.
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13
 (R
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at
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at
a
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üb
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r, 
18
13
)
x
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x
x
x
4,
5,
6,
7,
8,
9
co
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Sc
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ar
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ia
(D
en
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 &
 S
ch
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m
ül
le
r, 
17
75
)
x
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x
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5,
6
ve
ry
 c
om
m
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Sc
ot
op
te
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(H
uf
na
ge
l, 
17
67
)
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5,
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ve
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 c
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55
 sp
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S.
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 &
 S
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17
75
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7
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Se
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 1
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Sv
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až
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.7
.2
01
1 
(R
Š)
Sc
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x 
m
oe
ni
at
a
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co
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li,
 1
76
3)
x
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8
ra
re
Ji
rm
an
ec
, 1
9.
8.
20
11
 (R
Š&
M
Z)
; J
irm
an
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,
28
.7
.2
01
2 
(R
Š&
M
Z)
O
rt
ho
na
m
a 
ob
st
ip
at
a
(F
ab
ric
iu
s, 
17
94
)
x
x
x
8,
9,
10
,1
1,
12
ra
re
K
ob
je
gl
av
a,
 1
5.
11
.1
99
5 
(R
Š)
; L
ok
vi
ca
, 4
.1
1.
19
96
(R
Š)
; G
or
ja
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ko
, 5
.1
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19
96
 (M
Z)
; K
op
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18
.9
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01
4 
(R
Š&
M
Z&
C
M
)
Radovan Øtanta, Matjaæ Zadrgal: Contribution to the knowledge of Lepidoptera fauna of the Karst
123
Xa
nt
ho
rh
oe
 fl
uc
tu
at
a
(L
in
na
eu
s, 
17
58
)
x
x
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x
4,
5,
6,
7,
9
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Xa
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 1
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ra
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C
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uf
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17
67
)
x
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3,
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B
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 2
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99
 (R
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 &
 S
ch
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m
ül
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r, 
17
75
)
x
6
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 ra
re
Ji
rm
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, 4
.6
.2
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1 
(R
Š&
M
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C
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ra
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m
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(L
in
na
eu
s, 
17
58
)
x
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x
x
x
5,
6,
7,
8,
9,
10
co
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(H
üb
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r, 
17
99
)
x
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6,
7
ra
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Sk
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o,
 2
6.
6.
20
11
 (R
Š)
; J
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ec
, 2
8.
7.
20
12
(R
Š&
M
Z)
; J
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, 1
4.
6.
20
13
 (R
Š&
M
Z)
Ep
ir
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at
a
(M
ül
le
r, 
17
64
)
x
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x
4,
5,
6,
7,
9
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m
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Ep
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 ri
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ta
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üb
ne
r, 
18
13
)
x
x
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7
ve
ry
 ra
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K
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18
.7
.2
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4 
(R
Š&
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; J
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8.
5.
20
15
 (R
Š&
M
Z&
B
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SG
)
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ir
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 g
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(D
en
is
 &
 S
ch
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m
ül
le
r, 
17
75
)
x
x
x
x
x
5,
6,
7,
8,
9
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m
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Eu
ph
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9)
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7,
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9
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Lo
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 4
.9
.2
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(R
Š&
M
Z&
B
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19
.8
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(R
Š&
M
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; J
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8.
7.
20
12
(R
Š&
M
Z)
Eu
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ric
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Sc
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, 1
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2)
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6,
7,
8,
9
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Te
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Š)
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5.
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11
 (R
Š)
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K
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.9
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Š)
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9.
6.
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13
(R
Š&
M
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C
M
)
Ea
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ila
 b
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(D
en
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 &
 S
ch
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m
ül
le
r, 
17
75
)
x
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4
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m
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An
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en
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 &
 S
ch
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m
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r, 
17
75
)
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4,
5
ve
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 ra
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O
pa
tje
 se
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.4
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9 
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Š&
M
Z)
; K
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Je
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5.
19
96
 (R
Š&
M
L)
; L
ip
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 K
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9.
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11
 (R
Š)
; P
on
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ve
, 3
0.
3.
20
14
 (R
Š&
M
Z)
Pe
lu
rg
a 
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m
ita
ta
(L
in
na
eu
s, 
17
58
)
x
9
ve
ry
 ra
re
K
la
rič
i, 
26
.9
.1
99
5 
(R
Š)
Pe
nn
ith
er
a 
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m
at
a
(H
üb
ne
r, 
18
22
)
x
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10
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no
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Th
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en
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 &
 S
ch
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17
75
)
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4,
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6,
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9,
10
ve
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 c
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Th
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ch
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75
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10
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 ra
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Ji
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9.
10
.2
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3 
(R
Š&
M
Z)
Th
er
a 
ju
ni
pe
ra
ta
(L
in
na
eu
s, 
17
58
)
x
x
x
x
x
10
,1
1
ve
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om
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on
Acta entomologica slovenica, 24 (2), 2016
124
C
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 fu
lv
at
a
(F
or
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, 1
77
1)
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6
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Lo
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 2
5.
6.
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10
 (R
Š&
M
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, 4
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1
(R
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rh
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Sv
. T
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6.
6.
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12
 (R
Š)
;
Ji
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, 9
.6
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3 
(R
Š&
M
Z)
C
os
m
or
ho
e 
oc
el
la
ta
(L
in
na
eu
s, 
17
58
)
x
x
x
x
x
4,
5,
6,
8,
9
no
t c
om
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C
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17
67
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17
75
)
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8,
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10
no
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1)
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5,
6
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26
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at
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17
75
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9,
10
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at
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(L
in
na
eu
s, 
17
58
)
x
x
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x
11
,1
2,
1
ve
ry
 c
om
m
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K
la
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.1
.1
99
8 
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Š)
; T
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lj,
 7
.1
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7 
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Š)
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(D
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 S
ch
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m
ül
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r, 
17
75
)
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x
10
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12
ve
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 c
om
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at
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li,
 1
76
3)
x
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x
5,
6
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m
m
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As
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(H
uf
na
ge
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17
67
)
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x
x
4,
5,
6,
7,
8,
9
no
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Eu
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 1
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6
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Lo
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 2
5.
6.
20
10
 (R
Š&
M
Z&
B
Z)
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lia
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am
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(H
uf
na
ge
l, 
17
67
)
x
6
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 ra
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Ji
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.6
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3 
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 &
 S
ch
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r, 
17
75
)
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5,
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7
co
m
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Ph
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17
67
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6,
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Li
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K
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7.
19
95
 (R
Š&
M
Z)
; S
ko
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10
.6
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1 
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Š)
; P
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br
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 1
2.
7.
20
13
 (R
Š&
M
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;
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la
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.6
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4 
(R
Š)
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 1
76
3)
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x
x
4,
5
co
m
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Lo
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- S
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.4
.2
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7 
(R
Š&
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Z&
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Z&
SG
);
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.5
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3 
(R
Š&
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.3
.2
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4 
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Š)
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K
om
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5.
4.
20
14
(R
Š&
M
Z)
Radovan Øtanta, Matjaæ Zadrgal: Contribution to the knowledge of Lepidoptera fauna of the Karst
125
Tr
ip
ho
sa
 d
ub
ita
ta
(L
in
na
eu
s, 
17
58
)
x
x
x
x
x
3,
4,
5,
6,
7,
10
no
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om
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9.
10
.2
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3 
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Š&
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Z)
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at
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Pa
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at
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(D
en
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 &
 S
ch
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m
ül
le
r, 
17
75
)
x
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x
4,
5,
6,
8,
9
no
t c
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or
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ta
lb
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en
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 S
ch
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m
ül
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r, 
17
75
)
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x
3,
4,
5,
6,
7,
8,
9
no
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 S
ch
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17
75
)
x
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x
x
5,
6,
7,
8
no
t c
om
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O
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 se
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, 6
.6
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98
7 
(R
Š&
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Z)
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ut
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,
9.
8.
19
94
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Š)
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irm
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8.
7.
20
12
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Š&
M
Z)
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l
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ge
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l s
tru
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Š)
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La
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ar
pe
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5)
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x
x
5,
6,
7,
8,
9
no
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om
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.8
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4 
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Š)
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 1
0.
9.
20
10
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Š&
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Z&
JD
); 
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6.
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13
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Š&
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Z)
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Š)
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or
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e 
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lli
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ap
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ta
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er
ric
h-
Sc
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er
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83
8)
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6,
9
ra
re
Lo
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 2
5.
6.
20
10
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Š&
M
Z&
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6.
20
11
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Š&
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Z)
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rh
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Sv
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om
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2 
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Š)
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3 
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Š&
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67
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99
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5.
19
97
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SG
);
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8.
20
09
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Š)
; J
irm
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.5
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01
5
(R
Š&
M
Z&
B
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SG
)
M
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e 
pa
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ta
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et
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3)
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10
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 ra
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 3
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20
11
 (R
Š&
M
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ir,
 1
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10
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01
3
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Acta entomologica slovenica, 24 (2), 2016
126
Pe
ri
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m
a 
hy
dr
at
a
(T
re
its
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ke
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82
9)
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6
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9.
5.
20
11
 (R
Š&
M
Z)
; K
op
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a,
 3
0.
5.
20
11
(R
Š)
; K
ob
je
gl
av
a 
- J
el
en
ca
, 9
.5
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01
2 
(R
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M
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;
Ji
rm
an
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4.
6.
20
13
 (R
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M
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Pe
ri
zo
m
a 
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bu
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en
is
 &
 S
ch
iff
er
m
ül
le
r, 
17
75
)
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8
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 (M
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8.
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11
(R
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6,
7,
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V
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4
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17
58
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x
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5.
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88
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6.
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11
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Eu
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6
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 1
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 (R
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20
13
 (R
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M
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Eu
pi
th
ec
ia
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nu
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ta
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ne
r, 
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)
x
x
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7
ve
ry
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Ji
rm
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7.
20
12
 (R
Š&
M
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 - 
in
sp
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tio
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ni
ta
l s
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 (R
Š)
; G
or
ja
ns
ko
 - 
Po
dk
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ne
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7.
20
09
 (R
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M
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; G
or
ja
ns
ko
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9.
4.
20
04
 (M
Z)
Eu
pi
th
ec
ia
 in
tu
rb
at
a
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üb
ne
r, 
18
17
)
x
7
ve
ry
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Ji
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8.
7.
20
12
 (R
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M
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 - 
in
sp
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tio
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ta
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 (R
Š)
Eu
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th
ec
ia
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et
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ia
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oe
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78
1)
x
6
ra
re
Ji
rm
an
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.6
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01
1 
(R
Š&
M
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Eu
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th
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ia
 li
na
ri
at
a
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en
is
 &
 S
ch
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m
ül
le
r, 
17
75
)
x
8
ve
ry
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Li
pa
 n
a 
K
ra
su
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.8
.2
00
9 
(R
Š)
Eu
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th
ec
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lu
m
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at
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80
9)
x
6
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Sk
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 1
0.
6.
20
11
 (R
Š)
Eu
pi
th
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ia
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en
os
at
a
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ab
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s, 
17
87
)
x
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5
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 1
9.
5.
19
87
 (R
Š&
M
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Eu
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th
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hi
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er
er
iB
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at
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 1
89
3
x
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Eu
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8
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20
12
 (R
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11
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01
2 
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M
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; K
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 1
5.
5.
20
13
(R
Š&
M
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Eu
pi
th
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ia
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bb
re
vi
at
a
St
ep
he
ns
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1
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x
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3,
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5
ra
re
K
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19
.3
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99
5 
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Š&
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rv
av
i P
ot
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18
.4
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01
3 
(R
Š&
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Radovan Øtanta, Matjaæ Zadrgal: Contribution to the knowledge of Lepidoptera fauna of the Karst
127
Eu
pi
th
ec
ia
 d
od
on
ea
ta
G
ue
né
e,
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85
8
x
x
x
x
x
4,
5
ve
ry
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om
m
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Eu
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th
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ia
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us
ill
at
a
(D
en
is
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 S
ch
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m
ül
le
r, 
17
75
)
x
x
x
8,
9
ve
ry
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om
m
on
Eu
pi
th
ec
ia
 e
ri
ce
at
a
(R
am
bu
r, 
18
33
)
x
x
x
x
x
8,
9,
10
ve
ry
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om
m
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Eu
pi
th
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nt
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B
oi
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84
0
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5
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20
04
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B
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1.
5.
20
13
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Š&
M
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Eu
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th
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lin
at
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H
er
ric
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Sc
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ff
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86
1
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6
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Ja
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Š&
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ge
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Sc
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ff
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84
8
x
4
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5.
4.
20
14
 (R
Š&
M
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Eu
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th
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ch
ri
da
ta
Sc
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tz
e 
&
 P
in
ke
r, 
19
68
x
4
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9 
e.
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Eu
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ta
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86
1
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x
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9
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Li
pa
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9.
19
95
 (R
Š&
M
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; K
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01
1 
(R
Š)
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a
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8.
3.
20
12
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8.
4.
20
12
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Š&
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3 
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5.
4.
20
14
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Š&
M
Z)
Eu
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th
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di
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ta
(H
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r, 
18
13
)
x
4
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Po
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2 
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3
(R
Š&
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Eu
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ff
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84
8
x
5
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99
9 
(R
Š&
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 - 
in
sp
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ni
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Š)
Eu
pi
th
ec
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en
ta
ur
ea
ta
(D
en
is
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 S
ch
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m
ül
le
r, 
17
75
)
x
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x
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9
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 2
7.
9.
19
97
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 2
8.
4.
20
12
(R
Š&
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Eu
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m
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St
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x
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Eu
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ta
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(R
Š&
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is
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x
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20
12
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Z)
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ne
at
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M
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86
2
x
8
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Te
m
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ca
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1.
8.
20
08
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Š)
Eu
pi
th
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ra
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ta
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Sc
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ff
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, 1
86
1
x
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7
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 ra
re
Šk
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a,
 1
8.
7.
19
95
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Š&
M
Z)
Eu
pi
th
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tr
ic
at
a
(Z
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9)
x
5
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3.
5.
19
97
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Š&
M
Z)
Acta entomologica slovenica, 24 (2), 2016
128
Eu
pi
th
ec
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bs
in
th
ia
ta
(C
le
rc
k,
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75
9)
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8
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ry
 ra
re
Ji
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9.
8.
20
11
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Š&
M
Z)
Eu
pi
th
ec
ia
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ss
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ila
ta
D
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ay
, 1
85
6
x
x
6,
7
ra
re
G
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ko
, 5
.7
.2
00
9 
(M
Z)
; L
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, 2
5.
6.
20
10
(R
Š&
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B
Z)
Eu
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xi
gu
at
a
(H
üb
ne
r, 
18
13
)
x
x
x
3,
4
no
t c
om
m
on
Eu
pi
th
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te
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ta
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e 
V
ill
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s, 
17
89
)
x
x
x
x
x
8,
9
ve
ry
 c
om
m
on
Eu
pi
th
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m
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ra
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at
a
B
ru
an
d,
 1
85
0
x
x
x
x
x
8,
9,
10
ve
ry
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N
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TO
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O
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A
E 
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4)
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au
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in
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eu
s, 
17
58
)
x
x
x
x
x
8,
9
ve
ry
 c
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m
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Th
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 S
ch
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le
r, 
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75
)
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x
x
7,
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er
a 
cu
rt
ul
a
(L
in
na
eu
s, 
17
58
)
x
x
4,
5
ra
re
Za
gr
aj
ec
, 2
6.
4.
19
93
 (R
Š&
M
Z)
; K
ob
je
gl
av
a 
-
Je
le
nc
a,
 6
.5
.2
01
4 
(R
Š&
M
Z)
C
lo
st
er
a 
an
as
to
m
os
is
(L
in
na
eu
s, 
17
58
)
x
5
ve
ry
 ra
re
O
pa
tje
 se
lo
, 2
.5
.1
98
7 
(R
Š&
M
Z)
N
ot
od
on
ta
 tr
ito
ph
us
(D
en
is
 &
 S
ch
iff
er
m
ül
le
r, 
17
75
)
x
x
4,
5,
7
ve
ry
 ra
re
O
pa
tje
 se
lo
, 2
.5
.1
98
7 
(R
Š&
M
Z)
; K
os
ta
nj
ev
ic
a 
na
K
ra
su
, 1
1.
7.
19
87
 (R
Š&
M
Z)
; G
or
ja
ns
ko
, 2
4.
4.
19
94
(R
Š&
M
Z)
D
ry
m
on
ia
 d
od
on
ae
a
(D
en
is
 &
 S
ch
iff
er
m
ül
le
r, 
17
75
)
x
x
x
x
x
4,
5,
6,
7
co
m
m
on
D
ry
m
on
ia
 ru
fic
or
ni
s(
H
uf
na
ge
l, 
17
66
)
x
x
x
x
x
3,
4,
5
co
m
m
on
D
ry
m
on
ia
 q
ue
rn
a
(D
en
is
 &
 S
ch
iff
er
m
ül
le
r, 
17
75
)
x
x
4,
7
ve
ry
 ra
re
K
os
ta
nj
ev
ic
a 
na
 K
ra
su
, 1
.7
.1
98
9 
(R
Š&
M
Z)
;
K
ob
je
gl
av
a 
- J
el
en
ca
, 2
5.
4.
19
93
 (R
Š&
M
Z)
Ph
eo
si
a 
tr
em
ul
a
(C
le
rc
k,
 1
75
9)
x
5
ve
ry
 ra
re
D
ut
ov
lje
, 7
.5
.1
99
4 
(R
Š&
M
Z)
; K
ob
je
gl
av
a 
-
Za
vo
di
, 8
.5
.2
00
1 
(R
Š&
M
Z)
Pa
ra
dr
ym
on
ia
 v
itt
at
a
(S
ta
ud
in
ge
r, 
18
92
)
x
x
x
x
x
5,
6,
7
no
t c
om
m
on
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at
a 
in
 th
e 
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ha
pt
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 re
ga
rd
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te
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ie
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Pt
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a 
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le
rc
k,
 1
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9)
x
x
x
x
4,
5,
6,
8
ra
re
O
pa
tje
 se
lo
, 1
3.
8.
19
88
 (R
Š&
M
Z)
; O
pa
tje
 se
lo
,
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6.
19
91
 (R
Š&
M
Z)
; K
ob
je
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av
a 
- J
el
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ca
,
16
.5
.1
99
6 
(R
Š&
M
L)
; K
ob
je
gl
av
a 
- J
el
en
ca
,
6.
5.
20
14
 (R
Š&
M
Z)
Radovan Øtanta, Matjaæ Zadrgal: Contribution to the knowledge of Lepidoptera fauna of the Karst
129
Pt
ilo
ph
or
a 
pl
um
ig
er
a
(D
en
is
 &
 S
ch
iff
er
m
ül
le
r, 
17
75
)
x
x
x
x
11
,1
2
co
m
m
on
D
ut
ov
lje
, 1
5.
11
.1
99
5 
(R
Š)
; G
or
ja
ns
ko
, 1
3.
11
.2
01
0
(R
Š)
Pt
ilo
do
n 
ca
pu
ci
na
(L
in
na
eu
s, 
17
58
)
x
x
4,
7
ve
ry
 ra
re
K
os
ta
nj
ev
ic
a 
na
 K
ra
su
, 1
9.
7.
19
88
 (R
Š&
M
Z)
;
Za
gr
aj
ec
, 2
6.
4.
19
93
 (R
Š&
M
Z)
Pt
ilo
do
n 
cu
cu
lli
na
(D
en
is
 &
 S
ch
iff
er
m
ül
le
r, 
17
75
)
x
x
x
x
x
4,
5,
6,
7,
8,
9
no
t c
om
m
on
G
lu
ph
is
ia
 c
re
na
ta
(E
sp
er
, 1
78
5)
x
7
ve
ry
 ra
re
K
os
ta
nj
ev
ic
a 
na
 K
ra
su
, 1
5.
7.
19
87
 (R
Š&
M
Z)
;
K
la
rič
i, 
19
.7
.1
98
7 
(R
Š&
M
Z)
Fu
rc
ul
a 
fu
rc
ul
a
(C
le
rc
k,
 1
75
9)
x
7
ve
ry
 ra
re
K
os
ta
nj
ev
ic
a 
na
 K
ra
su
, 1
5.
7.
19
87
 (R
Š&
M
Z)
;
O
pa
tje
 se
lo
, 1
3.
7.
19
88
 (R
Š&
M
Z)
Fu
rc
ul
a 
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fid
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(B
ra
hm
, 1
78
7)
x
5
ve
ry
 ra
re
O
pa
tje
 se
lo
, 2
7.
5.
19
88
 (R
Š&
M
Z)
D
ic
ra
nu
ra
 u
lm
i(
D
en
is
 &
 S
ch
iff
er
m
ül
le
r, 
17
75
)
x
x
x
x
x
3,
4,
5
co
m
m
on
Ph
al
er
a 
bu
ce
ph
al
a
(L
in
na
eu
s, 
17
58
)
x
x
x
5,
6,
7
ra
re
Ph
al
er
a 
bu
ce
ph
al
oi
de
s(
O
ch
se
nh
ei
m
er
, 1
81
0)
x
x
x
7,
8
ra
re
O
pa
tje
 se
lo
, 2
.8
.1
98
7 
(R
Š&
M
Z)
; Z
ag
ra
je
c,
26
.7
.2
00
8 
(R
Š&
M
Z)
; T
rs
te
lj,
 2
3.
7.
20
09
(R
Š&
M
Z&
TL
); 
K
ra
jn
a 
va
s, 
18
.7
.2
01
4 
(R
Š&
M
Z)
Pe
ri
de
a 
an
ce
ps
(G
oe
ze
, 1
78
1)
x
x
x
x
x
3,
4,
5
co
m
m
on
St
au
ro
pu
s f
ag
i(
Li
nn
ae
us
, 1
75
8)
x
x
x
x
x
4,
5,
6,
7,
9
no
t c
om
m
on
K
la
rič
i, 
13
.9
.2
01
5 
(R
Š)
 - 
se
co
nd
 g
en
er
at
io
n,
 tw
o
m
al
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 o
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er
ve
d
H
ar
py
ia
 m
ilh
au
se
ri
(F
ab
ric
iu
s, 
17
75
)
x
x
x
x
x
3,
4,
5
no
t c
om
m
on
K
la
rič
i, 
27
.9
.2
01
3 
(R
Š&
M
Z)
 - 
un
us
ua
l d
at
e,
 o
ne
m
al
e 
ob
se
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ed
Sp
at
al
ia
 a
rg
en
tin
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(D
en
is
 &
 S
ch
iff
er
m
ül
le
r, 
17
75
)
x
x
x
x
x
4,
5,
6,
7,
8
co
m
m
on
ER
EB
ID
A
E 
(8
3)
N
ud
ar
ia
 m
un
da
na
(L
in
na
eu
s, 
17
61
)
x
x
6
ra
re
D
at
a 
in
 th
e 
C
ha
pt
er
 re
ga
rd
in
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in
te
re
st
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g 
sp
ec
ie
s
M
ilt
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ta
 m
in
ia
ta
(F
or
st
er
, 1
77
1)
x
x
7
ve
ry
 ra
re
K
os
ta
nj
ev
ic
a 
na
 K
ra
su
, 2
6.
7.
19
89
 (R
Š&
M
Z)
;
Za
gr
aj
ec
, 2
6.
7.
20
08
 (R
Š&
M
Z)
C
yb
os
ia
 m
es
om
el
la
(L
in
na
eu
s, 
17
58
)
x
x
x
x
6
ra
re
O
pa
tje
 se
lo
, 1
1.
6.
19
88
 (R
Š&
M
Z)
; K
om
en
 -
Ja
žm
er
ca
, 5
.6
.2
01
0 
(R
Š&
M
Z&
B
Z)
; J
irm
an
ec
,
14
.6
.2
01
3 
(R
Š&
M
Z)
Acta entomologica slovenica, 24 (2), 2016
130
Pe
lo
si
a 
m
us
ce
rd
a
(H
uf
na
ge
l, 
17
66
)
x
9
ve
ry
 ra
re
K
la
rič
i, 
27
.9
.2
01
3 
(R
Š&
M
Z)
Li
th
os
ia
 q
ua
dr
a
(L
in
na
eu
s, 
17
58
)
x
x
x
x
x
6,
7,
8,
9,
10
co
m
m
on
Ei
le
m
a 
de
pr
es
sa
(E
sp
er
, 1
78
7)
x
x
x
x
x
6,
7,
8,
9,
10
co
m
m
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Ei
le
m
a 
lu
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a
(Z
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en
, 1
81
7)
x
x
x
x
x
6,
7,
8
no
t c
om
m
on
K
op
riv
a,
 2
6.
6.
20
11
 (R
Š)
; K
op
riv
a,
 5
.7
.2
01
3 
(M
Z)
;
Po
db
re
že
, 1
2.
7.
20
13
 (R
Š&
M
Z)
; K
la
rič
i, 
18
.6
.2
01
4
(R
Š)
Ei
le
m
a 
co
m
pl
an
a
(L
in
na
eu
s, 
17
58
)
x
x
x
x
x
6,
7,
8,
9,
10
ve
ry
 c
om
m
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Ei
le
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an
ie
l, 
19
39
)
x
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8,
9,
10
no
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om
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at
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 th
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ha
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Ei
le
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üb
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r, 
18
08
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x
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x
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x
5,
6,
7,
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 c
om
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84
7)
x
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x
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x
5,
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8,
9
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m
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le
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(H
uf
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ge
l, 
17
66
)
x
x
x
x
x
4,
5,
6,
7,
9
co
m
m
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Am
at
a 
ph
eg
ea
(L
in
na
eu
s, 
17
58
)
x
x
x
x
x
6,
7
ve
ry
 c
om
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Am
at
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ta
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3)
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6
no
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om
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at
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D
ys
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xe
s a
nc
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a
(L
in
na
eu
s, 
17
67
)
x
x
x
x
x
6,
7,
8
co
m
m
on
D
ys
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xe
s f
am
ul
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(F
re
ye
r, 
18
36
)
x
x
6,
7,
8,
9
no
t c
om
m
on
K
os
ta
nj
ev
ic
a 
na
 K
ra
su
, 2
5.
7.
19
87
 (R
Š&
M
Z)
;
Za
gr
aj
ec
, 1
0.
9.
20
10
 (R
Š&
M
Z&
JD
); 
K
om
en
,
24
.9
.2
01
3 
(R
Š&
M
Z)
; K
la
rič
i, 
18
.6
.2
01
4 
(R
Š)
Ph
ra
gm
at
ob
ia
 fu
lig
in
os
a
(L
in
na
eu
s, 
17
58
)
x
x
x
x
x
3,
4,
5,
6,
7,
8,
9,
10
co
m
m
on
V
oj
šč
ic
a,
 3
1.
3.
19
94
 (R
Š&
M
Z)
Ph
ra
gm
at
ob
ia
 lu
ct
ife
ra
(D
en
is
 &
 S
ch
iff
er
m
ül
le
r, 
17
75
)
x
x
x
4,
5
ra
re
O
pa
tje
 se
lo
, 2
.5
.1
98
7 
(R
Š&
M
Z)
; K
os
ta
nj
ev
ic
a 
na
K
ra
su
, 6
.5
.1
98
9 
(R
Š&
M
Z)
; K
la
rič
i, 
21
.4
.2
01
0
(R
Š&
M
Z)
; J
irm
an
ec
, 8
.5
.2
01
5 
(R
Š&
M
Z&
B
Z&
SG
)
Sp
ilo
so
m
a 
lu
te
a
(H
uf
na
ge
l, 
17
66
)
x
5,
7
ve
ry
 ra
re
K
os
ta
nj
ev
ic
a 
na
 K
ra
su
, 2
3.
5.
19
87
 (R
Š&
M
Z)
;
O
pa
tje
 se
lo
, 1
.7
.1
98
9 
(R
Š&
M
Z)
Radovan Øtanta, Matjaæ Zadrgal: Contribution to the knowledge of Lepidoptera fauna of the Karst
131
Sp
ilo
so
m
a 
lu
br
ic
ip
ed
a
(L
in
na
eu
s, 
17
58
)
x
x
x
5,
6,
8
ra
re
K
ob
je
gl
av
a 
- J
el
en
ca
, 1
9.
5.
19
87
 (R
Š&
M
Z)
; O
pa
tje
se
lo
, 2
.8
.1
98
7 
(M
Z)
; B
rje
 p
ri 
K
om
nu
, 2
2.
5.
19
93
(R
Š&
M
Z)
D
ia
ph
or
a 
m
en
di
ca
(C
le
rc
k,
 1
75
9)
x
5
ve
ry
 ra
re
O
pa
tje
 se
lo
, 2
.5
.1
98
7 
(R
Š&
M
Z)
D
ia
ph
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a 
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sa
(H
üb
ne
r, 
18
31
)
x
x
x
x
x
3,
4,
5
ra
re
K
os
ta
nj
ev
ic
a 
na
 K
ra
su
, 7
.4
.1
98
9 
(R
Š&
M
Z)
;
K
la
rič
i, 
9.
5.
19
92
 (R
Š)
; D
ut
ov
lje
, 2
4.
4.
19
94
(R
Š&
M
Z)
; K
ob
je
gl
av
a 
- J
el
en
ca
, 9
.5
.2
01
2
(R
Š&
M
Z)
; P
od
br
ež
e,
 2
5.
4.
20
13
 (M
Z)
Rh
yp
ar
ia
 p
ur
pu
ra
ta
(L
in
na
eu
s, 
17
58
)
x
x
x
5,
6,
7
ra
re
Lo
kv
ic
a,
 2
8.
6.
19
96
 (R
Š)
; S
ko
po
, 2
6.
6.
20
11
 (R
Š)
D
ia
cr
is
ia
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nn
io
(L
in
na
eu
s, 
17
58
)
x
x
x
x
x
5,
6,
7,
8,
9
co
m
m
on
Ar
ct
ia
 v
ill
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a
(L
in
na
eu
s, 
17
58
)
x
x
x
x
x
4,
5,
6
co
m
m
on
Sp
ir
is
 sl
ov
en
ic
a
(D
an
ie
l, 
19
39
)
x
x
x
x
x
5,
6,
7
co
m
m
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at
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in
 th
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C
ha
pt
er
 re
ga
rd
in
g 
in
te
re
st
in
g 
sp
ec
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s
Eu
pl
ag
ia
 q
ua
dr
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ct
ar
ia
(P
od
a,
 1
76
1)
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x
x
x
x
7,
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 (R
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 S
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)
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61
)
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 (R
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 (R
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67
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Acta entomologica slovenica, 24 (2), 2016
132
Eu
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67
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67
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Radovan Øtanta, Matjaæ Zadrgal: Contribution to the knowledge of Lepidoptera fauna of the Karst
133
M
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58
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Acta entomologica slovenica, 24 (2), 2016
134
H
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on
Radovan Øtanta, Matjaæ Zadrgal: Contribution to the knowledge of Lepidoptera fauna of the Karst
135
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Acta entomologica slovenica, 24 (2), 2016
136
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Radovan Øtanta, Matjaæ Zadrgal: Contribution to the knowledge of Lepidoptera fauna of the Karst
137
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58
)
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8
ve
ry
 ra
re
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os
ta
nj
ev
ic
a 
na
 K
ra
su
, 2
5.
8.
19
89
 (R
Š&
M
Z)
Acta entomologica slovenica, 24 (2), 2016
138
C
uc
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lia
 b
la
tta
ri
ae
(E
sp
er
, 1
79
0)
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co
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 5
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2 
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 l.
 2
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01
 (R
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; T
rs
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22
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1 
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 l.
 2
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20
00
 (R
Š)
 - 
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l l
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on
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ro
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ul
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 c
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 Š
to
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20
05
 (R
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 -
la
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 o
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 c
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 L
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 1
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no
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 re
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 ly
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R
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18
33
x
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99
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. l
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6.
7.
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98
(R
Š)
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ip
a 
na
 K
ra
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20
03
 e
. l
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20
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 (R
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ll 
la
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ae
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ch
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 v
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i(
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re
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pa
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a 
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ra
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- l
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e 
(R
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rs
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.2
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1 
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 l.
 1
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20
00
 (R
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; K
ob
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-
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 6
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.2
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4 
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20
15
 (R
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l
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ap
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at
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D
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im
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 u
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a
(F
re
ye
r, 
18
35
)
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6
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ry
 ra
re
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 p
ri 
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om
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20
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 (M
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pr
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 c
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ta
(D
en
is
 &
 S
ch
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er
m
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le
r, 
17
75
)
x
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7
ra
re
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os
ta
nj
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ic
a 
na
 K
ra
su
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19
94
 (R
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;
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oj
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 1
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19
95
 (R
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M
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ip
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ra
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,
10
.7
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99
8 
(R
Š)
; T
rs
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lj,
 2
3.
7.
20
09
 (R
Š&
M
Z&
TL
)
Pr
ae
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ia
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rm
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St
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ng
er
, 1
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2
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9
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re
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 4
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01
0 
(R
Š&
M
Z&
B
Z)
Am
ph
ip
yr
a 
py
ra
m
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ea
(L
in
na
eu
s, 
17
58
)
x
x
x
x
x
6,
7,
8,
9,
10
,1
1
ve
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 c
om
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Am
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 1
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9)
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15
.9
.1
99
5 
(M
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ph
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(F
ab
ric
iu
s, 
17
87
)
x
x
x
x
7,
8,
9,
10
no
t c
om
m
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K
os
ta
nj
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ic
a 
na
 K
ra
su
, 1
9.
8.
19
94
 (R
Š&
M
Z)
;
G
or
ja
ns
ko
 - 
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dk
la
ne
c,
 3
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00
9 
(R
Š&
M
Z)
;
Ji
rm
an
ec
, 1
9.
8.
20
11
 (R
Š&
M
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; G
or
ja
ns
ko
 -
M
av
hi
nj
e,
 2
.1
0.
20
11
 (R
Š)
As
te
ro
sc
op
us
 sp
hi
nx
(H
uf
na
ge
l, 
17
66
)
x
x
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11
,1
2
no
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om
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on
Br
ac
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 n
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os
a
(E
sp
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78
5)
x
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4
no
t c
om
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Lo
ke
v,
 1
6.
3.
20
12
 (R
Š&
M
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; K
rv
av
i P
ot
ok
,
18
.4
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3 
(R
Š&
B
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SG
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(D
en
is
 &
 S
ch
iff
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r, 
17
75
)
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co
m
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M
eg
an
ep
hr
ia
 b
im
ac
ul
os
a
(L
in
na
eu
s, 
17
67
)
x
x
x
x
x
9,
10
no
t c
om
m
on
V
oj
šč
ic
a,
 9
.1
0.
19
95
 (R
Š)
; G
or
ja
ns
ko
 - 
M
av
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nj
e,
29
.9
.2
01
1 
(R
Š)
; L
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ev
, 3
0.
9.
20
11
 (R
Š&
M
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;
K
om
en
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4.
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20
13
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Š&
M
Z)
Radovan Øtanta, Matjaæ Zadrgal: Contribution to the knowledge of Lepidoptera fauna of the Karst
139
Al
lo
ph
ye
s o
xy
ac
an
th
ae
(L
in
na
eu
s, 
17
58
)
x
x
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x
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10
,1
1,
12
ve
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 c
om
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Py
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(H
uf
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ge
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17
66
)
x
x
8,
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ve
ry
 ra
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O
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, 8
.9
.1
99
2 
(R
Š)
; D
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.6
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99
4
(M
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; L
ip
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 K
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8.
8.
19
94
 (R
Š)
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hi
ni
a 
ca
rd
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(H
üb
ne
r, 
17
90
)
x
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7,
8
no
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 th
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 re
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H
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(D
en
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 &
 S
ch
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17
75
)
x
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6,
7,
8,
9
ra
re
O
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tje
 se
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, 1
3.
8.
19
88
 (R
Š&
M
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; D
ut
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,
31
.3
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4 
(R
Š&
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; D
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99
4 
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Š&
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 v
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17
66
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co
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18
13
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20
11
 (R
Š)
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la
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, 1
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19
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95
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; M
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7 
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ab
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17
75
)
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17
75
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 V
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 (M
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ra
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20
01
 (R
Š&
B
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;
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 2
8.
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20
11
 (R
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75
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 1
9.
5.
19
97
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2 
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7,
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9,
10
no
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El
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17
90
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5,
6,
7
no
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H
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66
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94
 (R
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6.
9.
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95
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la
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3 
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Š&
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sp
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sa
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am
bu
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18
34
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9
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87
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8.
7.
19
95
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Š&
M
Z)
; T
rs
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 1
6.
7.
20
10
(R
Š)
Acta entomologica slovenica, 24 (2), 2016
140
C
ar
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ri
na
 c
la
vi
pa
lp
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Sc
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17
63
x
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75
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 (R
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.8
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1 
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3 
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75
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7
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75
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1
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17
66
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6
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sp
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5)
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6,
7
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re
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 1
0.
9.
19
95
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Z)
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0
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Š&
M
Z&
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Z)
; K
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 2
0.
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20
11
 (R
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9.
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11
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Š)
D
yp
te
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(L
in
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eu
s, 
17
58
)
x
x
x
x
5,
6,
7,
8,
9
no
t c
om
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Tr
ac
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in
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58
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x
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x
5,
6,
7,
8,
9
no
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om
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M
or
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o 
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ra
(L
in
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eu
s, 
17
58
)
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x
7
ve
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 ra
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K
os
ta
nj
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a 
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 K
ra
su
, 4
.7
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99
6 
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Š)
; P
on
ik
ve
,
29
.6
.2
01
3 
(R
Š&
M
Z&
C
M
)
Po
ly
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ae
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sp
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7)
x
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x
6,
7,
8,
9
ve
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 c
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Th
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uf
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66
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x
x
7,
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9,
10
ve
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 1
75
9)
x
x
7
ve
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 ra
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pa
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9.
7.
19
88
 (R
Š&
M
Z)
; K
ob
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gl
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a 
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le
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6.
5.
19
96
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Š&
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L)
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hl
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 h
yp
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ic
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D
en
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 &
 S
ch
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m
ül
le
r, 
17
75
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x
x
x
4,
5,
8,
9,
10
ra
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G
or
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 1
0.
9.
19
95
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Š&
M
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;
Za
gr
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3.
10
.2
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8 
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Š)
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3.
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12
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Š)
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in
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17
58
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on
Radovan Øtanta, Matjaæ Zadrgal: Contribution to the knowledge of Lepidoptera fauna of the Karst
141
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 (R
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 (R
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 (R
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 2
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 (R
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 (M
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 (R
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 (R
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17
58
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66
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Ji
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 &
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10
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2 
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Š)
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20
13
 (R
Š&
M
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nt
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sp
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, 1
79
1)
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3
ra
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Š)
; D
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5.
3.
19
94
(R
Š&
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Š&
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Acta entomologica slovenica, 24 (2), 2016
142
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 (M
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17
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17
58
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 1
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ve
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m
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98
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; B
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 (R
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 (R
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 (R
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 (R
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Radovan Øtanta, Matjaæ Zadrgal: Contribution to the knowledge of Lepidoptera fauna of the Karst
143
Li
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95
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58
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75
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Acta entomologica slovenica, 24 (2), 2016
144
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 (R
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 (R
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 1
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)
Radovan Øtanta, Matjaæ Zadrgal: Contribution to the knowledge of Lepidoptera fauna of the Karst
145
G
or
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94
 (R
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 (R
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Acta entomologica slovenica, 24 (2), 2016
146
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58
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75
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58
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61
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58
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12
 (R
Š)
Radovan Øtanta, Matjaæ Zadrgal: Contribution to the knowledge of Lepidoptera fauna of the Karst
147
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üb
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18
09
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89
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75
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 1
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96
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11
 (R
Š&
M
Z)
; K
op
riv
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 1
5.
5.
20
13
(R
Š&
M
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ad
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sp
er
, 1
78
8)
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ra
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av
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- J
el
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5.
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96
 (R
Š&
M
L)
;
Po
ni
kv
e,
 1
2.
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20
13
 (R
Š)
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ad
en
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rp
le
xa
(D
en
is
 &
 S
ch
iff
er
m
ül
le
r, 
17
75
)
x
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ra
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os
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nj
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ic
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 K
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87
 (R
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;
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or
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 (M
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sp
iti
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D
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 &
 S
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le
r, 
17
75
)
x
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10
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om
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od
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 1
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9,
10
ve
ry
 c
om
m
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M
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m
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in
na
eu
s, 
17
61
)
x
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ve
ry
 ra
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 p
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2.
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93
 (R
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28
.8
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 &
 S
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r, 
17
75
)
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5,
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10
no
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aw
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9)
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10
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1
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6,
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9
ra
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os
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 K
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8.
19
87
 (R
Š&
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;
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 se
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7.
6.
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94
 (R
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or
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2.
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20
09
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; Z
ag
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 2
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 (R
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en
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 &
 S
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17
75
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,
11
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ab
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17
87
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no
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om
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in
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eu
s, 
17
67
)
x
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x
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9,
10
,
11
co
m
m
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M
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m
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 ri
pa
ri
a
(R
am
bu
r, 
18
29
)
x
x
x
x
4,
5,
6,
7,
8,
9
ra
re
O
pa
tje
 se
lo
, 2
1.
9.
19
90
 (R
Š&
M
Z)
; K
ob
je
gl
av
a,
26
.4
.1
99
7 
(R
Š&
M
Z)
; Z
ag
ra
je
c,
 2
2.
5.
20
09
(R
Š&
M
Z)
; J
irm
an
ec
, 3
.9
.2
01
5 
(R
Š&
M
Z&
C
M
)
Acta entomologica slovenica, 24 (2), 2016
148
Le
uc
an
ia
 o
bs
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et
a
(H
üb
ne
r, 
18
03
)
x
x
5,
6,
8
ve
ry
 ra
re
K
os
ta
nj
ev
ic
a 
na
 K
ra
su
, 1
5.
8.
19
87
 (R
Š&
M
Z)
;
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ob
je
gl
av
a 
- J
el
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ca
, 1
6.
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19
96
 (R
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L)
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uc
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 p
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s(
H
üb
ne
r, 
18
24
)
x
x
8,
9
ra
re
K
os
ta
nj
ev
ic
a 
na
 K
ra
su
, 2
5.
8.
19
89
 (R
Š&
M
Z)
;
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la
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i, 
7.
9.
19
93
 (R
Š&
M
Z)
; B
rje
 p
ri 
K
om
nu
,
25
.9
.2
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9 
(R
Š&
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, 1
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10
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Pa
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 &
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17
75
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x
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no
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uf
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66
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x
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x
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ve
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rt
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 &
 S
ch
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17
75
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5
no
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rt
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ab
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s, 
17
75
)
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x
2,
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ve
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 c
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rt
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en
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 &
 S
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le
r, 
17
75
)
x
x
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x
2,
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ve
ry
 c
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m
on
O
rt
ho
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a 
gr
ac
ili
s(
D
en
is
 &
 S
ch
iff
er
m
ül
le
r, 
17
75
)
x
x
x
x
x
3,
4
ra
re
K
os
ta
nj
ev
ic
a 
na
 K
ra
su
, 1
9.
3.
19
90
 (R
Š&
M
Z)
;
D
ut
ov
lje
, 3
1.
3.
19
94
 (R
Š&
M
Z)
; K
la
rič
i, 
26
.3
.1
99
5
(R
Š&
M
Z)
; K
rv
av
i P
ot
ok
, 1
8.
4.
20
13
 (R
Š&
B
Z&
SG
)
O
rt
ho
si
a 
op
im
a
(H
üb
ne
r, 
18
09
)
x
x
x
x
3,
4,
5
ra
re
D
ut
ov
lje
, 2
5.
3.
19
94
 (R
Š&
M
Z)
; K
la
rič
i, 
26
.3
.1
99
5
(R
Š&
M
Z)
; K
az
lje
, 2
8.
3.
20
12
 (R
Š&
M
Z)
; K
rv
av
i
Po
to
k,
 1
8.
4.
20
13
 (R
Š&
B
Z&
SG
)
O
rt
ho
si
a 
go
th
ic
a
(L
in
na
eu
s, 
17
58
)
x
x
x
x
x
3,
4,
5
co
m
m
on
An
or
th
oa
 m
un
da
(D
en
is
 &
 S
ch
iff
er
m
ül
le
r, 
17
75
)
x
x
x
x
x
2,
3,
4
ve
ry
 c
om
m
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Pe
ri
gr
ap
ha
 ro
ri
da
Fr
iv
al
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y,
 1
83
5
x
3,
4
ra
re
D
at
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 th
e 
C
ha
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ga
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te
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s
Pe
ri
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ic
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i 1
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6)
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4
ve
ry
 ra
re
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i P
ot
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8.
4.
20
13
 (R
Š&
B
Z&
SG
)
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ir
a 
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ns
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s(
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nn
ae
us
, 1
75
8)
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x
x
x
x
3,
4,
5
co
m
m
on
Eu
xo
a 
te
m
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a
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üb
ne
r, 
18
08
)
x
x
x
x
9
ve
ry
 ra
re
K
la
rič
i, 
7.
9.
19
93
 (R
Š&
M
Z)
; K
os
ta
nj
ev
ic
a 
na
K
ra
su
, 2
2.
9.
19
93
 (R
Š&
M
Z)
; L
ip
a 
na
 K
ra
su
,
10
.9
.1
99
5 
(R
Š&
M
Z)
; K
az
lje
, 1
7.
9.
20
11
 (R
Š&
M
Z)
Eu
xo
a 
co
s(
H
üb
ne
r, 
18
24
)
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8
ve
ry
 ra
re
Te
m
ni
ca
, 2
9.
8.
20
08
 (M
Z)
Ag
ro
tis
 c
in
er
ea
(D
en
is
 &
 S
ch
iff
er
m
ül
le
r, 
17
75
)
x
5
ve
ry
 ra
re
K
ob
je
gl
av
a,
 1
8.
5.
19
96
 (M
Z)
; K
ob
je
gl
av
a 
- J
el
en
ca
,
6.
5.
20
14
 (R
Š&
M
Z)
Radovan Øtanta, Matjaæ Zadrgal: Contribution to the knowledge of Lepidoptera fauna of the Karst
149
Ag
ro
tis
 se
ge
tu
m
(D
en
is
 &
 S
ch
iff
er
m
ül
le
r, 
17
75
)
x
x
x
x
x
3,
4,
5,
6,
7,
8,
9,
10
,1
1
co
m
m
on
Ag
ro
tis
 e
xc
la
m
at
io
ni
s(
Li
nn
ae
us
, 1
75
8)
x
x
x
x
x
5,
6,
7,
8,
9,
10
co
m
m
on
Ag
ro
tis
 ip
si
lo
n
(H
uf
na
ge
l, 
17
66
)
x
x
x
x
x
3,
4,
5,
6,
7,
8,
9,
10
,1
1
co
m
m
on
Ag
ro
tis
 b
ig
ra
m
m
a
(E
sp
er
, 1
79
0)
x
x
x
8,
9
ra
re
O
pa
tje
 se
lo
, 2
5.
8.
19
89
 (R
Š&
M
Z)
; K
la
rič
i, 
7.
9.
19
95
(R
Š&
M
Z)
; J
irm
an
ec
, 1
9.
8.
20
11
 (R
Š&
M
Z)
; L
ok
ev
,
25
.8
.2
01
4 
(R
Š)
Pe
ri
dr
om
a 
sa
uc
ia
(H
üb
ne
r, 
18
08
)
x
x
x
x
x
8,
9,
10
,1
1
co
m
m
on
Ax
yl
ia
 p
ut
ri
s(
Li
nn
ae
us
, 1
76
1)
x
x
x
6,
7,
8
ra
re
O
ch
ro
pl
eu
ra
 p
le
ct
a
(L
in
na
eu
s, 
17
61
)
x
x
x
x
x
5,
6,
7,
8,
9
no
t c
om
m
on
C
he
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ot
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 m
ar
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ta
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a
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ill
er
s, 
17
89
)
x
x
9,
10
ve
ry
 ra
re
V
oj
šč
ic
a,
 1
6.
10
.1
99
3 
(M
Z)
; J
irm
an
ec
, 5
.9
.2
01
3
(R
Š&
M
Z)
N
oc
tu
a 
pr
on
ub
a
(L
in
na
eu
s, 
17
58
)
x
x
x
x
x
5,
6,
7,
8,
9,
10
,
11
ve
ry
 c
om
m
on
N
oc
tu
a 
fim
br
ia
ta
(S
ch
re
be
r, 
17
59
)
x
x
x
x
x
6,
7,
8,
9,
10
co
m
m
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tu
a 
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re
ni
ca
B
ie
bi
ng
er
, S
pe
id
el
 &
 H
an
ig
k,
 1
98
3
x
x
x
x
x
9,
10
ra
re
Za
gr
aj
ec
, 3
0.
9.
20
10
 (R
Š)
; K
az
lje
, 1
7.
9.
20
11
(R
Š&
M
Z)
; S
ko
po
, 1
2.
10
.2
01
1 
(R
Š)
; J
irm
an
ec
,
5.
9.
20
13
 (R
Š&
M
Z)
; K
la
rič
i, 
27
.9
.2
01
3 
(R
Š&
M
Z)
N
oc
tu
a 
in
te
rp
os
ita
(H
üb
ne
r, 
17
90
)
x
x
x
x
7,
8,
9
ra
re
Li
pa
 n
a 
K
ra
su
, 1
0.
7.
19
98
 (R
Š)
; K
op
riv
a,
 1
3.
8.
20
11
(R
Š)
; J
irm
an
ec
, 1
9.
8.
20
11
 (R
Š&
M
Z)
; J
irm
an
ec
,
5.
9.
20
13
 (R
Š&
M
Z)
N
oc
tu
a 
co
m
es
H
üb
ne
r, 
18
13
x
x
x
x
x
5,
6,
7,
8,
9,
10
co
m
m
on
N
oc
tu
a 
in
te
rj
ec
ta
H
üb
ne
r, 
18
03
x
x
x
x
7,
8,
9
no
t c
om
m
on
N
oc
tu
a 
ja
nt
hi
na
D
en
is
 &
 S
ch
iff
er
m
ül
le
r, 
17
75
x
x
x
x
x
6,
7,
8,
9,
10
no
t c
om
m
on
N
oc
tu
a 
te
rt
ia
M
en
tz
er
 &
 a
l.,
 1
99
1
x
8
Ji
rm
an
ec
, 1
9.
8.
20
11
 (R
Š&
M
Z)
Ep
ile
ct
a 
lin
og
ri
se
a
(D
en
is
 &
 S
ch
iff
er
m
ül
le
r, 
17
75
)
x
x
x
x
8,
9
no
t c
om
m
on
Acta entomologica slovenica, 24 (2), 2016
150
Ly
co
ph
ot
ia
 p
or
ph
yr
ea
(D
en
is
 &
 S
ch
iff
er
m
ül
le
r, 
17
75
)
x
x
5,
6,
7
ra
re
Li
pa
 n
a 
K
ra
su
, 2
9.
5.
19
97
 (R
Š)
; L
ok
vi
ca
, 2
5.
5.
20
02
(R
Š&
B
Z&
SG
); 
K
op
riv
a,
 1
0.
7.
20
13
 (R
Š)
D
ia
rs
ia
 ru
bi
(V
ie
w
eg
, 1
79
0)
x
5
ve
ry
 ra
re
O
pa
tje
 se
lo
, 9
.5
.1
98
7 
(M
Z)
Eu
gn
or
is
m
a 
de
pu
nc
ta
(L
in
na
eu
s, 
17
61
)
x
9
ve
ry
 ra
re
Ji
rm
an
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, 3
.9
.2
01
5 
(R
Š&
M
Z&
C
M
)
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st
ia
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-n
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m
(L
in
na
eu
s, 
17
58
)
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x
x
x
x
5,
6,
7,
8,
9,
10
co
m
m
on
Xe
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um
(H
uf
na
ge
l, 
17
66
)
x
x
x
x
6,
7
ra
re
G
or
ja
ns
ko
 - 
Po
dk
la
ne
c,
 3
.7
.2
00
9 
(R
Š&
M
Z)
;
Te
m
ni
ca
, 2
1.
7.
20
09
 (R
Š&
M
Z)
; K
op
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 6
.7
.2
01
4
(R
Š&
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M
)
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ia
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ig
m
at
ic
a
(H
üb
ne
r, 
18
13
)
x
x
x
x
8,
9
no
t c
om
m
on
Xe
st
ia
 c
as
ta
ne
a
(E
sp
er
, 1
79
8)
x
x
x
x
x
9,
10
ve
ry
 c
om
m
on
Xe
st
ia
 x
an
th
og
ra
ph
a
(D
en
is
 &
 S
ch
iff
er
m
ül
le
r, 
17
75
)
x
x
x
x
x
8,
9,
10
ve
ry
 c
om
m
on
C
er
as
tis
 ru
br
ic
os
a
(D
en
is
 &
 S
ch
iff
er
m
ül
le
r, 
17
75
)
x
x
x
x
x
3,
4,
5
co
m
m
on