NATURA SLOVENIAE 25(1): 37-45 Prejeto / Received: 16. 6. 2023 SHORT COMMUNICATION Sprejeto / Accepted: 25. 6. 2023 DOI: 10.14720/ns.25.1.37-45 Biotehniška fakulteta Univerze v Ljubljani in Nacionalni inštitut za biologijo, Ljubljana, 2023 Addition to the knowledge on the moth fauna (Insecta: Lepidoptera) of Bosnia and Herzegovina and Croatia Toni KOREN1*, Dejan KULIJER2 1Association Hyla, Lipovac I no. 7, HR-10000 Zagreb, Croatia; E-mail: toni.koren@hhdhyla.hr 2National museum of Bosnia and Herzegovina, Zmaja od Bosne 3, 71000 Sarajevo, Bosnia and Herzegovina; E-mail: dejan.kulijer@gmail.com *Corresponding author Abstract. Additions to the moth fauna of Bosnia and Herzegovina and Croatia are given. We report on first observations of five species and on reconfirmation of one species after eight decades. In Croatia, we recorded the species Eublemma cochylioides (Guenée, 1852) for the first time, while Eublemma himmighoffeni (Millière, 1867) has been confirmed in the country after 85 years. We report on the first observation of five moth species for Bosnia and Herzegovina: Triodia adriaticus (Osthelder, 1931), Eumera regina Staudinger, 1892, Dryobotodes monochroma (Esper, 1790), Lithophane lapidea (Hübner, [1808]) and Nycteola columbana (Turner, 1925). Even though these records could be expected due to species general distributions, they fill the knowledge of moth diversity of both countries and the wider region. Key words: Hepialidae, Geometridae, Erebidae, Nolidae, Noctuidae, distribution, diversity Izvleček. Prispevek k favni nočnih metuljev (Insecta: Lepidoptera) Bosne in Hercegovine in Hrvaške – V prispevku podajamo nove podatke o nočnih metuljih v Bosni in Hercegovini ter na Hrvaškem, s prvimi opažanji petih vrst in ponovno potrditvijo ene vrste po več kot osem desetletjih. Na Hrvaškem smo prvič zabeležili vrsto Eublemma cochylioides (Guenée, 1852), medtem ko je bila Eublemma himmighoffeni (Millière, 1867) ponovno najdena po 85 letih. V Bosni in Hercegovini smo našli pet novih vrst za državo: Triodia adriaticus (Osthelder, 1931), Eumera regina Staudinger, 1892, Dryobotodes monochroma (Esper, 1790), Lithophane lapidea (Hübner, [1808]) in Nycteola columbana (Turner, 1925). Čeprav je bilo te vrste mogoče pričakovati glede na njihovo splošno razširjenost, naši podatki dopolnjujejo poznavanje raznolikosti nočnih metuljev v obeh državah in v regiji. Ključne besede: Hepialidae, Geometridae, Erebidae, Nolidae, Noctuidae, razširjenost, raznovrstnost Introduction In the beginning of the 20th century, an impressive work was published that covered the previously unknown moth fauna of Bosnia and Herzegovina (Rebel 1904a) and which today serves as a landmark for all who research the butterfly and moth fauna of the country. Afterwards, only a few papers dealing with the moths of Bosnia and Herzegovina were published 38 Toni KOREN & Dejan KULIJER: Addition to the knowledge on the moth fauna ... / SHORT COMMUNICATION NATURA SLOVENIAE 25(1): 37-45 (Schawerda 1918; Stauder 1925, 1926, 1929, 1930, 1933), most of which have been consolidated in the revision of the list (Lelo 2004). Still, in comparison with the neighbouring countries, the moth diversity of Bosnia and Herzegovina is far from sufficient (Plant & Jakšić 2018) and records of additional species of almost all moth families are expected. In Croatia the moth fauna, especially Macroheterocera, has been studied in greater detail, and for the larger families published and unpublished checklists exist (Kučinić 1997; Koren 2012; Mihoci 2012; Gumhalter 2019) rending it easier to put any records in the comparative perspective. During the recent years, several new and/or interesting moth species have been recorded in Bosnia and Herzegovina (Hanjalić & Lelo 2015; Koren & Kulijer 2020; Koren & Martinović 2020; Beshkov & Nahirnić-Beshkova 2021; Koren et al. 2021, 2022a), as well as in Croatia (Mrnjavčić Vojvoda et al. 2014; Koren & Gomboc 2015; Koren & Kulijer 2020). In this work the records of additional six species are presented, five new to Bosnia and Herzegovina and one new to the fauna of Croatia, with one reconfirmation of the species after over eight decades. Materials and methods This report covers our observations from the 2020–2022 period in Bosnia and Herzegovina and Croatia (Tab. 1). The study in Bosnia and Herzegovina was mainly conducted on the Klek peninsula and the nearby mainland, while the study in Croatia was conducted within the Klek settlement and the mainland south of the border with Bosnia and Herzegovina. Two main light-trapping methods were used. The main method applied was the pyramid- shaped collecting tent comprising a metal frame and two 15W UV lamps (604 mm) connected to a 12 V battery and covered with a white canvas. Six traps were used, distanced about ten meters apart. The second method implied the usage of two 6W 12V portable heath moth traps with actinic UV-A lamps (350–400 nm) which were set in the dusk and left on site and collected the following morning. Three portable heath moth traps were used per locality and night. In addition, in Klek settlement, Croatia, a 100W mercury light bulb put on a terrace was used to attract moths. The android application and digital platform Biologer were used to record field data during this research (Popović et al. 2020). The specimens were identified and are stored in the collection of T. Koren. For each record, the exact locality, coordinates, altitude, and dates are provided. Toni KOREN & Dejan KULIJER: Addition to the knowledge on the moth fauna ... / SHORT COMMUNICATION 39 NATURA SLOVENIAE 25(1): 37-45 Table 1. The list of localities with methods used and new records of moth species in Bosnia and Herzegovina and Croatia. Abbreviations: DK – Dejan Kuljer, TK – Toni Koren. Coordinates are in WGS84 decimal degrees. Tabela 1. Seznam lokalitet z uporabljenimi metodami in novimi podatki za nočne metulje Bosne in Hercegovine in Hrvaške. Okrajšave: DK – Dejan Kuljer, TK – Toni Koren. Koordinate so v WGS84 decimalnih stopinjah. No. Location Method Date (Observer) Species 1 House at Opuće settlement, Neum, BIH; 42.927186° N, 17.572877° E, 40 m a.s.l. portable heath moth traps 1.10.2021 (DK) Triodia adriaticus 2 Opuće settlement, northern part, Neum, BIH; 42.928529° N, 17.572879° E, 35 m a.s.l. portable heath moth traps 16.10.2021 (DK) 19.10.2021 (DK) Dryobotodes monochroma Lithophane lapidea 3 Konštar locality, Neum city, BIH; 42.930888° N, 17.658310° E, 175 m a.s.l. portable heath moth traps 14.9.2020 (DK) Eumera regina Nycteola columbana 4 Gradina locality, Neum city, BIH; 42.932013° N, 17.684761° E, 200 m a.s.l. portable heath moth traps 16.9.2020 (DK) Eumera regina 5 Klek settlement, CRO; 42.946881° N, 17.56341° E, 81 m a.s.l. mercury light bulb 9.10.2022 (DK) Eublemma cochylioides 6 Slopes on the eastern side of the bay, grasslands, and overgrown agricultural land, Zaton, Dubrovnik, CRO; 42.696625° N, 18.048319° E, 55 m a.s.l. pyramid-shaped tents 27.5.2021 (TK) Eublemma himmighoffeni Results and discussion In total, the records of seven moth species are presented, five being recorded for the first time in Bosnia and Herzegovina, one in Croatia and one rediscovered in Croatia after 85 years. The species were registered at six different localities (Fig. 1). The species Triodia adriaticus (Osthelder, 1931) (fam. Hepialidae) was found for the first time in Bosnia and Herzegovina, Opuće settlement (Loc. 1; Tab. 1, Fig. 1). This species inhabits parts of south-eastern Europe, from Istria in Croatia, across the Adriatic coastline to Macedonia, Albania (Beshkov 1994), Ionian Sea coast, Korfu and Peloponnesus in Greece (de Freina and Witt 1990). This species is usually numerous in favourable habitats and can be attracted to light sources in high numbers. No previous records of this species exist for Bosnia and Herzegovina, but its occurrence was expected due to the probably continuous distribution along the Adriatic coastline (Koren 2020). The species Eumera regina Staudinger, 1892 (fam. Geometridae) was found for the first time for Bosnia and Herzegovina, at two different localities, Konštar (Loc. 3; Tab. 1, Fig. 1) and Gradina (Loc. 4; Tab. 1, Fig. 1). This is a unique and striking Geometridae moth that cannot be confused with any other species due to the forewing terminal area being yellowish pink with two white dots (Fig. 2a) (Skou & Sihvonen 2015). Within Europe, the distribution of this species is restricted to the Balkan Peninsula and ranges from Dalmatia (Croatia) to Greece (Skou & Sihvonen 2015). No previous records of this species exist for Bosnia and Herzegovina, but its 40 Toni KOREN & Dejan KULIJER: Addition to the knowledge on the moth fauna ... / SHORT COMMUNICATION NATURA SLOVENIAE 25(1): 37-45 occurrence was expected due to recent records of E. regina in the Neretva River Delta (Koren, unpubl.), just several kilometres west of the localities where the species was found in Bosnia and Herzegovina. It was recorded in a habitat typical of this species in the region, karst grasslands surrounded by maquis (Fig. 2b). Figure 1. Distribution of the newly recorded moth species in Bosnia and Herzegovina & Croatia. Slika 1. Razširjenost novo zabeleženih vrst nočnih metuljev v Bosni in Hercegovini in na Hrvaškem. The species Dryobotodes monochroma (Esper, 1790) (fam. Noctuidae) was found for the first time in Bosnia and Herzegovina (Rebel 1904a; Lelo 2004), recorded on two different dates at Opuće settlement (Loc. 2; Tab. 1, Fig. 1). This is a Holo-Ponto-Mediterranean species with its northern distribution limit in Central Europe (Ronkay et al. 2001). The map presented in the Noctuidae Europaeae series (Ronkay et al. 2001) is misleading, as it includes the whole Balkan Peninsula and its countries. For Croatia and Bosnia and Herzegovina, this does not hold true. In Croatia, this species is, according to the newly gathered data (Koren, unpubl.) and historical records (Rebel 1912; Schwingenschuss & Wagner 1926; Stauder 1926), present only on the Mediterranean coastline, while in Bosnia and Herzegovina there have been no records of this species so far (Rebel 1904a; Lelo 2004). Thus, the presence of this species in the Mediterranean part of Bosnia and Herzegovina was expected. The identification was confirmed by the examination of male genital structures in order to distinguish it from the very similar Dryobotodes servadeii Parenzan, 1982 (Ronkay et al. 2001). Toni KOREN & Dejan KULIJER: Addition to the knowledge on the moth fauna ... / SHORT COMMUNICATION 41 NATURA SLOVENIAE 25(1): 37-45 Figure 2. Photos of three moth species, found in Bosnia and Herzegovina and Croatia: a) Eumera regina from Gradina, b) habitat of E. regina at Gradina, c) Eublemma cochylioides from Klek settlement, d) Eublemma himmighoffeni from Zaton. For the localities, see Tab. 1. Photo: T. Koren (a, d), D. Kulijer (b, c). Slika 2. Fotografije treh vrst nočnih metuljev, najdenih v Bosni in Hercegovini in na Hrvaškem: a) Eumera regina z lokalitete Gradina, b) habitat vrste E. regina na lokaliteti Gradina, c) Eublemma cochylioides iz naselja Klek, d) Eublemma himmighoffeni z lokacije Zaton. Za lokalitete glej Tab. 1. Foto: T. Koren (a, d), D. Kulijer (b, c). The second species of the family Noctuidae, Lithophane lapidea (Hübner, [1808]), was recorded at the second locality near Opuće settlement (Loc. 2; Tab. 1, Fig. 1), which is the first observation for Bosnia and Herzegovina. This is a Ponto-Mediterranean species in Europe occurring in France, northern and central Italy and across the Balkan Peninsula (Ronkay et al. 2001). In neighbouring Croatia, it is a fairly common species along the Adriatic coastline from Istria in the north (Koren 2022) down to the southern part of the country. Thus, the presence in the Mediterranean part of Bosnia and Herzegovina was expected. The species Nycteola columbana (Turner, 1925) (fam. Nolidae) was found at Konštar locality (Loc. 3; Tab. 1, Fig. 1) for the first time in Bosnia and Herzegovina (Rebel 1904b; Lelo 2004). This is an Eurasiatic species, distributed in southern parts of Europe, northern Africa and the Middle East (Fibiger et al. 2009). It is very similar to other members of the genus Nycteola and the correct identification was confirmed by examining the genital structures (Fibiger et al. 2009). 42 Toni KOREN & Dejan KULIJER: Addition to the knowledge on the moth fauna ... / SHORT COMMUNICATION NATURA SLOVENIAE 25(1): 37-45 The species Eublemma cochylioides (Guenée, 1852) (fam. Erebidae) (Fig. 2c) was found for the first time in Croatia at Klek settlement (Loc. 5; Tab. 1, Fig. 1). This is a variable species with the forewing colour ranging from yellow to pink. Especially notable are the specimens with the half of the forewing coloured yellow and half pink as is the case with the specimen from Klek (Fig. 2c). This individual was observed and photographed only when reaching a mercury bulb positioned at the external side of the house. Due to the unique appearance, its identification was possible even without catching the individual. The species E. cochylioides is a tropical species with disjunct distribution in Europe, Africa, Middle East to SE Asia, all to Australia (Fibiger et al. 2010). Recently, the spreading of this species has been noted in northern Europe and recorded up to Great Britain (Hatton et al. 2022). This species inhabits open, steppe-like habitats. Adults fly from September to November. Larvae feed on different Asteraceae species (Fibiger et al. 2010). This is not only the first record in Croatia, but also the northernmost record along the Adriatic coast. While the migrating behaviour is known in this species (Hatton et al. 2022), it is possible that it was just overlooked during previous surveys and is indeed native to the area. The second species of the Erebidae family, Eublemma himmighoffeni (Millière, 1867) (Fig. 2d), was found at Zaton in Croatia (Loc. 6; Tab. 1, Fig. 1). This is a small and easily recognizable member of the genus Eublemma due to its forewing colouration. For this species, only one reliable record exists in the literature for Croatia, Gruž (Gravosa) in Dubrovnik city (Schwingenschuss & Wagner 1926). Stauder (1927) mentioned the presence of this species in Dalmatia, but without exact locations. The second record of the species, also from Gruž, is available on the web portal lepiforum.de with the following data: Dalmatia, Gravosa - Zaton, 1.–8.10.1936 H. Fabigan (Lepiforum e.V. 2021). While the new record from Croatia reported herewith is not far from the previously known area of this species around Dubrovnik, it represents the first confirmed record of the species in the country after 85 years. The known flight period of this species is from the end of June till the end of August (Fibiger et al. 2010) and up to mid-September (Beshkov 1994), but our record from Croatia shows that this species is active even earlier, at the end of May. It inhabits dry grassland habitats near the coast (Fibiger et al. 2010), which concurs with our observations (Tab. 1). Conclusions Many species are being recorded in the Mediterranean part of the Bosnia and Herzegovina region, mostly the Neum and Klek peninsula areas. Here, the political border of Croatia is discontinued, with part of the coastline and the sea belonging to Bosnia and Herzegovina. Thus, it is the hotspot for recording Mediterranean species in the country, such as the ones presented in this work, that have not been recorded in the country so far. It is most likely that many of these species are present also within the mainland Herzegovina, as is the case with Mediterranean butterfly species (Lelo 2008; Koren et al. 2019; Koren et al. 2022b), but this should be investigated with additional surveys. Toni KOREN & Dejan KULIJER: Addition to the knowledge on the moth fauna ... / SHORT COMMUNICATION 43 NATURA SLOVENIAE 25(1): 37-45 The Klek Peninsula and the small territory around Neum are the only parts of Bosnia and Herzegovina with characteristic Eumediterranean and maritime habitats and therefore highly valuable for the overall biodiversity of the country. For several true Mediterranean plant species and habitat types, this is the sole area where they occur on the territory of Bosnia and Herzegovina (Kutleša & Lakušić 1964; Maslo & Milanović 2022). Due to the restricted range in the country, tourism development and related urbanization and construction of infrastructure, Mediterranean habitats and the species that inhabit them are under significant and growing threat. References Beshkov S. 1994. A Contribution to the knowledge of the Lepidoptera fauna of Albania 2. Some findings of a collecting trip in September 1993. Atalanta 26(1-2): 365-399. Beshkov S, Nahirnić-Beshkova A. 2021. Contribution to knowledge of the Balkan Lepidoptera II (Lepidoptera, Macrolepidoptera). Natura Montenegrina. 42: 1-44. doi: 10.37828/em.2021.42.1 de Freina JJ, Witt TJ. 1990. Die Bombyces und Sphinges der Westpalaearktis (Insecta, Lepidoptera), Band 2, Cossoidea: Cossidae, Limacodidae, Megalopygidae. Hepialoidea: Hepialidae. Pyraloidea: Thyridae. Zygaenoidea: Epipyropidae, Heterogynidae, Edition FW. ed. München (DE): Edition Forschung und Wissenschaft Verlag GmbH. Fibiger M, Ronkay L, Steiner A, Zilli A. 2009. Pantheinae, Dilobinae, Acronictinae, Eustrotiinae, Nolinae, Bagisarinae, Acontiinae, Metoponiinae, Heliothinae and Bryophilinae. Noctuidae Europaeae. Vol. 11. Sorø (DK): Entomological Press. Fibiger M, Ronkay L, Yela JL, Zilii A. 2010. Rivulinae, Boletobiinae, Hypenodinae, Araeopteroninae, Eublemminae, Herminiinae, Hypeninae, Phytometrinae, Euteliinae, and Micronoctuidae, including Supplement to Volumes 1-11. Noctuidae Europaeae, Vol. 12. Sorø (DK): Entomological Press. Gumhalter D. 2019. First checklist of pyraloid moths (Lepidoptera, Pyraloidea) in Croatia. Zootaxa. 4604(1): 59-102. doi: 10.11646/ZOOTAXA.4604.1.3 Hanjalić J, Lelo S. 2015. Hrvatska golupka, Hemaris croatica (Esper, 1800) (Lepidoptera, Sphingidae), nova vrsta u fauni leptira Bosne i Hercegovine. Prilozi fauni Bosne i Hercegovine. 11: 29-34. Hatton DH, Chainey JE, Spence JM. 2022. Eublemma cochylioides (Guenée, 1852) (Lepidoptera, Erebidae) in Hertfordshire – new to Great Britain. Entomologist's Record and Journal of Variation. 134(6): 282-286. Koren T. 2012. Distributional checklist of lappet moths (Lepidoptera, Lasiocampidae) of Croatia. Entomologia Croatica. 16(1-4): 81-104. Koren T. 2020. Butterflies and moths (Insecta, Lepidoptera) of the Lokrum island, southern Dalmatia. Natura Croatica. 29(2): 227-240. doi: 10.20302/NC.2020.29.29 Koren T. 2022. The diversity of moths (Lepidoptera, Heterocera) of significant landscape Donji Kamenjak and Medulin Archipelago, Istria, Croatia. Annales, Series Historia Naturalis. 32(1): 237- 260. doi: 10.19233/ASHN.2022.25 44 Toni KOREN & Dejan KULIJER: Addition to the knowledge on the moth fauna ... / SHORT COMMUNICATION NATURA SLOVENIAE 25(1): 37-45 Koren T, Gomboc S. 2015. First record of Chersotis rectangula ([Denis and Schiffermüller], 1775) in Croatia with new data for Chersotis multangula (Hübner, 1803) (Lepidoptera, Noctuidae). Šumarski list. 139(9-10): 441-445. Koren T, Kulijer D. 2020. Additions to the Crambidae (Insecta, Lepidoptera) fauna of Croatia and Bosnia & Herzegovina. Acta Entomologica Slovenica. 28(2): 141-148. Koren T, Martinović M. 2020. A contribution to the knowledge of the moth diversity of Bosnia and Herzegovina. Entomologist's Record and Journal of Variation. 132: 124-128. Koren T, Kulijer D, Martinović M. 2021. Baptria tibiale (Esper, 1804) (Lepidoptera, Geometridae), a new genus and species for the fauna of Bosnia and Herzegovina. Acta Entomologica Serbica. 26(2): 81-84. Koren T, Dender D, Ilić B, Martinović M. 2019. On the distribution of the Lattice Brown, Kirinia roxelana (Cramer, 1777) in Southern Croatia (Lepidoptera, Nymphalidae). Acta Entomologica Serbica. 24(1): 41-45. doi: 10.5281/zenodo.3266098 Koren T, Kulijer D, Martinović M, Lauš B, Kranželić D. 2022a. New records of the bat hawkmoth Hyles vespertilio (Esper, 1780) in Croatia, Bosnia and Herzegovina and Serbia. Acta Entomologica Serbica. 27(1): 1-11. https://doi.org/10.5281/zenodo.6390464 Koren T, Laus B, Burić I, Štih A, Kulijer D, Dender D, Kranželić D, Schmidt B, Verovnik R. 2022b. New records of Tarucus balkanicus at the northern limit of its distribution along the Adriatic coast (Lepidoptera, Papilionoidea, Lycaenidae). Fragmenta Entomologica. 54(1): 143-150. doi: 10.13133/2284-4880/768 Kučinić M. 1997. Faunističke, ekološke i zoogeografske značajke sovica (Insecta, Lepidoptera) Hrvatske [magistarski rad]. [Zagreb (HR)]: Prirodoslovno-matematički fakultet Zagreb. Kutleša L, Lakušić R. 1964. Flora and vegetation of the Klek Peninsula. Godišnjak Biološkog instituta Univerziteta u Sarajevu. 17: 61-115. Lelo S. 2004. Revizija Rebelovog popisa leptira Bosne i Hercegovine. Sarajevo (HR): Coron’s d.o.o. Lelo S. 2008. Dnevni leptiri Bosne i Hercegovine (Lepidoptera, Papilionoidea i Hesperioidea). Ključ za determinaciju vrsta sa osnovnim monografskim podacima. Sarajevo (HR): Prirodno-matematički fakultet Univerziteta u Sarajevu. Lepiforum e.V. 2021. Lepiforum e.V. Bestimmung von Schmetterlingen und ihren Präimaginalstadien; [accessed 4. 11. 2021]. https://www.lepiforum.org/ Maslo S, Milanović Đ. 2022. New records of two grass species of the tribe Andropogoneae (Poaceae) in Bosnia and Herzegovina. Phytologia Balcanica. 28(1): 69-74. doi: 10.7546/PhB.28.2022.6 Mihoci I. 2012. Raznolikost grbica (Lepidoptera, Geometridae) Hrvatske i ekološka uvjetovanost njihove visinske rasprostranjenosti [PhD]. [Zagreb (HR)]: University of Zagreb, Faculty of Science Zagreb. Mrnjavčić Vojvoda A, Mihoci I, Vajdić M, Kučinić M. 2014. Antitype suda (Geyer, 1832) (Lepidoptera, Noctuidae), new species of noctuid fauna of Croatia, found in the Biokovo Nature Park. Natura Croatica. 23(2): 379-388. Plant CW, Jakšić P. 2018. A provisional checklist and bibliography of the Pyraloidea of the Balkan Peninsula. Atalanta. 49(1–4): 219-263. Toni KOREN & Dejan KULIJER: Addition to the knowledge on the moth fauna ... / SHORT COMMUNICATION 45 NATURA SLOVENIAE 25(1): 37-45 Popović M, Vasić N, Koren T, Burić I, Živanović N, Kulijer D, Golubović A. 2020. Biologer: an open platform for collecting biodiversity data. Biodiversity Data Journal 8: e53014. doi: 10.3897/BDJ.8.e53014 Rebel H. 1904a. Studien über die Lepidopterenfauna der Balkanländer. II. Teil. Bosnien und Herzegowina. Annalen des k. k. Naturhistorischen Hofmuseum. 19: 97-377. Rebel H. 1904b. Sistematski spisak Lepidoptera (Lepirova) Bosne i Hercegovine. Glasnik Zemaljskog muzeja Bosne i Hercegovine. 16: 375-416. Rebel H. 1912. Lepidopteren aus dem Gebiete des Monte Maggiore in Istrien. I. Nachtrag. Jahresberichte Wiener entomologischer Verein. 22: 227-240. Ronkay L, Yela JL, Hreblay M. 2001. Hadeninae II. Noctuidae Europaeae. Vol. 5. Sorø (DK): Entomological Press. Schawerda K. 1918. Achter Nachtrag zur Lepidopterenfauna Bosniens und der Herzegowina. Verhandlungen der Zoologisch-Botanischen Gesellschaft in Wien. 68: 19-36. Schwingenschuss L, Wagner F. 1926. Beitrag zur Macro-Lepidopteren-Fauna Süddalmatiens insbesondere der Umgebung Gravosa’s. Zeitschrift des Österreichischen Entomologischen Vereins. 11-12: 1-3, 9-13, 26-29, 53-54, 67-72, 74-80, 81-86, 45-50, 62-64, 68-72. Skou P, Sihvonen P. 2015. Ennominae I. In: Hausmann A, editor. The Geometrid Moths of Europe, Vol. 5. Leiden (NL): Brill. Stauder H. 1925. Die Schmetterlingsfauna der illyro-adriatischen Festland- und Inselzone (Faunula Illyro-Adriatica). Zeitschrift für wissenschaftliche Insektenbiologie. 20(8): 191-226. Stauder H. 1926. Die Schmetterlingsfauna der illyro-adriatischen Festland- und Inselzone (Faunula Illyro-Adriatica). Zeitschrift für wissenschaftliche Insektenbiologie. 21(8/9): 179-190, 223-238. Stauder H. 1927. Die Schmetterlingsfauna der illyro-adriatischen Festland- und Inselzone (Faunula Illyro-Adriatica). Zeitschrift für wissenschaftliche Insektenbiologie. 21(10): 30-45, 74-92. Stauder H. 1929. Die Schmetterlingsfauna der illyro-adriatischen Festland- und Inselzone (Faunula Illyro-Adriatica). Entomologische Zeitung. 9: 39-41, 53-58, 65-72, 88-94, 115-117, 131-137, 157- 158, 177-178, 194-199, 213-218, 233-236, 251-252, 272-273, 292-293, 299-306, 318-325, 359- 364, 379-384, 397-403, 420-423, 437-442. Stauder H. 1930. Die Schmetterlingsfauna der illyro-adriatischen Festland- und Inselzone (Faunula Illyro-Adriatica). Entomologische Zeitung. 10: 9-10, 36-37, 52-57, 75-77, 105-108, 133-136, 153- 155, 201-204, 249-252, 275-276, 294-297, 309-310, 351-352, 374-377, 436-437. Stauder H. 1933. Die Schmetterlingsfauna der illyro-adriatischen Festland- und Inselzone (Faunula Illiyro-Adriatica). Entomologische Zeitung. 13: 18-20. © 2023 Toni Koren & Dejan Kulijer. To je prostodostopen članek, objavljen pod določili licence Creative Commons Priznanje avtorstva 4.0 Mednarodna, ki dovoljuje neomejeno rabo, razširjanje in kopiranje v kakršnemkoli mediju ter obliki, pod pogojem, da sta navedena avtor in vir. This is an open-access article distributed under the terms of the Creative Commons Attribution 4.0 International License, which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited.