HACQUETIA 4/2 • 2005, 121–149


SynTAxonomy And nomEnClATUrE of THE
CommUnITIES of THE AllIAnCES Astero 
Alpini-seslerion cAlcAriAe 
And seslerion 
tAtrAe 
In SlovAkIA


Ján Kliment1, Radmila BělohlávKová2, Dana BeRnátová1, ivan JaRolímeK3, 
anton PetRíK4, Jozef ŠiBíK3, Jana Uhlířová5 & milan valachovič3 

Abstract 

the article brings the most important results of the syntaxonomical revision of the communities of the alliances Astero-Seslerion calcariae and Seslerion tatrae from the territory of Slovakia: descriptions of the new suballiances (Astero-Seslerienion calcariae, Pulsatillo slavicae-Caricenion humilis), associations (Gentiano clusii-Festucetum 
versicoloris, Diantho nitidi-Seslerietum tatrae) and subassociations (Minuartio langii-Festucetum pallentis campanuletosum carpaticae and campanuletosum xylocarpae, Seslerietum tatrae luzuletosum rubellae and biscutelletosum laevigatae, 
Seslerio tatrae-Festucetum versicoloris ranunculetosum pseudomontani and inops). the authors validate the name 
Minuartio langii-Festucetum pallentis (Sillinger 1933) mucina in mucina et maglocký 1985 nom. nud. they propose the new name for the association Arctostaphyletum fatrense Sillinger 1933 (Seslerio albicantis-Arctostaphyletum 
uvae-ursi). 

Izvleček 
v članku so obravnavani najpomembnejši rezultati sintaksonomske revizije zvez Astero-Seslerion calcariae in 
Seslerion tatrae na Slovaškem. opisane so: nove podzveze (Astero-Seslerienion calcariae, Pulsatillo slavicae-Caricenion humilis), asociacije (Gentiano clusii-Festucetum versicoloris, Diantho nitidi-Seslerietum tatrae) in subasociacije 
(Minuartio langii-Festucetum pallentis campanuletosum carpaticae in campanuletosum xylocarpae, Seslerietum tatrae 
luzuletosum rubellae in biscutelletosum laevigatae, Seslerio tatrae-Festucetum versicoloris ranunculetosum pseudomontani 
in inops). avtorji so pravilno opisali asociacijo Minuartio langii-Festucetum pallentis (Sillinger 1933) mucina in 
mucina et maglocký 1985 nom. nud. Predlagajo novo ime za asociacijo Arctostaphyletum fatrense Sillinger 1933 
(Seslerio albicantis-Arctostaphyletum uvae-ursi). 

key words: syntaxonomical revision, new syntaxa, high mountain calcareous grasslands, Elyno-Seslerietea, Western carpathians 
ključne besede: sintaksonomska revizija, novi sintaksoni, visokogorski travniki na apnencu, Elyno-Seslerietea, 
Zahodni Karpati 

1 Botanical Garden of the University of Komenský, SK-038 15 Blatnica, Slovak Republic, e-mail: kliment@rec.uniba. 
sk, bernatova@rec.uniba.sk 
2 institute of Botany, academy of Sciences of czech Republic, cZ-282 43 Průhonice, czech Republic, e-mail: belohlavkova@ibot.cas.cz 
3 institute of Botany, Slovak academy of Sciences, Dúbravská cesta 14, SK-845 23 Bratislava, Slovak Republic, e-mail: 
ivan.jarolimek@savba.sk (corresponding author), jozef.sibik@savba.sk, milan.valachovic@savba.sk 
4 Botanical Garden of the University of Komenský, Botanická 3, SK-841 04 Bratislava, Slovak Republic, e-mail: petrika@ 
rec.uniba.sk 
5 Slovak national museum-natural history museum, vajanského nábrežie 2, P.o. Box 13, SK-810 06 Bratislava 16, 
e-mail: uhlirova@snm.sk 

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intRoDUction


the class Elyno-Seslerietea Br.-Bl. 1948 associates 
mainly natural, neutrophilous to basiphilous grass 
communities on the calcite and dolomite substratum in the montane to the alpine belt of european 
mountain ranges (Pyrenees, apennines, the alps, 
carpathians, mountains of the Balkan Peninsula). 
the communities occupy very shallow to medium 
deep, skeletal, humus and calcite rich soils. in the 
Slovak part of the carpathians the class is represented only by the order Seslerietalia coerulae Br.-Bl. 
in Br.-Bl. et Jenny 1926. it includes three alliances, 
which differ in exposition to the sun, wind and also by the thickness and duration of the snow cover. 
the alliance Astero alpini-Seslerion calcariae hadač 
ex hadač et al. 1969 comprises open communities 
on shallow, carbonate skeleton and humus rich 
soils on the steep sunny calcareous and dolomite 
slopes, fissured rocks, small rocky terraces and stable screes with short term snow in the montane (to 
subalpine) belt. Within the alliance Seslerion tatrae 
Pawłowski 1935 corr. Klika 1955 are ordered mainly close communities occupying medium deep 
soils with a favourable water regime at leeward 
habitats. in winter they are covered by thick and 
long term snow. communities are situated in the 
alpine (rarely also in subalpine) belt. the alliance 
Caricion firmae Gams 1936 represents dwarf shrub-
grass communities on rocky, north oriented habitats exposed to strong winds in the alpine, less in 
the subalpine belts, in winter with thin and short 
time snow. the alliances Astero alpini-Seslerion 
calcariae and Seslerion tatrae are limited by their 
distribution to the territory of the Western carpathians. 

this work is based on complex processing of 
relevé data of the communities of the class Elyno-
Seslerietea from the carbonate mountains of the 
Western carpathians and involves the most important results of their syntaxonomical revision. contentually it relates to the recent summary paper on 
the West carpathian communities of the alliance 
Caricion firmae (Šibík & al. 2004). 

mateRial anD methoDS 

Processed data set contains 844 phytocoenological 
relevés of the class Elyno-Seslerietea from the limestone areas of the Western carpathians, also partially (the association Seslerio tatrae-Festucetum versicoloris) from its Polish part (Fig. 1). all relevés were 


Figure 1: Area of the occurrence of the plant communities 
of the West Carpathian endemic alliances Astero-Seslerion 
calcariae and Seslerion tatrae (marked by ellipse). Grey area 
stands for orographical units in the Carpathians. 
Slika 1: Območje pojavljanja rastlinskih združb, ki jih uvrščamo 
v endemične zveze zahodnih Karpatov Astero-Seslerion calcariae in Seslerion tatrae (označeno z elipso). Siva 
barva označuje orografske enote v Karpatih. 

obtained by methods of the Zürich-montpelliér 
school (Braun-Blanquet 1964); individual authors 
used different scales of abundance: the 7-degree 
Braun-Blanquet scale, its modified 9-degree version (Barkman & al. 1964), the 10-degree Domin 
or 11-degree Domin-hadač scale (cf. Klika 1948; 
hadač & al. 1969). to obtain comparable data for 
numerical classification all relevés were transformed into the nine-degree ordinal scale (van den 
maarel 1979). Subspecies and some narrowly delimited species were included into the closest higher or broadly defined taxa. taxa determined only 
to the rank of genus and the most of mosses (with 
exception of equal presence in data) were excluded. numerical classifications were performed by 
the nclaS program from the SYn-taX 2000 (Podani 2001). ß - flexible method (ß = –0.25) and Jaccard’s, Ružička’s and Wishart’s similarity coefficients and euclidean distance were used. the hypotheses obtained were evaluated by comparison 
and analysis of phytocoenological tables processed 
by the FYtoPacK program (Jarolímek & Schlosser 
1997). the results of numerical classification were 
used for approximative orientation in large data 
sets. the arrangement of the tables does not strictly 
follow these results. 

the nomenclature of taxa follows the checklist 
of non vascular and vascular plants of Slovakia 
(marhold & hindák 1998); the exceptions com

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J. Kliment et al.: Syntaxonomy and nomenclature of tHe communitieS of tHe allianceS Astero Alpini-seslerion CAlCAriAe … in Slovakia 
prise the author’s citation. in the tables subspecies 
(without the name of species) are marked by asterisks (*). the value of frequency (in %) and mean 
value of abundance in the relevant column (upper 
index) for every taxon is calculated. Sources to the 
columns contain: short citation of source (in unpublished data only the name of author/authors of 
the relevé), number of relevés and their location 
on the level of orographic units according to the 
base map for Database of Fauna of Slovakia in scale 

1: 500 000. Full names of syntaxa with author’s citation are given only for the first mention of the 
name in the text. 
Diagnostic taxa of the class Elyno-Seslerietea and 
lower syntaxa are used in accordance with the synoptic table prepared for the new fourth volume of 
the series of the books Plant communities of Slovakia. other syntaxa are defined following the newest 
relevant publications (Kliment & al. 2004, Šibík & 
al. 2004, Petrík & al. 2005). the names of syntaxa 
are abbreviated in the tables 1–3 as follows: ac = Arabidion coerulae, as = Astero-Seslerion calcariae, at = 
Asplenietea trichomanis, Be = Brometalia erecti, ca = Calamagrostion arundinaceae, cc = Caricetea curvulae, 
cf = Caricion firmae, ch = Pulsatillo-Caricenion humilis, 
cK = Carici rupestris-Kobresietea, cv = Calamagrostietalia villosae, cy = Cystopteridion, eS = Elyno-Seslerietea, 
FB = Festuco-Brometea, fc = Festucion carpaticae, fv = Festucion versicoloris, Fv = Festucetalia valesiacae, lv = Loiseleurio-Vaccinietea, mU= Mulgedio-Aconitetea, oe= 
Oxytropido-Elynion, Pc = Potentilletalia caulescentis, pc 
= Potentillion caulescentis, pt = Papaverion tatrici, Sc = 
Seslerietalia coerulae, sf = Seslerio-Festucion pallentis, st 
= Seslerion tatrae, tf = Trisetion fusci. Diagnostic important taxa of the individual communities are presented in bold type. 

in descriptions of communities following abbreviations were used: art. = art of the code of phytocoenological nomenclature (icPn; Weber & al. 
2000), ass. = association, bas.: basionym, c = characteristic taxon (in tables), cf. = confer (compare), 
d = differential taxon (in tables), dom., D = dominant species (D – in tables), incl. = inclusive, ined. 
= ineditus (unpublished data), nom. corr. = corrected name, nom. ined. = ineffectively published 
name, nom. nov. = nomen novum, nom. nud. = nomen nudum, oFn = original form of a name, p. p. = 
pro parte (partly), pseud. = pseudonym, r. = relevé, 
reg. = regionally characteristic taxon, sp. div. = species diversae (various species), subass. = subassociation, subdom. = subdominant taxon, syn. = synonym, syntax. syn. = syntaxonomical synonym, transgr., t = transgressive taxon (t – in tables). 

the communities well known from the literature are characterised only shortly by essential information; less known and newly described syntaxa 
are characterised in more detail. 

ReSUltS anD DiScUSSion 

elyno-seslerietea Br.-Bl. 1948 

seslerietalia coerulae Br.-Bl. in Br.-Bl. et Jenny 1926 

Syn.: Seslerietalia tatrae hadač 1962 (art. 8)
Syntax. syn.: Seslerietalia tatrae hadač ex hadač et 
al. 1969 p. p. (excl. Delphinion elati hadač ex hadač 
et al. 1969)


hadač & al. (1969) reclassified chionophilous 
communities of the alliance Seslerion tatrae – within 
the class Aconito-Cardaminetea hadač 1956 – into 
the order Seslerietalia tatrae. he included into this 
order also the alliance Delphinion elati, which was 
later regarded by mucina & maglocký (1985) as a 
part of the order Adenostyletalia alliariae Br.-Bl. 1930 
(class Mulgedio-Aconitetea hadač et Klika in Klika 
1948). Recent syntaxonomical revision confirms 
the validity of ordering of the alliance Seslerion tatrae into the class Elyno-Seslerietea. mutual comparison of the alliances Astero-Seslerion calcariae, Seslerion 
tatrae and Caricion firmae shows a strong differences 
in their floristic composition, reflecting the peculiarities of their habitat conditions. chionophobia 
partly delimitates the stands of the alliances Astero-
Seslerion calcariae and Caricion firmae against the alliance Seslerion tatrae, even though it is caused by different ecological factors – the alliance Astero-Seslerion occupies sunny habitats, the alliance Caricion 
firmae occupies habitats exposed to strong winds. 
consequently these alliances have nearly no common diagnostic (differential) taxa (tab. 1). on the 
other hand, with regard to optimal occurrence in 
the alpine belt, the alliances Seslerion tatrae and Caricion firmae are floristically closer by the presence of 
high mountain taxa. at the same time they differ 
just in the thickness and duration of snow. a division of the class Elyno-Seslerietea into the two orders 
(Seslerietalia coerulae, Seslerietalia tatrae) based on 
snow conditions without any diagnostic taxa is 
hardly reasonable. therefore we prefer the older 
conception (holub & al. 1967), based on division 
of the class Elyno-Seslerietea in the Western carpathians into three alliances (Astero-Seslerion calcariae, 
Seslerion tatrae, Caricion firmae) within only one order Seslerietalia coerulae (Fig. 2). 

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124 

Figure 2: Syntaxonomical and synecological scheme of the plant communities belonging to the class Elyno-Seslerietea in the Slovakia. The (sub)species inside ellipses represent 
diagnostic taxa (characteristic and differential); the studied (sub)alliances are marked by thicker outlines. The arrows indicate dynamics and close syngenetic relationships 
between individual syntaxa.
Slika 2: Sintaksonomska in sinekološka shema rastlinskih združb, ki jih uvrščamo v razred Elyno-Seslerietea na Slovaškem. (Pod)vrste v elipsah predstavljajo diagnostične taksone 
(značilne in razlikovalne), obravnavane (pod)zveze so označene z debelejšo črto. Puščice označujejo dinamske in singenetske povezave med posameznimi sintaksoni. 


J. Kliment et al.: Syntaxonomy and nomenclature of tHe communitieS of tHe allianceS Astero Alpini-seslerion CAlCAriAe … in Slovakia 
Astero alpini-seslerion calcariae 
Hadač ex Hadač et 
al. 1969 nom. invers. propos. 

(tab. 1, columns 1–5) 
Syn.: Seslerieto-Asterion alpini hadač 1962 (art. 8); 
Astereto-Seslerion calcariae hadač et Smola 1962 (art. 
8); Astero serpentimontani-Seslerion hadač 1962 corr. 
mucina 1981 (art. 43) 
Pseud.: Seslerion coerulae auct. carpat. occid. p. p., 
non Br.-Bl. in Br.-Bl. et Jenny 1926 
Phantom: Seslerio tatrae-Asterion alpini hadač 1962 
(coldea 1997) 
characteristic taxa: Sesleria albicans (dom.), Aster 
alpinus (Fig. 3), Dianthus praecox subsp. praecox, Festuca tatrae (transgr.), Minuartia langii 
Differential taxa: Asplenium ruta-muraria, Campanula carpatica, Jovibarba globifera, Kernera saxatilis, Leontodon incanus 
nomenclatural type: Astereto-Seslerietum calcariae hadač et al. 1969, lectotypus (art. 20) 

hadač (1962) described the alliance Seslerio-Asterion alpini based on the list of characteristic species only, hence invalidly; the name was validated 
in the collective work from the Belianske tatry mts 
(hadač & al. 1969). he included into the alliance 
open neutrophile to basiphile communities occupying unstable, skeleton rich soils at small rocky 
terraces in the montane to subalpine belts. these 
habitats are protected from the wind and they are 
without snow in the winter. later (mucina & maglocký 1985), the content of the alliance was broadened by the other West carpathian mountain grass 
communities on calcite or dolomite slopes, which 
originally had been ordered within the alliance Seslerion coerulae. in accordance to articles 10b and 20 

Figure 3: Aster alpinus – characteristic taxon of the alliance 
Astero-Seslerion calcariae. 
Slika 3: Aster alpinus – značilnica zveze Astero-Seslerion cal-
cariae. 
of icPn we propose reversion of the sequence of 
the taxa names. 

communities of the alliance Astero-Seslerion calcariae are characteristised by the presence of large 
number of endemic taxa of vascular plants, especially of endemic [cW] and subendemic [cWs] 
taxa of the West carpathian mountains or their 
sub-regions and carpathian endemics [c] and sub-
endemics [cs] (see tab. 1). however, the presence 
of these taxa, together with peculiar floro- and syngenesis, differentiates the West carpathian phytocoenoses from similar ones from the alps, ordered 
within the alliance Seslerion coerulae, even though 
they are unified in some recent surveys (cf. Grabherr & al. 1993). 

Strong differences in the floristical composition 
of individual communities conditioned by distribution in different vegetation belts allow for two suballiances to be distinguished: Astero alpini-Seslerienion 
calcariae and Pulsatillo slavicae-Caricenion humilis. 

Astero alpini-seslerienion calcariae 
suball. nov. hoc 
loco 

(tab. 1, columns 1–3) 
characteristic taxa: Bupleurum ranunculoides 
Differential taxa: Androsace chamaejasme, Astragalus 
australis, Campanula tatrae, Festuca versicolor subsp. 
versicolor 

nomenclatural type: identical with the type of the 
alliance name 

the nominate suballiance corresponds in its 
content to the original diagnosis of the alliance Astero-Seslerion calcariae (hadač & al. 1969). it involves 
stands occurring in the supramontane to subalpine 
belts, mainly in the Belianske tatry mts. against the 
following suballiance it is differed by high mountain taxa, which usually do not decrease to the montane belt. Dominants of stands are grasses Festuca 
*versicolor and Sesleria albicans, rarely Festuca tatrae. 

associations: Astero alpini-Seslerietum calcariae hadač et al. 1969, Diantho praecoci-Festucetum versicoloris 
hadač et al. 1969, Gentiano clusii-Festucetum versicoloris Bělohlávková ass. nov. 

Astero alpini-seslerietum calcariae Hadač et al. 1969 

(tab. 1, column 1) 
characteristic taxon: Oxytropis campestris subsp. tatrae 
Differential taxa: Gypsophila repens (subdom.), Leontopodium alpinum; Fulgensia bracteata, Hymenostylium 
recurvirostrum, Squamarina gypsacea 

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Hacquetia 4/2 • 2005 

constant companions: Sesleria albicans (dom.), Aster 
alpinus, Bupleurum ranunculoides, Campanula tatrae, 
Dianthus *praecox, Euphrasia salisburgensis, Festuca tatrae, F. *versicolor, Galium anisophyllon, Helianthemum 
grandiflorum, Jovibarba globifera, Libanotis pyrenaica, 
Primula auricula subsp. hungarica, Ranunculus breyninus, Saxifraga paniculata, Thymus pulcherrimus subsp. sudeticus, Trisetum alpestre; Tortella tortuosa 
nomenclatural type: hadač & al. 1969: 67, r. 106, 
lectotypus hoc loco 

an open, species rich community dominated by 
Sesleria albicans occupies very shallow, skeleton and 
humus rich, slightly to medium moist soils on small 
terraces at steep, sunny, south oriented calcareous 
rocks in the subalpine belt in the Belianske tatry 
mts, ca 1490–1750 m a. s. l. habitats are protected 
from wind, exposed to direct sunshine, in winter 
without snow. the community is close to the communities of the alliance Potentillion caulescentis Br.-
Bl. in Br.-Bl. et Jenny 1926 em. Sutter 1969, mainly 
to the association Leontopodio-Asteretum alpini Šmarda ex Šmarda et al. 1971 by a low cover of herb 
layer (30–60 %), similar floristic composition and 
also by synecology. 

Diantho praecoci-Festucetum versicoloris 
Hadač et al. 
1969 

(tab. 1, column 2) 
Syn.: Festucetum tatrae Šmarda 1956 (art. 31) 
Syntax. syn.: Tortello tortuosae-Festucetum tatrae hadač 
et al. 1969 
non: Festucetum tatrae Szafer et al. 1923 corr. 1927 
Differential taxa: Festuca carpatica, Gentianella lutescens, Hieracium murorum, Linum extraaxillare, Silene 
vulgaris subsp. prostrata (Gaudin) chater et Walters, Trifolium pratense subsp. kotulae 
constant companions: Festuca *versicolor (dom.), F. 
tatrae (subdom.-dom.), Carex sempervirens subsp. 
Tatrorum (Zapał.) Pawł., Dianthus *praecox, Galium 
anisophyllon, Helianthemum grandiflorum, Libanotis 
pyrenaica, Lotus corniculatus var. alpicola Beck, Phyteuma orbiculare, Polygala amara subsp. brachyptera, 
Ranunculus breyninus, Scabiosa lucida, Trisetum alpestre; Tortella tortuosa 
nomenclatural type: hadač & al. 1969: 72–73, r. 
90, lectotypus hoc loco 

Subspecies Festuca *versicolor dominates in a species rich, open community. in some places species 

F. tatrae occurs with similar cover. the community 
occupies sheer, south oriented (SW-Se) talus cones 
in the subalpine belt in the Belianske tatry mts, 
1450–1775 m a. s. l. Shallow, rocky, humus rich and 
dense rooted soils are, during winter, without or 
with only thin snow cover and they are exposed to 
the sun radiation. 

hadač & al. (1969), based on four relevés from 
the Dolina Siedmich prameňov valley, described 
the association Tortello-Festucetum tatrae with constant subdominant presence of the subspecies Festuca *versicolor. it is in synmorphology, synecology 
and floristic composition very close to the association Diantho-Festucetum versicoloris. the authors 
characterized these stands as more thermophile 
with regard to the lower elevation of most of the 
localities. Despite these differences the syntaxonomical revision proves their ordering within the 
variability of the association Diantho-Festucetum versicoloris. their relevé 221 can be identified with this 
association without restrictions, while others differ 
mainly negatively. the lower number of species to 
a certain extent relates with the small sample area 
(2 m2 only) of two of analysed stands (r. 256, 258). 

Gentiano clusii-Festucetum versicoloris Bělohlávková 
ass. nov. hoc loco 

(tab. 2; tab. 1, column 3) 
Differential taxa: Androsace lactea, Bartsia alpina, 
Hedysarum hedysaroides, Potentilla crantzii, Ranunculus alpestris, Rhodiola rosea, Saxifraga moschata subsp. 
dominii, Sesleria tatrae, Vicia oreophila 
constant companions: Festuca *versicolor (dom.), 
Euphrasia salisburgensis, Galium anisophyllon, Gentiana clusii, Helianthemum grandiflorum, Jovibarba globifera, Phyteuma orbiculare, Poa alpina, Primula *hungarica, Saxifraga paniculata, Scabiosa lucida, Thymus 
*sudeticus, Trisetum alpestre 
nomenclatural type: tab. 2, r. 7, holotypus 

an open community with prevalence of the subspecies Festuca *versicolor, concurrently with an 
amount of dicotyledonous herbs, which attract attention by their various coloured flowers during 
flowering period. taxa Primula *hungarica and Gentiana clusii, more rarely Draba aizoides and Pulsatilla 
slavica subject striking yellow-blue spring aspect 
(Fig. 4). the more varicoloured summer aspect is 
conditioned mainly by species Androsace lactea, Bartsia alpina, Campanula cochleariifolia, Dianthus *praecox, Helianthemum grandiflorum, Phyteuma orbiculare, 
Potentilla crantzii, Ranunculus breyninus, Saxifraga 
paniculata, Scabiosa lucida and Vicia oreophila. Beside 
the dominant fescue the grasses Sesleria tatrae, S. al

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J. Kliment et al.: Syntaxonomy and nomenclature of tHe communitieS of tHe allianceS Astero Alpini-seslerion CAlCAriAe … in Slovakia 
Figure 4: Early vernal aspect of the community Gentiano 
clusii-Festucetum versicoloris. (Krivánska Malá Fatra Mts 

– rock formation “Sviňa”; altitude 1 555 m a. s. l.; May 27th 
2005). 
Slika 4: Zgodnjepomladanski aspekt asociacije Gentiano 
clusii-Festucetum versicoloris. (gorovje Krivánska Malá Fatra 
– gora »Sviňa«; nadmorska višina 1 555 m; 27. maj 2005). 
bicans, Poa alpina and Trisetum alpestre also participate in stands. the moss layer is developed scarcely. the average number of species per relevé is 30 
(min. 23, max. 37). 

the community occupies fissures and small terraces on steep, sunny to slightly shadowed calcareous rock in the subalpine belt. Dense roots of dominant fescue help the development of humus protorendzina in fissures and on terraces. these 
habitats are without snow or with a thin layer of 
short lasting snow during winter (Fig. 5); in early 
spring they are exposed to large variations of temperature. 

Based on recent knowledge on floristic composition, ecology and distribution, two variants can be 
distinguished within the association: a more mesophilous variant with Rhodiola rosea (tab. 2, r. 1–9; 
Differential taxa: Rhodiola rosea, Allium senescens 
subsp. montanum, Anemone narcissiflora, Hedysarum 
hedysaroides, Parnassia palustris, Potentilla crantzii, Vi-


Figure 5: Winter aspect of the community Gentiano clusii-
Festucetum versicoloris – highlighted by arrow (Krivánska 
Malá Fatra Mts – rock formation “Sviňa”; altitude 1 555 m a. 

s. l.; February 10th 2005; all photos were taken by J. Šibík). 
Slika 5: Zimski aspekt asociacije Gentiano clusii-Festucetum 
versicoloris – označeno s puščico (gorovje Krivánska Malá 
Fatra – gora »Sviňa«; nadmorska višina 1 555m; 10. februar 
2005; vse fotografije J. Šibík). 
ola biflora, Tortella tortuosa) occurs mainly in south 
oriented choppy rocks in the highest parts of the 
Krivánska malá Fatra mts, 1540–1625 m a. s. l. 
Stands of the variant with Minuartia langii (tab. 2, 

r. 10–14) are more xerophilous, with a higher portion of petrophytes and without mosses. this is reflected in the group of differential taxa (Carex ornithopoda, Hieracium villosum, Minuartia langii, Pulsatilla slavica, Rhodax alpestris, Veronica fruticans). the 
variant was found in the south and south-east oriented rocks of the choč mt. at altitude 1470–1500 
m a. s. l. (Švandová-Ursínyová 1966). 
pulsatillo slavicae-caricenion humilis 
Uhlířová sub-
all. nov. hoc loco 

(tab. 1, columns 4–5) 
characteristic taxa: Acinos alpinus, Allium ochroleucum, Bromus monocladus (reg.), Buphthalmum salicifolium (reg.), Coronilla vaginalis, Daphne arbuscula 
(reg.), Erysimum witmannii, Globularia cordifolia 
(reg.), Koeleria tristis (reg.), Pulsatilla slavica, Rhodax 
rupifragus 
Differential taxa: Carex humilis, Anthericum ramosum, 
Asperula cynanchica, A. tinctoria, Cyanus triumfettii 

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(subsp. dominii, subsp. axillaris), Festuca pallens, Hieracium bupleuroides, Polygonatum odoratum, Seseli osseum, Teucrium montanum, Tithymalus cyparissias, Vincetoxicum hirundinaria 
nomenclatural type: Caricetum humilis carpaticum 
Sillinger 1933 

thesuballiancerepresentsphytocoenosesoccurring in the montane belt. they usually occupy only 
small areas forming a vegetation complex with relic 
stands of the Scots Pine or european larch (Larix 
decidua subsp. polonica). typical components and diagnostic species of the communities of this suballiance are xero-thermophilous and prealpine species 
(Fig. 2). many of them survive there as relicts from 
the drier and warmer postglacial and/or interglacial 
periods. they often live in the ecological limit of 
their distribution and some of them reach there the 
altitudinallimitintheWesterncarpathians.thefloristic spectrum of these communities is supplemented by calciphilous oreophytes and dealpine species 
with optimal occurrence on the relatively colder 
habitats with higher air humidity. Carex humilis, Sesleria albicans, Festuca pallens, F. tatrae, Carex approximata 
Bell. ex all. and C. *tatrorum represent the most frequent dominants and subdominants. 

the suballiance associates phytocoenoses ordered by Sillinger (1933) into the “group of 
xerophilous and subxerophilous associations on 
calcareous rocks and fine scree (the alliance Seslerion coerulae p. p.)”, except for the association Calamagrostietum variae carpaticum Sillinger 1933, and 
also later described communities on sunny calcite 
and dolomite slopes in the montane to supramontane belt. the association Diantho nitidi-Caricetum 
tatrorum (syn.: Seslerio-Semperviretum fatrense Sillinger 
1933), distributed over large areas in the supra-
montane to subalpine belt, has a transition syntaxonomical position between the alliances Astero-Seslerion calcariae and Seslerion tatrae. 

associations: Seslerio albicantis-Arctostaphyletum 
uvae-ursi (Sillinger 1933) nom. nov., Minuartio langii-Festucetum pallentis (Sillinger 1933) mucina ex 
Kliment et al. nom. nov., Seslerio variae-Caricetum approximatae Bernátová et Kliment 1982, Globulario 
cordifoliae-Caricetum humilis Bernátová et Uhlířová 
1994, Pulsatillo slavicae-Caricetum humilis (Sillinger 
1933) mucina ex Uhlířová et Bernátová 2004, Festuco tatrae-Caricetum humilis Uhlířová et Petrík nom. 
ined. [a description of this association will be published in another paper], Seslerio calcariae-Festucetum 
tatrae Sillinger 1933, Diantho nitidi-Caricetum tatrorum (Sillinger 1933) Kliment et al. 2005. 

seslerioalbicantis-Arctostaphyletumuvae-ursi 
(Sillinger 
1933) nom. nov. hoc loco 

(tab. 1, column 4)
Bas.: Arctostaphyletum fatrense Sillinger 1933 (art. 
34a)
characteristic taxon: Arctostaphylos uva-ursi (dom.
)
constant companions: Sesleria albicans (subdom.)
, 
Festuca tatrae, Gentiana clusii, Phyteuma orbiculare, 
Primula *hungarica, Pulsatilla slavica, Scabiosa lucida, 
Thesium alpinum, Thymus *sudeticus
nomenclatural type: Sillinger 1933: 218, r. 3, lectotypus hoc loco


a less known grass- dwarf shrub community with 
dominant species Arctostaphylos uva-ursi, forms in 
suitable places large, floristically relatively poor 
stands (on average 24 species per relevé) on ridges, 
terraces and tops of calcite-dolomite cliffs in montane to subalpine belt. its physiognomy is complemented beside the constantly occurring grasses 
(Sesleria albicans, Festuca tatrae) by several lively flowering herbs, such as Coronilla vaginalis, Gentiana 
clusii, Gentianella fatrae, Phyteuma orbiculare, Primula 
*hungarica, Pulsatilla slavica and Scabiosa lucida. 
Ridge climate is manifested in the constant presence of calciphile oreophytes and humicolous species like Hieracium bifidum, Vaccinium vitis-idaea, V. 
myrtillus, Luzula luzuloides and Homogyne alpina. lichens, mainly species Cetraria islandica and species 
of the genus Cladonia, reach the higher cover. the 
community was described from the nízke tatry 
mts (Sillinger 1933), later it was found and ana-
lysed also in the veľká Fatra mts (Bernátová & 
Uhlířová ined.). 

Minuartio langii-Festucetum pallentis 
(Sillinger 1933) 
mucina ex kliment et al. hoc loco 

(tab. 1, column 5) 
Bas.: Festucetum pallentis carpaticum Sillinger 1933 
(art. 34a) 
Syn.: Festuca *pallens-Minuartia laricifolia-ass. Sillinger 1933 pro syn. (art. 3a, 43); Pulsatillo slavici-Festucetum pallentis (Sillinger 1933) mucina et Petrík 
in Petrík 1977 nom. ined. (art. 1); Minuartio langii-
Festucetum pallentis (Sillinger 1933) mucina in mucina et maglocký 1985 nom. nud. 
characteristic taxon: Festuca pallens (transgr., dom.) 
Differential taxon: Campanula carpatica 
constant companions: Asplenium ruta-muraria, Jovibarba globifera subsp. glabrescens, Saxifraga paniculata, 
Seseli osseum, Sesleria albicans, Tithymalus cyparissias; 
Homalothecium philippeanum, Tortella tortuosa 

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J. Kliment et al.: Syntaxonomy and nomenclature of tHe communitieS of tHe allianceS Astero Alpini-seslerion CAlCAriAe … in Slovakia 
nomenclatural type: Sillinger 1933: 181, r. 3, lectotypus 
nomenclatural note: in mucina & maglocký (1985: 
187) the name Minuartio langii-Festucetum pallentis 
(Sillinger 1933) mucina nom. nov. was published 
withspecificationofbazionym,butrelevantworkwas 
not cited in the references (art. 2b). therefore the 
validisation of this name is necessary and proper. 

a detailed description of the community was 
published by Sillinger (1933), later also by Petrík 
(1977, 1978). it occupies steep, south to south-east 
oriented calcite-dolomite slopes or rocky towers, 
small terraces and jags in the montane belt, approx. 700–1250 m a. s. l. Shallow, generally with 
discontinuous a-horizon soils are skeleton and humus rich, crumby, with alkaline soil reaction. open 
stands of the species Festuca pallens form a species 
medium rich community with regular presence of 
several calciphile dealpine and prealpine taxa and 
subthermophile elements. 

Basedonrecogniseddifferencesinfloristiccomposition two subassociations were distinguished: 

Minuartio langii-Festucetum pallentis campanuletosum 
carpaticae subass. nov. hoc loco 

(tab. 1, column 5a) 
Differential taxa: Campanula carpatica, Acinos alpinus, Allium ochroleucum, Asplenium trichomanes, Bupleurum falcatum, Cyanus triumfettii, Erysimum witmannii, Hieracium bupleuroides, Polygonatum odoratum, 
Pulsatilla slavica, Thymus *sudeticus, Vincetoxicum hirundinaria; Anomodon viticulosus, Encalypta streptocarpa, E. vulgaris, Fissidens dubius, Orthotrichum anomalum, Rhytidium rugosum 
nomenclatural type: identical with the type of the 
association name 

afloristically richer (average 36 taxa per relevé), 
more hydrophilic community with abundant presence of mosses was found in the nízke tatry mts 
and the Slovenský raj mts. 

Minuartio langii-Festucetum pallentis campanuletosum 
xylocarpae subass. nov. hoc loco 

(tab. 1, column 5b) 
Differential taxa: Campanula xylocarpa, Daphne arbuscula, Erysimum odoratum, Pulsatilla subslavica; Cladonia pyxidata, Peltigera rufescens 
nomenclatural type: valachovič & mucina 2004, 
tab. 1, r. 7, holotypus 

a more xerophile subassociation with an average number of taxa per relevé of only 27 was found 
in calcareous cliffs of the muránska planina mts. 

seslerion tatrae Pawłowski 1935 corr. klika 1955 

(tab. 1, columns 6–8)
Syn.: Seslerion bielzii Pawłowski 1935 (art. 43); Seslerion tatrae Pawłowski 1956 (art. 8, 31), hadač et 
Smola 1962 (art. 8, 31); Seslerion tatrae Pawłowski 
emend. hadač 1962 (art. 8, 31)
Syntax. syn.: Seslerion bielzii Pawłowski 1935 emend. 


a. nyárády 1967 p. p. (excl. Festucion marmarossicae
Pawłowski et Walas 1949)
Pseud.: Seslerion coerulae sensu auct. carpat. occid. 
p. p., non Br.-Bl. in Br.-Bl. et Jenny 1926 
non: Festuco saxatilis-Seslerion bielzii (Pawłowski et 
Walas 1949) coldea 1984 
Phantom: Seslerion tatrae hadač 1962 (mucina & 
maglocký 1984, 1985) 
characteristic taxa: Sesleria tatrae (dom.), Astragalus 
alpinus, A. australis (transgr.), Euphrasia tatrae, Gentiana verna, Tephroseris capitata, Trifolium *kotulae 
Differential taxa: Achillea millefolium subsp. alpestris, 
Anthoxanthum alpinum, Avenula versicolor, Ligusticum 
mutellina, Linum extraaxillare, Parnassia palustris, Phleum hirsutum, Potentilla aurea, Primula elatior 
nomenclatural type: Seslerietum bielzii Domin 1929, 
lectotypus (art. 20) 
the alliance Seslerion tatrae merges basiphilous 
and mesophilous, species rich, flowery communities of short grasses, which prefer lee slopes and 
habitats with long-term snow on carbonate bedrock 
in the subalpine and alpine belt. their distribution 
is limited to high mountains of the Western carpathians (Krivánska malá Fatra, veľká Fatra, nízke, 
Západné and Belianske tatry). this fact is stressed 
by name-giving species – West carpathian subendemic Sesleria tatrae. 

togetherwiththisspeciesseveralmoreendemic 
taxa were found in analysed stands. endemics and 
subendemics of the Western carpathians and the 
carpathians are highlighted in the table (tab. 1). 

Pawłowski (1935) divided the West carpathian 
high mountain communities of the order Seslerietalia coerulae to the individual alliance based on the 
sharp differences in their structure against the relevant communities from the alps and mountains 
of Balkan Peninsula. in accordance with knowledge of that time he proposed for this alliance the 
name Seslerion bielzii [taxon Sesleria tatrae was originally distinguished in hierarchic level of form as 

S. bielzii f. tatrae Degen; till Deyl (1946) reclassified 
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Hacquetia 4/2 • 2005 

it up to level of species]. these changes in taxonomic evaluation were for the first time reflected 
by Klika (1955), who published in a survey of syntaxa of czechoslovakia the name Seslerion (Bielzii) 
tatrae Pawł. 1935. Pawłowski (1956) used directly 
the name Seslerion tatrae in characteristics of the 
vegetation units of the tatry mts, but the name of 
alliance and subordered associations he published 
without author’s citations. 

Diantho nitidi-seslerietum tatrae 
Bělohlávková ass. 
nov. hoc loco 

(tab. 3; tab. 1, column 6) 
Syn.: Polygono vivipari-Seslerietum tatrae (Šmarda 
1956) Bělohlávková 1980 (art. 1) 
non: Seslerietum tatrae Šmarda 1956 
Differential taxa: Dianthus nitidus subsp. nitidus, 
Knautia arvensis, Polygonatum verticillatum, Ranunculus nemorosus, Viola canina 
constant companions: Sesleria tatrae (dom.), Carex 
*tatrorum (subdom.-dom.), Bartsia alpina, Bellidiastrum michelii, Campanula serrata, Carlina acaulis, Festuca *versicolor, Galium anisophyllon, Leucanthemum 
vulgare agg., Pedicularis verticillata, Phyteuma orbiculare, Primula elatior, Scabiosa lucida, Soldanella carpatica 

nomenclatural type: tab. 3, r. 2, holotypus 

the community is represented by short grass, 
nearly completely closed, floristically and physiognomically various stands with more or less equal 
presence of the (sub-) species Sesleria tatrae and 
Carex *tatrorum, locally also Festuca *versicolor. their 
grassy look is vivified during the vegetation period 
by varicoloured flowers of many species. mosses 
and lichens are presented only sparsely. 

typical habitats of the community are plain to 
slightly concave slopes with gentle inclination, usually at the foot of steeper slopes or under the top 
cliffs in the subalpine belt of the malá Fatra mts. 
During winter they are protected by thick snow, 
which holds over up to the end of may. Sufficiency 
of soil moisture is ensured both by the melting 
snow and also by running water from higher situated parts of slopes and channels among rocks. 

these relatively sizeable differences in synecology and floristic composition we classify at the level 
of variants with regard to the limited area and small 
number of relevés. 

typical variant (tab. 3, r. 1–22) covers typical 
hydro- and chionophilous stands of the association 
with the higher presence of acidophilous and neu

trophilous species, which are adapted to the short 
vegetation period. 

homogeneous large stands of the variant with 
Allium *montanum (tab. 3, r. 23–29; Differential 
taxa: Allium *montanum, Carduus glaucinus, Gentiana clusii, Hieracium villosum, Jovibarba globifera, 
Libanotis pyrenaica, Polygala *brachyptera, Ranunculus 
breyninus, Valeriana tripteris) occupy regular south to 
south-east oriented slopes with inclination 30–40° 
below or among rocky walls at altitude approx. 
1520–1630 m. 

seslerietum tatrae 
domin 1929 nom. corr. hoc loco 

(tab. 1, column 7) 
oFn: Seslerietum bielzii Domin 1929 (art. 43) [Domin 
1929: 9] 
Syn.: Seslerietum bielzii Deyl 1936 (art. 31, 43); Seslerietum tatrae Šmarda 1956 (art. 31), Bareš et hadač 
1958 (art. 2b, 31), hadač et Smola 1962 (art. 2b, 
31) 
characteristic taxa: Cardaminopsis halleri subsp. tatrica, Tephroseris capitata (transgr.) 
Differential taxa: Anemone narcissiflora, Festuca picturata, Geranium sylvaticum, Homogyne alpina, Oreogeum montanum, Viola lutea subsp. sudetica 
constant companions: Sesleria tatrae (dom.), Carex 
*tatrorum (subdom.), Achillea *alpestris, Alchemilla 
sp. div., Anthoxanthum alpinum, Bartsia alpina, Bistorta vivipara, Campanula tatrae, Galium anisophyllon, 
Helianthemum grandiflorum, Leontodon hispidus, Ligusticum mutellina, Myosotis alpestris, Parnassia palustris, Phleum hirsutum, Phyteuma orbiculare, Potentilla 
aurea, Primula elatior, Ranunculus breyninus, Scabiosa 
lucida, Soldanella carpatica 

closed, two-layered, species rich community of 
short grasses dominated by species Sesleria tatrae 
and Carex *tatrorum. there are many versicolour 
flowering herbs among the constantly presented 
species. the favourable water regime is reflected in 
higher cover of mosses. 

the community occupies shallow depressions 
on steep south oriented and from wind protected 
slopes in the subalpine to alpine belt approx. 
(1650) 1750–2050 m a. s. l. most of the localities 
are concentrated in altitude between 1900–2000 m 

a. s. l. in these habitats snow holds over for 7–7.5 
months (hadač & Smola 1962). 
the high constancy of the species Achillea *alpestris, Anemone narcissiflora, Campanula tatrae, Crepis 
mollis, Festuca picturata, Geranium sylvaticum, Luzula 
luzuloides subsp. rubella and Oreogeum montanum to

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J. Kliment et al.: Syntaxonomy and nomenclature of tHe communitieS of tHe allianceS Astero Alpini-seslerion CAlCAriAe … in Slovakia 
gether with the respectable presence of species Avenula versicolor indicates a close relation of the association to the communities of the order Calamagrostietalia villosae (class Mulgedio-Aconitetea – see 
Fig. 2). Within the alliance Seslerion tatrae it is closest to the associations Trifolio kotulae-Caricetum tatrorum and Seslerio tatrae-Festucetum versicoloris. transitions among them are frequent. 

Domin (1929) for the first time validly described 
high mountain chionophilous stands dominated 
by species Sesleria tatrae as individual associations, 
with regard to the taxonomical knowledge of the 
time (cf. Domin & Podpěra 1928), with the name 
Seslerietum bielzii. Šmarda (1956) used for the first 
time the correct name of taxon (Sesleria tatrae) in 
the published relevé of this community. this is perhaps why, in the later period in syntaxonomical literature, the name Seslerietum tatrae Šmarda 1956 
was used (cf. hadač & al. 1969; Šomšák & al. 1980; 
Unar & al. 1984, 1985; mucina & maglocký 1985). 
Some authors mentioned the name Seslerietum bielzii Domin 1929 only as a synonym. Based on results 
of the syntaxonomical revision we can state that 
Domin’s relevé from havran mt. in the Belianske 
tatry mts in its floristical composition completely 
conforms to the actual conception of the association Seslerietum tatrae. in accordance to art. 43 icPn 
we propose the correction of the original name. 

Based on floristic composition, soil ecology and 
chorology two subassociations were distinguished: 

seslerietum tatrae luzuletosum rubellae 
subass. nov. 
hoc loco 

(tab. 1, column 7a) 
Differential taxa: Achillea *alpestris, Campanula elliptica, Cortusa matthioli, Crepis conyzifolia, C. mollis, Cyanus mollis, Hedysarum hedysaroides, Linum extraaxillare, Luzula *rubella, Myosotis alpestris, Phyteuma spicatum, Ranunculus pseudomontanus, R. thora, Thymus 
alpestris, Trommsdorfia uniflora 
nomenclatural type: identical with the type of the 
association name 

more mesophilous subassociation occupies medium deep (to deep), fresh moist to moist, loamy to 
slightly clayey soils on calcite, dolomite, calcareous 
clay or marl bedrock. Skeleton is accumulated in 
the low part of the soil profile. components of 
mountain tall-herb communities and alpine grasslands differ this subassociation against the following 
one. its centre of distribution lies in the Belianske 
tatry mts, it was also found scarcely in the nízke 

tatrymts.thestanddocumentedby(Šmarda1956) 
on the south slope of Bujačí mt., 1920 m a. s. l. takes 
a transition position between both subassociations. 

seslerietum tatrae biscutelletosum laevigatae subass. 
nov. hoc loco 

(tab. 1, column 7b) 
Differential taxa: Biscutella laevigata, Anthyllis *alpestris, Hieracium villosum, Selaginella selaginoides, Thymus pulcherrimus 

nomenclatural type: Unar & al. 1984, tab. 19, r. 3, 
holotypus 

Stands of the subassociation occupy medium 
deep soils on calcite and dolomite bedrock with a 
higher portion of coarse grained skeleton. they 
are distributed in the Západné tatry mts. 

seslerio tatrae-Festucetum versicoloris 
Pawłowski et 
Stecki 1927 nom. corr. hoc loco 

(tab. 1, column 8)
oFn: Festuca varia-Sesleria bielzii-ass. Pawłowski et 
Stecki 1927 (art. 43)
Syn.: Festucetum variae calcicolum Szafer et al. 1923 


p. p. (art. 34a, 43); Festuca varia-Sesleria bielzii-ass. 
Szafer et al. 1927 (art. 2b), Pawłowski 1928 (art. 
2b); Varietum tatricum Szafer et al. 1927 pro syn. 
(art. 2b, 3a), Pawłowski et Stecki 1927 pro syn. (art. 
3a, 34a), Pawłowski 1928 pro syn. (art. 2b, 3a); Sesleria bielzii-Festuca versicolor-ass. Br.-Bl. 1930 p. p. 
(art. 43), vierhapper 1930 (art. 2b, 43), Pawłowski 
1935 (art. 3a, 31, 43); Versicoloretum tatricum vierhapper 1930 pro syn. (art. 2b, 3a, 34a), Sillinger 
1933 (art. 34a), Pawłowski 1935 (art. 34a), Pawłowski 
1956 (art. 2b, 31, 34a); Festucetum versicoloris tatrense 
Deyl 1936 (art. 2b, 34a); Festucetum versicoloris 
Šmarda 1956 (art. 31) 
non: Festucetum versicoloris Domin 1929; Festucetum 
versicoloris Domin 1933; Diantho-Festucetum versicoloris hadač et al. 1969 
Pseud.: Seslerietum bielzii sensu Br.-Bl. 1930 p. p. , 
non Domin 1929 
Differential taxa: Androsace chamaejasme, Campanula alpina, Carex atrata, C. firma, Cerastium eriophorum, Erigeron hungaricus, Minuartia pauciflora (Kit. 
ex Kanitz) Dvořáková (syn.: M. gerardii auct. car-
pat. non Willd.), M. sedoides, Pedicularis oederi, Salix 
alpina, S. reticulata, Saxifraga *dominii, Selaginella 
selaginoides, Silene acaulis subsp. longiscapa; Cetraria 
islandica 
constant companions: Festuca *versicolor (dom.), 
131 


Hacquetia 4/2 • 2005 

Sesleria tatrae (subdom.), Carex *tatrorum (subdom.), 
Bistorta vivipara, Campanula tatrae, Galium anisophyllon, Parnassia palustris, Phyteuma orbiculare, Poa alpina, Saxifraga paniculata, Scabiosa lucida, Soldanella 
carpatica, Thymus pulcherrimus; Tortella tortuosa 
nomenclatural type: Szafer & al. 1923, tab. 11, r. 2, 
lectotypus (Unar & al. 1985: 34) 

a short stalk, open, species rich community 
with important presence of grasses and sedges. a 
detailed description was published by Pawłowski & 
Stecki (1927) and Pawłowski (1935). the community occupies prevailingly south oriented rocky 
slopes, stabilized carbonate screes, small rocky terraces and cornices of steep rocky walls in the subalpine and alpine belt. Soils below the stands are 
very shallow to shallow, humus and skeleton rich, 
with neutral to alkaline reaction (in younger development stadiums up to strong alkaline), in the upper horizon slightly acid due to decalcification. 
Snow is not too thick, it melts early in spring. the 
soil surface together with vegetation is consequently exposed to severe fluctuations of temperature. 
the influence of wind is low because of wind protected habitats. 

in some syntaxonomical surveys (e. g. matuszkiewicz 1982, mucina & maglocký 1985) this community was mentioned as Seslerio tatrae-Festucetum 
versicoloris (Szafer et al. 1923) 1927. Szafer & al. 
(1927: 66) otherwise proposed for the association 
Festucetum variae calcicolum nomen novum (Festuca 
varia-Sesleria Bielzii-assoziation), however without 
an unambiguous bibliographical citation of the 
place of the valid publishing of the rejected name: 
“Früher haben wir diese assoziation ‘Festucetum 
variae calcicolum’ genannt. Um missverständnisse 
zu vermeiden, ziehen wir die nun gewählte, mehr 
präzise Benennung vor.” 

the association is closely related to the communities of the alliance Caricion firmae, in the highest 
localities to the communities of the alliance Oxytropido-Elynion Br.-Bl. 1949 (class Carici rupestris-Kobresietea bellardii ohba 1974 – see Fig. 2). it is distributed on calcite, dolomite and marl limestone 
bedrock in the Západné and Belianske tatry mts. 
Based on floristic differences two subassociation 
were distinguished: 

seslerio tatrae-Festucetum versicoloris ranunculetosum 
pseudomontani subass. nov. hoc loco 

(tab. 1, column 8a)
Differential taxa: Ranunculus pseudomontanus, An

thoxanthum alpinum, Bartsia alpina, Bistorta major, 
Botrychium lunaria, Cerastium arvense subsp. glandulosum, Festuca supina, Gentiana verna, Ligusticum 
mutellina, Linum extraaxillare, Luzula luzuloides, Myosotis alpestris, Pedicularis verticillata, Phleum hirsutum, 
Primula elatior subsp. tatrensis, Rhodiola rosea, Thymus 
pulcherrimus, Vaccinium vitis idaea 
nomenclatural type: identical with the type of association name 

the subassociation is represented by large, species rich stands (on average 49 taxa per relevé), 
which occupy gravel to coarse-grained screes on 
south-west to south-east oriented slopes with inclination 20–35 °. a group of differential taxa is 
formed by species of alpine meadows and carbonate screes. Stands in the highest elevations in the 
Západné and Belianske tatry, approx. 1880–2140 
m a. s. l., were divided within individual variant by 
Erigeron hungaricus (Differential taxa: Erigeron hungaricus, Avenula versicolor, Ligusticum mutellinoides, 
Lloydia serotina, Luzula spicata subsp. mutabilis, Oreogeum montanum, Pulsatilla scherfelii, Rhodax alpestris, 
Salix retusa, Tephroseris capitata, Cetraria cucullata, C. 
nivalis, Cladonia rangiferina, Thamnolia vermicularis). 
in the subalpine belt the community beside the 
screes occupies also steep, weathering rock walls 
with various orientation. consequently, the stands 
are enriched by species which prefer a rocky background, such as Crepis jacquinii, Dianthus *praecox, 
Euphrasia salisburgensis, Gentiana clusii, Trisetum alpestre, Veronica aphylla and V. fruticans. 

seslerio tatrae-Festucetum versicoloris inops subass. 
nov. hoc loco 

(tab. 1, column 8b)
nomenclatural type: Unar & al. 1984, tab. 16, r. 9, 
holotypus


Floristically poorer stands with average number 
of species 37 were found on small terraces of calcite rocks in the subalpine belt in the červené vrchy 
mts. in comparison to the previous subassociation 
several species with stronger affinity to the rocky 
substratum are presented with higher frequency: 
Asplenium ruta-muraria, Pinguicula alpina and Saxifraga wahlenbergii. the more frequent occurrence 
of some mosses (Lophozia bantriensis, Meesia uliginosa) can be conditioned by insufficient determination of cryptogam flora in the beginning of phytocoenological studies, so it was not regarded in the 
differentiation of communities. 

132 


J. Kliment et al.: Syntaxonomy and nomenclature of tHe communitieS of tHe allianceS Astero Alpini-seslerion CAlCAriAe … in Slovakia 
acKnoWleDGementS


authors are indebted to Ján Ripka from Daphne 
institute of applied ecology, Bratislava, for processing of the map. this work was supported by 
grants from Grant agency veGa 1/0045/03 and 
1/2347/05 and also by the project no. 07G0208 
a040 Biodiverzita 002. 

ReFeRenceS 

Bareš, m. & hadač, e. 1958: Poznámky o snehových pomeroch v Doline siedmich prameňov vo 
vzťahu k vegetácii a lavínam. Sborn. Prác o tatransk. nár. Parku 2: 20–29. 

Barkman, J. J., Doing, h. & Segal, S. 1964: Kritische 
Bemerkungen und vorschläge zur quantitativen vegetationsanalyse. acta Bot. neerl. 13: 
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J. Kliment et al.: Syntaxonomy and nomenclature of tHe communitieS of tHe allianceS Astero Alpini-seslerion CAlCAriAe … in Slovakia 
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768. 
aPPenDiX 

Sources to Tab. 1: 

1 
hadač & al. 1969, p. 67 (Astereto-Seslerietum calcariae), 5 r.; Petrík ined., 10 r., the Belianske tatry mts. 

2 
hadač & al. 1969, p. 72 (Diantho-Festucetum versicoloris), 5 r., p. 76 (Tortello-Festucetum tatrae), r. 
221, 257; Šmarda 1956, p. 24 (Festucetum tatrae), 
1 r.; Petrík ined., 4 r., the Belianske tatry mts. 

3 
Kliment & al., tab. 2 (Gentiano clusii-Festucetum 
versicolor), 14 r. 

4 
Sillinger 1933, p. 218 (Arctostaphyletum fatrense), 
3 r.; Bernátová & Uhlířová ined., 3 r., veľká 
Fatra; Bernátová & obuch ined., 2 r., the nízke 
tatry mts. 

5a Sillinger 1933, p. 181 (Festucetum pallentis carpaticum), 9 r.; hajdúk 1958, tab. 14, r. 4 (Festucetum pallentis carpaticum typicum); Petrík 1977, 
tab. 3 (Pulsatillo slavici-Festucetum pallentis), r. 
10–16; Petrík 1978, tab. 3 (Festucetum pallentis 
carpaticum), 23 r.; Petrík & al. 1982, tab. 1, r. 11 
(Festucetum pallentis carpaticum). 

5b valachovič & mucina 2004, tab. 1, r. 1–3, 5, 7–9 
(Minuartio langii-Festucetum palentis), r. 11, 12, 
15 (Seslerio-Festucetum tatrae). 

6 
Kliment & al., tab. 3 (Diantho nitidi-Seslerietum 
tatrae), 29 r. 

7a Domin 1929, p. 8 (Seslerietum bielzii), 1 r.; Šmarda 
1956, r. 20 (Seslerietum tatrae), 1 r.; hadač & al. 
1969, p. 108 (Seslerietum tatrae), r. 23, 25, 124, 
152; Petrík ined., 1 r., the nízke tatry mts, 11 r., 
the Belianske tatry mts. 

7b Šomšák & al. 1980, tab. 2 (Carduo glauci-Caricetum tatrorum), r. 11, 12, tab. 3 (Seslerietum tatrae), r. 1, 3; Unar & al. 1984, tab. 18 (Carduo 
glauci-Caricetum tatrorum), r. 3, 6, tab. 20 (Seslerietum tatrae), 5 r.; Petrík ined., 1 r., the Západné 
tatry mts. 

8a Szafer & al. 1923, tab. 11 (Festucetum variae calcicolum), r. 2, 4, 5; Pawłowski & Stecki 1927, 
tab. 4 (Festuca varia-Sesleria Bielzii-ass.), r. 3–10; 
Domin 1929, p. 8 (Festucetum versicoloris), 1 r.; 
Braun-Blanquet 1930, p. 106 (Sesleria bielzii-Festuca versicolor-ass.), r. 3; Pawłowski 1935, tab. 1 
(Versicoloretum tatricum), r. 1–10; Šmarda & al. 
1971, tab. 8 (Festucetum versicoloris calcicolum), r. 
57, tab. 10 (Carduo glauci-Caricetum tatrorum), r. 
91, 97; Šomšák & al. 1980, tab. 2 (Carduo glauci-
Caricetum tatrorum), r. 6; Unar & al. 1984, tab. 16 
(Festucetum versicoloris calcicolum), r. 2, 7. 

8b Unar & al. 1984, tab. 16, r. 3, 4, 6, 8–10. 

localities of relevés: 
Tab. 2, Gentiano clusii-Festucetum versicoloris:


(r. 1–9 = the Krivánska malá Fatra mts, r. 10–14 = 
the chočské vrchy mts) 
1. 
the Steny mt., outcrops of marl limestone below 
the ridge trace; 1570 m a. s. l., Se, 60 °, 12 m2, 
e1: 90 %, e0: 5 %, 23. 7. 2003, Bernátová & 
Uhlířová. 
2. 
the chleb mt., west rocky slope of amphitheatre with rocky outcrops; 1625 m a. s. l., W, 85 °, 
25 m2, e1: 75 %, e0: 10 %, 18. 8. 1975, 
Bělohlávková. 
3. 
ridge Koniarky – the malý Kriváň 
mt., cliff in 
north slope; 1550 m a. s. l., nW, 80 °, 25 m2, e1: 
75 %, e0: 15 %, 8. 8. 1977, Bělohlávková. 
4. 
narrow ridge between the chleb mt. and the 
hromové mt.; 1620 m a. s. l., S, 80 °, 14 m2, e1: 
60 %, e0: 10 %, 22. 8. 2001, Bernátová & Kliment. 
135 


Hacquetia 4/2 • 2005 

5. 
ridge Koniarky – the malý Kriváň 
mt., cliff in 
north slope; 1540 m a. s. l., n, 75 °, 25 m2, e: 
85 %, e0: 15 %, 17. 8. 1978, Bělohlávková. 
6. 
ridge Koniarky – the malý Kriváň 
mt., cliff near 
the ridge trace (the first from the saddle below 
the Koniarky), 1550 m a. s. l., WSW, 80 °, 25 m2, 
e1: 85 %, e0: 2 %, 16. 8. 1975, Bělohlávková. 
7. 
ridge Koniarky – the malý Kriváň 
mt., cliff near 
the ridge trace (second from the saddle below 
the Koniarky); 1555 m a. s. l., S, 90 °, 25 m2, e: 
70 %, e0: 5 %, 17. 8. 1975, Bělohlávková. 
8. 
ridge Koniarky – the malý Kriváňmt., cliff above 
the saddle below the Koniarky; 1550 m a. s. l., 
SSe, 70 °, 25 m2, e1: 75 %, e0: 2 %, 26. 7. 1973, 
Bělohlávková. 
9. 
ridge Koniarky – the malý Kriváň 
mt., cliff on 
north slope; 1555 m a. s. l., SSW, 50 °, 25 m2, e: 
80 %, e0: 5 %, 17. 8. 1975, Bělohlávková. 
10.the veľký choč mt. (1607,7 m); 1470 m a. s. l., 
SSe, 45 °, 15 m2, e1: 75 %, 29. 7. 1965 (ŠvandováUrsínyová 1966, r. 62). 
11.the veľký choč mt.; 1480 m a. s. l., SSe, 45 °, 
8 m2, e1: 75 %, e0: 5 %, 29. 7. 1965 (ŠvandováUrsínyová 1966, r. 64). 
12.the veľký choč mt.; 1470 m a. s. l., eSe, 65 °, 
10 m2, e1: 75 %, 29. 7. 1965 (Švandová-Ursínyová 1966, r. 63). 
13.the veľký choč mt., rocky wall; 1478 m a. s. l., 
ne, 80 °, 12 m2, e1: 85 %, 29. 7. 1965 (ŠvandováUrsínyová 1966, r. 66). 
14.the veľký choč mt.; 1500 m a. s. l., S, 45 °, 21 m2, 
e1: 30 %, 29. 7. 1965 (Švandová-Ursínyová 1966, 
r. 67). 
Tab. 3, Diantho nitidi-Seslerietum tatrae 

(r. 1–29 = the Krivánska malá Fatra mts) 
1. the malý Kriváň 
mt. (1670,9 m), north slope; 
1510 m a. s. l., n, 20 °, 25 m2, e1: 95 %, 11. 6. 
1986 (milová & Urbanová 1989: 302 ut Polygono 
vivipari-Seslerietum tatrae). 
2. ridge between the malý Kriváň 
mt. – the Koniarky mt., north slope; 1520 m a. s. l., nW, 30 °, 
25 m2, e1: 98 %, e0: 2 %, 22. 7. 1973, 
Bělohlávková. 
3. ridge between the malý Kriváň 
mt. – the Koniarky mt., north-west slope; 1500 m a. s. l., nW, 
25 °, 25 m2, e1: 100 %, 28. 6. 1984 (milová & 
Urbanová 1989: 303 ut Polygono vivipari-Seslerietum tatrae). 
4. ridge between the hromové mt. – the Severné 
Steny mt., west slope; 1600 m a. s. l., W, 30 °, 
1

25 m2, e1: 100 %, e0: 10 %, 14. 8. 1973, 
Bělohlávková.

 5. the Pekelník mt. (1609,3 m), north slope below 
the summit; 1575 m a. s. l., n, 25 °, 25 m2, e1: 
95 %, e0: 3 %, 10. 8. 1973, Bělohlávková. 
6. chrapáky, plain topic of the ridge in the north 
slope, enclave in dwarf pine; 1475 m a. s. l., n, 
2°, 25 m2, e: 95 %, 9. 8. 1973, Bělohlávková.
1 
1

 7. chrapáky, near the spring on north slope; 1400 
m a. s. l., nW, 25 °, 25 m2, e1: 95 %, e0: 10 %, 9. 
8. 1973, Bělohlávková. 
8. the malý Kriváň 
mt., ridge to the saddle Priehyb, south slopes; 1595 m a. s. l., SSW, 35 °, 
25 m2, ec: 85 %, e1: 80 %, e0: 5 %, 18. 7. 2001, 
Šibík & Krajčiová-Šibíková. 49°10'56,8" n. l.,
1

18°59'25,1" e. l. 

9. 
the malý Kriváňmt., east slope; 1624 m a. s. l., e, 
30 °, 25 m2, ec: 100 %, e1: 100 %, e0: 1 %, 31. 7. 
2002, Bernátová, Šibík & Uhlířová. 49°11'02,0" 
n. l., 18°59'52,5" e. l. 
10.the malý Kriváň 
mt., ridge part; 1649 m a. s. l., 
nnW, 30 °, 16 m2, ec: 100 %, e1: 100 %, e0: 5 %, 

31. 7. 2002, Bernátová, Šibík & Uhlířová. 
49°10'56,8" n. l., 18°59'58,1" e. l. 
11.the malý Kriváň 
mt., below the summit; 1631 m 

a. s. l., nnW, 30 °, 16 m2, ec: 100 %, e1: 100 %, 
e0: 0 %, 31. 7. 2002, Bernátová, Šibík & Uhlířová. 
49°11'01,2" n. l., 18°59'50,7" e. l. 
12.saddle vráta (1462 m), north slope, enclave in 
dwarf pine; 1400 m a. s. l., nW, 25 °, 25 m2, e1: 
95 %, e0: 20 %, 17. 7. 1973, Bělohlávková. 

13.saddle vráta, north slope, near the tourist trace; 
1375 m a. s. l., nnW, 20 °, 25 m2, e1: 98 %, e0: 
5 %, 21. 7. 1973, Bělohlávková. 

14.the veľký Kriváň 
mt., shallow depression on 
south slope; 1625 m a. s. l., S, 30 °, 25 m2, e1: 
100 %, e0: 10 %, 11. 8. 1973, Bělohlávková. 

15.the veľký Kriváňmt., south slope; 1570 m a. s. l., 
S, 25 °, 25 m2, e1: 98 %, e0: 5 %, 25. 7. 1973, 
Bělohlávková. 

16.the veľký Kriváňmt., channel on the west slope; 
1500 m a. s. l., SW, 35 °, 25 m2, e1: 98 %, 11. 8. 
1973, Bělohlávková. 

17.the chleb mt. (1645,6 m), south-west slope 
above the trace; 1522 m a. s. l., SSW, 40 °, 20 m2, 
e1: 95 %, 28. 7. 2002, Bernátová & Uhlířová. 
49°11,352' n. l., 19°02,520' e. l. 

18.the hromové mt. (1636,2 m), south slope; 1624 
m a. s. l., SSW, 30 °, 25 m2, e1: 90 %, 29. 7. 2002, 
Bernátová & Uhlířová. 49°11,387' n. l., 19°03,302' 

e. l. 
136 


J. Kliment et al.: Syntaxonomy and nomenclature of tHe communitieS of tHe allianceS Astero Alpini-seslerion CAlCAriAe … in Slovakia 
19.the hromové mt., west from r. 18; 1620 m a. s. l.
, 
25.the chleb mt., south slope; 1610 m a. s. l., SSe, 
SSe, 30 °, 25 m2, e1: 100 %, 29. 7. 2002, Berná-40 °, 25 m2, e1: 95 %, 30. 7. 2002, Bernátová & 
tová & Uhlířová. 49°11,409' n. l., 19°03,414' e. l. Uhlířová. 
20.the veľký Kriváň 
mt., shallow channel on south-26.the chleb mt., south slope; 1596 m a. s. l., S, 
west slope; 1575 m a. s. l., SW, 35 °, 25 m2, e1: 40°, 24 m2, e1: 100 %, 29. 7. 2002, Bernátová & 
95 %, 11. 8. 1973, Bělohlávková. Uhlířová. 49°11,244' n. l., 19°03,149' e. l. 
21.ridge between the malý Kriváň 
mt. – the Ko-27.the chleb mt., channel on south-east slope; 
niarky mt., south slope; 1550 m a. s. l., Se, 30 °, 1524 m a. s. l., SSe, 35°, 24 m2, e1: 90 %, 30. 7. 
25 m2, e1: 90 %, 17. 8. 1973, Bělohlávková. 2002, Bernátová & Uhlířová. 49°11,199' n. l., 
22.the 
chleb mt., depression with boulders on 19°03,193' e. l. 
south slope; 1300 m a. s. l., eSe, 35 °, 25 m2, e1: 28.the chleb mt., SSe slope below rocky walls; 
95 %, e0: 1 %, 17. 8. 1973, Bělohlávková. 1572 m a. s. l., 30 °, 24 m2, e1: 90 %, 30. 7. 2002, 
23.the chleb mt., south slope below the summit; 
Bernátová & Uhlířová. 49°11,220' n. l., 19°03,134' 
1632 m a. s. l., S, 40 °, 16 m2, e1: 100 %, 28. 7. e. l. 
2002, Bernátová & Uhlířová. 49°11,259' n. l., 29.the Stienky mt., Se slope below the summit; 
19°03,117' e. l. 1612 m a. s. l., Jv, 20 °, 24 m2, e1: 85 %, e0: 15 %, 
24.the chleb mt., south slope; 1625 m a. s. l., SSe, 
1. 8. 2002, Bernátová & Uhlířová. 49°11,521' n. 
30 °, 25 m2, e1: 100 %, 29. 7. 2002, Bernátová & l., 19°03,673' e. l. 
Uhlířová. 49°11,271' n. l., 19°03,114' e. l. 
Recieved 13. 4. 2005 
Revision recieved 23. 9. 2005 
accepted 7. 10. 2005 

137 


Hacquetia 4/2 • 2005 

Table 1: Analysed communities of the class Elyno-Seslerietea from the Western Carpathians.
Tabela 1: Obravnavane združbe razreda Elyno-Seslerietea iz zahodnih Karpatov.
1 – Astero alpini-Seslerietum calcariae;2 – Diantho praecoci-Festucetum versicoloris;3 – Gentiano clusii-Festucetum versicoloris;4 – Seslerio albicantis-Arctostaphyletum uvae-ursi;5 – Minuartio langii-Festucetum pallentis, 5a 


– subass. campanuletosum carpaticae, 5b – subass. campanuletosum xylocarpae;6 – Diantho nitidi-Seslerietum 
tatrae;7 – Seslerietum tatrae, 7a – subass. luzuletosum rubellae, 7b – subass. biscutelletosum laevigatae;8 – Seslerio 
tatrae-Festucetum versicoloris, 8a – subass. ranunculetosum pseudomontani, 8b – subass. inops 
1–5 – Astero-Seslerion calcariae; 6–8 – Seslerion tatrae 
Number of column 1 2 3 4 5a 5b 5 6 7a 7b 7 8a 8b 8 
Number of relevés 15 12 14 10 41 10 51 29 18 12 30 28 6 34 
Average number of taxa 37 39 30 24 36 27 34 38 51 43 48 49 37 47 

Diagnostic taxa of the associations 
pc Oxytropis *tatrae [C] C1 47 5 17 3 . . . . . . . . . 4 2 . 3 2 
pc Gypsophila repens d1 93 5 42 4 7 2 10 2 . . . . . . . . . . 
pc Leontopodium alpinum d1 80 3 17 2 . . 5 3 . 4 2 . . . . . . . 
Fulgensia bracteata (E0) d1 47 2 . . . . . . . . . . . . . 
Squamarina gypsacea (E0) d1 47 3 . . . . . . . . . . . . . 
Hymenostylium recurvirostrum (E0) d1 40 3 . . . . . . . . . . . . . 
Gentianella lutescens d2 20 2 67 2 . . . . . 31 2 11 2 25 2 17 2 36 2 . 29 2 
Silene vulgaris d2 . 58 2 . . 7 2 . 6 2 3 2 11 2 . 7 2 . . . 
fc Festuca carpatica [C] d2 . 42 3 . . . . . 7 2 33 5 25 2 30 4 7 2 . 6 2 
Hieracium murorum d2 . 42 2 . . 2 2 10 2 4 2 10 2 . 17 2 7 2 4 1 . 3 1 
Bartsia alpina d3,8a . . 71 3 30 2 . . . 90 3 61 3 75 2 67 3 57 2 . 47 2 
ca Vicia oreophila d3 . . 64 2 . . . . 7 2 . . . . . . 
fv Hedysarum hedysaroides d3,7a . . 50 3 . . . . . 39 3 . 23 3 36 3 50 3 38 3 
fv Potentilla crantzii d3 . . 50 3 . . . . 55 3 6 2 8 3 7 3 61 3 33 4 56 3 
tf Rhodiola rosea d3,8a . . 50 3 . . . . 24 3 39 2 42 2 40 2 39 2 . 32 2 
cf Androsace lactea d3 . 17 2 50 2 . . . . 3 2 . . . 4 2 17 3 6 3 
ac, cf Ranunculus alpestris d3 . . 43 2 . . . . . 6 2 . 3 2 11 2 33 3 15 2 
fv Saxifraga *dominii [CW] d3,8 . . 29 2 . . . . . . . . 61 2 50 2 59 2 
Arctostaphylos uva-ursi C4 . . . 100 8 . . . . . . . . . . 
Festuca pallens t4 . . . 10 2 100 6 100 6 100 6 . . . . . . . 
cy Campanula carpatica [C] d5,5a . . . . 83 2 . 67 2 . . . . . . . 
Sc,ES Dianthus *nitidus [CW] d6 . . 7 2 . . . . 90 3 . . . . . . 
Knautia arvensis d6 . . 7 2 . 20 2 . 16 2 66 3 . 8 3 3 3 . . . 
Ranunculus nemorosus d6 . . . . . . . 41 2 . . . . . . 
Viola canina d6 . . 7 2 . . . . 24 2 . . . . . . 
Polygonatum verticillatum d6 . . . . . . . 21 2 . . . . . . 
Cardaminopsis *tatrica [C] C7 . . . . . . . . 83 3 50 3 70 3 14 2 17 3 15 2 
st Tephroseris capitata t7 . 17 4 . . . . . . 67 3 83 2 73 2 29 2 17 1 26 2 
Homogyne alpina d7 . . . 10 2 . . . 34 2 72 4 92 2 80 3 43 2 17 3 38 2 
MU Geranium sylvaticum d7 . 8 2 . . . . . 34 2 89 3 67 3 80 3 11 2 . 9 2 
Cv Festuca picturata d7 . . . . . . . . 89 5 58 4 77 4 4 2 . 3 2 
Cv Anemone narcissiflora d7 . . 29 2 . . . . 17 3 89 3 50 3 73 3 32 2 50 2 35 2 
Oreogeum montanum d7 . . . . . . . . 78 3 42 3 63 3 25 2 17 3 24 2 
Viola *sudetica d7 . 8 2 . . . . . . 28 4 8 6 20 5 . . . 
Cetraria islandica (E0) d8 . . 7 2 10 5 . 10 2 2 2 3 3 6 2 42 5 20 4 86 3 83 3 85 3 
Sc,ES Selaginella selaginoides d8,7b . . . . . . . 24 2 11 2 58 2 30 2 71 2 83 3 74 2 
fv Pedicularis oederi d8 . . . . . . . . 17 2 42 3 27 3 71 2 83 3 74 2 
ac Salix reticulata d8 . . 7 2 . . . . . 22 2 . 13 2 61 3 83 3 65 3 
fv Silene *longiscapa d8 . . . . . . . . 6 2 8 3 7 3 64 3 67 3 65 3 
cf Carex firma d8 13 3 . 21 3 . . . . 7 3 . . . 54 2 50 3 53 2 
oe Carex atrata d8 . . . . . . . . 6 3 8 2 7 3 43 2 67 2 47 2 
cf Salix alpina d8 . . 21 2 . . . . 17 4 11 3 17 2 13 3 39 3 67 4 44 3 
fv Cerastium eriophorum d8 . . . . . . . . 6 2 8 3 7 3 50 2 17 2 44 2 
fv Minuartia sedoides d8 . . . . . . . . . . . 36 2 33 3 35 2 

138 


J. Kliment et al.: Syntaxonomy and nomenclature of tHe communitieS of tHe allianceS Astero Alpini-seslerion CAlCAriAe … in Slovakia 

Number of column 1 2 3 45a5b 5 67a7b 78a8b 8 
Number of relevés 15 12 14 10 41 10 51 29 18 12 30 28 6 34 
Average number of taxa 37 39 30 24 36 27 34 38 51 43 48 49 37 47 

fv Minuartia pauciflora [C] d8...........213333 24 3 
CC Campanula alpina d8 . . . . . . . . . . .182502 24 2 
Differential taxa of the subassociations 
ch Erysimum witmannii [C] d5a.... 68 3 .553. . . . . . . 
ch Pulsatilla slavica [CW] d5a . . 365 902 66 2 .532. . . . . . . 
ES Thesium alpinum d5a 132 172 72 602 61 2 102512282282173232 . . . 
Sc,ES Thymus pulcherrimus [C] d5a,7b,8a 933 504 642 803 56 3 204 493 213 62 83 3 37 3 75 3 . 623 
ch Allium ochroleucum d5a . . .402 51 2 .412. . . . . . . 
Fv Bupleurum falcatum d5a . . . . 51 2 .412. . . . . . . 
AT Asplenium trichomanes d5a . . . . 44 2 102372. . . . . . . 
ch Acinos alpinus d5a . . . . 41 3 202372. . . . . . . 
Encalypta vulgaris (E0) d5a 473 173 . . 63 3 103533. . . . . . . 
Encalypta streptocarpa (E0) d5a272 . . . 54 2 202472 . . . . 45173 64 
Rhytidium rugosum (E0) d5a 202 583 293 102 44 2 10637374 . . .252 .212 
Fissidens dubius (E0) d5a 7282 . . 44 2 .352. . . . . . . 
Orthotrichum anomalum (E0) d5a.... 41 2 203372. . . . . . . 
Anomodon viticulosus (E0) d5a.... 29 2 .242. . . . . . . 
pc Campanula xylocarpa [CW] d5b..... 60 3 123. . . . . . . 
Fv Erysimum odoratum d5b . . . . . 60 2 122. . . . . . . 
Fv Pulsatilla subslavica [CW] d5b..... 40 2 82. . . . . . . 
ch Daphne arbuscula [CW] d5b..... 20 6 46. . . . . . . 
Cladonia pyxidata (E0) d5b .82. . . 70 3 143 . 63 . 33322333322 
Peltigera rufescens (E0) d5b..... 40 2 82. . . .73.63 
ac,oe Myosotis alpestris d7a,8a .82. . . . .72 83 3 332 633 50 2 . 412 
Cv Campanula elliptica d7a .503. . . . .452 78 2 .47242 . 32 
Cv Luzula *rubella d7a,8a . 173 72203 . . . 243 72 3 172 503 29 2 . 242 
Thymus alpestris d7a .17375 . . . .522 72 3 8247311 3 . 93 
Cv Crepis mollis d7a .82. . . . .142 67 3 172473 42 . 32 
Phyteuma spicatum d7a . . . . . . . . 61 2 .372. . . 
Cv Trommsdorfia uniflora d7a . . . . . . .31 50 4 .30442 . 32 
Cv Crepis conyzifolia d7a . . . . . . . . 39 4 .234. . . 
Ranunculus thora d7a . . . . . . . . 33 3 . 203 11 3 . 93 
Ranunculus pseudomontanus [Cs] d7a,8a.......213 28 3 . 173 50 2 . 412 
Cv Cyanus mollis [Cs] d7a . . . . .2024232 22 3 .133. . . 
Cortusa matthioli d7a . . . . . . . . 22 3 .13373 . 63 
Sc,ES Biscutella laevigata d7b . 172 . . 102 . 8245311 2 67 2 332 362 334 352 
Sc,ES Hieracium villosum d7b 472252212 . 23 . 23342 . 67 2 272 11 2 172 122 
Sc,ES Anthyllis *alpestris d7b 202 422 362 303 202 203 202 594 172 50 3 302 143 . 123 
pt Cerastium *glandulosum [CW] d8a........334502403 64 2 . 532 
Sc,ES Pedicularis verticillata d8a 532333212 . . . . 693672422572 61 2 173 532 
LV Vaccinium vitis-idaea d8a . . 292404 . . . 453282422332 61 3 . 503 
MU Bistorta major d8a . 83 72 . . . . 104443582502 39 2 . 322 
Botrychium lunaria d8a . 332 . . 22 . 22142612252472 39 2 . 322 
oe Erigeron hungaricus [C] d8a72.72........ 32 3 . 263 
CC Festuca supina d8a . . . . . . . .172.102 25 2 . 212 

Astero alpini-Seslerienion calcariae 

Bupleurum ranunculoides C 732 582 213 . . . . . . . .11 2.92 
CK,ES Festuca *versicolor [Cs] d 874 1007 1007 204 . . . 724 333 503 403 1007 1008 1007 
st Astragalus australis d 473 423 76 . . . . 36 63 82 73 42503123 
oe Androsace chamaejasme d,d8 272 172 72 . . . . . 172332232753332 68 3 
CK,Cv Campanula tatrae [CW] d 602 583 . . . . . . 612832702713673713 

Pulsatillo slavicae-Caricenion humilis 

Rhodax rupifragus C... 30254 . 44 ......
.
Coronilla vaginalis C... 502. . . ......
.
Globularia cordifolia C... 404. . . ......
.
Koeleria tristis C... 102. . . ......
.


139 


Hacquetia 4/2 • 2005 

Number of column 1 2 3 45a5b 5 67a7b 78a8b 8 
Number of relevés 15 12 14 10 41 10 51 29 18 12 30 28 6 34 
Average number of taxa 37 39 30 24 36 27 34 38 51 43 48 49 37 47 

Fv Carex humilis d... 403 343 202 313 ......
.
FB Tithymalus cyparissias d .423 . 202 662 302 592 ......
.
Pc Hieracium bupleuroides d,d5a . . . 202 632 202 552 ......
.
FB Anthericum ramosum d... 105 292 202 272 ......
.
Fv Teucrium montanum d... 102 273 202 253 ......
.
Polygonatum odoratum d,d5a . . . 203 412 . 332 ......
.
Asperula tinctoria d... 202 122 . 102 ......
.
Fv Seseli osseum d... . 1003 703 943 ......
.
FB Cyanus triumfettii d,d5a . . . . 632 202 552 ......
.
Vincetoxicum hirundinaria d,d5a . . . . 592 202 512 ......
.
Fv Asperula cynanchica d... . 492 402 472 ......
.


Astero alpini-Seslerion calcariae 

Sesleria albicans C 1005 583 503 1004 834 906 844 ....... 
Festuca tatrae [CWs] t 672 755 . 602 52 202 82 .63.33. . . 
Aster alpinus C 933 332 364 303 124 . 104 . . . .42.32 
Dianthus *praecox [CW] C 673 834 573 . 373 202 332 . . . .11 1. 91 
Minuartia langii C 72 173 363 302 613 302 553 72. . . . . . 
Jovibarba globifera d 672 172 712 502 932 702 882 172. 8232. . . 
Pc Asplenium ruta-muraria d 532 . 142 . 982 902 962 . . . . .50292 
pc Kernera saxatilis d 272 172 . 102 242 . 202 ....... 
sf Leontodon incanus d 133 . . 302 292 202 272 ....... 

Seslerion tatrae 

Sesleria tatrae [CWs] C,d3 . 17 2 71 3 .... 1006 1008 1007 1007 1005 834 975 
Trifolium *kotulae [C] C,d2 72 50 2 ..... 72392332372 72 . 62 
Gentiana verna C,d8a .17272 . . . . 172 333 672 472 612 . 502 
Astragalus alpinus C 75172143 . . . . .284 .17474 . 64 
Euphrasia tatrae [C] C....... . 283 332 302 252 172 242 
Potentilla aurea d .82. . . . . 413 944 1003 973 612 333 562 
Parnassia palustris d .173292 . . . . 412 612 583 602 862 833 852 
Cv Achillea *alpestris d,d7a .172212 . . . . 593 943 252 673 252 . 212 
Cv Linum extraaxillare [Cs] d,d2,7a,8a 13 2 75 4 ..... 172 673 254 503 293 . 243 
MU Primula elatior d,d8a . . 71102. . . 623 1003 673 873 502 . 412 
Ligusticum mutellina d,d8a . . . . . . . 483 785 675 735 503 . 413 
Anthoxanthum alpinum d,d8a . .73. . . . 453 943 923 933 392 . 322 
Cv Phleum hirsutum d,d8a .172. . . . . 552 783 673 733 362 . 292 
CC Avenula versicolor d....... . 282 504 373 293 173 263 
Seslerietalia coerulae, Elyno-Seslerietea 

Scabiosa lucida 72 923 713 702 292 303 292 973 503 753 603 712 332 652 
Ranunculus breyninus 1003 833 504 202 23 103 43 312 503 1002 703 543 502 533 
Helianthemum grandiflorum 603 753 863 203 293 202 273 453 784 584 704 643 333 593 
Phyteuma orbiculare 202 673 1002 702 322 302 312 903 893 1003 933 932 503 852 
Galium anisophyllon 802 923 792 502 72 . 62 662 673 502 603 933 333 823 
Polygala *brachyptera 402 673 142 202 372 502 392 312 11 2 172 132 72 . 62 
Euphrasia salisburgensis 602 582 712 302 292 203 272 342 62 82 72 292 . 242 
Carex *tatrorum [CW] 533 675 362 403 . . . 1007 945 1005 975 825 674 795 
Bellidiastrum michelii 202 252 292 202 . . . 723 392 672 502 573 333 533 
Veronica fruticans 332332432. . . . . . 823272. 62 
cf Arenaria tenella [CW] 20282. . . . . . . 813111 2. 92 
Colymbada alpestris 27283. .72.62. . . . . . . 
Gentiana clusii 132 .862602 . . .173 . . .142 .122 
Carduus glaucinus . 582 . 302 542 302 492 414 172 422 272 42 . 32 
Gentianella fatrae [CW] ..432402...172...... 
cb Knautia kitaibelii [CWs] . . . 302122 . 102453 6317410311 2 . 92 
cf Pinguicula alpina . . . . . . .32. . .11 2673212 

Asplenietea trichomanis 

Pc Saxifraga paniculata 803 422 1004 . 883 902 883 212 . 82 32 573 673 593 

140 


J. Kliment et al.: Syntaxonomy and nomenclature of tHe communitieS of tHe allianceS Astero Alpini-seslerion CAlCAriAe … in Slovakia 
Number of column 1 2 3 4 5a 5b 5 6 7a 7b 7 8a 8b 8 
Number of relevés 15 12 14 10 41 10 51 29 18 12 30 28 6 34 
Average number of taxa 37 39 30 24 36 27 34 38 51 43 48 49 37 47 
Pc Primula *hungarica [CWs] 
pc Crepis jacquinii 
pc Trisetum alpestre 
pc Campanula cochleariifolia 
pc Draba aizoides 
cy Valeriana tripteris 
cy Asplenium viride 
cy Cystopteris fragilis 
Festuco-Brometea 
Genista pilosa 
Viola hirta 
sf Chamaecytisus hirsutus 
Colymbada scabiosa 
Fv Campanula *divergentiformis 
Fv Potentilla arenaria 
Fv Allium flavum 
Mulgedio-Aconitetea 
Cv Astrantia major 
Cv Solidago *minuta 
Aconitum firmum [C] 
Cv Pyrethrum clusii 
Cv Campanula serrata [C] 
Cv Cirsium erisithales 
Senecio subalpinus 
Gentiana asclepiadea 
Acetosa arifolia 
tf Taraxacum alpinum 
tf Cerastium fontanum 
Poa chaixii 
Caricetea curvulae 
Agrostis pyrenaica 
Pulsatilla scherfelii 
Luzula sudetica 
Hieracium alpinum 
Oreochloa disticha 
Juncus trifidus 
Salicetea herbaceae 
ac Leontodon pseudotaraxaci [C] 
Luzula *obscura [C] 
ac Salix retusa 
ac,pt Saxifraga wahlenbergii [CW] 
Carici rupestris-Kobresietea 
Bistorta vivipara 
Dryas octopetala 
oe Saussurea alpina 
Ligusticum mutellinoides 
Dianthus glacialis 
fv Agrostis alpina 
oe Sedum atratum 
oe Carex capillaris 
oe Oxytropis carpatica [C] 
CK Lloydia serotina 
fv Erigeron uniflorus 
CK Luzula *mutabilis 
93 4 
13 3 
100 4 
7 2 
47 2 
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. 
. 
. 
. 
. 
. 
. 
. 
. 
17 3 
17 2 
33 3 
. 
. 
50 2 
50 2 
17 3 
. 
. 
. 
. 
17 2 
17 3 
. 
17 1 
33 1 
. 
6 2 
15 3 
18 2 
3 2 
12 2 
. 
9 2 
. 
. 
. 
. 
. 
. 
. 
. 
6 2 
6 2 
9 2 
. 
. 
. 
. 
. 
. 
6 2 
. 
. 
12 3 
15 2 
12 2 
12 2 
15 2 
12 2 
44 2 
3 3 
21 2 
12 2 
76 2 
41 3 
15 2 
24 3 
6 2 
9 2 
12 2 
6 3 
6 3 
18 2 
12 2 
15 2 

141 


Hacquetia 4/2 • 2005 

Number of column 1 2 3 4 5a 5b 5 6 7a 7b 7 8a 8b 8 
Number of relevés 15 12 14 10 41 10 51 29 18 12 30 28 6 34 
Average number of taxa 37 39 30 24 36 27 34 38 51 43 48 49 37 47 
Other taxa 
Carlina acaulis 7 2 50 2 . 30 2 5 2 . 4 2 66 3 39 3 42 3 40 3 4 1 17 1 6 1 
Lotus corniculatus 7 2 83 3 . 10 2 7 2 . 6 2 59 3 28 3 58 3 40 3 4 2 . 3 2 
Gymnadenia conopsea 7 1 33 2 . 20 2 2 2 . 2 2 10 2 33 2 42 2 37 2 7 2 . 6 2 
Linum catharticum 7 1 42 3 . 10 2 10 2 10 2 10 2 10 2 . 8 2 3 2 . 17 2 3 2 
Rhodax alpestris 33 3 . 29 5 10 2 . . . . 6 5 . 3 5 25 2 . 21 2 
Leucanthemum vulgare agg. . 33 2 50 2 . 12 2 . 10 2 97 3 50 2 33 2 43 2 7 4 . 6 4 
Cardaminopsis arenosa agg. . 33 2 29 2 . 24 2 40 3 27 2 38 2 . . . 14 2 17 1 15 2 
Pimpinella major . 58 3 . . 2 2 . 2 2 10 2 22 3 25 3 23 3 . . . 
Laserpitium latifolium . 17 4 . 10 2 34 2 20 4 31 2 . 6 3 . 3 3 . . . 
Allium *montanum 27 2 25 2 43 2 10 3 10 3 . 8 3 24 3 . . . . . . 
Hieracium bifidum 7 2 8 2 7 5 50 2 15 2 20 2 16 2 34 2 . . . . . . 
Thalictrum minus 7 2 8 2 . . 10 2 20 2 12 2 3 2 . . . . . . 
Libanotis pyrenaica 67 2 67 3 21 3 . 2 2 . 2 2 17 2 . . . . . . 
Digitalis grandiflora . 33 2 . . 2 2 20 2 6 2 . . . . . . . 
Fragaria vesca . 8 2 7 2 . 10 2 30 2 14 2 . . . . . . . 
Campanula rapunculoides . 8 2 . . 20 2 10 2 18 2 . . . . . . . 
Carex digitata . 17 3 . 30 2 37 2 . 29 2 . . . . . . . 
Clematis alpina . 8 2 . 10 2 2 2 20 2 6 2 . . . . . 17 2 3 2 
Achillea stricta . . . 10 2 15 2 30 2 18 2 . . . . . . . 
Cotoneaster tomentosus . . . 20 2 12 2 10 2 12 2 . . . . . . . 
Sorbus aria . . . 20 2 10 2 10 1 10 2 . . . . . . . 
Securigera varia . . . . 44 2 30 3 41 2 . . . . . . . 
Sedum album . . . . 39 3 20 2 35 3 . . . . . . . 
Dianthus carthusianorum . . . . 34 2 10 3 29 2 . . . . . . . 
Calamagrostis varia . . . . 24 2 . 20 2 7 2 6 2 17 3 10 2 . . . 
Galium mollugo agg. . . . . 2 2 10 2 4 2 3 2 6 2 . 3 2 . . . 
Hypericum maculatum . . . . . 10 1 2 1 21 3 33 2 8 2 23 2 . . . 
Heracleum sphondylium . 8 3 . . . 10 2 2 2 38 2 28 2 8 2 20 2 . . . 
Poa alpina . 33 2 64 2 . . . . 52 3 44 2 50 2 47 2 64 2 50 2 62 2 
Leontodon hispidus . 17 2 . . . . . 34 4 72 4 50 3 63 3 14 2 17 2 15 2 
Alchemilla sp. div. . 8 2 . . . . . 48 3 83 3 50 2 70 2 64 2 . 53 2 
Arabis alpina . 17 2 . . . . . . 6 2 . 3 2 14 2 17 2 15 2 
Trifolium badium . 8 3 . . . . . . 6 2 8 3 7 3 4 2 . 3 2 
Pinus mugo . . 14 2 10 2 . . . . 6 2 8 2 7 2 4 2 17 2 6 2 
Viola biflora . . 36 2 . . . . 28 3 39 4 17 2 30 3 39 2 17 3 35 2 
Swertia perennis . . 7 2 . . . . 10 4 44 3 42 2 43 3 43 2 67 3 47 3 
Vaccinium myrtillus . . . 20 4 . . . 38 2 44 3 42 2 43 3 21 3 33 4 24 3 
Soldanella carpatica [CW] . . . . . . . 62 2 94 3 83 2 90 3 71 2 67 2 71 2 
Luzula sylvatica . . . . . . . 24 2 11 2 . 7 2 11 2 . 9 2 
Dianthus *alpestris . . . . . . . . 17 2 33 2 23 2 7 2 . 6 2 
oe Astragalus frigidus . . . . . . . . 17 3 . 10 3 11 4 . 9 4 
oe Astragalus norvegicus . . . . . . . . 17 3 . 10 3 4 2 . 3 2 
Trifolium orbelicum . . . . . . . . 17 2 . 10 2 4 2 . 3 2 
Veronica aphylla . . 7 2 . . . . . . 8 2 3 2 29 2 . 24 2 
Traunsteinera globosa . . . . . . . 28 2 6 3 17 2 10 2 . . . 
Coeloglossum viride . . . . . . . 3 2 28 2 8 2 20 2 . . . 
Saxifraga aizoides . . . . . . . 7 2 . . . 29 2 33 3 29 2 
Viola alpina . . 7 2 . . . . . . . . 11 2 17 3 12 2 
LV Vaccinium gaultherioides . . . . . . . . . . . . 33 4 6 4 
Bryophytes and lichens 
Tortella tortuosa 93 4 100 3 43 3 50 3 71 3 100 4 76 4 10 3 28 4 33 3 30 3 61 3 83 5 65 3 
Rhytidiadelphus triquetrus . 8 2 7 2 10 2 2 2 . 2 2 . 17 4 . 10 4 21 2 . 18 2 
Ctenidium molluscum 7 2 8 2 . . . 10 2 2 2 . . 8 3 3 3 7 2 . 6 2 
Thuidium abietinum 7 2 17 2 . . 17 2 10 2 16 2 . . . . 4 2 . 3 2 
Myurella julacea 40 2 . . . 2 2 . 2 2 . . . . 7 3 33 4 12 3 

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J. Kliment et al.: Syntaxonomy and nomenclature of tHe communitieS of tHe allianceS Astero Alpini-seslerion CAlCAriAe … in Slovakia 
Number of column 1 2 3 4 5a 5b 5 6 7a 7b 7 8a 8b 8 
Number of relevés 15 12 14 10 41 10 51 29 18 12 30 28 6 34 
Average number of taxa 37 39 30 24 36 27 34 38 51 43 48 49 37 47 

Schistidium apocarpum s. l. 47 4 . 7 3 10 2 39 2 10 2 33 2 3 2 . . . 4 2 . 3 2 
Ditrichum flexicaule 40 2 . 7 3 40 2 54 3 50 3 53 3 . . 8 2 3 2 . . . 
Homalothecium philippeanum 47 2 25 2 . 20 2 68 3 40 3 63 3 . . . . . . . 
Cladonia sp. 40 3 8 2 14 3 10 3 . 10 2 2 2 . . . . . . . 
Bryum argenteum 27 2 33 1 . . 44 2 40 2 43 2 . . . . . . . 
Tortula ruralis 7 3 . 14 3 . 27 2 10 2 24 2 . . . . . . . 
Grimmia sp. 53 3 . . . 2 2 10 2 4 2 . . . . . . . 
Distichium capillaceum 47 2 . . . 15 2 . 12 2 . . . . . . . 
Hypnum cupressiforme . 8 2 14 3 . 20 2 10 3 18 2 . . . . . . . 
Pleurozium schreberi . 17 4 . . . . . 7 5 . 8 2 3 2 11 2 . 9 2 
Campylium protensum . 8 2 . . . . . . 6 5 8 2 7 4 4 2 17 2 6 2 
Sanionia uncinata . 8 2 7 2 . . . . 10 3 6 2 33 3 17 2 18 2 17 2 18 2 
Hylocomium splendens . . 7 3 . . . . 14 4 . . . 25 4 33 4 26 4 
Cladonia rangiferina . . . . . 10 6 2 6 . . 17 3 7 3 18 2 . 15 2 
Mnium marginatum . . . . . . . . 6 5 . 3 5 4 2 33 3 9 2 
Cladonia furcata . . . . . . . . . 8 2 3 2 7 3 33 3 12 3 
Lophozia bantriensis . . . . . . . . . 8 2 3 2 . 50 2 9 2 
Plagiochila asplenioides . . . . . . . . . 25 2 10 2 . 33 2 6 2 
Cetraria cucullata . . . . . . . . . . . 21 2 17 3 21 2 
Cetraria nivalis . . . . . . . . . . . 18 2 . 15 2 
Thamnolia vermicularis 7 2 . . . . . . . . . . 18 2 . 15 2 
Meesia uliginosa . . . . . . . . . . . . 67 2 12 2 

Number of accesoric taxa 
E1 3 25 2 13 41 21 58 37 24 6 28 31 6 34 
E03315 2 6201331 723 526291440 

Explanations: [C] = Carpathian endemit, [Cs] = Carpathian subendemit, [CW] = endemit of the Western Carpathians and 
their subregions, [CWs] = West Carpathian subendemit 

Taxa with rare occurrence in the described subassociations:
5a – Minuartio langii-Festucetum pallentis campanuletosum carpaticae:
E1: Acer pseudoplatanus 22,FB Arabis hirsuta 102,FB Arenaria serpyllifolia 22, Carex ornithopoda 202,Be Carlina biebersteinii 22,


Clinopodium vulgare 22, Convolvulus arvensis 22, Cotoneaster integerrimus 22, Epipactis atrorubens 102, Galium austriacum 22, Fv Galium glaucum 52, Geranium robertianum 22, FB Geranium sanguineum 52, Gymnadenia odoratissima 22, Fv 
Hippocrepis comosa 52,AT Hylotelephium maximum 172, Hypericum perforatum 52, Fv Inula ensifolia 52, Larix decidua 52, 
Lembotropis nigricans 72, Melampyrum sylvaticum 52,Fv Melica ciliata 22, M.nutans 52, Origanum vulgare 72, Picea abies 22, 
Pilosella officinarum 22, FB Pimpinella saxifraga 22, Pinus sylvestris 172, Poa molineri 103, Fv Potentilla heptaphylla 22, P. 
thuringiaca 22, Rhinanthus serotinus 172, Rosa pendulina 22, Rubus saxatilis 22, Sedum sexangulare 22, Silene nemoralis 52, S. 
pseudotites 152, Spiraea media 72, FB Stachys recta 153, Taraxacum sp. 22, Vicia cracca 22. 

E0: 
Barbilophozia barbata 22, Bartramia ithyphylla 22, Bryum capillare 202, Campylium chrysophyllum 22, Dicranum scoparium 52, Distichium inclinatum 22, Ditrichum heteromallum 22, Encalypta rhaptocarpa 52, Grimmia pulvinata 222, Homalothecium sericeum 103, Hypnum sp. 22, Leskea polycarpa 292, Porella platyphylla 52, Pseudoscleropodium purum 22, Solorina saccata 222, Tortella inclinata 243, Tortula intermedia 102, Tortula sp. 122, Trichostomum crispulum 52, Weissia condensata 72. 

5b 
– Minuartio langii-Festucetum campanuletosum xylocarpae: 

E1: 
Acer pseudoplatanus 101, Aconitum variegatum 101, Briza media 102, Calamagrostis arundinacea 102, Campanula persicifolia 102, C. trachelium 101, Chamaecytisus ratisbonensis 102, Galium schultesii 203, Geranium robertianum 102, Mycelis 
muralis 101, Origanum vulgare 102,FB Pimpinella saxifraga 102, Poa nemoralis 102,AT Polypodium vulgare 102, Rhinanthus 
minor 102, Rubus idaeus 103, Sempervivum matricum 102, Silene sillingerii [CW] 102, Silene sp. 101, FB Teucrium chamaedrys 102, Tithymalus epithymoides 102. 

E0: 
Brachythecium sp. 102, Bryoerythrophyllum recurvirostrum 103, Cladonia fimbriata 202, C. squamosa 102, Collema polycarpon 102, Ditrichum sp.104, Endocarpon sp.102, Leptogium lichenoides 103, Neckera crispa 102, Plagiopus oederi 102, Pseudoscleropodium purum 102, Toninia caerulea 102, Tortula intermedia 102. 

7a – Seslerietum tatrae luzuletosum rubellae:
E1: Agrostis capillaris 62, Allium schoenoprasum subsp. alpinum 62, Angelica sylvestris 112, Avenella flexuosa 63, Cv Bupleurum 


143 


Hacquetia 4/2 • 2005 

longifolium 112, Callianthemum coriandrifolium 62, Cv Gentiana punctata 112, ca Hieracium prenanthoides 112, Hieracium 
sp. 112, Hylotelephium argutum [C] 62, Leucanthemum rotundifolium [Cs] 62, Lilium martagon 62, Orchis mascula subsp. 
signifera 112, Plantago atrata subsp. carpatica [C] 112, Pyrola minor 112, P. rotundifolia 62, Rhinanthus pulcher 114, Senecio 
abrotanifolius subsp. carpaticus 62, Taraxacum sp. 283, Tephroseris crispa 63, Tofieldia calyculata 62, Trisetum flavescens 
subsp. tatricum [C] 63, Trollius altissimus 112, MU Veratrum album subsp. lobelianum 172. 

E0: Brachythecium glareosum 112, B. velutinum 224, Bryum algovicum 65, B. capillare 62, Campylium calcareum 65, Desmatodon 
latifolius 62, Eurhynchium praelongum 65, E. schleicheri 65, Fissidens taxifolius 61, Homalothecium lutescens 65, Lophocolea 
bidentata 62, Mnium spinosum 114, M. thomsonii 63, Peltigera aphthosa 62, Plagiomnium cuspidatum 114, Pohlia wahlenbergii 61, Porella platyphylla 62, Pseudoleskea incurvata 62, Ptychodium plicatum 62, Rhizomnium punctatum 114, Rhodobryum 
roseum 62, Rhytidiadelphus squarrosus 61, Tortula intermedia 65. 

7b 
– Seslerietum tatrae biscutelletosum laevigatae: 

E1: Cv Bupleurum longifolium 82, Hieracium sp. 82, Listera ovata 82, Taraxacum fontanum 252, T. nigricans [CW] 172, Veronica 
alpina 82. 

E0: 
Desmatodon latifolius 173, Dicranum scoparium 82, Heterocladium dimorphum 173, Homalothecium sericeum 83, Rhizomnium punctatum 172. 

8a – Seslerio tatrae-Festucetum versicoloris ranunculetosum pseudomontani: 

E1: 
Adoxa moschatellina 42, oe,fv Antennaria carpatica subsp. carpatica [C] 112, Avenella flexuosa 42, Calluna vulgaris 71, oe, 
fv Carex fuliginosa 143, cf Chamorchis alpina 72, Coeloglossum viride 142, Cystopteris regia 42, pt Delphinium oxysepalum 
[CW] 42, Doronicum stiriacum 42, Erigeron alpinus 42,CK Gentiana nivalis 42, Gentianopsis ciliata 41, Geum rivale 42, Gymnadenia odoratissima 112, Hieracium caesium 72, H.pilosum 43, Luzula campestris 42, L.multiflora 42,oe Oxytropis halleri 74, 
Phleum rhaeticum 42, Picea abies 42, CC Primula minima 73, Pritzelago alpina 42, Pyrola carpatica [C] 112, Ranunculus 
acris 42, Saxifraga adscendens 42, ac S. androsacea 42, S. hieraciifolia 42, fv S. oppositifolia 42, Trollius altissimus 42. 

E0: 
Alectoria ochroleuca 112, Biatorella fossarum 42, Brachythecium glareosum 72, Bryum sp. 43, Campylium chrysophyllum 42, 
Campylopus subulatus 43, CC Cladonia arbuscula 112, C. bellidifolia 43, C. deformis 43, C. gracilis 142, C. symphycarpa 73, 
Heterocladium dimorphum 42, Hypnum revolutum 42, Hypogymnia vittata 42, Lecanora epibryon 42, Mnium spinosum 43, 
Ochrolechia upsaliensis 72, Physconia muscigena 42, Plagiobryum demissum 43, Pogonatum urnigerum 42, Pohlia cruda 42, 
Polytrichum alpinum 42, Pseudoleskeella catenulata 42, Racomitrium canescens 72, Rhytidiadelphus squarrosus 72, Solorina 
bispora 42, Thuidium erectum 42, Th. tamariscinum 42, Vulpicida tubulosus 73. 

8b 
– Seslerio tatrae-Festucetum versicoloris inops: 

E1: 
Hieracium fritzei 172, Huperzia selago 173, Pilosella officinarum 171, ac Saxifraga androsacea 171, S. hieraciifolia 171, fv S. 
oppositifolia 172. 

E0: 
Anthelia juratzkana 172, Blepharostoma trichophyllum 172, Brachythecium reflexum 172, Bryum sp.332, Cladonia symphycarpa 172, Lophozia longiflora 171, Mannia pilosa 172, Mnium stellare 172, Orthothecium chryseon 172, Peltigera sp. 173, Pohlia 
cruda 332, Psoroma hypnorum 172, Solorina saccata 172, Timmia norvegica 172. 

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J. Kliment et al.: Syntaxonomy and nomenclature of tHe communitieS of tHe allianceS Astero Alpini-seslerion CAlCAriAe … in Slovakia 
Table 2: Gentiano clusii-Festucetum versicoloris Bělohlávková ass. nov.; variant with Rhodiola rosea (r. 1–9); vari-
ant with Minuartia langii (r. 10–14)
.
Tabela 2: Gentiano clusii-Festucetum versicoloris Bělohlávková ass. nov.; varianta z vrsto Rhodiola rosea (popisi 
1–9), varianta z vrsto Minuartia langii (popisi 10–14)
.


Number of relevé 0 0 0 0 0 0 0 0 0 1 1 1 1 1 
1 2 3 4 5 6 7 8 9 0 1 2 3 4 
Number of taxa 2 3 2 2 2 2 2 2 2 2 2 3 2 2 C 
8 7 4 4 7 3 4 4 6 5 7 1 9 3 % 

Diagnostic taxa of association 
ES Festuca *versicolor D 4 3 3 3 3 4 3 3 3 3 3 3 3 a 100 
st Sesleria tatrae d a ++ . . . 1 1 1 . a a 1 1 71 
Bartsia alpina d + 1 1 + a 1 + . 1 . 1 . 1 . 71 
Cv Vicia oreophila d ++ . . . 1 ++ . . + + ++ 64 
cf Androsace lactea d ++ . . 1 +++ . . . . + . 50 
cf Ranunculus alpestris d . + . . 1 . . . . + . + ++ 43 
Differential taxa of variants 
tf Rhodiola rosea a 1 . + + . ++ + . . . . . 50 
Potentilla crantzii . + 1 1 . a a + + . . . . . 50 
Hedysarum hedysaroides . + 1 . 1 1 1 1 + . . . . . 50 
Allium *montanum 1 . . + + a 1 . a . . . . . 43 
Tortella tortuosa (E0) 1 1 1 1 . . 1 1 . . . . . . 43 
Viola biflora +++ . 1 + . . . . . . . . 36 
Parnassia palustris + . + . . . . + + . . . . . 29 
Saxifraga moschata . . + . + . + . + . . . . . 29 
Cv Anemone narcissiflora . . . . +++ . 1 . . . . . 29 
as Minuartia langii . . . . . . . . . 1 + 1 + 1 36 
as Pulsatilla slavica . . . . . . . . . a a 3 a + 36 
ES Veronica fruticans . . . + . . . . . ++ + ++ 43 
Carex ornithopoda . . . . . . . . . 1 1 + + 1 36 
Rhodax alpestris . . . . . . . . . 3 3 + 1 . 29 
ES Hieracium villosum . . . . . . . . . ++ + . . 21 
Astero-Seslerion calcariae 
Jovibarba globifera d ++ . 1 . + . + . ++ + ++ 71 
Dianthus *hungaricus t . . . . + 1 1 1 + 1 + . . a 57 
Sesleria albicans C . . ++ + 1 1 1 1 . . . . . 50 
Aster *glabratus C . . + . . a a a 1 . . . . . 36 
Bupleurum ranunculoides C . . . . . 1 + . 1 . . . . . 21 
Seslerietalia coerulae, Elyno-Seslerietea 
Phyteuma orbiculare ++ 1 r ++ 1 + 1 ++ 1 + 1 100 
Gentiana clusii . +++ +++ 1 . 1 1 + 1 + 86 
Helianthemum grandiflorum + 1 . + . + a + 1 1 + + 1 a 86 
Galium anisophyllon ++ 1 + ++ . + . ++ + . + 79 
Euphrasia salisburgensis . + . . . +++ + 1 + 1 1 1 71 
Scabiosa lucida + a 1 . . 1 + . 1 . + + + 1 71 
Thymus *sudeticus . ++ . + . . + . ++ 1 ++ 64 
Ranunculus breyninus 1 . . 1 . . . . . 1 1 a a 1 50 
Gentianella fatrae . + . . . . . . . ++ + ++ 43 
Carex *tatrorum . . + . . . + . . + . + + . 36 
Anthyllis *alpestris . + . . . + . 1 . ++ . . . 36 

145 


Hacquetia 4/2 • 2005 

Number of relevé 000000000 11111 
123456789 01234 

Bellidiastrum michelii .+.....+1 ...+. 29 
Pedicularis verticillata .+.+..... ...1. 21 
cf Salix alpina .+1+..... ..... 21 
st Astragalus alpinus .1.1..... ..... 14 
Hieracium villosum ......... +++.. 21 
cf Carex firma ......... .++a. 21 
Polygala *brachyptera ......... +...+ 14 

Asplenietea trichomanis 

Pc Saxifraga paniculata 1aa1a1+a1 ++11a 100 
Pc Trisetum alpestre ++++.++. . 1a1+1 79 
Pc Primula *hungarica . . . . 1aaa+ aa+aa 71 
Pc Campanula cochleariifolia ++.+. .++. . .1+1 57 
pc Draba aizoides ....++.+. .+11. 43 
Pc Crepis jacquinii .+1...... ..+.. 21 

Asplenium ruta-muraria ......... +.+.. 14
cy Asplenium viride ......... ..r+. 14


Other taxa 

Poa alpina +1+1++1+. . . . .+ 64 
Leucanthemum vulgare agg. ++......1 ++.++ 50 
Cardaminopsis arenosa agg. +..++.... ..+.. 29 
Rhytidium rugosum (E0) +1a.....1 ..... 29 
Libanotis pyrenaica a+....1.. ..... 21 
Cv Achillea *alpestris ......... .+1.+ 21 

Rarely occurring taxa: 

E1: 
Androsace chamaejasme + (5); Anthoxanthum alpinum 1 (9); Astragalus australis 2b (1); Bistorta major + (1); Campanula 
serrata + (9); Cystopteris fragilis r (4), + (11); Delphinium oxysepalum + (5); Dianthus nitidus + (14); Dryas octopetala 1 (2); 
Erigeron hungaricus + (1); Fragaria vesca + (10); Gentiana verna + (5); Gypsophila repens + (5); Hieracium bifidum 2a (12); 
Knautia arvensis + (1); Luzula luzuloides + (1); Pinus mugo r (10), + (13); Primula elatior r (1); Salix retusa + (2); Sedum 
alpestre + (4); Swertia perennis + (5); Thesium alpinum + (12); Thymus alpestris 2a (4); Veronica aphylla + (2); Viola alpina 

+ (12); V. canina + (8). 
E0: Dicranum scoparium + (3), 1 (5); Hylocomium splendens 1 (5); Hypnum cupressiforme + (1), 1 (2); Marchantia polymorpha 
+ (11); Rhytidiadelphus triquetrus + (1); Sanionia uncinata + (3); Schistidium apocarpum 1 (5); Tortula ruralis 1 (6, 9). 
– Cetraria islandica + (13); Cladonia sp. + (1), 1 (4). 
146 


J. Kliment et al.: Syntaxonomy and nomenclature of tHe communitieS of tHe allianceS Astero Alpini-seslerion CAlCAriAe … in Slovakia 
Table 3: Diantho nitidi-Seslerietum tatrae Bělohlávková ass. nov.; variant typical (r. 1-22); variant with Allium 
*montanum (r. 23-29) 
Tabela 3: Diantho nitidi-Seslerietum tatrae Bělohlávková ass. nov.; tipična varianta (popisi 1-22); varianta z vrsto 

Allium *montanum (popisi 23-29) 
Number of relevé 0 0 0 0 0 0 0 0 0 1 1 1 1 1 1 1 1 1 1 2 2 2 2 2 2 2 2 2 2 
1 2 3 4 5 0 6 7 8 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 
Number of taxa 3 2 3 3 3 2 3 3 4 4 3 4 4 3 4 3 4 4 4 2 3 3 3 4 3 4 4 3 5 C 
3 7 0 9 5 4 8 3 1 9 9 2 0 8 1 1 4 6 7 4 8 4 8 3 4 0 9 8 2 % 

Diagnostic taxa of association 
st Sesleria tatrae D a b a 3 3 b b 4 b 3 3 3 b b 3 3 3 3 b 3 3 3 a a b b b b b 100 
ES Dianthus nitidus d 1 1 1 . 1 1 + 1 1 1 + 1 a 1 1 . + + + + 1 . + 1 1 1 + 1 1 90 
Knautia arvensis d + + + a . . . . . . + 1 . + + 1 + 1 + a 1 a . 1 . + a . + 66 
Ranunculus nemorosus d + + + + . 1 . . . . . 1 1 . . . . + 1 . . . . + . . + . + 41 
Polygonatum verticillatum d . . . . . . . . + + . . . . . 1 . . . . . . + + . + . . . 21 
Differential taxa of variants 
MU Primula elatior + + r + + . . 1 . + . a 1 1 + 1 1 + 1 . + 1 . . . . . . + 62 
Potentilla crantzii a a . . . . . 1 + + + 1 + 1 1 1 1 a 1 + . . . . . + . . . 55 
Poa alpina a a + + 1 . + . + + . + + . a . + . . + 1 . . . . . . . 1 52 
Ligusticum mutellina + . + + 1 + + 1 1 + 1 . . 1 + . . b 1 . . . . . . . . . . 48 
Alchemilla sp. div. + + + 1 . . + 1 . + . a a . . . a 1 a . 1 + . . . . . . . 48 
Anthoxanthum alpinum . . . + 1 a a a . . + 1 1 . . . 1 + 1 + a . . . . . . . . 45 
Bistorta vivipara a . + . 1 . . + 1 + 1 + + + 1 . + . . . + . . . . . . . . 45 
Vaccinium vitis-idaea + + + 1 1 + + . + + + . . . . . . + . . + . . + . . . . . 45 
Potentilla aurea . . + 1 + . 1 a + + 1 . . + . . 1 + + . . . . . . . . . . 41 
Parnassia palustris . . + 1 + . + . + + + + + + . + . . . . . . + . . . . . . 41 
Vaccinium myrtillus . . . . a . a r + + + . . + + + . + + . . . . . . . . . . 38 
Homogyne alpina + . . . + . . 1 1 1 1 . . + + . . + + . . . . . . . . . . 34 
Viola canina . . . + . . + . . . . + + + + . . . . . + . . . . . . . . 24 
Cv Luzula luzuloides . . . 1 + . a . . . . + + 1 . . . . . . . a . . . . . . . 24 
Luzula sylvatica . . . . + . . . . r + + + . . + . . . . . + . . . . . . . 24 
ES Selaginella selaginoides . . + . . 1 + . + + 1 . + . . . . . . . . . . . . . . . . 24 
Ranunculus pseudomontanus . . . . 1 . . + + . . . . . . . . 1 . . + 1 . . . . . . . 21 
Hypericum maculatum . . . . . . + a . . . . + . . . 1 . 1 . + . . . . . . . . 21 
Cv Anemone narcissiflora . . . . . . + . . . . . . 1 1 1 . . . + . . . . . . . . . 17 
Allium *montanum . . . . . . . . . . . . . . . . . . . . . . + 1 a + + + + 24 
ES Carduus glaucinus . . . . . . . . . . . . . . 1 + + . . a . + + a a m a a a 41 
ES Ranunculus breyninus . . . . . . . . + . . . . . . . . . + . . . + 1 1 + + + 1 31 
ES Thesium alpinum . . . . . . . . . . . . . . . . . . + . . . + + + + + + + 28 
ES Hieracium villosum . . . . . . . . . . . + + . + . + . . . + . . + + + . + 1 34 
ES Polygala *brachyptera . . . . . . . . + . . + + . + . . . . . . . . + . + 1 + + 31 
Libanotis pyrenaica . . . . . . . . . . . . . . . . . . . . . . + . . + + + 1 17 
Jovibarba globifera . . . . . . . . . . . . . . . . . . . . . . . + + + . + + 17 
ES Gentiana clusii . . . . . . . . . . . + . . . . . . . . . . + 1 a . 1 . . 17 
AT Valeriana tripteris . . . . . . . . . . . . . . . . . . . . . . + . + + . . . 10 
Seslerion tatrae 
Cv Achillea *alpestris d 1 1 1 1 1 + + a . + + 1 . + 1 . 1 . . . . . . . . . + + + 59 
Cv Phleum hirsutum d + + + + + . . . . + . + + . 1 + . + + . . + . + . . . + + 55 
Gentiana verna t . . . + . . . . . . . . . . . . . + . . . . . + . . + + . 17 
Cv Linum extraaxillare d . . . + . . . . . . . . . + . 1 . . . . . . . 1 + . . . . 17 

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Number of relevé 0000000001111111111222 2222222 
1234506780123456789012 3456789 % 

Trifolium *kotulae C ................. ++ ... ....... 
7
Astragalus australis t ...................... ...... b 
3


Seslerietalia coerulae, Elyno-Seslerietea 
Carex *tatrorum 333313ab54433b33444333 4545454 100 
Scabiosa lucida 11+a1aa111a1a11++11+a+ a1 .++11 97 
Phyteuma orbiculare 11+1 .+++1+11111+1++ .1. +1+++11 90 
Bellidiastrum michelii +++++1 .a11a1+ ...... +1 . 111+1++ 72 
Festuca *versicolor 111aa1a .++b11aa ...... + 3ba1+ .+ 72 
Pedicularis verticillata aaa11++ .1.1+ .. +1 .+.+.. +11++ .1 69 
Galium anisophyllum ++ ..... a+++++++ .1++ .1. .++ .+++ 66 
Anthyllis *alpestris ... +... ++1b11 .+.... 1++ 1bbaaa . 59 
Helianthemum grandiflorum ....... 1.1+ .. +11a ..... a11 .+++ 45 
Knautia kitaibelii ....... +1a ...... +1+ ... a111+1+ 45 
Biscutella laevigata r.... 11 ... 1.. 1++ .... +. +111 .1. 45 
Euphrasia salisburgensis r.+... +... +++ ......... .. +++ .+ 34 
Thymus *sudeticus ....... 11+11 ........ 1. ....... 21 
Gentianella fatrae r........... +...... +++ ....... 17 

cf 
Salix alpina ......... a1 ........... .aa+ ... 17


cf 
Carex firma ..... 1................ +...... 
7


Mulgedio-Aconitetea 

Cv 
Campanula serrata ... ++++++++ .++++bbb .+. ++ .++ .. 69 

Cv 
Campanula elliptica ++ .+........ +.. +1+1 .. 1 +... +++ 45 
Geranium sylvaticum ... +... +.r.+.1.. +a+ ... .... +.r 34 

Cv 
Cirsium erisithales ............... ++ r+1 .+ ... ++++ 34


tf 
Rhodiola rosea ... 1......... +.. 1a+ ... ... +.. + 24
Senecio subalpinus ................ +++ .. + ....... 14


Cv 
Crepis mollis ............ +.+.++ .... ....... 14
Astrantia major ........ +.. ++ ......... ....... 10
Gentiana asclepiadea ............... +.+... + ....... 10
Bistorta major .... a........ a... r.... ....... 10
Solidago *minuta ....... 1........ +.+... ....... 10


Other taxa 
Leucanthemum margaritae +++1111++11+1 . +1aa11a+ +1++a+1 100 
Bartsia alpina aa1111+11111 .11a .++11+ 1++++ .+ 90 
Carlina acaulis ++ ...... 1++ .... ++aba1+ +11a1+b 66 
Soldanella carpatica ++ .++ .++11111+1+ .. +... ++ .... + 62 
Lotus corniculatus +.+1+ .. 1+ ..... +.baa .11 +.. +1+1 59 
Thymus alpestris ..... ++ ..... +.+.b1+ .. + +1+1+++ 52 
Heracleum sphondylium ........ +.... r.. ++++ .+ ... +++1 38 
Cardaminopsis arenosa agg. ....... +r+.++++ .+..... ++ ... +. 38 
Leontodon hispidus ......... 1...... bbb .1+ .+.++ .a 34 
Hieracium bifidum ....... r+r+....... +... .++1 .11 34 
Gentianella lutescens ......... ++ ........... 1++++++ 31 
Tragopogon orientalis .............. ++ .++ .. + .+.. ++ . 28 
Viola biflora .... 1.... +... a+1 r+.... .... +.. 28 
Dryas octopetala .. +..... b1b ......... a. bbm .... 28 
Hieracium sp. +.... +..... +.+.1... 1++ ....... 28 
Traunsteinera globosa r.r+.............. +++1 ...... 1 28 

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Number of relevé 
0000000001111111111222 2222222 
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Pc 
Saxifraga paniculata .... +............ 1+ .. + +..... 1 21
Hieracium lachenalii .+.... +.... +.1+ ..... +. ....... 21
Vicia cracca 11+ ................. +1 ....... 17
Trifolium pratense ... +++1 ......... 1..... ....... 17
Agrostis capillaris ................ +++++ . ....... 17
Orchis *signifera ................. ++ ... .+... ++ 17
Tofieldia calyculata .. r....... aa .. +....... .... a.. 17
Hieracium caesium ............. +........ .. +.1+ . 14
Antennaria dioica ..... ++ ............. +. .... +.. 14
Botrychium lunaria ........ +.. +...... +... ...... + 14
Hylocomium splendens (E0) .... 1.1...... a1 ....... ....... 14


Rarely occurring taxa: 

E1: 
Acetosa arifolia + (17); Allium victorialis + (7, 9); Androsace lactea + (29); Avenella flexuosa + (18, 19); Briza media + (21); 
Calamagrostis arundinacea 1 (22); C. varia + (25, 28); Campanula cochleariifolia + (25); Carex flacca + (27); Cerastium 
holosteoides + (17); Coeloglossum viride + (10); Crepis jacquinii + (29); Cruciata glabra + (18, 20, 24); Cyanus mollis + 
(16); Deschampsia cespitosa + (18), r (19); Euphrasia picta + (7); Festuca carpatica + (10, 23); Galium album + (25); Geum 
rivale + (17); Gymnadenia conopsea + (16, 23, 27); Hieracium murorum + (18, 20, 27); Hippocrepis comosa + (27); Huperzia 
selago + (10); Lilium martagon + (20); Linum catharticum + (13, 27, 28); Luzula multiflora + (7); Minuartia langii + 
(20, 26); Myosotis alpestris + (2, 6); Phleum rhaeticum + (5, 14, 16); Picea abies + (9), r (11); Pilosella lactucella + (7); 
Pinguicula alpina + (27); Plantago media + (5); Ranunculus acris + (13); Rhinanthus pulcher 2a (29); Rh. serotinus + (4); 
Salix silesiaca + (23); Saxifraga aizoides + (10, 27); Silene *prostrata + (21); Swertia *alpestris + (3), 2a (10, 11); Taraxacum 
officinale agg. + (27); Thalictrum aquilegiifolium r (26)+ Th. minus + (9); Trifolium repens + (5); Trommsdorfia uniflora r 
(8); Veronica chamaedrys + (12, 13); Vicia oreophila + (24, 28); V. sylvatica + (21). 

E0: 
Amblystegium varium + (29); Dicranum scoparium 1 (2, 7); Plagiochila porelloides + (10); Pleurozium schreberi 2a (4, 12); 
Pohlia cruda + (4); Rhytidium rugosum 2a (12), 1 (15); Sanionia uncinata 1 (7, 13), + (10); Schistidium apocarpum + (22); 
Tortella tortuosa 1 (10, 29), + (22). – Cetraria islandica 1 (4). 

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