HACQUETIA 5/1 • 2006, 113–130



moss FlorA oF THE ProKlETIJE moUnTAIns 


(sErBIA, monTEnEgro)


Andrej MARTInČIČ* 

Abstract 

The author presents the results of his own research of flora of mosses (Musci) of the part of Prokletije Mts. that 
lies in Serbia and in Montenegro. He mentions 268 species for the part, which so far remained briofloristically practically unknown; among them 10 species are new records for Serbia and 44 for Montenegro. In his 
horological analysis, the author specifically emphasises the large proportion of »northern elements« (arctic 
alpine, subarctic-subalpine, boreal-montane, boreal-temperate), which migrated into the Prokletije Mts. in 
the Pleistocene. 
Key words: Bryophyta 
(Musci) flora, Prokletije Mts. (Serbia, Montenegro), horological analysis, red-listed species 

Izvleček 

Avtor predstavlja rezultate lastnih raziskovanj flore listnatih mahov (Musci) tistega dela gorskega masiva Prokletije, ki leži v državi Srbija in Črna gora. Za predel, ki je bil doslej briofloristično praktično nepoznan, navaja 
268 vrst; med njimi je prvič zabeleženih 10 vrst za Srbijo in 44 za Črno goro. V horološki analizi avtor posebej 
poudarja velik delež »severnih elementov« (arktično-alpinski, subarktično-subalpinski, borealno-montanski, 
borealno-temperatni), ki so se priselili v Prokletije v pleistocenu. 
Ključne besede: mahovna flora (Musci), Prokletije (Srbija, Črna gora), horološka analiza, vrste na Rdečem 
seznamu 

InTRoDUCTIon author was therefore able to collect relatively rich 

material in the period between 1961 – 197 , which, 
The region of former yugoslavia is briofloristically despite being incomplete, throws new light on the 
very unevenly researched. The most information floristic and phytogeographical situation of this 
that throws light on moss flora relates to the terri-mountain range. Most of the moss material was coltory of Slovenia and littoral parts (comp. Pavletić lected in the higher regions, above the timberline. 
1955, MaRtinčič 1968, Düll & al. 1999). Among the 
least investigated parts are certain mountain ranges 
in Montenegro, Macedonia and Serbia (kosovo). THE STUDy AREA 
oneofsuchregionsisalsotheProkletijeMts.There 
was only a little information on the moss flora avail-Prokletije Mts. is a massif extending over the wid-
able so far for the part that lies in the region of er border region between Serbia, Montenegro 
Serbia and Montenegro (Figure 1). This data was (republics of former yugoslavia) and Albania. It 
contributed by BauMgaRtneR (1915), szePesfalvy in represents the southeastern end of the Dinaric 
csiki, JavoRka & küMMeRle (1926), RuDski (19 6, mountains which closes with the river Drim in the 
1949), MaRtinčič (196 ) and Poelt (1976). The south, i.e. in Albania. In the former yugoslav re


* Zaloška 78a, SI-1000 Ljubljana 
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Hacquetia 5/1 • 2006, 113–130 

gion it extends between Lake Skadar and Vrmoša 
in the west, and Beli Drim in the east. In the north, 
it ends with Hajla, Žljeb and Mokra planina. The 
valleys of rivers Pećka Bistrica, Dečanska Bistrica 
and some others, as well as the Plavska kotlina basin cut the »yugoslav« part of Prokletije Mts. into 
several distinctive groups. The highest peak is Djeravica at 2656 m and most of the peaks are higher 
than 2400 m, among them Marjaš (25 0 m), Žuti 
kamen (2522 m), karanfil (2480 m), Starac (2426 
m). geologically speaking there is great diversity. 
After Marković (1958) a big part of the »yugoslav« 
Prokletije Mts. consists of Permian-Carbon slates 
and Palaeozoic slate limestones. In places, they are 
covered with thick layers of Triassic limestones and 
dolomites. Cretaceous and Jurassic limestone and 
in places flysch are represented to a smaller extent. 
Acid eruptive rocks are very much localised. In the 
Pleistocene, Prokletije Mts. were exposed to a very 
strong influence of glaciation. The ice covered not 
only the peaks, but extended all the way down to 
the valleys, where there were also glaciers. Today, 
the traces of former glaciers are visible everywhere, 

especially in basins, glacier lakes and glacier valleys 
and moraines. 

Collecting localities 

Localities (Figure 1) are equipped with the number 
of the base unit of the Central-European floristic 
mapping system (MTB), which was used for the 
entire territory of former yugoslavia. The denotation in parenthesis is from the UTM network ( 4T 
world part), and for subdivisions we applied the 
grid of 10×10 km. Most of the localities have the 
abbreviation of the state in which they are located: 
Serbia (s) or Montenegro (m). Four of the localities, namely the Čakor mountain pass and the peaks 
of Devojački krš, Starac, Pasji vrh, are on the border 
between the two states, and as the moss material 
was collected on both sides, it is impossible to determine its origin. Regretfully, ecological denotation 
of particular localities remains highly inadequate. 
Altitude most often indicates only the belt in which 
the moss material on a certain locality was collect-

Figure 1: Map of the Prokletije Mts. (part in Serbia, Montenegro) with position of investigated Localities (numbered circles). 
Slika 1: Skica srbsko-črnogorskega dela Prokletij z vrisanimi raziskanimi lokalitetami (oštevilčeni krogci). 
114 



anDReJ MaRtinčič: Moss floRa of tHe PRokletiJe Mountains (seRBia, MontenegRo) 


ed. geological – petrographic data are often very 
generalized – there are relatively few peaks with a 
homogenous petrographic composition, as it literally changes at every step on most of the peaks. 

1: 
Mt. Vaganica, 1900 m, 2000 m; alpine grasslands and rock crevices; paleozoic slate limestones. MTB 285 (Dn12). s. 
2: 
Boge – slope above valley of Bogska reka river, 
1500 m, 1700 m; triassic limestone. MTB 286 
(Dn2 ). s. 
: 
Valley of Bogska reka river, 1100–1 00 m; triassic limestone. MTB 286 (Dn2 ). s. 

4: 
Mt. Hajla: Dramodol, 2100 m; alpine grasslands and rock crevices; limestone. MTB 286 
(Dn2 ). s. 
5: 
Mt. Hajla: Škreli above Boge, 1600 m, 1800 m; 
limestone. MTB 286 (Dn2 ). s. 
6: 
Mt. Hajla, 1900 m, 2270 m; alpine grasslands 
and rock crevices; triassic limestone. MTB 286 
(Dn2 ). s. 
7: 
Mt. Žljeb, 1700 m, 2000 m; alpine grasslands 
and rock crevices; triassic limestone. MTB 287 
(Dn ). s. 
8: 
Mt. Vizitor, 1100 m, 1500 m, 1750 m, 1900 m; 
forests, subalpine and alpine grasslands, 
springs; limestone. MTB 85 (Dn01). m. 
9: 
Mt. Babina gora, 1600–1700 m; limestone. 
MTB 86 (Dn22). s. 
10: Mountain pass Čakor, 1850 m; paleozoic slate 
limestones. MTB 86 (Dn12). s-m. 
11: Mt. 
Devojački krš, 1900 m, 2000 m; alpine 
grasslands and rock crevices; paleozoic slate 
limestones. MTB 86 (Dn12). s-m. 
12: near the spring of Istočka reka river at the 
town of Peć, 600 m; limestone. MTB 288 ( 
Dn5 ). s. 
1 : Mt. Malje 
nedjinat, 1700–2200 m; subalpine 
and alpine grasslands, rock crevices; triassic 
limestone. MTB 86 (Dn22). s. 

14: Mt. 
Marjaš, 1900–2400 m; alpine grasslands 
and rock crevices; paleozoic slate limestones. 
MTB 86 (Dn21). s. 
15: Mt. Planinica, 1900–2000 m; alpine grasslands; 
limestone. MTB 86 (Dn12). s. 
16: Mt. Starac, 1900–2400 m; alpine grasslands, 
rock crevices, springs; paleozoic slates. MTB 
86 (Dn22). s-m. 
17: Mt. Žuti kamen, 1800–2400 m; alpine grasslands, rock crevices; triassic limestone. MTB 
86 (Dn22). s. 
18: gorge Rugovska klisura, 500 m; rocks; limestone. MTB 87 (Dn 2). s. 
19: Springs Alipašini izvori near gusinje, 1000 m; 
springs, rock crevices; triassic limestone. MTB 
484 (Dn01). m. 
20: Valley grbaja, 1000 m, 1200 m; triassic limestone. MTB 484 (Dn00). m. 
21: Mt. karanfil-slopes, 1450–1600 m; forests; triassic limestone. MTB 484 (Dn00). m. 
22: Mt. Popadija, 1700–2000 m; alpine grasslands, 
rock crevices; triassic limestone and cretaceous 
flysch. MTB 484 (Cn90). m. 
2 : Mt. Trojan, 1900–2180 m; alpine grasslands, 
rock crevices; triassic limestone. MTB 484 
(Cn91). m. 

24: Mt. Volušnica, 1600 m; subalpine grasslands; 
flysch. MTB 484 (Cn90). m. 
25: near gusinje, 1000 m; limestone. MTB 485 
(Dn01). m. 
26: Savina voda in valley Ropojana, 1200 m; springs; 
triassic limestone. MTB 485 (Dn00). m. 
27: Vragonos above river Treskavička reka , 1500 m; 
stand of Pinus leucodermis; flint conglomerate. 
MTB 485 (Dn11). m. 
28: Mt. 
Djeravica, 2200–2600 m; alpine grasslands, rock crevices; eruptive-gabro. MTB 486 
(Dn20-21). s. 
29: Mt. Pasji vrh, 1900–2400 m; alpine grasslands; 
limestone. MTB 486 (Dn21). s-m. 
0: Around lake Ridsko jezero, 2000 m; rocks, 
rock crevices; flint conglomerate. MTB 486 
(Dn21). m. 

 1: Valley of river Treskavička reka, 1400–1500 m; 
flint conglomerate. MTB 486 (Dn11). m. 
2: Mt. Treskavica under Veliki Rid, 1600–1900 m; 
flint conglomerate. MTB 486 (Dn11). m.


 : Valley of river Dečanska Bistrica, 900–1200 m; 
paleozoic slate. MTB 487 (Dn 1). s. 

 4: Valley of river Ločanska Bistrica, 1 00 m; paleozoic slate. MTB 487 (Dn 0). s. 

 5: Mt. Maja kurvala, 1700–1900 m; paleozoic limestone. MTB 487 (Dn 1). s. 

 6: Valley Ropojana, 1200 m; triassic limestone. 
MTB 584 (Dn00). m. 

 7: Mt. karanfil-peak, 2000–2400 m; alpine grasslands, rock crevices; triassic limestone. MTB 
584 (Dn00).m. 

 8: Mt. Belić, 1600–2 00 m; forests, subalpine and 
alpine grasslands, rock crevices; triassic limestone. MTB 585 (Dn00). m. 

For the sake of completeness we included data from the literature as well (BauMgaRtneR 1915, 
szePesfalvy 1926, RuDski 19 6, 1949, Poelt 1976). 
The source BReiDleR (1888) was not considered, 


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Hacquetia 5/1 • 2006, 113–130 


because it is not clear if the localities in this work 

e. g. Vila, kokura, Širokar, Mojan, lake Rikavac belong to the Prokletije Mts. 
nomenclature and taxonomy follows the work 
MaRtinčič (200 ): Annotated check-list of mosses 
of Slovenia, which is mostly based on the work of 
DieRssen (2001). The map of study area (Figure 1) 
is equipped with the grid of 10×10 km from UTM 
network. names of localities are from map “Prokletije, mountain map” (1966). The specimens are 
preserved in the Herbarium of Department of Biology (Biotechnical faculty) University of Ljubljana (LJU). 

RESULTS AnD DISCUSSIon 

Floristical results 

Based on the data acquired so far, the flora of 
mosses (Musci) of the »yugoslav« part of Prokletije 
Mts. comprises 268 species. This represents 42 % 
of moss species recorded till now in Serbia-Montenegro. The folowing 16 species are new to Serbia-
Montenegro: Brachythecium erythrorhizon, B. glaciale, 

B.oedipodium,B.trachypodium,Bryumblindii,B.muehlenbeckii, B. sauteri, Didymodon icmadophylus, Grimmia 
affinis, G. decipiens, Lescuraea saxicola, Orthotrichum 
alpestre, Platydictya jungermannioides, Racomitrium 
aciculare, Rhynchostegiella tenuicaulis and Scorpidium 
cossonii. In Serbia are for the first time reported: 
Brachythecium glaciale, B. oedipodium, Bryum blindii, 
B. sauteri, Didymodon icmadophilus, Grimmia affinis, 
Racomitrium aciculare, Rhynchostegiella tenuicaulis, 
Timmia bavarica, T. norvegica and 44 species are new 
to Montenegro (marked with * in List of taxa). The 
number of recorded moss species for Prokletije 
Mts. is not definite and with further research, especially of the lower part for which we have very little 
data, the species number will be increased. 
List of taxa: 

Amblystegium confervoides 
(Brid.) B., S. & g.: bortemp – Planinica, 1500 m. 
Amblystegium serpens 
(Hedw.) B., S. & g.: ubikv – valley of river Bogska reka, 1 00 m; Belić, 1700 m. 

*Amblystwgium subtile 
(Hedw.) B., S. & g.: temp-subkont – valley grbaja, 1200 m; valley Ropojana: 
near Savina voda, 1200 m; Vragonos above 
river Treskavička reka, 1500 m; valley of river 
Treskavička reka, 1400 m; Volušnica, 1600 m. 

Amphidium mougeotii 
(B. & S.) Schimp.: bor-mont 

– Djeravica, 2500 m; SW above Dečani, 1600 m 
(BauMgaRtneR 1915). 
Andreaea rothii Web. & Mohr subsp. rothii: bor-temp 

- Djeravica, 2500 m. 
Andreaea rupestris 
Hedw. var. rupestris: bor-mont 
– Djeravica, 2500 m; Starac, 2100 m. 
Anomodon attenuatus 
(Hedw.) Hüben.: subtemp – Vizitor, 1700 m; valley grbaja, 1200 m; Vragonos 
above river Treskavička reka, 1500 m; valley 
Ropojana, 1200 m. 

Anomodon viticulosus 
(Hedw.) Hook. & Tayl.: bortemp – at spring of river Istočka reka near Peć, 
600 m; valley of river Bogska reka near kučište, 
1100 m; springs Alipašini izvori near gusinje, 
1000 m; valley grbaja, 1200 m; valley Ropojana, 
1200 m; Belić, 1650 m. 

Antitrichia curtipendula 
(Hedw.) Brid.: bor-temp – 
Starac, 1800 m; valley Ropojana, 1200 m. 
Atrichum undulatum (Hedw.) P. Beauv.: bor-temp. 

– Vizitor, 1500 m; Hajla: Dramodol, 2100 m; 
Babina gora, 1600 m; Malje nedjinat, 1900 m; 
Vragonos above river Treskaviička reka, 1500 m; 
Dečani near Peć (szePesfalvy 1926 – 87, 
Dn42). 
Aulacomnium palustre (Hedw.) Schwaegr.: subbor 

– Starac, 2100 m; Djeravica, 1900 m. 
Barbula convoluta Hedw.: ubikv – Žljeb, 2000 m. 
Barbula unguiculata 
Hedw.: ubikv – Trojan, 1900 m; 
Belič, 2 00 m; pass Čakor (szePesfalvy 1926); 
near Peć (szePesfalvy 1926 – 87, Dn42). 

Bartramia halleriana 
Hedw.: bor-mont – Babina gora, 
1600 m; valley of river Treskavička reka, 1400 m; 
valley Ropojana, 1200 m. 

Bartramia ithyphylla 
Brid.: bor-mont – Vizitor, 
1900 m; Žljeb, 2000 m; Devojački krš, 1900 m; 
Starac, 2 00 m; Planinica, 1900 m; Babina gora, 1600 m; Malje nedjinat, 1900 m; Djeravica, 
2500 m; Treskavica under Veliki Rid, 1800 m; 
Belić, 1900 m; Žljeb, 1700 m. 

var. strigosa 
(Wahlenb.) C. Hartm. – Pasji vrh, 
2100 m. 

var. breviseta 
(Lindb.) kindb. – Maja kurvala, 
1900 m. 

Bartramia pomiformis 
Hedw.: bor-temp – Starac, 
2100 m; Vragonos above river Treskavička reka, 
1 00 m. 

Blindia acuta (Hedw.) B., S. & g.: bor-mont – Starac, 
2100 m. 
*Brachythecium albicans 
(Hedw.) B., S. & g.: bortemp – Vizitor, 1900 m. 
*Brachythecium erythrorhizon 
B., S. & g.: subarkt-subalp – at lake Ridsko jezero, 2000 m. 


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Brachythecium geheebii 
Milde: temp-subkont 
– valley grbaja, 1200 m; Belić, 1700 m; karanfil, 
1500 m. 

*Brachythecium glaciale B., S. & g.: subarkt-subalp 

– Planinica, 2000 m; Žljeb, 1500 m; Trojan, 
1900 m; 
Brachythecium glareosum 
(Spruce) B., S. & g.: bortemp 
– Žljeb, 2000 m; Vizitor, 1500 m; gorge 
Rugovska klisura near Peć, 500 m; valley of river Treskavička reka, 1500 m; karanfil 1450 m; 
Belić, 2 00 m; Djeravica; valley grbaja, 1200 m; 
Hajla: Dramodol, 2100 m. 

*Brachythecium 
mildeanum 
(Schimp.) Schimp. ex 
Milde: temp 
– Vizitor, 1900 m; valley grbaja, 
1200 m; Vragonos above river Treskavička reka, 
1500 m; Pasji vrh, 2100 m; at lake Ridsko jezero, 
2000 m; Belić, 1650 m. 

Brachythecium oedipodium 
(Mitt.) Jaeg.: temp 
– Planinica, 2000 m; Pasji vrh, 2100 m. 
Brachythecium plumosum 
(Hedw.) B., S. & g.: bortemp – Treskavica under V. Rid, 1700 m. 
Brachythecium populeum 
(Hedw.) B., S. & g.: temp 

– Planinica, 2000 m; valley of river Treskavička 
reka, 1400 m. 
Brachythecium reflexum 
(Starke) B., S. & g.: bor-mont 

– Boge, 1600 m; Hajla: kota 2278; karanfil, 
2050 m. 
Brachythecium 
rivulare 
B., S. & g.: bor-temp 
– Vizitor, 
1700 m; Starac, 2100 m; gorge Rugovska klisura 
near Peć, 500 m; springs Alipašini izvori near 
gusinje, 1000 m; valley grbaja, 1200 m; Maja 
kurvala, 1800 m; valley Ropojana: at Savina voda, 
1200 m; Babina gora, 1600 m; Vizitor, 1900 m. 

Brachythecium rutabulum 
(Hedw.) B., S. & g.: temp 

– valley of river Bogska reka, 1100 m; valley 
grbaja, 1200 m; Belić, 1650 m. 
Brachythecium salebrosum 
(Web. & Mohr) B., S. & 
g.: subbor 
– Boge 1700 m; Žljeb, 1500 m; Vizitor, 
1000 m, 1500 m; Planinica, 1900 m; Popadija, 
1900 m; Djeravica, 1900 m; karanfil, 2000 m; 
Belić, 1900 m; Žljeb, 1500 m; pass Čakor, 
1500 m (Szepesfalvy 1926). 

Brachythecium starkei 
(Brid.) B., S. & g.: bor-mont 
– 
Vizitor, 1750 m; Starac, 2100 m; Belić, 2 00 m. 
Brachythecium trachypodium 
(Brid.) B., S. & g.: subarkt-subalp – Starac, 2 00 m. 
Brachythecium velutinum 
(Hedw.) B., S. & g.: temp 

– valley of river Bogska reka, 1200 m; Vizitor, 1700 m; Planinica, 1900 m; valley grbaja, 
1200 m; Treskavica under V. Rid, 1800 m; valley 
of river Ločanska Bistrica, 1 00 m; valley Ropojana, 1200 m; Belić, 1400 m, 1900 m; Volušnica, 
1600 m; Babina gora, 1600 m. 
Bryoerythrophyllum recurvirostrum 
(Hedw.) Chen: 
bor-temp 
– valley of river Bogska reka, 1100 m; 
Hajla: Dramodol, 2100 m; Žljeb, 1500 m, 
2000 m; Planinica, 2000 m; Malje nedjinat, 
1900 m; gorge Rugovska klisura near Peć, 
500 m; springs Alipašini izvori near gusinje, 
1000 m; valley of river Treskavička reka, 1400 m; 
karanfil, 2000 m. 

Bryum alpinum 
With.: temp 
– Vizitor, 1900 m; Starac, 
2 00 m; Malje nedjinat, 1900 m. 

Bryum argenteum 
Hedw.: ubikv. – pass Čakor, 1500 m 
(szePesfalvy 1926) 

*Bryum blindii 
B., S. & g.: temp 
– Babina gora, 
1600 m; Starac, 2100 m; valley grbaja, 1200 m. 

Bryum caespiticium 
Hedw.: ubikv 
– valley of river 
Bogska reka, 1100 m; Devojački krš, 1800 m; 
Žuti kamen, 1700 m; Popadija, 1800 m; Pasji 
vrh, 2100 m. 

Bryum capillare 
Hedw.: ubikv 
– Vizitor, 1800 m; 
Babina gora, 1600 m; Devojački krš, 1800 m; 
valley Ropojana, 1200 m; karanfil, 2000 m. 

Bryum creberrimum 
Tayl.: bor-temp 
– Hajla: Škreli 
above Boge, 1800 m; valley of river Bogska reka, 
1100 m; Babina gora, 1600 m; Devojački krš, 
1800 m; Malje nedjinat, 1900 m; valley grbaja, 
1200 m; valley of river Treskavička reka, 1400 m; 
at lake Ridsko jezero, 2000 m. 

Bryum funckii 
Schwaegr.: temp 
– Devojački krš, 
1800 m. 

*Bryum imbricatum 
(Schwaegr.) B. & S.: temp 
– Hajla: kota 2278; Planinica, 2000 m; Starac, 1900 m; 
Devojački krš, 1800 m; Belić, 2 00 m. 

Bryum intermedium 
(Brid.) Bland.: subbor 
– Hajla: 
Dramodol, 2100 m. 

*Bryum 
muehlenbeckii 
B., S. & g.: bor-mont 
– Vizitor, 
1900 m; Belić, 2200 m. 

Bryum pallescens 
Schleich. ex Schwaegr.: ubikv 

– Babina gora, 1600 m; Devojački krš, 1800 m; 
valley grbaja, 1200 m; Popadija, 1800 m; along 
river Ločanska Bistrica, 1 00 m. 
Bryum pseudotriquetrum 
(Hedw.) gaertn., Meyer & 
Scherb. var. pseudotriquetrum: ubikv 
– Vaganica, 
2000 m; Vizitor, 1900 m; Babina gora, 1600 m; 
Devojački krš, 1800 m; Starac, 2100 m; Popadija, 1700 m; springs Alipašini izvori near gusinje, 1000 m; valley of river Treskavička reka, 
1400 m; Treskavica, under Veliki Rid, 1600 m; 
Djeravica, 2100 m; Maja kurvala, 1800 m; Pasji 
vrh, 2 00 m; Belić, 1900 m. 

*var. bimum 
(Brid.) Hartm.: temp 
– Malje nedjinat, 
1800 m; Starac, 2100 m; springs Alipašini izvori 
near gusinje, 1000 m; valley of river Treskavička 
reka, 1400 m. 


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*Bryum sauteri 
B., S. & g.: temp 
– Djeravica, 2 00 m; 
valley of river Treskavička reka, 1400 m. 

Bryum schleicheri 
Lam. & DC.: bor-mont 
– Vizitor, 
1900 m; Starac, 2100 m; Vragonos above river 
Treskavička reka, 1500 m; Djeravica, 2200 m; 
Treskavica under Veliki Rid, 1800 m. 

var. latifolium 
(Schwaegr.) Schimp.: bor-mont 
– Babina gora, 1600 m; Devojački krš, 1800 m; Starac, 2100 m; springs Alipašini izvori near gusinje, 1000 m; Djeravica, 2000 m; Maja kurvala, 
1800 m. 

Bryum turbinatum 
(Hedw.) Turn.: ubikv 
– Pasji vrh, 
2200 m. 

Bryum weigelii Spreng.: bor-mont – Starac, 2200 m. 

Buxbaumia viridis 
(Moug. ex Lam.& DC.) Brid. ex 
Moug. & nestl.: bor-mont 
– at river Ločanska Bistrica, 1 00 m. 

*Calliergon stramineum 
(Brid.) kindb.: bor-mont 
– 
Vizitor, 1750 m; Starac, 2100 m; at lake Ridsko 
jezero, 2000 m. 

var. patens (Lindb.) Broth. – Vizitor, 1750 m. 

Calliergonella cuspidata 
(Hedw.) Loeske: temp 

– Hajla: Dramodol, 2100 m; Vizitor, 1500 m; 
Malje nedjinat, 1700 m; gorge Rugovska klisura 
near Peć, 500 m; Treskavica under Veliki Rid, 
1600 m; springs Alipašini izvori near gusinje, 
1000 m; at lake Ridsko jezero, 2000 m; Starac, 
2100 m. 
Campyliadelphus chrysophyllus 
(Brid.) kanda: ubikv 

– Popadija, 1900 m. 
*Campyliadelphus elodes 
(Lindb.) kanda: temp 
– Vizitor, 1600 m; gorge Rugovska klisura near Peć, 
500 m. 

Campylium stellatum 
(Hedw.) Lange & C. Jens.: bortemp 
– Hajla, 2000 m; Vizitor, 1900 m; Malje 
nedjinat, 1700 m; Popadija, 1800 m; Pasji vrh, 
2100 m; karanfil, 2000 m; Belić, 2000 m. 

var. protensum 
(Brid.) Bryhn ex grout: bor-temp 

– Starac, 2100 m. 
Campylophyllum calcareum 
(Crundw. & nyholm) 
Hedenäs: temp 
– valley of river Dečanska Bistrica, 1100 m. 

*Campylophyllum halleri 
(Hedw.) Fleisch.: bor-mont 

– Boge: at alpine house, 1700 m; Devojački krš, 
1900 m; Belić, 1650 m. 
Ceratodon purpureus 
(Hedw.) Brid.: ubikv 
– valley 
of river Bogska reka, 1 00 m; Hajla: Dramodol, 2100 m; Babina gora, 1600 m; Devojački 
krš, 1900 m; Malje nedjinat, 1800 m; Starac, 
2100 m; valley grbaja, 1000 m; Trojan, 2000 m; 
Vragonos above river Treskavička reka, 1500 m; 
at river Ločanska Bistrica, 1 00 m; Maja kurvala, 1800 m; Žljeb, 1700 m. 

Cinclidotus aquaticus 
(Hedw.) B. & S.: temp 
– gorge 
Rugovska klisura near Peć, 500 m. 

Cinclidotus fontinaloides 
(Hedw.) P. Beauv.: meridtemp 
– valley grbaja, 1200 m; valley Ropojana: 
near Savina voda, 1200 m. 

Cirriphyllum piliferum 
(Hedw.) grout: bor-temp 

– Planinica, 2000 m; valley grbaja, 1200 m; valley Ropojana: at Savina voda, 1200 m; karanfil, 
1450 m. 
Cirriphyllum tommasinii 
(Sendtn. ex Boul.) grout: 
temp-subkont 
– valley of river Bogska reka, 
1 00 m; Vizitor, 1900 m; Popadija, 1800 m; valley of river Treskavička reka, 1400 m; karanfil, 
2000 m; Belić, 1650 m, 1900 m; valley grbaja, 
1200 m. 

Climacium dendroides 
(Hedw.) Web. & Mohr: subbor 

– springs Alipašini izvori near gusinje, 1000 m. 
Cratoneuron filicinum (Hedw.) Spruce: ubikv 
var. filicinum 
– valley of Bogska reka, 1 00 m; Vizitor, 
1900 m; Devojački krš, 1900 m; springs Alipašini 
izvori near gusinje, 1000 m; gorge Rugovska 
klisura near Peć, 500 m; Volušnica, 1600 m; 
Trojan, 1900 m; valley of river Dečanska Bistrica, 1100 m; karanfil, 1600 m; Belić, 2000 m; 
Peklen (szePesfalvy 1926 – 87, Dn42). 

var. atrovirens 
(Brid.) ochyra – Vizitor, 1000– 
1500 m; valley of river Ločanska Bistrica, 1 00 m; 
valley Ropojana: at Savina voda, 1200 m. 

Ctenidium molluscum (Hedw.) Mitt.: bor-temp 
– valley 
of river Bogska reka, 1 00 m; Hajla: kota 2278; 
Vizitor, 1900 m; Planinica, 2000 m; gorge Rugovska klisura near Peć, 500 m; springs Alipašini 
izvori near gusinje, 1000 m; valley grbaja, 
1200 m; Popadija, 1800 m; Pasji vrh, 1800 m; 
valley of river Ločanska Bistrica, 1 00 m; valley Ropojana, 1200 m; karanfil, 1450 m; Belić, 
2 00 m; Žljeb, 1700 m. 

Cynodontium polycarpon 
(Hedw.) Schimp.: bor-mont 

– Starac, 2100 m. 
Desmatodon heimii 
(Hedw.) Mitt.: bor-temp 
– Dečani 
near Peć (szePesfalvy 1926 – 487, Dn41) 
Desmatodon latifolius 
(Hedw.) Brid.: subarkt-subalp 
– Boge, 1700 m; Hajla: kota 2279, 1900 m; 
Devojački krš, 1900 m; Starac, 2100 m; Žuti kamen, 2400 m; Marjaš, 2 00 m; Malje nedjinat, 
2200 m; Djeravica, 2500 m. 

Dichodontium pellucidum 
(Hedw.) Schimp. bor-mont 

– Hajla: kota 2278; Žljeb, 2000 m; Babina gora, 
1700 m; valley of Treskavička reka, 1500 m; at 
lake Ridsko jezero, 2000 m; karanfil, 2000 m; 
Belić, 2 00 m. 
*Dicranella grevilleana 
(Brid.) Schimp.: bor-mont 
– 
Trojan, 2000 m; at lake Ridsko jezero, 2000 m. 


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*Dicranella humilis 
Ruthe: bor-mont 
– Devojački krš, 
2000 m. 

Dicranoweisia crispula 
(Hedw.) Milde: bor-mont 

– Babina gora, 1700 m; Starac, 2400 m; Pasji 
vrh, 2200 m; Djeravica, 2500 m; Maja kurvala, 
1700 m; Popadija, 1800 m; SE from Plav along 
road to Dečani, 1800 m (BauMgaRtneR 1915). 
Dicranum fuscescens 
Sm.: bor-mont 
– Starac, 2200 m; 
Pasji vrh, 2 00 m; Djeravica, 2 00 m; at lake 
Ridsko jezero, 2000 m. 

Dicranum 
muehlenbeckii 
B., S. & g.: bor-mont 
– Pasji 
vrh, 2 00 m. 

Dicranum polysetum 
Sw.: bor-mont 
– Babina gora, 
1600 m; Devojački krš, 2000 m; Malje nedjinat: 
kota 2 41; Planinica, 1900 m; Starac 2100 m; 
Pasji vrh, 2 00 m; valley of river Ločanska Bistrica, 1 00 m; Maja kurvala, 1700 m. 

Dicranum scoparium 
Hedw.: subbor 
– Hajla: Škreli 
above Boge, 1800 m; valley of river Bogska reka, 
1 00 m; Vizitor, 1900 m; Starac, 2100 m; valley grbaja, 1200 m; karanfil, 1400 m; Trojan, 
1900 m; valley Ropojana, 1200 m; Vragonos 
above river Treskavička reka, 1500 m; Pasji vrh, 
2 00 m; at lake Ridsko jezero, 2000 m; Belić, 
2200 m; pass Čakor (szePesfalvy 1926); near 
Peć (szePesfalvy 1926 – 87, Dn42); Dečani 
near Peć (szePesfalvy 1926 – 487, Dn41); Maja 
Rusolija, 1900–2200 m (RuDski 1949 – 287, 
Dn4 ). 

Dicranum tauricum 
Sap.: temp 
-Vizitor, 1900 m; 
Babina gora, 1600 m; Planinica, 1900 m; Starac 2100 m; Malje nedjinat, 1700 m; Treskavica 
above Veliki Rid, 1600 m; at lake Ridsko jezero, 
2000 m; along river Ločanska Bistrica, 1 00 m; 
Maja kurvala, 1700 m. 

Didymodon acutus 
(Brid.) k. Saito: merid-temp 
– Vizitor, 1900 m; along river Dečanska Bistrica, 
1200 m; Belić, 2 00 m. 

Didymodon fallax (Hedw.) Zander: ubikv 
– valley of 
river Bogska reka, 1 00 m; Hajla: Dramodol, 
2100 m; Devojački krš, 1900 m; gorge Rugovska 
klisura near Peć, 500 m; Belić, 2000 m. 

Didymodon icmadophilus 
(Schimp.) k. Saito: subarktsubalp – Hajla: Dramodol, 2100 m. 
Distichium capillaceum 
(Hedw.) B., S. & g.: bor-mont 

– Boge, 1700 m; Hajla: kota 19 ; Planinica, 
2000 m; Devojački krš, 2000 m; Starac, 2 00 m; 
Malje nedjinat, 1900 m; gorge Rugovska klisura 
near Peć, 500 m; Popadija, 1800 m; karanfil, 
1450 m, 2000 m; valley Ropojana, 1200 m; Stubica, 1400 m (RuDski 1949 – 287, Dn4 ); Maja 
Rusolija, 1900–2200 m (RuDski 1949 – 287, 
Dn4 ). 
Distichium inclinatum 
(Hedw.) B., S. & g.: subarktsubalp – Belić, 2 00 m. 

*Ditrichum crispatissimum (C. Müll.) Par.: bor-temp – 
valley of river Bogska reka, 1200 m; gorge Rugovska klisura near Peć, 500 m; Belić, 1650 m; 
valley Ropojana: near Savina voda, 1200 m; Popadija, 1800 m; Djeravica. 

Ditrichum 
flexicaule 
(Hedw.) Hampe: bor-temp 
– 
Boge, 1700 m; Hajla: kota 2278; Žljeb, 2000 m; 
Vizitor, 1700 m; Devojački krš, 1900 m; Starac, 
2100 m; Malje nedjinat, 1900 m; Žuti kamen, 
1800 m; gorge Rugovska klisura near Peć, 500 m; 
springs Alipašini izvori near gusinje, 1000 m; 
valley grbaja, 1200 m; Popadija, 1800 m; Trojan, 1900 m; Djeravica, 2400 m; valley Ropojana: near Savina voda, 1200 m; karanfil, 1450 m, 
1600 m; Belić, 1650 m, 2000 m; koprivnik, 
2000 m(szePesfalvy 1926 – 87, Dn 2). 

*Dryptodon patens(Hedw.) Brid.: bor-mont– Treskavica under Veliki Rid, 1700 m; at lake Ridsko jezero, 2000 m. 

Encalypta affinis 
Hedw. f.: arkt-alp 
– Žljeb, 2000 m; 
Devojački krš, 2000 m; Žuti kamen, 1900 m; 
Starac, 2100 m; Pasji vrh, 2100 m. 

Encalypta alpina 
Sm.: arkt-alp 
– Hajla: kota 2278; 
Planinica, 1900 m; Popadija, 1800 m; Belić, 
2 00 m. 

Encalypta ciliata 
Hedw.: bor-mont 
– Babina gora, 
1700 m; Marjaš, 1900 m; Vragonos above river 
Treskavička reka, 1500 m; valley Ropojana, 
1200 m; koprivnik above Peć, 2000 m (szePesfalvy 1926 – 87, Dn 2). 

Encalypta rhaptocarpa 
Schwaegr.: subarkt-subalp 

– Devojački krš, 2000 m; Marjaš, 2 00 m; Malje 
nedjinat, 1900 m. 
Encalypta streptocarpaHedw.: bor-temp– valley of river 
Bogska reka, 1 00 m; Hajla: Dramodol, 2100 m; 
Žljeb, 2000 m; Vizitor, 1500 m; Malje nedjinat, 
2100 m; Devojački krš, 2000 m; Marjaš, 2000 m; 
Starac, 2100 m; Planinica, 1900 m; springs 
Alipašini izvori near gusinje, 1000 m; karanfil, 
1600 m; Popadija, 1900 m; Pasji vrh 2200 m; valley Ropojana, 1200 m; Belić, 1650 m, 1900 m; 
Stubica, 1400 m (RuDski 1949 – 287, Dn4 ). 

Encalypta vulgaris 
Hedw.: merid-temp 
– Hajla: kota 
2278; valley of river Bogska reka near kučište, 
1100 m; Devojački krš, 1900 m; Starac, 2200 m; 
Malje nedjinat, 1900 m; karanfil, 2050 m; Belić, 
1900 m; Djeravica. 

Entodon concinnus 
(De not.) Par.: bor-mont 
– valley 
Ropojana, 1200 m. 
Entosthodon fascicularis 
(Hedw.) C. Müll.: temp 
– 
Žljeb, 1700 m. 


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Eucladium verticillatum 
(Brid.) B., S. & g.: meridtemp 
– at spring of river Istočka reka near 
Peć, 
500 m; gorge Rugovska klisura near Peć, 500 m; 
near gusinje, 1000 m; Belić, 1700 m; Peklen 
(szePesfalvy 1926 – 87, Dn42) 

*Eurhynchium angustirete 
(Broth.) T. kop.: tempsubkont 
– Hajla: Škreli above Boge, 1800 m; valley of river Bogska reka, 1 00 m; Babina gora, 
1600 m; Vragonos above river Treskavička reka, 
1500 m; Treskavica under V. Rid, 1600 m; karanfil, 1600 m. 

Eurhynchium crassinervium 
(Wils.) Schimp.: temp 

– Planinica, 1900 m; Belić, 2 00 m. 
Eurhynchium pulchellum 
(Hedw.) Jenn. var. pulchellum: subbor 
– valley of river Bogska reka, 1 00 m; 
Vragonos above river Treskavička reka, 1500 m; 
Žljeb, 1200 m. 

*var. diversifolium 
(B.,S.&g.)C.Jens.: subarkt-subalp 

– Boge, 1700 m; Vizitor, 1700 m; Devojački krš, 
1900 m; Planinica, 1900 m; Starac, 2100 m; valley grbaja, 1200 m; Trojan, 1900 m; valley Ropojana, 1200 m; Belić, 1900 m; Žljeb, 1700 m. 
Eurhynchium schleicheri 
(Hedw. f.) Jur.: merid-temp 

– valley grbaja, 1200 m; Vragonos above river 
Treskavička reka, 1500 m; valley Ropojana, 
1200 m; karanfil, 1450 m; Belić, 2000 m. 
Eurhynchium speciosum 
(Brid.) Jur.: temp 
– gorge 
Rugovska klisura near Peć, 500 m; valley of river 
Treskavička reka, 1400 m. 

Fissidens adianthoides 
Hedw.: bor-temp 
– near Peć 
(szePesfalvy 1926 – 87, Dn42) 

Fissidens dubius 
P. Beauv.: temp 
– Hajla, 2270 m; 
Planinica, 2000 m; Popadija, 1900 m; Volušnica, 
1600 m; gorge Rugovska klisura near Peć, 500 m; 
Pasji vrh, 1900 m; karanfil, 2000 m, 1450 m; valley Ropojana, 1200 m; Belić, 1700 m, 1900 m, 
2 00 m. 

Fissidens taxifolius 
Hedw. ssp. taxifolius: merid-temp 

– Vizitor, 1700 m; karanfil, 2000 m; at spring of 
Istočka reka near Peć, 600 m. 
Fontinalis antipyretica Hedw.: bor-temp 
– Starac, 
1800 m. 
Funaria hygrometrica 
Hedw.: ubikv 
– at spring of 
river Istočka reka near Peć, 600 m; gorge Rugovska klisura near Peć, 500 m; springs Alipašini 
izvori near gusinje, 1000 m; Vragonos above 
valley of river Treskavička reka, 1500 m; Žljeb, 
1700 m; pass Čakor, 1500 m (szePesfalvy 1926). 

Funaria muehlenbergii 
Turn.: merid-temp 
– at spring 
of river Istočka reka near Peć, 600 m; Hajla: kota 2278; Belić, 1800 m. 

Grimmia 
affinis Hornsch.: bor-temp 
– Djeravica, 
2 00 m. 

Grimmia alpestris (Web. & Mohr) Schleich. ex 
Hornsch.: bor-mont 
– Starac 2100 m; Pasji vrh 
1900 m; Treskavica under Veliki Rid 1600 m; 
Maja kurvala 1800 m. 

Grimmia anodon 
B. & S.: bor-temp 
– valley Ropojana: 
near Savina voda, 1200 m; Belić, 2000 m. 

Grimmia caespiticia 
(Brid.) Jur.: subarkt-subalp 
– Pasji vrh, 1700 m; Djeravica, 2 00 m. 

*Grimmia decipiens (k.F. Schultz)Lindb.: temp 
– 
Treskavica under Veliki Rid, 1700 m. 

Grimmia elongata kaulf.: subarkt-subalp 
– Starac, 
2400 m (Poelt 1976). 

Grimmia funalis 
(Schwaegr.) B. & S.: bor-mont 
– Starac, 2 00 m. 

Grimmia hartmanii 
Schimp. var. hartmanii: subbor 

– Babina gora; Maja kurvala 1800 m; SE from 
Plav along road to Dečani, 1600 m (BauMgaRtneR 1915). 
*Grimmia montana 
B. & S.: temp 
– Starac, 2200 m; 
Maja kurvala, 1800 m; Pasji vrh, 1700 m. 

Grimmia ovalis 
(Hedw.) Lindb.: subbor 
– valley of 
river Bogska reka 1 00 m; Djeravica. 

Grimmia pulvinata 
(Hedw.) Sm. var. pulvinata: 
merid-temp 
– at spring of river Istočka reka near 
Peć, 600 m; Vizitor, 1500 m; springs Alipašini 
izvori near gusinje, 1000 m; Dečani near Peć 
(szePesfalvy 1926 – 487, Dn41). 

Gymnostomum aeruginosum 
Sm.: subbor 
– Starac, 
1900 m; Vizitor, 1900 m. 

Gymnostomum calcareum 
nees & Hornsch.: meridtemp – gorge Rugovska klisura near Peć, 500 m. 

Herzogiella seligeri (Brid.) Iwats.: subtemp 
– valley of river Bogska reka, 1 00 m; Babina gora, 
1600 m; Treskavica under V. Rid; valley of river 
Ločanska Bistrica, 1 00 m. 

Heterocladium dimorphum 
(Brid.) B., S. & g.: bormont 
– Vizitor, 1900 m; Planinica, 2000 m; Starac, 2100 m; Malje nedjinat, 1800 m; Volušnica, 
1600 m; Popadija, 1800 m; Pasji vrh, 1900 m. 

*Homalia besseri 
Lob.: temp-subkont 
– at spring of 
river Istočka reka near Peć, 600 m; valley Ropojana, 1200 m. 

Homalothecium lutescens 
(Hedw.) Robins.: merid-temp 

– valley grbaja, 1200 m. 
Homalothecium philippeanum 
(Spruce) B., S. & 
g.: temp-subkont 
– valley of river Bogska reka 
at kučište, 1100 m; Hajla: Dramodol, 2100 m; 
Vizitor, 1700 m; Devojački krš, 1900 m; Malje 
nedjinat, 1700 m; valley grbaja, 1200 m; Volušnica, 1600 m; Djeravica, 1900 m; valley of 
river Dečanska Bistrica, 1100 m; valley Ropojana, 1200 m; karanfil, 1450 m, 2000 m; Belić, 
1650 m, 2 00 m; Žljeb, 1700 m. 


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Homalothecium sericeum 
(Hedw.) B., S. & g.: meridtemp 
– Planinica, 1900 m; springs Alipašini izvori 
near gusinje, 1000 m; Djeravica, 2000 m; valley 
Ropojana, 1200 m; Devojački krš, 2000 m; near 
Peč (szePesfalvy 1926 – 87, Dn42); Dečani 
near Peć (szePesfalvy 1926 – 487, Dn41); pass 
Čakor (szePesfalvy 1926). 

Hygroamblystegium tenax 
(Hedw.) Jenn.: temp 
– 
gorge Rugovska klisura near Peć, 500 m. 

*Hygrohypnum duriusculum 
(De not.) Jamieson: 
subarkt-subalp 
– Starac, 2100 m; at lake Ridsko 
jezero, 2000 m. 

Hygrohypnum luridum 
(Hedw.) Jenn.: bor-temp 

– Babina gora, 1600 m; gorge Rugovska klisura 
near Peć, 500 m. 
Hylocomium splendens (Hedw.) B., S. & g.: subbor 

– valley of river Bogska reka, 1 00 m; Babina gora, 1600 m; Planinica, 2000 m; Starac, 2100 m; 
valley grbaja, 1200 m; Vragonos above river 
Treskavička reka, 1500 m; Treskavica under 
V. Rid, 1700 m; Djeravica, 2000 m; Pasji vrh, 
2000 m; Maja kurvala, 1800 m; valley Ropojana, 
1200 m; Belić, 1800 m. 
Hypnum callichroum 
Brid.: subarkt-subalp 
– Starac, 
2100 m. 

Hypnum cupressiforme 
Hedw.: ubikv 
– at spring of 
river Istočka reka near Peć, 600 m; valley of 
river Bogska reka, 1 00 m; Hajla: Dramodol, 
2100 m; Vizitor, 1400 m; Babina gora, 1600 m; 
Vragonos above river Treskavička reka, 1500 m; 
Treskavica under V. Rid, 1700 m; Belić, 2000 m; 
pass Čakor, 1500 m (szePesfalvy 1926). 

Hypnum jutlandicum 
Holmen & Warncke: temp-subatl 

– at spring of river Istočka reka near Peć, 600 m; 
near Peć (szePesfalvy 1926 – 87, Dn42). 
Hypnum pallescens 
(Hedw.) P. Beauv.: temp-subkont 
– 
Boge: at alpine house, 1700 m; Marjaš, 2100 m. 
Hypnum recurvatum 
(Lindb. & H. Arnell) kindb.: 
subarkt-subalp 
– Hajla: kota 2278; Malje nedjinat, 1800 m; Volušnica, 1600 m. 

Hypnum revolutum 
(Mitt.) Lindb.: subarkt-subalp 
– 
Žuti kamen, 1900 m; between Marjaš and Pasji 
vrh; Djeravica, 2400 m; Belić, 2 00 m. 

Hypnum vaucheri 
Lesq.: bor.mont 
– Hajla: kota 
19 , 1900 m; Malje nedjinat, 1800 m; Belić, 
2 00 m. 

Isopterygiopsis pulchella 
(Schimp.) Iwats.: bor-mont 
– 
Babina gora, 1600 m; Starac, 1900 m; Planinica, 
1900 m; Volušnica, 1600 m; Djeravica, 2000 m; 
Treskavica under V. Rid, 1800 m. 

Isothecium alopecuroides 
(Dubois) Isov.: bor-temp 

– Babina gora, 1600 m; Vizitor, 1900 m; Starac, 
1900 m; Volušnica, 1600 m; Trojan, 1900 m; 
Vragonos above river Treskavička reka, 1500 m; 
at lake Ridsko jezero, 2000 m; along river 
Ločanska Bistrica, 1 00 m; valley Ropojana, 
1200 m; Belić, 1800 m; near Peć (szePesfalvy 
1926 – 87, Dn42). 

Isothecium myosuroides 
Brid.: bor-temp 
– valley of river 
Treskavička reka, 1400 m. 

Kiaeria falcata 
(Hedw.) I. Hagen: arkt-alp 
– at lake 
Ridsko jezero, 2000 m. 

Leptodon smithii 
(Hedw.) Web. & Mohr: atl-medit 
– 
at spring of river Istočka reka near Peć, 600 m. 

Lescuraea mutabilis 
(Brid.) Lindb. ex I. Hagen: tempsubkont 
– Starac, 1900 m; Djeravica, 2000 m; 
Treskavica under Veliki Rid, 1700 m. 

*Lescuraea saxicola 
(B., S. & g.) Milde: bor-mont 

– Starac, 2100 m; karanfil, 1600 m. 
Leskea polycarpa 
Hedw.: temp 
– valley of river 
Treskavička reka, 1400 m. 
Leucodon sciuroides 
(Hedw.) Schwaegr.: subtemp 
– at 
spring of river Istočka reka near Peć, 
600 m; 
valley of river Bogska reka, 1 00 m; Vizitor, 
1700 m; Babina gora, 1600 m; Marjaš, 1700 m; 
valley grbaja, 1200 m; Volušnica, 1600 m; 
Vragonos above river Treskavička reka, 1500 m; 
Treskavica under Veliki Rid, 1900 m; Belić, 
1650 m. 

var. morensis 
(Schwaegr.) De not.: subatl-submed 
– at 
spring of river Istočka reka near Peć, 600 m; valley of river Treskavička reka, 1400 m. 

Metaneckera menziesii 
(Hook.) Steere: submed 
– valley of river Bogska reka, 1 00 m; Starac, 1800 m; 
valley grbaja, 1200 m; valley Ropojana, 1200 m; 
Belić, 1650 m. 

Mnium ambiguum 
H. Müll.: bor-mont 
– Babina gora, 
1600 m; Starac, 2100 m; valley grbaja, 1200 m. 

Mnium marginatum 
(Dicks.) P. Beuv.: bor-temp 
– Hajla: Škreli above Boge, 1800 m; Vizitor, 1500 m; 
Djeravica, 2000 m; valley Ropojana, 1200 m; 
Belić, 1900 m. 

Mnium spinosum 
(Voit.) Schwaegr.: bor-mont 
– Hajla: Dramodol, 2100 m; Malje nedjinat, 2200 m; 
Marjaš, 2 00 m. 

Mnium stellare 
Hedw.: bor-temp 
– Planinica, 1900 m; 
Marjaš, 2 00 m; Malje nedjinat, 1900 m; valley grbaja, 1200 m; Pasji vrh, 1800 m; valley of 
river Dečanska Bistrica, 1200 m; Maja kurvala, 
1800 m; karanfil, 2000 m; Hajla: kota 2278; 
Belić, 2 00 m. 

Mniumthomsonii 
Schimp.: bor-mont 
– Vizitor, 1900 m; 
Babina gora, 1600 m; Planinica, 2000 m; Starac, 2 00 m; Malje nedjinat, 1900 m; Popadija, 
1800 m; Volušnica, 1600 m; Trojan, 1900 m; 
Belić, 2 00 m; Žljeb, 1700 m. 


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Myurella julaceae (Schwaegr.) B., S. & g.: subarkt-subalp – Devojački krš, 1900 m; Pasji vrh 2100 m. 

Neckera complanata 
(Hedw.) Hüb.: bor-temp – valley of river Bogska reka, 1 00 m; Devojački 
krš, 1900 m; valley of river Dečanska Bistrica, 
1100 m; valley Ropojana, 1200 m. 

Neckera crispa 
Hedw.: temp – gorge Rugovska 
klisura near Peć, 500 m; near Peć (szePesfalvy 
1926 – 87, Dn42); Crveni krš, 900 m ( 288, 
Dn5 ). 

Oncophorus virens 
(Hedw.) Brid.: subarkt-subalp 

– Starac, 2100 m; Pasji vrh, 2400 m; at lake Ridsko jezero, 2000 m. 
*Orthothecium intricatum 
(Hartm.) B., S. & g.: bormont – Starac, 2100 m; karanfil, 1450 m; Belić, 
2000 m. 

Orthothecium rufescens 
(Brid.) B., S. & g.: bor-mont 

– gorge Rugovska klisura near Peć, 500 m. 
Orthotrichum affine 
Brid.: temp – Babina gora, 
1600 m. 
Orthotrichum 
alpestre 
Hornsch. ex B., S. & g.: subarkt-subalp – Vizitor, 1500 m, on Fagus moesiaca. 
Orthotrichum anomalum 
Hedw.: subtemp – Vizitor, 
1500 m; pass Čakor, 1500 m (szePesfalvy 1926) 
Orthotrichum cupulatum 
Brid.: temp – Starac, 1900 m; 
Malje nedjinat, 1800 m; valley Ropojana, 
1200 m; Vragonos above river Treskavička reka, 
1500 m; Vizitor, 1500 m. 

Orthotrichum diaphanum 
Brid.: merid-temp. - Peklen 
(szePesfalvy 1926 – 87, Dn42). 
Orthotrichum rupestre 
Schleich. ex Schwaegr.: bortemp – Starac, 2100 m. 
Orthotrichum stramineum 
Hornsch. ex Brid.: temp 

– Vizitor, 1500 m, on Fagus moesiaca. 
Palustriella commutata (Hedw.) ochyra 
var. commutata: bor-temp – Vaganica, 2000 m; Vizitor, 
1900 m; Devojački krš, 1900 m; Starac, 2100 m; 
Malje nedjinat, 1800 m, 1000 m; at lake Ridsko 
jezero, 2000 m; valley of river Dečanska Bistrica, 
1100 m; Maja kurvala, 1800 m. 

var. falcata 
(Brid.) ochyra: bor-temp – Popadija, 
1700 m. 

*var. fluctuans 
(Bruch & al.) ochyra: bor-temp – 
gorge Rugovska klisura near Peć, 500 m; springs 
Alipašini izvori near gusinje, 1000 m. 

Palustriella decipiens 
(De not.) ochyra: subarktsubalp – Vaganica, 2000 m; Hajla: Dramodol, 
2100 m; Vizitor, 1800 m; Starac, 2100 m; Malje 
nedjinat, 1800 m; springs Alipašini izvori near 
gusinje, 1000 m; Popadija, 1700 m; Trojan, 
1900 m; Pasji vrh, 2200 m; Belić, 1900 m; Babino brdo, 1600 m, 

Paraleucobryum enerve 
(Thed.) Loeske: arkt-alp – 

Starac, 2 00 m; Djeravica, 2200 m; Babina gora, 
1600 m. 

Paraleucobryum sauteri 
(B., S. & g.) Loeske: tempsubkont – karanfil, 1450 m. 

Philonotis caespitosa 
Jur.: bor-temp – Vizitor, 1900 m; 
Djeravica, 2400 m. 

Philonotis calcarea 
(B. & S.) Schimp.: bor-temp – Vizitor, 1900 m; valley of river Bogska reka, 1 00 m; 
Malje nedjinat, 1800 m; gorge Rugovska klisura 
near Peć, 500 m; springs Alipašini izvori near 
gusinje, 1000 m; Popadija, 1700 m; valley of 
river Dečanska Bistrica, 1100 m. 

Philonotis fontana 
(Hedw.) Brid.: bor-temp – Vaganica, 2000 m; Babina gora, 1600 m; Devojački 
krš, 1900 m; Starac, 2100 m; Malje nedjinat, 
1800 m; Djeravica, 2 00 m; Pasji vrh, 2200 m; 
valley of river Treskavička reka, 1400 m; Maja 
kurvala, 1900 m. 

Philonotis marchica 
(Hedw.) Brid.: merid-temp 

– Treskavica under Veliki Rid, 1600 m; gorge 
Rugovska klisura near Peć, 500 m; Peklen 
(Szepesfalvy 1926 – 87, Dn42). 
Philonotis seriata 
Mitt.: subarkt-subalp – Vizitor, 
1900 m; Starac, 2100 m; Maja kurvala, 1800 m; 
Štedin (RuDski 19 6 – 287, Dn ). 

*Philonotis tomentella 
Mol.: ark-alp – Devojački krš, 
1900 m; Starac, 2100 m; Malje nedjinat, 1700 m; 
Popadija, 1700 m; valley of river Treskavička 
reka, 1400 m; Djeravica, 2000 m; Treskavica under Veliki Rid, 1600 m; karanfil, 1600 m; Belić, 
1900 m. 

Plagiobryum zierii 
(Hedw.) Lindb.: subarkt-subalp 

– Žljeb, 2000 m. 
Plagiomnium affine 
(Bland.) T. kop.: temp – Starac, 
1900 m; Vragonos above river Treskavička reka, 
1500 m; Treskavica under Veliki Rid, 1700 m; 
valley of river Ločanska Bistrica, 1 00 m. 

Plagiomnium cuspidatum 
(Hedw.) T. kop.: bortemp – Vizitor, 1500 m; springs Alipašini izvori 
near gusinje, 1000 m; valley grbaja, 1200 m; 
Vragonos above river Treskavička reka, 1500 m; 
valley Ropojana: near Savina voda, 1 00 m; karanfil, 1600 m. 

Plagiomnium elatum 
(B. & S.) T. kop.: bor-temp -
Malje nedjinat, 1700 m; Vragonos above river 
Treskavička reka, 1500 m. 

Plagiomnium medium 
(B. & S.) T. kop.: bor-mont – 
Planinica, 2000 m; Babina gora, 1600 m; valley 
grbaja, 1200 m. 

Plagiomnium rostratum 
(Schrad.) T. kop.: bor-temp 

– Vaganica, 2000 m; valley of river Bogska reka, 
1 00 m; Malje nedjinat, 1800 m; gorge Rugovska klisura near Peć, 500 m; Trojan, 1900 m; 
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valley grbaja, 1200 m; valley Ropojana, 1200 m; 
karanfil, 1450 m; Belić, 1400 m, 1800 m; Žljeb, 
1700 m. 

Plagiomnium undulatum 
(Hedw.) T. kop.: temp 

– valley of river Bogska reka, 1 00 m; springs 
Alipašini izvori near gusinje, 1000 m; valley 
Ropojana, 1200 m. 
Plagiopus oederiana 
(Sw.) Crum & Anders.: bor-mont 

– Vizitor, 1900 m; valley of river Bogska reka 
near kučište, 1100 m; gorge Rugovska klisura 
near Peć, 500 m; valley of river Treskavička 
reka, 1400 m; Maja kurvala, 1800 m; Belić, 
1400 m, 1900; near Peć (szePesfalvy 1926 – 
87, Dn42). 
var. alpina (Schwaerg.) Möller – Marjaš, 2000 m. 
Plagiothecium cavifolium 
(Brid.) Iwats.: bor-mont 

– Vizitor, 1750 m; Babina gora, 1600 m. 
Plagiothecium curvifolium 
Schlieph. ex Limpr.: temp 
– Žljeb, 1500 m. 
Plagiothecium denticulatum 
(Hedw.) B., S. & g.: bortemp – Vizitor, 1900 m; Babina gora, 1600 m; 
Planinica, 1900 m; Starac, 2100 m; Treskavica 
under V. Rid, 1800 m; Djeravica, 1800 m. 

*var. obtusifolium(Turn.) Moore – Planinica, 1900 m; 
at lake Ridsko jezero, 2000 m. 

Plagiothecium laetum 
B., S. & g.: bor-mont – Babina 
gora, 1600 m; Planinica, 2000 m; Maja kurvala, 
1800 m; Žljeb, 1500 m. 

Plagiothecium nemorale (Mitt.) Jaeg.: temp – Vizitor, 
1500-1700 m. 

Plagiothecium platyphyllum 
Mönkm.: bor-mont – Starac, 2100 m (rev. Pilous) 

Plagiothecium ruthei 
Limpr.: bor-temp – Planinica, 
1900 m. 

*Platydictya jungermannioides 
(Brid.) Crum: subarktsubalp – Vizitor, 1700 m. 

Pleurozium schreberi 
(Brid.) Mitt.: bor-temp – Starac, 
2100 m; Marjaš, 2100 m; valley of river Ločanska 
Bistrica, 1 00 m. 

Pogonatum aloides 
(Hedw.) P. Beauv.: bor-temp 

– Babina gora, 1600 m; pass Čakor, 1500 m 
(Szepesfalvy 1926). 
Pogonatum urnigerum 
(Hedw.) P. Beauv.: bor-mont – 
Babina gora 1700 m; Devojački krš, 2000 m; Starac, 2100 m; Vragonos above river Treskavička 
reka, 1500 m; Maja kurvala, 1800 m; Belić, 
1600 m. 

Pohlia cruda 
(Hedw.) Lindb.: bor-temp – Žljeb, 
2000 m; Vizitor, 1800 m; Babina gora, 1600 m; 
Devojački krš, 1800m; Planinica, 1900 m; Starac, 
2 00 m; between Marjaš and Pasji vrh, 2100 m; 
valley grbaja, 1200 m; Popadija, 1800 m; Djeravica, 2100 m; valley of river Treskavička reka, 

1400 m; Maja kurvala, 1800 m; valley Ropojana, 
1200 m; karanfil, 2000 m; Belić, 1650 m. 

Pohlia elongata 
Hedw. var. elongata: bor-mont – Djeravica, 2500 m. 

Pohlia nutans 
(Hedw.) Lindb.: subbor – Devojački 
krš, 1800 m; Starac, 2000 m; Pasji vrh, 1900 m; 
at lake Ridsko jezero, 2000 m. 

Polytrichum alpinum 
Hedw.: subarkt-subalp – Vizitor, 1700 m; Babina gora, 1700 m; Devojački 
krš, 2000 m; Starac, 2100 m; Planinica, 1900 m; 
Marjaš, 2000 m; at lake Ridsko jezero, 2000 m; 
Djeravica, 2 00 m; Maja kurvala, 1800 m; Belić, 
2000 m; Žljeb, 1700 m.. 

Polytrichum commune 
Hedw. var. commune: subbor 

– Starac, 2100 m; at lake Ridsko jezero, 2000 m; 
Djeravica, 2200 m; Stubica, 1400 m (RuDski 
1949 – 287, Dn4 ). 
Polytrichum formosum 
Hedw.: bor-temp – Hajla, 
2270 m; Vizitor, 1700 m; Babina gora, 1700 m; 
Planinica, 2000 m; Popadija, 1800 m; Vragonos 
above river Treskavička reka, 1500 m; Treskavica under Veliki Rid, 1900 m; valley of river 
Treskavička reka, 1400 m; karanfil, 1600 m. 

Polytrichum juniperinum 
Hedw.: bor-temp – Boge, 
1700 m; Hajla: Dramodol, 2100 m; Hajla 
2270 m; Vizitor, 1900 m; Babina gora, 1700 m; 
Malje nedjinat, 2200 m; Starac, 2400 m; valley grbaja, 1200 m; Popadija, 1900 m; Trojan, 
2000 m; karanfil 2000 m; Djeravica, 2500 m; 
Belić, 2200 m. 

*Polytrichum longisetum 
Sw. ex Brid.: bor-temp – Vizitor, 1900 m. 

Polytrichum piliferum 
Hedw.: bor-temp – Devojački 
krš, 2000 m; Starac, 2 00 m; near gusinje, 
1000 m; Pasji vrh, 2400 m. 

Pseudoleskea incurvata 
(Hedw.) Loeske: bor-mont – 
Hajla: kota 2278, 1900 m; Boge, 1700 m; valley 
of river Bogska reka, 1 00 m; Vizitor, 1900 m; 
Devojački krš, 1900 m; Planinica, 2000 m; Žuti 
kamen, 1700 m; Marjaš, 2000 m; Volušnica, 
1600 m; Popadija, 1800 m; Trojan, 1900 m; valley grbaja, 1200 m; Djeravica, 2200 m; Treskavica under Veliki Rid, 1900 m; Pasji vrh, 2000 m; 
valley Ropojana: near Savina voda, 1200 m; karanfil, 1450 m, 2000 m; Belić, 1650 m, 2000 m, 
2 00 m; Žljeb, 1700 m. 

*Pseudoleskea patens 
(Lindb.) kindb.: bor-mont – 
Hajla: Dramodol, 2100 m; Babina gora, 1600 m; 
Starac, 2000 m; at lake Ridsko jezero, 2000 m. 

*Pseudoleskea radicosa 
(Mitt.) Macoun & kindb. 
var. radicosa: subarkt-subalp – Vizitor, 1700 m; 
Planinica, 1900 m; Starac, 1900 m; valley of river Treskavička reka, 1400 m; Pasji vrh, 2000 m; 


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valley Ropojana, 1200 m; karanfil, 1600 m, 
2000 m; Žljeb, 1500 m; Djeravica, 2000 m. 

Pseudoleskea saviana 
(De not.) Latz.: southalp-balk 

– Vizitor, 1750 m; Volušnica, 1600 m; valley of 
river Treskavička reka, 1400 m; Treskavica under Veliki Rid, 1800 m; Maja kurvala, 1700 m; 
karanfil, 1600 m; Belić, 1900 m. 
Pseudoleskeella catenulata 
(Schrad.) kindb.: bor-mont 

– Hajla: Dramodol, 2100 m; Vizitor, 1900 m; 
Žuti kamen, 1800 m; Trojan, 1900 m; valley 
grbaja, 1200 m. 
Pseudoleskeella nervosa 
(Brid.) nyh.: bor-mont 
– valley of river Bogska reka, 1 00 m; Boge, 1600 m; 
Vizitor, 1700 m; valley Ropojana: near Savina 
voda, 1200 m; valley of river Ločanska Bistrica, 
1 00 m; Belić, 2100 m; Volušnica, 1600 m. 

Pterigynandrum filiforme 
Hedw.: bor-mont 
– valley of 
river Bogska reka near kučište, 1100 m; Hajla: 
Dramodol, 2100 m; Vizitor, 1700 m; Babina 
gora, 1600 m; Malje nedjinat, 1800 m; between 
Marjaš and Pasji vrh; Starac, 2100 m; valley grbaja, 1200 m; Popadija, 1800 m; Volušnica, 1600 m; 
Vragonos above river Treskavička reka, 1500 m; 
Djeravica, 2100 m; valley of river Ločanska 
Bistrica, 1 00 m; Maja kurvala, 1800 m; valley 
Ropojana, 1200 m; Belić, 1650 m. 

Pterygoneurum ovatum 
(Hedw.) Dix.: merid-temp 

– Planinica, 1900 m. 
Ptychodium plicatum 
(Web. & Mohr) Schimp.: subarkt-subalp 
– Boge, 1700 m; Hajla: kota 2278; 
Žljeb, 2000 m; Planinica, 1900 m; valley grbaja, 
1200 m; Popadija, 1800 m; karanfil, 1450 m, 
2000 m; Belić, 1900 m, 2 00 m. 

Racomitrium aciculare 
(Hedw.) Brid.: bor-temp 
– 
Babina gora, 1700 m; Starac, 2100 m. 
Racomitrium canescens 
(Hedw.) Brid.: bor-mont 

– Treskavica under Veliki Rid, 1700 m; Popadija, 1800 m; Trojan, 1900 m; karanfil, 1600 m; 
Belić, 1650 m. 
Racomitrium elongatum 
Frisvoll: bor-temp 
– Žljeb, 
1500 m; Treskavica under Veliki Rid, 1700 m; 
Trojan, 2000 m. 

Racomitrium ericoides 
(Brid.) Brid.: subbor 
– Trojan, 
2000 m; Belić, 1650 m. 
*Racomitrium heterostichum 
(Hedw.) Brid.: bor-temp 

– Hajla: Dramodol, 2100 m; Vizitor, 1900 m; 
Devojački krš, 1900 m; Starac, 2100 m; Marjaš, 
2400 m; Popadija, 1800 m; Treskavica under 
Veliki Rid, 1900 m; Djeravica, 2500 m; at lake 
Ridsko jezero, 2000 m; Maja kurvala, 1800 m. 
*Racomitrium sudeticum 
(Funck) B. & S.: subarkt-subalp 
– Vizitor, 1750 m; Starac, 2 00 m; Djeravica, 
2400 m; at lake Ridsko jezero, 2000 m. 

Rhabdoweisia fugax 
(Hedw.) B., S. & g.: bor-mont 

– Starac, 2 00 m. 
Rhizomnium pseudopunctatum 
(B. & S.) T. kop.: bor
mont – Starac, 2100 m; Djeravica, 
2400 m. 
Rhizomnium punctatum 
(Hedw.) T. kop.: bor-temp 

– Vaganica, 1900 m; Vizitor, 1900 m; Babina 
gora, 1600 m; Starac, 2100 m; Malje nedjinat, 
1800 m; Popadija, 1700 m; Vragonos above 
river Treskavička reka, 1500 m; valley of river 
Treskavička reka, 1400 m; Treskavica above Veliki Rid, 1700 m; Pasji vrh, 2000 m; pass Čakor, 
1500m (szePesfalvy 1926). 
*Rhodobryum roseum 
(Hedw.) Limpr.: bor-temp 
– Vizitor, 1900 m; Planinica, 2000 m. 

Rhynchostegiella tenella 
(Dicks.) Limpr. var. tenella: 
subatl-submed 
– Starac, 1800 m; valley grbaja, 
1200 m; Belić, 1700 m. 

Rhynchostegiella tenuicaulis 
(Spruce) kartt.: temp 

– Babina gora, 1600 m. 
Rhynchostegium murale 
(Hedw.) B., S. & g.: temp 
– valley Ropojana, 1200 m; karanfil, 2000 m. 
Rhynchostegium riparioides 
(Hedw.) Card.: temp 
– gorge Rugovska klisura near Peć, 500 m; 
springs Alipašini izvori near gusinje, 1000 m; 
valley Ropojana: Savina voda, 1200 m. 
Rhytidiadelphus loreus 
(Hedw.) Warnst.: bor-temp 
– 
Starac, 1900 m; at lake Ridsko jezero, 2000 m. 

Rhytidiadelphus squarrosus 
(Hedw.) Warnst.: bortemp 
– Vragonos above river Treskavička reka, 
1500 m; Belić, 1800 m. 

Rhytidiadelphus triquetrus 
(Hedw.) Warnst.: bortemp 
– Boge, 1700 m; Hajla: Škreli above Boge, 
1800 m; Vizitor, 1900 m; Babina gora, 1600 m; 
Starac, 2000 m; valley grbaja, 1200 m; Vragonos 
above river Treskavička reka, 1500 m; Popadija, 1800 m; valley Ropojana, 1200 m; karanfil, 
1600 m; Belić, 2 00 m. 

Rhytidium rugosum (Hedw.) kindb.: bor-temp 
– valley 
of river Bogska reka, 1 00 m; Marjaš, 1800 m; 
Djeravica, 1900 m. 

Saelania glaucescens 
(Hedw.) Broth.: subarkt-subalp 

– Popadija, 1800 m; Pasji vrh, 200 m; 
Sanionia uncinata 
(Hedw.) Loeske: bor-mont 
– Vizitor, 1750 m; Devojački krš, 1900 m; Starac, 
2100 m; Babina gora, 1600 m; Malje nedjinat, 1700 m; Volušnica, 1600 m; valley of river 
Treskavička reka, 1400 m; Djeravica, 1900 m; 
valley Ropojana, 1200 m; Maja kurvala, 1700 m; 
Belić, 1900 m; Žljeb, 1500 m; Maja Rusolija, 
1900–2200 m (Rudski 1949 – 287, Dn4 ). 

Schistidium apocarpum 
(Hedw.) B. & S.: bor-temp 

– at spring of river Istočka reka near Peć, 
600 m; Boge, 1600 m; Hajla: kota 19 , 1900 m; 
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Vizitor, 1100 m, 1700 m; Planinica, 1900 m; 
Devojački krš, 2000 m; Starac, 2000 m; Malje 
nedjinat, 1800 m; gorge Rugovska klisura near 
Peć, 500 m; springs Alipašini izvori near gusinje, 1000 m; valley grbaja, 1200 m; along river 
Ločanska Bistrica, 1200 m; valley Ropojana, 
1200 m; Belić, 1650 m, 2 00 m; pass Čakor, 
1500 m (szePesfalvy 1926). 

var. homodictyon 
(Dix.) Crundw. & nyholm – Hajla: 
kota 2278. 

Schistidium atrofuscum 
(Schimp.) Limpr.: bor-mont – 
Marjaš, 2200 m; Malje nedjinat, 2200 m; Belić, 
1700 m. 

Schistidium brunnescens 
Limpr.: temp – Hajla: kota 
2278; Planinica, 1900 m. 

Schistidium rivulare 
(Brid.) Podp. subsp. rivulare: 
bor-mont - Babina gora, 1600 m. 

Scleropodium purum 
(Hedw.) Limpr.: temp – Dečani 
near Peć (szePesfalvy 1926 – 487, Dn41); near 
Peć (szePesfalvy 1926 – 87, Dn42). 

*Scorpidium cossonii 
(Schimp.) Hedenäs: bor-mont 

– Vizitor, 1750 m; Starac, 2200 m. 
Seligeria pusilla 
(Hedw.) B., S. & g.: bor-temp – gorge 
Rugovska klisura near Peć, 500 m. 
*Sphagnum capillifolium 
(Ehrh.) Hedw.: bor-temp - at 
lake Ridsko jezero, 2000 m; Starac, 2100 m. 
*Sphagnum cuspidatum 
Ehrh. ex Hoffm.: bor-temp – 
at lake Ridsko jezero, 2000 m; Starac, 2100 m. 
*Sphagnum denticulatum 
Brid.: bor-temp – Vizitor, 
1750 m; Starac 2100 m. 
*Sphagnum fallax 
(klinggr.) klinggr.: bor-temp – at 
lake Ridsko jezero, 2000 m; Starac 2100 m; Vizitor, 1750 m. 

*Sphagnum girgensohnii 
Russ.: bor-mont – at lake Ridsko jezero, 2000 m. 

*Sphagnum magellanicum 
Brid.: bor-mont – Vizitor, 
1750 m. 

*Sphagnum palustre L.: bor-temp – Vizitor, 1750 m. 

*Sphagnum platyphyllum 
(Lindb. ex Braithw.) Sull. 
ex Warnst.: bor-mont – Starac, 2100 m. 

*Sphagnum teres 
(Schimp.) Aongstr.: bor-mont – at 
lake Ridsko jezero, 2000 m; Starac, 2100 m 

Syntrichia intermedia 
Brid.: merid-temp – at spring of 
river Istočka reka near Peć, 600 m; Malje nedjinat, 1600 m. 

Syntrichia norvegica 
Web.: subarkt-subalp – Boge, 
1600 m; Hajla: kota 2278; Vizitor, 1900 m; Malje 
nedjinat, 1900 m; Volušnica, 1600 m; Treskavica under Veliki Rid, 1600 m; karanfil, 1600 m, 
2000 m. 

Syntrichia ruralis 
(Hedw.) Web. & Mohr subsp. ruralis: bor-temp – Hajla: Dramodol, 2100 m; Hajla: 
Škreli above Boge, 1800 m; valley of river Bogs

ka reka, 1 00 m; Vizitor, 1700 m; Babina gora, 
1600 m; Planinica, 2000 m; Starac, 2100 m; 
Malje nedjinat, 1600 m; valley grbaja, 1200 m; 
Volušnica 1600 m; Popadija, 1900 m; Trojan, 
1900 m; springs Alipašini izvori near gusinje, 
1000 m; Vragonos above Treskavička reka, 
1500 m; along river Ločanska Bistrica, 1 00 m; 
Maja kurvala, 1900 m; valley Ropojana, 1200 m; 
karanfil, 1450 m, 2000 m; Belić, 1650 m, 
2200 m; pass Čakor, 1500 m (szePesfalvy 1926). 

Tayloria froelichiana (Hedw.) Mitt. ex Broth.: subarkt-subalp – Malje nedjinat, 1900 m. 

Tetraphis pellucida 
Hedw.: bor-temp – Babina gora, 
1700 m. 

Thamnobryum alopecurum (Hedw.) gang. var. alopecurum: temp –Vragonos above river Treskavička 
reka, 1500 m. 

Thuidium abietinum(Hedw.) B., S. & g.: ubikv – valley 
of river Bogska reka, 1 00 m; Hajla: Dramodol, 
2100 m; Vizitor, 1500 m; Devojački krš, 1900 m; 
valley grbaja, 1200 m; Belić, 1700 m; Peklen 
(szePesfalvy 1926 – 87, Dn42). 

Thuidium delicatulum 
(Hedw.) Mitt.: temp – near 
Peć (szePesfalvy 1926 – 87, Dn42); Dečani 
near Peć (szePesfalvy 1926 – 487, Dn41). 

Thuidium philiberti 
Limpr.: temp – gorge Rugovska 
klisura near Peć, 500 m; valley grbaja, 1200 m; 
valley Ropojana, 1200 m; Belić, 1400 m. 

*var. pseudotamarisci (Limpr.) Mönkm. – karanfil, 1450 m; valley Ropojana, 1200 m; Belić, 
1650 m. 

Thuidium tamariscinum 
(Hedw.) B., S. & g.: temp 

– Dečani near Peć (Szepesfalvy 1926 – 487, 
Dn41); Maja Rusolija, 1900–2200 m (RuDski 
1949 – 287, Dn4 ). 
*Timmia austriaca 
Hedw.: subarkt-subalp – Boge, 
1700 m; Vizitor, 1900 m; Planinica, 2000 m; Devojački krš, 1900 m; Starac, 2100 m; Malje nedjinat, 1900 m; Popadija, 1800 m; Pasji vrh, 2000 m; 
Djeravica, 2 00 m; Maja kurvala, 1900 m; karanfil, 1450, 2000 m; Belić, 1650, 2 00 m. 

Timmia bavarica Hessl.: subarkt-subalp – Vizitor, 
1900 m; Devojački krš, 2000 m; Starac, 2200 m; 
karanfil, 2100 m; Belić, 1900 m; Žljeb, 1500 m. 

Timmia norvegica 
Zetterst.: subarkt-subalp – Boge, 
1700 m; Hajla: kota 2278; valley Ropojana, 
1200 m; Belić, 1900 m. 

Tortella inclinata 
(Hedw. f.) Limpr.: temp – Belić, 
1650 m. 

Tortella tortuosa 
(Hedw.) Limpr.: bor-temp – at spring 
of river Istočka reka near Peć, 600 m; Boge, 
1500 m; Hajla: Dramodol, 2100 m; Vizitor, 
1900 m; Žuti kamen, 2000 m; Planinica, 1900 m; 


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Starac, 2100 m; Malje nedjinat, 1700 m; gorge 
Rugovska klisura near Peć, 500 m; valley grbaja, 1200 m; Popadija, 1900 m; springs Alipašini 
izvori near gusinje, 1000 m; valley Ropojana, 
1200 m; karanfil, 1600 m; Belić, 1800 m; Žljeb, 
1700 m. 

Tortula mucronifolia 
Schwaerg.: bor-mont 
– Malje 
nedjinat, 1700 m; near Peć (szePesfalvy 1926 

– 87, Dn42) 
Tortula muralis 
Hedw.: temp 
– at spring of river 
Istočka reka near Peć, 600 m; Vizitor, 1500 m; 
along river Dečanska Bistrica, 900 m; valley 
Ropojana, 1200 m; pass Čakor, 1500 m (szePesfalvy 1926). 

var. aestiva 
Hedw.: temp. – at spring of river Istočka 
reka near Peć, 600 m. 

Tortula obtusifolia 
(Schwaegr.) Math.: temp 
– Starac, 
1900 m; Popadija, 1800 m. 

Tortula subulata 
Hedw.: subtemp 
– valley of river 
Bogska reka near kučište, 1100 m; Hajla: Dramodol, 2100 m; Žljeb, 2000 m; Devojački krš, 
2000 m; Starac, 2200 m; Babina gora, 1600 m; 
Malje nedjinat, 2000 m; Popadija, 1900 m; 
Trojan, 1900 m; Volušnica, 1600 m; Djeravica, 
2 00 m; Treskavica under Veliki Rid, 1900 m; 
Maja kurvala, 1800 m; along river Ločanska Bistrica, 1 00 m; karanfil, 2000 m; Belić, 2 00 m; 
pass Čakor, 1500 m (szePesfalvy 1926). 

*var. angustata 
(Schimp.) Lindb.: merid-temp 
– 
Treskavica under Veliki Rid, 1600 m. 

var. graeffii 
Warnst. – valley of river Bogska reka, 
1100 m; Starac, 1900 m; along river Dečanska 
Bistrica, 1100 m. 

Trichostomum crispulum 
Bruch: merid-temp 
– gorge 
Rugovska klisura, 500 m; Volušnica, 1600 m; 
valley Ropojana, 1200 m. 

Warnstorfia exannulata 
(B., S. & g.) Loeske: bortemp 
– Starac, 2100 m; at lake Ridsko jezero, 
2000 m. 

Weisia condensa 
(Voit.) Lindb.: merid-temp 
– Popadija, 1700 m; Malje nedjinat, 2 00 m. 

Weisia controversa 
Hedw.: merid-temp 
– Devojački 
krš, 1900 m; Popadija, 1900 m; valley Ropojana, 
1200 m; Žljeb, 1700 m. 

Chorological analysis 

The basis for the classification of geoelements was 
the work of Düll (1999), but his classifications were 
modified in many respects when considering the 
work of Hill & PReston (1998) and the author’s 
own findings (MaRtinčič 1966). Unlike the condi

tions in vascular plants, the chorological analysis of 
the entire distribution area of mosses is extremely 
difficult and often even impossible. This is because 
the moss species’ distribution areas often extend 
over more than one continent or even on both 
hemispheres and are extremely disjunctive. When 
discussing the European species it is therefore 
more appropriate to define the floral elements 
based only on the European or Eurasian distribution area. The most important starting point here 
are the zonal vegetation belts – arctic, boreal, temperate, meridional (comp. szMaJDa 1989, MaRtinčič 
1966) – or zonobiomes (Hill & PReston 1998), 
with consideration of altitudinal belts. Changes in 
classifications as given by Düll occurred especially 
in assessment of the concepts of oceanic 
and continental, used by Düll to name geoelements, with 
which he thwarts the principle of zonal climatic 
vegetation belts. 

The results of the chorological analysis are presented in Figure 2. Phytogeographically the most 
important group is the arctic-alpine geoelement 
(arctic-alpine, subarctic-subalpine). It comprises 5 
species (1 ,1 %), the largest part of which are the 
subarctic-subalpine species, i.e. the species which 
do not reach into the real Arctic within Europe. 
Denotation »subalpine« is not the most suitable, 
as these species grow mostly in the alpine belt on 
the Balkan Peninsula – in the Prokletije Mts. and 
Macedonian mountains (comp. MaRtinčič 1966). 
HoRvat (195 ) already stressed that Prokletije Mts., 

..
..
..
.
..
..
..
..
Figure 2: Distribution of mosses of the Prokletije Mts. over 
chorological elements (in %). Numbers above bars – number 
of species. 

Slika 2: Razporeditev horoloških elementov v flori listnatih 
mahov Prokletij. Številke nad stolpci – število vrst. 

........
........
......
........
............
..........
.....
. 


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anDReJ MaRtinčič: Moss floRa of tHe PRokletiJe Mountains (seRBia, MontenegRo) 


apart from the Macedonian mountains, are an important diversity centre for flowering plants and 
lichens of the Alpine-arctic origin. The same is evident in moss flora. Contrary to the situation in the 
Macedonian mountains, however (MaRtinčič 1966, 
1980), where most of the arctic-alpine moss species grow only on silicate bedrock, in the Prokletije 
Mts. grow on silicate, crystalloid slates, as well as 
on Triassic limestones. It can be said for the entire 
group of arctic-alpine species that it migrated into 
the Prokletije Mts. in the Pleistocene. The largest 
number of species migrated from the Alps towards 
the southeast across the Dinaric crests. Unfavourable climatic conditions today, however, prevent 
the growth of most of these elements in the region 
between the Alps and the Prokletije Mts. The only 
exception are the silicate mountains Vranica in 
Bosnia. This is the reason why many arctic (subarctic)-alpine(subalpine) species in the western parts 
of the Balkan Peninsula exhibit a characteristic disjunct distribution or part of it: the Alps (Slovenia) 

– Vranica (Bosnia) – Prokletije Mts. (Serbia, Montenegro) – Šar planina (Macedonia). 
The boreal-montane element is represented 
with an even higher number of species as an arctic-
alpine element. It comprises as many as 60 species 
(22,4 %). The vertical range of thriving is relatively 
large. Most often, the species thrive in the regions 
above the timberline, the upper altitudinal line is 
between 2200 and 2500 m. only rarely and only 
isolated species reach below the 1000 m line. In 
comparison with the Alps there is therefore a perceptible increase in altitude for the lowest of the 
localities. 

Considering also the subboreal species (1 

– 4,8 %) we can determine that the flora of mosses in the Prokletije Mts. comprises as many as 
40. % species which can be called the »northern 
element«. The diversity centre of these species in 
Europe is in Scandinavia and in the mountains of 
Central Europe. Although we can notice an explicit decrease in the number of these species toward 
the south, they give the Prokletije Mts., contrary to 
the situation in flowering plants, an explicitly Alpine (Central-European) character. 
This picture is complemented by another 66 
(24,6 %) species of the boreal-temperate element 
(sensu Hill & PReston 1998). The species growing 
in Europe, above all in the boreal and temperate 
zonobiome are considered a boreal-temperate element, which corresponds to the boreal and Central-Europen floral region (sensu Meusel). Also 
worth mentioning is the group of the temperate 

element: temperate, temperate subcontinental (= 
Central-European), subtemperate, which comprises altogether 57 species (21, %). Among them, 
there are 4 species (16 %) that are truly temperate. The temperate-subcontinental element is represented with 9 species ( .4 %) – the species whose 
distribution area in Europe does not reach into 
the Atlantic parts in the west. Some of them have 
a relatively clear Central-European distribution. of 
other 7 species (7.5 %) we consider 17 (6. %) 
to belong to the meridional-temperate element 
(south-temperate sensu Hill & PReston 1998). All 
other species, with the exception of Leptodon smithii 
(atl-med), Metaneckera menziesii and Rhynchostegiella 
tenella (subatl-submed), Leucodon sciuroides var. morensis (submed), are the species commonly distributed in Europe. 

nature conservation value of the 
bryoflora 

Prokletije is included in the Site Register of the 
briologically important parts of Europe on account 
of the considerable diversity of moss flora and chorological types of species (MaRtinčič 1995). Comparison with Sabovljević’s calculations of the proportion of the number of species and the size of a 
particular region (saBovlJević (2004: 2 ) confirms 
that Prokletije is in fact one of the richest regions 
in the Balkan peninsula. Its significance is even 
greater considering that especially the higher regions are very well preserved. For the time being, 
human influence remains relatively small and for 
the most part limited to traditional management, 
pasture. only in certain areas in the lower regions 
is the use of forests more significant. The impact 
of tourism is negligible as it was practically non-existent because of the region’s borderline position 
between Albania and former yugoslavia. For all 
these reasons, briophytic flora still has a completely 
natural character. And as the habitats are so well 
preserved there are no signs of reduction in the 
population size of individual species or of any real 
threat to them. 

However, when taking into consideration the 
Red list for Serbia-Montenegro (saBovlJević et al. 
2004) we ascertain that 46 species from the list of 
the moss flora of Prokletije are considered endangered. 11 of these species fall into the category of 
critically endangered species (CR), 17 into the category of endangered species and 18 into the category of vulnerable species. A further 20 species are 


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Hacquetia 5/1 • 2006, 113–130 


in the low risk category. nevertheless, when dealing with such definitions we should be aware that 
the list is based on the revised IUCn criteria. These 
are founded above all on quantitative signs, on the 
number of the sites known in a certain area (see 
HallingBäck et al. 1998). In our case, this is seen 
especially in the critically endangered category and 
in the category of endangered species, where all of 
the 16 species found in Prokletije are included and 
which are new to Serbia-Montenegro. The inadequacy of quantitative criteria when determining 
the threat status in countries which have not been 
floristically adequately researched has already been 
stressed by PaPP & eRzBeRgeR (2005). 

nine species are included in the Red Data Book 
of European Bryophytes (ECCB 1995). Among 
them, one species, Buxbaumia viridis is vulnerable 
(VU),fivespeciesareintherare(R)category: Brachythecium geheebii, Bryum blindii, Dicranella humilis, 
Grimmia caespiticia, Paraleucobryum sauteri; and three 
of them are regionally threatened (RT): Campylium 
elodes, Pseudoleskea saviana, Tayloria froelichiana. 

PoVZETEk 

mahovna flora Prokletij (srbija, Črna gora) 

ozemlje nekdanje Jugoslavije je v brioflorističnem 
pogledu zelo neenakomerno raziskano. Med najmanj raziskanimi predeli so nekatera gorstva v Črni 
gori, Makedoniji in Srbiji (kosovo). Eno takih območij so Prokletije. Za del, ki leži na ozemlju držav 
Srbija, Črna gora, so bili doslej na razpolago le maloštevilni podatki o mahovni flori. 

Preučevano območje 

Prokletije so masiv, ki se razprostira v širšem obmejnem pasu med državami Srbija in Črna gora (republiki nekdanje Jugoslavije) ter Albanija (sl. 1). 
Predstavljajo jugovzhodni zaključek Dinarskega 
gorstva, ki se na jugu, v Albaniji, zaključuje z reko 
Drim. V nekdanjem jugoslovanskem delu se razprostirajo med Skadarskim jezerom in Vrmošo na 
zahodu ter Belim Drimom na vzhodu. na severu se 
zaključijo s Hajlo, Žljebom in Mokro planino. najvišji vrh je Djeravica (2656 m), večje število vrhov 
pa presega 2400 m. V geološkem pogledu vlada 
velika pestrost. Velik del »jugoslovanskih« Prokletij 
je zgrajen iz permokarbonskih skrilavcev ter paleozojskih skrilastih apnencev. Čeznje so ponekod 

debele plasti triadnih apnencev in dolomitov. kisle 
eruptivne kamnine so močno lokalizirane. V pleistocenu so bile Prokletije izpostavljene zelo močnemu vplivu poledenitev. Led je pokrival ne samo 
vrhove, temveč je segal daleč navzdol, obstajali pa 
so tudi veliki dolinski ledeniki. 

nahajališča (razvidna so v angleškem besedilu) 
so opremljena s številko osnovnega polja sistema 
srednjeevropskega florističnega kartiranja (MTB), 
ki smo ga uporabili za celotno ozemlje nekdanje 
Jugoslavije. oznaka v oklepaju pa izhaja iz mreže 
UTM ( 4T del) in podrazdelitve na mrežo 10 × 
10 km. Večina lokacij je opremljena tudi s kratico 
države, v kateri ležijo: Srbija (s) ali Črna gora (m). 

Horološka analiza 

na podlagi dosedanjih podatkov obsega flora listnatih mahov (Musci) »jugoslovanskega« dela Prokletij 268 vrst, kar predstavlja 42 % od skupnega 
števila doslej v Srbiji in Črni gori ugotovljenih vrst 
iz te skupine. 10 vrst je novih za ozemlje Srbije in 
44 za Črno goro. 

Podlago za opredelitev geoelementov je sicer 
predstavljalo delo Dülla (1999), vendar smo njegove opredelitve v mnogočem modificirali na podlagi dela Hill & PReston (1998) ter lastnih dognanj 
(MaRtinčič 1966). V nasprotju z razmerami pri semenkah je horološka opredelitev celotnega areala 
pri mahovih zelo težavna, marsikdaj celo nemogoča. Areali mahovnih vrst se namreč pogosto raztezajo na več kontinentih ali celo na obeh poloblah 
in so močno disjunktni. Zato je za vrste, ki uspevajo 
v Evropi, mnogo bolj smiselna opredelitev flornih 
elementov le na podlagi evropskega ali evrazijskega 
areala. Pri tem predstavljajo najpomembnejše izhodišče zonalno potekajoči vegetacijski pasovi – ark-
tični, borealni, temperatni, meridionalni (prim. sz-
MaJDa 1989, MaRtinčič 1966) oz. zonobiomi pri Hill 
& PReston (1998), z upoštevanjem višinskih pasov. 

Rezultate horološke analize prikazuje slika 2. Fitogeografsko najpomembnejšo skupino predstavlja 
arktično-alpinski geoelement (arktično-alpinski, 
subarktično-subalpinski). obsega 5 vrst (1 ,1 %), 
od tega odpade največji del na subarktično-subalpinske vrste, torej take, ki v Evropi ne segajo v 
pravo Arktiko. Že HoRvat (195 ) poudarja, da so 
Prokletije, poleg makedonskih gorstev, pomemben 
center razširjenosti cvetnic in lišajev alpsko-arktičnega porekla. omenjeno je razvidno tudi pri mahovni flori. Za celotno skupino lahko rečemo, da 
se je priselila v Prokletije v pleistocenu. 


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Še številneje je zastopan borealno-montanski 
element. obsega kar 60 vrst (22,4 %). najpogosteje uspevajo v predelih nad gozdno mejo, zgornja 
višinska meja je med 2200–2500 m. Le redko in 
izolirano sežejo pod 1000 m. V primerjavi z Alpami 
je viden občuten dvig nadmorske višine najnižjih 
nahajališč. ob upoštevanju še subborealnih vrst 
(1 –4,8 %) lahko ugotovimo, da je v flori listnatih mahov v Prokletijah kar 40, % vrst, ki jih lahko označimo kot »severni element«. V nasprotju z 
razmerami pri cvetnicah dajejo v Prokletijah izrazit 
alpski (srednjeevropski) pečat. 

navedeno podobo izpopolnjuje še 66 (24,6 %) 
vrst borealno-temperatnega elementa (sensu Hill 
& PReston 1998). Sem štejemo vrste, ki uspevajo 
v Evropi predvsem v borealnem in temperatnem 
conobiomu, kar ustreza borealni in srednjeevropski florni regiji (sensu Meusel). omembe vredna 
je še skupina temperatni element: temperatni, 
temperatno-subkontinentalni (= srednjeevropski), 
subtemperatni, ki obsega skupno 57 vrst (21, %). 
Med preostalimi 7 (7,5 %) vrstami prištevamo 17 
(6, %) vrst v meridionalno-temperatni element 
(south-temperate sensu Hill & PReston 1998). Vse 
druge so z izjemo vrst Leptodon smithii (atl-med), 
Metaneckera menziesii in Rhynchostegiella tenella (subatl-submed), Leucodon sciuroides var. morensis (submed) v Evropi splošno razširjene (ubikv). 

naravovarstvena ocena 

Prokletije so vključene v Seznam briološko pomembnih območij Evrope (MaRtinčič 1995). Po 
vrstni pestrosti predstavljajo eno najbogatejših območij na Balkanu. njihov pomen je še toliko večji, 
ker je vpliv človeka, zlasti v višjih legah, minimalen, 
ohranjeni so vsi pomembni habitati, zaradi obmejne lege pa tudi v bližnji prihodnosti ni pričakovati 
večjih škodljivih posegov zaradi komunikacije, urbanizacije, turizma. ne glede na rečeno, pa na podlagi Rdečega seznama ogroženih mahov Srbije in 
Črne gore spada 46 vrst med ogrožene: 11 vrst je v 
kategoriji kritično ogrožene, 17 je ogroženih in 18 
ranljivih. nadaljnjih 20 vrst spada med neznatno 
ogrožene. opredelitev je izvedena na podlagi novih kriterijev IUCn, ki upoštevajo za mahove v večini primerov zgolj število nahajališč (HallingBäck 
et al. 1998) in ne ogroženosti njihovih habitatov. 
Pri floristično slabše raziskanih območjih je starejša 
klasifikacija mnogo bolj ustrezna. 

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Recieved 18. 1. 2006 
Revision recieved 15. 5. 2006 
Accepted 2. 6. 2006 


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