FOLIA BIOLOGICA ET GEOLOGICA 64/1, 5–100, LJUBLJANA 2023 VEGETATION OF MOIST ROCK CREVICES AND MOIST (SLOPE) DEBRIS IN THE LIŠČAK GORGE (BAČA VALLEY, JULIAN ALPS) VEGETACIJA VLAŽNIH SKALNIH RAZPOK IN VLAŽNEGA POBOČNEGA GRUŠČA V GRAPI LIŠČAKA (BAŠKA DOLINA, JULIJSKE ALPE) In memory of Prof. Stanko Buser (1932-2006) / V spomin prof. Stanku Buserju (1932-2006) Igor DAKSKOBLER1 & Andrej MARTINČIČ2 http://dx.doi.org/10.3986/fbg0100 ABSTRACT Vegetation of moist rock crevices and moist (slope) debris in the Liščak gorge (Bača Valley, Julian Alps) We studied the phytosociology of communities of moist rock crevices and slope debris in the Liščak gorge in the southern Julian Alps. Based on the relevés from this and sev- eral other gorges in the Julian Alps and their foothills we described new associations Violo biflorae-Astrantietum car- niolicae, Palustriello commutati-Astrantietum carniolicae, Veronico urticifoliae-Violetum biflorae, Palustriello commu- tati-Veronicetum urticifoliae, Veronico urticifolia-Saxifrage- tum cuneifolii, Calamagrostio variae-Asteretum bellidiastri and Lamio orvalae-Lunarietum redivivae, and a new alliance Astrantio carniolicae-Paederotion luteae. Key words: vegetation, Asplenietea trichomanis, Arunco- -Petasition albi, Lamio orvalae-Sambucetum nigrae, Natura 2000, Slovenia IZVLEČEK Vegetacija vlažnih skalnih razpok in vlažnega pobočnega grušča v grapi Liščak (Baška dolina, Julijske Alpe) Fitocenološko smo proučili združbe vlažnih skalnih razpok in pobočnega grušča v grapi Liščak v južnih Julijskih Alpah. Na podlagi popisov iz te grape in iz nekaterih drugih grap v Julijskih Alpah s prigorjem smo opisali nove asociaci- je Violo biflorae-Astrantietum carniolicae, Palustriello com- mutati-Astrantietum carniolicae, Veronico urticifoliae-Viole- tum biflorae, Palustriello commutati-Veronicetum urticifoli- ae, Veronico urticifolia-Saxifragetum cuneifolii, Calamagro- stio variae-Asteretum bellidiastri in Lamio orvalae-Lunarie- tum redivivae ter novo zvezo Astrantio carniolicae-Paedero- tion luteae. Ključne besede: vegetacija, Asplenietea trichomanis, Arunco-Petasition albi, Lamio orvalae-Sambucetum nigrae, Natura 2000, Slovenija 1 Scientific Research Centre of the Slovenian Academy of Sciences and Arts, 1Institute of Biology, Regional unit Tolmin, Brunov drevored 13, SI-5220 Tolmin, Igor.Dakskobler@zrc-sazu.si 2 Zaloška 78 a, SI-1000 Ljubljana, andrej.martincic@siol.net DAKSKOBLER & MARTINČIČ: VEGETATION OF MOIST ROCK CREVICES AND MOIST (SLOPE) DEBRIS IN THE LIŠČAK GORGE 6 FOLIA BIOLOGICA ET GEOLOGICA 64/1 – 2023 The Liščak is an about 3-km-long creek in the Kneška Grapa gorge and the Bača Valley in the southern Julian Alps. It starts at the elevation of around 1000 m under Mt. Ploha (1270 m) in the Tolmin–Bohinj range and flows into the Kneža river at 331 m a.s.l. At the straight line distance of around 2500 m it overcomes an almost 700 m difference in height. Numerous tributaries flow into the stream from both banks at an even steeper gradient than that of the main gorge, which is very narrow, with only a few small expansions where gravel and debris accummulate. The geological bedrock is ex- tremely diverse, consisting of dolomite, limestone, chert, marlstone and claystone (Buser 1986, 1987). As it faces southeast, the climate is relatively warm and humid, with an annual precipitation average of above 2000 mm (Zupančič 1998) and the mean annual tem- perature of around 7–8 °C (Cegnar 1998). Around 90% of the land cover in the Liščak basin (3.42 km2) is forest. The largest forest stand areas are classified into the following associations: Seslerio autu- mnalis-Fagetum, Lamio orvalae-Fagetum, Saxifrago cu- neifolii-Fagetum, Fraxino orni-Ostryetum and Veratro nigri-Fraxinetum excelsioris. Not a single house in the entire Liščak basin is in- habited anymore; there are still remains of old mills, but no artificial barriers or other direct human impact. Because of numerous geological and geomorphologi- cal specifics the stream and its gorge are protected as a valuable natural feature of regional or local impor- tance (Rojšek 1986, 1991, https://www.naravovarstve- ni-atlas.si/web/DefaultNvaPublic.aspx.). Our first investigations into the vegetation of the Liščak basin date back to 1986 – 1988. After a longer period we revisited the gorge in 2015 and subsequently, mainly in 2018, 2019, 2020 and 2021, we made more than 200 phytosociological and floristic relevés, some of them also under the guidance and assistance of Peter Razpet. An overall description of flora and forest vegetation will be the subject of another paper. Here, we focus exclusively on the vegetation of moist rock crevices and tall herbs on moist slope debris (Figure 1). 1 INTRODUCTION 2 METHODS Our relevés were made using the Central-European phytosociological approach (Braun-Blanquet 1964) and entered into the FloVegSi database (T. Seliškar, Vreš & A. Seliškar 2003). We collected mosses and liverworts, which Andrej Martinčič, the co-author, de- termined in the laboratory. The relevés in Tables 1–13 were processed using hierarchical classification, un- weighted average linkage method – UPGMA and Wis- hart’s similarity ratio. We transformed the combined cover-abundance values into ordinal scale (1–9) ac- cording to van der Maarel (1979). Numerical com- parisons were performed with the SYN-TAX 2000 pro- gram package (Podani 2001). Our aim was to classify the relevés from Liščak and its immediate vicinity (Ve- like Luti) into a syntaxonomic system with previously described communities. As this was not always possi- ble, some of our tables include relevés from other gorg- es in western and northwestern Slovenia, where we re- cently studied chasmophytic vegetation (Dakskobler et al. 2021, Dakskobler & Martinčič 2020, 2021a). This served as the basis for our description of several new associations. Most of the communities of moist rock crevices in the submontane and montane belt in the Julian Alps and their foothills are dominated by vascular plants Pinguicula alpina, Astrantia carniolica, Viola biflora, Aster bellidiastrum, Veronica urticifolia, Valeriana tripteris, V. saxatilis, Saxifraga cuneifolia, Saxifraga aizoides, Asplenium viride, A. trichomanes, and mosses and liverworts Orthothecium rufescens, Pa- lustriella commutata, Hymenostylium recurvirostrum and Conocephalum conicum. Despite considerable flo- ristic similarity they cannot be classified into a single association. In our descriptions of new communities at the rank of association we therefore took into account the constancy and medium cover of the dominant spe- cies in our relevés, because this is the key trait that al- lows us to identify these communities also in the field. We apply the same criteria for certain meadow, shrub and forest communities, where the dominant species of the highest stand layer determines into which asso- ciation a community is classified. In the classification of species into phytosociologi- cal groups (groups of diagnostic species) we mainly refer to the Flora alpina (Aeschimann et al. 2004a,b), but rely also on our own experience. The nomencla- tural sources for the names of vascular plants were the Mala flora Slovenije (Martinčič et al. 2007) and the FloVegSi database. Hodgetts et al. (2020) was the no- menclatural source for the names of mosses and liver- worts. Šilc & Čarni (2012), Mucina et al. (2016) and DAKSKOBLER & MARTINČIČ: VEGETATION OF MOIST ROCK CREVICES AND MOIST (SLOPE) DEBRIS IN THE LIŠČAK GORGE 7FOLIA BIOLOGICA ET GEOLOGICA 64/1 – 2023 Figure 1: Approximate localities of relevés of moist rock crevices and slope debris in the basin of Liščak and its immediate surroundings (the stands of the subassociation Campanulo cespitosae-Saxifragetum aizoidis achnatheretosum calamagrostis are coloured in red). Slika 1: Približna nahajališča fitocenoloških popisov združb vlažnih skalnih razpok in pobočnega grušča v povodju Liščaka in njegovi neposredni soseščini (z rdečo so pobarvani sestoji subasociacije Campanulo cespitosae-Saxifragetum aizoidis achnatheretosum calamagrostis). DAKSKOBLER & MARTINČIČ: VEGETATION OF MOIST ROCK CREVICES AND MOIST (SLOPE) DEBRIS IN THE LIŠČAK GORGE 8 FOLIA BIOLOGICA ET GEOLOGICA 64/1 – 2023 Dakskobler & Martinčič (2020) served as nomen- clatural sources for the names of the syntaxa. Geo- graphic coordinates of relevés are determined accord- ing to the Slovenian geographic coordinate system D 48 (zone 5) based on Gauss-Krüger projection and the Bessel ellipsoid. 3 RESULTS AND DISCUSION 3.1 Moist rock crevice communities with domi- nant Pinguicula alpina and (or) Astrantia carni- olica Based on their floristic similarity the phytosociologi- cal relevés of moist rock crevices in Liščak grouped into several clusters (Figure 2) in which we identified several communities. Table 1 comprises 13 relevés with dominant vas- cular plant Pinguicula alpina and mosses Palustriella commutata and Hymenostylium recurvirostrum that can be classified into the association Astrantio carnio- licae-Pinguiculetum alpinae. Its diagnostic species are Pinguicula alpina, Astrantia carniolica, Hymenostyli- um recurvirostrum, Palustriella commutata and Peta- sites paradoxus. We recently published a more de- tailed description of this association (Dakskobler & Figure 2: Dendrogram of relevés of moist rock crevices in the Liščak gorge and its immediate vicinity (UPGMA, 1-similarity ratio). Slika 2: Dendrogram popisov vlažnih skalnih razpok v grapi Liščak in njeni okolici (UPGMA, 1-similarity ratio). Martinčič 2020). Stands from the Liščak and its vi- cinity (downstream from the confluence of the Liščak and Kneža / Knešca – Velike Luti) are very similar to the stands from other areas in the Soča Valley, but possibly occur at slightly higher elevations, from 300 m to 620 m a.s.l. They are classified into the new variant with Calamagrostis varia. The geological bed- rock is dolomite with chert and limestone with chert and (or) marlstone, and the aspect is predominantly sunny. In the relevé cluster on the left side of the dendro- gram in Figure 2 the dominant vascular plant was Astrantia carniolica and the dominant mosses were Pa- lustriella commutata and Hymenostylium recurvirostre. Pingicula alpina occurs in only a few relevés and has low medium cover, so these stands (Table 3) do not be- long to the previously described association. Slightly Asplenietea D iss im ila rit y 0,95 0,90 0,85 0,80 0,75 0,70 0,65 0,60 0,55 0,50 0,45 0,40 0,35 0,30 0,25 0,20 0,15 0,10 0,05 0,00 1 3 13 29 41 6 2 55 50 64 45 48 46 51 18 19 43 49 8 9 10 11 32 36 33 34 37 47 63 39 56 44 57 58 59 61 4 62 35 38 5 30 31 42 12 17 14 15 16 40 52 53 54 7 20 21 25 22 24 27 23 26 28 60 DAKSKOBLER & MARTINČIČ: VEGETATION OF MOIST ROCK CREVICES AND MOIST (SLOPE) DEBRIS IN THE LIŠČAK GORGE 9FOLIA BIOLOGICA ET GEOLOGICA 64/1 – 2023 similar communities of moist rock crevices in the pre- Alpine and Dinaric phytogeographical region were classified into the association Astrantio carniolicae- -Primuletum carniolicae. Almost as a rule, the domi- nant species was Primula carniolica, which has no known localities in the southern Julian Alps (Daksko- bler & Martinčič 2020). To facilitate classification of these relevés into the syntaxonomical system we made another table with the relevés with dominant vascular plant Astrantia carniolica from other regions in the Julian Alps. In part, we discussed these communities in recently pub- lished articles (Dakskobler et al. 2021, Dakskobler & Martinčič 2021a) and concluded that they cannot be classified into the association Astrantio-Pinguicule- tum nor into the association Astrantio-Primuletum carniolicae. Table 2 comprises 18 relevés with species composi- tion that is significantly different from the species composition of the Liščak relevés (Table 3), because their localities have a much larger altitudinal range. Viola biflora is very common in these relevés, but as it is not dominant in most of the relevés and therefore does not meet this criterion to be able to serve as the name-giving species of the association (Violetum bi- florae), but it can be used in name of it, as Violo bi- florae-Astrantietum carniolicae. It comprises the rele- vés of moist rock crevices with dominant Astrantia carniolica that cannot be classified into associations Astrantio-Pinguiculetum alpinae or Astrantio-Primule- tum carniolicae. Its diagnostic species are Astrantia carniolica, Viola biflora, Paederota lutea, Carex brachy- stachys, Adenostyles glabra, and Heliosperma pusillum. The nomenclatural type of the new association, holo- typus, is relevé 3 in Table 2. We distinguish four vari- ants: var. Palustriella commutata, var. Rhodothamnus chamaecistus, var. typica and var. Viola biflora. The latter comprises the last three relevés in Table 2, which show similarity with stands of the association Veronico urticifoliae-Violetum biflorae, which will be presented in more detail below. The elevation of relevés of this association spans 540 m to 1350 m (submontane-alti- montane belt), the geological bedrock is dolomite or limestone, and the aspect of their localities (Figure 3) is almost always shady. Compared to the stands in Table 2 stands in Table 3 occur at lower elevations (300–785 m), and are char- acterised by the absence of certain subalpine-alpine species, including montane-(sub) alpine species Viola biflora and Heliosperma pusillum. According to domi- Figure 3: Localities of stands of the association Violo biflorae-Astrantietum carniolicae Slika 3: Nahajališča sestojev subasociacije Violo biflorae-Astrantietum carniolicae DAKSKOBLER & MARTINČIČ: VEGETATION OF MOIST ROCK CREVICES AND MOIST (SLOPE) DEBRIS IN THE LIŠČAK GORGE 10 FOLIA BIOLOGICA ET GEOLOGICA 64/1 – 2023 nant species and results of hierarchical classification (Table 11, Figure 10), we classify this stands into the new association Palustriello commutati-Astrantietum carniolicae ass. nov. hoc loco. Its nomenclatural type, holotypus, is relevé 14 in Table 3. The species composi- tion of relevés 1–4 is different, so they are for now clas- sified into the variant Palustriello-Astrantietum carni- olicae var. Carex mucronata. Its differential species are Carex mucronata, Aster bellidiastrum, Hydrogonium croceum (Barbula crocea) and Valeriana saxatilis. The locality of these relevés is the right bank of the Kneža downstream of the confluence with the Liščak, at the beginning of the ravine of Velike Luti; the geological bedrock is dolomite with chert and the aspect is dis- tinctly shady; the average elevation is 320 m. In the stands of this variant we collected and determined also two relatively rare moss species. Trichostomum crispu- lum is a meridional-temperate species of which we have relatively limited recent data for Slovenia (Mar- tinčič 2018: 58). Microlejeunea ulicina is distributed mainly in North America, North Africa, the Mediter- ranean and western Europe to Norway. It occurs mainly on tree bark and decaying wood on very airy, moist areas. In Slovenia it is red-listed as vulnerable (Martinčič 2016: 109). Other relevés in Table 3 belong to the typical vari- ant Palustriello-Astrantietum carniolicae var. typica and their stands are characterised by abundant occur- rence of Palustriella commutata and Hymenostylium recurvirostre. The elevation of the relevés is 300 m to 785 m, the aspect is shady and sunny, the geological bedrock is predominantly limestone with admixture of marlstone and chert. 3.2 Moist rock crevice communities with domi- nant Veronica urticifolia, Saxifraga cuneifolia or S. aizoides Table 4 comprises communities of moist rocks in the Liščak gorge in which Pinguicula alpina or Astrantia carniolica are either absent or non-dominant, but the moss layer is very rich. These stands are classified into the new association Palustriello commutati-Veronice- tum urticifoliae. Its nomenclatural type, holotypus, is relevé 1 in Table 4. The diagnostic species of the asso- ciation are Palustriella comutata, Conocephalum coni- cum, Veronica urticifolia, Asplenium trichomanes, Ga- leobdolon flavidum and Senecio ovatus. The elevation of the relevés spans 350 m to 950 m, the geological bed- Figure 4: Distribution of Veronica urticifolia in Slovenia (FloVegSi database) Slika 4: Razširjenost vrste Veronica urticifolia v Sloveniji po podatkih baze FloVegSi DAKSKOBLER & MARTINČIČ: VEGETATION OF MOIST ROCK CREVICES AND MOIST (SLOPE) DEBRIS IN THE LIŠČAK GORGE 11FOLIA BIOLOGICA ET GEOLOGICA 64/1 – 2023 rock is mixed, composed of limestone, marlstone and chert, in places also claystone; the aspect is predomi- nantly shady. Veronica urticifolia is a south-European montane species, a character species of the suballiance Abieti- -Piceenion (Aeschimann et al. 2004b: 216). In Slovenia it is widespread in most submontane-montane-subal- pine areas. It occurs in many forest, especially spruce and beech as well noble hardwood communities. Its sites are often shady and slightly moist rocks, both cal- careous and non-calcareous, and in the broadest sense it can be considered also a character species of com- munities of moist rock crevices. Stands of the association Palustriello commutati- -Veronicetum urticifoliae characteristically occur in a distinctly forested environment and their species com- position therefore comprises a good number of charac- ter species of alliances Tilio-Acerion, Aremonio-Fagion and order Fagetalia sylvaticae, i.e. species that are fre- quent in the surrounding forest communities. Table 4 also includes an additional relevé with an exclusively moss community, which is provisionally classified into the association Cratoneuretum commutati. Table 5 comprises stands classified into the recently described association Campanulo cespitosae-Saxifrage- tum aizoidis (Dakskobler & Martinčič 2020). It is characterised by the occurrence of the subalpine-alpine species Saxifraga aizoides, a character species of spring communities (Montio-Cardaminetea) in the moist shady rocks in gorges of the submontane and lower montane belt. The diagnostic species of the association are Saxifraga aizoides, Campanula cespitosa, Molinia arundinacea, Palustriella commutata, Calamagrostis varia and Marchantia quadrata (Preissia quadrata). But for the last (Marchantia quadrata) the relevés from the Liščak gorge comprise all of the listed species. Their ecology, however, is different. They were found at eleva- tions spanning 725 m to 785 m, the geological bedrock is mainly limestone with admixture of claystone and chert, and the aspect is generally sunny. This is a dis- tinctly erosion area (the nearest toponym is Pod Jamo) next to the Luknova Grapa gorge (this gorge starts under Mt. Luken and flows into the Liščak at Sopota). One relevé is from the right bank of the Liščak, the lo- cality is at the Gradnikova Grapa gorge (under Mt. Gradnik), also in an erosion area, where the bedrock consists of limestone, marlstone and chert (see Figure 1). The sites are characterised by unconsolidated rock, with rock fragments that are not solid, but broken, loose and slightly moist. The species composition shows Figure 5: Approximate localities of stands of the association Veronico urticifoliae-Saxifragetum cuneifolii on the map of Slovenia Slika 5: Približna nahajališča sestojev asociacije Veronico urticifoliae-Saxifragetum cuneifolii na zemljevidu Slovenije DAKSKOBLER & MARTINČIČ: VEGETATION OF MOIST ROCK CREVICES AND MOIST (SLOPE) DEBRIS IN THE LIŠČAK GORGE 12 FOLIA BIOLOGICA ET GEOLOGICA 64/1 – 2023 a good share of character species of screes and dry grasslands. These relevés can undoubtedly be classified into a new, drier form of this community, into the sub- association Campanulo-Saxifragetum aizoidis ach- natheretosum calamagrostis. Its differential species are Achnatherum calamagrostis, Tortella tortuosa, Saxifra- ga crustata, Hieracium glaucum and Calamintha einse- leana, which are good indicators of these site conditions on still slightly moist, debris covered, predominantly sunny rocks. The nomenclatural type of the new subas- sociation, holotypus, is relevé 5 in Table 5. Table 6 comprises relevés of slightly moist, shady rock crevices in the forest belt, where Saxifraga cune- ifolia occurs, whereas the previously discussed domi- nant species of moist rocks (except for Veronica urtici- folia) are usually absent. Saxifraga cuneifolia frequently occurs in stony, shady, montane spruce, beech and fir- beech forests. Such stands were found also in the gorge of the Liščak, but our table comprises also similar rele- vés from other areas. Their approximate localities are shown in Figure 5. These stands are classified into the new associa- tion Veronico urticifoliae-Saxifragetum cuneifolii. Its diagnostic species are Saxifraga cuneifolia, Veronica urticifolia, Asplenium trichomanes, Exertotheca crispa (Neckera crispa) and Valeriana tripteris. The nomen- clatural type, holotypus, of the new association is rele- vé 5 in Table 6. The distribution and sites of the south-European montane species Saxifraga cuneifolia in Slovenia was described in detail several years ago (Dakskobler 2015). Its distribution is very similar to the distribution of Veronica urticifolia, which is also a character species of spruce forests; they occur on similar sites and fre- quently in the same communities, including commu- nities of shady rocks on mixed calcareous-silicate bed- rock. Stands of the association Veronico urticifoliae- -Saxifragetum cuneifolii are slightly similar to the stands of the association Palustriello-Veronicetum urti- cifoliae, but the latter have a very different composition and medium coverage of the moss layer. The elevation of the localities ranges between 150 m and 1060 m, and the geological bedrock is varied as well: limestone, claystone, marlstone, limestone with chert and marl, rarely also dolomite and breccia. Stands of this associa- tion were found on all, albeit predominantly shady as- pects. Most of the localities are situated in gorges and depressions in the beech belt. This is reflected also in the species composition, which features a number of companion species characteristic for beech, maple, beech-oak and spruce forests. We distinguish two variants: var. typica, which has no special differential species, and var. Galeobdolon flavidum (its differential species include Fissidens dubius and Phyllitis scolopen- drium), which characterises slightly moister rocks with more moss species and more companion species of beech forests. 3.3 Moist rock crevice communities with domi- nant Aster bellidiastrum and (or) Viola biflora The south-European montane species Aster bellidia- strum is a character species of subalpine-alpine calcar- eous grasslands from the class Elyno-Seslerietea (Ae- schimann et al. 2004b: 432), but frequently occurs also in communities of moist rock crevices (Daksko- bler & Martinčič 2020), including those that we have already described or those that we are to describe herein: Astrantio-Pinguiculetum alpinae, Paederoto lu- teae-Astrantietum carniolicae, Campanulo cespitosae- -Saxifragetum aizoidis and Veronico urticifoliae-Viole- tum biflorae. Table 7 comprises the phytosociological relevés (including a few relevés from the Liščak gorge) in which this species is the dominant vascular plant. Most of these relevés were made on riparian rocks that are periodically exposed to water from creeks or the river, in part also on rock walls next to watercourses in gorges and ravines, some also outside the pre-Alpine– Alpine region (Figures 6 and 7). Stands in Table 7 are classified into the new asso- ciation Calamagrostio variae-Asteretum bellidiastri ass. nov. hoc loco. Its diagnostic species are Aster bellidia- strum and Calamagrostis varia, the only vascular plants with a constancy of more than 50% in 28 rele- vés; their counterparts among mosses are Ctenidium molluscum and Tortella tortuosa. The nomenclatural type, holotypus, of the new as- sociation is relevé 7 in Table 7. We distinguish two vari- ants, var. Brachythecium rivulare (differential species include other hygrophilous mosses: Hygrohypnum luri- dum, Didymodon spadiceus, Pedinophyllum interrup- tum, Plagiumnium rostratum, and Veronica urticifolia among vascular plants) on slightly moister sites, and var. Tortella tortuosa on slightly drier sites. Within this variant (Table 7) we can exclude the subvariant with Sesleria caerulea and the subvariant with Petasites pa- radoxus. One of the characteristics of the stands of this association is a substantial number of so-called acci- dental species, which occur due to the position of their sites near watercourses. Here we determined altogether 138 species, more than in any other discussed chasmo- phytic community. Due to the immediate vicinity of watercourses they become colonised by plants that are otherwise characteristic for forest, grassland and ru- deral communities. However, none of these species on DAKSKOBLER & MARTINČIČ: VEGETATION OF MOIST ROCK CREVICES AND MOIST (SLOPE) DEBRIS IN THE LIŠČAK GORGE 13FOLIA BIOLOGICA ET GEOLOGICA 64/1 – 2023 Figure 6: Distribution of Aster bellidiastrum in Slovenia (FloVegSi database). Slika 6: Razširjenost vrste Aster bellidiastrum v Sloveniji po podatkih v bazi FloVegSi. Figure 7: Localities of stands of the association Calamagrostio variae-Asteretum bellidiastri on the map of Slovenia. Slika 7: Nahajališča sestojev asociacije Calamagrostio variae-Asteretum bellidiastri na zemljevidu Slovenije. DAKSKOBLER & MARTINČIČ: VEGETATION OF MOIST ROCK CREVICES AND MOIST (SLOPE) DEBRIS IN THE LIŠČAK GORGE 14 FOLIA BIOLOGICA ET GEOLOGICA 64/1 – 2023 these riparian rocks has a high frequency and they were recorded only on one or a few relevés. The elevation of the localities ranges from 235 m (at Bača under the hamlet of Slatne) to 780 m (Gačnik in Trebuša) or 785 m (under the Sopot waterfall in Liščak). The geological bedrock is limestone, dolomite, limestone or dolomite with chert, in places also with marl, rarely breccia or conglomerate. Several localities have both a shady and sunny aspect. In the stands of this association we col- lected and determined also two, relatively rare moss species, namely Trichostomum crispulum (on riparian rocks by the Bača River under the hamlet of Slatne, quadrant 9849/1) and Palustriella decipiens (riparian rocks by the Soča River at Srpenica, at 315 m a.s.l., which is one of the lowland localities of this species, otherwise distributed mainly in the subalpine-alpine belt – comp. Dakskobler & Martinčič 2021b: 63). Viola biflora is an Arctic-alpine species, a charac- ter species of the class Betulo-Alnetea viridis (Aeschi- mann et al. 2004a: 441). Its distribution in Slovenia is very similar to the distribution of Aster bellidiastrum (Figure 8). It is relatively frequent in communities of moist rock crevices (Dakskobler & Martinčič 2020), in our case mainly in the stands of the association Violo biflorae-Astrantietum carniolicae. In Table 8 we ar- ranged the relevés of moist rock crevices and rock shel- ters, where this species is the dominant vascular plant in terms of constancy and medium coverage. Thus it can serve as the name-giving species and these stands are therefore classified into the new association Vero- nico urticifoliae-Violetum biflorae. Its diagnostic spe- cies are Viola biflora, Veronica urticifolia and Cono- cephalum conicum. The nomenclatural type of the new association, holotypus, is relevé 11 in Table 8. Even though there are some differences between the relevés in this table, it would be difficult to describe them at the rank of lower synsystematic units. Relevés 12–15 are classified into the variant with Astrantia carniolica and show certain similarity with the stands of the vari- ant Violo biflorae-Astrantietum carniolicae var. Viola biflora. The second variant, evident from Table 8, is the variant with Palustriella commutata (relevés 16–25 in Table 8). Within this variant relevés 18 and 19 in Table 8 could be treated as the subvariant with Primula auri- cula. In our paper on the vegetation of the Prodar Gorge (Dakskobler & Martinčič 2021) we classified these two relevés into the provisional association Pri- mulo auriculae-Violetum biflorae. As a valid descrip- tion of such an association would require more relevés, Figure 8: Distribution of Viola biflora in Slovenia (FloVegSi database). Slika 8: Razširjenost vrste Viola biflora v Sloveniji po podatkih v bazi FloVegSi. DAKSKOBLER & MARTINČIČ: VEGETATION OF MOIST ROCK CREVICES AND MOIST (SLOPE) DEBRIS IN THE LIŠČAK GORGE 15FOLIA BIOLOGICA ET GEOLOGICA 64/1 – 2023 we provisionally assign these relevés to the association Veronico-Violetum biflorae. Other relevés in Table 9 are classified into the variant typica. We did not find the localities of Viola biflora in the Liščak gorge, nor did we find localities of the stands of this association, which are otherwise common to the Vintgar Gorge at Podhom (Dakskobler et al. 2021). Approximate localities of these stands are shown in Figure 9. They were found at elevations ranging from 210 m (the Idrijca valley at Stopnik) to 1000 m (Curk Waterfall under Mt. Krn), most often on dolomite as well as on dolomite with chert, limestone, limestone with chert and (or) marlstone, even on rock debris. Most localities have a shady aspect. Table 9 comprises ten phytosociological relevés that grouped separately from the relevés of the associa- tion Veronico-Violetum biflorae (in hierarchical classi- fication) and cannot be classified into this association. Due to the insufficient number of relevés and their considerable diversity they are classified into only pro- visionally described syntaxa. Relevés 1–9 in Table 9 are provisionally classified into the association Paederoto luteae-Violetum biflorae nom. prov. Its diagnostic spe- cies are Viola biflora and Paederota lutea. Relevés 1–4 in this Table (we made them at the Curk Waterfall by the Kozjak stream under Mt. Krnčica) are classified into the variant with Trisetum argenteum. Its differen- tial species include Saxifraga aizoides and Marchantia quadrata (Preissia quadrata). Relevé 5 (from the Kozi- jska Grapa gorge in the Trebuša Valley) is classified into the variant with Saxifraga rotundifolia. Relevés 6 and 7 (the first was made under Mt. Mangart and the second in the forest reserve Apica above Zapoden) are classified into the variant with Cystopteris fragilis (its differential species is also Conocephalum conicum). Relevé 8 from Gozdec in the Kanin Mountains is clas- sified into the variant with Campanula carnica (its dif- ferential species is also Cyclamen purpurascens) and relevé 9 (from the Lopučnica valley) into the variant with Saxifraga sedoides (its differential species include Cystopteris montana and Heliosperma pusillum). With its full floristic composition relevé 10 stands out the most from other relevés with dominant Viola biflora. It was made by a small concavity under Mt. Mangart. It is provisionally classified into the association Cerastio subtriflorae-Violetum biflorae nom. prov. Its diagnostic species are Viola biflora, Cerastium subtriflorum, Fe- stuca nitida, Doronicum glaciale and Sanonia uncinata. Figure 9: Approximate localities of stands of the association Veronico urticifoliae-Violetum biflorae on the map of Slovenia Slika 9: Približna nahajališča sestojev asociacije Veronico urticifoliae-Violetum biflorae na zemljevidu Slovenije DAKSKOBLER & MARTINČIČ: VEGETATION OF MOIST ROCK CREVICES AND MOIST (SLOPE) DEBRIS IN THE LIŠČAK GORGE 16 FOLIA BIOLOGICA ET GEOLOGICA 64/1 – 2023 3.4 Other chasmophytic communities above the Liščak gorge Table 10 comprises eleven phytosociological relevés of chasmophytic communities in the Liščak gorge that cannot be classified into any of the associations de- scribed herein. As we do not have a sufficient number of relevés to adequately classify them into a syntaxo- nomic system, they were given provisional names. The relevés vary between each other and in our opinion only relevés 9–11 in Table 10 can be assigned to the al- liance Astrantio carniolicae-Paeaderotion luteae. Rele- vés 9 and 10 are provisionally classified into the asso- ciation Valeriano tripteridis-Veronicetum urticifoliae nom. prov. (see Dakskobler & Martinčič 2023), and relevé 11 into the provisional association Palustriello commutati-Phyllitidetum scolopendrii nom. prov. The elevation of the relevés ranges between 350 m and 530 m, the aspect is shady, and the geological bedrock is limestone with admixture of chert or marlstone. For the time being, relevés 1–5 are classified into the alli- ance Physoplexido comosae-Saxifragion petraeae, more specifically relevé 1 into the provisional association Sileno hayekianae-Campanuletum carnicae nom. prov., relevés 2 and 3 into the provisional association Seslerio caeruleae-Sedetum albi nom. prov., and relevés 4 and 5 into the provisional association Arabido alpinae-Sede- tum albi nom. prov. They were made at elevations be- tween 770 m and 1100, on slopes above the gorge rath- er than in the gorge, on sunny aspects; the bedrock is predominantly limestone. Relevés 6–8 are provision- ally classified into the association Tortello tortuosae- -Asplenietum trichomanis nom. prov. These relevés were made on sunny aspects at elevations between 410 m and 560 m; the geological bedrock is limestone ad- mixed with claystone or marlstone. 3.5 Survey of described syntaxa and argumenta- tion for the alliance Astrantio carniolicae- Paederotion luteae Chasmophytic communities on moist calcareous or mixed calcareous-silicate bedrock in the submontane- altimontane (subalpine) belt in the Southeastern Alps and in the northern part of the Dinaric Alps have so far been classified into the alliance Cystopteridon fragi- lis Richard 1972. Recently (Dakskobler in Martin- čič 2020) we listed the reasons why they could also be classified into a vicariant alliance of the alliance Physo- plexido comosae-Saxifragion petraeae Mucina et Theu- rillat 2015. We proposed it be named Astrantio carnio- licae-Paederotion luteae nom. prov. The material pro- cessed for this paper (see Table 11) provided sufficient grounds for its valid description. Astrantio carniolicae-Paederotion luteae all. nov. hoc loco. The nomenclatural type, holotypus, of the new al- liance is the association Astrantio carniolicae-Primule- tum carniolicae Dakskobler & Martinčič 2020 (Dakskobler & Martinčič 2020, Table 1). It is described as an alliance of moist rock crevices in the Southeastern Alps and the northern part of the Dinaric Alps. The diagnostic species of the new alliance are di- vided into phytogeographical-ecological and ecological. The former comprise Astrantia carniolica, Paede- rota lutea and Primula carniolica, to a smaller extent also diagnostic species of alliances Physoplexido como- sae-Saxifragion petraeae and Aremonio-Fagion as well as other southeastern-Alpine–northern-Dinaric spe- cies that individually occur in their stands. These suf- ficiently differentiate them from similar communities of the alliance Cystopteridion. The ecological diagnostic species are Aster bellidi- astrum, Hydrogonium croceum (Barbula crocea), Carex brachystachys, Cystopteris fragilis, Asplenium viride, Eucladium verticillatum, Fissidens dubius, Hyme- nostylium recurvirostrum, Apopellia endiviifolia (Pellia endiviifolia), Palustriella commutata, Marchantia qua- drata (Preissia quadrata), Pingiucula alpina, Orthothe- cium rufescens, Saxifraga aizoides, Tofieldia calyculata, Valeriana saxatilis, V. tripteris and Viola biflora. These are character species of various alliances and classes, but differentiate the studied stands against communi- ties from the vicariant alliance Physoplexido comosae- -Saxifragion petraeae. In addition to its holotype association and the be- low-listed associations discussed in this paper, the new alliance also includes associations Ranunculo traunfell- neri-Paederoteum luteae Surina 2005, Primuletum car- niolicae Accetto 2008, Neckero crispae-Campanuletum justinianae Accetto 1995, Triseto argentei-Leontodonte- tum brumatii Dakskobler, Seliškar et Vreš 2012, Phyte- umato columnae-Primuletum carniolicae Dakskobler et Martinčič 2020 and Primulo carniolicae-Potentille- tum clusianae Dakskobler & Martinčič 2020. The synthetic table (Table 11) comprises eight col- umns that represent only the communities of moist rocks validly described in this paper. With hierarchical classification we obtained the dendrogram in Figure 10. DAKSKOBLER & MARTINČIČ: VEGETATION OF MOIST ROCK CREVICES AND MOIST (SLOPE) DEBRIS IN THE LIŠČAK GORGE 17FOLIA BIOLOGICA ET GEOLOGICA 64/1 – 2023 The syntaxa that are the most similar in terms of floristic similarity, which takes into account the con- stancy of all recorded species, are Astrantio carnioli- cae-Pinguiculetum alpinae, Palustriello commutati- -Astrantietum carniolicae, Violo biflorae-Astrantietum carniolicae and Veronico urticifoliae-Violetum biflorae, and syntaxa Palustriello commutati-Veronicetum urti- cifoliae and Veronico urticifoliae-Saxifragetum cuneifo- lii. The syntaxa that stand out the most are Calamagro- stio variae-Asteretum bellidiastri and Campanulo cespi- tosae-Saxifragetum aizoidis achnatheretosum calama- grostis. Communities with dominant Pinguicula alpi- na, Astrantia carniolica and (or) Viola biflora, and communities with dominant Veronica urticifolia and (or) Saxifraga cuneifolia are more similar. Some of the described syntaxa could be grouped based solely on their floristic similarity, but based on the dominant species with the highest medium coverage criterion this is no longer possible. Table 11 also shows that all described associations comprise a sufficient number of diagnostic species to be classified into the alliance Astrantio-Paederotion luteae. 3.6 Classification of the researched communi- ties into the syntaxonomical system Asplenietea trichomanis (Br.-Bl. in Meier et Br.-Bl. 1934) Oberd. 1977 Potentilletalia caulescentis Br.-Bl. in Br.-Bl. et Jenny 1926 Figure 10: Dendrogram of communities of moist rock crevices in the Julian Alps and their foothills (UPGMA, 1-similarity ratio). Slika 10: Dendrogram združb vlažnih skalnih razpok v Julijskih Alpah s prigorjem (UPGMA, 1-similarity ratio). Legend (Legenda): AcPa Astrantio carniolicae-Pinguiculetum alpinae (Table 1) PcAc Palustriello commutati-Astrantietum carniolicae (Table 3) VbAc Violo biflorae-Astrantietum carniolicae (Table 2) VuVb Veronico urticifoliae-Violetum biflorae (Table 8) PcVu Palustriello commutati-Veronicetum urticifoliae (Table 4) VuSc Veronico urticifoliae-Saxifragetum cuneifolii (Table 6) CvAb Calamagrostio variae-Asteretum bellidiastri (Table 7) CcSaac Campanulo cespitosae-Saxifragetum aizoidis achnatheretosum calamagrostis (Table 5) DAKSKOBLER & MARTINČIČ: VEGETATION OF MOIST ROCK CREVICES AND MOIST (SLOPE) DEBRIS IN THE LIŠČAK GORGE 18 FOLIA BIOLOGICA ET GEOLOGICA 64/1 – 2023 Astrantio carniolicae-Paederotion luteae all. nov. hoc loco Astrantio carniolicae-Pinguiculetum alpinae Dak- skobler et Martinčič 2020 var. Calamagrostis varia Violo biflorae-Astrantietum carniolicae ass. nov. hoc loco var. Hymenostyilium recurvirostrum var. Rhodothamnus chamaecistus var. typica var. Viola biflora Palustriello commutati-Astrantietum carniolicae ass. nov. hoc loco var. Carex mucronata var. typica Palustriello commutati-Veronicetum urticifoliae ass. nov. hoc loco Campanulo cespitosae-Saxifragetum aizoidis Dak- skobler et Martinčič 2020 achnatheretosum calamagrostis subas. nov. hoc loco Veronico urticifoliae-Saxifragetum cuneifolii ass. nov. hoc loco var. typica var. Galeobdolon flavidum Calamagrostio variae-Asteretum bellidiastri ass. nov. hoc loco var. Brachythecium rivulare var. Tortella tortuosa Veronico urticifoliae-Violetum biflorae ass. nov. hoc loco var. typica var. Astrantia carniolica var. Palustriella commutata subvar. Primula auricula Paederoto luteae-Violetum biflorae nom. prov. var. Trisetum argenteum var. Saxifraga rotundifolia var. Campanula carnica var. Saxifraga sedoides Cerastio subtriflorae-Violetum biflorae nom. prov. Valeriano tripteridis-Veronicetum urticifoliae nom. prov. Palustriello commutati-Phyllitidetum scolopendrii nom. prov. Physoplexido comosae-Saxifragion petraeae Mucina et Theurillat 2015 Sileno hayekianae-Campanuletum carnicae nom. prov. Seslerio caeruleae-Sedetum albi nom. prov. Arabido alpinae-Sedetum albi nom. prov. Tortello tortuosae-Asplenietum trichomanis nom. prov. Montio-Cardaminetea Br.-Bl. & Tx. ex Klika et Hadač 1944 Montio-Cardaminetalia Pawłowski et al. 1928 Cratoneurion commutati Koch 1928 Cratoneuretum commutati Aichinger 1933 3.7 Tall-herb community with dominant Lunar- ia rediviva on colluvial-deluvial soils in gorges and ravines One of the specifics of the vegetation in the Liščak gorge is also tall herbs on colluvial soils, at the foot of very steep slopes, especially on the right bank of the creek in its lower and medium course. The dominant tall herb species here is Lunaria rediviva. Having re- corded similar communities also elsewhere in western Slovenia we prepared a table (Table 12) with 40 relevés and arranged it with hierarchical classification. Most of the relevés in the table can be classified into the new association Lamio orvalae-Lunarietum redivivae ass. nov. hoc loco. Its diagnostic species are Lunaria redivi- va, Galeobdolon flavidum, Sambucus nigra, Phyllitis scolopendrium, Lamium orvala, Cardamine pentaphyl- los, Plagiomnium undulatum and Thamnobryum alo- pecurum. In a recent article we already described it as a provisional association (Dakskobler & Martinčič 2021). Lunaria rediviva is a species characteristic for noble hardwood forests from the alliance Tilio-Aceri- on. It occurs in most parts of Slovenia (Figure 11), in noble hardwood communities as well as in riparian woodland, for example in stands of associations Lamio orvale-Salicetum albae, Lamio orvalae-Alnetum inca- nae and Lamio orvalae-Alnetum glutinosae. On smaller, 10 m2 to 100 m2 large areas, its stands are syndynamically related in particular to communi- ties of noble hardwood forests from the alliance Tilio- -Acerion (associations Fraxino orni-Aceretum pseudo- platani, Veratro nigri-Fraxinetum, Hacquetio-Fraxine- tum, Lamio orvalae-Aceretum, Omphalodo-Aceretum) and mesophilous beech communities from the alliance Aremonio-Fagion (Ornithogalo pyrenaici-Fagetum, La- mio orvalae-Fagetum, Omphalodo-Fagetum, Isopyro- -Fagetum). In places, successional development pro- ceeds across elderberry shrubs from the subassociation Lamio orvalae-Sambucetum nigrae Poldini et Vidali 1995 lunarietosum redivivae (Table 13), which is de- scribed below. Approximate localities of recorded stands of this association are shown in Figure 12. They were made in the Alpine, pre-Alpine, sub-Mediterranean, Dinar- ic and pre-Dinaric phytogeographical regions. The DAKSKOBLER & MARTINČIČ: VEGETATION OF MOIST ROCK CREVICES AND MOIST (SLOPE) DEBRIS IN THE LIŠČAK GORGE 19FOLIA BIOLOGICA ET GEOLOGICA 64/1 – 2023 Figure 11: Distribution of Lunaria rediviva (FloVegSi database). Slika 11: Razširjenost vrste Lunaria rediviva v Sloveniji (podatkovna baza FloVegSi). elevation of the relevés ranges from 100 m (Petnik gorge at Branik, see also Dakskobler & Poldini 2021) to 1050 m (Kacenpoh gorge at Podbrdo). The average number of species per relevé is 21, and the number of species ranges between 9 (rockfall materi- al in the Doblarca gorge) and 45 (the relevé at the con- tact of the colluvium and alluvium in the Prodar gorge at Podbrdo), depending on the size of the com- munity surface area and certain ecological factors (rockiness, rock debris or gravel, admixture of non- calcareous rocks). Even though our relevés comprise also Aruncus di- oicus they cannot be classified into the association Arunco vulgaris-Lunarietum redivivae Sádlo et Petřík in Chytrý 2009, because its stands belong in a group of ruderal communities and are classified into the alli- ance Impatienti noli-tangere-Stachyion sylvaticae and class Galio-Urticetea (Sádlo & Petřík 2009). The studied association is classified into the alliance Arun- co-Petasition, which comprises tall herb communities on stony, nutrient-rich soils on steep slopes in the montane and upper montane belt in the Alps. The no- menclatural type, holotypus, of the new association Lamio orvalae-Lunarietum redivivae is relevé 8 in Table 12. Relevé 39 in Table 12 (the locality under Mt. Črna Gora above mountain pasture Za Liscem in the vicin- ity of Mt. Črna Prst) is classified into the provisional subassociation Doronico austriaci-Adenostyletum alli- ariae lunarietosum redivivae nom. prov., and relevé 40 in Table 12 (the bottom of the frost hollow above Pra- dol between Mt. Mija and Mt. Ljubija) into the provi- sional association Lunario redivivae-Saxifragetum ro- tundifoliae nom. prov. The studied communities with dominant Lunaria rediviva are classified into higher syntaxonomic units as follows: Mulgedio-Aconitetea Hadač et Klika in Klika et Hadač 1944 Adenostyletalia alliariae Br.-Bl. 1926 Adenostylion alliariae Br.-Bl. 1926 Doronico austriaci-Adenostyletum alliariae Horvat ex Horvat et al. 1974 lunarietosum redivivae nom. prov. Lunario redivivae-Saxifragetum rotundifoliae nom. prov. Petasito-Chaerophylletalia Morariu 1967 Arunco-Petasition albi Br.-Bl. et Sutter 1977 Lamio orvalae-Lunarietum redivivae ass. nov. hoc loco DAKSKOBLER & MARTINČIČ: VEGETATION OF MOIST ROCK CREVICES AND MOIST (SLOPE) DEBRIS IN THE LIŠČAK GORGE 20 FOLIA BIOLOGICA ET GEOLOGICA 64/1 – 2023 Figure 12: Localities of stands of the association Lamio orvalae-Lunarietum redivivae on the map of Slovenia. Slika 12: Nahajališča sestojev asociacije Lamio orvalae-Lunarietum redivivae na zemljevidu Slovenije. 3.8 Shrub communities of Sambucus nigra and Lamium orvala (Lamio orvalae-Sambucetum ni- grae) in Slovenia As tall herbs with dominant Lunaria rediviva (Lamio orvalae-Lunarietum redivivae) are one of the specifics of the vegetation at Liščak and are often syndynami- cally related to Sambucus nigra shrubs, we hierarchi- cally classified 25 relevés of these shrubs and arranged them in Table 13 (their approximate localities are shown in Figure 13). Most, at least 24 of them, can be classified into the association Lamio orvalae-Sambuce- tum nigrae, which was described by Poldini & Vidali (1995), who classified it into the class Querco-Fagetea and order Prunetalia spinosae. They divided it into two geographical variants: var. geogr. Helleborus odorus – the Friulian race (razza friulana), and var. geogr. Helle- bous istriacus – the Karstic race (razza carsica), and listed Lamium orvala as the only diagnostic species. Some of our relevés were made in gaps of Illyrian beech forests from the alliance Aremonio-Fagion, most often on potential sites of associations Lamio orvalae- -Fagetum, Arunco-Fagetum or Ornithogalo-Fagetum, and some on very stony, gravelly sites (colluvium, rockfall material) under rock walls, where we find po- tential sites of noble hardwood communities from the alliance Tilio-Acerion. Floristically, they are clearly different from the Friuli relevés, most notably in the frequency and high coverage of Lunaria rediviva in most of the relevés. Diagnostic species of the association are Sambucus nigra, Lamium orvala, Phyllitis scolopendrium and Geranium robertianum. Anemone trifolia is the geo- graphical differential species (the first three relevés in Table 13 from Mt. Donačka Gora and relevé 25 from Istria do not belong in this geographical variant). Relevés 1–22 in Table 13 are classified into the new subassociation lunarietosum redivivae subass. nov. hoc loco. Its differential species are Lunaria rediviva, Circaea lutetiana and Cardamine pentaphyllos. Its no- menclatural type, holotypus, is relevé 11 in Table 13. The elevation of the localities ranges from 230 m to 780 m (from the colline to the lower montane belt) and the aspect is predominantly shady. The soil is colluvial-delluvial, in places also Chromic Cambisols or eutric soils. We distinguish several variants. The localities of the stands of the variant with Impatiens noli-tangere (relevés 1–6) are in the old growth forest remnant under Mt. Donačka Gora, in the Zapoška Grapa gorge under Mt. Porezen, and in the Vintgar Gorge at Pod- hom. The differential species of this variant are also Milium effusum and Urtica dioica, the latter mainly on account of high medium coverage. The listed spe- DAKSKOBLER & MARTINČIČ: VEGETATION OF MOIST ROCK CREVICES AND MOIST (SLOPE) DEBRIS IN THE LIŠČAK GORGE 21FOLIA BIOLOGICA ET GEOLOGICA 64/1 – 2023 cies indicate nutrient-rich sites. The localities of the stands of the variant with Cardamine trifolia (its dif- ferential species are also Fraxinus excelsior and Tilia platyphyllos), relevés 7–14, are in the Avšček Gorge (Banjšice) on the sites of associations Ornithogalo-Fa- getum and Lamio orvalae-Fagetum. Relevés of the stands of the variant with Scopolia carniolica (relevés 15–20) were made in the gorges of Avšček, Raskovec (Žirovnica near Žiri), Pekel at Postaja and in the Vratarska Grapa gorge (the latter two are on the northern rim of the Banjšice Plateau above the Idrijca Valley). The stands of this variant occur on aceretal sites, in the vicinity of stands of associations Veratro nigri-Fraxinetum or Hacquetio-Fraxinetum. Relevés 21 and 22 are classified into the variant with Leuco- jum vernum (its locality is a small gorge at the village of Svino in the Kobarid area). Relevés 23 and 24 do not belong in the subassociation lunarietosum redivi- vae; it is classified only at the rank of variant, namely var. Adenostyles glabra. Its locality is Kašana in the Zadlaščica Gorge, and its site is the rockfall material under the wall. Urtica dioica has high medium cover- age. Relevé 25 in Table 13 also does not belong in the subassociation lunarietosum, possibly not even into the association Lamio orvalae-Sambucetum, because it does not comprise its diagnostic species. For the time being it is still treated in the framework of this association as a special variant with Ruscus aculeatus (the differential species is also Primula vulgaris). In terms of species composition this stand is still more similar to the stands of this association than to the stands of the association Bryonio dioicae-Sambuce- tum nigrae, which was also described by Poldini & Vidali (1995). Its locality is Branski Bošk in a shady gorge under the village of Korte in Istria, in the vicin- ity of localities of the association Ornithogalo-Carpi- netum betuli. According to our findings, the association Lamio orvalae-Sambucetum nigrae is classified into higher syntaxonomic units as follows: Querco-Fagetea Br.-Bl. et Vlieger in Vlieger 1937 (Car- pino-Fagetea sylvaticae Jakucs ex Passarge 1968) Fagetalia sylvaticae Pawłowski 1928 Tilio-Acerion Klika 1955 Lamio orvalae-Sambucetum nigrae Poldini et Vi- dali 1995 lunarietosum redivivae subass. nov. var. Impatiens noli-tangere var. Cardamine trifolia var. Scopolia carniolica var. Leucojum vernum var. Adenostyles glabra (prov.) var. Ruscus aculeatus (prov.) Figure 13: Approximate localities of stands of the association Lamio orvalae-Sambucetum nigrae on the map of Slovenia. Slika 13: Nahajališča sestojev asociacije Lamio orvalae-Sambucetum nigrae na zemljevidu Slovenije. DAKSKOBLER & MARTINČIČ: VEGETATION OF MOIST ROCK CREVICES AND MOIST (SLOPE) DEBRIS IN THE LIŠČAK GORGE 22 FOLIA BIOLOGICA ET GEOLOGICA 64/1 – 2023 The Liščak gorge in the Southern Julian Alps is a natu- ral feature of local importance, mainly on account of its geomorphological and geological specifics. Its water- shed area is almonst entirely forested, uninhabited and with little direct human impact. There are no artificial barriers on the creek, only remnants of former mills and a few signs of former charcoal production and floating of wood. In addition to numerous localities of protected Taxus baccata and endemic Aconitum angu- stifolium, communities of moist rock crevices also fea- ture among the botanical curiosities of this gorge. They are classified into the following associations: Astrantio carniolicae-Pingiuculetum alpinae, Palustriello commu- tati-Astrantietum carniolica, Palustriello commutati-Ve- ronicetum urticifoliae, Veronico urticifoliae-Saxifrage- tum cuneifolii, Calamagrostio varie-Asteretum bellidia- stri and Campanulo cespitosae-Saxifragetum aizoidis. All but the first and the last were described as new. The most important in view of nature conserva- tion are the stands of the first (Astrantio-Pinguicule- tum, localities of protected Pinguicula alpina), the sec- ond (Palustriello-Astrantietum carniolica, localities of two relatively rare mosses Trichostomum crispulum and Microlejeunea ulicina in the ravine of Velike Luti in the Kneška Grapa gorge), and the last (Campanulo- -Saxifragetum) association, namely because of the spe- cific character of the erosion area by the Luknova Grapa gorge, where the frigidophilous and hygrophi- lous subalpine-alpine species Saxifraga aizoides and the thermophilous scree species Achnatherum calama- grostis occur together. When describing the communi- ties of moist rock crevices at Liščak we also analysed our relevés from other gorges in the Julian Alps and their foothills, and described two new associations, Violo biflorae-Astrantietum carniolicae and Veronico urticifoliae-Violetum biflorae, which, however, do not have any localities in Liščak. Based on the newly de- scribed communities analysed herein and similar communities in the northern-Dinaric and pre-Alpine part of Slovenia we also described a new alliance Astrantio carniolicae-Paederotion luteae, into which we assign the studied communities. A special feature of the Liščak gorge is also tall herbs with Lunaria rediviva and Lamium orvala on colluvium (gravel) at the foot of steep slopes. Based on the relevés from this and other gorges in Slovenia we classify such stands into the new association Lamio or- valae-Lunarietum redivivae. One of its character spe- cies is also the shrub Sambucus nigra, which is occa- sionally, on similar sites in the next successional stage, a dominant species in the stands of the association Lamio orvalae-Sambucetum nigrae. Such stands are known from the neighbouring Friuli-Venezia Giulia. We documented them with a phytosociological table for Slovenia and described a new subassociation with Lunaria rediviva (lunarietosum redivivae). Associations Violo biflorae-Astrantietum carnioli- cae, Calamagrostio variae-Asteretum bellidiastri, Vero- nico urticifoliae-Violetum biflorae, Lamio orvalae-Lu- narietum redivivae and Lamio orvalae-Sambucetum nigrae are new to the vegetation of the Triglav National Park (our descriptions take into account also the rele- vés from Trenta, Loška Koritnica, Možnica, Zadlaščica, Voje, Kot, the Pokljuka ravine and Vintgar. Chasmophytic communities described in this paper belong to the Natura 2000 habitat type 8210 Cal- careous rocky slopes with chasmophytic vegetation and 7220* Petrifying springs with tufa formation (Cra- toneurion), and tall herbs with Lunaria rediviva belong in Natura 2000 habitat type 6430 Hydrophilous tall herb fringe communities of plains and of the montane to alpine levels. 4. CONCLUSIONS 5 POVZETEK Liščak je okoli 3 km dolg potok v Kneški grapi in Baški dolini v južnih Julijskih Alpah. Izvira na nadmorski vi- šini okoli 1000 m pod goro Ploha (1270 m) v Tolminsko- -Bohinjskem pogorju in se izliva v reko Knežo na nad- morski višini 331 m. Na zračni razdalji okoli 2500 m premaga višinsko razliko skoraj 700 m. Z obeh bregov se v potok izlivajo številni pritoki s še večjim padcem kot ga ima glavna grapa. Ta je zelo ozka, le z redkimi manjšimi razširitvami, kjer se kopiči prod in pobočni grušč. Geološka podlaga je zelo pisana: dolomit, apne- nec, roženec, laporovec, glinavec (Buser 1986, 1987). Ker je grapa obrnjena proti jugovzhodu, je podnebje razmeroma toplo in humidno, z letnim povprečjem padavin več kot 2000 mm (Zupančič 1998) in srednjo letno temperaturo okoli 7-8 °C (Cegnar 1998). V povodju Liščaka (3,42 km2) je gozdnatost okoli 90 %. Največje površine gozdnih sestojev uvrščamo v na- slednje asociacije: Seslerio autumnalis-Fagetum, Lamio orvalae-Fagetum, Saxifrago cuneifolii-Fagetum, Fraxino orni-Ostryetum in Veratro nigri-Fraxinetum excelsioris. DAKSKOBLER & MARTINČIČ: VEGETATION OF MOIST ROCK CREVICES AND MOIST (SLOPE) DEBRIS IN THE LIŠČAK GORGE 23FOLIA BIOLOGICA ET GEOLOGICA 64/1 – 2023 V celotnem povodju Liščaka ni več nobene nase- ljene hiše, na potoku so še ostanki mlinov, ni pa nobe- nih umetnih pregrad in drugih neposrednih človeko- vih vplivov. Zaradi številnih geoloških in geomorfolo- ških posebnosti sta potok in njegova grapa zavarovana kot naravna vrednota regionalnega oz. lokalnega po- mena (Rojšek 1986, 1991, https://www.naravovarstve- ni-atlas.si/web/DefaultNvaPublic.aspx.). Vegetacijo v povodju Liščaka smo začeli raziskova- ti leta 1986, nadaljevali v letih 1987 in 1988. Po dolgem času smo to grapo ponovno obiskali leta 2015 in s po- močjo in vodenjem Petra Razpeta predvsem v letih 2018, 2019, 2020 in 2021 v njej naredili več kot 200 fito- cenoloških in florističnih popisov. Podobo celotnega rastlinstva in rastja bomo opisali v drugem članku. V tem se posvečamo izključno rastju vlažnih skalnih razpok in visokemu steblikovju na vlažnem pobočnem grušču (slika 1). Popise iz Liščaka in njegove neposredne okolice (Velike Luti) smo želeli umestiti v sintaksonomski sy- stem. Pri nekaterih popisih je bilo to mogoče, pri drugih ne, zato smo v nekatere preglednice uvrstili tudi popise iz nekaterih drugih grap, kjer zadnja leta proučujemo rastje skalnih razpok (Dakskobler et al. 2021, Daksko- bler & Martinčič 2020, 2021) in na tej podlagi opisali nekatere nove asociacije. V združbah vlažnih skalnih razpok podgorskega in gorskega pasu v Julijskih Alpah s prigorjem večinoma prevladujejo naslednje cevnice Pinguicula alpina, Astrantia carniolica, Viola biflora, Aster bellidiastrum, Veronica urticifolia, Valeriana trip- teris, V. saxatilis, Saxifraga cuneifolia, Saxifraga aizoi- des, Asplenium viride, A. trichomanes in mahovi in je- trenjaki Orthothecium rufescens, Palustriella commuta- ta, Hymenostylium recurvirostrum in Conocephalum co- nicum. Kljub precejšnji floristični podobnosti jih ne moremo uvrstiti le v eno samo asociacijo. Zato smo pri opisih novih združb na rangu asociacije upoštevali tudi stalnost in srednje zastiranje prevladujočih vrst na popi- snih ploskvah, saj prav po tem znaku te združbe lahko prepoznamo tudi na terenu. Podobno ravnamo v neka- terih primerih tudi pri traviščnih, grmiščnih in gozdnih združbah, ko je za uvrstitev v določeno asociacijo odlo- čilna prevladujoča vrsta najvišje sestojne plasti. Po napisanem pristopu smo v grapi Liščaka prepo- znali naslednje asociacije: Astrantio carniolicae-Pingiu- culetum alpinae, Palustriello commutati-Astrantietum carniolica, Palustriello commutati-Veronicetum urtici- foliae, Veronico urticifoliae-Saxifragetum cuneifolii, Calamagrostio varie-Asteretum bellidiastri in Campa- nulo cespitosae-Saxifragetum aizoidis. Vse razen prve in zadnje smo opisali kot nove. Naravovarstveno najbolj vredni so sestoji prve (Astrantio-Pinguiculetum, nahajališča zavarovane vrste Pinguicula alpina), druge (Palustriello-Astrantie- tum carniolica, nahajališča dveh razmeroma redkih mahovnih vrst Trichostomum crispulum in Microlejeu- nea ulicina v soteski Velike Luti v Kneški grapi) in za- dnje (Campanulo-Saxifragetum), zaradi posebnosti erozijskega območja ob Luknovi grapi, kjer skupaj uspevata hladno- in vlagoljubna subalpinsko-alpinska vrsta Saxifraga aizoides in toploljubna vrsta melišč Achnatherum calamagrostis. Ob opisovanju združb vlažnih skalnih razpok ob Liščaku smo v pretres in obravnavo vključili tudi naše popise iz drugih grap v Julijskih Alpah in njihovem prigorju in opisali še dve novi asociaciji Violo biflorae-Astrantietum carniolicae in Veronico urticifoliae-Violetum biflorae, ki pa v Li- ščaku nimata svojih nahajališč. Na podlagi v teh članku novo opisanih združb in podobnih združb v severnodinarskem in predalpskem delu Slovenije smo opisali tudi novo zvezo Astrantio carniolicae-Paederotion luteae, kamor obravnavane asociacije tudi uvrščamo. Njene diagnostične vrste de- limo na fitogeografsko-ekološke in ekološke. Prve so Astrantia carniolica, Paederota lutea in Primula carni- olica, v manjši meri tudi diagnostične vrste zvez Physo- plexido comosae-Saxifragion petraeae in Aremonio-Fa- gion ter druge jugovzhodnoalpsko-severnodinarske vrste, ki se posamično pojavljajo v njihovih sestojih. Te jih zadostno razlikujejo od podobnih združb iz zveze Cystopteridion. Ekološke diagnostične vrste so Aster bellidiastrum, Hydrogonium croceum (Barbula crocea), Carex brachystachys, Cystopteris fragilis, Asplenium vi- ride, Eucladium verticillatum, Fissidens dubius, Hyme- nostylium recurvirostrum, Apopellia endiviifolia (Pellia endiviifolia), Palustriella commutata, Marchantia qua- drata (Preissia quadrata), Pingiucula alpina, Orthothe- cium rufescens, Saxifraga aizoides, Tofieldia calyculata, Valeriana saxatilis, V. tripteris in Viola biflora. Naštete so sicer značilnice različnih zvez in razredov, a prouče- ne združbe razlikujejo od združb bolj suhega skalovja iz zveze Physoplexido comosae-Saxifragion petraeae. Posebnost grape Liščaka je tudi visoko steblikovje s srebrenko (Lunaria rediviva) in velecvetno mrtvo ko- privo (Lamium orvala) na koluviju (grušču) ob dnu strmih pobočij. Takšne sestoje na podlagi popisov tudi iz drugih grap uvrščamo v novo asociacijo Lamio orva- lae-Lunarietum redivivae. Ena izmed njenih značilnic je tudi črni bezeg (Sambucus nigra), ki je ponekod na podobnih rastiščih v naslednji sukcesijski stopnji pre- vladujoča vrsta zgornje plasti v sestojih asociacije Lamio orvalae-Sambucetum nigrae. Takšne sestoje po- znajo v sosednji deželi Furlaniji Julijski krajini, s fito- cenološko tabelo smo jih dokumentirali tudi v Sloveni- ji in opisali novo subasociacijo s srebrenko (lunarieto- sum redivivae). DAKSKOBLER & MARTINČIČ: VEGETATION OF MOIST ROCK CREVICES AND MOIST (SLOPE) DEBRIS IN THE LIŠČAK GORGE 24 FOLIA BIOLOGICA ET GEOLOGICA 64/1 – 2023 Asociacije Violo biflorae-Astrantietum carniolicae, Calamagrostio variae-Asteretum bellidiastri, Veronico urticifoliae-Violetum biflorae, Lamio orvalae-Lunarie- tum redivivae in Lamio orvalae-Sambucetum nigrae so novost v rastju Triglavskega narodnega parka, saj smo pri njihovem opisu upoštevali tudi popise iz Trente, Loške Koritnice, Možnice, Zadlaščice, Voj, Pokljuške soteske, Kota in Vintgarja. V tem članku opisane združbe skalnih razpok so- dijo v Natura 2000 habitatna tipa 8210 Karbonatna skalnata pobočja z vegetacijo skalnih razpok in 7220* Lehnjakotvorni izviri (Cratoneurion), visoko stebli- kovje s srebrenko pa v Natura 2000 habitatni tip 6430 Nižinske in montanske do alpinske hidrofilne združbe z visokim steblikovjem. ACKNOWLEDGEMENTS Peter Razpet, Erik Kragelj, Karin Rutar, Branko Zupan, Iztok Sajko, Prof. Dr. Boštjan Surina and Mag. Boško Čušin helped the younger author in the field surveying of the described communities. Iztok Sajko prepared Figure 1 for print. Dr. Branko Vreš is the caretaker of the FloVegSi database and, together with Brane An- derle, Mag. Andrej Seliškar, Prof. Dr. Boštjan Surina, Branko Dolinar, Janez Mihael Kocjan and Branko Zupan, the co-author of the distribution maps in Fig- ures 4, 7, 9 and 12. Mag. Daniel Rojšek provided the data on the nature conservation status of Liščak. 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DAKSKOBLER & MARTINČIČ: VEGETATION OF MOIST ROCK CREVICES AND MOIST (SLOPE) DEBRIS IN THE LIŠČAK GORGE 26 FOLIA BIOLOGICA ET GEOLOGICA 64/1 – 2023 Figure 14: Kneška Grapa gorge at Velike Luti Slika 14: Kneška grapa pri koritih Velike Luti Figure 15: The Liščak gorge (Kneška Grapa gorge, Bača Valley). Slika 15: Liščak (Kneška grapa, Baška dolina). DAKSKOBLER & MARTINČIČ: VEGETATION OF MOIST ROCK CREVICES AND MOIST (SLOPE) DEBRIS IN THE LIŠČAK GORGE 27FOLIA BIOLOGICA ET GEOLOGICA 64/1 – 2023 Figure 16: Researched sites in the Liščak gorge: moist rocks, debris and talus. Slika 16: Proučena rastišča v grapi Liščaka, vlažno skalovje, pobočni grušč in vršaji. Figure 17 a: Stand of the association Astrantio carniolicae-Pinguiculetum alpinae in the Liščak gorge. Slika 17 a: Sestoj asociacije Astrantio carniolicae-Pinguiculetum alpinae v grapi Liščak. DAKSKOBLER & MARTINČIČ: VEGETATION OF MOIST ROCK CREVICES AND MOIST (SLOPE) DEBRIS IN THE LIŠČAK GORGE 28 FOLIA BIOLOGICA ET GEOLOGICA 64/1 – 2023 Figure 17 b: Stand of the association Astrantio carniolicae-Pinguiculetum alpinae in the Kneška Grapa gorge (Velike Luti). Slika 17 b: Sestoj asociacije Astrantio carniolicae-Pinguiculetum alpinae v Kneški grapi (Velike Luti). Figure 18: Stand of the association Violo biflorae-Astrantietum carniolicae, Govci above the Trebuša Valley. Slika 18: Sestoj asociacije Violo biflorae-Astrantietum carniolicae, Govci nad dolino Trebuše. DAKSKOBLER & MARTINČIČ: VEGETATION OF MOIST ROCK CREVICES AND MOIST (SLOPE) DEBRIS IN THE LIŠČAK GORGE 29FOLIA BIOLOGICA ET GEOLOGICA 64/1 – 2023 Figure 19: Stand of the variant Palustriello commutati-Astrantietum carniolicae var. Carex mucronata, Velike Luti in the Kneška Grapa gorge. Slika 19: Sestoj variante Palustriello commutati-Astrantietum carniolicae var. Carex mucronata, Velike Luti v Kneški grapi. Figure 20: Stand of the association Palustriello commutati-Astrantietum carniolicae in the Liščak gorge. Slika 20: Sestoj asociacije Palustriello commutati-Astrantietum carniolicae v grapi Liščaka. DAKSKOBLER & MARTINČIČ: VEGETATION OF MOIST ROCK CREVICES AND MOIST (SLOPE) DEBRIS IN THE LIŠČAK GORGE 30 FOLIA BIOLOGICA ET GEOLOGICA 64/1 – 2023 Figure 21: Stand of the association Palustriello-Veronicetum urticifoliae in the Liščak gorge. Slika 21: Sestoj asociacije Palustriello-Veronicetum urticifoliae v grapi Liščaka. Figure 22: Stand of the association Cratoneuretum commutati s. lat. in the Liščak gorge. Slika 22: Sestoj asociacije Cratoneuretum commutati s. lat. v grapi Liščaka. DAKSKOBLER & MARTINČIČ: VEGETATION OF MOIST ROCK CREVICES AND MOIST (SLOPE) DEBRIS IN THE LIŠČAK GORGE 31FOLIA BIOLOGICA ET GEOLOGICA 64/1 – 2023 Figure 23: Stand of the subassociation Campanulo cespitosae-Saxifragetum aizoidis achnatheretosum calamagrostis, Luknova Grapa gorge (Za Jamo). Slika 23: Sestoj subasociacije Campanulo cespitosae-Saxifragetum aizoidis achnatheretosum calamagrostis, Luknova grapa (Za Jamo). Figure 24: Stand of the association Veronico urticifoliae-Saxifragetum cuneifolii in the Liščak gorge. Slika 24: Sestoj asociacije Veronico urticifoliae-Saxifragetum cuneifolii v grapi Liščaka. DAKSKOBLER & MARTINČIČ: VEGETATION OF MOIST ROCK CREVICES AND MOIST (SLOPE) DEBRIS IN THE LIŠČAK GORGE 32 FOLIA BIOLOGICA ET GEOLOGICA 64/1 – 2023 Figure 25: Stand of the association Calamagrostio variae-Asteretum bellidiastri in the Kneška Grapa gorge. Slika 25: Sestoj asociacije Calamagrostio variae-Asteretum bellidiastri v Kneški grapi. Figure 26: Stand of the association Calamagrostio variae-Asteretum bellidiastri in the Vintgar gorge. Slika 26: Sestoj asociacije Calamagrostio variae-Asteretum bellidiastri v Vintgarju. DAKSKOBLER & MARTINČIČ: VEGETATION OF MOIST ROCK CREVICES AND MOIST (SLOPE) DEBRIS IN THE LIŠČAK GORGE 33FOLIA BIOLOGICA ET GEOLOGICA 64/1 – 2023 Figure 27: Strand of the association Veronico urticifoliae-Violetum biflorae in the Vintgar Gorge. Slika 27: Sestoj asociacije Veronico urticifoliae-Violetum biflorae v Vintgarju. Figure 28: Stand of the syntaxon Seslerio caeruleae-Sedetum albi nom. prov. in the spring area of the Liščak gorge. Slika 28: Sestoj sintaksona Seslerio caeruleae-Sedetum albi v povirju Liščaka. DAKSKOBLER & MARTINČIČ: VEGETATION OF MOIST ROCK CREVICES AND MOIST (SLOPE) DEBRIS IN THE LIŠČAK GORGE 34 FOLIA BIOLOGICA ET GEOLOGICA 64/1 – 2023 Figure 29 a: Stand of the association Lamio orvalae-Lunarietum redivivae in the Vintgar gorge. Slika 29 a: Sestoj asociacije Lamio orvalae-Lunarietum redivivae v Vintgarju. Figure 29 b: Stand of the association Lamio orvalae-Lunarietum redivivae in the Liščak gorge. Slika 29 b: Sestoj asociacije Lamio orvalae-Lunarietum redivivae v grapi Liščaka. DAKSKOBLER & MARTINČIČ: VEGETATION OF MOIST ROCK CREVICES AND MOIST (SLOPE) DEBRIS IN THE LIŠČAK GORGE 35FOLIA BIOLOGICA ET GEOLOGICA 64/1 – 2023 Figure 30: Stand of the association Lamio orvalae-Sambucetum nigrae in the Avšček gorge. Slika 30: Sestoj asociacije Lamio orvalae-Sambucetum nigrae v grapi Avščka. Foto (Photo): I. Dakskobler DAKSKOBLER & MARTINČIČ: VEGETATION OF MOIST ROCK CREVICES AND MOIST (SLOPE) DEBRIS IN THE LIŠČAK GORGE 36 FOLIA BIOLOGICA ET GEOLOGICA 64/1 – 2023 Ta bl e 1 (P re gl ed ni ca 1 ): A st ra nt io ca rn io lic ae -P in gu ic ul et um a lp in ae v ar . C al am ag ro st is v ar ia Su cc es si ve n um be r of r el ev é (Z ap or ed na š te vi lk a po pi sa ) 1 2 3 4 5 6 7 8 9 19 11 12 13 14 D at ab as e nu m be r of r el ev é (D el ov na š te vi lk a po pi sa ) 28 22 76 28 22 59 28 22 61 28 22 66 28 22 73 28 52 77 28 52 81 28 52 78 28 52 79 28 52 82 28 71 63 28 72 15 28 53 58 28 72 08 El ev at io n in m (N ad m or sk a vi ši na v m ) 59 0 56 0 56 5 62 5 62 0 32 5 33 5 33 0 33 0 33 5 45 5 30 0 30 0 32 0 A sp ec t ( Le ga ) SE W W W S SW SW W SW SW SW SW S W N E Sl op e in d eg re es (N ag ib v s to pi nj ah ) 90 90 90 90 80 90 90 90 90 90 90 90 90 80 Pa re nt m at er ia l ( M at ič na p od la ga ) LM C h LM C h LM C h D C h D C h D C h D C h D C h D C h D C h LM C h LC lC h D C h D C h So il (T la ) Li Li Li Li Li Li Li Li Li Li Li Li Li Li St on in es s i n % (K am ni to st v % ) 10 0 10 0 10 0 10 0 10 0 10 0 10 0 10 0 10 0 10 0 10 0 10 0 10 0 10 0 C ov er o f h er b la ye r in % (Z as ti ra nj e ze liš čn e pl as ti v % ): 30 30 30 40 30 30 30 40 35 40 40 50 25 40 C ov er o f m os s l ay er in % (Z as ti ra nj e m ah ov ne p la st i v % ): 40 50 70 60 60 40 40 30 30 60 50 30 20 30 N um be r of s pe ci es (Š te vi lo v rs t) 9 21 18 19 24 15 12 10 15 18 15 15 10 22 R el ev é ar ea (V el ik os t p op is ne p lo sk ve ) m 2 10 5 10 10 10 10 20 10 10 5 10 10 10 10 D at e of ta ki ng r el ev é (D at um p op is a) 8/21/2020 8/21/2020 8/21/2020 8/21/2020 8/21/2020 4/9/2021 4/9/2021 4/9/2021 4/9/2021 4/9/2021 8/11/2021 6/18/2021 4/14/2021 6/18/2021 Lo ca lit y (N ah aj al iš če ) Liščak-Zališčar jeva grapa Liščak-Luknova grapa Liščak Liščak Liščak Velike Luti Velike Luti Velike Luti Velike Luti Velike Luti Liščak-Sopot Knežica-Mlinar Velike Luti Velike Luti Q ua dr an t ( K va dr an t) 97 49 /3 97 49 /3 97 49 /3 97 49 /3 97 49 /3 98 49 /1 98 49 /1 98 49 /1 98 49 /1 98 49 /1 98 49 /1 98 49 /1 98 49 /1 98 49 /1 C oo rd in at e G K Y (D -4 8) m 41 12 38 41 15 53 41 15 65 41 16 26 41 15 89 41 10 53 41 08 76 41 10 29 41 10 11 41 08 46 41 13 59 41 10 65 41 10 58 41 07 34 C oo rd in at e G K X (D -4 8) m 5118090 5118074 5118055 5118286 5118274 5116536 5116758 5116573 5116584 5116789 5117686 5116408 5116428 5116795 D ia gn os ti c sp ec ie s of th e as so ci at io n (D ia gn os ti čn e vr st e as oc ia ci je ) Pr . Fr . A P Pi ng ui cu la a lp in a E1 1 1 1 1 + 2 3 3 3 3 3 2 2 2 14 10 0 A P H ym en os ty liu m re cu rv ir os tr um E0 + 2 3 3 2 2 3 1 2 3 3 1 1 1 14 10 0 A P Pa lu st ri el la c om m ut at a E0 3 3 2 2 4 3 2 2 2 2 2 3 1 2 14 10 0 A P A st ra nt ia c ar ni ol ic a E1 3 . . + + 1 1 . r . 3 3 + 1 10 71 T R Pe ta si te s p ar ad ox us E1 . . . . . 1 r + r . . + . r 6 43 A P A st ra nt io ca rn io lic ae -P ae de ro ti on lu te ae A st er b el lid ia st ru m E1 . 1 1 + + + + + r . 1 1 1 1 12 86 A po pe lli a en di vi ifo lia (P el lia e nd iv iif ol ia ) E0 1 1 1 1 . . . 1 . + + . . + 8 57 C ar ex b ra ch ys ta ch ys E1 . 2 1 2 . 1 . . + . + . . + 7 50 O rt ho th ec iu m r uf es ce ns E0 . 1 + + . . . . + . . . 1 2 6 43 Eu cl ad iu m v er tic ill at um E0 + + . . + . 1 . . + 1 . . . 6 43 To fie ld ia c al yc ul at a E1 . . . r + . + . . + . 1 . . 5 36 V al er ia na tr ip te ri s E1 . . 1 . 2 . . . . . + . . + 4 29 M ar ch an tia q ua dr at a (P re is si a qu ad ra ta ) E0 . + . . + . . . . + . . . . 3 21 Fi ss id en s d ub iu s E0 . . + . . . . + . . . . . + 3 21 Ju ng er m an ni a at ro vi re ns E0 . . . + . . + . . . . . . . 2 14 DAKSKOBLER & MARTINČIČ: VEGETATION OF MOIST ROCK CREVICES AND MOIST (SLOPE) DEBRIS IN THE LIŠČAK GORGE 37FOLIA BIOLOGICA ET GEOLOGICA 64/1 – 2023 Su cc es si ve n um be r of r el ev é (Z ap or ed na š te vi lk a po pi sa ) 1 2 3 4 5 6 7 8 9 19 11 12 13 14 Pr . Fr . H yd ro go ni um c ro ce um (B ar bu la c ro ce a) E0 . . . . . . . . . 1 . . . 1 2 14 Sa xi fr ag a ai zo id es E1 . + . . . . . . . . . . . . 1 7 V al er ia na sa xa til is E1 . . . . . . . . . . . . . 2 1 7 D if fe re nt ia l s pe ci es o f t he v ar ia nt (R az li ko va ln ic a va ri an te ) EP C al am ag ro st is v ar ia E1 + 1 1 2 2 + + . r 1 . + + + 12 86 Ps Sp Ph ys op le xi do co m os ae -S ax if ra gi on p et ra ea e C am pa nu la c es pi to sa E1 + . . 2 + . . . . . + . . . 4 29 H ie ra ci um p or ri fo liu m E1 . . . . r . . . . . . . . . 1 7 PC Po te nt ill et al ia ca ul es ce nt is Sa xi fr ag a cr us ta ta E1 . r . . . . . . . . . . . . 1 7 A sp le ni et ea tr ic ho m an is A sp le ni um r ut a- m ur ar ia E1 . + + . . . . . . + . . . . 3 21 A sp le ni um tr ic ho m an es E1 . + . . . . . . . r . . . . 2 14 T R T hl as pi et ea ro tu nd ifo lii H ie ra ci um b ifi du m E1 . + 1 + + r . . r + . . . . 7 50 A de no st yl es g la br a E1 . + 1 + . . . . . . 1 . . . 4 29 M C M on ti o- C ar da m in et ea C on oc ep ha lu m c on ic um E0 . . . . 1 . . . . . . . . + 2 14 A ne ur a pi ng ui s E0 . . . . . . . . + . . . . . 1 7 C D C ar ic et al ia d av al lia na e C am py lo ph yl lo ps is c al ca re a (C am py liu m c al ca re a) E0 . . + . . . . . . . . . . . 1 7 C am py liu m st el la tu m E0 . . . . . . . + . . . . . . 1 7 C ar ex le pi do ca rp a E1 . . . . . . . . . . . + . . 1 7 ES El yn o- Se sl er ie te a Se sl er ia c ae ru le a E1 . . . . . + 1 1 + 2 . . + 1 7 50 M A M ol in io -A rr he na th er et ea A ng el ic a sy lv es tr is E1 . . . + . r . . . . . . . . 2 14 BA Be tu lo -A ln et ea Sa lix a pp en di cu la ta E2 a . . . . . r . + . + . . . . 3 21 EA Ep ilo bi et ea a ng us ti fo lii Eu pa to ri um c an na bi nu m E1 . . . . + . . . . . r + . . 3 21 Tu ss ila go fa rf ar a E1 . . . . . . . . . r . . . . 1 7 EP Er ic o- Pi ne te a, F es tu co -B ro m et ea M ol in ia a ru nd in ac ea E1 + + . . . 1 + 2 1 . + + . 2 9 64 Er ic a ca rn ea E1 . . . . + . . . r . . + r . 4 29 Bu ph th al m um sa lic ifo liu m E1 . . . . 1 . . . . r . . . . 2 14 C ar ex o rn ith op od a E1 . + . . . . . . . + . . . . 2 14 C ir si um e ri si th al es E1 . . . . . . . . . + . . . . 1 7 V P Va cc in io -P ic ee te a V er on ic a ur tic ifo lia E1 . . + + . . . . . . . . . . 2 14 A po se ri s f oe tid a E1 . . . . . . . . . . . + . . 1 7 G en tia na a sc le pi ad ea E1 . . . + . . . . . . . . . . 1 7 A F A re m on io -F ag io n, E ry th ro ni o- C ar pi ni on C yc la m en p ur pu ra sc en s E1 . . . . + . . . . . . . . . 1 7 Pr im ul a vu lg ar is E1 . . . . + . . . . . . . . . 1 7 DAKSKOBLER & MARTINČIČ: VEGETATION OF MOIST ROCK CREVICES AND MOIST (SLOPE) DEBRIS IN THE LIŠČAK GORGE 38 FOLIA BIOLOGICA ET GEOLOGICA 64/1 – 2023 TA Ti lio -A ce ri on A ru nc us d io ic us E1 . . . . . . . . . . + . . . 1 7 FS Fa ge ta lia sy lv at ic ae G al iu m la ev ig at um E1 . + + . + . . . . . + . + r 6 43 Sa lv ia g lu tin os a E1 . + + + + . . . . . . . . . 4 29 M yc el is m ur al is E1 . . . . + . . . . . . . . . 1 7 Fa gu s s yl va tic a E1 . . . . . r . . . . . . . . 1 7 Q P Q ue rc et al ia p ub es ce nt i- pe tr ae ae C ar ex fl ac ca E1 . . . . . + . 1 1 . . 1 . . 4 29 O st ry a ca rp in ifo lia E2 a . + . . . . . . . . . . . . 1 7 Q F Q ue rc o- Fa ge te a H ed er a he lix E1 . . . + + . . . . . . . . . 2 14 C ar ex d ig ita ta E1 . . + . . . . . . . . . . . 1 7 C le m at is v ita lb a E1 . . . . . . . . . . . + . . 1 7 M L M os se s (M ah ov i) Ex er to th ec a c ri sp a (N ec ke ra c ri sp a) E0 . . . . 1 . . . . . . . . . 1 7 Se lig er a tr ifa ri a E0 . . . . . . . + . . . . . . 1 7 D ic ho do nt iu m p el lu ci du m E0 . . . . . . . . . . . . . 1 1 7 D id ym od on fa lla x E0 . . . . . . . . . . . . . + 1 7 To rt el la to rt uo sa E0 . . . . . . . . . . . . . + 1 7 Le ge nd - Le ge nd a L Li m es to ne - ap ne ne c C h C he rt - ro že ne c D D ol om ite - do lo m it C l C la ys to ne - gl in av ec M M ar ls to ne - la po ro ve c Li L ith os ol - ka m ni šč e Pr . P re se nc e (n um be r of r el ev és in w hi ch th e sp ec ie s i s p re se nt ed ) - š te vi lo p op is ov , v k at er ih s e po ja vl ja v rs ta Fr . F re qu en cy in % - fr ek ve nc a v % Su cc es si ve n um be r of r el ev é (Z ap or ed na š te vi lk a po pi sa ) 1 2 3 4 5 6 7 8 9 19 11 12 13 14 Pr . Fr . DAKSKOBLER & MARTINČIČ: VEGETATION OF MOIST ROCK CREVICES AND MOIST (SLOPE) DEBRIS IN THE LIŠČAK GORGE 39FOLIA BIOLOGICA ET GEOLOGICA 64/1 – 2023 Ta bl e 2 (P re gl ed ni ca 2 ): V io lo b if lo ra e- A st ra nt ie tu m ca rn io lic ae Su cc es si ve n um be r of r el ev é (Z ap or ed na š te vi lk a po pi sa ) 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 D at ab as e nu m be r of r el ev é (D el ov na š te vi lk a po pi sa ) 257807 269661 273173 282738 282739 258917 269646 273153 274072 274078 278144 286378 269643 269647 277320 288035 268929 277777 El ev at io n in m (N ad m or sk a vi ši na v m ) 93 0 88 0 11 75 56 5 56 5 98 5 11 30 11 30 12 85 13 00 13 50 12 00 92 5 11 30 11 82 54 0 55 0 69 7 A sp ec t ( Le ga ) N N W N W N W N W N N N W N N N W N E N N N W N N EE N Sl op e in d eg re es (N ag ib v s to pi nj ah ) 80 80 80 80 80 80 85 80 80 95 90 90 70 50 10 0 80 85 95 Pa re nt m at er ia l ( M at ič na p od la ga ) L L D D D D L D L D D D L L L D D D D So il (T la ) Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li St on in es s i n % (K am ni to st v % ) 10 0 10 0 10 0 10 0 10 0 10 0 10 0 10 0 10 0 10 0 10 0 10 0 10 0 10 0 10 0 10 0 10 0 10 0 C ov er o f h er b la ye r in % (Z as ti ra nj e ze liš čn e pl as ti v % ): 30 50 30 30 40 40 40 25 40 20 40 30 60 80 30 30 25 30 C ov er o f m os s l ay er in % (Z as ti ra nj e m ah ov ne p la st i v % ): 30 30 30 20 20 80 60 10 20 10 10 10 70 40 10 20 40 10 N um be r of s pe ci es (Š te vi lo v rs t) 16 14 24 16 14 11 24 27 28 32 24 19 27 23 14 22 22 24 R el ev é ar ea (V el ik os t p op is ne p lo sk ve ) m 2 10 10 10 10 10 4 10 10 15 20 15 10 10 10 10 10 10 15 D at e of ta ki ng r el ev é (D at um p op is a) 8/11/2015 6/21/2017 9/19/2018 6/22/2020 6/22/2020 7/3/2015 6/21/2017 9/21/2018 7/27/2018 7/27/2018 6/14/2019 7/26/2021 6/21/2017 6/21/2017 7/10/2019 7/22/2021 10/9/2017 6/24/2019 Lo ca lit y (N ah aj al iš če ) Zjabci Trenta-Be- ladovec Kot-Kotarica Vintgar Vintgar Trenta-Zadnji- ca Trenta-Lepoč Zadnjica- Kanjavec Loška Koritni- ca-Jalovec Loška Koritni- ca-Jalovec Trenta-Zapo- tok Komna-Pekel Trenta-Be- ladovec Trenta-Lepoč Strmec-Man- gart Gorenja Trebuša Govci Otuška-Pri mlinu Voje izvir Krope Q ua dr an t ( K va dr an t) 9648/3 9648/2 9549/3 9650/2 9650/2 9648/2 9648/2 9648/2 9548/3 9548/3 9648/1 9748/2 9648/2 9648/2 9547/4 9949/3 9949/2 9649/3 C oo rd in at e G K Y (D -4 8) m 400276 404242 414666 430243 430248 407689 404708 408369 397748 397754 399390 407214 404237 404699 392642 410883 420057 414612 C oo rd in at e G K X (D -4 8) m 5131801 5136645 5140324 5139553 5139570 5137099 5135878 5137042 5143135 5143115 5138583 5127432 5136661 5135876 5143224 5097256 5103034 5130398 D ia gn os ti c sp ec ie s of th e as so ci at io n (D ia gn os ti čn e vr st e as oc ia ci je ) Pr . Fr . A P A st ra nt ia c ar ni ol ic a E1 2 3 2 2 3 3 1 + 2 + 3 1 3 3 2 2 + 1 18 10 0 A P V io la b if lo ra E1 + 1 1 + 2 + r + 1 1 1 + 1 + + 2 2 1 18 10 0 A P C ar ex b ra ch ys ta ch ys E1 1 1 + + 1 + . + + + r + . . 1 . + r 14 78 T R A de no st yl es g la br a E1 + + 1 + 1 . + r + r . . 1 3 . . . + 12 67 A P Pa ed er ot a lu te a E1 1 + 2 . . + + 1 1 1 1 . . + . 1 + . 12 67 A P H el io sp er m a pu si llu m E1 . 1 1 . . . + . 1 + . + + . . . . . 7 39 D if fe re nt ia l s pe ci es o f l ow er u ni ts (R az li ko va ln ic e ni žj ih e no t) A P Pa lu st ri el la c om m ut at a E0 1 . 2 2 2 4 + . . . . . . . . + . . 7 38 A P H ym en os ty liu m re cu rv ir os tr um E0 + . 1 . 1 4 1 . . . . + . . . . . + 7 38 EP R ho do th am nu s c ha m ae ci st us E1 . . . . . . . + 1 1 + + . . . 1 . . 6 33 DAKSKOBLER & MARTINČIČ: VEGETATION OF MOIST ROCK CREVICES AND MOIST (SLOPE) DEBRIS IN THE LIŠČAK GORGE 40 FOLIA BIOLOGICA ET GEOLOGICA 64/1 – 2023 Su cc es si ve n um be r of r el ev é (Z ap or ed na š te vi lk a po pi sa ) 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 Pr . Fr . A P M ar ch an tia q ua dr at a (P re is si a qu ad ra ta ) E0 . . . . . . . + + + . + . . . . . . 4 22 A P A st ra nt io ca rn io lic ae -P ae de ro ti on lu te ae O rt ho th ec iu m r uf es ce ns E0 1 2 2 2 1 . 2 + 1 + + + 1 1 1 2 . + 16 89 A sp le ni um v ir id e E1 r . + 1 . . + + + + 1 . + + . 1 1 1 13 72 A st er b el lid ia st ru m E1 . . + . . . + 1 + + . . + + 1 1 + + 11 61 Fi ss id en s d ub iu s E0 . . . . . . 1 . + + + + 1 . . + + + 9 50 V al er ia na sa xa til is E1 + . . . . . . + 1 1 + 1 . . 1 1 . . 8 44 C ys to pt er is re gi a E1 . . 1 . . . r . 1 + . + . . . . . + 6 33 C ys to pt er is fr ag ili s E1 . . . 1 + . 2 . . . + . . + . . . + 6 33 Pi ng ui cu la a lp in a E1 r . . . . . + . . . . + . . . + . . 4 22 V al er ia na tr ip te ri s E1 . . . + . . . . . . . . . 1 . . 1 . 3 17 To fie ld ia c al yc ul at a E1 . . . . . + . r . . . . . . . 1 . . 3 17 H yd ro go ni um c ro ce um (B ar bu la c ro ce a) E0 . . . . . . . . + + . . . . . + . . 3 17 A po pe lli a en di vi ifo lia (P el lia e nd iv iif ol ia ) E0 . . . . . + . . . . . . . . . . . + 2 11 C yr to m ni um h ym en op hy llo id es E0 . . . . . . . . . . . . + . . + . . 2 11 Ps Sp Ph ys op le xi do co m os ae -S ax if ra gi on p et ra ea e C am pa nu la z oy si i E1 . . . . . . . + + . . . . . r . . . 3 17 C am pa nu la c ar ni ca E1 . . . . . + . . . . . . + . . . . . 2 11 H ie ra ci um p or ri fo liu m E1 . . . . . . . . r . . . . . . . . . 1 6 C am pa nu la c es pi to sa E1 . . . . . . . . . + . . . . . . . . 1 6 Ph yt eu m a sc he uc hz er i s ub sp . c ol um na e E1 . . . . . . . . . . . . . . . + . . 1 6 PC Po te nt ill et al ia ca ul es ce nt is C am pa nu la c oc hl ea ri ifo lia E1 . . 1 . . . . + + . . 1 . + 1 . . . 6 33 A ra bi s s te llu la ta E1 . . . . . . . . . + + + . . + . . . 4 22 Pr im ul a au ri cu la E1 . + . . . . r . r . r . . . . . . . 4 22 Po te nt ill a ca ul es ce ns E1 + . . . . . . . . . r . . . . . . . 2 11 H ie ra ci um h um ile E1 . . . . . . . + . . . . . . . . . . 1 6 Po te nt ill a cl us ia na E1 . . . . . . . + . . . . . . . . . . 1 6 A T A sp le ni et ea tr ic ho m an is A sp le ni um r ut a- m ur ar ia E1 . . . . . . . . + + + . . . . . + + 5 28 A sp le ni um tr ic ho m an es E1 . + . . . . . . . . . . . . . . 1 + 3 17 M oe hr in gi a m us co sa E1 . . . + . . . . . . . . . . . . . . 1 6 K er ne ra sa xa til is E1 . . . . . . . . . . . . . . . r . . 1 6 A C A ra bi de ta lia ca er ul ea e So ld an el la m in im a E1 . . + . . . . . . . . . + + . . . . 3 17 T R T hl as pi et ea ro tu nd ifo lii H ie ra ci um b ifi du m E1 . . . . . . . + . . . . + . + . . r 4 22 G ym no ca rp iu m ro be rt ia nu m E1 . . . . . . . + . . + 1 . . . . . + 4 22 G yp so ph ila re pe ns E1 . . . . . . . + r + . . . . . . . . 3 17 Fe st uc a ni tid a E1 . . + . . . . . . . . . . 1 . . . . 2 11 C ys to pt er is m on ta na E1 . . . . . . r . . . . . . 1 . . . . 2 11 Sa xi fr ag a ca es ia E1 . . . . . . . . . r . . . . + . . . 2 11 A ra bi s a lp in a E1 . . . . . . r . . . . . . . . . . . 1 6 A th am an ta c re te ns is E1 . . . . . . . r . . . . . . . . . . 1 6 Bi sc ut el la la ev ig at a E1 . . . . . . . . r . . . . . . . . . 1 6 DAKSKOBLER & MARTINČIČ: VEGETATION OF MOIST ROCK CREVICES AND MOIST (SLOPE) DEBRIS IN THE LIŠČAK GORGE 41FOLIA BIOLOGICA ET GEOLOGICA 64/1 – 2023 C er as tiu m c ar in th ia cu m E1 . . . . . . . . + . . . . . . . . . 1 6 Tr is et um a rg en te um E1 . . . . . . . . . + . . . . . . . . 1 6 Pe ta si te s p ar ad ox us E1 . . . . . . . . . . . . . . + . . . 1 6 M C M on ti o- C ar da m in et ea . . . . . . . . . . . . . . . . . . C on oc ep ha lu m c on ic um E0 + 1 . . + . 1 . . . + . . . . 1 3 1 8 44 C ra to ne ur on fi lic in um E0 . . + . . 1 . . . . . . . . . . . . 2 11 Pt yc ho st om um p se ud ot ri qu et ru m (B ry um p se ud ot ri qu et ru m ) E0 . . 1 . . . . . . . . . . . . . . . 1 6 O xy rr hy nc hi um sc hl ei ch er i E0 . . . . . . . . . . . + . . . . . . 1 6 C ol ol ej eu ne a ca lc ar ea E0 . . . . . . . . . . . + . . . . . . 1 6 C D C ar ic et al ia d av al lia na e C am py liu m st el la tu m E0 . . . . . . 1 . . . . . . . . . . . 1 6 ES El yn o- Se sl er ie te a Se sl er ia c ae ru le a E1 1 1 + . . . . 1 r . + . . 1 . . . . 7 39 C ar ex fi rm a E1 . . + . . . . + r r r . . . . . . . 5 28 La se rp iti um p eu ce da no id es E1 . + . . . . . r . . . . . + . 1 . . 4 22 C ar ex fe rr ug in ea E1 . . + . . . + . r . . . . . . . . . 3 17 A ch ill ea a tr at a E1 . . . . . . . . r . . . . . . . . . 1 6 Ph yt eu m a or bi cu la re E1 . . + . . . . . . . . . . . . . . . 1 6 R an un cu lu s h yb ri du s E1 . . . . . . . r . . . . . . . . . . 1 6 G lo bu la ri a co rd ifo lia E1 . . . . . . . . . + . . . . . . . . 1 6 Se la gi ne lla se la gi no id es E1 . . . . . . . . . . . . . + . . . . 1 6 BA Be tu lo -A ln et ea Sa lix a pp en di cu la ta E2 a + . . . . . . r . . . . . . . . . . 2 11 M uA M ul ge di o- A co ni te te a Sa xi fr ag a ro tu nd ifo lia E1 . . . + . . + . . . . . . . . . . + 3 17 A co ni tu m ly co ct on um s ub sp . r an un cu lif ol iu m E1 . . . . . . . . . . . . + . . . . . 1 6 C ha er op hy llu m h ir su tu m E1 . . . . . . . . . . . . . 1 . . . . 1 6 A co ni tu m d eg en ii su bs p. p an ic ul at um E1 . . . . . . . . . . . . . . . . . r 1 6 EP Er ic o- Pi ne te a C al am ag ro st is v ar ia E1 1 + . + . + . + + + r . 1 + . 1 r . 12 67 R ho do de nd ro n hi rs ut um E1 . . + . . . . . + . + + . . . . . . 4 22 Er ic a ca rn ea E1 . . . . . . . . . . + . . . . . . . 1 6 A sp er ul a ar is ta ta E1 . . . . . . . . . . . . . . + . . . 1 6 V P Va cc in io -P ic ee te a V er on ic a ur tic ifo lia E1 . + . 1 1 . . . . . . . 1 + . . + + 7 39 Sa xi fr ag a cu ne ifo lia E1 . . . . . . . . . . + . + . . . . + 3 17 O xa lis a ce to se lla E1 . . . . . . . . . . . . . + . . + . 2 11 H om og yn e sy lv es tr is E1 . . . . . . . . . . . . . . . . . r 1 6 A F A re m on io -F ag io n C ar da m in e tr ifo lia E1 . . . + . . . . . . . . . . . . . . 1 6 A ne m on e tr ifo lia E1 . . . . . . . . . . r . . . . . . . 1 6 C yc la m en p ur pu ra sc en s E1 . . . . . . . . . . . . . . . . + . 1 6 TA Ti lio -A ce ri on G er an iu m ro be rt ia nu m E1 . . . + . . . . . . . . . + . . + . 3 17 Ph yl lit is sc ol op en dr iu m E1 . . . + + . . . . . . . . . . . . . 2 11 Su cc es si ve n um be r of r el ev é (Z ap or ed na š te vi lk a po pi sa ) 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 Pr . Fr . DAKSKOBLER & MARTINČIČ: VEGETATION OF MOIST ROCK CREVICES AND MOIST (SLOPE) DEBRIS IN THE LIŠČAK GORGE 42 FOLIA BIOLOGICA ET GEOLOGICA 64/1 – 2023 A ru nc us d io ic us E1 . . . . + . . . . . . . . . . . + . 2 11 FS Fa ge ta lia sy lv at ic ae G al eo bd ol on fl av id um E1 . . . . . . . . . . . . + 1 . 1 + + 5 28 Fa gu s s yl va tic a E1 . . . . . . . . . r . . . . . . . . 1 6 M yc el is m ur al is E1 . . . . . . . . . . . . . . . r . . 1 6 G al iu m la ev ig at um E1 . . . . . . . . . . . . . . . . r . 1 6 Q F Q ue rc o- Fa ge te a C ar ex d ig ita ta E1 . . . . + . . . . . . . + . . + . . 3 17 M L M os es (M ah ov i) C te ni di um m ol lu sc um E0 . . . . . . 1 . . + 1 . 2 2 . . + . 6 33 To rt el la to rt uo sa E0 . . . . . . . + 1 + . . . . + . . . 4 22 Pl ag io m ni um u nd ul at um E0 . . . . + . . . . . . . + . . . . . 2 11 Pe di no ph yl lu m in te rr up tu m E0 . . . . . . . . . + . + . . . . . . 2 11 M ni um th om so ni i E0 . . . . . . . . . + . . 1 . . . . . 2 11 Pl ag io ch ila p or el lo id es E0 . . . . . . . . . + . . + . . . . . 2 11 Ex er to th ec a cr is pa (N ec ke ra c ri sp a) E0 . . . . . . . . . . . . . . . . 3 1 2 11 Po hl ia w ah le nb er gi i E0 . . 1 . . . . . . . . . . . . . . . 1 6 Lo ph oz ia sp . E0 . . . . . . + . . . . . . . . . . . 1 6 Br yu m sp . E0 . . . . . . + . . . . . . . . . . . 1 6 M et zg er ia c on ju ga ta E0 . . . . . . . . . + . . . . . . . . 1 6 Se rp ol es ke a co nf er vo id es (A m bl ys te gi um c on fe rv oi de s) E0 . . . . . . . . . + . . . . . . . . 1 6 C am py lo ph yl lu m h al le ri E0 . . . . . . . . . . . . 1 . . . . . 1 6 Lo ph oz ia o bt us a E0 . . . . . . . . . . . . 1 . . . . . 1 6 A m bl ys te gi um se rp en s E0 . . . . . . . . . . . . + . . . . . 1 6 D id ym od on a cu tu s E0 . . . . . . . . . . . . + . . . . . 1 6 M yu re lla ju la ce a E0 . . . . . . . . . . . . + . . . . . 1 6 T ha m no br yu m a lo pe cu ru m E0 . . . . . . . . . . . . . . . . + . 1 6 Le ge nd - Le ge nd a L Li m es to ne - ap ne ne c D D ol om ite - do lo m it Li L ith os ol - ka m ni šč e Pr . P re se nc e (n um be r of r el ev és in w hi ch th e sp ec ie s i s p re se nt ed ) - š te vi lo p op is ov , v k at er ih s e po ja vl ja v rs ta Fr . F re qu en cy in % - fr ek ve nc a v % Su cc es si ve n um be r of r el ev é (Z ap or ed na š te vi lk a po pi sa ) 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 Pr . Fr . DAKSKOBLER & MARTINČIČ: VEGETATION OF MOIST ROCK CREVICES AND MOIST (SLOPE) DEBRIS IN THE LIŠČAK GORGE 43FOLIA BIOLOGICA ET GEOLOGICA 64/1 – 2023 Ta bl e 3 (P re gl ed ni ca 3 ): Pa lu st ri el lo co m m ut at i-A st ra nt ie tu m ca rn io lic ae Su cc es si ve n um be r of r el ev é (Z ap or ed na š te vi lk a po pi sa ) 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 D at ab as e nu m be r of r el ev é (D el ov na š te vi lk a po pi sa ) 287209 287210 287211 287213 287154 275015 287207 287168 287216 287161 287165 287162 278123 281764 281768 281770 281772 282277 El ev at io n in m (N ad m or sk a vi ši na v m ) 32 0 32 0 32 0 32 0 34 0 35 0 35 0 45 0 30 0 45 0 45 0 45 0 40 0 75 0 75 0 77 0 78 5 59 0 A sp ec t ( Le ga ) N N N N N W N N W SE SW N W N W SE N W SW SW S S se Sl op e in d eg re es (N ag ib v s to pi nj ah ) 95 85 70 80 80 90 80 90 95 90 90 80 95 70 70 70 95 80 Pa re nt m at er ia l ( M at ič na p od la ga ) DCh DCh DCh DCh LMCh LMCh LMCh LMCh LClCh LMCh LMCh LMCh LMCh LMCh LMCh LMCh LM LMCh So il (T la ) Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li St on in es s i n % (K am ni to st v % ) 10 0 10 0 10 0 10 0 10 0 10 0 10 0 10 0 10 0 10 0 10 0 10 0 10 0 10 0 10 0 10 0 10 0 10 0 C ov er o f h er b la ye r in % (Z as ti ra nj e ze liš čn e pl as ti v % ): 35 30 30 30 25 30 30 30 30 30 20 40 35 60 30 60 30 20 C ov er o f m os s l ay er in % (Z as ti ra nj e m ah ov ne p la st i v % ): 20 10 10 5 40 70 60 40 70 50 30 70 40 60 30 60 50 70 N um be r of s pe ci es (Š te vi lo v rs t) 32 17 19 14 14 20 13 20 25 25 20 12 18 23 8 11 9 12 R el ev é ar ea (V el ik os t p op is ne p lo sk ve ) m 2 10 10 10 10 10 10 10 10 10 20 10 10 10 20 40 10 6 10 D at e of ta ki ng r el ev é (D at um p op is a) 6/18/2021 6/18/2021 6/18/2021 6/18/2021 8/11/2021 7/1/2018 6/18/2021 8/11/2021 6/18/2021 8/11/2021 8/11/2021 8/11/2021 6/26/2019 6/24/2020 6/24/2020 6/24/2020 6/24/2020 8/21/2020 Lo ca lit y (N ah aj al iš če ) Velike Luti Velike Luti Velike Luti Velike Luti Lisec-Knežica Liščak Liščak Liščak-Sopot Velike Luti Liščak Liščak-Sopot Liščak-Sopot Liščak-Drsela Liščak-Sopot Liščak-Sopot Liščak-Sopot Liščak-Sopot Liščak-Zališčar jeva grapa Q ua dr an t ( K va dr an t) 9849/1 9849/1 9849/1 9849/1 9849/1 9849/1 9849/1 9849/1 9849/1 9849/1 9849/1 9849/1 9849/1 9749/3 9749/3 9749/3 9749/3 9749/3 C oo rd in at e G K Y (D -4 8) m 410755 410761 410760 410786 410331 410897 410888 411283 411032 411350 411332 411306 411130 411706 411722 411718 411716 411241 C oo rd in at e G K X (D -4 8) m 5116796 5116801 5116817 5116820 5117078 5117034 5117016 5117644 5116513 5117654 5117654 5117650 5117352 5118602 5118604 5118620 5118648 5118083 D ia gn os ti c sp ec ie s of th e as so ci at io n (D ia gn os ti čn e vr st e as oc ia ci je ) Pr . Fr . A P A st ra nt ia c ar ni ol ic a E1 3 2 2 1 1 3 3 1 1 2 2 1 3 1 1 1 + 2 18 10 0 A P Pa lu st ri el la c om m ut at a E0 2 . . . 3 4 3 3 3 3 2 4 2 3 3 4 3 1 15 83 EP C al am ag ro st is v ar ia E1 . . + r + . + + 1 1 1 + . + + 1 1 + 14 78 EP M ol in ia a ru nd in ac ea E1 1 + 1 1 . . . . + . . . . 1 1 . 1 . 8 44 D ia gn os ti c sp ec ie s of lo w er u ni ts (D ia gn os ti čn e vr st e ni žj ih e no t) A P A st er b el lid ia st ru m E1 2 2 1 1 2 . . . . . . . . . . . . . 5 28 A P H yd ro go ni um c ro ce um (B ar bu la c ro ce a) E0 + 1 2 + + . . . . . + . . . . . . . 6 33 ES C ar ex m uc ro na ta E1 + + + 1 . . . . . . . . . . . . . . 4 22 A P V al er ia na sa xa til is E1 + + + r . . . . . . . . . . . . . . 4 22 DAKSKOBLER & MARTINČIČ: VEGETATION OF MOIST ROCK CREVICES AND MOIST (SLOPE) DEBRIS IN THE LIŠČAK GORGE 44 FOLIA BIOLOGICA ET GEOLOGICA 64/1 – 2023 Su cc es si ve n um be r of r el ev é (Z ap or ed na š te vi lk a po pi sa ) 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 Pr . Fr . A P A st ra nt io ca rn io lic ae -P ae de ro ti on lu te ae H ym en os ty liu m re cu rv ir os tr um E0 1 . . . . 2 . 1 3 1 1 3 3 . 1 1 2 1 12 67 C ar ex b ra ch ys ta ch ys E1 . . + + . + + . + 1 + . + + . + + . 11 61 A po pe lli a en di vi ifo lia (P el lia e nd iv iif ol ia ) E0 1 + . . . 1 . 1 1 1 1 + . 1 . . . 1 10 56 O rt ho th ec iu m r uf es ce ns E0 + 1 . + + . + . . 1 1 . 1 + . . . . 9 50 Pi ng ui cu la a lp in a E1 1 + + . . . . . + + . . . . . . . . 5 28 Eu cl ad iu m v er tic ill at um E0 . . . . . + 1 1 + . . . + . . . . . 5 28 Pa ed er ot a lu te a E1 1 + + . . . . . . . . . . . . . 2 . 4 22 Fi ss id en s d ub iu s E0 1 . . . . . . . . + + . . . . . . . 3 17 To fie ld ia c al yc ul at a E1 + . + + . . . . . . . . . . . . . . 3 17 A sp le ni um v ir id e E1 . . . . . . . . + 2 . . + . . . . . 3 17 V al er ia na tr ip te ri s E1 . . . . + . . . . . + . . . . . . . 2 11 M ar ch an tia q ua dr at a (P re is si a qu ad ra ta ) E0 . . . . . 1 . . . . . . . . . . . . 1 5 Ps Sp Ph ys op le xi do co m os ae -S ax if ra gi on p et ra ea e H ie ra ci um p os pi ch al ii E1 . . . . . . . . . + . . . . . . . . 1 6 Sa xi fr ag a cr us ta ta E1 . . . . . . . . . . . . . . . . . + 1 6 H ie ra ci um p or ri fo liu m E1 . . . . . . . . . . . . . . . . . + 1 6 A T A sp le ni et ea tr ic ho m an is A sp le ni um tr ic ho m an es E1 . . . . . + + + . . + . + . . . . . 5 28 K er ne ra sa xa til is E1 . . . . . . . . + . . . . . . . . . 1 6 Po ly po di um in te rj ec tu m E1 . . . . . . . . + . . . . . . . . . 1 6 A sp le ni um r ut a- m ur ar ia E1 . . . . . . . . . + . . . . . . . . 1 6 T R T hl as pi et ea ro tu nd ifo lii A de no st yl es g la br a E1 . . . . . . . + . 1 + 2 . 3 1 3 2 . 8 44 H ie ra ci um b ifi du m E1 . . + . . . . . + . . . . . . . . + 3 17 G ym no ca rp iu m ro be rt ia nu m E1 + . . . . . . . . . . . . . . . . . 1 6 Pe uc ed an um v er tic ill ar e E1 . . . r . . . . . . . . . . . . . . 1 6 A ch na th er um c al am ag ro st is E1 . . . . + . . . . . . . . . . . . . 1 6 Pe ta si te s p ar ad ox us E1 . . . . . . . . + . . . . . . . . . 1 6 M C M on ti o- C ar da m in et ea C on oc ep ha lu m c on ic um E0 + . . . . 1 . 1 1 2 + . + 1 . 1 . 4 10 56 O xy rr hy nc hi um h ia ns E0 . . . . . . + + . . . . + + . . . . 4 22 C ra to ne ur on fi lic in um E0 + + . . . . . . . . . . 3 . . . . 3 17 Br ac hy th ec iu m r iv ul ar e E0 + . . . . . . . + . . . . + . . . . 3 17 Pt yc ho st om um p se ud ot ri qu et ru m (B ry um p se ud ot ri qu et ru m ) E0 + . . . . . . . . + . . . . . . . . 2 11 O xy rr hy nc hi um sc hl ei ch er i E0 . . . . . + . . . . . . . . . . . . 1 6 G ym no st om um a er ug in os um E0 . . . . . . 2 . . . . . . . . . . . 1 6 R hy nc ho st eg iu m r ip ar io id es (P la ty hy pn id iu m r ip ar io id es ) E0 . . . . . . . . . . . . . . + . . . 1 6 C D C ar ic et al ia d av al lia na e C am py liu m st el la tu m E0 . . . . . + . . . . . . . . . . . . 1 6 ES El yn o- Se sl er ie te a Se sl er ia c ae ru le a E1 1 1 + + . . . . . 1 . . . . . + . . 6 33 Be to ni ca a lo pe cu ro s E1 r + . . . . . . . . . . . . . . . . 2 11 M A M ol in io n, M ol in io -A rr he na th er et ea A ng el ic a sy lv es tr is E1 r . . . . . . . + + . . . . . . . . 3 17 DAKSKOBLER & MARTINČIČ: VEGETATION OF MOIST ROCK CREVICES AND MOIST (SLOPE) DEBRIS IN THE LIŠČAK GORGE 45FOLIA BIOLOGICA ET GEOLOGICA 64/1 – 2023 C re pi s p al ud os a E1 . . . . . . . . . . . . + 1 . 1 . . 3 17 C al th a pa lu st ri s E1 . . . . . . . . . r . . . . . . . . 1 6 BA Be tu lo -A ln et ea Sa lix a pp en di cu la ta E2 a . . . . + . . . . . . . . . . . . . 1 6 M uA M ul ge di o- A co ni te te a A co ni tu m a ng us tif ol iu m E1 . . . . . . . . . . . . . + . . . . 1 6 C ha er op hy llu m h ir su tu m E1 + . . . . . . . . . . . . . . . . . 1 6 Se ne ci o ov at us E1 . . . . . . . . . . . + . . . . . . 1 6 EA Ep ilo bi et ea a ng us ti fo lii , T ri fo lio -G er an ie te a Eu pa to ri um c an na bi nu m E1 . . . . . . . . . . . . . 1 + + + . 4 22 V er ba sc um la na tu m E1 . . . . . . . . + . . . . . . . . . 1 6 So la nu m d ul ca m ar a E1 . . . . . . . . . . . . . + . . . . 1 6 EP Er ic o- Pi ne te a Bu ph th al m um sa lic ifo liu m E1 . . . . . . . . . + r . . . . . . + 3 17 C ir si um e ri si th al es E1 . . . . . . . . . . . . . r . . . . 1 6 V P Va cc in io -P ic ee te a V er on ic a ur tic ifo lia E1 . . . + + + + + 1 + . . + 1 . + . + 11 61 G en tia na a sc le pi ad ea E1 r . + . . . . . . + . . . . . . . . 3 17 H om og yn e sy lv es tr is E1 . . . . . r . . . . . . . . . . . . 1 6 A po se ri s f oe tid a E1 . . . . . . . . + . . . . . . . . . 1 6 Sa xi fr ag a cu ne ifo lia E1 . . . . . . . . . . . . + . . . . . 1 6 A F A re m on io -F ag io n, E ry th ro ni o- C ar pi ni on C ar da m in e tr ifo lia E1 . . . . . . . . . . . + . + . . . . 2 11 A ne m on e tr ifo lia E1 r . . . . . . . . . . . . . . . . . 1 6 La m iu m o rv al a E1 . . . . . . . . + . . . . . . . . . 1 6 TA Ti lio -A ce ri on A ru nc us d io ic us E1 . . . . . . . + + + . + 1 . . . . . 5 28 G er an iu m ro be rt ia nu m E1 . . . . . . . + + . . . . + . . . . 3 17 Ph yl lit is sc ol op en dr iu m E1 . . . . . . . + . . + . . . . . . . 2 11 U lm us g la br a E1 . . . . . . . . . + + . . . . . . . 2 11 Po ly st ic hu m a cu le at um E1 . . . . . . . . . . + . + . . . . . 2 11 A ce r p se ud op la ta nu s E1 . . . . . . . . . . . + . . . . . . 1 6 A I A ln io n in ca na e K na ut ia d ry m ei a su bs p. in te rm ed ia E1 . . . . + . . . + . . . . . . . . . 2 11 C ar da m in e im pa tie ns . . . . . . . . . . . . . + . . . . 1 6 FS Fa ge ta lia sy lv at ic ae G al iu m la ev ig at um E1 + + . + + . + + . + + . . . . . . . 8 44 G al eo bd ol on fl av id um E1 . + . . . 1 + 1 + . . 1 + . . . . . 7 39 Br ac hy po di um sy lv at ic um E1 . . . . . . . + . . . 1 + . . . . . 3 17 Pe ta si te s a lb us E1 . . . . . + 1 . . . . . . . . . . . 2 11 M yc el is m ur al is E1 . . . . . . . . + . . . . + . . . . 2 11 Fr ax in us e xc el si or E1 . . . . . r . . . . . . . . . . . . 1 6 D ry op te ri s f ili x- m as E1 . . . . . . . . + . . . . . . . . . 1 6 Sa lv ia g lu tin os a E1 . . . . . . . . . . + . . . . . . . 1 6 Q P Q ue rc et al ia p ub es ce nt i- pe tr ae ae C ar ex fl ac ca E1 . . + . . . . . . . . . . + . . . . 2 11 Su cc es si ve n um be r of r el ev é (Z ap or ed na š te vi lk a po pi sa ) 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 Pr . Fr . DAKSKOBLER & MARTINČIČ: VEGETATION OF MOIST ROCK CREVICES AND MOIST (SLOPE) DEBRIS IN THE LIŠČAK GORGE 46 FOLIA BIOLOGICA ET GEOLOGICA 64/1 – 2023 Q F Q ue rc o- Fa ge te a H ed er a he lix E1 . . . . . r . + . . . . . . . . . . 2 11 C ar ex d ig ita ta E1 . . . . . . . . . . . . + . . . . + 2 11 Po te nt ill a er ec ta E1 . . + . . . . . . . . . . . . . . . 1 6 M L M os se s (M ah ov i) Pl ag io m ni um ro st ra tu m E0 + . . . . + . + . . . . . . . . . . 3 17 H yg ro hy pn um lu ri du m E0 + + . . . . . . . . . . . . . . . . 2 11 M ni um m ar gi na tu m E0 . . . . . + . + . . . . . . . . . . 2 11 Ju ng er m an ni a at ro vi re ns E0 + . . . . . . . . . . . . . . . . . 1 6 Lo ph oz ia sp . E0 + . . . . . . . . . . . . . . . . . 1 6 D ic ho do nt iu m p el lu ci du m E0 + . . . . . . . . . . . . . . . . . 1 6 Tr ic ho st om um c ri sp ul um E0 + . . . . . . . . . . . . . . . . . 1 6 C hi on ol om a te nu ir os tr e (O xy st eg us te nu ir os tr e) E0 . + . . . . . . . . . . . . . . . . 1 6 M ic ro le je un ea u lic in a E0 . . + . . . . . . . . . . . . . . . 1 6 A lle ni el la c om pl an at a (N ec ke ra c om pl an at a) E0 . . . . + . . . . . . . . . . . . . 1 6 C te ni di um m ol lu sc um E0 . . . . . . . . . + . . . . . . . . 1 6 Pl ag io ch ila p or el lo id es E0 . . . . . . . . . . + . . . . . . . 1 6 Le ge nd - Le ge nd a L Li m es to ne - ap ne ne c C h C he rt - ro že ne c D D ol om ite - do lo m it C l C la ys to ne - gl in av ec M M ar ls to ne - la po ro ve c Li L ith os ol - ka m ni šč e Pr . P re se nc e (n um be r of r el ev és in w hi ch th e sp ec ie s i s p re se nt ed ) - š te vi lo p op is ov , v k at er ih s e po ja vl ja v rs ta Fr . F re qu en cy in % - fr ek ve nc a v % Su cc es si ve n um be r of r el ev é (Z ap or ed na š te vi lk a po pi sa ) 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 Pr . Fr . DAKSKOBLER & MARTINČIČ: VEGETATION OF MOIST ROCK CREVICES AND MOIST (SLOPE) DEBRIS IN THE LIŠČAK GORGE 47FOLIA BIOLOGICA ET GEOLOGICA 64/1 – 2023 Table 4 (Preglednica 4): Palustriello commutati-Veronicetum urticifoliae, Cratoneuretum commutati Successive number of relevé (Zaporedna številka popisa) 1 2 3 4 5 6 7 8 9 10 1 Database number of relevé (Delovna številka popisa) 27 50 14 27 50 16 28 26 95 28 27 18 28 71 43 28 17 69 28 27 01 28 27 02 28 71 48 28 71 67 28 71 74 Elevation in m (Nadmorska višina v m) 350 350 530 650 940 770 540 540 870 450 350 Aspect (Lega) N N N EE SE E SW S E N W N W W S N W N W Slope in degrees (Nagib v stopinjah) 70 90 90 85 85 80 80 80 80 95 90 Parent material (Matična podlaga) LM C h LM C h LC h D C hC l LC lC h LM C h LC h LC h C l LM C h LM C h Soil (Tla) Li Li Li Li Li Li Li Li Li Li Hy Stoniness in % (Kamnitost v %) 100 100 100 100 100 100 100 100 100 100 100 Cover of herb layer in % (Zastiranje zeliščne plasti v %): 40 40 30 30 30 50 30 10 10 20 Cover of moss layer in % (Zastiranje mahovne plasti v %): 30 80 30 40 40 30 70 90 80 80 80 Number of species (Število vrst) 17 21 28 20 23 9 25 16 13 20 3 Relevé area (Velikost popisne ploskve) m2 10 10 10 15 20 6 15 15 5 20 10 Date of taking relevé (Datum popisa) 7/ 1/ 20 18 7/ 1/ 20 18 9/ 15 /2 02 0 9/ 15 /2 02 0 8/ 10 /2 02 1 6/ 24 /2 02 0 9/ 15 /2 02 0 9/ 15 /2 02 0 8/ 10 /2 02 1 8/ 11 /2 02 1 6/ 18 /2 02 1 Locality (Nahajališče) Li šč ak Li šč ak Li šč ak -Z al iš ča r je va g ra pa Li šč ak Li šč ak p ov ir je Li se c- So po t Li šč ak Li šč ak Li šč ak p ov ir je Li se c- So po t Li šč ak Quadrant (Kvadrant) 98 49 /1 98 49 /1 97 49 /3 97 49 /3 97 49 /3 97 49 /3 97 49 /3 97 49 /3 97 49 /3 98 49 /1 98 49 /1 Coordinate GK Y (D-48) m 41 09 09 41 08 78 41 13 47 41 11 49 41 17 39 41 17 15 41 14 61 41 14 64 41 17 06 41 12 90 41 09 26 Coordinate GK X (D-48) m 51 17 04 4 51 17 00 8 51 17 94 6 51 18 05 5 51 18 91 6 51 18 61 3 51 17 96 5 51 17 98 4 51 18 79 9 51 17 61 4 51 17 05 3 Diagnostic species of the association (Dignostične vrste asociacije) Pr. Fr. AP Palustriella commutata E0 2 2 1 3 1 1 3 5 4 4 10 100 3 MC Conocephalum conicum E0 2 2 2 3 3 + + + + 1 10 100 VP Veronica urticifolia E1 3 3 + 1 + 1 + + + + 10 100 AT Asplenium trichomanes E1 1 + 2 1 1 + + + + + 10 100 FS Galeobdolon flavidum E1 . 1 + 1 1 . 2 . 1 1 7 70 MuASenecio ovatus E1 . . + 1 + . + + . . 5 50 AP Astrantio carniolicae-Paederotion luteae Hymenostylium recurvirostrum E0 1 . + . . . 2 1 . 1 5 50 Apopellia endiviifolia (Pellia endiviifolia) E0 . . 1 + . . + + + + 6 60 Fissidens dubius E0 . . + . . . 1 + . 1 4 40 Valeriana tripteris E1 . . + . r . . + . . 3 30 Astrantia carniolica E1 1 . . . . . . + . . 2 20 Cystopteris fragilis E1 . . . 1 2 . . . . . 2 20 Asplenium viride E1 + . . . . . . . . . 1 10 Eucladium verticillatum E0 . . + . . . . . . . 1 10 4 Carex brachystachys E1 . . + . . . . . . . 1 10 Paederota lutea E1 . . . . . 1 . . . . 1 10 Orthothecium rufescens E0 . . . . . . 1 . . . 1 10 PsSp Physoplexido comosae-Saxifragion petraeae Campanula carnica E1 . . . . r 1 . . + . 3 30 PC Potentilletalia caulescentis Saxifraga crustata E1 . . . . . . . + . . 1 10 AT Asplenietea trichomanis Asplenium ruta-muraria E1 . . + . . + + . . . 3 30 DAKSKOBLER & MARTINČIČ: VEGETATION OF MOIST ROCK CREVICES AND MOIST (SLOPE) DEBRIS IN THE LIŠČAK GORGE 48 FOLIA BIOLOGICA ET GEOLOGICA 64/1 – 2023 TR Thlaspietea rotundifolii Adenostyles glabra E1 + . . + . 1 + . + . 5 50 Hieracium bifidum E1 . . + . . . . . . . 1 10 Gymnocarpium robertianum E1 . . . r . . . . . . 1 10 MC Montio-Cardaminetea Oxyrrhynchium schleicheri E0 . . 1 + + . 1 . . . 4 40 Oxyrrhynchium hians E0 . . + . . . + + . 1 4 40 Brachythecium rivulare E0 . . . + + . . . . . 2 20 Rhynchostegium riparioides E0 . . . . 1 . . . 1 . 2 20 + Cratoneuron filicinum E0 . . . . . . . . + . 1 10 CD Caricetalia davallianae Calliergonella lindbergii E0 . . + . . . . . . . 1 10 ES Elyno-Seslerietea Sesleria caerulea E1 . . . . . 2 . . . . 1 10 EA Epilobietea angustifolii Rubus idaeus E1 . + . . . . . . . . 1 10 TG Trifolio-Geranietea Campanula rapunculoides E1 . . + . . . + . . . 2 20 Digitalis grandiflora E1 . . + . . . . . . . 1 10 EP Erico-Pinetea, Festuco-Brometea Calamagrostis varia E1 . . . . + . . . . + 2 20 Cirsium erisithales E1 . . . . + . . . . . 1 10 Buphthalmum salicifolium E1 . . . . . . + . . . 1 10 VP Vaccinio-Piceetea Oxalis acetosella E1 + . + . . . . . . + 3 30 Saxifraga cuneifolia E1 . + . . . . 1 . . . 2 20 Solidago virgaurea E1 . . . + r . . . . . 2 20 Calamagrostis arundinacea E1 . + . . . . . . . . 1 10 AF Aremonio-Fagion, Erythronio-Carpinion Cardamine trifolia E1 + + . . . . 1 1 + 1 6 60 Lamium orvala E1 + . . . . . . . + . 2 20 Primula vulgaris E1 . . . . + . . . . . 1 10 TA Tilio-Acerion Geranium robertianum E1 . . . 1 + . . . . + 3 30 Aruncus dioicus E1 . + . . . . + . . . 2 20 Phyllitis scolopendrium E1 . 1 . . . . . . . 1 2 20 Polystichum setiferum E1 . + . . . . . . . . 1 10 Polystichum x wirtgenii E1 . + . . . . . . . . 1 10 Ulmus glabra E1 . + . . . . . . . . 1 10 Circaea x intermedia E1 . . . . + . . . . . 1 10 FS Fagetalia sylvaticae Mycelis muralis E1 . + . . + . + + . . 4 40 Galium laevigatum E1 . + . . . . + . . + 3 30 Petasites albus E1 + 1 . . . . . . . . 2 20 Fraxinus excelsior E1 + + . . . . . . . . 2 20 Salvia glutinosa E1 . . . + + . . . . . 2 20 Festuca altissima E1 + . . . . . . . . . 1 10 Asarum europaeum subsp. caucasicum E1 . . + . . . . . . . 1 10 Campanula trachelium E1 . . . . + . . . . . 1 10 Poa nemoralis E1 . . . . + . . . . . 1 10 Sambucus nigra E2a . . . . . . . . . + 1 10 QP Quercetalia pubescenti-petraeae Sesleria autumnalis E1 . . + + + . . . . . 3 30 Arabis turrita E1 . . + + . . . . . . 2 20 QF Querco-Fagetea Hedera helix E1 1 + + . . . + + . . 5 50 Carex digitata E1 1 . + + . . + . . . 4 40 Veratrum nigrum E1 + . . . . . . . . . 1 10 Clematis vitalba E1 . . . + . . . . . . 1 10 ML Mosses (Mahovi) Plagiomnium rostratum E0 . . + . . . + + . + 4 40 Thamnobryum alopecurum E0 . 2 . . . . + . . 1 3 30 Successive number of relevé (Zaporedna številka popisa) 1 2 3 4 5 6 7 8 9 10 Pr. Fr. 1 DAKSKOBLER & MARTINČIČ: VEGETATION OF MOIST ROCK CREVICES AND MOIST (SLOPE) DEBRIS IN THE LIŠČAK GORGE 49FOLIA BIOLOGICA ET GEOLOGICA 64/1 – 2023 Mnium thomsonii E0 . . 1 . . . . . . 1 2 20 Plagiomnium undulatum E0 . 2 . . . . . . . . 1 10 Tortella tortuosa E0 . . + . . . . . . . 1 10 Mnium stellare E0 . . . 1 . . . . . . 1 10 Pedinophyllum interruptum E0 . . . . . . . . . + 1 10 Legend - Legenda L Limestone - apnenec Ch Chert - roženec D Dolomite - dolomit Cl Claystone - glinavec M Marlstone - laporovec Li Lithosol - kamnišče Hy Hygromorphic soil - higromorfna tla Pr. Presence (number of relevés in which the species is presented) - število popisov, v katerih se pojavlja vrsta Fr. Frequency in % - frekvenca v % Successive number of relevé (Zaporedna številka popisa) 1 2 3 4 5 6 7 8 9 10 Pr. Fr. 1 DAKSKOBLER & MARTINČIČ: VEGETATION OF MOIST ROCK CREVICES AND MOIST (SLOPE) DEBRIS IN THE LIŠČAK GORGE 50 FOLIA BIOLOGICA ET GEOLOGICA 64/1 – 2023 Table 5 (Preglednica 5): Campanulo cespitosae-Saxifragetum aizoidis achnatheretosum calamagrostis Successive number of relevé (Zaporedna številka popisa) 1 2 3 4 5 6 7 8 9 10 Database number of relevé (Delovna številka popisa) 28 17 76 28 27 04 28 27 05 28 27 09 28 27 06 28 27 08 28 27 14 28 27 07 28 27 11 28 27 15 Elevation in m (Nadmorska višina v m) 740 750 740 725 740 730 750 735 785 750 Aspect (Lega) SE SSW S S W S SW S SW NE Slope in degrees (Nagib v stopinjah) 70 80 80 90 70 80 80 70 75 80 Parent material (Matična podlaga) LM C h LC hC l LC hC l LC hC l LC hC l LC hC l LC hC l LC hC l LC hC l LC hC l Soil (Tla) Li Li Li Li Li Li Li Li Li Li Stoniness in % (Kamnitost v %) 100 100 100 100 100 100 100 100 100 100 Cover of shrub layer in % (Zastiranje grmovne plasti v %): Cover of herb layer in % (Zastiranje zeliščne plasti v %): 40 30 30 40 40 60 30 50 60 20 Cover of moss layer in % (Zastiranje mahovne plasti v %): 5 10 5 20 20 10 20 15 20 10 Number of species (Število vrst) 17 19 10 13 12 14 9 18 20 15 Relevé area (Velikost popisne ploskve) m2 10 10 10 10 10 10 10 10 10 10 Date of taking relevé (Datum popisa) 6/ 24 /2 02 0 9/ 15 /2 02 0 9/ 15 /2 02 0 9/ 15 /2 02 0 9/ 15 /2 02 0 9/ 15 /2 02 0 9/ 15 /2 02 0 9/ 15 /2 02 0 9/ 15 /2 02 0 9/ 15 /2 02 0 Locality (Nahajališče) Li se c- G ra dn ik ov a gr ap a Li šč ak -L uk no va gr ap a Li šč ak -L uk no va gr ap a Li šč ak -L uk no va gr ap a Li šč ak -L uk no va gr ap a Li šč ak -L uk no va gr ap a Li šč ak -L uk no va gr ap a Li šč ak -L uk no va gr ap a Li šč ak -L uk no va gr ap a Li šč ak -L uk no va gr ap a Quadrant (Kvadrant) 97 49 /3 98 49 /1 98 49 /1 98 49 /1 98 49 /1 98 49 /1 98 49 /1 98 49 /1 98 49 /1 98 49 /1 Coordinate GK Y (D-48) m 41 15 16 41 20 04 41 19 87 41 19 51 41 19 91 41 19 61 41 20 03 41 19 79 41 20 52 41 20 19 Coordinate GK X (D-48) m 51 18 54 9 51 17 84 1 51 17 84 3 51 17 83 8 51 17 84 4 51 17 83 9 51 17 84 5 51 17 84 5 51 17 84 8 51 17 82 6 Diagnostic species of the association (Diagnostične vrste asociacije) Pr. Fr. MC Saxifraga aizoides E1 + 2 1 2 2 2 2 2 3 1 10 100 PcSp Campanula cespitosa E1 1 1 + 1 1 1 + 1 1 1 10 100 EP Molinia arundinacea E1 1 1 . + 1 . . 2 + . 6 60 AP Palustriella commutata E0 + . . . 1 . . 1 . . 3 30 EP Calamagrostis varia E1 . . . . . . + . . . 1 10 Differential species of subassociation (Razlikovalnice subasociacije) TR Achnatherum calamagrostis E1 + 1 2 2 3 4 2 1 1 1 10 100 ML Tortella tortuosa E0 1 1 + 1 2 2 2 1 1 . 9 90 TR Hieracium glaucum E1 . + + + + + + + + + 9 90 PC Saxifraga crustata E1 + + 1 1 . + . + . 1 7 70 TG Calamintha einseleana E1 . + + 1 1 1 . + r . 7 70 AP Astrantio carniolicae-Paederotion luteae Hymenostylium recurvirostrum E0 + . . . . . . . + . 2 20 Apopellia endiviifolia (Pellia endiviifolia) E0 . + . + . . . . + . 3 30 Hydrogonium croceum (Barbula crocea) E0 . . . + . . . . + . 2 20 Aster bellidiastrum E1 . . . . + . . . . . 1 10 PcSp Physoplexido comosae-Saxifragion petraeae Hieracium porrifolium E1 1 1 + + . . 1 + + 1 8 80 TR Thlaspietea rotundifolii Hieracium bifidum E1 . . . . . . . + r 1 3 30 Trisetum argenteum E1 + . . . . . . + + . 3 30 Petasites paradoxus E1 . . . . . + . . . . 1 10 Adenostyles glabra E1 . . . . . . . . . + 1 10 Hieracium dollineri E1 . . . . . . . . . + 1 10 DAKSKOBLER & MARTINČIČ: VEGETATION OF MOIST ROCK CREVICES AND MOIST (SLOPE) DEBRIS IN THE LIŠČAK GORGE 51FOLIA BIOLOGICA ET GEOLOGICA 64/1 – 2023 ES Elyno-Seslerietea Sesleria caerulea E1 . . . . . . . + 1 1 3 30 Carex ferruginea E1 . . . . . . . + . . 1 10 FB Festuco-Brometea Bromopsis erecta E1 1 . . . . + . . . . 2 20 Carlina acaulis E1 . . . . . + . . r . 2 20 Brachypodium rupestre E1 + . . . . . . . . . 1 10 Carex humilis E1 + . . . . . . . . . 1 10 Buphthalmum salicifolium E1 . + . . . . . . . . 1 10 Pimpinella saxifraga E1 . + . . . . . . . . 1 10 Thymus praecox E1 . + . . . . . . . . 1 10 TG Trifolio-Geranietea Hypericum perforatum E1 . . . . . . . + r . 2 20 Libanotis daucifolia E1 + . . . . + . . . . 2 20 EA Epilobietea angustifolii Eupatorium cannabinum E1 . r . . . . . + . . 2 20 Tussilago farfara E1 . . . . 1 . . + . . 2 20 BA Betulo-Alnetea Salix appendiculata E2a . . . + . . + . . . 2 20 EP Erico-Pinetea Epipactis atrorubens E1 + . . . . . . . . . 1 10 Erica carnea E1 . . . . . . . . . + 1 10 VP Vaccinio-Piceetea Picea abies E2a . r . . . . . . . . 1 10 Larix decidua E2a . . r . . . . . . . 1 10 Veronica urticifolia E1 . . . . . . . . . + 1 10 TA Tilio-Acerion Geranium robertianum E1 . . . + . + . . . . 2 20 EC Erythronio-Carpinion Primula vulgaris E1 . r . . . . . . . . 1 10 QP Quercetalia pubescenti-petraeae Ostrya carpinifolia E2a . + 1 + + . . 1 + . 6 60 Fraxinus ornus E3a . . . . + . . . . . 1 10 Fraxinus ornus E2a + . . . . + + . 1 + 5 50 Sesleria autumnalis E1 . + . . . . . . . + 2 20 Carex flacca E1 + . . . . . . . . . 1 10 ML Mosses (Mahovi) Pohlia wahlenbergii E0 . + . + . + . . + . 4 40 Didymodon vinealis E0 . . . . . . . . + . 1 10 Ctenidium molluscum E0 . . . . . . . . . 1 1 10 Legend - Legenda L Limestone - apnenec Ch Chert - roženec Cl Claystone - glinavec M Marlstone - laporovec Li Lithosol - kamnišče Pr. Presence (number of relevés in which the species is presented) - število popisov, v katerih se pojavlja vrsta Fr. Frequency in % - frekvenca v % Successive number of relevé (Zaporedna številka popisa) 1 2 3 4 5 6 7 8 9 10 DAKSKOBLER & MARTINČIČ: VEGETATION OF MOIST ROCK CREVICES AND MOIST (SLOPE) DEBRIS IN THE LIŠČAK GORGE 52 FOLIA BIOLOGICA ET GEOLOGICA 64/1 – 2023 Table 6 (Preglednica 6): Veronico urticifoliae-Saxifragetum cuneifolii Successive number of relevé (Zaporedna številka popisa) 1 2 3 4 5 6 7 8 9 10 11 Database number of relevé (Delovna številka popisa) 21 51 09 26 11 82 26 17 97 26 17 98 26 82 60 26 90 96 26 90 98 27 99 22 26 10 06 26 17 96 26 12 27 Elevation in m (Nadmorska višina v m) 160 450 300 300 710 850 825 590 260 240 320 Aspect (Lega) N W N W N N E E W W W N N E N N E Slope in degrees (Nagib v stopinjah) 80 90 85 80 80 90 90 80 80 100 95 Parent material (Matična podlaga) L L Br L C l C l LC h L L L L Soil (Tla) Li Li Li Li Li Li Li Li Li Li Li Stoniness in % (Kamnitost v %) 100 100 100 100 100 100 100 100 100 100 100 Cover of herb layer in % (Zastiranje zeliščne plasti v %): 30 17 20 15 40 20 30 30 30 20 10 Cover of moss layer in % (Zastiranje mahovne plasti v %): 30 20 10 10 60 30 20 20 20 30 10 Number of species (Število vrst) 20 17 13 12 13 10 16 11 17 14 8 Relevé area (Velikost popisne ploskve) m2 10 10 10 10 5 5 10 20 10 10 10 Date of taking relevé (Datum popisa) 4/ 3/ 20 07 4/ 29 /2 01 6 5/ 3/ 20 16 5/ 3/ 20 16 9/ 4/ 20 17 5/ 17 /2 01 7 5/ 17 /2 01 7 4/ 11 /2 02 0 4/ 22 /2 01 6 5/ 3/ 20 16 4/ 29 /2 01 6 Locality (Nahajališče) Sr nj ak N a V rh u Pl av e- So pe t Pl av e- So pe t Z ap oš ka g ra pa K ac en po h- K ob la K ac en po h K ac en po h Pl av e- St rm ec Pl av e- So pe t N a V rh u- St rm ec Quadrant (Kvadrant) 99 47 /4 99 47 /4 99 47 /3 99 47 /3 98 49 /2 97 49 /4 97 49 /4 97 49 /4 99 47 /3 99 47 /3 99 47 /3 Coordinate GK Y (D-48) m 39 14 58 39 03 53 38 98 92 38 98 55 42 12 65 42 01 53 42 01 33 42 02 27 39 03 27 38 99 77 39 02 29 Coordinate GK X (D-48) m 50 98 74 2 50 99 66 0 51 00 08 2 51 00 08 8 51 13 48 8 51 20 93 4 51 20 99 1 51 19 98 3 51 00 05 5 51 00 23 4 50 99 85 5 Diagnostic species of the association (Diagnostične vrste asociacije) VP Saxifraga cuneifolia E1 2 1 1 + 3 2 2 2 1 2 1 AT Asplenium trichomanes E1 1 1 1 1 1 1 1 1 + 1 1 VP Veronica urticifolia E1 1 . + 1 1 + + + 2 + + ML Exertotheca crispa (Neckera crispa) E0 + 2 2 1 3 2 2 2 . . . AP Valeriana tripteris E1 . + . . 1 . + . . . . Differential species of lower units (Razlikovalnice nižjih enot) FS Galeobdolon flavidum E1 . . + . . . . . + + . PA Fissidens dubius E0 . + . . 1 . . . . . + TA Phyllitis scolopendrium E1 . . . . . . . + . . . AP Astrantio carniolicae-Paederotion luteae Palustriella commutata E0 . . + . . . . . + 1 . Cystopteris fragilis E1 . . . . + + . . . . . Orthothecium rufescens E0 + . . . . . . . . . . Carex brachystachys E1 . . . . . . + . . . . Apopellia endiviifolia (Pellia endiviifolia) E0 . . + . . . . . + + . Hydrogonium croceum (Barbula crocea) E0 . . . . . . . . . . . Aster bellidiastrum E1 r . . . . . . . . . . Hymenostylium recurvirostrum E0 . . . . . 2 . . . . . Asplenium viride E1 . . . . . . + + . . . Eucladium verticillatum E0 . . . . . . . . . 2 . Paederota lutea E1 . . . . . . + . . . . Viola biflora E1 . . . . . . + . . . . Jungermannia atrovirens E0 . . . . . . . . . . . Astrantia carniolica E1 . . . . . . . . . . . DAKSKOBLER & MARTINČIČ: VEGETATION OF MOIST ROCK CREVICES AND MOIST (SLOPE) DEBRIS IN THE LIŠČAK GORGE 53FOLIA BIOLOGICA ET GEOLOGICA 64/1 – 2023 Table 6 (Preglednica 6): Veronico urticifoliae-Saxifragetum cuneifolii Successive number of relevé (Zaporedna številka popisa) 1 2 3 4 5 6 7 8 9 10 11 Database number of relevé (Delovna številka popisa) 21 51 09 26 11 82 26 17 97 26 17 98 26 82 60 26 90 96 26 90 98 27 99 22 26 10 06 26 17 96 26 12 27 Elevation in m (Nadmorska višina v m) 160 450 300 300 710 850 825 590 260 240 320 Aspect (Lega) N W N W N N E E W W W N N E N N E Slope in degrees (Nagib v stopinjah) 80 90 85 80 80 90 90 80 80 100 95 Parent material (Matična podlaga) L L Br L C l C l LC h L L L L Soil (Tla) Li Li Li Li Li Li Li Li Li Li Li Stoniness in % (Kamnitost v %) 100 100 100 100 100 100 100 100 100 100 100 Cover of herb layer in % (Zastiranje zeliščne plasti v %): 30 17 20 15 40 20 30 30 30 20 10 Cover of moss layer in % (Zastiranje mahovne plasti v %): 30 20 10 10 60 30 20 20 20 30 10 Number of species (Število vrst) 20 17 13 12 13 10 16 11 17 14 8 Relevé area (Velikost popisne ploskve) m2 10 10 10 10 5 5 10 20 10 10 10 Date of taking relevé (Datum popisa) 4/ 3/ 20 07 4/ 29 /2 01 6 5/ 3/ 20 16 5/ 3/ 20 16 9/ 4/ 20 17 5/ 17 /2 01 7 5/ 17 /2 01 7 4/ 11 /2 02 0 4/ 22 /2 01 6 5/ 3/ 20 16 4/ 29 /2 01 6 Locality (Nahajališče) Sr nj ak N a V rh u Pl av e- So pe t Pl av e- So pe t Z ap oš ka g ra pa K ac en po h- K ob la K ac en po h K ac en po h Pl av e- St rm ec Pl av e- So pe t N a V rh u- St rm ec Quadrant (Kvadrant) 99 47 /4 99 47 /4 99 47 /3 99 47 /3 98 49 /2 97 49 /4 97 49 /4 97 49 /4 99 47 /3 99 47 /3 99 47 /3 Coordinate GK Y (D-48) m 39 14 58 39 03 53 38 98 92 38 98 55 42 12 65 42 01 53 42 01 33 42 02 27 39 03 27 38 99 77 39 02 29 Coordinate GK X (D-48) m 50 98 74 2 50 99 66 0 51 00 08 2 51 00 08 8 51 13 48 8 51 20 93 4 51 20 99 1 51 19 98 3 51 00 05 5 51 00 23 4 50 99 85 5 Diagnostic species of the association (Diagnostične vrste asociacije) VP Saxifraga cuneifolia E1 2 1 1 + 3 2 2 2 1 2 1 AT Asplenium trichomanes E1 1 1 1 1 1 1 1 1 + 1 1 VP Veronica urticifolia E1 1 . + 1 1 + + + 2 + + ML Exertotheca crispa (Neckera crispa) E0 + 2 2 1 3 2 2 2 . . . AP Valeriana tripteris E1 . + . . 1 . + . . . . Differential species of lower units (Razlikovalnice nižjih enot) FS Galeobdolon flavidum E1 . . + . . . . . + + . PA Fissidens dubius E0 . + . . 1 . . . . . + TA Phyllitis scolopendrium E1 . . . . . . . + . . . AP Astrantio carniolicae-Paederotion luteae Palustriella commutata E0 . . + . . . . . + 1 . Cystopteris fragilis E1 . . . . + + . . . . . Orthothecium rufescens E0 + . . . . . . . . . . Carex brachystachys E1 . . . . . . + . . . . Apopellia endiviifolia (Pellia endiviifolia) E0 . . + . . . . . + + . Hydrogonium croceum (Barbula crocea) E0 . . . . . . . . . . . Aster bellidiastrum E1 r . . . . . . . . . . Hymenostylium recurvirostrum E0 . . . . . 2 . . . . . Asplenium viride E1 . . . . . . + + . . . Eucladium verticillatum E0 . . . . . . . . . 2 . Paederota lutea E1 . . . . . . + . . . . Viola biflora E1 . . . . . . + . . . . Jungermannia atrovirens E0 . . . . . . . . . . . Astrantia carniolica E1 . . . . . . . . . . . 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 31 32 33 26 27 51 26 27 52 26 90 95 25 59 04 26 08 39 26 08 43 27 36 94 27 52 63 26 14 25 28 71 78 28 71 77 28 26 96 28 26 94 27 36 94 27 81 24 27 76 94 27 32 15 28 26 97 28 27 00 28 71 39 27 17 04 27 17 09 660 660 865 440 350 550 600 350 350 350 350 525 500 600 400 390 1060 515 530 800 340 360 SW W SE E S N W N N E N N W N E SW W N W N W N EE E N W SE SE W N E N W N W N 90 90 75 85 85 80 80 85 100 85 90 85 95 80 80 80 80 80 80 90 80 90 LM C h LM C h C l L L L LC h D L LC h LC h LC h LC h LC h C M C h LM L LC h LC h L M LM Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 20 20 30 20 20 30 30 30 60 20 10 20 20 30 30 40 30 30 30 30 40 40 5 15 0 30 30 30 50 50 20 30 10 30 60 50 40 60 40 30 390 10 40 30 9 9 12 11 18 29 18 19 12 18 24 25 23 18 16 22 19 20 25 28 14 19 5 5 5 10 10 10 10 15 10 10 10 10 10 10 10 10 10 10 15 15 10 20 9/ 10 /2 01 6 9/ 10 /2 01 6 5/ 17 /2 01 7 3/ 18 /2 01 5 4/ 4/ 20 16 4/ 4/ 20 16 9/ 5/ 20 18 4/ 17 /2 01 9 4/ 11 /2 01 6 6/ 18 /2 02 1 6/ 18 /2 02 1 9/ 15 /2 02 0 9/ 15 /2 02 0 9/ 5/ 20 18 6/ 26 /2 01 9 6/ 26 /2 01 9 9/ 18 /2 01 8 9/ 15 /2 02 0 9/ 15 /2 02 0 8/ 10 /2 02 1 7/ 12 /2 01 8 7/ 12 /2 01 8 D ri se lp oh D ri se lp oh K ac en po h- K ob la A vš če k A vš če k A vš če k Li šč ak -D rs el a G ru dn ic a- K lin e A vš če k Li šč ak Li šč ak Li šč ak -Z al iš ča r je va g ra pa Li šč ak Li šč ak -D rs el a Li šč ak -D rs el a Li šč ak -D rs el a Li šč ak -M le čn i r ob Li šč ak -Z al iš ča r je va g ra pa Li šč ak Li šč ak -S op ot R og -I dr ija K os i- Id ri ja 97 49 /4 97 49 /4 97 49 /4 99 48 /1 99 48 /1 99 48 /1 98 49 /1 98 48 /4 99 48 /1 98 49 /1 98 49 /1 97 49 /3 98 49 /1 98 49 /1 98 49 /1 98 49 /1 98 49 /1 98 49 /1 97 49 /3 97 49 /3 98 47 /4 98 47 /4 41 88 07 41 88 22 42 01 99 40 09 78 40 07 46 40 07 22 41 15 87 40 63 37 40 04 73 41 09 11 41 09 18 41 13 64 41 13 70 41 15 87 41 12 06 41 10 77 41 22 46 41 13 83 41 14 40 41 17 28 39 54 13 39 51 92 51 19 24 4 51 19 27 8 51 20 98 9 51 03 08 9 51 03 46 2 51 03 02 7 51 17 17 0 51 09 95 1 51 03 66 6 51 17 02 5 51 17 04 4 51 17 93 2 51 17 88 4 51 17 17 0 51 17 43 4 51 17 27 0 51 17 04 6 51 17 91 1 51 17 94 0 51 18 64 4 51 11 08 6 51 10 91 0 Pr. Fr. + + 1 1 1 1 1 1 3 2 1 2 1 1 + + + 1 1 1 2 1 33 100 1 2 1 1 . 1 2 1 1 + + 2 + 2 2 + 1 1 1 + 1 2 32 97 + 1 + . . r + + . 1 . + 1 + 2 2 1 1 1 . 1 2 27 82 . . . . + + + 2 + 1 1 + . + 2 + 3 1 . + 1 . 23 70 . . + . . 2 . . . . . + + . + . + 1 1 + + . 13 39 . . . + + + + + + 1 . 2 1 + . + + 2 1 . 1 + 19 58 . . . . + 1 1 . 2 1 1 1 2 1 . 1 . 2 1 + 1 1 18 55 . . . + + . + + . 1 1 + . + 1 + . . . . . . 11 33 . . . . . . + . . 1 1 . . + . . . . . . . . 7 22 . . . . . . 1 . . . . . . 1 . . . . . . . r 5 15 . . . . + + . . . . . . 2 . . . . . . . . . 4 12 . . . . . . . . . . + . + . . . . . + . . . 4 12 . . . . . . . . . . . . . . . . . . . . . . 3 9 . . . . . . . . . . . . 1 . . . . . . . 1 2 3 9 . . . . . . . . . . . . + . . . . . . . . . 2 6 . . . . . . . . . . . . 2 . . . . . . . . . 2 6 . . . . . . . . . . . . . . . . . . . . . . 2 6 . . . . . . . . . . + . . . . . . . . . . . 2 6 . . . . . . . . . . . . . . . . . . . . . . 1 3 . . . . . . . . . . . . . . . . . . . . . . 1 3 . . . . . . . . . . + . . . . . . . . . . . 1 3 . . . . . . . . . . . . . . . + . . . . . . 1 3 DAKSKOBLER & MARTINČIČ: VEGETATION OF MOIST ROCK CREVICES AND MOIST (SLOPE) DEBRIS IN THE LIŠČAK GORGE 54 FOLIA BIOLOGICA ET GEOLOGICA 64/1 – 2023 PsSp Physoplexido comosae-Saxifragion petraeae Phyteuma scheuchzeri subsp. columnae E1 1 1 1 + . . . . 1 + . Saxifraga petraea E1 . + . . . . . . . . r Campanula carnica E1 . . . . . . . . . . . PC Potentilletalia caulescentis Saxifraga crustata E1 . . . . . . + . . . . AT Asplenietea trichomanis Moehringia muscosa E1 + + . + . + . . . . . Asplenium ruta-muraria E1 . . . . . . 1 . + . . Polypodium vulgare E1 . . . . . + . . . . . Ceterach javorkeanum E1 . r . . . . . . . . . Sedum album E1 . . . . . . . . . . . Sedum hispanicum E1 . . . . . . . . . . . Polypodium interjectum E1 . . . . . . . . . . . TR Thlaspietea rotundifolii Hieracium bifidum E1 . + . . . + . . . . . Adenostyles glabra E1 . . . . . . . . . . . MC Montio-Cardaminetea Conocephalum conicum E0 + . . . + . . . + + . Gymnostomum aeruginosum E0 . . . 1 . . . . 1 . . Cratoneuron filicinum E0 . . . . . . . . + . . ES Elyno-Seslerietea Sesleria caerulea E1 . . . . . . + . . . . Sesleria tenuifolia subsp. kalnikensis E1 . r . . . . . . . . . BA Betulo-Alnetea Salix appendiculata E1 . . . . . . . . . . . MuA Mulgedio-Aconitetea Senecio ovatus E1 . . . . . . . . r . . Phyteuma ovatum E1 . . . . . . . + . . . Saxifraga rotundifolia E1 . . . . . . . . . . . Aconitum degenii subsp. paniculatum E1 . . . . . . . . . . . SSC Sambuco-Salicion capreae Salix caprea E1 . . . . . . . . . . . TG Trifolio-Geranietea Campanula rapunculoides E1 . . . . . . . . . . . Hypericum perforatum E1 . . . . . . . . . . . Digitalis grandiflora E1 . . . . . . . . . . . Laserpitium siler E1 . . . . . . . . . . . EP Erico-Pinetea, Festuco-Brometea Calamagrostis varia E1 . . . . . . . . . . . Carex ornithopoda E1 . . . . . . . . . . . Cirsium erisithales E1 . . . . . . . . . . . VP Vaccinio-Piceetea Oxalis acetosella E1 . . . + . . . . + . . Solidago virgaurea E1 . . . . . . . . . . . Calamagrostis arundinacea E1 . . . . + . . . . . . Hieracium murorum E1 + . . . . . . . . . . Aposeris foetida E1 . . . . . . . . . . . Homogyne sylvestris E1 . . . . . . . . . . . AF Aremonio-Fagion, Erythronio-Carpinion Cyclamen purpurascens E1 + r . + . + . . . . . Cardamine trifolia E1 1 . . . . . . . . . . Lamium orvala E1 . . . . . . . . . r . Cardamine enneaphyllos E1 r . . . . . . . . . . Anemone trifolia E1 . . . . . . . . + . . Scopolia carniolica E1 . . . . . . . . . . . Primula vulgaris E1 . . . . . . . . . . . TA Tilio-Acerion, Alnion incanae Aruncus dioicus E1 . . . . + . + . . . . Geranium robertianum E1 . . . . . . . . . . . Tephroseris pseudocrispa E1 . . r . + . . . r . . Ulmus glabra E1 . r . . . . . . . . . Acer pseudoplatanus E1 . . . . . . . . + . . Successive number of relevé (Zaporedna številka popisa) 1 2 3 4 5 6 7 8 9 10 11 DAKSKOBLER & MARTINČIČ: VEGETATION OF MOIST ROCK CREVICES AND MOIST (SLOPE) DEBRIS IN THE LIŠČAK GORGE 55FOLIA BIOLOGICA ET GEOLOGICA 64/1 – 2023 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 31 32 33 Pr. Fr. . . . . . . . . . . . . . . . . . . . . . . 6 18 . . . 1 1 1 . . + . . . . . . . . . . . . . 6 18 . . 1 . . . . . . . . . . . . . . . . . . . 1 3 . . . . . . . . . . . . . . . . . . . . . . 1 3 . . + . + + . + . . . . . . . . 1 . . . . . 9 27 . . + r . + . . . . + . . . . . 1 . . . . . 7 21 . . . . . . . + . . . . . . . . . . . . . . 2 6 . . . . . . . . . . . . . . . . . . . . . . 1 3 . . + . . . . . . . . . . . . . . . . . . . 1 3 . . . . . . . + . . . . . . . . . . . . . . 1 3 . . . . . . . + . . . . . . . . . . . . . . 1 3 . . + . . + . . . . . . . . + . . . . . . + 6 18 . . . . . . . . . . . . . . . . . . + 1 . . 2 6 . . . + . . + . . . . 1 . + . 1 . + 1 + 2 2 14 42 . . . . . . . . . . . . . . . . . . . . . . 2 6 . . . . . . . . . . . . . . . . . . . . . . 1 3 . . . . . . . . . . . . . . . . + . . 1 . . 3 9 . . . . . . . . . . . . . . . . . . . . . . 1 3 . . . . . r . . . . . . . . . . . . . . . . 1 3 . . . . . . . . . . . + + . . . . 1 + . . . 5 15 . . . . . . . . . . . . . . . . . . . . . . 1 3 . + . . . . . . . . . . . . . . . . . . . . 1 3 . . . . . . . . . . . . . . . . . . . + . . 1 3 . . . . . . . . . . . . . . . r . . . . . . 1 3 . . . . . . . . . . . . r . . . + . . + . r 4 12 . . . . . + . . . . . . . . . . . . . . . . 1 3 . . . . . . . . . . . . . . . . . . + . . . 1 3 . . . . . . . . . . . . . . . . . . . r . . 1 3 . . . . . + . . . + . . . . . . . . . + . . 3 9 . . . . . + . . . . . . . . . . . . . . . . 1 3 . . . . . . . . . . . . . . . . . . . + . . 1 3 + . . . . . . . + + + . . . + + . . 1 + . . 10 30 . . + . . . . . . . . . + . . . + . + + . . 5 15 . . . . . . . . . . . . . . . . . . . + . . 2 6 . . . . . . . . . . . . . . . . . . . . . . 1 3 . . . . . . . . . . . . . . . + . . . . . . 1 3 . . . . . . . . . . . . . . . . . . . r . . 1 3 + . + . . . . . . . . . . . . . . + . + + + 10 30 . . . . . . . . + . . . . . . + . . + . . . 4 12 . . . . . . . . . . . . . . . + . . . . . . 2 6 . . . . . . . . . . . . . . . . . . . . . . 1 3 . . . . . . . . . . . . . . . . . . . . . . 1 3 . . . . . r . . . . . . . . . . . . . . . . 1 3 . . . . . . . . . . . . . . . . . . + . . . 1 3 . . . . . . . + . . . . . . . . . . . . + + 5 15 . . . . r . . . + . . + . . . . . . . + . 1 5 15 . . . . . . . . . . + . . . . . . . . . . . 4 12 . . . . . . . . . + . . . . . + . . . . . . 3 9 . . . . . . . . . . + . . . . . + . . . . . 3 9 DAKSKOBLER & MARTINČIČ: VEGETATION OF MOIST ROCK CREVICES AND MOIST (SLOPE) DEBRIS IN THE LIŠČAK GORGE 56 FOLIA BIOLOGICA ET GEOLOGICA 64/1 – 2023 Polystichum setiferum E1 . . . . . . . . . . . Polystichum aculeatum E1 . . . . . . . . . . . Chrysosplenium alternifolium E1 . . . . . . . . . . . FS Fagetalia sylvaticae Mycelis muralis E1 . . + + . . . . . . . Galium laevigatum E1 . . . . . . . . . . . Salvia glutinosa E1 + . . . . . . . . . . Prenanthes purpurea E1 . r . . . . . . . . . Asarum europaeum subsp. caucasicum E1 . . . . . . . + . . . Cardamine pentaphyllos E1 . . . . . . . . . . . Mercurialis perennis E1 . . . . . . . . . . . Allium ursinum E1 . . . . . . . . . + . Euphorbia dulcis E1 . . . . . . . . . . . Sambucus nigra E1 . . . . . . . . . . . Poa nemoralis E1 . . . . . . . . . . . Festuca altissima E1 . . . . . . . . . . . Actaea spicata E1 . . . . . . . . . . . QP Quercetalia pubescenti-petraeae Arabis turrita E1 . . . . . . . . . . . Sesleria autumnalis E1 . . . . . . . . . . . Ostrya carpinifolia E1 . . . . . . . . . . . Fraxinus ornus E1 . . . . . . . . . . . QP Querco-Fagetea Hedera helix E1 . + 1 . . . . . . 1 + Carex digitata E1 1 + . . . . . + + . . Veratrum nigrum E1 + . r r . . . . . . . Hepatica nobilis E1 + r . . . . . . . . . Clematis vitalba E1 r . . . . . . . . . . Festuca heterophylla E1 . . . . . . . . . . . ML Mosses (Mahovi) Ctenidium molluscum E0 . . . . 1 . . + . . 1 Isothecium alopecuroides E0 . . 1 1 . . . . . + . Oxyrrhynchium hians E0 . . . . . . . . 1 r . Mnium marginatum E0 + 1 . . . . . . . . 1 Tortella tortuosa E0 . . . . + . . . . . . Thamnobryum alopecurum E0 . . . . . . . . . . . Mnium thomsonii E0 . . . . . . . . . . . Anomodon viticulosus E0 . . . . . . . . . . . Plagiomnium undulatum E0 . . . . . . . . . . . Pseudanomodon attenuatus (Anomodon attenuatus) E0 . . . . . . . 2 . . . Chionoloma tenuirostre (Oxystegus tenuirostre) E0 + . . . . + . . . . Encalypta streptocarpa E0 + . . . . . . . . . . Brachythecium rutabulum E0 . . . . . . . . . . . Plagiomnium rostratum E0 . . . . . . . . . . . Pedinophyllum interruptum E0 . . . . . . . . . . . Bryum sp. E0 . . . . . . 1 . . . . Homalothecium philippeanum E0 . . . . . . . . . . . Alleniella complanata (Neckera complanata) E0 . . . . . . . . . . . Mnium spinulosum E0 . . . . . . . . . . . Weisia sp. E0 . . . . . . . . . . . Brachythecium salebrosum E0 . . . . . . . . . . . Amblystegium serpens E0 . . . . . . . . . . . Legend - Legenda L Limestone - apnenec Br Breccia - breča Ch Chert - roženec D Dolomite - dolomit Cl Claystone - glinavec M Marlstone - laporovec Li Lithosol - kamnišče Pr. Presence (number of relevés in which the species is presented) - število popisov, v katerih se pojavlja vrsta Fr. Frequency in % - frekvenca v % Successive number of relevé (Zaporedna številka popisa) 1 2 3 4 5 6 7 8 9 10 11 DAKSKOBLER & MARTINČIČ: VEGETATION OF MOIST ROCK CREVICES AND MOIST (SLOPE) DEBRIS IN THE LIŠČAK GORGE 57FOLIA BIOLOGICA ET GEOLOGICA 64/1 – 2023 . . . . . . . . . + + . . . + . . . . . . . 3 9 . . . . . . . . . + + . . . . + . . . . . . 3 9 . . . . . . + . . . . . . + . . . . . . . . 2 6 . . . r + + + . . . . + + + + . + + 1 . . r 14 42 . . . . . + . + . . + + + . . . . + + . + + 9 27 . . . . . . . . . . . + . . r . . + 1 . . . 5 15 . + . . . . . . . + . . + . . . . . . + . . 5 15 . + . . . . . . . . + . . . . . + . + . . . 5 15 . . . . . r + + . . . . . + . . . . . . . . 4 12 . . . . . + + . . . . . . + . . . . . . . . 3 9 . . . . . . . . . . . . . . . . . . . . . . 1 3 . . . . . + . . . . . . . . . . . . . . . . 1 3 . . . . . . . . . . + . . . . . . . . . . . 1 3 . . . . . . . . . . . . . . . . + . . . . . 1 3 . . . . . . . . . . . . . . . . r . . . . . 1 3 . . . . . . . . . . . . . . . . . . + . . . 1 3 . . . . . . . . . . . + . . + + + . . . . . 4 12 . . . . . . . . . . . . . . . . . . + + . . 2 6 . . + . . . . . . . . . . . . . . . . . . . 1 3 . . . . . . . . . . . . . . . . . . . + . . 1 3 . . . . + r . + + 1 . + + . + + + . + + 1 . 17 52 + + . . . . . . . + . + + . + + . + + + . + 15 45 . . . . . r . . . + + . . . . . . . . . . . 6 18 . . . . . . . . . . . . . . . . . . . . . . 2 6 . . . . . . . . . . . . . . . . . . . . . . 1 3 . . . . . . . . . . . . . . . . . . . + . . 1 3 . . . + 1 1 1 2 . . . 2 + 1 2 + . 1 . + . 1 16 48 . 1 . 1 1 + 1 . . . 1 . . 1 . 1 . . 1 + . . 13 39 r . . . . . . . . . + 1 . . . 1 . 2 . . . 1 7 21 . . . . . . . . . . . 1 + . . . . . 1 . . . 6 18 . . . . . + . . . . . . + . . . . . . + + 1 6 18 . . . + + . . + 1 1 1 . . . . . . . . . . . 6 18 + 1 . . + + . . 1 . . . . . . . . . . . . . 5 15 . . . . 1 . 1 1 . . . . . 1 . . 1 . . . . . 5 15 . . . . . . 1 . . . 1 1 . 1 . + . . . . . . 5 15 r . . . + + . . . . . . . . . . . . . . . . 4 12 . . . . . . . . . . . . . . . . . . . . . . 2 6 . . . . . . . . . . . . . . . . . + . . . . 2 6 . . . . . . . 1 . . . + . . . . . . . . . . 2 6 . . . . . . . . . . . + . . . . . 1 . . . . 2 6 . . . . . . . . . . . + . . . . . + . . . . 2 6 . . . . . . . . . . . . . . . . . . . . . . 1 3 . . . . 1 . . . . . . . . . . . . . . . . . 1 3 . . . . . . . 1 . . . . . . . . . . . . . . 1 3 . . . . . . . . . . . + . . . . . . . . . . 1 3 . . . . . . . . . . . . + . . . . . . . . . 1 3 . . . . . . . . . . . . . . . 1 . . . . . . 1 3 . . . . . . . . . . . . . . . . . + . . . . 1 3 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 31 32 33 Pr. Fr. DAKSKOBLER & MARTINČIČ: VEGETATION OF MOIST ROCK CREVICES AND MOIST (SLOPE) DEBRIS IN THE LIŠČAK GORGE 58 FOLIA BIOLOGICA ET GEOLOGICA 64/1 – 2023 Table 7 (Preglednica 7): Calamagrostio variae-Asteretum bellidiastri Successive number of relevé (Zaporedna številka popisa) 1 2 3 4 5 6 7 8 9 Database number of relevé (Delovna številka popisa) 26 80 41 27 22 25 27 22 26 27 22 23 27 22 30 27 49 91 28 27 32 27 49 89 27 55 10 Elevation in m (Nadmorska višina v m) 260 235 235 240 320 270 560 275 275 Aspect (Lega) N E N E N E N E N N W N N W N E W Slope in degrees (Nagib v stopinjah) 80 80 80 90 70 35 15 20 80 Parent material (Matična podlaga) C o L LC h LC h LC h L D L Br Soil (Tla) Li Li Li Li Li Li Li Li Li Stoniness in % (Kamnitost v %) 100 100 100 100 100 100 100 100 100 Cover of shrub layer in % (Zastiranje grmovne plasti v %): . 5 2 . . . . 5 . Cover of herb layer in % (Zastiranje zeliščne plasti v %): 35 40 50 30 40 30 40 30 30 Cover of moss layer in % (Zastiranje mahovne plasti v %): 70 60 50 30 50 40 30 40 70 Number of species (Število vrst) 15 16 29 17 24 7 22 18 11 Relevé area (Velikost popisne ploskve) m2 5 10 10 20 10 15 10 15 15 Date of taking relevé (Datum popisa) 8/ 6/ 20 17 8/ 24 /2 01 8 8/ 24 /2 01 8 8/ 24 /2 01 8 8/ 23 /2 01 8 6/ 29 /2 01 8 6/ 22 /2 02 0 6/ 29 /2 01 8 5/ 14 /2 01 9 Locality (Nahajališče) Sl at ne -B ač a Sl at ne -B ač a Sl at ne -B ač a Sl at ne -B ač a K lo nt e- Ba ča K os ta nj ev ic a- Id ri ja V in tg ar K os ta nj ev ic a- Id ri ja M ag oz d- So ča Quadrant (Kvadrant) 98 49 /1 98 49 /1 98 49 /1 98 49 /1 98 49 /1 98 47 /4 96 50 /2 98 47 /4 97 47 /1 Coordinate GK Y (D-48) m 41 06 81 41 05 99 41 05 75 41 06 63 41 47 86 39 29 32 43 02 04 39 27 48 38 99 40 Coordinate GK X (D-48) m 51 13 04 6 51 13 10 0 51 13 11 1 51 13 04 4 51 13 82 6 51 08 90 8 51 39 50 1 51 08 95 7 51 27 01 3 Diagnostic species of association (Diagnostične vrste asociacije) AP Aster bellidiastrum E1 3 3 2 2 2 2 3 2 3 EP Calamagrostis varia E1 + + + + 1 1 1 2 + Differential species of lower units (Razlikovalnice nižjih enot) ML Brachythecium rivulare E0 2 2 2 2 . 2 . 2 . ML Hygrohypnum luridum E0 + + + + . . . 1 1 ML Didymodon spadiceus E0 1 1 1 1 . . . . 1 ML Pedinophyllum interruptum E0 + + + . . + + . . ML Plagiomnium rostratum E0 . . + . + . + . . ML Brachythecium rutabulum E0 . . . . 2 . 2 . 3 ML Tortella tortuosa E0 . . . . 1 . . . . TR Hieracium bifidum E1 . . . . + . . . . AP Astrantio carniolicae-Paederotion luteae Apopellia endiviifolia (Pellia endiviifolia) E0 . . + . . + . . 1 Hydrogonium croceum (Barbula crocea) E0 . . 1 . 2 . . . . Valeriana tripteris E1 . . . + . . + . . Selaginella helvetica E1 + . 1 r . . . . . Paederota lutea E1 . . . . . . . . + Carex brachystachys E1 . . . . . . . . . Cystopteris fragilis E1 + + + . . . . . . Tofieldia calyculata E1 . . . . . . . . . Jungermannia atrovirens E0 . . . + + + . . . Hymenostylium recurvirostrum E0 . . . + . + . . . DAKSKOBLER & MARTINČIČ: VEGETATION OF MOIST ROCK CREVICES AND MOIST (SLOPE) DEBRIS IN THE LIŠČAK GORGE 59FOLIA BIOLOGICA ET GEOLOGICA 64/1 – 2023 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 28 72 14 27 39 67 28 86 97 27 10 60 28 88 69 28 88 74 28 89 31 28 89 32 28 52 80 28 53 57 27 76 43 27 82 24 28 53 57 28 60 24 27 77 83 28 53 62 28 17 73 27 50 17 28 72 14 295 780 370 270 350 350 355 350 330 300 354 530 300 315 580 295 785 350 295 SE N W N N E N S N E N N SW 0 S W 0 N E E SS W SW N S E 90 15 70 20 90 90 80 75 90 70 80 90 95 80 30 80 95 80 90 LC lC h D L D L LC l L L D C h D C h L D A D C h L D D C h C M C h C M C h LC lC h Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 5 5 5 5 . 5 . . 5 2 . . 2 . . 5 . 5 5 30 30 40 40 20 10 20 30 30 25 30 20 25 40 40 20 40 30 30 5 30 30 10 5 5 20 20 10 30 10 30 30 50 20 30 30 30 5 13 26 28 27 21 16 12 17 16 14 9 18 14 11 9 7 9 15 13 10 15 20 50 20 10 10 10 10 10 10 10 10 10 5 10 6 10 10 6/ 18 /2 02 1 7/ 2/ 20 18 8/ 25 /2 02 1 4/ 29 /2 01 8 8/ 26 /2 02 1 8/ 26 /2 02 1 9/ 2/ 20 21 9/ 2/ 20 21 4/ 9/ 20 21 4/ 14 /2 02 1 6/ 9/ 20 19 9/ 20 /2 01 9 4/ 14 /2 02 1 5/ 20 /2 02 1 6/ 24 /2 01 9 4/ 14 /2 02 1 6/ 24 /2 02 0 7/ 1/ 20 18 6/ 18 /2 02 1 K ne ži ca -M lin ar G ač ni k N ad iž a- Ja m ni ko va vo da G or en ja T re bu ša - Tr eb uš ic a N ad iž a- Lo gj e N ad iž a- G ra de c N ad iž a- Le gr ad a N ad iž a- Le gr ad a V el ik e Lu ti V el ik e Lu ti To lm in ka -S re dn ji- ca Tr en ta V el ik e Lu ti Sr pe ni ca -S oč a V oj e- M os tn ic a Lo je -K ne ži ca Li šč ak -S op ot Li šč ak K ne ži ca -M lin ar 98 49 /1 99 49 /1 97 46 /4 99 48 /4 97 46 /4 97 46 /4 97 46 /4 97 46 /4 98 49 /1 98 49 /1 97 48 /3 96 48 /1 98 49 /1 97 47 /1 96 49 /3 98 49 /1 97 49 /3 98 49 /1 98 49 /1 41 10 53 41 21 78 37 83 66 40 96 81 37 88 35 37 88 17 37 87 15 37 87 22 41 09 93 41 10 32 40 19 96 40 07 14 41 10 32 38 63 45 41 45 58 41 10 47 41 17 36 41 08 31 41 10 53 51 16 38 0 51 01 82 9 51 22 58 6 51 00 93 5 51 22 25 6 51 22 24 2 51 22 32 2 51 22 29 7 51 16 60 6 51 16 48 0 51 22 07 3 51 35 77 0 51 16 48 0 51 28 00 1 51 29 15 2 51 16 36 6 51 18 63 2 51 16 97 2 51 16 38 0 Pr. Fr. 2 1 1 2 2 + 1 1 1 3 2 1 3 3 2 1 2 2 2 28 100 1 + + 1 1 . + + . r r + r . + 1 + . 1 24 86 . . 1 . . . . . . . . . . . . . . . . 7 25 . . 2 . . . . . . . . . . 2 . . . . . 8 29 . . . . . . . . . . . . . 1 . . . . . 6 21 . . . . . . . . . . . . . . . . . . . 5 18 . . + . . . . . . . . . . . . . . . . 4 14 . 1 . . . . . . . . . . . . . . . . . 4 14 . 1 . 1 1 1 2 2 1 3 1 1 3 1 2 2 . . . 15 54 . . + + . . + + + 1 . . 1 . r . + . . 10 36 . . + . . . . . . . . . . . . . . 1 . 5 18 . . + . . . . . . . . 2 . . . . . 1 . 5 18 . . + . . . + + . . . . . . . . . . . 5 18 . + . . . . . . . . . . . . . . . . . 4 14 . + . + . . . . . . . + . . . . . . . 4 14 . . . + . . . . 2 . . + . . . . . . . 3 11 . . . . . . . . . . . . . . . . . . . 3 11 . . . + . . 1 . . . . . . . . . . . . 2 7 . . 1 . . . . . . . . . . . . . . . . 4 14 . . . . . . . . 1 . . . . . . . 1 . . 4 14 DAKSKOBLER & MARTINČIČ: VEGETATION OF MOIST ROCK CREVICES AND MOIST (SLOPE) DEBRIS IN THE LIŠČAK GORGE 60 FOLIA BIOLOGICA ET GEOLOGICA 64/1 – 2023 Palustriella commutata E0 . . . . . . . . . Eucladium verticillatum E0 . . + . . . . . . Viola biflora E1 . . . . . . + . . Astrantia carniolica E1 . . . . . . . . . Fissidens dubius E0 . . . . . . . . . Valeriana saxatilis E1 . . . . . . . . . Pinguicula alpina E1 . . . . . . . . . Marchantia quadrata (Preissia quadrata) E0 . . . . . . . . . PcSp Physoplexido comosae-Saxifragion petraeae Hieracium pospichalii E1 + . . . . . . . . Campanula cespitosa E1 . . . . . . . . . Campanula carnica E1 . . . . . . . . . Hieracium porrifolium E1 . . . . . . . . . Athamanta turbith E1 . . . . . . . . . Micromeria thymifolia E1 . . . . . . . . . Phyteuma scheuchzeri subsp. columnae E1 . . . . . . . . . Campanula zoysii E1 . . . . . . . . . PC Potentilletalia caulescentis Saxifraga hostii E1 . . . . . . . . . Festuca stenantha E1 . . . . . . . . . Potentilla caulescens E1 . . . . . . . . . AT Asplenietea trichomanis Asplenium ruta-muraria E1 . . . . . . . . + Asplenium trichomanes E1 . . . . . . . . . MC Montio-Cardaminetea Conocephalum conicum E0 3 1 2 + + . . . . Cratoneuron filicinum E0 . . 1 . . . . . . Oxyrrhynchium hians E0 . + . . . . . . . Calliergonella cuspidata E0 . . + . . . . . . Cololejeunea calcarea E0 . . . + . . . . . Fontinalis antipyretica var. antipyretica E0 . . . . . . . 1 . Palustriella decipiens E0 . . . . . . . . . CD Caricetalia davallianae Carex flava agg. E1 . . . . . . . . . Carex lepidocarpa E1 . . . . . . . . . Campylium stellatum E1 . . . . . . . . . TR Thlaspietea rotundifolii Trisetum argenteum E1 . . . . . . . . 1 Petasites paradoxus E1 . . . . . . . . . Achnatherum calamagrostis E1 . . . . . . . + . Adenostyles glabra E1 . . . . . . + . . Hieracium glaucum E1 . . . . . . . . + Poa compressa E1 . . . . . . . . . Gymnocarpium robertianum E1 . . . . . . . . . Peucedanum verticillare E1 . . . . . . . . . Hieracium piloselloides E1 . . . . . . . . . Soldanella minima E1 . . . . . . . . . ES Elyno-Seslerietea Sesleria caerulea E1 . . . . . . . . . Carex ornithopodoides E1 . . . . . . . . . Carex ferruginea E1 . . . . . . . . . Carex mucronata E1 . . . . . . . . . Erigeron glabratus E1 . . . . . . . . . BA Betulo-Alnetea Salix appendiculata E2a . + . . . . . . . MuA Mulgedio-Aconitetea Chaerophyllum hirsutum E1 . . . . r . + . . Petasites hybridus E1 . . . . . . . + . EA Epilobietea angustifolii Eupatorium cannabinum E1 . r + . . . . + . Tussilago farfara E1 . . . . + . . 1 . Cardamine hirsuta E1 . . . . r . . . . Carex oederi E1 . . . . . . . . . Successive number of relevé (Zaporedna številka popisa) 1 2 3 4 5 6 7 8 9 DAKSKOBLER & MARTINČIČ: VEGETATION OF MOIST ROCK CREVICES AND MOIST (SLOPE) DEBRIS IN THE LIŠČAK GORGE 61FOLIA BIOLOGICA ET GEOLOGICA 64/1 – 2023 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 Pr. Fr. . . . . . . . . . . . . . . . . 1 1 . 2 7 . . . . . . . . . . . . . . . . . . . 1 4 . . . . . . . . . . . . . . . . . . . 1 4 . + . . . . . . . . . . . . . . . . . 1 4 . . + . . . . . . . . . . . . . . . . 1 4 . . . 2 . . . . . . . . . . . . . . . 1 4 . . . . . . . . 1 . . . . . . . . . . 1 4 . . . . . . . . . . . + . . . . . . . 1 4 . 1 . + . . . . + 1 . . 1 . . . . . . 6 21 . . . . + + + + . . 2 + . . . . . . . 6 21 . . + . . . . . . + . . + . r . . . . 4 14 . . . . . + . + . . + + . . . . . . . 4 14 . . . . + . + + . . . . . . . . . . . 3 11 . + . . + . . . . . . . . . . . . . . 2 7 . . . + . . . . . . . . . . . . . . . 1 4 . . . . . . . . . . . + . . . . . . . 1 4 . . . . + . . r . . . . . . . . . . . 2 7 . . . . . . . . . . . + . . 1 . . . . 2 7 . . . . . . . . . . . + . . . . . . . 1 4 . . . . . + . + + . . . . . . . . . . 4 14 . . . . + + . . + . . . . . . . . . . 3 11 . . 1 . . . . . . . . . . . . . . 1 . 7 25 . . + . . . . . . . . . . . . . . . . 2 7 . . . . . . . . . . . . . . . . . . . 1 4 . . . . . . . . . . . . . . . . . . . 1 4 . . . . . . . . . . . . . . . . . . . 1 4 . . . . . . . . . . . . . . . . . . . 1 4 . . . . . . . . . . . . . 1 . . . . . 1 4 . . . + . . . . . . . . . . . . . . . 1 4 . + . . . . . . . . . . . . . . . . . 1 4 . . . . . . . . . . . . . + . . . . . 1 4 . . . . . . . . . 2 1 1 2 1 . . . . . 6 21 + . + + 1 . . . . . . . . . . + . . + 6 21 . . + . 1 . . . . . + . . . . . . . . 4 14 . . . . . . . . . . . . . . . . 1 1 . 3 11 . . + . . . . . . . . . . . . . . . . 2 7 . + . . . . . . . . . + . . . . . . . 2 7 . . . . + + . . . . . . . . . . . + . 3 11 . . . . + . . + . . . . . . . . . . . 2 7 . . . 1 . . . . . . . . . . . . . . . 1 4 . . . . . . . . . . . . . . + . . . . 1 4 . . 2 2 2 1 + 1 1 . . . . . . . . . . 7 25 . . . . . . . . . + . + + . . . . . . 3 11 . . . + . . . . . . . . . . . . . . . 1 4 . . . + . . . . . . . . . . . . . . . 1 4 . . . . . . . . . . . + . . . . . . . 1 4 1 + 1 + . + . . + . . . . . . + . + 1 10 36 . . + . . + . . . . . . . . . . . . . 4 14 . 1 . . . . . . . . . . . . . . . . . 2 7 . . . . . . . . . . . . . . . . r . . 4 14 . + . . . . . . . . . . . . . . . . . 3 11 . . . . . . . . . . . . . . . . . . . 1 4 . . . + . . . . . . . . . . . . . . . 1 4 DAKSKOBLER & MARTINČIČ: VEGETATION OF MOIST ROCK CREVICES AND MOIST (SLOPE) DEBRIS IN THE LIŠČAK GORGE 62 FOLIA BIOLOGICA ET GEOLOGICA 64/1 – 2023 Fragaria vesca E1 . . . . . . . . . TG Trifolio-Geranietea Campanula rapunculoides E1 . . . . . . + . . Hypericum perforatum E1 . . . . . . . . . PaT Poo alpinae-Trisetetalia Poa alpina E1 . . . . . . . . . Mo Molinion Taraxacum sect. Palustria E1 . . . . . . . . . Filipendula ulmaria E1 . . . . . . + . . Crepis paludosa E1 . . . . . . . . . MA Molinio-Arrhenatheretea Deschampsia cespitosa E1 + . . . r . r . . Prunella vulgaris E1 . . . . . + . 1 . Leontodon hispidus E1 + . r . . . . . . Agrostis stolonifera E1 . + . . . . . . . Angelica sylvestris E1 . . r . . . . . . Galium mollugo E1 . . . . . . + . . Taraxacum sect. Ruderalia E1 . . . . . . . + . Dactylis glomerata E1 . . . . . . . . . FB Festuco-Brometea Buphthalmum salicifolium E1 . . . . . . . . . Thymus praecox E1 . . . . . . . . . Plantago media E1 . . . . . . . . . Brachypodium rupestre E1 . . . . . . . . . Koeleria pyramidata E1 . . . . . . . . . Linum catharticum E1 . . . . . . . . . Euphorbia cyparissias E1 . . . . . . . . . Taraxacum sect. Erythrosperma E1 . . . . . . . . . Centaurea dichroantha E1 . . . . . . . . . SM Stellarietea mediae Galinsoga ciliata E1 . . . . r . . . . EP Erico-Pinetea Carex ornithopoda E1 . 1 + . . . . . . Erica carnea E1 . . . . . . . . . Leontodon incanus E1 . . . . . . . . . Rubus saxatilis E1 . . . . . . + . . Euphrasia cuspidata E1 . . . . . . . . . Molinia arundinacea E1 . . . . . . . . . VP Vaccinio-Piceetea Veronica urticifolia E1 1 1 1 1 + 1 1 . . Oxalis acetosella E1 . . r . + . . . . Clematis alpina E1 . . . . . . . . . Calamagrostis arundinacea E1 . . . . . . . . . TA Tilio-Acerion Phyllitis scolopendrium E1 . . + . . . + . . Aruncus dioicus E1 . . . . . 1 . . . Geranium robertianum E1 . + . + + . . . . Polystichum x wirtgenii E1 . . . . . . . . . AI Alnion incanae Rubus caesius E1 . + . . . . . r . Alnus glutinosa E2a . . r . . . . + . Salix eleagnos E2a . . r . . . . . . Knautia drymeia subsp. intermedia E1 . . . . + . . . . Frangula alnus E2a . . . . . . . . . Viburnum opulus E2a . . . . . . . . . FS Fagetalia sylvaticae Galeobdolon flavidum E1 + . . + r . . . . Galium laevigatum E1 . . . . + . . . . Mycelis muralis E1 . . r . . . + . . Salvia glutinosa E1 . . + . . . . . . Brachypodium sylvaticum E1 . . + . r . . . . Tilia cordata E2a . + . . . . . . . Viola reichenbachiana E1 . r . . . . . . . Successive number of relevé (Zaporedna številka popisa) 1 2 3 4 5 6 7 8 9 DAKSKOBLER & MARTINČIČ: VEGETATION OF MOIST ROCK CREVICES AND MOIST (SLOPE) DEBRIS IN THE LIŠČAK GORGE 63FOLIA BIOLOGICA ET GEOLOGICA 64/1 – 2023 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 Pr. Fr. . . . . . . . + . . . . . . . . . . . 1 4 . . + . + . . + . . . . . . . . . . . 4 14 . . . . + . . . . . . . . . . . . . . 1 4 . . . . . . . . . . . . . . + . . . . 1 4 . . . . . . . . . r . . r . . . . . . 2 7 . . . . . . . . . . . . . . . . . . . 1 4 . . . . . . . . . . . . . . . . 1 . . 1 4 . + . . . . . . . . . . . . . . . . . 4 14 . . . . . . . . . . . . . . 1 . . . . 3 11 . . . . . . . . . . . . . . . . . . . 2 7 . . . . . . . . . . . . . . . . . . . 1 4 . . . . . . . . . . . . . . . . . . . 1 4 . . . . . . . . . . . . . . . . . . . 1 4 . . . . . . . . . . . . . . . . . . . 1 4 . + . . . . . . . . . . . . . . . . . 1 4 + . + . + + + . + . . . . . . . . . + 7 25 . 1 . . . . . . . . . . . . . . . . . 1 4 . + . . . . . . . . . . . . . . . . . 1 4 . + . . . . . . . . . . . . . . . . . 1 4 . + . . . . . . . . . . . . . . . . . 1 4 . . . + . . . . . . . . . . . . . . . 1 4 . . . . . . . . . . r . . . . . . . . 1 4 . . . . . . . . . . . . . . . + . . . 1 4 . . . . . . . . . . + . . . . . . . . 1 4 . . . . . . . . . . . . . . . . . . . 1 4 . + . + . . . . . 1 . . 1 . . . . . . 6 21 + . . . . . . . + . . . . . . . . . + 3 11 . . . + . . . . + . . . . . . . . . . 2 7 . . . . . . . . . . . . . . . . . . . 1 4 . + . . . . . . . . . . . . . . . . . 1 4 . . . . . . . . + . . . . . . . . . . 1 4 + + 1 . . . . + . . . . . . . . . 1 + 13 46 . . . . . . . . . . . . . . . . . . . 2 7 . . . . . r . . . . . . . . . + . . . 2 7 . . . . . . . . . . . . . . . . . + . 1 4 + . . . + . . . . . . . . . . . . . + 5 18 + . . . . . + . . . . . . . . . . + + 5 18 . . . . . . . . . . . . . . . . . . . 3 11 . . . . . . . . . . . . . . . . . + . 1 4 . . . . . . . + . . . . . . . . . . . 3 11 . . . . . . . . . . . . . . . . . . . 2 7 . . . . . . . . . . . . . . . . . . . 1 4 . . . . . . . . . . . . . . . . . . . 1 4 . . . + . . . . . . . . . . . . . . . 1 4 . . . + . . . . . . . . . . . . . . . 1 4 . . . . . . . . . . . . . . . . . + . 4 14 + . . . + . . . . . . . . . . . . . + 4 14 . . . . . . . . . . . . . . . . . 1 . 3 11 . . + . + . . . . . . . . . . . . . . 3 11 . . . . . . . . . . . . . . . . . . . 2 7 . . . . . . . . . . . . . . . . . . . 1 4 . . . . . . . . . . . . . . . . . . . 1 4 DAKSKOBLER & MARTINČIČ: VEGETATION OF MOIST ROCK CREVICES AND MOIST (SLOPE) DEBRIS IN THE LIŠČAK GORGE 64 FOLIA BIOLOGICA ET GEOLOGICA 64/1 – 2023 Epilobium montanum E1 . . . . r . . . . Ranunculus lanuginosus E1 . . . . . . + . . Fagus sylvatica E2a . . . . . . . . . Campanula trachelium E1 . . . . . . . . . QP Quercetalia pubescenti-petraeae Ostrya carpinifolia E2a . . . . . . . . . Fraxinus ornus E1 . . . . . . . . . Clematis vitalba E1 . . . . . . . r . Carex flacca E1 . . . . . . . . . QF Querco-Fagetea Hedera helix E1 . . . + . . . r . Hieracium racemosum E1 . . r . . . . . . Hieracium umbellatum E1 . . r . . . . . . Viola riviniana E1 . . . . . . . + . Carex digitata E1 . . . . . . . . . ML Mosses (Mahovi) Ctenidium molluscum E0 1 . 1 . 1 . 1 1 . Schistidium apocarpum E0 . . . . 1 1 . . Cirriphyllum crassinervium E0 . . . 1 . . 1 . . Encalypta streptocarpa E0 . . . . . . . . . Dichodontium pellucidum E0 . . + . . . . 1 . Sciuro-hypnum starkei (Brachythecium starkei) E0 . 1 . . . . . . . Trichostomum crispulum E0 . + . . . . . . . Amblystegium serpens E0 . . + . . . . . . Jungermannia sp. E0 . . . + . . . . . Mesoptychia collaris (Leiocolea collaris) E0 . . . . + . . . . Thamnobryum alopecurum E0 . . . . . . + . . Campylopus sp. E0 . . . . . . . . . Hypnum cupressiforme E0 . . . . . . . . . Didymodon insulanus E0 . . . . . . . . . Dichodontium flavescens E0 . . . . . . . . . Lophozia sp. E0 . . . . . . . . . Exertotheca crispa (Neckera crispa) E0 . . . . . . . . . Plagiumnium elatum E0 . . . . . . . . . Hylocomium splendens E0 . . . . . . . . . Legend - Legenda L Limestone - apnenec Br Brecia - breča Ch Chert - roženec Co Conglomerate - konglomerat D Dolomite - dolomit Cl Claystone - glinavec M Marlstone - laporovec Li Lithosol - kamnišče Pr. Presence (number of relevés in which the species is presented) - število popisov, v katerih se pojavlja vrsta Fr. Frequency in % - frekvenca v % Successive number of relevé (Zaporedna številka popisa) 1 2 3 4 5 6 7 8 9 DAKSKOBLER & MARTINČIČ: VEGETATION OF MOIST ROCK CREVICES AND MOIST (SLOPE) DEBRIS IN THE LIŠČAK GORGE 65FOLIA BIOLOGICA ET GEOLOGICA 64/1 – 2023 Epilobium montanum E1 . . . . r . . . . Ranunculus lanuginosus E1 . . . . . . + . . Fagus sylvatica E2a . . . . . . . . . Campanula trachelium E1 . . . . . . . . . QP Quercetalia pubescenti-petraeae Ostrya carpinifolia E2a . . . . . . . . . Fraxinus ornus E1 . . . . . . . . . Clematis vitalba E1 . . . . . . . r . Carex flacca E1 . . . . . . . . . QF Querco-Fagetea Hedera helix E1 . . . + . . . r . Hieracium racemosum E1 . . r . . . . . . Hieracium umbellatum E1 . . r . . . . . . Viola riviniana E1 . . . . . . . + . Carex digitata E1 . . . . . . . . . ML Mosses (Mahovi) Ctenidium molluscum E0 1 . 1 . 1 . 1 1 . Schistidium apocarpum E0 . . . . 1 1 . . Cirriphyllum crassinervium E0 . . . 1 . . 1 . . Encalypta streptocarpa E0 . . . . . . . . . Dichodontium pellucidum E0 . . + . . . . 1 . Sciuro-hypnum starkei (Brachythecium starkei) E0 . 1 . . . . . . . Trichostomum crispulum E0 . + . . . . . . . Amblystegium serpens E0 . . + . . . . . . Jungermannia sp. E0 . . . + . . . . . Mesoptychia collaris (Leiocolea collaris) E0 . . . . + . . . . Thamnobryum alopecurum E0 . . . . . . + . . Campylopus sp. E0 . . . . . . . . . Hypnum cupressiforme E0 . . . . . . . . . Didymodon insulanus E0 . . . . . . . . . Dichodontium flavescens E0 . . . . . . . . . Lophozia sp. E0 . . . . . . . . . Exertotheca crispa (Neckera crispa) E0 . . . . . . . . . Plagiumnium elatum E0 . . . . . . . . . Hylocomium splendens E0 . . . . . . . . . Legend - Legenda L Limestone - apnenec Br Brecia - breča Ch Chert - roženec Co Conglomerate - konglomerat D Dolomite - dolomit Cl Claystone - glinavec M Marlstone - laporovec Li Lithosol - kamnišče Pr. Presence (number of relevés in which the species is presented) - število popisov, v katerih se pojavlja vrsta Fr. Frequency in % - frekvenca v % 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 Pr. Fr. . . . . . . . . . . . . . . . . . . . 1 4 . . . . . . . . . . . . . . . . . . . 1 4 . . . + . . . . . . . . . . . . . . . 1 4 . . + . . . . . . . . . . . . . . . . 1 4 + . . + . + . . . r . . r . . . . . + 4 14 . . + . + + . . . . . . . . . . . . . 3 11 . . . . . + . . . . . . . . . . . . . 2 7 . . . 1 . . . . . . . . . . . . . . . 1 4 + . . . . . . . . . . . . . . . . . + 4 14 . . . . . . . . . . . . . . . . . . . 1 4 . . . . . . . . . . . . . . . . . . . 1 4 . . . . . . . . . . . . . . . . . . . 1 4 . . . . . . . . . . . . . . . . . + . 1 4 + 1 . 1 . 1 3 2 . 1 . + 1 2 . . . . + 16 57 . 1 . . . 1 . . . . . . . . . . . . . 4 18 . . . . . . . . . . . . . 2 . . . . . 3 11 . . . . . . . . . 1 . + 1 . . . . . . 3 11 . . . . . . . . . . . . . . . . . . . 2 7 . . . . . . . . . . . . . . . . . . . 1 4 . . . . . . . . . . . . . . . . . . . 1 4 . . . . . . . . . . . . . . . . . . . 1 4 . . . . . . . . . . . . . . . . . . . 1 4 . . . . . . . . . . . . . . . . . . . 1 4 . . . . . . . . . . . . . . . . . . . 1 4 . + . . . . . . . . . . . . . . . . . 1 4 . . 1 . . . . . . . . . . . . . . . . 1 4 . . + . . . . . . . . . . . . . . . . 1 4 . . + . . . . . . . . . . . . . . . . 1 4 . . + . . . . . . . . . . . . . . . . 1 4 . . . . + . . . . . . . . . . . . . . 1 4 . . . . . . . . . . . . . + . . . . . 1 4 . . . . . . . . . . . . . + . . . . . 1 4 DAKSKOBLER & MARTINČIČ: VEGETATION OF MOIST ROCK CREVICES AND MOIST (SLOPE) DEBRIS IN THE LIŠČAK GORGE 66 FOLIA BIOLOGICA ET GEOLOGICA 64/1 – 2023 Table 8 (Preglednica 8): Veronico urticifoliae-Violetum biflorae Successive number of relevé (Zaporedna številka popisa) 1 2 3 4 5 6 7 8 9 Database number of relevé (Delovna številka popisa) 21 47 87 26 16 49 25 88 18 22 70 07 26 10 72 28 09 06 25 49 12 26 09 39 25 88 19 Elevation in m (Nadmorska višina v m) 450 550 665 210 590 465 780 550 665 Aspect (Lega) W N W W N E N N W N W N W N W Slope in degrees (Nagib v stopinjah) 90 10 80 80 80 70 95 95 95 Parent material (Matična podlaga) LM D eb D C h D D D D L D D C h Soil (Tla) Li Li Li Li Li Li Li Li Li Stoniness in % (Kamnitost v %) 100 20 100 100 100 10 100 100 100 Cover of shrub layer in % (Zastiranje grmovne plasti v %): . . . . . . . . . Cover of herb layer in % (Zastiranje zeliščne plasti v %): 30 80 60 30 50 60 30 30 30 Cover of moss layer in % (Zastiranje mahovne plasti v %): 40 50 60 50 80 80 10 10 40 Number of species (Število vrst) 9 17 15 13 14 9 16 14 12 Relevé area (Velikost popisne ploskve) m2 10 10 5 5 10 2 10 10 5 Date of taking relevé (Datum popisa) 5/ 9/ 20 01 5/ 28 /2 01 6 10 /1 2/ 20 15 5/ 14 /2 00 9 4/ 20 /2 01 6 5/ 4/ 20 20 5/ 5/ 20 14 4/ 15 /2 01 6 10 /1 2/ 20 15 Locality (Nahajališče) M at ev ž- Id ri ja Z ak oj šk a gr ap a Z ak oj šk a gr ap a St op ni k- G aš pe r Id rš ek K oz je k- V rš e M ož ni ca Ž ir ov ni ca - R as ko ve c Z ak oj šk a gr ap a Quadrant (Kvadrant) 98 47 /2 98 49 /2 98 49 /2 99 49 /1 99 50 /4 99 48 /2 96 47 /1 99 50 /4 98 49 /2 Coordinate GK Y (D-48) m 39 65 18 41 82 09 41 89 79 41 45 34 42 92 93 40 85 56 38 97 96 43 04 28 41 88 82 Coordinate GK X (D-48) m 51 14 00 8 51 14 29 4 51 14 42 4 51 05 56 1 50 96 50 6 51 05 33 5 51 38 67 4 50 96 26 4 51 14 40 7 Diagnostic species of the association (Diagnostične vrste asociacije) AP Viola biflora E1 1 3 2 3 3 4 3 2 2 MC Conocephalum conicum E0 3 3 4 1 2 4 + 1 2 VP Veronica urticifolia E1 1 + + . 1 . r + . AP Astrantio carniolicae-Paederotion luteae Orthothecium rufescens E0 . . . 2 1 + 1 1 2 Asplenium viride E1 . . . . + + 1 + + Hymenostylium recurvirostrum E0 . . 1 . . . . . 1 Palustriella commutata E0 . 1 . . . . . . . Cystopteris fragilis E1 . . . . . . + + . Carex brachystachys E1 . . . . . . + + + Apopellia endiviifolia (Pellia endiviifolia) E0 . . . . . . . . . Aster bellidiastrum E1 . . . 1 + 1 . . . Astrantia carniolica E1 . . . . . . + . . Valeriana tripteris E1 . . + . . . . . . Paederota lutea E1 . . . 1 . + + . . Fissidens dubius E0 . . . . . . . . . Jungermannia atrovirens E0 . . . . . . . . . Hydrogonium croceum (Barbula crocea) E0 . . . . . . . . . Pinguicula alpina E1 . . . . . . . 1 . Valeriana saxatilis E1 . . . . . . . . . Eucladium verticillatum E0 . . . . . . . . . Primula carniolica E1 . . . . . . . . . PcSp Physoplexido comosae-Saxifragion petraeae Campanula carnica E1 r . . . . . r . . Phyteuma scheuchzeri subsp. columnae E1 . . . + + . . . . DAKSKOBLER & MARTINČIČ: VEGETATION OF MOIST ROCK CREVICES AND MOIST (SLOPE) DEBRIS IN THE LIŠČAK GORGE 67FOLIA BIOLOGICA ET GEOLOGICA 64/1 – 2023 Table 8 (Preglednica 8): Veronico urticifoliae-Violetum biflorae Successive number of relevé (Zaporedna številka popisa) 1 2 3 4 5 6 7 8 9 Database number of relevé (Delovna številka popisa) 21 47 87 26 16 49 25 88 18 22 70 07 26 10 72 28 09 06 25 49 12 26 09 39 25 88 19 Elevation in m (Nadmorska višina v m) 450 550 665 210 590 465 780 550 665 Aspect (Lega) W N W W N E N N W N W N W N W Slope in degrees (Nagib v stopinjah) 90 10 80 80 80 70 95 95 95 Parent material (Matična podlaga) LM D eb D C h D D D D L D D C h Soil (Tla) Li Li Li Li Li Li Li Li Li Stoniness in % (Kamnitost v %) 100 20 100 100 100 10 100 100 100 Cover of shrub layer in % (Zastiranje grmovne plasti v %): . . . . . . . . . Cover of herb layer in % (Zastiranje zeliščne plasti v %): 30 80 60 30 50 60 30 30 30 Cover of moss layer in % (Zastiranje mahovne plasti v %): 40 50 60 50 80 80 10 10 40 Number of species (Število vrst) 9 17 15 13 14 9 16 14 12 Relevé area (Velikost popisne ploskve) m2 10 10 5 5 10 2 10 10 5 Date of taking relevé (Datum popisa) 5/ 9/ 20 01 5/ 28 /2 01 6 10 /1 2/ 20 15 5/ 14 /2 00 9 4/ 20 /2 01 6 5/ 4/ 20 20 5/ 5/ 20 14 4/ 15 /2 01 6 10 /1 2/ 20 15 Locality (Nahajališče) M at ev ž- Id ri ja Z ak oj šk a gr ap a Z ak oj šk a gr ap a St op ni k- G aš pe r Id rš ek K oz je k- V rš e M ož ni ca Ž ir ov ni ca - R as ko ve c Z ak oj šk a gr ap a Quadrant (Kvadrant) 98 47 /2 98 49 /2 98 49 /2 99 49 /1 99 50 /4 99 48 /2 96 47 /1 99 50 /4 98 49 /2 Coordinate GK Y (D-48) m 39 65 18 41 82 09 41 89 79 41 45 34 42 92 93 40 85 56 38 97 96 43 04 28 41 88 82 Coordinate GK X (D-48) m 51 14 00 8 51 14 29 4 51 14 42 4 51 05 56 1 50 96 50 6 51 05 33 5 51 38 67 4 50 96 26 4 51 14 40 7 Diagnostic species of the association (Diagnostične vrste asociacije) AP Viola biflora E1 1 3 2 3 3 4 3 2 2 MC Conocephalum conicum E0 3 3 4 1 2 4 + 1 2 VP Veronica urticifolia E1 1 + + . 1 . r + . AP Astrantio carniolicae-Paederotion luteae Orthothecium rufescens E0 . . . 2 1 + 1 1 2 Asplenium viride E1 . . . . + + 1 + + Hymenostylium recurvirostrum E0 . . 1 . . . . . 1 Palustriella commutata E0 . 1 . . . . . . . Cystopteris fragilis E1 . . . . . . + + . Carex brachystachys E1 . . . . . . + + + Apopellia endiviifolia (Pellia endiviifolia) E0 . . . . . . . . . Aster bellidiastrum E1 . . . 1 + 1 . . . Astrantia carniolica E1 . . . . . . + . . Valeriana tripteris E1 . . + . . . . . . Paederota lutea E1 . . . 1 . + + . . Fissidens dubius E0 . . . . . . . . . Jungermannia atrovirens E0 . . . . . . . . . Hydrogonium croceum (Barbula crocea) E0 . . . . . . . . . Pinguicula alpina E1 . . . . . . . 1 . Valeriana saxatilis E1 . . . . . . . . . Eucladium verticillatum E0 . . . . . . . . . Primula carniolica E1 . . . . . . . . . PcSp Physoplexido comosae-Saxifragion petraeae Campanula carnica E1 r . . . . . r . . Phyteuma scheuchzeri subsp. columnae E1 . . . + + . . . . 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 28 16 03 28 27 29 28 16 07 28 16 08 28 16 09 28 16 13 24 60 74 26 21 34 27 33 32 28 11 16 28 08 07 28 18 22 28 74 30 24 72 06 25 88 14 28 18 20 560 575 588 545 540 580 492 776 460 450 650 1000 840 620 625 1000 N N W N W N W N N W N W N W W E N N E N N E SW W N N SW 70 90 95 90 90 95 90 10 100 95 80 90 85 80 85 60 D D D D D D LC h D eb LM C h L D eb LM LM D C h D C h D eb Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li 100 100 100 100 100 100 90 30 100 100 100 100 100 100 100 70 . . . . . . . . . . . . . 20 . . 40 30 30 30 50 30 40 60 30 60 30 30 30 40 40 50 30 20 30 20 20 20 30 60 40 30 50 20 20 20 50 30 20 20 14 24 25 23 24 13 17 18 19 9 13 35 17 20 20 5 10 10 10 10 5 2 10 10 10 10 10 50 5 10 5/ 21 /2 02 0 6/ 22 /2 02 0 5/ 21 /2 02 0 5/ 21 /2 02 0 5/ 21 /2 02 0 5/ 21 /2 02 0 4/ 28 /2 01 2 7/ 9/ 20 16 8/ 8/ 20 18 5/ 23 /2 02 0 5/ 7/ 20 20 6/ 29 /2 02 0 8/ 15 /2 02 1 4/ 28 /2 00 0 10 /1 2/ 20 15 6/ 29 /2 02 0 V in tg ar V in tg ar V in tg ar V in tg ar V in tg ar V in tg ar D ri se lp oh H ob a- Ba ča Pr od ar je va g ra pa Pr od ar je va g ra pa K oz je k D re žn iš ke R av ne - C ur k Šp ič no ko va g ra pa sl ap Z ak oj šk a gr ap a Z ak oj šk a gr ap a D re žn iš ke R av ne - C ur k 95 50 /4 96 50 /2 96 50 /2 96 50 /2 96 50 /2 96 50 /2 97 49 /4 97 49 /4 98 49 /2 98 49 /2 99 48 /2 97 47 /2 98 49 /2 98 49 /2 98 49 /2 97 47 /2 43 09 44 43 01 51 43 08 40 43 07 75 43 06 37 43 02 76 41 87 23 42 24 78 41 87 85 41 87 05 40 85 13 39 52 09 42 09 37 41 87 75 41 87 75 39 52 17 51 39 95 1 51 39 39 8 51 39 89 7 51 39 82 0 51 39 77 3 51 39 60 1 51 18 20 2 51 20 50 6 51 17 44 2 51 17 59 9 51 04 98 1 51 26 20 0 51 17 11 8 51 14 40 0 51 14 41 2 51 26 19 3 Pr. Fr. 3 2 2 2 3 3 2 3 3 3 3 2 2 2 2 2 25 100 2 2 1 1 1 + + 3 1 1 1 + 1 1 2 1 25 100 + 1 1 1 1 1 + + 1 + . + 1 1 + . 20 80 1 1 2 1 2 1 2 . 1 2 1 . . 1 1 + 19 76 + 1 + + 1 + . . r + r . + + . . 16 64 + 1 . 1 1 + 1 . . 1 . 2 + . . + 12 48 . . . . 1 . 1 2 3 3 3 1 1 1 1 1 12 48 . + 1 1 1 1 . + . . + + + . . . 11 44 . . . + 1 . 1 . + + . . . + + . 10 40 . . . . + . 2 1 . + . + + . + 1 8 32 . . 2 2 1 + . . . . + . . . . . 8 32 . . 1 2 1 + . . . . r . . . . . 6 24 . + . . . + . . . . r . . 1 2 . 6 24 . . . . . . + . . r + . . . . . 6 24 . + . . . 1 . . + . . . . + + . 5 20 . . . + + + . . . + . . . . . . 4 16 . . . + + . + . . . . . . . . . 3 12 . . . . . . . . . . . . . . . . 1 4 . . . 1 . . . . . . . . . . . . 1 4 . . . . . . 1 . . . . . . . . . 1 4 . . . . . . . . . . + . . . . . 1 4 . . . . . . + . . . . . . . . . 3 12 . . . . . . . . . . r . . . . . 3 12 DAKSKOBLER & MARTINČIČ: VEGETATION OF MOIST ROCK CREVICES AND MOIST (SLOPE) DEBRIS IN THE LIŠČAK GORGE 68 FOLIA BIOLOGICA ET GEOLOGICA 64/1 – 2023 PC Potentilletalia caulescentis Primula auricula E1 . . . . . . . . . Potentilla caulescens E1 . . . . . . . . . AT Asplenietea trichomanis Asplenium trichomanes E1 + + . . . . 1 + . Asplenium ruta-muraria E1 . . . . . . . . . Kernera saxatilis E1 . . . . . . . . . Moehringia muscosa E1 . . . . . . . . . TR Thlaspietea trotundifolii Adenostyles glabra E1 . + 2 . . . . . 1 Gymnocarpium robertianum E1 . 1 . . . . . . 1 Hieracium bifidum E1 . . . . . . r . . Trisetum argenteum E1 . . . . . . . . . MC Montio-Cardaminetea Oxyrrhynchium hians E0 . . . . . . . . . Oxyrrhynchium schleicheri E0 . . . . . . . . . Flexitrichum flexicaule (Ditrichum flexicaule) E0 . . . . . . . . . Rhynchostegium riparioides (Platyhypnidium riparioides) E0 . . . 1 . . . . . Fontinalis antipyretica subsp. antipyretica E0 . . . + . . . . . Cololejeunea calcarea E0 . . . . . . . . . CD Caricetalia davallianae Campylium stellatum E0 . . . . 1 . . . . ES Elyno-Seslerietea Sesleria caerulea E1 . . . . . . . . + Carex ferruginea E1 . . . . . . . . . Mo Molinion, Molino-Arrhenatheretea Angelica sylvestris E1 . . . . . . . . . Caltha palustris E1 . . + r . . . . . Crepis paludosa E1 . . 1 . . . . . . TG Trifolio-Geranietea Campanula rapunculoides E1 . . . . . . . . . EA Epilobietea angustifolii Rubus idaeus E1 . + . . . . . . . Solanum dulcamara E1 . . . . . . . . . MuA Mulgedio-Aconitetea Chaerophyllum hirsutum E1 . . 2 + . . . . . Saxifraga rotundifolia E1 . . + . r . . . . Senecio ovatus E1 . . . . . . . . . Thalictrum aquilegiifolium E1 . 1 . . . . . . . Phyteuma ovatum E1 . . . . . . . . . EP Erico-Pinetea Calamagrostis varia E1 . . + . . . . + + Aquilegia nigricans E1 . . . . . . . . . Carex ornithopoda E1 r . . . . . . . . Rhodothamnus chamaecistus E1 . . . . . . + . . Carex alba E1 . . . . . . . . . VP Vaccinio-Piceetea Oxalis acetosella E1 1 2 . . + . . . . Saxifraga cuneifolia E1 1 . + . . . . . . Solidago virgaurea E1 . . . . . . . . . Clematis alpina E1 . + . . . . . . . Abies alba E1 . . . . . . . . . Homogyne sylvestris E1 . . . . . . . . . AF Aremonio-Fagion Lamium orvala E1 . . . . . . . . . Cardamine trifolia E1 . . . . + . . . . Anemone trifolia E1 . + . . . . . . . Cardamine enneaphyllos E1 . . . . . + . . . Cyclamen purpurascens E1 . . . . . . . r . Euphorbia carniolica E1 . . . . . . . . + Scopolia carniolica E1 . . . . . . . . . Omphalodes verna E1 . . . . . . . . . Primula vulgaris E1 . . . . . . . . . TA Tilio-Acerion Geranium robertianum E1 . 1 + . . . . . + Successive number of relevé (Zaporedna številka popisa) 1 2 3 4 5 6 7 8 9 DAKSKOBLER & MARTINČIČ: VEGETATION OF MOIST ROCK CREVICES AND MOIST (SLOPE) DEBRIS IN THE LIŠČAK GORGE 69FOLIA BIOLOGICA ET GEOLOGICA 64/1 – 2023 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 Pr. Fr. . . . . . . . . 1 1 . . . . . . 2 8 . . . + . . . . . . . . . . . . 1 4 + + r . . 1 + + 1 . . + + . . 1 14 56 . + . . . + . . . . r . . . . . 3 12 . . . + . . . . . . . . . . . . 1 4 . . . . . . . + . . . . . . . . 1 4 . . + r . . 1 + + + . . . 1 1 2 12 48 . . . . + + . . . . . . . + . . 5 20 . . . . . . . . + . . . 1 . . . 3 12 . . . . . . . . . . . r . . . + 2 8 1 1 . . . . . . . + . . . . + 1 5 20 1 . . + + . . . . . . . . . . . 3 12 . + . . . . . . . . . . . . . . 1 4 . . . . . . . . . . . . . . . . 1 4 . . . . . . . . . . . . . . . . 1 4 . . . . . + . . . . . . . . . . 1 4 + . . . . . . . . . . . . . . . 2 8 . . . + . . 1 . . . . . 1 . . . 4 16 . . . . . . . . . . r . . . . . 1 4 . + . . . . + . . . . . . r . . 3 12 . . . . . . . . . . . . . . . . 2 8 . . . . . . . . . . . . . . . + 2 8 + . + + . . . . . . . . . . . . 3 12 . . . . . . . . . . . . . . . . 1 4 . . . . . . . + . . . . . . . . 1 4 . . . . . . 1 + 1 + . . . . . . 6 24 . . . . . . . . . . 1 . . . . r 4 16 . . . . . . r . . . r . . . . + 3 12 . . . . . . . . . . . . . + . . 2 8 . . . . + . . . . . . . . . . . 1 4 . . . . . + + . . . r . . 1 1 + 9 36 . . + + + . . . . . . . . . . . 3 12 . . . . . . . . . . . . . . . . 1 4 . . . . . . . . . . . . . . . . 1 4 . . . + . . . . . . . . . . . . 1 4 + . . . . . . . + . . . . . . . 5 20 . 1 . . . . . . . . . . . . + . 4 16 . . . . . . r + . . . . . r . . 3 12 . . . . . . . . . . . . . . . . 1 4 . . . . + . . . . . . . . . . . 1 4 . . . . . . . . . . . . . r . . 1 4 + . r . . . . . . . . . . . . + 3 12 . . . . . . . . . . . . . + . . 2 8 . . . . . . . . . . . . . . . . 1 4 . . . . . . . . . . . . . . . . 1 4 . . . . . . . . . . . . . . . . 1 4 . . . . . . . . . . . . . . . . 1 4 . . . . . . . . . . . . . 1 . . 1 4 . . . . . . . . . . . . . + . . 1 4 . . . . . . . . . . . . . . r . 1 4 + + . + 1 . . . . . . . . + . + 9 36 DAKSKOBLER & MARTINČIČ: VEGETATION OF MOIST ROCK CREVICES AND MOIST (SLOPE) DEBRIS IN THE LIŠČAK GORGE 70 FOLIA BIOLOGICA ET GEOLOGICA 64/1 – 2023 Aruncus dioicus E1 . . . . . . . . . Polystichum aculeatum E1 . . . . . . . . . Acer pseudoplatanus E2a . . . . . . . . . Euonymus latifolia E2a . . . . . . . . . AI Alnion incanae Festuca gigantea E1 . . + . . . . . . FS Fagetalia sylvaticae Galeobdolon flavidum E1 r + . . 1 . . r . Mycelis muralis E1 . . . . . . r r . Salvia glutinosa E1 . . . . . . . . . Actaea spicata E1 . . . r . . . . . Sambucus nigra E2a . . . . . . . . . Mercurialis perennis E1 . 1 . . . . . . . Dryopteris filix-mas E1 . r . . . . . . . Phyteuma spicatum subsp. coeruleum E1 . . . . . + . . . Cardamine pentaphyllos E1 . . . . . + . . . Campanula trachelium E1 . . . . . . . r . Galium laevigatum E1 . . . . . . . . . Poa nemoralis E1 . . . . . . . . . Fagus sylvatica E2a . . . . . . . . . Fraxinus excelsior E2a . . . . . . . . . Lonicera alpigena E2a . . . . . . . . . Petasites albus E1 . . . . . . . . . Daphne mezereum E2a . . . . . . . . . QP Quercetalia pubescenti-petraeae Ostrya carpinifolia E1 . . . . . . . . . Euonymus verrucosa E2a . . . . . . . . . QF Querco-Fagetea Aegopodium podagraria E1 . 1 . . . . . . . Carex digitata E1 . . . . . . . . . Corylus avellana E2a . . . . . . . . . ML Mosses (Mahovi) Mnium thomsonii E0 . . . . 2 . + . . Tortella tortuosa E0 . . . . . . . . . Plagiochila porelloides E0 . . . . + . . . . Exertotheca crispa (Neckera crispa) E0 . . . . . . + . . Pedinophyllum interruptum E0 . . . . . . . . . Ctenidium molluscum E0 . . . . . . . . . Plagiomnium rostratum E0 . . . . . . . . . Lophozia sp. E0 . . . . . . . . . Reboulia hemisphaerica E0 . . + . . . . . . Brachythecium rutabulum E0 . . . 1 . . . . . Dichodontium pellucidum E0 . . . 1 . . . . . Mnium lycopodioides E0 . . . . . . . . . Fissidens taxifolius E0 . . . . . . . . . Alleniella complanata (Neckera complanata) E0 . . . . . . . . . Ptychostomum capillare (Bryum capillare) E0 . . . . . . . . . Encalypta streptocarpa E0 . . . . . . . . . Amblystegium serpens E0 . . . . . . . . . Philonotis caespitosa E0 . . . . . . . . . Entodon concinnus E0 . . . . . . . . . Legend - Legenda L Limestone - apnenec Ch Chert - roženec D Dolomite - dolomit Deb Debris - grušč M Marlstone - laporovec Li Lithosol - kamnišče Pr. Presence (number of relevés in which the species is presented) - število popisov, v katerih se pojavlja vrsta Fr. Frequency in % - frekvenca v % 1-11 Veronico urticifoliae-Violetum biflorae var. typica 12-15 Veronico urticifoliae-Violetum biflorae var. Astrantia carniolica 16-25 Veronico urticifoliae-Violetum biflorae var. Palustriella commutata 18-19 Veronico urticifoliae-Violetum biflorae var. Palustriella commutata subvar. Primula auricula Successive number of relevé (Zaporedna številka popisa) 1 2 3 4 5 6 7 8 9 DAKSKOBLER & MARTINČIČ: VEGETATION OF MOIST ROCK CREVICES AND MOIST (SLOPE) DEBRIS IN THE LIŠČAK GORGE 71FOLIA BIOLOGICA ET GEOLOGICA 64/1 – 2023 . . . + + + r . . + . . . + + . 7 28 . . + . . . . . . . . . . . . . 1 4 . . . . . . . . . . . . . + . . 1 4 . . . . . . . . . . . . . + . . 1 4 . . . . . . . . . . . . . . . . 1 4 . . . . + . . . + + . . . 1 + 2 10 40 . + . . . . . + + . r . . . . . 6 24 . . . . . . + . + + + . . + . . 5 20 . . . . . . . . . . . . + . . . 2 8 . . . . . . . . . . . . . + . + 2 8 . . . . . . . . . . . . . . . . 1 4 . . . . . . . . . . . . . . . . 1 4 . . . . . . . . . . . . . . . . 1 4 . . . . . . . . . . . . . . . . 1 4 . . . . . . . . . . . . . . . . 1 4 . . . . . . r . . . . . . . . . 1 4 . . . . . . . . . . . . + . . . 1 4 . . . . . . . . . . . . . + . . 1 4 . . . . . . . . . . . . . + . . 1 4 . . . . . . . . . . . . . + . . 1 4 . . . . . . . . . . . . . + . . 1 4 . . . . . . . . . . . . . + . . 1 4 + . . . . . . . . . . . . . . . 1 4 . . . . . . . . . . . . . + . . 1 4 . . . . . . . . . . . . . . . . 1 4 . . . r . . . . . . . . . . . . 1 4 . . . . . . . . . . . . . + . . 1 4 . . . . . 1 . . . . . . . . . . 3 12 . . . . + + . . . . . . . + . . 3 12 . . . . . + . . . . . . . . . . 2 8 . . . . . . + . . . . . . . . . 2 8 + + . . . . . . . . . . . . . . 2 8 + . . . + . . . . . . . . . . . 2 8 . + . . . . . . . . . . . + . . 2 8 . . . . 1 1 . . . . . . . . . . 2 8 . . . . . . . . . . . . . . . . 1 4 . . . . . . . . . . . . . . . . 1 4 . . . . . . . . . . . . . . . . 1 4 1 . . . . . . . . . . . . . . . 1 4 + . . . . . . . . . . . . . . . 1 4 + . . . . . . . . . . . . . . . 1 4 . + . . . . . . . . . . . . . . 1 4 . . . . . 1 . . . . . . . . . . 1 4 . . . . . . . . . . . . . . + . 1 4 . . . . . . . . . . . . . . . 1 1 4 . . . . . . . . . . . . . . . + 1 4 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 Pr. Fr. DAKSKOBLER & MARTINČIČ: VEGETATION OF MOIST ROCK CREVICES AND MOIST (SLOPE) DEBRIS IN THE LIŠČAK GORGE 72 FOLIA BIOLOGICA ET GEOLOGICA 64/1 – 2023 Table 9 (Preglednica 9): Paederoto luteae-Violetum biflorae, Cerastio subtriflorae-Violetum biflorae Successive number of relevé (Zaporedna številka popisa) 1 2 3 4 5 6 7 8 9 10 Database number of relevé (Delovna številka popisa) 28 18 17 28 18 18 28 18 19 28 18 21 28 05 38 28 24 50 28 29 05 28 69 86 25 77 39 28 21 01 Elevation in m (Nadmorska višina v m) 1000 1000 1000 1000 424 1850 1460 1330 1689 1850 Aspect (Lega) SW SW SW SW NE NW N SE NE W Slope in degrees (Nagib v stopinjah) 90 90 90 100 80 30 90 85 1 45 Parent material (Matična podlaga) LM LM LM LM D Li Br Li Deb Deb Soil (Tla) Li Li Li Li Li Li Li Li Li Li Stoniness in % (Kamnitost v %) 100 100 100 100 80 100 100 100 30 10 Cover of herb layer in % (Zastiranje zeliščne plasti v %): 60 30 35 35 40 60 30 30 80 90 Cover of moss layer in % (Zastiranje mahovne plasti v %): 40 40 30 20 80 50 60 20 40 60 Number of species (Število vrst) 13 13 14 13 15 9 25 24 12 18 Relevé area (Velikost popisne ploskve) m2 10 10 10 10 4 4 10 20 3 2 Date of taking relevé (Datum popisa) 6/ 29 /2 02 0 6/ 29 /2 02 0 6/ 29 /2 02 0 6/ 29 /2 02 0 4/ 27 /2 02 0 7/ 14 /2 02 0 10 /9 /2 02 0 8/ 19 /2 02 1 8/ 7/ 20 15 7/ 29 /2 02 0 Locality (Nahajališče) D re žn iš ke R av ne - C ur k D re žn iš ke R av ne - C ur k D re žn iš ke R av ne - C ur k D re žn iš ke R av ne - C ur k K oz ijs ka g ra pa - Tr eb uš a M an ga rt A pi ca - L op e G oz de c Lo pu čn ic a M an ga rt Quadrant (Kvadrant) 97 47 /2 97 47 /2 97 47 /2 97 47 /2 99 48 /2 95 47 /4 96 48 /1 96 46 /4 96 48 /4 95 47 /4 Coordinate GK Y (D-48) m 39 52 59 39 52 50 39 52 41 39 52 17 40 86 24 39 57 71 39 91 69 38 36 88 40 55 43 39 57 97 Coordinate GK X (D-48) m 51 26 19 3 51 26 19 4 51 26 19 9 51 26 19 8 51 05 35 8 51 44 63 2 51 38 19 1 51 33 74 2 51 30 33 5 51 44 54 5 Diagnostic species of syntaxa (Diagnostične vrste sintaksonov) Pr. Fr. AP Viola biflora E1 2 1 3 1 3 3 2 2 4 4 10 100 AP Paederota lutea E1 2 2 . + . + 1 1 + . 7 70 TR Trisetum argenteum E1 1 1 + + . . . . . . 4 40 MC Saxifraga aizoides E1 2 1 + . . . . . . . 3 30 AP Marchantia quadrata (Preissia quadrata) E0 1 1 1 . . . . . . . 3 30 MuASaxifraga rotundifolia E1 . . . . 1 . . . . . 1 10 AP Cystopteris fragilis E1 . . . . . 2 3 + . . 3 30 MC Conocephalum conicum E0 . . . + . 3 2 . . . 3 30 PcSp Campanula carnica E1 . . . + . . . 1 . . 2 20 AF Cyclamen purpurascens E1 . . . . . . . 1 . . 1 10 AP Heliosperma pusillum E1 . . . . . . 1 . 1 + 3 30 AC Saxifraga sedoides E1 . . . . . . . . 3 . 1 10 TR Cystopteris montana E1 . . . . . . + . 1 . 2 20 TR Festuca nitida E1 . . . . . . + . . 1 2 20 AC Sanionia uncinata E0 . . . . . . . . . 3 1 10 AC Doronicum glaciale E1 . . . . . . . . . 2 1 10 ES Cerastium subtriflorum E1 . . . . . . . . . 1 1 10 AP Astrantio carniolicae-Paederotion luteae Palustriella commutata E0 2 1 1 . 1 . 1 . . 2 6 60 Hymenostylium recurvirostrum E0 2 1 3 1 4 . . . . . 5 50 Aster bellidiastrum E1 2 1 2 . . . 1 . . . 4 40 Carex brachystachys E1 1 + + . . . . + . . 4 40 Orthothecium rufescens E0 . . . + + . 1 . 2 . 4 40 Apopellia endiviifolia (Pellia endiviifolia) E0 + . 1 1 . . . + . . 4 40 Fissidens dubius E0 . . . . + . + . . . 2 20 Asplenium viride E1 . . . . . . + . . + 2 20 DAKSKOBLER & MARTINČIČ: VEGETATION OF MOIST ROCK CREVICES AND MOIST (SLOPE) DEBRIS IN THE LIŠČAK GORGE 73FOLIA BIOLOGICA ET GEOLOGICA 64/1 – 2023 Valeriana saxatilis E1 + . . . . . . . . . 1 10 Astrantia carniolica E1 . . . . 1 . . . . . 1 10 Pinguicula alpina E1 . . . . . . + . . . 1 10 Valeriana tripteris E1 . . . . . . . + . . 1 10 PcSp Physoplexido comosae-Saxifragion petraeae Moehringia villosa E1 + . . . . . . . . . 1 10 PcSp Potentiletalia caulescentis Saxifraga crustata E1 . + . r . . . . . . 2 20 AT Asplenietea trichomanis Asplenium trichomanes E1 . + . 1 . . . + . . 3 30 Asplenium ruta-muraria E1 . . . . . . . 1 . . 1 10 Moehringia muscosa E1 . . . . . . . + . . 1 10 TR Thlaspietea trotundifolii Adenostyles glabra E1 + . + . . . . + . . 3 30 MC Montio-Cardaminetea Oxyrrhynchium schleicheri E0 . . . . + . + . . . 2 20 Flexitrichum flexicaule (Ditrichum flexicaule) E0 . . . . . . 1 . . + 2 20 Oxyrrhynchium hians E0 . . . . + . . . . . 1 10 Ptychostomum pseudotriquetrum (Bryum pseudotriquetrum) E0 . . . . . . . . . + 1 10 CD Caricetalia davallianae Campylium stellatum E0 . . . . . . + . . + 2 20 Parnassia palustris E1 . . . . . . . . + . 1 10 ES Elyno-Seslerietea Erigeron glabratus E1 . . . + . . . . . . 1 10 Sesleria caerulea E1 . . . . . . . + . . 1 10 Polygonum viviparum E1 . . . . . . . . . + 1 10 Mo Molinion Caltha palustris E1 . . . . + . . . . . 1 10 MuA Mulgedio-Aconitetea Ranunculus platanifolius E1 . . . . . . . + . . 1 10 EP Erico-Pinetea Calamagrostis varia E1 . . + . . . . + + . 3 30 VP Vaccinio-Piceetea Veronica urticifolia E1 . . r . . . . 1 . . 2 20 Dryopteris expansa E1 . . . . . . + . . . 1 10 AF Aremonio-Fagion Cardamine enneaphyllos E1 . . . . + . . . . . 1 10 Lamium orvala E1 . . . . + . . . . . 1 10 TA Tilio-Acerion Geranium robertianum E1 . + r + + . . . . . 4 40 FS Fagetalia sylvaticae Salvia glutinosa E1 . + + + . . . . . . 3 30 Cardamine pentaphyllos E1 . . . . + . . . . . 1 10 Mycelis muralis E1 . . . . + . . . . . 1 10 Galeobdolon flavidum E1 . . . . . . . + . . 1 10 ML Mosses (Mahovi) Mnium marginatum E0 . . . . . 1 + . + . 3 30 Tortella tortuosa E0 . . . . . . 1 + . . 2 20 Plagiomnium rostratum E0 . . . . . 3 . . . . 1 10 Brachythecium tenuicaule E0 . . . . . + . . . . 1 10 Pedinophyllum interruptum E0 . . . . . + . . . . 1 10 Pohlia wahlenbergii E0 . . . . . + . . . . 1 10 Campylophyllum halleri E0 . . . . . . 1 . . . 1 10 Didymodon fallax E0 . . . . . . 1 . . . 1 10 Lophozia sp. E0 . . . . . . 1 . . . 1 10 Mnium thomsonii E0 . . . . . . 1 . . . 1 10 Aneura pinguis E0 . . . . . . + . . . 1 10 Rhynchostegium murale E0 . . . . . . + . . . 1 10 Isothecium alopecuroides E0 . . . . . . . 1 . . 1 10 Anomodon viticulosus E0 . . . . . . . + . . 1 10 Encalypta streptocarpa E0 . . . . . . . + . . 1 10 Exertotheca crispa (Neckera crispa) E0 . . . . . . . + . . 1 10 Schistidium apocarpum E0 . . . . . . . + . . 1 10 Successive number of relevé (Zaporedna številka popisa) 1 2 3 4 5 6 7 8 9 10 Pr. Fr. DAKSKOBLER & MARTINČIČ: VEGETATION OF MOIST ROCK CREVICES AND MOIST (SLOPE) DEBRIS IN THE LIŠČAK GORGE 74 FOLIA BIOLOGICA ET GEOLOGICA 64/1 – 2023 Dermatocarpon miniatum E0 . . . . . . . + . . 1 10 Plagiochila asplenioides E0 . . . . . . . . 2 . 1 10 Ctenidium molluscum E0 . . . . . . . . 1 . 1 10 Scapania nemorea E0 . . . . . . . . 1 . 1 10 Climacium dendroides E0 . . . . . . . . . 1 1 10 Pohlia cruda E0 . . . . . . . . . + 1 10 Polytrichastrum alpinum E0 . . . . . . . . . + 1 10 Timmia austriaca E0 . . . . . . . . . + 1 10 Tritomaria quinquedentata E0 . . . . . . . . . + 1 10 Barbilophozia hatcheri E0 . . . . . . . . . + 1 10 Legend - Legenda L Limestone - apnenec Br Breccia - breča D Dolomite - dolomit Deb Debris - grušč M Marsltone - laporovec Li Lithosol - kamnišče Pr. Presence (number of relevés in which the species is presented) - število popisov, v katerih se pojavlja vrsta Fr. Frequency in % - frekvenca v % 1-8 Paederoto luteae-Violetum biflorae nom. prov. 9 Cerastio subtriflorae-Violetum biflorae Successive number of relevé (Zaporedna številka popisa) 1 2 3 4 5 6 7 8 9 10 Pr. Fr. DAKSKOBLER & MARTINČIČ: VEGETATION OF MOIST ROCK CREVICES AND MOIST (SLOPE) DEBRIS IN THE LIŠČAK GORGE 75FOLIA BIOLOGICA ET GEOLOGICA 64/1 – 2023 Table 10 (Preglednica 10): Asplenietea trichomanis Liščak Successive number of relevé (Zaporedna številka popisa) 1 3 2 4 5 6 7 8 9 10 11 Database number of relevé (Delovna številka popisa) 28 17 71 28 71 44 27 32 19 26 96 23 28 71 32 27 76 61 28 52 57 28 52 58 28 26 91 28 26 99 28 71 76 Elevation in m (Nadmorska višina v m) 770 945 1060 1105 1090 410 560 555 530 525 350 Aspect (Lega) S SW W SW SSW SW SSE SSE NW N NW Slope in degrees (Nagib v stopinjah) 80 90 80 90 90 80 95 95 60 80 70 Parent material (Matična podlaga) L LC lC h L L LR LM LC l LC l LC h LC h LM C h Soil (Tla) Li Li Li Li Li Li Li Li Li Li Li Stoniness in % (Kamnitost v %) 100 100 80 100 100 100 100 100 100 100 100 Cover of herb layer in % (Zastiranje zeliščne plasti v %): 30 20 40 20 30 70 30 20 40 40 40 Cover of moss layer in % (Zastiranje mahovne plasti v %): 5 5 10 5 5 5 5 5 30 40 20 Number of species (Število vrst) 10 10 19 13 17 25 13 4 20 37 24 Relevé area (Velikost popisne ploskve) m2 10 20 10 10 20 10 10 10 10 15 20 Date of taking relevé (Datum popisa) 6/ 24 /2 02 0 8/ 10 /2 02 1 9/ 18 /2 01 8 6/ 9/ 20 17 8/ 10 /2 02 1 6/ 26 /2 01 9 4/ 8/ 20 21 4/ 8/ 20 21 9/ 15 /2 02 0 9/ 15 /2 02 0 6/ 18 /2 02 1 Locality (Nahajališče) Li sč ak -S op ot Li šč ak p ov ir je Li šč ak -M le čn i r ob Li šč ak -G ra nt ar sk a po lja na Li šč ak -G ra nt ar sk a po lja na Li šč ak -D rs el a Li šč ak -L is ec Li šč ak Li šč ak Li šč ak Li šč ak Quadrant (Kvadrant) 97 49 /3 97 49 /3 98 49 /1 97 49 /3 97 49 /3 98 49 /1 97 49 /3 97 49 /3 98 49 /1 98 49 /1 98 49 /1 Coordinate GK Y (D-48) m 41 17 10 41 17 54 41 22 19 41 20 53 41 20 42 41 11 54 41 15 33 41 15 39 41 13 74 41 14 06 41 09 18 Coordinate GK X (D-48) m 51 18 61 4 51 18 93 6 51 16 78 8 51 18 96 9 51 18 97 2 51 17 37 1 51 18 07 0 51 18 07 2 51 17 85 8 51 17 91 0 51 17 04 2 Diagnostic species of syntaxa (Diagnostične vrste sintaksonov) Pr. AT Sedum album E1 1 + 1 1 3 . . . . . . 5 ES Sesleria caerulea E1 1 1 3 . . . . . + . . 4 PcSp Campanula carnica E1 1 . . . 1 . . . . . . 2 PcSp Silene hayekiana E1 + . . . . . . . . . . 1 TR Arabis alpina E1 . . . + + . . . . . . 2 AT Asplenium trichomanes E1 + + + + + 2 3 2 2 1 . 10 AP Fissidens dubius E0 + . . . . . + . 1 1 1 5 FS Galeobdolon flavidum E1 . . . . . . . . 1 1 3 3 AP Valeriana tripteris E1 . . . . . . . . + 1 . 2 TA Phyllitis scolopendrium E1 . . . . . . . . . + 2 2 AP Palustriella commutata E0 . . . . . . . . . . 1 1 PsSp Physoplexido comosae-Saxifragion petraeae Athamanta turbith E1 . . 1 . . . . . . . . 1 AP Astrantio carniolicae-Paederotion luteae Cystopteris fragilis E1 . . . 1 + . . . . + . 3 Carex brachystachys E1 . . . . . + . . . + . 2 Orthothecium rufescens E0 . . . . . . + . . 1 . 2 Eucladium verticillatum E0 . . . . . . + . . . . 1 Asplenium viride E1 . . . . . . . . . 1 . 1 Hydrogonium croceum (Barbula crocea) E0 . . . . . . . . . + . 1 Hymenostylium recurvirostrum E0 . . . . . . . . . + . 1 Apopellia endiviifolia (Pellia endiviifolia) E0 . . . . . . . . . + . 1 Jungermannia atrovirens E0 . . . . . . . . . + . 1 DAKSKOBLER & MARTINČIČ: VEGETATION OF MOIST ROCK CREVICES AND MOIST (SLOPE) DEBRIS IN THE LIŠČAK GORGE 76 FOLIA BIOLOGICA ET GEOLOGICA 64/1 – 2023 AT Asplenietea trichomanis Asplenium ruta-muraria E1 . . . + + + . . . + . 4 Polypodium vulgare E1 . . + . . . . . . . . 1 Festuca stenantha E1 . . . . r . . . . . . 1 Moehringia muscosa E1 . . . . . . + . . . . 1 TR Thlaspietea rotundifolii Gymnocarpium robertianum E1 . . . . . + . . . + . 2 Achnatherum calamagrostis E1 . + . . . . . . . . . 1 Hieracium bifidum E1 . . . . . + . . . . . 1 Adenostyles glabra E1 . . . . . . . . . 1 . 1 MC Montio-Cardaminetea Conocephalum conicum E0 . . . . . + + . . + . 3 Gymnostomum aeruginosum E0 . 1 . . + . . . . . . 2 Oxyrrhynchium hians E0 . . . . . . . . . 1 + 2 Oxyrrhynchium schleicheri E0 . . . . . . + . . . . 1 Brachythecium rivulare E0 . . . . . . . . 1 . . 1 Cololejeunea calcarea E0 . . . . . . . . . 1 . 1 ES Elyno-Seslerietea Festuca calva E1 . . . + 1 . . . . . . 2 Erigeron glabratus E1 . . + . . . . . . . . 1 FB Festuco-Brometea Dianthus monspessulanus E1 . + + . + . . . . . . 3 Allium carinatum subsp. pulchellum E1 . . + . . . . . . . . 1 Thymus praecox E1 . . + . . . . . . . . 1 Bromopsis erecta E1 . . . . r . . . . . . 1 Buphthalmum salicifolium E1 . . . . . . . . . + . 1 TG Trifolio-Geranietea Campanula rapunculoides E1 r + . . . + + . . . . 4 Digitalis grandiflora E1 . . . . . + . . . . . 1 Hypericum perforatum E1 . . . . . + . . . . . 1 Laserpitium latifolium E1 + . . . . . . . . . . 1 Libanotis daucifolia E1 + . . . . . . . . . . 1 Silene nutans E1 . + . . . . . . . . . 1 Achillea distans E1 . . . . + . . . . . . 1 Mo Molinion Angelica sylvestris E1 . . . . . . . . . + . 1 EP Erico-Pinetea Calamagrostis varia E1 . . . . . . r . . . + 2 EA Epilobietea angustifolii Rubus idaeus E1 . . . . + . . . . . . 1 Bromopsis benekenii E1 . . . . . + . . . . . 1 MuA Mulgedio-Aconitetea Senecio ovatus E1 . . . . + . . . 1 + . 3 Aconitum angustifolium E1 . . . . . . . . . + . 1 VP Vaccinio-Piceetea Oxalis acetosella E1 . . . . . . . + + + + 4 Veronica urticifolia E1 + . . . . . . . . + . 2 Solidago virgaurea E1 . . . . . . . . . + . 1 AF Aremonio-Fagion Cardamine trifolia E1 . . . . . . . . + 1 . 2 Cyclamen purpurascens E1 . . . . . . . + + . . 2 TA Tilio-Acerion Geranium robertianum E1 . . . + + + . . . . . 3 Tephroseris pseudocrispa E1 . . . . . + . . 1 . . 2 Polystichum setiferum E1 . . . . . + . . . . 1 2 Acer pseudoplatanus E1 . . . . . . . . . . + 1 Aruncus dioicus E1 . . . . . . . . . . + 1 Polystichum aculeatum E1 . . . . . . . . . . + 1 FS Fagetalia sylvaticae Salvia glutinosa E1 . + . + + + . . 1 + . 6 Galium laevigatum E1 . . . . . + . . . + + 3 Poa nemoralis E1 . . . + . 1 . . . . . 2 Successive number of relevé (Zaporedna številka popisa) 1 3 2 4 5 6 7 8 9 10 11 Pr. DAKSKOBLER & MARTINČIČ: VEGETATION OF MOIST ROCK CREVICES AND MOIST (SLOPE) DEBRIS IN THE LIŠČAK GORGE 77FOLIA BIOLOGICA ET GEOLOGICA 64/1 – 2023 Asarum europaeum subsp. caucasicum E1 . . . . . + . . . + . 2 Fagus sylvatica E1 . . . + . . . . . . . 1 Myosotis sylvatica E1 . . . + . . . . . . . 1 Brachypodium sylvaticum E1 . . . . . + . . . . . 1 Campanula trachelium E1 . . . . . + . . . . . 1 Circaea lutetiana E1 . . . . . + . . . . . 1 Dryopteris filix-mas E1 . . . . . + . . . . . 1 Mycelis muralis E1 . . . . . + . . . . . 1 Petasites albus E1 . . . . . . . . . . + 1 QP Quercetalia pubescenti-petraeae Sesleria autumnalis E1 . . . . . . . . + + . 2 Primula veris subsp. columnae E1 . . . + . . . . . . . 1 Arabis turrita E1 . . . . . + . . . . . 1 Fraxinus ornus E1 . . . . . . . . . . + 1 QP Querco-Fagetea Hedera helix E1 . . . . . + . . + . + 3 Carex digitata E1 . . . . . . . . 1 + . 2 Vinca minor E1 . . . . . . r . . . . 1 Taxus baccata E2a . . . . . . . . + . . 1 Hepatica nobilis E1 . . . . . . . . . . + 1 Veratrum nigrum E1 . . . . . . . . . . + 1 ML Mosses (Mahovi) Ctenidium molluscum E0 . . . . . . . + 1 1 1 4 Isothecium alopecuroides E0 . 1 . . . . . . 1 . 1 3 Tortella tortuosa E0 . . 1 . . 1 + . . . . 3 Mnium marginatum E0 . . . . . . + . 1 1 1 4 Plagiomnium rostratum E0 . . . . . . . . + 1 . 2 Homalothecium lutescens E0 . . 1 . . . . . . . . 1 Schistidium apocarpum E0 . . 1 . . . . . . . . 1 Encalypta streptocarpa E0 . . . 1 . . . . . . . 1 Anomodon viticulosus E0 . . . . 1 . . . . . . 1 Chionoloma tenuirostre (Oxystegus tenuirostre) E0 . . . . . . + . . . . 1 Didymodon insulanus E0 . . . . . . . . . + . 1 Rhynchostegium murale E0 . . . . . . . . . + . 1 Plagiomnium undulatum E0 . . . . . . . . . . 1 1 Thamnobryum alopecurum E0 . . . . . . . . . . 1 1 Exertotheca crispa (Neckera crispa) E0 . . . . . . . . . . + 1 Plagiochila porelloides E0 . . . . . . . . . . + 1 Legend - Legenda 1 Sileno hayekianae-Campanuletum carnicae nom. prov. 2-3 Seslerio caeruleae-Sedetum albi nom. prov. 4-5 Arabido alpinae-Sedetum albi nom. prov. 6-8 Tortello tortuosae-Asplenietum trichomanis nom. prov. 9-10 Valeriano tripteridis-Veronicetum urticifoliae nom. prov. 11 Palustriello commutati-Phyllitidetum scolopendrii nom. prov. L Limestone - apnenec Ch Chert - roženec Cl Claystone - glinavec M Marlstone - laporovec Li Lithosol - kamnišče Pr. Presence (number of relevés in which the species is presented) - število popisov, v katerih se pojavlja vrsta Fr. Frequency in % - frekvenca v % Successive number of relevé (Zaporedna številka popisa) 1 3 2 4 5 6 7 8 9 10 11 Pr. DAKSKOBLER & MARTINČIČ: VEGETATION OF MOIST ROCK CREVICES AND MOIST (SLOPE) DEBRIS IN THE LIŠČAK GORGE 78 FOLIA BIOLOGICA ET GEOLOGICA 64/1 – 2023 Table 11: Communities of moist rock crevices in NW and W Slovenia Preglednica 11: Združbe vlažnih skalnih razpok v severozahodni in zahodni Sloveniji Successive number (Zaporedna številka) 1 2 3 4 5 6 7 8 Number of relevés (Število popisov) 14 18 18 25 10 33 28 10 Sign for syntaxa (Oznaka sintaksonov) AcPa PcAc VbAc VuVb PcVu VuSc CvAb CsSaac Number of species (Število vrst) 55 78 101 119 72 112 138 53 Astrantio carniolicae-Paederotion luteae Pinguicula alpina E1 100 28 22 4 . . 4 . Hymenostylium recurvirostrum E0 100 67 38 48 50 6 14 20 Palustriella commutata E0 100 83 38 48 100 22 7 30 Aster bellidiastrum E1 86 28 61 32 . 6 100 10 Astrantia carniolica E1 71 100 100 24 20 3 4 . Apopellia endiviifolia (Pellia endiviifolia) E0 57 56 11 32 60 9 18 30 Carex brachystachys E1 50 61 78 40 10 12 11 . Orthothecium rufescens E0 43 50 89 76 10 12 . . Eucladium verticillatum E0 43 28 . 4 10 6 4 . Tofieldia calyculata E1 36 17 17 . . . 7 . Valeriana tripteris E1 29 11 17 24 30 39 18 . Marchantia quadrata (Preissia quadrata) E0 21 5 22 . . . 4 . Fissidens dubius E0 14 17 33 20 40 55 4 . Hydrogonium croceum (Barbula crocea) E0 14 33 17 12 . 9 18 20 Jungermannia atrovirens E0 14 6 . 16 . 3 14 . Valeriana saxatilis E1 7 22 44 4 . . 4 . Saxifraga aizoides E1 7 . . . . . . . Paederota lutea E1 . 22 67 24 10 3 14 . Asplenium viride E1 . 17 72 64 10 6 . . Viola biflora E1 . . 100 100 . 3 4 . Heliosperma pusillum E1 . . 39 . . . . . Cystopteris fragilis E1 . . 33 44 20 15 11 . Cystopteris regia E1 . . 33 . . . . . Cyrtomnium hymenophylloides E0 . . 11 . . . . . Primula carniolica E1 . . . 4 . . . . Selaginella helvetica E1 . . . . . . 14 . Physoplexido comosae-Saxifragion petraeae Campanula cespitosa E1 29 . 6 . . . 21 100 Hieracium porrifolium E1 7 6 6 . . . 14 80 Hieracium pospichalii E1 . 6 . . . . 21 . Campanula zoysii E1 . . 17 . . . 4 . Campanula carnica E1 . . 11 12 30 3 14 . Phyteuma scheuchzeri subsp. columnae E1 . . 6 12 . 18 4 . Saxifraga petraea E1 . . . . . 18 . . Athamanta turbith E1 . . . . . . 11 . Micromeria thymifolia E1 . . . . . . 7 . Potentilletalia caulescentis Saxifraga crustata E1 7 6 . . 10 3 . 70 Campanula cochleariifolia E1 . . 33 . . . . . Arabis stellulata E1 . . 22 . . . . . Primula auricula E1 . . 22 8 . . . . Potentilla caulescens E1 . . 11 4 . . 4 . Hieracium humile E1 . . 6 . . . . . Potentilla clusiana E1 . . 6 . . . . . Festuca stenantha E1 . . . . . . 7 . Saxifraga hostii E1 . . . . . . 7 . Asplenietea trichomanis Asplenium ruta-muraria E1 21 6 28 12 30 21 14 . Asplenium trichomanes E1 14 28 17 56 100 97 11 . Kernera saxatilis E1 . 6 6 4 . . . . Polypodium interjectum E1 . 6 . . . . . . Moehringia muscosa E1 . . 6 4 . 27 . . Polypodium vulgare E1 . . . . . 6 . . Sedum album E1 . . . . . 3 . . Sedum hispanicum E1 . . . . . 3 . . Ceterach javorkeanum E1 . . . . . 3 . . DAKSKOBLER & MARTINČIČ: VEGETATION OF MOIST ROCK CREVICES AND MOIST (SLOPE) DEBRIS IN THE LIŠČAK GORGE 79FOLIA BIOLOGICA ET GEOLOGICA 64/1 – 2023 Thlaspietea rotundifolii Hieracium bifidum E1 50 17 22 12 10 18 36 30 Petasites paradoxus E1 43 6 6 . . . 21 10 Adenostyles glabra E1 29 44 67 48 50 6 11 10 Gymnocarpium robertianum E1 . 6 22 20 10 . 11 . Achnatherum calamagrostis E1 . 6 . . . . 14 100 Peucedanum verticillare E1 . 6 . . . . 7 . Soldanella minima E1 . . 17 . . . 4 . Hieracium bifidum E1 . . 22 . . . . . Gypsophila repens E1 . . 11 . . . . . Festuca nitida E1 . . 11 . . . . . Cystopteris montana E1 . . 11 . . . . . Saxifraga caesia E1 . . 6 . . . . . Arabis alpina E1 . . 6 . . . . . Athamanta cretensis E1 . . 6 . . . . . Biscutella laevigata E1 . . 6 . . . . . Cerastium carinthiacum E1 . . 6 . . . . . Trisetum argenteum E1 . . 6 8 . . 21 30 Hieracium glaucum E1 . . . . . . 7 90 Poa compressa E1 . . . . . . 7 . Centaurea dichroantha E1 . . . . . . 4 . Hieracium piloselloides E1 . . . . . . 4 . Hieracium dollineri E1 . . . . . . . 10 Montio-Cardaminetea Conocephalum conicum E0 14 56 44 100 100 42 25 . Aneura pinguis E0 7 . . . . . 4 . Oxyrrhynchium hians E0 . 22 . 20 40 21 4 . Cratoneuron filicinum E0 . 17 11 . 10 3 7 . Brachythecium rivulare E0 . 17 . . 20 . 25 . Ptychostomum pseudotriquetrum (Bryum pseudotriquetrum) E0 . 11 6 . . . . . Oxyrrhynchium schleicheri E0 . 6 6 12 40 . . . Rhynchostegium riparioides (Platyhypnidium riparioides) E0 . 6 . 4 20 . . . Gymnostomum aeruginosum E0 . 6 . . . 6 . . Cololejeunea calcarea E0 . . 6 4 . . 4 . Flexitrichum flexicaule (Ditrichum flexicaule) E0 . . . 4 . . . . Fontinalis antipyretica subsp. antipyretica E0 . . . 4 . . 4 . Palustriella decipiens E0 . . . . . . 4 . Calliergonella cuspidata E0 . . . . . . 4 . Saxifraga aizoides E1 . . . . . . . 100 Caricetalia davallianae Campylium stellatum E0 7 6 6 8 . . 4 . Carex lepidocarpa E1 7 . . . . . 4 . Campylophyllopsis calcarea (Campylium calcareum) E0 7 . . . . . . . Calliergonella lindbergii E0 . . . . 10 . . . Carex flava agg. E1 . . . . . . 4 . Elyno-Seslerietea Sesleria caerulea E1 50 33 39 16 10 9 25 30 Carex mucronata E1 . 22 . . . . 4 . Betonica alopecuros E1 . 11 . . . . . . Carex firma E1 . . 28 . . . . . Laserpitium peucedanoides E1 . . 22 . . . . . Carex ferruginea E1 . . 17 4 . . 4 10 Achillea atrata E1 . . 6 . . . . . Globularia cordifolia E1 . . 6 . . . . . Selaginella selaginoides E1 . . 6 . . . . . Ranunculus hybridus E1 . . 6 . . . . . Phyteuma orbiculare E1 . . 6 . . . . . Sesleria tenuifolia subsp. kalnikensis E1 . . . . . 3 . . Carex ornithopodoides E1 . . . . . . 11 . Erigeron glabratus E1 . . . . . . 4 . Poo alpinae-Trisetetalia Poa alpina E1 . . . . . . 4 . Successive number (Zaporedna številka) 1 2 3 4 5 6 7 8 DAKSKOBLER & MARTINČIČ: VEGETATION OF MOIST ROCK CREVICES AND MOIST (SLOPE) DEBRIS IN THE LIŠČAK GORGE 80 FOLIA BIOLOGICA ET GEOLOGICA 64/1 – 2023 Molinio-Arrhenatheretea Angelica sylvestris E1 14 17 . 12 . . 4 . Crepis paludosa E1 . 17 . 8 . . 4 . Caltha palustris E1 . 6 . 8 . . . . Taraxacum sect. Palustria E1 . . . . . . 7 . Filipendula ulmaria E1 . . . . . . 4 . Deschampsia cespitosa E1 . . . . . . 14 . Prunella vulgaris E1 . . . . . . 11 . Leontodon hispidus E1 . . . . . . 7 . Agrostis stolonifera E1 . . . . . . 4 . Dactylis glomerata E1 . . . . . . 4 . Galium mollugo E1 . . . . . . 4 . Taraxacum sect. Ruderalia E1 . . . . . . 4 . Betulo-Alnetea Salix appendiculata E2a 21 6 11 . . 3 36 20 Mulgedio-Aconitetea Senecio ovatus E1 . 6 . 12 50 15 . . Aconitum angustifolium E1 . 6 . . . . . . Chaerophyllum hirsutum E1 . 6 . . . . . . Saxifraga rotundifolia E1 . . 17 16 . 3 . . Aconitum degenii subsp. paniculatum E1 . . 6 . . 3 . . Aconitum lycoctonum subsp. ranunculifolium E1 . . 6 . . . . . Chaerophyllum hirsutum E1 . . 6 24 . . 14 . Thalictrum aquilegiifolium E1 . . . 8 . . . . Phyteuma ovatum E1 . . . 4 . 3 . . Petasites hybridus E1 . . . . . . 7 . Epilobietea angustifolii, Sambuco-Salicion capreae Eupatorium cannabinum E1 21 22 . . . . 14 20 Tussilago farfara E1 7 . . . . . 11 20 Solanum dulcamara E1 . 6 . 4 . . . . Verbascum lanatum E1 . 6 . . . . . . Rubus idaeus E1 . . . 4 10 . . . Salix caprea E2a . . . . . 3 . . Cardamine hirsuta E1 . . . . . . 4 . Carex oederi E1 . . . . . . 4 . Fragaria vesca E1 . . . . . . 4 . Galinsoga ciliata E1 . . . . . . 4 . Trifolio-Geranietea Campanula rapunculoides E1 . . . 12 20 3 14 . Digitalis grandiflora E1 . . . . 10 3 . . Calamintha einseleana E1 . . . . . . . 70 Hypericum perforatum E1 . . . . . 3 4 20 Laserpitium siler E1 . . . . . 3 . . Libanotis daucifolia E1 . . . . . . . 20 Erico-Pinetea Calamagrostis varia E1 86 78 67 36 20 9 86 10 Molinia arundinacea E1 64 44 . . . . 4 60 Erica carnea E1 29 . 6 . . . 11 10 Buphthalmum salicifolium E1 14 11 . . 10 . 25 10 Carex ornithopoda E1 14 . . 4 . 3 21 . Cirsium erisithales E1 7 6 . . 10 3 . . Rhodothamnus chamaecistus E1 . . 33 4 . . . . Rhododendron hirsutum E1 . . 22 . . . . . Asperula aristata E1 . . 6 . . . . . Aquilegia nigricans E1 . . . 12 . . . . Carex alba E1 . . . 4 . . . . Epipactis atrorubens E1 . . . . . . . 10 Leontodon incanus E1 . . . . . . . 7 Euphrasia cuspidata E1 . . . . . . . 4 Rubus saxatilis E1 . . . . . . . 4 Festuco-Brometea . . . . . . . . Brachypodium rupestre E1 . . . . . . 4 10 Successive number (Zaporedna številka) 1 2 3 4 5 6 7 8 DAKSKOBLER & MARTINČIČ: VEGETATION OF MOIST ROCK CREVICES AND MOIST (SLOPE) DEBRIS IN THE LIŠČAK GORGE 81FOLIA BIOLOGICA ET GEOLOGICA 64/1 – 2023 Thymus praecox E1 . . . . . . 4 10 Euphorbia cyparissias E1 . . . . . . 4 . Koeleria pyramidata E1 . . . . . . 4 . Linum catharticum E1 . . . . . . 4 . Plantago media E1 . . . . . . 4 . Taraxacum sect. Erythrosperma E1 . . . . . . 4 . Carlina acaulis E1 . . . . . . . 20 Bromopsis erecta E1 . . . . . . . 20 Carex humilis E1 . . . . . . . 10 Pimpinella saxifraga E1 . . . . . . . 10 Vaccinio-Piceetea Veronica urticifolia E1 14 61 39 80 100 82 46 10 Aposeris foetida E1 7 6 . . . 3 . . Gentiana asclepiadea E1 7 17 . . . . . . Saxifraga cuneifolia E1 . 6 17 16 20 100 . . Homogyne sylvestris E1 . 6 6 4 . 3 . . Oxalis acetosella E1 . . 11 20 30 30 7 . Solidago virgaurea E1 . . . 12 20 15 . . Clematis alpina E1 . . . 4 . . 7 . Abies alba E1 . . . 4 . . . . Calamagrostis arundinacea E1 . . . . 10 6 4 . Hieracium murorum E1 . . . . . 3 . . Larix decidua E2a . . . . . . . 10 Picea abies E2a . . . . . . . 10 Aremonio-Fagion, Erythronio-Carpinion Cyclamen purpurascens E1 7 6 6 4 . 30 . . Primula vulgaris E1 7 . . 4 10 3 . 10 Cardamine trifolia E1 . 6 6 8 60 12 . . Lamium orvala E1 . 6 . 12 20 6 . . Anemone trifolia E1 . . 6 4 . 3 . . Scopolia carniolica E1 . . . 4 . 3 . . Dentaria enneaphyllos E1 . . . 4 . 3 . . Euphorbia carniolica E1 . . . 4 . . . . Omphalodes verna E1 . . . 4 . . . . Tilio-Acerion Aruncus dioicus E1 7 28 11 28 20 15 18 Geranium robertianum E1 . 17 17 36 30 15 11 20 Phyllitis scolopendrium E1 . 11 11 . 20 3 18 Polystichum aculeatum E1 . 11 . 4 . 9 . Acer pseudoplatanus E1 . 6 . 4 . 9 . Ulmus glabra E1 . 11 . . 10 9 . Euonymus latifolia E2a . . . 4 . . . Circaea x intermedia E1 . . . . 10 . . Polystichum setiferum E1 . . . . 10 9 . Polystichum x wirtgenii E1 . . . . 10 . 4 Tephroseris pseudocrispa E1 . . . . . 12 . Chrysosplenium alternifolium E1 . . . . . 6 . Alnion incanae Knautia drymeia subsp. intermedia E1 . 11 . . . . 4 . Cardamine impatiens E1 . 6 . . . . . . Rubus caesius E1 . . . . . . 11 . Alnus glutinosa E2a . . . . . . 7 . Salix eleagnos E2a . . . . . . 4 . Frangula alnus E2a . . . . . . 4 . Viburnum opulus E2a . . . . . . 4 . Fagetalia sylvaticae Galium laevigatum E1 43 44 6 4 30 27 14 . Salvia glutinosa E1 29 6 . 20 20 15 11 . Fagus sylvatica E1 7 . 6 . . . 4 . Mycelis muralis E1 7 11 6 24 40 42 11 . Galeobdolon flavidum E1 . 39 28 40 70 58 14 . Brachypodium sylvaticum E1 . 17 . . . . 7 . Successive number (Zaporedna številka) 1 2 3 4 5 6 7 8 DAKSKOBLER & MARTINČIČ: VEGETATION OF MOIST ROCK CREVICES AND MOIST (SLOPE) DEBRIS IN THE LIŠČAK GORGE 82 FOLIA BIOLOGICA ET GEOLOGICA 64/1 – 2023 Petasites albus E1 . 11 . 4 20 . . . Fraxinus excelsior E1 . 6 . 4 20 . . . Dryopteris filix-mas E1 . 6 . 4 . . . . Sambucus nigra E2a . . . 8 10 3 . . Poa nemoralis E1 . . . 4 10 3 . . Campanula trachelium E1 . . . 4 10 . 4 . Cardamine pentaphyllos E1 . . . 4 . 12 . . Mercurialis perennis E1 . . . 4 . 9 . . Actaea spicata E1 . . . 8 . 3 . . Daphne mezereum E2a . . . 4 . . . . Lonicera alpigena E2a . . . 4 . . . . Phyteuma spicatum subsp. coeruleum E1 . . . 4 . . . . Asarum europaeum subsp. caucasicum E1 . . . . 10 15 . . Festuca altissima E1 . . . . 10 3 . . Prenanthes purpurea E1 . . . . . 12 . . Allium ursinum E1 . . . . . 3 . . Euphorbia dulcis E1 . . . . . 3 . . Epilobium montanum E1 . . . . . . 4 . Ranunculus lanuginosus E1 . . . . . . 4 . Tilia cordata E2a . . . . . . 4 . Viola reichenbachiana E1 . . . . . . 4 . Quercetalia pubescenti-petraeae Carex flacca E1 29 11 . . . . . 10 Ostrya carpinifolia E2a 7 . . 4 . 3 14 60 Euonymus verrucosa E2a . . . 4 . . . . Sesleria autumnalis E1 . . . . 30 6 . 20 Arabis turrita E1 . . . . 20 12 . . Fraxinus ornus E3a . . . . . . . 10 Fraxinus ornus E2a . . . . . 3 . 50 Querco-Fagetea Hedera helix E1 14 11 . . 50 52 14 . Carex digitata E1 7 11 17 4 40 45 4 . Clematis vitalba E1 7 . . . 10 3 . . Potentilla erecta E1 . 6 . . . . . . Aegopodium podagraria E1 . . . 4 . . . . Corylus avellana E2a . . . 4 . . . . Veratrum nigrum E1 . . . . 10 18 . . Hepatica nobilis E1 . . . . . 6 . . Festuca heterophylla E1 . . . . . 3 . . Hieracium racemosum E1 . . . . . . 4 . Hieracium umbellatum E1 . . . . . . 4 . Viola riviniana E1 . . . . . . 4 . Mosses (Mahovi) Tortella tortuosa E0 7 . 22 12 10 18 54 90 Exertotheca crispa (Neckera crispa) E0 7 . 11 8 . 70 4 . Dichodontium pellucidum E0 7 . . 4 . . 7 . Didymodon fallax E0 7 . . . . . . . Seligera trifaria E0 7 . . . . . . . Plagiomnium rostratum E0 . 17 . 8 40 6 14 . Hygrohypnum luridum E0 . 11 . . . . 29 . Mnium marginatum E0 . 11 . . . 18 . . Ctenidium molluscum E0 . 6 33 8 . 48 57 10 Plagiochila porelloides E0 . 6 11 8 . . . . Lophozia sp. E0 . 6 6 8 . . 4 . Alleniella complanata (Neckera complanata) E0 . 6 . 4 . 3 . . Dichodontium pellucidum E0 . 6 . . . . . . Trichostomum crispulum E0 . 6 . . . . 4 . Chionoloma tenuirostre (Oxystegus tenuirostre) E0 . 6 . . . . . . Microlejeunea ulicina E0 . 6 . . . . . . Mnium thomsonii E0 . . 11 12 20 15 . . Plagiomnium undulatum E0 . . 11 . 10 15 . . Amblystegium serpens E0 . . 6 4 . 3 4 . Successive number (Zaporedna številka) 1 2 3 4 5 6 7 8 DAKSKOBLER & MARTINČIČ: VEGETATION OF MOIST ROCK CREVICES AND MOIST (SLOPE) DEBRIS IN THE LIŠČAK GORGE 83FOLIA BIOLOGICA ET GEOLOGICA 64/1 – 2023 Thamnobryum alopecurum E0 . . 6 . 30 18 4 . Bryum sp. E0 . . 6 . . 3 . . Pohlia wahlenbergii E0 . . 6 . . . . 40 Campylophyllum halleri E0 . . 6 . . . . . Didymodon acutus E0 . . 6 . . . . . Lophozia obtusa E0 . . 6 . . . . . Metzgeria conjugata E0 . . 6 . . . . . Serpoleskea confervoides (Amblystegium confervoides) E0 . . 6 . . . . . Myurella julacea E0 . . 6 . . . . . Pedinophyllum interruptum E0 . . . 8 10 6 18 . Brachythecium rutabulum E0 . . . 4 . 6 14 . Encalypta streptocarpa E0 . . . 4 . 6 7 . Fissidens taxifolius E0 . . . 4 . . . . Mnium lycopodioides E0 . . . 4 . . . . Philonotis caespitosa E0 . . . 4 . . . . Reboulia hemisphaerica E0 . . . 4 . . . . Entodon concinnus E0 . . . 4 . . . . Ptychostomum capillare (Bryum capillare) E0 . . . 4 . . . . Mnium stellare E0 . . . . 10 . . . Isothecium alopecuroides E0 . . . . . 39 . . Anomodon viticulosus E0 . . . . . 15 . . Pseudanomodon attenuatus (Anomodon attenuatus) E0 . . . . . 12 . . Chionoloma tenuirostre (Oxystegus tenuirostre) E0 . . . . . 6 . . Brachythecium salebrosum E0 . . . . . 3 . . Homalothecium philippeanum E0 . . . . . 3 . . Mnium spinulosum E0 . . . . . 3 . . Weisia sp. E0 . . . . . 3 . . Didymodon spadiceus E0 . . . . . . 21 . Schistidium apocarpum E0 . . . . . . 18 . Cirriphyllum crassinervium E0 . . . . . . 11 . Mesoptychia collaris (Leiocolea collaris) E0 . . . . . . 4 . Campylopus sp. E0 . . . . . . 4 . Hypnum cupressiforme E0 . . . . . . 4 . Pseudoleskeella catenulata E0 . . . . . . 4 . Rhynchostegium murale E0 . . . . . . 4 . Jungermannia sp. E0 . . . . . . 4 . Sciuro-hypnum starkei (Brachythecium starkei) E0 . . . . . . 4 . Didymodon insulanus E0 . . . . . . 4 . Dichodontium flavescens E0 . . . . . . 4 . Plagiomnium elatum E0 . . . . . . 4 . Hylocomium splendens E0 . . . . . . 4 . Didymodon vinealis E0 . . . . . . . 10 Legend / Legenda AcPa Astrantio carniolicae-Pinguiculetum alpinae (Table 1) PcAc Palustriello commutati-Astrantietum carniolicae (Table 3) VbAc Violo biflorae-Astrantietum carniolicae (Table 2) VuVb Veronico urticifoliae-Violetum biflorae (Table 8) PcVu Palustriello commutati-Veronicetum urticifoliae (Table 4) VuSc Veronico urticifoliae-Saxifragetum cuneifolii (Table 6) CvAb Calamagrostio variae-Asteretum bellidiastri (Table 7) CcSaac Campanulo cespitosae-Saxifragetum aizoidis achnatheretosum calamagrostis (Table 5) Successive number (Zaporedna številka) 1 2 3 4 5 6 7 8 DAKSKOBLER & MARTINČIČ: VEGETATION OF MOIST ROCK CREVICES AND MOIST (SLOPE) DEBRIS IN THE LIŠČAK GORGE 84 FOLIA BIOLOGICA ET GEOLOGICA 64/1 – 2023 Table 12 (Preglednica 12): Lamio orvalae-Lunarietum redivivae Successive number of relevé (Zaporedna številka popisa) 1 2 3 4 5 6 7 8 9 10 11 12 13 Database number of relevé (Delovna številka popisa) 23 84 17 25 84 52 28 71 66 26 29 39 26 29 60 27 38 72 28 52 95 26 32 13 27 43 52 26 29 14 26 83 66 24 27 69 24 81 05 Elevation in m (Nadmorska višina v m) 630 370 450 150 240 430 350 620 440 310 224 225 430 Aspect (Lega) N N SE W E N SE N E N SE N S N E Slope in degrees (Nagib v stopinjah) 40 20 45 5 5 10 30 20 20 5 30 15 30 Parent material (Matična podlaga) L D eb LM C h D eb D eb D eb D eb Ta D eb R o D eb D eb D eb Soil (Tla) Co Co Co Co Co Co Co Co Co Re Co Co Co Stoniness in % (Kamnitost v %) 30 60 20 10 20 80 80 60 40 20 10 20 80 Cover of shrub layer in % (Zastiranje grmovne plasti v %): 20 5 . 10 10 . . . . . . . . Cover of herb layer in % (Zastiranje zeliščne plasti v %): 90 60 70 80 70 70 70 70 50 80 70 80 80 Cover of moss layer in % (Zastiranje mahovne plasti v %): 10 . 20 . 10 20 20 30 20 . 40 . 30 Number of species (Število vrst) 19 12 13 24 21 30 31 28 20 9 19 24 14 Relevé area (Velikost popisne ploskve) m2 30 20 30 20 10 20 100 20 20 10 10 20 20 Date of taking relevé (Datum popisa) 8/ 9/ 20 10 4/ 9/ 20 15 8/ 11 /2 02 1 10 /1 1/ 20 16 10 /1 1/ 20 16 7/ 23 /2 01 8 4/ 9/ 20 21 9/ 22 /2 01 6 4/ 20 /2 01 8 10 /1 3/ 20 16 8/ 22 /2 01 7 4/ 8/ 20 12 5/ 1/ 20 13 Locality (Nahajališče) M rz li po to k A vš če k Li šč ak -S op ot D ob al re c D ob al re c M at ev že va g ra pa - Id ri ja Li šč ak Ž ag a- Su še c St ru žn ik ar je va gr ap a D ob al re c Ši ro ka d ra ga -B ač a Z ad la šč ic a- M ed ve do va g la va Pr ad ol Quadrant (Kvadrant) 97 48 /3 99 48 /1 98 49 /1 98 48 /3 98 48 /3 98 47 /2 98 49 /1 97 46 /2 99 49 /1 98 48 /3 98 49 /1 97 48 /3 97 46 /4 Coordinate GK Y (D-48) m 39 79 61 40 07 82 41 12 77 39 87 88 39 86 98 39 64 57 41 08 35 38 32 89 41 47 39 39 86 06 40 99 23 40 35 14 38 02 42 Coordinate GK X (D-48) m 51 21 40 0 51 03 36 6 51 17 63 2 51 09 34 7 51 10 31 8 51 13 54 0 51 16 99 0 51 28 20 9 51 04 72 4 51 11 02 3 51 13 28 4 51 18 09 1 51 21 26 6 Diagnostic species of the associations (Diagnostične vrste asociacij) TA Lunaria rediviva E1 4 4 4 4 4 3 4 4 3 4 4 4 4 FS Galeobdolon flavidum E1 2 1 1 . 1 2 2 + . . 1 . . FS Sambucus nigra E2a + + + r + + . + + . . + + TA Phyllitis scolopendrium E1 + 1 + 2 3 2 2 1 + . 2 1 1 AF Lamium orvala E1 . . . . . . + + + . . + . FS Cardamine pentaphyllos E1 . 2 . 1 + 2 2 1 . . . + 2 ML Plagiomnium undulatum E0 1 . . + . 1 + 3 + . . . 1 ML Thamnobryum alopecurum E0 . . . . . 1 1 + 1 2 3 1 1 MuADoronicum austriacum E1 . . . . . . . . . . . . . MuAAdenostyles alliariae E1 . . . . . . . . . . . . . MuACicerbita alpina E1 . . . . . . . . . . . . . MuASaxifraga rotundifolia E1 . . . . . . . + . . . . + ArP Arunco-Petasition albi, Petasito-Chaerophylletalia Senecio ovatus E1 . . . + . . . + + . 1 . . Chaerophyllum hirsutum E1 + . . . . . . 1 . . . . . Athyrium filix-femina E1 . . . . . . . r + . . . . Silene dioica E1 . . . . . . . . . . . . . Milium effusum E1 . . . . . . . . . . . . . MuA Mulgedio-Aconitetea Aconitum degenii subsp. paniculatum E1 . . . . . . . . . . . . . Carduus personata E1 . . . . . . . . . . . . . Myrrhis odorata E1 . . . . . . . . . . . . . DAKSKOBLER & MARTINČIČ: VEGETATION OF MOIST ROCK CREVICES AND MOIST (SLOPE) DEBRIS IN THE LIŠČAK GORGE 85FOLIA BIOLOGICA ET GEOLOGICA 64/1 – 2023 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 31 32 33 34 35 36 37 38 39 40 24 81 09 27 01 53 24 79 04 27 82 46 24 27 28 27 23 78 24 86 42 28 10 91 25 50 12 25 50 15 28 15 20 28 27 64 28 27 68 26 77 45 27 23 81 28 08 90 28 52 96 28 71 79 24 61 14 26 22 32 25 82 23 27 33 64 27 43 42 27 46 99 26 46 84 26 95 90 24 81 15 475 200 280 105 225 527 410 575 240 270 470 570 555 390 500 500 350 350 10301050 815 463 385 440 450 1350 470 0 N S N N E W SW 0 SE E SW SW N N E N W N N W S S N W SE S SW E N W N E 0 N W N 0 0 25 5 10 20 10 0 5 25 40 35 60 50 45 60 10 45 60 20 30 5 5 40 0 10 40 0 D eb D eb D eb D eb D eb D eb G r D D eb D eb Ta D eb D eb D eb LC h D eb LM C h LM C h L D eb G r G r D C hC l G r D eb L D eb Co Co Co Re Co Co Co Co Co Co Co Co Co Co Co Co Co Co Re Co Co Co Eu Co Co Re Li 60 80 100 20 30 10 20 30 40 80 10 20 20 90 100 20 30 20 70 10 10 20 5 10 30 10 30 . . . . 2 . . . 5 5 . . . 20 . . . . . . . . . . . . . 80 70 70 100 70 95 95 100 70 80 80 80 80 70 50 70 70 80 70 100 100 80 70 60 70 100 70 20 40 50 . 30 . . . . . . . . . 30 20 20 30 10 . . 15 . 10 . . 90 20 20 16 26 18 36 18 19 16 21 24 22 23 17 19 33 18 10 23 17 37 45 18 11 26 16 39 20 20 10 30 30 50 5 10 20 20 10 30 30 30 30 50 20 20 30 20 30 30 20 15 10 20 100 5/ 1/ 20 13 3/ 20 /2 01 7 4/ 18 /2 01 3 4/ 15 /2 01 9 4/ 1/ 20 12 8/ 17 /2 01 8 6/ 12 /2 01 3 5/ 21 /2 02 0 4/ 7/ 20 14 4/ 7/ 20 14 3, 6, 20 20 6/ 22 /2 02 0 6/ 22 /2 02 0 7/ 13 /2 01 7 8/ 17 /2 01 8 5/ 4/ 20 20 4/ 9/ 20 21 6/ 18 /2 02 1 5/ 18 /2 01 2 7/ 20 /2 01 6 6/ 5/ 20 15 8/ 10 /2 01 8 4/ 20 /2 01 8 3/ 30 /2 01 9 7/ 6/ 20 16 6/ 8/ 20 17 7/ 16 /2 00 1 Pr ad ol Z el p ot ok R aš a- G lo bo ča k Br an ik -P et ni k Lj ub in j-P ot oč ni ca Se vš čk ar -S op ot a K ob ile V in tg ar Br an ic a- Tr eb iž an i Br an ic a- Tr eb iž an i K oz je k V in tg ar V in tg ar V ol ar ja -M rz lic a Se vš čk ar -S op ot a K oz je k- V rš e Li šč ak Li šč ak M ar ni v rh K ac en po h- Sk al a Po kl ju šk a so te sk a Pr od ar je va g ra pa St ru žn ik Pr od ar je va g ra pa V in tg ar Č rn a go ra -p l. za Li sc em Pr ad ol 97 46 /4 99 47 /1 02 49 /2 01 48 /2 98 48 /2 98 48 /2 02 58 /1 96 50 /2 01 49 /3 01 49 /3 99 48 /2 96 50 /2 96 50 /2 97 48 /3 98 48 /2 99 48 /2 98 49 /1 98 49 /1 00 49 /4 97 49 /4 96 50 /1 98 49 /2 99 49 /1 98 49 /2 98 48 /1 97 49 /4 97 46 /4 38 06 78 38 88 38 41 58 05 40 59 86 40 45 69 40 79 83 52 77 83 43 00 75 41 56 67 41 57 40 40 84 83 43 02 73 43 04 47 39 74 17 40 79 66 40 84 72 41 08 43 41 08 94 41 88 14 41 99 50 42 58 54 41 87 83 41 47 38 41 87 02 39 70 70 41 85 03 38 03 06 51 21 04 0 51 05 27 0 50 70 84 4 50 79 49 4 51 14 53 5 51 14 98 0 50 71 50 8 51 39 33 5 50 74 34 3 50 74 16 6 51 05 40 6 51 39 58 8 51 39 73 3 51 20 48 8 51 14 92 6 51 05 41 3 51 16 98 2 51 17 04 7 50 88 99 1 51 21 56 8 51 37 30 6 51 17 54 5 51 04 90 4 51 17 62 9 51 13 24 4 51 22 49 1 51 21 26 6 Pr. Fr. 4 4 3 4 4 5 4 3 3 4 3 3 4 2 1 + 1 + 4 4 3 3 4 2 + 3 1 40 100 . 1 . . . . . 1 + 3 1 1 1 . 2 + 1 2 + + + + . . 1 . . 24 60 . . + . + + . . + + . + + 2 1 + . . . . + + + . + . . 24 60 + 3 . 1 + 1 . . . . . 2 1 + 2 . 1 1 . . . 1 . . . . . 24 60 + . 3 3 + + . . 4 + 3 3 1 + 3 3 3 4 . . . 1 + . 1 . . 22 55 2 1 . . . . . 2 . . 2 . . . . 1 . . 1 . 1 2 . . 1 . . 17 43 2 3 . . . . . . . . . . . . 2 3 . . . . . 1 . 1 . . 1 14 35 2 1 + 1 . . . . . . . . . . . . + . . . . + . . . . . 14 35 . . . . . . . . . . . . . . . . . . . . . . 1 . . 3 . 2 5 . . . . . . . . . . . . . . . . . . . . . . . . . 3 . 1 3 . . . . . . . . . . . . . . . . . . . . . . . . . + . 1 3 . . . . . . . . . . . + . . . 1 . . . . 1 . . . . . 4 6 15 . . . . . + . 1 . . + 1 1 1 . + . . . + 1 2 . . + . . 15 38 . . . . . + 2 1 . . . + 1 . . . . . . . 1 . . . . . . 8 20 + . . . . 1 . . . . . . . + . . . . + . 2 . + . . 2 + 10 25 . . . . . . + . . . . . . . . . . . + . + . . . . . . 3 8 . . . . . . . 2 . . . . . . . . . . . . + . . . . . . 2 5 . . . . . . . . . . 1 . + . . . . . . . . . . . . + . 3 8 . . . . . . . . . . . . . . . . . . . + + . . . . . . 2 5 . . . . . . . . . . . . . . . . . . . + 2 . . . . 1 . 3 8 DAKSKOBLER & MARTINČIČ: VEGETATION OF MOIST ROCK CREVICES AND MOIST (SLOPE) DEBRIS IN THE LIŠČAK GORGE 86 FOLIA BIOLOGICA ET GEOLOGICA 64/1 – 2023 Geum rivale E1 . . . . . . . . . . . . . Stellaria nemorum E1 . . . . . . . . . . . . . Aconitum lycoctonum E1 . . . . . . . . . . . + . Veratrum album E1 . . . . . . . . . . . . . Rumex alpinus E1 . . . . . . . . . . . . . Petasites hybridus E1 . . . . . . . . . . . . . Senecio cacaliaster E1 . . . . . . . . . . . . . Aconitum angustifolium E1 . . . . . . . . . . . . . TG Trifolio-Geranietea Campanula rapunculoides E1 . . . . . . . . . . . . . Clinopodium vulgare E1 . . . . . . . . . . . . . Vincetoxicum hirundinaria E1 . . . . . . . . . . . . . EA Epilobietea angustifolii Impatiens parviflora E1 . . . . . . . . . . + . . Eupatorium cannabinum E1 . . . . . . . . . . . . . Galeopsis speciosa E1 + . . . . 1 . . . . . . . Solanum dulcamara E1 + . . . . . . . . . . . . Rubus idaeus E2a . . . . . . . . . . . . . Fragaria vesca E1 . . . . . . . . . . + . . Barbarea vulgaris E1 . . . . . . . . . . . . . Arctium nemorosum E1 . . . . . . . . . . . . . Galeopsis pubescens E1 . . . . . . . . . . . . . Stachys sylvatica E1 . . . . . . . . . . . . . MC Montio-Cardaminetea Conocephalum conicum E0 + . + 1 1 . . + 1 . 1 . . Cratoneuron filicinum E0 . . + . . . . . . . . . . Oxyrrhynchium hians E0 . . + . . . . . . . . . . ES Elyno-Seslerietea Carex ferruginea E1 . . . . . . . . . . . . . Sesleria caerulea E1 . . . . . . . . . . . . . Mo Molinion Caltha palustris E1 . . . . . 1 . . . . . . + Angelica sylvestris E1 . . . . . + + . . . . . . Cirsium oleraceum E1 . . . . . . . . . . . . . Filipendula ulmaria E1 . . . . . . . . . . . . . MA Molinio-Arrhenatheretea Anthriscus sylvestris E1 . . . 1 . . . . . . . . . Geranium phaeum E1 . . . . . . . . . . . . . GU Galio-Urticetea, Stellarietea mediae Urtica dioica E1 . . . . . . . . . . . 1 1 Geum urbanum E1 . . . . . . . . . . + . . Glechoma hederacea E1 . . . . . . . . . . . . . Lamium maculatum E1 . . . . . . . . . + . . . Parietaria officinalis E1 . . . + . . . . . . . . . Erigeron annuus E1 . . . . . . . . . . . . . Impatiens glandulifera E1 . . . . . . . . . . + . . Veronica chamaedrys E1 . . . . . . . . . . . . . Stellaria media E1 . . . . . . . . . . . . . TR Thlaspietea rotundifolii Adenostyles glabra E1 . . . . . . . . + . . . . Gymnocarpium dryopteris E1 . . . . . . . . . . . . . Gymnocarpium robertianum E1 . . . . . . . . . . . . . Arabis alpina E1 . . . . . . . . . . . . . AP Astrantio-Paederotion luteae Cystopteris fragilis E1 + . . . + . . + . . . . . Fissidens dubius E0 . . . . . + . + . . . . . Palustriella commutata E0 . . 1 . . . . . . . . . . Astrantia carniolica E1 . . . . . . . . . . . . . Valeriana tripteris E1 . . . . . . . . . . . . . Apopellia endiviifolia (Pellia endiviifolia) E0 . . . + . . . . . . . . . Paederota lutea E1 . . . . . . . . . . . . . Orthothecium rufescens E0 . . . . . . . . . . . . . Viola biflora E1 . . . . . . . . . . . . . Successive number of relevé (Zaporedna številka popisa) 1 2 3 4 5 6 7 8 9 10 11 12 13 DAKSKOBLER & MARTINČIČ: VEGETATION OF MOIST ROCK CREVICES AND MOIST (SLOPE) DEBRIS IN THE LIŠČAK GORGE 87FOLIA BIOLOGICA ET GEOLOGICA 64/1 – 2023 . . . . . . . . . . . + . . . . . . . . 1 . . . . . . 2 5 . . . . . . . . . . . . . . . . . . . . 1 . . . . + . 2 5 . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 3 . . . . . . + . . . . . . . . . . . . . . . . . . . . 1 3 . . . . . . . . . . . . . . . . . . . . + . . . . . . 1 3 . . . . . . . . . . . . . . . . . . . . . . . + . . . 1 3 . . . . . . . . . . . . . . . . . . . . . . . . . 1 . 1 3 . . . . . . . . . . . . . . . . . . . . . . . . . . + 1 3 . . . . . . . . . . . . + . . . . . . . . . . . . . . 1 3 . . . . . . . . . . . + . . . . . . . . . . . . . . . 1 3 . . . . . . . . . . + . . . . . . . . . . . . . . . . 1 3 . . . . . 1 . . . . . . + . . . . . . . . . . . . . . 3 8 . . . . . . 1 . . . + . . . . . . . . + . . . . . . . 3 8 . . . . . . . . . . . . . . . . . . . . . . . . . . . 2 5 . . . . . . . . . . . . . . . . . . . + . . . . . . . 2 5 . . . . . . . . . . . . . . . . . . . + . . . . . + . 2 5 . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 3 . . . . . + . . . . . . . . . . . . . . . . . . . . . 1 3 . . . . . . + . . . . . . . . . . . . . . . . . . . . 1 3 . . . . . . . . . . . . . . . . . . . . . 1 . . . . . 1 3 . . . . . . . . . . . . . . . . . . . . 1 . . . . . . 1 3 . . . . . . . . . . . 1 . . . 1 + 2 + . . . . . + . + 14 35 . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 3 . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 3 . . . . . . . . . . + . . . . + . . . . . . . . . . . 2 5 . . . . . . . . . . . . . . . . . . . . . . . . . . + 1 3 + . . . . + . . . . . . . . . 1 . . . . + . . + . . . 7 18 . . . . . + . . + . . . 1 . . . . . . . . + . . . . . 6 15 . . . . . . 1 . . . . . . . . . . . . . + . . . . . . 2 5 . . . . . . . . . . . . . . . . . . . . 1 . . . . . . 1 3 . . . . . . . . . . . . . . . . . . . . . . . 2 . . . 2 5 . . . . . . 1 2 . . . . . . . . . . . . . . . . . . . 2 5 + . . . . . . . . . . . . + . . . . 2 3 2 2 . . . 1 + 10 25 . . . . . . . + . . . . . . . . . . . . . . . . . . . 2 5 . . . . . 2 . . . . . . . . 1 . . . . . . . . . . . . 2 5 . . . . . . . . . . . . . . . . . . . . 1 . . . . . . 2 5 . . . . . . . . . . . . + . . . . . . . . . . . . . . 2 5 . . . . . . . + . . . . . . . . . . . . . . . . . . . 1 3 . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 3 . . . . . . . . . + . . . . . . . . . . . . . . . . . 1 3 . . . . . + . . . . . . . . . . . . . . . . . . . . . 1 3 . . . . . . . . . . 1 1 . 2 . + . . . . . . . . . . + 6 15 . . . . . . . . . . . . . . . . . . 1 . . . . . . . + 2 5 . . . . . . . . . . + . . . . . . . . . . . . . . . . 1 3 . . . . . . . . . . . . . . . . . . . . . . . . . . 1 1 3 . . . . . . . . . . . . . . + . . . . . . . . . . . 1 5 13 . . . . . . . . . . . . . . . + . 1 . . . + . . . . . 5 13 . . . . . . . . . . . . . . . + + 2 . . . . . . . . . 4 10 . . . . . . . . . . + . . . . + . . . . . . . . . . . 2 5 . . . . . . . . . . . + . . . . . . . . . . . . . . + 2 5 . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 3 . . . . . . . . . . . . . . . + . . . . . . . . . . . 1 3 . . . . . . . . . . . . . . . . . . . . . + . . . . . 1 3 . . . . . . . . . . . . . . . . . . . . . . . . . . 1 1 3 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 31 32 33 34 35 36 37 38 39 40 Pr. Fr. DAKSKOBLER & MARTINČIČ: VEGETATION OF MOIST ROCK CREVICES AND MOIST (SLOPE) DEBRIS IN THE LIŠČAK GORGE 88 FOLIA BIOLOGICA ET GEOLOGICA 64/1 – 2023 AT Asplenietea trichomanis Asplenium trichomanes E1 . . . + + + + 1 . . . . . Polypodium vulgare E1 . . . . . + . + + . . . + Saxifraga petraea E1 . . . + . . . . . r . . . AI Alnion incanae Chrysosplenium alternifolium E1 . . . + . . . . . . . . . Cardamine impatiens E1 . . + . . . . . + . . . . Impatiens noli-tangere E1 . . . . . . . . . . . . . Festuca gigantea E1 . . . . . . . . . . . . . Rubus caesius E1 . . . . . . . . . . . . . Aesculus hippocastanum E2a . . . . . . . . . . . . . Matteuccia struthiopteris E1 . . . . . . . . . . . . . TA Tilio-Acerion Geranium robertianum E1 . . . 1 1 1 + . 1 + + . . Stellaria montana E1 . . . . . + 1 + + + . . . Aruncus dioicus E1 + . + . + . . . . . . . . Polystichum setiferum E1 . + 1 . + . + . 1 . + . . Adoxa moschatellina E1 . 1 . . . . + . . . . 1 . Acer pseudoplatanus E2a + . . . + . . + . . . . . Acer pseudoplatanus E1 . . . . . . + . . . . . . Ulmus glabra E1 . r . + + . + . . . . . . Arum maculatum E1 . + . . . . . . . . . 1 1 Polystichum aculeatum E1 . . . . . . + + . . . . . Thalictrum aquilegiifolium E1 . . . . . . . . . . . . + Dryopteris affinis E1 . . . . . + . . . . + . . Tephroseris pseudocrispa E1 . . . . . . . . . . . + . Isopyrum thalictroides E1 . . . . . . . . . . . . . Circaea x intermedia E1 . . . . . . . . . . . . . Polystichum braunii E1 . . . . . . . . . . . . . Anthriscus nitida E1 . . . . . . . . . . . . . Polystichum x bicknellii E1 . . . . . + . . . . . . . Corydalis solida E1 . . . . . . . . . . . . . Dryopteris remota E1 . . . . . . . . . . . . . Acer platanoides E2a . . . . . . . . . . . . . Hesperis candida E1 . . . . . . . . . . . . . Tephroseris longifolia E1 . . . . . . . . . . . . . Cardamine flexuosa E1 . . . . . . . . . . . . . Veronica montana E1 . . . . . . . . . . . . . EC Erythronio-Carpinion Galanthus nivalis E1 . . . . . . + . . . . 2 . Helleborus odorus E1 . . . . . . . . . . . . . Ornithogalum pyrenaicum E1 . . . . . . . . . . . . . AF Aremonio-Fagion Cardamine trifolia E1 3 . . 1 . + 1 . + . . . . Cardamine enneaphyllos E1 . . . . . . . . + . . . . Anemone trifolia E1 . + . . . 1 + . . . . + . Cyclamen purpurascens E1 . . . . + . . . . . . . . Geranium nodosum E1 . . . . + . . . . + . + . Scopolia carniolica E1 . . . . . . . . . . 2 . . Euphorbia carniolica E1 . . . . . . . . . . . . . Daphne laureola E2a . . . + . . . . . . . . . Hacquetia epipactis E1 . . . . . . . . . . . . . Epimedium alpinum E1 . . . . . . . . . . . . . Vicia oroboides E1 . . . . . . . . . . . . . Omphalodes verna E1 . . . . . . . . . . . . . FS Fagetalia sylvaticae Salvia glutinosa E1 . . . . . . . 1 . . + . . Dryopteris filix-mas E1 + . . . . 1 . . + . . . . Asarum europaeum subsp. caucasicum E1 + . . . . . + . . + + 1 . Brachypodium sylvaticum E1 . . + + . . + . . . + . . Corydalis cava E1 . . . . . . . . . . . 2 3 Mycelis muralis E1 . + . . . + . + . . . . . Successive number of relevé (Zaporedna številka popisa) 1 2 3 4 5 6 7 8 9 10 11 12 13 DAKSKOBLER & MARTINČIČ: VEGETATION OF MOIST ROCK CREVICES AND MOIST (SLOPE) DEBRIS IN THE LIŠČAK GORGE 89FOLIA BIOLOGICA ET GEOLOGICA 64/1 – 2023 . . . . . . . . + . + . . . . . . . . . . . . . . . . 7 18 . . . . . . . . . + . . . . . . . . . . . . . . . . + 6 15 . . . . . . . . . . . . . . . . . . . . . . . . . . . 2 5 + 1 . . . . . . . . . . . . . . . . + . + + 1 3 . . 1 9 23 . . . + . + . . + + . . . . . . . . . . . + . . . . . 7 18 + . . . . . . . . . . . . . . . . . . . 2 2 . . . 2 . 4 10 . . . . . + . . . . . . . . . . . . . . . . . . . . . 1 3 . . . . . . + . . . . . . . . . . . . . . . . . . . . 1 3 . . . . . . . . + . . . . . . . . . . . . . . . . . . 1 3 . . . . . . . . . . . . . . . . . . . . 2 . . . . . . 1 3 . + . . + 1 . + . . 1 1 1 + 2 + . . . 1 1 2 . . . . + 21 53 . 1 . + . . 1 . . . . . . . . . . . + . . 1 + 1 + . . 13 33 . . . . . . + . + + . + 2 . . . + . . . + + . . . . . 11 28 . + . . 2 . . . . . . . . 2 . . + 1 . . . . . . . . . 11 28 1 . . . + + . . . . . . . . + . . . + . . + . . . . 1 10 25 . . . . . . . . . 1 . . . . . . . . . . . . . . . . . 4 10 . . . + . . . . 1 + . . . . . + . . 1 . + . . . + . + 9 23 . . . . . + . . . . . . . . . + . . . . . . . . + . . 7 18 + + . . + + . . . . . . . . . . . . . . . . . . . . . 7 18 . . . . . . . . . . + . + . . + . . . . . + + . . . . 7 18 . . . . . . + . . . + + . . . . . . . . . . . . . . . 4 10 . . . . . . . . . + . . . . . . . . . . + . . . . . . 4 10 . . . . . . . . . . . . . . . . . . . . . + . . . . . 2 5 . . + + . . . . . . . . . . . . . . . . . . . . . . . 2 5 . . . . . . . . . . . . . . . . . . . . + + . . . . . 2 5 . . . . . . . . . . . . . . . . . . + . . . + . . . . 2 5 . . . . . . . . . . . . . . . . . . . . . 2 . . . + . 2 5 . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 3 + . . . . . . . . . . . . . . . . . . . . . . . . . . 1 3 + . . . . . . . . . . . . . . . . . . . . . . . . . . 1 3 . . . . + . . . . . . . . . . . . . . . . . . . . . . 1 3 . . . . . . 1 . . . . . . . . . . . . . . . . . . . . 1 3 . . . . . . . . . . . . . . . + . . . . . . . . . . . 1 3 . . . . . . . . . . . . . . . . . . . . + . . . . . . 1 3 . . . . . . . . . . . . . . . . . . . . + . . . . . . 1 3 . + + 1 + + . . . . . . . . + . . . . . . . . . . . . 8 20 . . . . . + . . . . . . . + . . . . . . . . . . + . . 3 8 . . . + . . . . . . . . . . . . . . . . . . . . . . . 1 3 . 1 . . . . . . + . . . . . . + 1 1 . . . . . . . . + 11 28 + 1 2 + 1 . + . . . . . . . . 1 . . . + . . . . . . + 10 25 . . . . . . . . . . + . . . . . . . . . . . . . + . . 6 15 . . . . . + . . . . . . . + . . . . . + . . . . + . . 5 13 . . . . . . . . . . . . . . . . . . . . . . . . 2 . . 4 10 . . . . . . . . . . . . . . . . . . . . . . 3 . . . . 2 5 . . . . . . . . . . + . . . . + . . . . . . . . . . . 2 5 . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 3 . . . . . r . . . . . . . . . . . . . . . . . . . . . 1 3 . . . . . . 1 . . . . . . . . . . . . . . . . . . . . 1 3 . . . . . . . + . . . . . . . . . . . . . . . . . . . 1 3 . . . . . . . . . . . . . . . . . . . . . . . . 1 . . 1 3 . . . + . 1 . 1 + 1 3 1 1 3 1 + + . . . + 1 . . . . . 16 40 . . . . . . . . . . . . . . . . . . 1 2 1 + + . + 2 + 11 28 . . + . . . . . . . 1 . . . . + + . . . . . . . + . . 10 25 . . . . . + . . . . . . + + + . . . . + . + . . . . . 10 25 3 2 . 3 2 . . . . . . . . . . . . . . . . . . . . 1 1 8 20 . . . + . . . . . . . . . . . + + . . + . + . . . . . 8 20 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 31 32 33 34 35 36 37 38 39 40 Pr. Fr. DAKSKOBLER & MARTINČIČ: VEGETATION OF MOIST ROCK CREVICES AND MOIST (SLOPE) DEBRIS IN THE LIŠČAK GORGE 90 FOLIA BIOLOGICA ET GEOLOGICA 64/1 – 2023 Fraxinus excelsior E2a 1 . . . + . . . . . . . . Petasites albus E1 . . . . . . + . . . . . . Dentaria bulbifera E1 . . . . . . + . . . . . . Circaea lutetiana E1 . . . . . . . r . . . . . Symphytum tuberosum E1 . . . . . . . . . . . . . Mercurialis perennis E1 . . . . . . . + . . . 1 . Allium ursinum E1 . . . . . . . . . . . . . Pulmonaria officinalis E1 . . . . . . . . . . . + . Fagus sylvatica E3 . . . . . . . . . . . . . Fagus sylvatica E1 . . . . . . + . . . . . . Festuca altissima E1 . . . . 1 . . . . . . . . Galium laevigatum E1 . . . . . . + . . . . . . Ranunculus lanuginosus E1 . . . . . . . . . . . . . Actaea spicata E1 + . . . . . . . . . . . . Myosotis sylvatica E1 . . . . . . . . . . . . . Phyteuma spicatum subsp. coeruleum E1 . . . . . . . . . . . . . Epilobium montanum E1 . . . . . . . . . . . . . Scrophularia nodosa E1 . . . . . . . . . . . . . Tilia cordata E1 . . . r . . . . . . . . . Lathyrus vernus E1 . . . . . . . . . . . + . Leucojum vernum E1 . . . . . . . . . . . . . Galeobdolon montanum E1 . . . . . . . . . . . . . Carex sylvatica E1 . . . . . . . . . . . . . Euphorbia dulcis E1 . . . . . . . . . . . . . Polygonatum multiflorum E1 . . . . . . . . . . . . . Galium odoratum E1 . . . . . . . . . . . . . Luzula nivea E1 . . . . . . . . . . . . . QP Quercetalia pubescenti-petraeae Ruscus aculeatus E1 . . . 1 + . . . . . . . . Arabis turrita E1 . . . . . . . . . . . . . Inula conyza E1 . . . . . . . . . . . . . QP Querco-Fagetea Hedera helix E1 . . . . + + + . . . + + . Aegopodium podagraria E1 . . . + . 1 . . . . + + . Clematis vitalba E1 . . . . . . + . . . . . . Ficaria verna E1 . . . . . . . . . . . . . Anemone nemorosa E1 . . . . . 1 . . . . . . . Carex digitata E1 . . . . . + . . . + . . . Corylus avellana E2a . . . r . . . . . . . . . Lathraea squamaria E1 . . . . . . . . . . . + . Moehringia trinervia E1 . . . . . . . . . . . . . Dactylis polygama E1 . . . r . . . . . . . . . Hepatica nobilis E1 . . . . . + . . . . . . . Paris quadrifolia E1 . . . . . . . . . . . . . Acer campestre E2a . . . . . . . . . . . . . Acer campestre E1 . . . . . . . . . . . . . Anemone ranunculoides E1 . . . . . . . . . . . . . Cerastium sylvaticum E1 . . . . . . . . . . . . . Gagea lutea E1 . . . . . . . . . . . . . Vinca minor E1 . . . . . . . . . . . . . Viola riviniana E1 . . . . . . . . . . . . . Rubus hirtus E1 . . . . . . . . . . . . . Veratrum nigrum E1 . . . . . . . . . . . . . RP Rhamno-Prunetea Cornus sanguinea E2a . . . . . . . . . . . . . Rubus fruticosus agg. E1 . . . . . . . . . . . . . EP Erico-Pinetea Cirsium erisithales E1 . . . . . . . . . . . . . Calamagrostis varia E1 . . . . . . . . . . . . . Molinia arundinacea E1 . . . . . . . . . . . . . Vaccinio-Piceetea Oxalis acetosella E1 + + . 1 . 1 + + + . . . . Successive number of relevé (Zaporedna številka popisa) 1 2 3 4 5 6 7 8 9 10 11 12 13 DAKSKOBLER & MARTINČIČ: VEGETATION OF MOIST ROCK CREVICES AND MOIST (SLOPE) DEBRIS IN THE LIŠČAK GORGE 91FOLIA BIOLOGICA ET GEOLOGICA 64/1 – 2023 . . . . + + . 1 . . . . . . . 1 . . . . . + . . + . . 8 20 . . . . . . . . . . . . 2 2 . . 1 1 . . . 1 + . + . . 8 20 + . . 1 + . . . 1 1 . . . . . . . . . . . . + . . . . 7 18 . . . . . . . . + 1 . . . . . . . . 1 . . 1 . . + . . 6 15 . . . . . . . . . + + . . . . . + . . . + + + . . . . 6 15 . . 1 . . . . 1 . . . . . . . . . . . . . . . . 1 . . 5 13 . + . . 3 4 3 . . . . . . . + . . . . . . . . . . . . 5 13 . . . + . + . . . . . . . . . . . . . . . . . . + . . 4 10 . . . . . . . . . . . . . . . . . . . . . . . . . . r 1 3 . . . . . . . . . . . . . . . . + . 1 . . . . . . . . 3 8 . . . . . . . . . . . 4 2 . . . . . . . . . . . . . . 3 8 . . . . . . . . . . . . . + . . . . . + . . . . . . . 3 8 . . . . . . . 1 . . . . + . . . . . . . 1 . . . . . . 3 8 . . . . . . . . . . . . . . . . . . . . . . + . . . . 2 5 . . . . . . . + . . . . . . . . . . . . + . . . . . . 2 5 . . . . . . . . . . 1 . . . . + . . . . . . . . . . . 2 5 . . . . . . . . . . . + . . . . . . + . . . . . . . . 2 5 . . . . . . . . . . . . + . . . . . + . . . . . . . . 2 5 . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 3 . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 3 + . . . . . . . . . . . . . . . . . . . . . . . . . . 1 3 . . . + . . . . . . . . . . . . . . . . . . . . . . . 1 3 . . . . . . + . . . . . . . . . . . . . . . . . . . . 1 3 . . . . . . . . . + . . . . . . . . . . . . . . . . . 1 3 . . . . . . . . . + . . . . . . . . . . . . . . . . . 1 3 . . . . . . . . . . . . . . . . . . . . . . + . . . . 1 3 . . . . . . . . . . . . . . . . . . . . . . . . . . + 1 3 . . . + . . . . . . . . . . . . . . . . . . . . . . . 3 8 . . . . . . . . . + . . . . . . . . . . . . . . . . . 1 3 . . . . . . . . . . . . + . . . . . . . . . . . . . . 1 3 . + . + 1 r . . . . . . . . 1 . . . . . . + . . . . . 11 28 . + . 1 . 2 . 1 . . . . . . + . . . . . . . . 1 . . . 10 25 . . . . . . . . . . . + . . + + . . . . . + . . + . . 6 15 . . . 1 1 1 . . + . . . . . . . . . . . . . . 1 . . . 5 13 . . + . . . . . . . . . . . . . . . . . . . . 1 . . . 3 8 . . . . . . . . . . . . . . . . + . . . . . . . . . . 3 8 . . . + . . . . . . . . . . . . . . . . . . . . . . . 2 5 . . + . . . . . . . . . . . . . . . . . . . . . . . . 2 5 . . . . . . . . . + . . . . . . . . . . . . + . . . . 2 5 . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 3 . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 3 + . . . . . . . . . . . . . . . . . . . . . . . . . . 1 3 . . . . . . . . + . . . . . . . . . . . . . . . . . . 1 3 . . . + . . . . . . . . . . . . . . . . . . . . . . . 1 3 . . . . + . . . . . . . . . . . . . . . . . . . . . . 1 3 . . . . . + . . . . . . . . . . . . . . . . . . . . . 1 3 . . . . . 1 . . . . . . . . . . . . . . . . . . . . . 1 3 . . . . . + . . . . . . . . . . . . . . . . . . . . . 1 3 . . . . . . . . . + . . . . . . . . . . . . . . . . . 1 3 . . . . . . . . . + . . . . . . . . . . . . . . . . . 1 3 . . . . . . . . . . + . . . . . . . . . . . . . . . . 1 3 . . . . . + . . . . . . . . . . . . . . . . . . . . . 1 3 . . . . . . + . . . . . . . . . . . . . . . . . . . . 1 3 . . . . . . . 1 . . + + . . . . . . . . . . . . . . . 3 8 . . . . . . . . . . . . . 1 . . . . . . . . . . . . . 1 3 . . . . . . . . . . . . . . . . . . . + . . . . . . . 1 3 . + . + . + . . . + . . . . + 1 . . + . . 1 . . 1 . 1 17 43 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 31 32 33 34 35 36 37 38 39 40 Pr. Fr. DAKSKOBLER & MARTINČIČ: VEGETATION OF MOIST ROCK CREVICES AND MOIST (SLOPE) DEBRIS IN THE LIŠČAK GORGE 92 FOLIA BIOLOGICA ET GEOLOGICA 64/1 – 2023 Veronica urticifolia E1 . . . . + . + . . . . . . Saxifraga cuneifolia E1 . . . . . + . . . . . . . Aposeris foetida E1 . . . . . . . . . . . . . Calamagrostis arundinacea E1 + . . . . . . . . . . . . Phegopteris connectilis E1 . . . . . . . + . . . . . Dryopteris dilatata E1 . . . . . . . . . . . . . Luzula luzuloides E1 . . . . . . . . . . . . . Clematis alpina E1 . . . . . . . . . . . . . ML Mosses (Mahovi) Ctenidium molluscum E0 . . . . + 1 . + 1 . . . . Brachythecium rutabulum E0 . . . . . . 1 . . . . 1 2 Isothecium alopecuroides E0 . . . . . 1 . . . . . + . Exertotheca crispa (Neckera crispa) E0 . . . . . . . + . . . . . Schistidium apocarpum E0 . . . . . . . . . . . . . Eurhynchium zetterstedtii E0 . . . . . . 1 . . . . . . Sciuro-hypnum populeum E0 . . . . . . . . . . . . . Plagiomnium rostratum E0 . . . . . . . . . . . . . Pseudanomodon attenuatus (Anomodon attenuatus) E0 . . . . . . . . . . . . . Didymodon insulanus E0 . . . . . . . . . . . . . Hygrohypnum luridum E0 . . . . . . . . . . . . . Plasteurhynchium striatulum E0 . . . . . . . . . . . . . Rhynchostegium murale E0 . . . . . . . . . . . . . Schistidium crassifolium E0 . . . . . . . . . . . . . Rhytidiadelphus loreus E0 . . . . . . . . . . . . . Hylocomiadelphus triquetrus (Rhytidiadelphus triquetrus) E0 . . . . . . . . . . . . . Hylocomium splendens E0 . . . . . . . . . . . . . Rhizomnium punctatum E0 . . . . . . . . . . . . . Peltigera sp. E0 . . . . . . . . . . . . . Plagiochila porelloides E0 . . . . . . . . . . . . . Sanionia uncinata E0 . . . . . . . . . . . . . Legend - Legenda L Limestone - apnenec Ch Chert - roženec D Dolomite - dolomit Deb Debris - grušč Gr Gravel - prod M Marlstone - laporovec Ro Rockslide - podorno gradivo Ta Talus - vršaj Co Colluvial-deluvial soil - koluvialno-deluvialna tla Eu Eutric brown soil - evtrična rjava tla Re rendzina - rendzina Li Lithosol - kamnišče Pr. Presence (number of relevés in which the species is presented) - število popisov, v katerih se pojavlja vrsta Fr. Frequency in % - frekvenca v % 1-38 Lamio orvalae-Lunarietum redivivae 39 Doronico austriaci-Adenostyletum alliariae lunarietosum redivivae 40 Lunario redivivae-Saxifragetum rotundifoliae nom. prov. Successive number of relevé (Zaporedna številka popisa) 1 2 3 4 5 6 7 8 9 10 11 12 13 DAKSKOBLER & MARTINČIČ: VEGETATION OF MOIST ROCK CREVICES AND MOIST (SLOPE) DEBRIS IN THE LIŠČAK GORGE 93FOLIA BIOLOGICA ET GEOLOGICA 64/1 – 2023 . . . . . . . . . . . + + . . . . . . . + . . . . . + 6 15 . . . . . . . . . . . . . . . . . . . . . . . . . . 2 2 5 . . . . . . . + . . . . . . . 1 . . . . . . . . . . . 2 5 . . . . . . . . . . . . . . . . . . . . . . . . . + . 2 5 . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 3 . . . . . . . . . . . . . . . . . . r . . . . . . . . 1 3 . . . . . . . . . . . . . . . . . . . . . . + . . . . 1 3 . . . . . . . . . . . . . . . . . . . . . . . . . . + 1 3 . . 3 1 . . . . . . . . . . . 2 + . 1 . . 1 . . + . 1 12 30 . 1 1 . . . . . . . . . . . . . . . . . . . . 1 1 . . 7 18 . . 1 + . . . . . . . . . . 2 . . . 1 . . . . . . . . 6 15 . . 1 . . . . . . . . . . . . . . . . . . . . . . . + 3 8 . . + . . . . . . . . . . . . . . . + . . . . . . . + 3 8 . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 3 . . . . . . . . . . . . . . . . . . . . . 1 . . . . . 1 3 . . . . . . . . . . . . . . . . . . . . . 1 . . . . . 1 3 . . . . . . . . . . . . . . . . . . . . . + . . . . . 1 3 . . . . . . . . . . . . . . . . . . . . . + . . . . . 1 3 . . . . . . . . . . . . . . . . . . . . . + . . . . . 1 3 . . . . . . . . . . . . . . . . . . . . . + . . . . . 1 3 . . . . . . . . . . . . . . . . . . . . . + . . . . . 1 3 . . . . . . . . . . . . . . . . . . . . . + . . . . . 1 3 . . . . . . . . . . . . . . . . . . . . . . . . . . 4 1 3 . . . . . . . . . . . . . . . . . . . . . . . . . . 1 1 3 . . . . . . . . . . . . . . . . . . . . . . . . . . 1 1 3 . . . . . . . . . . . . . . . . . . . . . . . . . . 1 1 3 . . . . . . . . . . . . . . . . . . . . . . . . . . + 1 3 . . . . . . . . . . . . . . . . . . . . . . . . . . + 1 3 . . . . . . . . . . . . . . . . . . . . . . . . . . + 1 3 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 31 32 33 34 35 36 37 38 39 40 Pr. Fr. DAKSKOBLER & MARTINČIČ: VEGETATION OF MOIST ROCK CREVICES AND MOIST (SLOPE) DEBRIS IN THE LIŠČAK GORGE 94 FOLIA BIOLOGICA ET GEOLOGICA 64/1 – 2023 Ta bl e 13 (P re gl ed ni ca 1 3) : L am io o rv al ae -S am bu ce tu m n ig ra e l un ar ie to su m re di vi va e Su cc es si ve n um be r of r el ev é (Z ap or ed na š te vi lk a po pi sa ) 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 D at ab as e nu m be r of r el ev é (D el ov na š te vi lk a po pi sa ) 248415 248416 249067 268269 281618 282737 258453 260838 260842 261426 261429 258459 258460 260840 260844 260845 260847 260937 285376 285839 284764 284765 280014 280015 264505 El ev at io n in m (N ad m or sk a vi ši na v m ) 76 0 75 0 76 5 78 0 59 0 57 0 37 0 37 0 37 0 36 0 38 0 36 0 35 0 37 0 54 5 54 5 53 0 55 0 31 0 23 0 26 0 25 5 51 0 50 0 12 5 A sp ec t ( Le ga ) N N W E E N W N W N N E N E N E N W N W N W N N E N N W N E N E N W N E SE SE N E Sl op e in d eg re es (N ag ib v s to pi nj ah ) 35 35 10 15 35 25 25 35 25 25 35 20 30 30 40 30 35 30 15 10 30 30 10 5 10 Pa re nt m at er ia l ( M at ič na p od la ga ) M M Ss L D eb D eb D eb LM LM D eb D eb LM LM LM LM LM LM D eb L D eb D eb D eb R o R o Fl So il (T la ) C C C C Eu C o C o C o C o C C C C C o C o Eu Eu Eu C C C o C C C o C o C o C o C o C o C o Eu St on in es s i n % (K am ni to st v % ) 0 0 0 10 40 30 60 10 10 20 30 5 5 0 10 10 10 5 30 40 70 70 70 50 20 C ov er o f t re ee la ye r in % (Z as ti ra nj e dr ev es ne p la st i v % ): . . . . . . . . . . . . . . . . . 15 . . . . . . 10 C ov er o f s hr ub la ye r in % (Z as ti ra nj e gr m ov ne p la st i v % ): 60 80 80 80 50 50 70 90 80 80 80 60 70 80 80 70 80 60 70 40 70 70 70 70 70 C ov er o f h er b la ye r in % (Z as ti ra nj e ze liš čn e pl as ti v % ): 70 60 50 70 10 0 90 40 70 70 70 70 70 40 70 50 60 60 80 80 70 70 60 60 50 90 C ov er o f m os s l ay er in % (Z as ti ra nj e m ah ov ne p la st i v % ): . . . 10 15 . . 5 10 50 40 . . 10 . 5 5 5 30 20 40 30 20 20 . N um be r of s pe ci es (Š te vi lo v rs t) 25 33 21 44 35 33 23 27 40 40 47 27 28 33 33 27 32 40 25 40 24 20 34 26 41 R el ev é ar ea (V el ik os t p op is ne p lo sk ve ) m 2 10 0 10 0 10 0 10 0 30 50 50 50 50 50 50 10 0 50 50 50 50 50 10 0 60 4 40 40 10 0 10 0 10 0 D at e of ta ki ng r el ev é (D at um p op is a) 5/21/2013 5/21/2013 7/10/2013 9/4/2017 5/21/2020 6/22/2020 4/9/2015 4/4/2016 4/4/2016 4/11/2016 4/11/2016 4/9/2015 4/9/2015 4/4/2016 4/4/2016 4/4/2016 4/4/2016 4/15/2016 4/15/2021 5/13/2021 3/17/2021 3/17/2021 4/13/2020 4/13/2020 4/12/2016 Lo ca lit y (N ah aj al iš če ) Donačka gora Donačka gora Donačka gora Zapoška grapa Vintgar Vintgar Avšček Avšček Avšček Avšček Avšček Avšček Avšček Avšček Avšček Avšček Avšček Žirovnica-Raskovec Postaja-Pekel Slap ob Idrijci-Vrata Svino Svino Zadlaščica-Kašana Zadlaščica-Kašana Korte-Branski bošk Q ua dr an t ( K va dr an t) 9760/1 9760/1 9760/2 9849/2 9650/2 9650/2 9948/1 9948/1 9948/1 9948/1 9948/1 9948/1 9948/1 9948/1 9948/1 9948/1 9948/1 9950/4 9848/4 9848/4 9747/3 9747/3 9748/4 9748/4 0548/1 C oo rd in at e G K Y (D -4 8) m 557450 557498 558289 421229 430213 430228 400794 400763 400842 400483 400490 400794 400790 400815 400728 400669 400710 430471 404710 408441 390120 390097 406223 406147 395989 C oo rd in at e G K X (D -4 8) m 5124660 5124689 5124846 5113779 5139509 5139520 5103353 5103391 5103287 5103648 5103603 5103484 5103563 5103410 5103047 5103056 5103059 5096318 5111479 5107856 5122874 5122892 5120614 5120541 5039051 D ia gn os ti c sp ec ie s of th e as so ci at io n (D ia gn os ti čn e vr st e as oc ia ci je ) Pr . Fr . FS Sa m bu cu s n ig ra E2 b 4 4 4 4 3 3 4 5 5 4 4 2 4 4 4 4 5 3 4 3 4 4 4 4 3 25 10 0 FS Sa m bu cu s n ig ra E2 a + + . . . . . 1 + . + . . . 1 . . + . 1 . . 1 + . 10 40 FS Sa m bu cu s n ig ra E1 . . . . . . + 1 . . . . . . 1 . . . . . . . . . . 3 12 A F La m iu m o rv al a E1 . . . 1 2 2 + 1 1 . + 2 1 + + + + + 1 3 . . 3 2 . 18 72 TA Ph yl lit is sc ol op en dr iu m E1 1 . . 1 2 1 1 1 1 2 2 1 1 . . . . + 1 1 4 3 1 2 . 18 72 TA G er an iu m ro be rt ia nu m E1 + 1 + 1 1 . + . . 1 1 . . . + + + 1 + 1 + . 1 . . 16 64 DAKSKOBLER & MARTINČIČ: VEGETATION OF MOIST ROCK CREVICES AND MOIST (SLOPE) DEBRIS IN THE LIŠČAK GORGE 95FOLIA BIOLOGICA ET GEOLOGICA 64/1 – 2023 Su cc es si ve n um be r of r el ev é (Z ap or ed na š te vi lk a po pi sa ) 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 Pr . Fr . D if fe re nt ia l s pe ci es o f s ub as so ci at io n an d va ri an ts (R az li ko va ln e vr st e su ba so ci ac ije in v ar ia nt ) TA Lu na ri a re di vi va E1 3 2 2 1 4 4 3 2 2 4 3 2 . 1 1 + + 1 2 . 2 1 . . . 20 80 FS C ir ca ea lu te tia na E1 1 1 . 1 + 1 1 + + . + . + . . + . . + . . . . 1 . 13 52 FS C ar da m in e pe nt ap hy llo s E1 . . . . 2 1 1 3 3 3 . 1 . . 1 1 2 2 1 . . . . r . 13 52 G eo gr af sk a ra zl ik ov al na v rs ta A F A ne m on e tr ifo lia E1 . . . . . . + + 1 1 1 + + 1 + + . . + + + + + . . 15 60 R az li ko va ln ic e ni žj ih e no t G U U rt ic a di oi ca (i nc . U . g al eo ps ifo lia ) E1 2 3 1 2 1 2 . . . . + . . . . . . . + + . . 2 2 2 12 48 A I Im pa tie ns n ol i- ta ng er e E1 1 2 . + 1 1 . . . . . . . . . . . . + . . . . . . 6 24 M uA M ili um e ff us um E1 . 1 + . 1 1 . . . . . . . . . . . . . . . . . . . 4 16 FS Fr ax in us e xc el si or E3 b . . . . . . . . . . . . . . . . . + . . . . . . . 1 4 FS Fr ax in us e xc el si or E2 a . . . . . . . + + + + + . + . . . . . . . . . . . 6 24 A F C ar da m in e tr ifo lia E1 . . . . . . 1 + + + + 1 + . . . . . . . . . . . . 7 28 TA Ti lia p la ty ph yl lo s E2 . . . . . . r + + + + . . . . . r . . . . . . . . 6 24 A F Sc op ol ia c ar ni ol ic a E1 . . . 1 . . . . + . . . . . 3 3 3 4 4 3 . . . . . 8 32 FS Le uc oj um v er nu m E1 . . . . . . . . . . . . . . . . . . . . 4 3 . . . 2 8 T R A de no st yl es g la br a E1 . . . + . . . . . . . . . . . . . . . . . . + + . 3 12 EC Pr im ul a vu lg ar is E1 . . . . . . . . . . . . . . . . . . . . . . + . 3 2 8 Q P Ru sc us a cu le at us E2 a . . . . . . . . . . . . r . . . . . . . . . . . 3 2 8 TA Ti lio -A ce ri on A ce r p se ud op la ta nu s E3 a . . . . . . . . . . . . . . + . . . . . . . . . . 1 4 A ce r p se ud op la ta nu s E2 r . 1 1 1 1 . + 1 + + 1 1 + . . . . . . . 1 + . . 14 56 A ce r p se ud op la ta nu s E1 . . . . . . . . 1 . + . + . + + + . . + . . . . . 7 28 A ru m m ac ul at um E1 1 1 . 1 . . 1 1 . . . + + + . . + . . . 2 1 . . 1 12 48 U lm us g la br a E3 b . . . . . . . . . . . . . . . . . + . . . . . . . 1 4 U lm us g la br a E2 r + + . . . 1 + 1 + . 3 1 . . . . + . + . . . . . 11 44 Po ly st ic hu m a cu le at um E1 r + . . 1 + . . . . . . . + . . + + . + 1 . + 1 . 11 44 A ru nc us d io ic us E1 . + . . . + . . + . + . . + + + + + . . . . . . . 9 36 Po ly st ic hu m se tif er um E1 + . . + . . 2 . 2 . + + 1 1 . . . . . . . . . . . 8 32 D ry op te ri s a ff in is E1 . + . + + . . . + . + . . . . . . . . . . . . . + 6 24 St el la ri a m on ta na E1 . . . + . 1 . . + . . . . . . . . . + . . + . . . 5 20 A do xa m os ch at el lin a E1 . . . . . . . . . . . . . . . . . . 1 . + + + . . 4 16 Po ly st ic hu m b ra un ii E1 . . . . . . . . . . 1 . . . . . . . . . + + . . . 3 12 A ce r p la ta no id es E2 + . + . . . . . . . . . . . . . . . . . . . . . . 2 8 Eu on ym us la tif ol ia E2 . . . + . . . . . . . . . . . . . + . . . . . . . 2 8 C ir ca ea x in te rm ed ia E1 . . . . + + . . . . . . . . . . . . . . . . . . . 2 8 Te ph ro se ri s p se ud oc ri sp a E1 . . . . + + . . . . . . . . . . . . . . . . . . . 2 8 Po ly st ic hu m x lu er ss en ii E1 . . . . . + . . . . . . . . . . . + . . . . . . . 2 8 Ju gl an s r eg ia E2 . . . . . . . . . + . . . . . . . . . + . . . . . 2 8 A ce r p la ta no id es E1 + . . . . . . . . . . . . . . . . . . . . . . . . 1 4 H es pe ri s c an di da E1 . + . . . . . . . . . . . . . . . . . . . . . . . 1 4 A nt hr is cu s n iti da E1 . . . + . . . . . . . . . . . . . . . . . . . . . 1 4 T ha lic tr um a qu ile gi ifo liu m E1 . . . . . . . . . . . . . . . . . + . . . . . . . 1 4 A I A ln io n in ca na e C hr ys os pl en iu m a lte rn ifo liu m E1 . . . . 1 + . . . . 1 . . . . . . . 1 . . + . . . 5 20 C ar da m in e im pa tie ns E1 . . . + + . . . . . . . . . . . . . . . . . . . . 2 8 DAKSKOBLER & MARTINČIČ: VEGETATION OF MOIST ROCK CREVICES AND MOIST (SLOPE) DEBRIS IN THE LIŠČAK GORGE 96 FOLIA BIOLOGICA ET GEOLOGICA 64/1 – 2023 V ib ur nu m o pu lu s E2 . . . . . . . . . . . . . . . . . + . . . . . . . 1 4 A ru m it al ic um E1 . . . . . . . . . . . . . . . . . . . . . . . . + 1 4 C ar ex p en du la E1 . . . . . . . . . . . . . . . . . . . . . . . . + 1 4 Eq ui se tu m te lm at ei a E1 . . . . . . . . . . . . . . . . . . . . . . . . + 1 4 A F A re m on io -F ag io n C yc la m en p ur pu ra sc en s E1 . . . . . . . . + + . . . . . . . + . . + . . . . 4 16 C ar da m in e en ne ap hy llo s E1 . . + . . . . . . . . . . . . . . + . + . . . . . 3 12 O m ph al od es v er na E1 . . . + . . . . . . . . . . . . . + . . . . . . . 2 8 H ac qu et ia e pi pa ct is E1 . . . . . . . . . . . . . . . . . + . + . . . . . 2 8 G er an iu m n od os um E1 . . . . . . . . . . . . . . . . . . + + . . . . . 2 8 V ic ia o ro bo id es E1 . . . . + . . . . . . . . . . . . . . . . . . . . 1 4 H el le bo ru s n ig er E1 . . . . . . . . . . . . . . . . . . . + . . . . . 1 4 EC Er yt hr on io -C ar pi ni on G al an th us n iv al is E1 . . . . . . + . . . . . . . . . . . . . . . 1 . . 2 8 H el le bo ru s o do ru s E1 . . . . . . . . + . . . . . . . . . . . 1 . . . . 2 8 Lo ni ce ra c ap ri fo liu m E2 a . . . . . . . . . . . . . . . . . . . . . . . . + 1 4 FS Fa ge ta lia sy lv at ic ae Sa lv ia g lu tin os a E1 + 1 + + + + . . . . + . . . + + + . + 1 . . 1 + 2 15 60 G al eo bd ol on fl av id um E1 . . . . 1 1 1 . + 1 1 + + . 1 + + + . 1 . . + . . 14 56 Fa gu s s yl va tic a E2 a + + . 1 . . . r . . + 1 + . r . r . . + . . . + . 11 44 Fa gu s s yl va tic a E1 . + . . . . . . . . 1 . . . . . . . + + . . . . . 4 16 D ry op te ri s f ili x- m as E1 2 1 1 1 . 1 . + + + . . . . + . . . . 1 . . . + . 11 44 M er cu ri al is p er en ni s E1 1 1 + . . . . . + 1 . . + . . . . . . . + + . + 3 10 40 A sa ru m e ur op ae um s ub sp . c au ca si cu m E1 . . . . . . + . + + 1 . + 1 . . . . . + . . + 1 . 9 36 G al iu m la ev ig at um E1 . . . . . . . . . 1 + + . + + + + . . + . . + . . 9 36 G al iu m o do ra tu m E1 + + 1 1 . . . + . . . + + 2 . + . . . . . . . . . 9 36 Sy m ph yt um tu be ro su m E1 + + + . . . . . . . + . + + + + . . . . . . . . 1 9 36 C ar da m in e bu lb ife ra E1 . + 2 . . . . 1 . . + 3 + 2 . . . . 1 . . . . . . 8 32 M yc el is m ur al is E1 . + . . . . + . . . 1 . . . + + + . . . . . + + . 8 32 Pe ta si te s a lb us E1 . . . + . . . + + . . 1 2 3 . . 1 . . . . . . . . 7 28 Fe st uc a al ti ss im a E1 + + . + . . r . . + 1 . . . . . . . . . . . . . . 6 24 R an un cu lu s l an ug in os us E1 . . . . 3 1 . . + . . . . . . . . . . . + + . . . 5 20 A ct ae a sp ic at a E1 . . . . + . . + . + + . . . . . + . . . . . . . . 5 20 Po ly go na tu m m ul tif lo ru m E1 + . . . . . . . + . . + . + . . . . . . . . . . . 4 16 Pu lm on ar ia o ff ic in al is E1 . . . . . . . + . . . + + . . . . . + . . . . . . 4 16 V io la re ic he nb ac hi an a E1 . . . . . . . . + . + . + + . . . . . . . . . . . 4 16 Sc ro ph ul ar ia n od os a E1 . + . + . . . . . . . . . . . . + . . . . . . . . 3 12 La th yr us v er nu s E1 . . . . . . . . . . . . . . + . . + . . . . . . + 3 12 C or yd al is c av a E1 2 . . . . . . . . . . . . . . . . . 3 . . . . . . 2 8 Br ac hy po di um sy lv at ic um E1 . . . + . . . . . . . . . . . . . . . . . . + . . 2 8 M yo so ti s s yl va tic a E1 . . . . + . . . . . . . . . . . . . . . . . + . . 2 8 La bu rn um a lp in um E2 . . . . . + . . . + . . . . . . . . . . . . . . . 2 8 Pa ri s q ua dr ifo lia E1 . . . . . . + + . . . . . . . . . . . . . . . . . 2 8 Sa ni cu la e ur op ae a E1 . . . . . . . . . + . . . . . . . . . . . . . . + 2 8 C ar ex sy lv at ic a E1 . . . . . . . . . . . . . + . . . . . . . . . . + 2 8 Eu ph or bi a du lc is E1 . . . . . . . . . . . . . . 1 . . . . . . . . . + 2 8 Su cc es si ve n um be r of r el ev é (Z ap or ed na š te vi lk a po pi sa ) 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 Pr . Fr . DAKSKOBLER & MARTINČIČ: VEGETATION OF MOIST ROCK CREVICES AND MOIST (SLOPE) DEBRIS IN THE LIŠČAK GORGE 97FOLIA BIOLOGICA ET GEOLOGICA 64/1 – 2023 D ap hn e m ez er eu m E2 a . . . . . . . . . . . . . . + . . . . . . . . + . 2 8 A lli um u rs in um E1 . . . . . . . . . . . . . . . . . + . . . . . . 1 2 8 La th yr us v er nu s s ub sp . f la cc id us E1 . . . . . . . . . . . . . . . . . . . . . . + + . 2 8 C am pa nu la tr ac he liu m E1 . . . + . . . . . . . . . . . . . . . . . . . . . 1 4 Ep ilo bi um m on ta nu m E1 . . . . + . . . . . . . . . . . . . . . . . . . . 1 4 M el ic a nu ta ns E1 . . . . . + . . . . . . . . . . . . . . . . . . . 1 4 Lo ni ce ra a lp ig en a E2 a . . . . . . . . . . . . . . . . . + . . . . . . . 1 4 Ph yt eu m a sp ic at um s ub sp . c oe ru le um E1 . . . . . . . . . . . . . . . . . + . . . . . . . 1 4 Eu ph or bi a am yg da lo id es E1 . . . . . . . . . . . . . . . . . . . . . . . . + 1 4 Q P Q ue rc et al ia p ub es ce nt i- pe tr ae ae O st ry a ca rp in ifo lia E2 . . . . . + . . + 1 + . . . . . . . . . . . . . . 4 16 Se sl er ia a ut um na lis E1 . . . . . . . . . . + . . . . + . . . . . . . + . 3 12 A ra bi s t ur ri ta E1 . . . . . . . . . . + . . . . . . . . . . . + . . 2 8 H yp er ic um m on ta nu m E1 . . . + . . . . . . . . . . . . . . . . . . . . . 1 4 C or nu s m as E2 a . . . . . . . . . + . . . . . . . . . . . . . . . 1 4 La th yr us v en et us E1 . . . . . . . . . . . . . + . . . . . . . . . . . 1 4 A sp ar ag us te nu ifo liu s E1 . . . . . . . . . . . . . . . . . . . + . . . . . 1 4 A sp ar ag us a cu tif ol iu s E2 a . . . . . . . . . . . . . . . . . . . . . . . . + 1 4 C ar ex fl ac ca E1 . . . . . . . . . . . . . . . . . . . . . . . . + 1 4 Fr ax in us o rn us E2 . . . . . . . . . . . . . . . . . . . . . . . . + 1 4 H el le bo ru s m ul tif id us s ub sp . i st ri ac us E1 . . . . . . . . . . . . . . . . . . . . . . . . + 1 4 O rc hi s p ur pu re a E1 . . . . . . . . . . . . . . . . . . . . . . . . r 1 4 Ta m us c om m un is E1 . . . . . . . . . . . . . . . . . . . . . . . . + 1 4 Q F Q ue rc o- Fa ge te a A eg op od iu m p od ag ra ri a E1 . . 2 + 1 1 . . . . + + . 2 . . . + . + 1 + . . . 11 44 H ed er a he lix E1 . . . . . . + . + + + + + + + . + . + . . . . . 1 11 44 C ar ex d ig ita ta E1 . . . . . . r . . + . + . + + + . . . . . . + + + 9 36 C le m at is v ita lb a E2 a . . . . . + . . + . + . + + + . + . . 1 . . . . + 9 36 Ru bu s h ir tu s E2 a . . + . . . . . + . . . 1 1 + . . . . . + . . + . 7 28 A ne m on e ne m or os a E1 . . + . . . . . 1 . . . 1 . + . . . + . . + . . . 6 24 C or yl us a ve lla na E2 a . . . 1 . . . + . . . . . . . . . + . + . . . . + 5 20 R an un cu lu s f ic ar ia E1 . . . . . . . . . . . . . . . . . . 2 . + + . . 1 4 16 H ep at ic a no bi lis E1 . . . . . . . . + . . . . . . . . . . + + + . . . 4 16 A ne m on e ra nu nc ul oi de s E1 . . . . . . . + 1 . . . . . . . . . . . . . . . . 2 8 A ce r c am pe st re E2 b . . . . . . . . . . + . . . . . . . . . . . . . + 2 8 C er as tiu m sy lv at ic um E1 . . . . . . . . . . . + . . . . . . . . + . . . . 2 8 Py ru s p yr as te r E2 a . r . . . . . . . . . . . . . . . . . . . . . . . 1 4 C ar pi nu s b et ul us E2 a . . . . . . . + . . . . . . . . . . . . . . . . . 1 4 V er at ru m n ig ru m E1 . . . . . . . . . . . . . . r . . . . . . . . . . 1 4 Lo ni ce ra x yl os te um E2 . . . . . . . . . . . . . . . . . . . + . . . . . 1 4 Pt er id iu m a qu ili nu m E1 . . . . . . . . . . . . . . . . . . . + . . . . . 1 4 A ce r c am pe st re E1 . . . . . . . . . . . . . . . . . . . . . . . . + 1 4 EP Er ic o- Pi ne te a C ir si um e ri si th al es E1 . . . . . . . . . . . . . . . . . + . . . . . . . 1 4 Bu ph th al m um sa lic ifo liu m E1 . . . . . . . . . . . . . . . . . . . . . . + . . 1 4 Su cc es si ve n um be r of r el ev é (Z ap or ed na š te vi lk a po pi sa ) 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 Pr . Fr . DAKSKOBLER & MARTINČIČ: VEGETATION OF MOIST ROCK CREVICES AND MOIST (SLOPE) DEBRIS IN THE LIŠČAK GORGE 98 FOLIA BIOLOGICA ET GEOLOGICA 64/1 – 2023 V P Va cc in io -P ic ee te a O xa lis a ce to se lla E1 + . . . . . + . + + 1 1 1 1 . + + + . + . . + + . 14 56 V er on ic a ur tic ifo lia E1 . . . . + . . . . . + . . . . . + + . . . . . . . 4 16 C al am ag ro st is a ru nd in ac ea E1 . . . . . . . . . . . . . + + + . . . . . . . . . 3 12 So lid ag o vi rg au re a E1 . . . + . . . . . . . . . . . . + . . . . . . + . 3 12 Sa xi fr ag a cu ne ifo lia E1 . . . . . . . . . + . . . . + . . . . . . . . . . 2 8 Lu zu la lu zu lo id es E1 . + . . . . . . . . . . . . . . . . . . . . . . . 1 4 Lu zu la p ilo sa E1 . . . . . . . . . . . + . . . . . . . . . . . . . 1 4 Pi ce a ab ie s E2 a . . . . . . . . . . . . . . . . . + . . . . . . . 1 4 Ro sa p en du lin a E2 a . . . . . . . . . . . . . . . . . + . . . . . . . 1 4 R P R ha m no -P ru ne te a Ru bu s f ru tic os us a gg . E2 a . . . . . . . + . + + . . . . + . . . . . . . . . 4 16 Eu on ym us e ur op ae a E2 a . . . . . . . . . + . . . . . . . . . . . . . . . 1 4 So rb us a uc up ar ia E2 . . . . . . . . . . . . . . . . . + . . . . . . . 1 4 Ro bi ni a ps eu do ac ac ia E2 . . . . . . . . . . . . . . . . . . . . . . . . 1 1 4 C ra ta eg us m on og yn a E2 . . . . . . . . . . . . . . . . . . . . . . . . + 1 4 Li gu st ru m v ul ga re E2 a . . . . . . . . . . . . . . . . . . . . . . . . + 1 4 Fi cu s c ar ic a E2 . . . . . . . . . . . . . . . . . . . . . . . . + 1 4 BA Be tu lo -A ln et ea Sa lix a pp en di cu la ta E2 a . . . + 1 . . . . . . . . . . . . . . . . . . . . 2 8 M uA M ul ge di o- A co ni te te a A th yr iu m fi lix -f em in a E1 1 + 1 . . . . + + + + + + 1 + + + . . . . . . . . 13 52 Se ne ci o ov at us E1 + . 1 1 + 1 . . . . + . . + + 1 1 + . . . . 1 + . 13 52 A co ni tu m d eg en ii su bs p. p an ic ul at um E1 . . . . + + . . . . . . . . . . . + . + . . . . . 4 16 D or on ic um a us tr ia cu m E1 . . . + . . . . . . . . . . . . . . . . + . . . . 2 8 Sa xi fr ag a ro tu nd ifo lia E1 . . . . + + . . . . . . . . . . . . . . . . . . . 2 8 V er at ru m a lb um E1 . . . . . . . . . . . . . . . . . + . 1 . . . . . 2 8 Pe ta si te s h yb ri du s E1 . . . + . . . . . . . . . . . . . . . . . . . . . 1 4 C ha er op hy llu m h ir su tu m E1 . . . . . . . . . . . . . . . . . + . . . . . . . 1 4 A co ni tu m ly co ct on um E1 . . . . . . . . . . . . . . . . . . . + . . . . . 1 4 EA Ep ilo bi et ea a ng us ti fo lii Ru bu s i da eu s E2 a . + + + . . . . . . . . . . + + 1 . . . . . . + . 7 28 H yp er ic um h ir su tu m E1 . . . . + + . . . . + . . . . + . . . . . . . . . 4 16 A tr op a be lla -d on na E1 . 1 . + . . . . . . . . . . . . + . . . . . . . . 3 12 Eu pa to ri um c an na bi nu m E1 . . . + . + . . . . . . . . . . . . . . . . . . . 2 8 St ac hy s s yl va tic a E1 . . . . . . . . . . . . + . . . + . . . . . . . . 2 8 Br om op si s b en ek en ii E1 . . . 1 . . . . . . . . . . . . . . . . . . . . . 1 4 So la nu m d ul ca m ar a E1 . . . + . . . . . . . . . . . . . . . . . . . . . 1 4 St ac hy s a lp in a E1 . . . . + . . . . . . . . . . . . . . . . . . . . 1 4 A rc tiu m n em or os um E1 . . . . . . . . . . . . . . + . . . . . . . . . . 1 4 C ir si um a rv en se E1 . . . . . . . . . . . . . . . . + . . . . . . . . 1 4 Tu ss ila go fa rf ar a E1 . . . . . . . . . . . . . . . . + . . . . . . . . 1 4 T G Tr ifo lio -G er an ie te a V er ba sc um la na tu m E1 . r . . . . . . . . . . . . . . . . . . . . . . . 1 4 Li liu m c ar ni ol ic um E1 . . . . . . . . . . . . . . . . . . . + . . . . . 1 4 Su cc es si ve n um be r of r el ev é (Z ap or ed na š te vi lk a po pi sa ) 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 Pr . Fr . DAKSKOBLER & MARTINČIČ: VEGETATION OF MOIST ROCK CREVICES AND MOIST (SLOPE) DEBRIS IN THE LIŠČAK GORGE 99FOLIA BIOLOGICA ET GEOLOGICA 64/1 – 2023 G U G al io -U rt ic et ea A lli ar ia p et io la ta E1 . + . . . . . . . . . . . . . . . . . . . . . . . 1 4 La m iu m m ac ul at um E1 . . . 1 . . . . . . . . . . . . . . . . . . . . . 1 4 C ar da m in e hi rs ut a E1 . . . . . . . . . . . . . . . . . . . . . . . . + 1 4 C he lid on iu m m aj us E1 . . . . . . . . . . . . . . . . . . . . . . . . + 1 4 Er ig er on a nn uu s E1 . . . . . . . . . . . . . . . . . . . . . . . . + 1 4 M A M ol in io -A rr he na th er et ea A ng el ic a sy lv es tr is E1 . . . . . . . . . . . . . . . + . . . . + + . . . 3 12 Ta ra xa cu m s ec t. Ru de ra lia E1 . + . . . . . . . . . . . . . + . . . . . . . . . 2 8 C al th a pa lu st ri s E1 . . . . . . . . . . . . . . . . . + . . . . . . . 1 4 T R T hl as pi et ea ro tu nd ifo lii G ym no ca rp iu m ro be rt ia nu m E1 . . . . . . . . . . . . . . . . . . . 1 . . . . . 1 4 A T A sp le ni et ea tr ic ho m an is A sp le ni um tr ic ho m an es E1 . . . + . . . . . . + . . . + . . . . + . . + . . 5 20 Po ly po di um v ul ga re E1 . . . . . . . . + . . . . . . . . . . . . . + + . 3 12 V al er ia na tr ip te ri s E1 . . . . . . . . . . . . . . + . . . . . . . + . . 2 8 C ys to pt er is fr ag ili s E1 . . . . + . . . . . . . . . . . . . . . . . . . . 1 4 Sa xi fr ag a pe tr ae a E1 . . . . . . . . . + . . . . . . . . . . . . . . . 1 4 M oe hr in gi a m us co sa E1 . . . . . . . . . + . . . . . . . . . . . . . . . 1 4 Pa ri et ar ia ju da ic a E1 . . . . . . . . . . . . . . . . . . . . . . . . + 1 4 M L M os se s (M ah ov i) T ha m no br yu m a lo pe cu ru m E0 . . . . . . 1 1 1 3 . . . . . . . . 2 . 1 2 + . 2 9 36 Is ot he ci um a lo pe cu ro id es E0 . . . + . . . . . 1 1 . . . . . . . 1 . 2 . 2 1 . 7 28 Pl ag io m ni um u nd ul at um E0 . . . . 1 1 . . + 1 . . . . . . . + . 2 . . . . . 6 24 C te ni di um m ol lu sc um E0 . . . . . . . . . + 1 . . . . . . . . 1 + . 2 . . 5 20 C on oc ep ha lu m c on ic um E0 . . . . 1 1 . . . . + . . . . . . + . . . . . . . 4 16 Ex er to th ec a cr is pa (N ec ke ra c ri sp a) E0 . . . . . . . . . . 1 . . . . . . . . 2 . . 1 1 . 4 16 Br ac hy th ec iu m r ut ab ul um E0 . . . . . . . . . 1 . . . + . . . . . . . 2 . . . 3 12 Po ly tr ic hu m fo rm os um E0 . + . . . . . . . . . . . + . . . . . + . . . . . 3 12 A tr ic hu m u nd ul at um E0 . + . . . . . . . . . . . + . . . . . . . . . . . 2 8 Fi ss id en s d ub iu s E0 . . . . . . . . + . . . . . . . . + . . . . . . . 2 8 Ps eu da no m od on a tt en ua tu s ( A no m od on a tt en ua tu s) E0 . . . . . . . . . . + . . . . . . . . . . . . . + 2 8 Fi ss id en s t ax ifo liu s E0 . . . . . . . . . . . . . + . . 1 . . . . . . . . 2 8 Eu rh yn ch iu m a ng us tir et e E0 . . . . . . . . . 1 . . . . . . . . . . . . . . . 1 4 T hu id iu m ta m ar is ci nu m E0 . . . . . . . . . + . . . . . . . . . . . . . . . 1 4 H yp nu m c up re ss ifo rm e E0 . . . . . . . . . . . . . . . . . . . 1 . . . . . 1 4 H om al ot he ci um lu te sc en s E0 . . . . . . . . . . . . . . . . . . . . . . 1 . . 1 4 Sc hi st id iu m a po ca rp um E0 . . . . . . . . . . . . . . . . . . . . . . 1 . . 1 4 Le ge nd - Le ge nd a L Li m es to ne - ap ne ne c C h C he rt - ro že ne c D eb D eb ri s - g ru šč Fl F ly sc h - f liš G r G ra ve l - p ro d Su cc es si ve n um be r of r el ev é (Z ap or ed na š te vi lk a po pi sa ) 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 Pr . Fr . DAKSKOBLER & MARTINČIČ: VEGETATION OF MOIST ROCK CREVICES AND MOIST (SLOPE) DEBRIS IN THE LIŠČAK GORGE 100 FOLIA BIOLOGICA ET GEOLOGICA 64/1 – 2023 M M ar ls to ne - la po ro ve c R o R oc ks lid e - p od or no g ra di vo Ss S an ds to ne - pe šč en ja k C C C hr om ic C am bi so ls - rj av a po ka rb on at na tl a C o C ol lu vi al -d el uv ia l s oi l - k ol uv ia ln o- de lu vi al na tl a Eu E ut ri c br ow n so il - e vt ri čn a rj av a tla R e re nd zi na - re nd zi na Li L ith os ol - ka m ni šč e Pr . P re se nc e (n um be r of r el ev és in w hi ch th e sp ec ie s i s p re se nt ed ) - š te vi lo p op is ov , v k at er ih s e po ja vl ja v rs ta Fr . F re qu en cy in % - fr ek ve nc a v % Su cc es si ve n um be r of r el ev é (Z ap or ed na š te vi lk a po pi sa ) 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 Pr . Fr .