107 Spiraea decumbens Koch subsp. tomentosa (Poech) Dostál, novelty for the flora of Slovenia and the Dinaric Alps Abstract In Govci under Mt. Poldanovec above the T rebuša Valley we found a new locality of southeastern-Alpine (Alpic) endemic Spiraea decumbens subsp. tomentosa, which is new to the flora of Slovenia and the Dinaric Alps. Its closest known localities are in the western Julian Prealps in Northeastern Italy. A small population of several ten shrublets occurs at elevations of about 570 m in shady, almost vertical dolomite rocks, in a stand of the endemic association Phyteumato columnae-Primuletum carniolicae. Despite a very small population we assume there are other localities in this area, where numerous gorges are very difficult to acces and have therefore not yet been sufficiently studied. Iz vleček V Govcih pod Poldanovcem nad dolino T rebuše smo našli novo nahajališče jugovzhodno-alpskega endemita Spiraea decumbens subsp. tomentosa, kar je novost za floro Slovenije in Dinarskega gorstva. Njegova najbližja znana nahajališča so v zahodnih Julijskih Predalpah v severovzhodni Italiji. Majhna populacija, nekaj deset grmičev, raste na nadmorski višini okoli 570 m v senčnem skoraj navpičnem dolomitnem skalovju, v sestoju endemične asociacije Phyteumato columnae- Primuletum carniolicae. Kljub zelo majhni populaciji domnevamo, da je nahajališč več, vendar so številne grape v tem območju zelo težko dostopne in še ne dovolj raziskane. Key words: flora, vegetation, endemic species, Spiraea decumbens, T rnovski Gozd plateau, Dinaric Alps, Slovenia. Ključne besede: flora, vegetacija, endemiti, Spiraea decumbens, T rnovski gozd, Dinarsko gorstvo, Slovenija. Corresponding author: Igor Dakskobler E-mail: igor.dakskobler@zrc-sazu.si Received: 1. 12. 2021 Accepted: 29. 1. 2022 1 Scientific Research Centre of the Slovenian Academy of Sciences and Arts, Institute of Biology, Regional unit Tolmin, Tolmin, Slovenia. Igor Dakskobler1 DOI: 10.2478/hacq-2022-0003 22/1 • 2023, 107–116 22/1 • 2023, 107–116 108 Igor Dakskobler Spiraea decumbens Koch subsp. tomentosa (Poech) Dostál, new for the flora of Slovenia Introduction Spiraea decumbens is an endemic species of the South- eastern Alps with known localities in northeastern Italy (provinces of Udine, Pordenone, Belluno and T reviso) and northwestern Slovenia (Kobarid municipality: Breginjski kot, and Bovec municipality: Učja valley) – Aeschimann et al. (2004: 726–728) and Pavlin et al. (2015). We dis- tinguish between two subspecies, type subspecies S. de- cumbens subsp. decumbens (sprawling spiraea, also white lace spiraea) and subspecies Spiraea decumbens subsp. to- mentosa, syn. S. decumbens subsp. hacquetii, Spiraea hac- quetii (which could be translated as Hacquet’s spiraea) – Poldini et al. (2001), Aeschimann et al. (2004), Pavlin et al. (2015), Pignatti (2017). The main difference between them is the hairiness of their leaves and inflorescences. The leaves, pedicels, inflorescence and fruits of the type subspecies are glabrous. The leaves of the subsp. tomentosa are tomentose on the upper side and below, often finely serrated only on the upper third; pedicels, receptacle and fruits are also hairy. The type subspecies occurs in Italian provinces of Udine and Pordenone (Friuli Venezia Giulia region) and in northwestern Slovenia, whereas the subspe- cies tomentosa has so far been known only in northeast- ern Italy, in the regions Friuli Venezia Giulia (provinces of Udine and Pordenone) and Veneto (provinces of Belluno and Treviso) – Poldini (2002), Aeschimann et al. (2004), Gobbo & Poldini (2005), E. Pignatti & S. Pignatti (2017). The type subspecies predominates in the eastern part of the species range (Julian Alps and Julian Prealps), but it is frequent also in the Carnic Alps (in the west all the way to the Lumiei valley) and the Carnic Prealps (up to Mt. Raut and Mt. Jouf di Maniago). Where the ranges of both subspecies overlap it is not uncommon for their populations to almost come into contact (Arzino Valley, Mt. Jouf di Maniago) – Martini, in litt. Fabrizio Martini (in litt.) finds that the distinction be- tween the two subspecies is not always unequivocal because of the atypical specimens (perhaps hybrids, introgressive forms) that also occur in the areas where the ranges of both subspecies overlap. According to detailed range maps for Friuli Venezia Giulia and based on his own experience Martini (in litt.) finds that the subspecies S. decumbens subsp. tomentosa occurs mainly in the western part of the species’ range (Carnic Alps to the west of the Tagliamento River, but individual sites are located also in the south- ern Carnic Alps (up to the Gleriis valley to the north of the town of Moggio Udinese) and in the western Julian Prealps. Melzer (1982) reported about the locality of this subspecies in the Julian Alps in the Val Saisera (Zajzera), under Mt. Jôf di Montasio (Montaž), on 1800 m a.s.l. (source: Schaeftlein, 1935, herbarium GZU). Currently known distribution of the type subspecies in Slovenia was published several years ago (Pavlin et al., 2015: 3, 8, 10). The data by grid squares are still valid, but recently (in the autumn of 2021) Marko Pavlin dis- covered a new locality on the sunny slopes of Mt. Stol, which is situated about 5 km more to the east than other localities under the Stol ridge. Spiraea decumbens occurs subspontaneously also in the Trenta Valley, near Mary’s chapel on the northern side of the road towards the Up- per Trenta Valley and Vršič, about 500 m from the side road exit to Zadnjica. The spiraea there was planted and spread subspontaneously in the rocks in the immedi- ate vicinity of the chapel (Simona Strgulc Krajšek, Filip Küzmič, written note and FloVegSi database). On 14 July 2021 we found a new locality of Spiraea de- cumbens in Govci above the T rebuša Valley, on the north- ern edge of the Trnovski Gozd plateau (9949/3), namely its tomentose subspecies S. decumbens subsp. tomentosa, which is new to the flora of Slovenia and the Dinaric Alps, and therefore deserves to be reported in more detail. Methods Spiraea decumbens subsp. tomentosa was found as we were looking for localities of Daphne blagayana in Govci above the Trebuša Valley. The relevés on its new locality and its surroundings were made using the standard Central- European phytosociological method (Braun-Blanquet, 1964) and entered into the FloVegSi database (T. Seliškar et al., 2003). The relevés were processed using hierarchical classification, unweighted average linkage method – UP- GMA and Wishart’s similarity ratio. We transformed the combined cover-abundance values into ordinal scale (1–9) according to van der Maarel (1979). Numerical compari- sons were performed with the SYN-TAX 2000 program package (Podani, 2001). Flora alpina (Aeschimann et al., 2004) and Flora d’Italia (Pignatti, 2017) were used for identifying differentiation characters between the two subspecies. While the opinion of the authors of the Flora alpina was that Spiraea decumbens subsp. hacquetii is its valid name, we follow the new edition of Flora d’Italia, us- ing as valid the name Spiraea decumbens subsp. tomentosa. For the proper Slovenian name of this taxon we propose the name “Hacquetova medvejka”, whereas the basionym of this taxon is Spiraea hacquetii Fenzel & Koch. The nomenclatural sources for the names of vascular plants are the Mala flora Slovenije (Martinčič et al., 2007) and FloVegSi database, the nomenclatural source for mosses is Hodgetts et al. (2020) and the nomenclatural sources for the names of syntaxa are Šilc & Čarni (2012), E. Pignatti & S. Pignatti (2014), Pavlin et al. (2015) and Mucina et al. (2016). 22/1 • 2023, 107–116 109 Igor Dakskobler Spiraea decumbens Koch subsp. tomentosa (Poech) Dostál, new for the flora of Slovenia Results and discussion New locality of subspecies Spiraea decumbens subsp. tomentosa 9949/3 (UTM 33TVL19) Slovenia: Primorska, T rnovski Gozd plateau, Govci above the T rebuša Valley, under Mt. Poldanovec, 570 m a.s.l., shady dolomite rocks in a small gorge, at the foot of its extremely steep, difficult-to-access upper (spring) area. Leg. et det. I. Dakskobler, 14. 7. 2021, herbarium LJS 12267. S. decumbens subsp. tomentosa occurs in the forested area under Mt. Poldanovec, in a small gorge that we pro- visionally call Ličerjeva Grapa (after the forester Franc Ličer) – Figures 1 and 3. T o the west of this gorge is a big- ger one called Špikova Grapa, and to the east of the lat- ter several more gorges, including Orlejška Grapa, which is the deepest. The locality is at the elevation of about Figure 1: Locality of Spiraea decumbens subsp. tomentosa in Govci under Mt. Poldanovec. Slika 1: Nahajališče podvrste Spiraea decumbens subsp. tomentosa v Govcih pod Poldanovcem. Figure 2: Natural localities of Spiraea decumbens in Slovenia (excluding its subspontaneous locality in the T renta Valley, quadrant 9648/2). Slika 2: Naravna nahajališča vrste Spiraea decumbens v Sloveniji (nje- nega subspontanega nahajališče v T renti, v kvadrantu 9648/2, nismo upoštevali).                                                                       Spiraea decumbens Spiraea tomentosa 22/1 • 2023, 107–116 110 Igor Dakskobler Spiraea decumbens Koch subsp. tomentosa (Poech) Dostál, new for the flora of Slovenia 570 m, the parent material (geological substrate) is dolo- mite, and the surrounding forest communities are classi- fied predominantly in associations Rhododendro hirsuti- Fagetum and Fraxino orni-Pinetum nigrae (Dakskobler, 1998, 1999, 2003). The spiraea grows in an almost verti- cal northwest-facing rock wall on the right bank of a nar- row gorge, covering only several square metres (Figure 4). The population comprises several ten shrublets (Figure 5). Based on its morphological characters – tomentose leaves, pedicel and receptacle – we determine it as S. decumbens subsp. tomentosa (Figure 6 and Figure 7, herbarium LJS 12267). Comparison with the photographs of the taxon Spiraea decumbens subsp. tomentosa from northeastern Italy, which were kindly forwarded to us by Fabrizio Mar- tini (photographs were taken by Claudio Peruzovich, Elio Polli, Manfredo Torelli and Giuliano Mainardis), shows that leaves of the specimens from Govci are not shiny, and according to this character they are very similar to the specimens photographed at the locality of Lake Verzegnis (Carnic Prealps). Figure 3: Gorge under Mt. Poldanovec, locality of Spiraea decumbens subsp. tomentosa. Photo: I. Dakskobler. Slika 3: Grapa pod Poldanovcem, kjer je nahajališče Hacquetove medvejke (Spiraea decumbens subsp. tomentosa). Foto: I. Dakskobler. Table 1 comprises companion species in the chasmo- phytic community. In addition to the relevé with Spiraea decumbens subsp. tomentosa we included also three relevés of chasmophytic communities in the immediate vicinity. Based on its species composition the stand with Hacquet’s spiraea under Mt. Poldanovec is classified into the associa- tion Phyteumato columnae-Primuletum carniolicae (Daks- kobler & Martinčič, 2020), provisionally into a new sub- unit (variant) with Spiraea decumbens subsp. tomentosa. Diagnostic species of the association are mainly Primula carniolica, Phyteuma scheuchzeri subsp. columnae and Paederota lutea. The first two are not known in Slovenia in communities with Spiraea decumbens subsp. decumbens (i.e. in the stands of associations Spiraeo-Potentilletum caulescentis, Aquilegio einseleanae-Spiraeetum decumben- tis and Spiraeo decumbentis-Seslerietum calcariae), and the third (Paederota lutea) occurs in some of the relevés – Pavlin et al. (2015). Phyteuma scheuchzeri subsp. colum- nae is presented in the stands of the association Spiraeo- Potentilletum caulescentis in NE Italy (Poldini, 1969; 22/1 • 2023, 107–116 111 Igor Dakskobler Spiraea decumbens Koch subsp. tomentosa (Poech) Dostál, new for the flora of Slovenia Figure 4: Population of Spiraea decumbens subsp. tomentosa under Mt. Poldanovec. Photo: I. Dakskobler. Slika 4: Populacija Hacquetove medvejke (Spiraea decumbens subsp. tomentosa) pod Poldanovcem. Foto: I. Dakskobler. Figure 6: Spiraea decumbens subsp. tomentosa, inflorescence. Photo: I. Dakskobler. Slika 6: Hacquetova medvejka (Spiraea decumbens subsp. tomentosa), posnetek socvetja. Foto: I. Dakskobler. Figure 5: Spiraea decumbens subsp. tomentosa under Mt. Poldanovec, close-up. Photo: I. Dakskobler. Slika 5: Hacquetova medvejka (Spiraea decumbens subsp. tomentosa) pod Poldanovcem, bližnji pogled. Foto: I. Dakskobler. Poldini, 1973). E. Pignatti & S. Pignatti (2014, 2016) published three association tables from the Dolomites in NE Italy, which comprise also Spiraea decumbens subsp. tomentosa: Potentilletum caulescentis (frequency 58%), Physoplexidi-Asplenietum seelosii (frequency 27%) and Saxifragetum burseranae (frequency 4 %). The species that are common in both their relevés with Spiraea tomentosa and our relevé from Govci include Paederota lutea, Phy- teuma scheuchzeri subsp. columnae, Valeriana saxatilis, As- ter bellidiastrum, Asplenium ruta-muraria, A. trichomanes, Sesleria caerulea, Carex mucronata and C. ferruginea. In the immediate vicinity of the locality of Spiraea de- cumbens subsp. tomentosa in Govci, but not in its stand, grows another endemic species, Aquilegia iulia, which is also unknown in other distribution ranges of Spiraea de- cumbens. Figure 7: Spiraea decumbens subsp. decumbens (Breginj – Prekopa, Leg. B. Vreš) – left and Spiraea decumbens subsp. tomentosa (Govci) – right. Photo: B. Vreš. Slika 7: Spiraea decumbens subsp. decumbens (Breginj – Prekopa, Leg. B. Vreš) – levo in Spiraea decumbens subsp. tomentosa (Govci) – desno. Foto: B. Vreš. 22/1 • 2023, 107–116 112 Igor Dakskobler Spiraea decumbens Koch subsp. tomentosa (Poech) Dostál, new for the flora of Slovenia Conclusions The differences between the flora and vegetation of the Julian Alps (as part of the Southeastern Alps) and the Trnovski Gozd Plateau (northeasternmost part of the Dinaric Alps) are not yet completely clear. Certain typi- cal Alpine (Alpic) species thus still grow on the north- ern edge of the Trnovski Gozd plateau, e.g. Campanula zoysii, Aquilegia iulia (southeastern-Alpine endemics) and Larix decidua. This area represents also the localities of other species that otherwise dominate mainly in the Alpine-Prealpine part of Slovenia: Campanula carnica, C. cochleariifolia, Cerastium subtriflorum, Draba aizoides, Dryas octopetala, Heliosperma alpestre, Pedicularis rosea, Potentilla clusiana, Ranunculus carinthiacus, R. hybridus, R. traunfellneri, Rhodothamnus chamaecistus, Saxifraga crustata, S. squarrosa, Senecio abrotanifolius (Dakskobler, 2019; Dakskobler & Vreš, 2021). It is therefore not sur- prising that the new locality of the southeastern-Alpine (Alpic) endemic Spiraea decumbens was discovered here, but its subspecies identity is unexpected, because most of the localities of S. decumbens subsp. tomentosa are situated to the west of the type subspecies, exclusively in north- eastern Italy. The population in Govci is very small and difficult to access, which prevents us from making more in-depth morphological or genetic comparisons with populations from Italy. Spiraea decumbens subsp. tomentosa was not observed in other gorges surveyed so far on the northern edge of the Trnovski Gozd plateau, under Stador, Stanov Rob, Poldanovec and Zeleni Rob. Because of the difficult and challenging terrain in Govci many gorges there are difficult to reach and we only succeeded in surveying a few of them. We assume that the population of Spiraea decumbens subsp. tomentosa on the northern edge of the T rnovski Gozd plateau is larger than the currently known population, and that other researchers, who are more ex- perienced climbers, are likely to find new localities of this subspecies. There are other montane species with a small population size (e.g. Botrychium simplex, Carex brunnes- cens, Nigritella archiducis-joannis) known in Slovenia, but we believe they are more threatened than the studied sub- species in Govci. The human impact in this part of the Trnovski Gozd plateau is insignificant. The threat factors consist mainly in natural disasters and climate change. Nevertheless, given its scarcity Spiraea decumbens subsp. tomentosa should be properly evaluated and considered in the Red List of threatened vascular flora of Slovenia (Anonymous, 2002). Povzetek Spiraea decumbens Koch subsp. tomentosa (Poech) Dostál, novost za floro Slovenije in Dinarskega gorstva Vrsta Spiraea decumbens je endemit Jugovzhodnih Alp z znanimi nahajališči v severovzhodni Italiji (province Udine, Pordenone, Belluno in Treviso) in severozahodni Sloveniji (občina Kobarid: Breginjski kot in občina Bo- vec: dolina Učje). Razlikujemo dve podvrsti, tipsko S. decumbens subsp. decumbens (polegla medvejka) in pod- vrsto Spiraea decumbens subsp. tomentosa, sin. S. decum- bens subsp. hacquetii (dlakava polegla medvejka, ker je bazionim te podvrste Spiraea hacquetii, zanjo predlaga- mo slovensko ime Hacquetova medvejka). Glavna razlika med njima je v dlakavosti listov in socvetja. Tipska pod- vrsta ima gole liste, cvetne peclje, socvetje in plodove. Podvrsta S. decumbens subsp. tomentosa ima na zgorn- ji in spodnji strani dlakave liste, ti so pogosto samo na zgornji tretjini drobno nazobčani, dlakavi so tudi cvetni peclji, cvetišče in plodovi. Tipska podvrsta uspeva v itali- janskih provincah Udine in Pordenone (dežela Furlanija Julijska krajina) in v severozahodni Sloveniji, podvrsta S. decumbens subsp. tomentosa pa je bila do zdaj znana le v severovzhodni Italiji, v deželah Furlanija Julijska kraji- na (pokrajini / provinci Udine in Pordenone) in Veneto / Benečija (pokrajini / provinci Belluno in Treviso). 14. 7. 2021 smo v Govcih nad dolino Trebuše, na severnem robu Trnovskega gozda (9949/3) našli novo nahajališče polegle medvejke in sicer njene dlakave podvrste (S. de- cumbens subsp. tomentosa), kar je novost za floro Sloveni- je in Dinarskega gorstva. Hacquetova medvejka raste v gozdnatem območju pod Poldanovcem, v manjši grapi, ki ji za zdaj še ne vemo imena, začasno jo imenujemo Ličerjeva grapa (po gozdarju Francu Ličerju) – sliki 1 in 3. Zahodno od nje je večja Špikova grapa, vzhodno od nje je več grap, med katerimi je najbolj globoka Orlejška grapa. Nahajališče je na nadmorski višini približno 570 m, geološka podlaga je dolomit, okoliške gozdne združbe uvrščamo predvsem v asociaciji Rhododendro hirsuti-Fa- getum in Fraxino orni-Pinetum nigrae. Hacquetova med- vejka raste v skoraj navpični, proti severozahodu obrn- jeni steni desnega brega ozke grape (prav tu se začenja njen zelo strm, težko prehoden zgornji, povirni del), na površini le nekaj kvadratnih metrov (slika 4). Populacija obsega nekaj deset grmičev (slika 5). Po morfoloških zna- kih, očitni dlakavosti listov, cvetnega peclja in cvetišča, jo določamo kot podvrsto S. decumbens subsp. tomentosa (sliki 6 in 7, herbarij LJS12267). Primerjava s fotogra- fijami taksona Spiraea decumbens subsp. tomentosa iz severovzhodne Italije, ki nam jih je prijazno posredoval 22/1 • 2023, 107–116 113 Igor Dakskobler Spiraea decumbens Koch subsp. tomentosa (Poech) Dostál, new for the flora of Slovenia prof. Fabrizio Martini (avtorji fotografij so Claudio Pe- ruzovich, Elio Polli, Manfredo Torelli in Giuliano Mai- nardis), kažejo, da listi primerkov iz Govcev niso bleščeči in so po tem znaku zelo podobni primerkom, ki so jih fotografirali na nahajališču Lago di Verzegnis (Karnij- ske Alpe). Spremljevalne vrste v združbi skalnih razpok kaže preglednica 1. Poleg popisa, v katerem je medvejka, smo v preglednico uvrstili še tri popise skalnih razpok, ki smo jih naredili v neposredni bližini. Po vrstni sestavi sestoj s poleglo medvejko pod Poldanovcem uvrščamo v asociacijo Phyteumato columnae-Primuletum carniolicae, začasno v novo podenoto (varianto) s podvrsto Spiraea decumbens subsp. tomentosa. Diagnostične vrste za to asociacijo so predvsem Primula carniolica, Phyteuma scheuchzeri subsp. columnae in Paederota lutea. Prvi dve ne poznamo v združbah, kjer v Sloveniji uspeva pod- vrsta Spiraea decumbens subsp. decumbens (to je v sesto- jih asociacij Spiraeo-Potentilletum caulescentis, Aquilegio einseleanae-Spiraeetum decumbentis in Spiraeo decumben- tis-Seslerietum calcariae), tretja (Paederota lutea) pa se po- javlja v nekaterih popisih. V neposredni bližini nahajališ- ča Hacquetove medvejke, a ne v njenem sestoju, raste še en endemit, Aquilegia iulia, ki ga prav tako ne poznamo v drugih območjih razširjenosti vrste Spiraea decumbens. Med Julijskimi Alpami (kot delom Jugovzhodnih Alp) in Trnovskim gozdom (skrajnim severovzhodnim delom Dinarskega gorstva) v rastju in rastlinstvu ni ostrih meja. Zato tukajšnje novo nahajališče jugovzhodnoalpskega endemita Spiraea decumbens ne preseneča, pač pa njego- va podvrstna pripadnost, saj ima podvrsta S. decumbens subsp. tomentosa večinoma nahajališča bolj zahodno od tipske podvrste, samo v severovzhodni Italiji. Resnejše morfološke ali genetske primerjave s populacijami iz Ita- lije za zdaj niso mogoče, saj je populacija v Govcih zelo majhna in težko dostopna. V drugih do zdaj pregledanih grapah na severnem robu Trnovskega gozda, pod Stador- jem, Stanovim robom, Poldanovcem in Zelenim robom, podvrste Spiraea decumbens subp. tomentosa nismo opazi- li. Vendar so številne grape v Govcih zelo težko prehodne in nam je uspelo priti le v nekaterih od njih. Domneva- mo, da je populacija Hacquetove medvejke na severnem robu Trnovskega gozda večja od te, ki jo poznamo zdaj in bodo plezalsko bolj vešči raziskovalci verjetno našli njena nova nahajališča. Ker so v tem delu Trnovskega gozda človekovi vplivi zelo majhni, ni zelo ogrožena. De- javnik ogrožanja so predvsem naravne ujme in podnebne spremembe. Kljub temu za podvrsto Spiraea decumbens subsp. tomentosa zaradi redkosti predlagamo za uvrstitev na rdeči seznam (Anonymous, 2002). Acknowledgements Prof. Dr. Andrej Martinčič very kindly determined se- veral moss species in the relevés included in Table 1. Prof. Dr. Fabrizio Martini kindly forwarded the data on the currently known distribution ranges of subspecies of Spiraea decumbens in northeastern Italy and the photo- graphs of the studied subspecies (by Claudio Peruzovich, Elio Polli, Manfredo Torelli and Giuliano Mainardis) – I am very grateful to all of them. Marko Pavlin, Ass. Prof. Dr. Simona Strgulc Krajšek and Dr. Filip Küzmič allowed me to cite their recent data that supplement the species range of Spiraea decumbens in Slovenia. Iztok Sajko pre- pared Figure 1 and Figure 2 for print. Dr. Branko Vreš is the caretaker of FloVegSi database and curator of the Her- barium LJS. 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Hacquetia, 11(1), 113–164. 22/1 • 2023, 107–116 115 Igor Dakskobler Spiraea decumbens Koch subsp. tomentosa (Poech) Dostál, new for the flora of Slovenia Number of relevé (Zaporedna številka popisa) 1 2 3 4 Pr. Fr. Database number of relevé (Delovna številka popisa) 286298 286306 286308 286307 Elevation in m (Nadmorska višina v m) 570 565 575 565 Aspect (Lega) NW NNW N NW Slope in degrees (Nagib v stopinjah) 85 70-100 85 90 Parent material (Matična podlaga) D D D D Soil (Tla) Li Li Li Li Stoniness in % (Kamnitost v %) 100 100 100 100 Cover of herb layer in % (Zastiranje zeliščne plasti v %): E1 30 30 30 30 Co ver of moss layer in % (Zastiranje maho vne plasti v %) E0 20 20 10 20 Number of species (Število vrst) 27 21 30 29 Relevé area (Velikost popisne ploskve) m 2 5 10 10 15 Date of taking relevé (Datum popisa) 14.07.2021 14.07.2021 14.07.2021 14.07.2021 Locality (Nahajališče) Govci Govci Govci Govci Quadrant (Kvadrant) 9949/3 9949/3 9949/3 9949/3 Coordinate GK Y (D-48) m 410544 410538 410521 410543 Coordinate GK X (D-48) m 5097496 5097510 5097522 5097507 Diagnostic species of the association (Diagnostične vrste asociacije) AP Paederota lutea E1 1 1 1 2 4 100 PcSp Phyteuma scheuchzeri subsp. columnae E1 + 1 1 1 4 100 AP Orthothecium rufescens E0 + 1 1 1 3 100 AP Primula carniolica E1 r + + . 3 75 AP Valeriana tripteris E1 1 . + + 3 75 AP Cystopteridion s. lat. (Astrantio-Paederotion luteae nom. prov.) Aster bellidiastrum E1 + 1 1 1 4 100 Valeriana saxatilis E1 + 1 1 2 4 100 Hydrogonium croceum (Barbula crocea)* E0 + + 1 1 4 100 Asplenium viride E1 + . + + 3 75 Fissidens dubius E0 + + . + 3 75 Jungermannia atrovirens* E0 + . . + 2 50 Pinguicula alpina E1 . . 1 + 2 50 Astrantia carniolica E1 . + . . 1 25 Viola biflora E1 . + . . 1 25 Tofieldia calyculata E1 . . 1 . 1 25 Marchantia quadrata (Preissia quadrata)* E0 . . + . 1 25 PcSp Physoplexido comosae-Saxifragion petraeae Spiraea decumbens subsp. tomentosa E1 1 . . . 1 25 AT Asplenietea trichomanis Asplenium ruta-muraria E1 + . + + 3 75 TR Thlaspietea rotundifolii Aquilegia iulia E1 . + r + 3 75 Gymnocarpium robertianum E1 + . . + 2 50 Adenostyles glabra E1 1 . + . 2 50 Hieracium bifidum E1 . . + + 2 50 MC Montio-Cardaminetea Conocephalum conicum E0 + . . 1 2 50 Palustriella commutata E0 . + . . Cololejeunea calcarea* E0 . . + . 1 25 Table 1: Rock crevice communities in Govci under Mt. Poldanovec. Tabela 1: Združbe skalnih razpok v Govcih pod Poldanovcem. 22/1 • 2023, 107–116 116 Igor Dakskobler Spiraea decumbens Koch subsp. tomentosa (Poech) Dostál, new for the flora of Slovenia Number of relevé (Zaporedna številka popisa) 1 2 3 4 Pr. Fr. ES Elyno-Seslerietea Carex ferruginea E1 + + + + 4 100 Sesleria caerulea E1 1 1 1 . 3 75 Carex mucronata E1 + . 1 + 3 75 Laserpitium peucedanoides E1 + . + . 2 50 Betonica alopecuros E1 . . + + 2 50 FB Festuco-Brometea Buphthalmum salicifolium E1 + r + . 3 75 EP Erico-Pinetea Rhododendron hirsutum E1 . + r + 3 75 Rhodothamnus chamaecistus E1 . + . + 2 50 Calamagrostis varia E1 . . + + 2 50 Carex ornithopoda E1 . + . . 1 25 Rubus saxatilis E1 . . . r 1 25 VP Vaccinio-Piceetea Homogyne sylvestris E1 . . . + 1 25 AF Aremonio-Fagion Cyclamen purpurascens E1 + . . . 1 25 TA Tilio-Acerion Geranium robertianum E1 . + + . 2 50 Acer pseudoplatanus E1 . . + + 2 50 FS Fagetalia sylvaticae Salvia glutinosa E1 + . 1 + 3 75 Galium laevigatum E1 + . . . 1 25 QP Quercetalia pubescenti-petraeae Fraxinus ornus E2a + . . . 1 25 QF Querco-Fagetea Carex digitata E1 . . . + 1 25 ML Mosses (Mahovi) Mesoptychia collaris (Lophozia collaris)* E0 + + . + 3 75 Ctenidium molluscum E0 + . + . 2 50 Cyrtomnium hymenophylloides* E0 + . . + 2 50 Riccardia palmata* E0 . + . . 1 25 Tortella tortuosa E0 . . + . 1 25 Legend / Legenda D Dolomite – dolomit Li Lithosol – kamnišče Pr. Presence (number of relevés in which the species is presented) – število popisov, v katerih se pojavlja vrsta Fr. Frequency in % – frekvenca v % * determined Andrej Martinčič / določil Andrej Martinčič