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ACTA ENTOMOLOGICA SLOVENICA
LJUBLJANA, DECEMBER 2006    Vol. 14, ot. 2: 131-156
REVIEW OF THE ANT FAUNA (HYMENOPTERA: FORMICIDAE) OF
CROATIA
Gregor BRAOKO
Biotechnical Faculty, Department of Biology, Veona pot 111, SI-1000 Ljubljana,
Slovenia e-mail: gregor.bracko@bf.uni-lj.si
Abstract - The ant fauna of Croatia is relatively poorly known. There are almost no recent literature data, while past investigations were mainly restricted to the coastal part of the country. This study represents general review of the ant fauna for the whole territory of Croatia. Published data and unpublished material from different collections were used and recent taxonomic revisions were considered. Ant list brings 140 species with their known localities. Twenty-three species are mentioned for the first time for Croatia. Most of the records come from the Mediterranean part of the country, where the diversity of species is greatest. Additional species are expected to be found in further investigations in that region as well as in other parts of the country.
Key words: ants, Formicidae, fauna, Croatia
Izvleoek - PREGLED FAVNE MRAVELJ (HYMENOPTERA: FORMICIDAE) HRVAOKE
Favna mravelj Hrvaoke je razmeroma slabo poznana. Novejoih podatkov iz literature skorajda ni, medtem ko so bile pretekle raziskave veoinoma omejene na obalni del draave. Ta otudija predstavlja splooen pregled favne mravelj celotnega ozemlja Hrvaoke. Uporabljeni so bili ae objavljeni podatki in neobjavljen material iz razlionih zbirk, upootevane so bile novejoe taksonomske revizije. Seznam mravelj obsega 140 vrst z njihovimi znanimi najdiooi. Triindvajset vrst je prvio omenjenih za Hrvaoko. Veoina podatkov izhaja iz mediteranskega dela draave, kjer je raznovrstnost najveoja. Pri nadaljnjih raziskavah lahko prioakujemo oe dodatne vrste s tega obmooja in tudi iz drugih delov draave.
Kljuone besede: mravlje, Formicidae, favna, Hrvaoka
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Introduction
Insects, the most diverse animal group, were an object of great interest in scientific research for ages. However, different insect groups have not been equally treated. Ants, for example, deserved much less attention than some other insects although they are very often encountered and found almost everywhere in the terrestrial habitats. Investigations dealing with ants increased considerably in recent years. Taxonomic studies were especially productive. In European ant fauna, revisions of some genera (e.g. Myrmica, Leptothorax (Temnothorax), Lasius, Formica) were done and many new species were described. Updated faunistic reviews of numerous European countries were also performed so that much better insight into European ant fauna is now possible.
Unfortunately, the investigations of ant fauna in Croatia in recent time were rather moderate. Majority of published data is older than 50 years and originates mainly from the coastal Croatia. Also the beginnings of myrmecological expeditions were limited to the coastal part of the country, especially Dalmatia. The history of investigations of ant fauna in Dalmatia was well documented by Nonveiller (1989). First published data for Croatia can be found in the famous Formicina austriaca (Mayr 1855). Among the collectors who contributed material for Mayr's work, Georg von Frauenfeld yielded most of the records for Dalmatia after his visit to the region a year earlier. At the end of the 19th and in the beginning of the 20th century, there were several authors who published their finds in Dalmatia (Gasperini 1887, 1889, Katuria 1891, Galvagni 1902, Mayr & Kohl 1908, Tölg & Fahringer 1911). Important contribution was made by Croatian author Korlevia (1890) who listed ants from Rijeka region, Gorski Kotar and Pregrada in Hrvatsko Zagorje. Part of his material is kept in the Croatian Natural History Museum in Zagreb. The most comprehensive work in the earlier 20th century is a review of the ant fauna of Venezia Giulia and Dalmatia (Müller 1923). This also very important taxonomical work brings many data on ant species from Italian, Slovenian and Croatian part of former Venezia Giulia and Dalmatia. It includes material collected by different colleagues and several literature sources, some already mentioned above. Later contributions by Soudek (1925) and Zimmermann (1934) for Dalmatia, and Zimmermann (1930) for Quarner islands must be mentioned, too. Since the investigations of ant fauna were oriented to the Mediterranean Croatia, the work by Vogrin (1955) is very important, as he included a wider territory of Croatia (and other parts of former Yugoslavia) in it. Müller (1957) brought some information on ants from Dugi otok and nearby islands. In the catalogue of Formicidae of Italy, Baroni Urbani (1971) also included Istria and Rijeka in the area of his study. Agosti & Collingwood (1987a) and Petrov & Collingwood (1992) were collecting ants here as well, but they listed species only for the whole territory of former Yugoslavia.
The Croatian Natural History Museum keeps an ant collection with a great amount of the specimens dating from the end of the 19th century. Some of this material is published in Korlevia (1890), Vogrin (1955) and Müller (1957). There are
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some ant specimens from Rovinj, Pula and Krk island held in the Slovenian Museum of Natural History as part of the Hymenoptera collection of Evgen Jaeger (1880-1960).
The present review of the ant fauna of Croatia is based on the literature sources and up to now unpublished collected material. Many records come from older literature and most of this material was not available for identification. Considering recent taxonomic revisions, there can be some changes in taxonomic position and distribution pattern of certain species. This review should also be a stimulus for other myrmecologists to contribute their eventual data to supplement the list of Croatian ants.
Material and methods
The data from the literature and unpublished material from different collections were used for the present review. Cedric Collingwood (Skipton, UK) sent his list of ants sampled in the territory of Croatia during the expedition to the Balkans in middle eighties. Ants from the author's collection were gathered by the author and Al Vrezec (National Institute of Biology, Ljubljana) sporadically in the past 10 years, using direct sampling and pitfall traps. Ants collected by Tonai Rafia (Split) in Dalmatia in years 2005 and 2006 are kept in the National Museum of Zadar. Paula Durbeoia and Lucija Oeria Jelaska (Department of Zoology, Faculty of Science in Zagreb) sampled ants in Gorski Kotar, Medvednica and Vis island, and they are deposited at the Department of Zoology. All mentioned material, except that of C. Collingwood, and ants from the Croatian Natural History Museum and Slovenian Museum of Natural History were checked by the author. Majority of specimens were identified using taxonomic keys in Kutter (1977), Agosti & Collingwood (1987b) and Seifert (1996). Ant classification and taxonomic nomenclature generally follow Bolton (1995, 2003).
Many older studied literature give Italian geographic names for the localities of recorded species. All the names used in the list of species here are Croatian and correspond to the Veliki atlas Hrvatske (2002). Abbreviation “i.” stands for island. For the position of less known localities see the Appendix.
The references for locality data are indicated by a number in parenthesis in the list of ant species (with the exception of some individual references which are given with the name of the author and year): (1) - Mayr 1855, (2) - Gasperini 1887, (3) -Gasperini 1889, (4) - Korlevia 1890, (5) - Galvagni 1902, (6) - Tölg & Fahringer 1911, (7) - Müller 1923, (8) - Soudek 1925, (9) - Zimmermann 1930, (10) -Zimmermann 1934, (11) - Vogrin 1955, (12) - Müller 1957, (13) - Baroni Urbani 1971, (14) - Buschinger et al. 1988, (15) - collection of the Croatian Natural History Museum, (16) - collection of the Slovenian Museum of Natural History, (17) -collection of the National Museum of Zadar (leg. T. Rafia), (18) - collection of P. Durbeoia and L. Oeria Jelaska, (19) - Collingwood pers. comm., (20) - author's collection.
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List of ant species
Subfamily AMBLYOPONINAE Forel, 1893
Amblyopone denticulata (Roger, 1859)
Localities in CRO: Limski kanal (13); Kaotela (7); Lapad (10)
Subfamily PONERINAE Lepeletier, 1835
Cryptopone ochracea (Mayr, 1855) Localities in CRO: Zemunik (7); i. Hvar (3)
Hypoponera eduardi (Forel, 1894)
Localities in CRO: Uoka (13); Solin, Dubrovnik (7)
Ponera coarctata (Latreille, 1802)
Localities in CRO: Savudrija, Rovinj, Veruda (7); Uoka (13); near Poreo (20); Orehovica
(4); Draga (15); Rijeka, Bakar (Csosz & Seifert 2003); Velika Kapela (19); Zemunik,
Kistanje, Kaotela (7); Dubrovnik, Lapad, Rijeka Dubrovaoka (10); Krapina (11); Narta,
Hlebine, Milanovac, near Slavonski Brod (20)
Note: Csosz & Seifert (2003) gave P. coarctata var. testacea Emery, 1895 a status of a
species. Some of the literature data can therefore refer to P. testacea.
Ponera testacea Emery, 1895
Localities in CRO: Bakar, Senj, Gospia, Kaotel Suaurac (Csosz & Seifert 2003); Bjelina
(17)
Note: See comment under P. coarctata.
Subfamily PROCERATIINAE Emery, 1895
Proceratium algiricum Forel, 1899
Localities in CRO: Kaotela, near Dubrovnik (7); Dubrovnik (10)
Proceratium melinum (Roger, 1860) Localities in CRO: Konavle (10)
Subfamily MYRMICINAE Lepeletier, 1835
Aphaenogaster epirotes (Emery, 1895)
Localities in CRO: Rovinj, Vodnjan (7); Pula (13) (20); Veaica (4 - indicated as A. subterranea); i. Rab, i. Cres (9); Zadar, Bokanjac (7); Lioane Ostrovioke, Vinioae, i. Oiovo (Okrug Gornji), Ovrljevo (17); Marjan (10) (17); Kaotel Lukoia, Leaevica, i. Hvar (Jelsa), i. Vis (7); i. Hvar (Brusje) (2); Seoca, i. Brao (Milna) (17); Podgora, i. Koroula, i. Kopiote (10); Donja Vruoica (20); Trsteno (8); Dubrovnik (19); i. Oipan, i. Kolooep, Srd, Grua, Lapad, Rijeka Dubrovaoka, near Oibaoa, i. Lokrum, Konavle (10)
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Note: Some authors treat this species as a subspecies or variety of A. obsidiana (Mayr, 1861). A. obsidiana s. str. seems to be absent from major part of the Balkans (Atanassov & Dlussky 1992).
Aphaenogaster finzii Müller, 1921
Localities in CRO: Teradige (7); i. Krk (Krk) (9); Aegar, Zemunik (7); Vrpolje, Ovrljevo, Arnovnica (17); Marjan, between Klis and Solin, Podgora (10); i. Brao (Milna) (17); Gornja Vruoica (20); i. Bisaoe, i. Suoac, i. Kolooep, Dubrovnik, Rijeka Dubrovaoka, Oibaoa (10)
Aphaenogaster ionia Santschi, 1933
Localities in CRO: i. Krk (16); i. Krk (near Krk, between Skrboiai and Malinska), i. Rab
(Rab) (9); Senj (11) (15); near Zadar (1); Nin, Bibinje, Babindub, i. Ugljan (7); i. Dugi
otok (Zaglav) (20); near Sinj (1); Trogir (10); i. Oiovo (17); Split (2) (19); Solin, Kaotel
Stari, i. Vis (7); i. Hvar (10) (20); i. Sv. Klement, i. Bioevo, i. Svetac (10); i. Koroula (7)
(8) (20); i. Otok, Orebia, i. Mljet, Dubrovnik, Grua, Lapad, i. Lokrum (10)
Note: This quite common species along the Adriatic coast is stated as A. testaceopilosa
var. balcanica Emery, 1898 in Müller (1923) and Zimmermann (1930). However, A.
balcanica is restricted to eastern Balkans, Aegean Islands and Turkey (Zimmermann
1934). The data in Mayr (1855) and Gasperini (1887) for A. senilis Mayr, 1853 also refer
to A. ionia.
Aphaenogaster muelleriana Wolf, 1915
Localities in CRO: i. Rab (Rab, Lopar) (9); Sukooan (17); Trsteno (8); Dubrovnik, Grua,
Lapad (10)
Aphaenogaster splendida (Roger, 1859) Localities in CRO: Split (7); Baovice (8); i. Vis (2)
Aphaenogaster subterranea (Latreille, 1798)
Localities in CRO: Savudrija, Rovinj, Vodnjan, Oijana (7); near Poreo, Limski kanal, Pula (20); Orehovica (4); Rijeka (13) (15); Bakar (9); Senj (11); Velika Kapela (19); i. Krk (16); i. Rab (Rab) (9); i. Cres (7) (9) (20); Kistanje (7); Oskoruono (20); Konavle (10); Sljeme (18); near Slavonski Brod (20)
Aphaenogaster subterraneoides Emery, 1881 Localities in CRO: i. Hvar (Vrbanj) (7)
Chalepoxenus muellerianus (Finzi, 1922)
Localities in CRO: i. Krk (near Baoka), i. Pag (Kolan), Ljubao (14); i. Vir (Torovi) (20);
near Dubrovnik (14)
Crematogaster ionia Forel, 1911 Localities in CRO: Osijek (19)
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Crematogaster schmidti (Mayr, 1853)
Localities in CRO: Livade, Lovran, Smiloia (7); Dugi otok (12); Ovrljevo, Podaopilje (17); i. Koroula (Koroula) (10) (20); Kuoiote (7); Oskoruono, Kuna (20); Orebia, Slano, i. Kolooep, near Grua, near Oibaoa, i. Lokrum, i. Daksa (10); Dubrovnik, Lapad (7) (10); Konavoski Dvori (20); Prekriaje (15); Naoice, near Slavonski Brod (20)
Crematogaster scutellaris (Olivier, 1792)
Localities in CRO: near Livade, Vrsar - Limski kanal, Oijana (7); near Poreo, Limski kanal, Pula (20); near Rijeka (1); Orehovica (4); Suoak (13); Rijeka, Cicilinovo, Crikvenica, Senj (11); i. Krk (9) (11) (16); i. Rab (11) (20); i. Rab (Rab, Sv. Ilija, Banjol, Dundo, Lopar, near Blasio, Rt Zidine), i. Goli, i. Cres (9); i. Cres (Vodice) (20); i. Unije, i. Susak, i. Kozjak, i. Greben (7); i. Pag (15) (20); near Zadar (1); Nin, Arbanasi (7); Sukooan (17); i. Piokera, i. Vir (Torovi), i. Dugi otok (Zaglav) (20); i. Dugi otok (12); Lioane Ostrovioke, Split (17); near Split, near Makarska (1); Solin, Kaotela, Trogir (7); Marjan (7) (10) (17); Solin, Dugopolje, Podgora (10); i. Brao (Kneza Ravan, Bol, Nereaioaa) (7); i. Hvar (2) (7) (11) (20); i. Sv. Klement (10); i. Vis (2); i. Vis (Komiaa) (5) (7); i. Vis (Brguljac) (18); i. Svetac (7); i. Mali & Veli Paraan, i. Bioevo, i. Lastovo, i. Kopiote, i. Veli Lastovljan, i. Mroara (10); i. Suoac (7) (10); i. Koroula (Koroula) (11) (20); Rijeka Dubrovaoka, Lapad (6)
Crematogaster sordidula (Nylander, 1849)
Localities in CRO: Piaan, Rovinj, Oijana, Veruda, Volosko, Lovran (7); Pula (16) (20); Limski kanal (20); Rijeka (11) (13); Cicilinovo, Orehovica, Veaica (4); Novi Vinodolski, Senj, i. Krk (Jurandvor) (11); i. Rab (Rab), i. Oruda, i. Looinj (Osorooica), i. Zabodarski, i. Trasorka, i. Kozjak, i. Grujica, i. Orjule, i. Sv. Petar, i. Greben (7); i. Cres, i. Rab (9); Modria (17); near Zadar (1); Krmoina (7); i. Vir (Vir) (20); i. Dugi otok (Sali), i. Tukoooak (12); Vinioae (17); Labin (2); Split (1) (2); Kaotela, between Klis and Solin (7); Marjan (10) (17); Podaopilje (17); Podgora, i. Brao (Bol) (10); i. Hvar (Brusje) (2); i. Vis (Komiaa), i. Bioevo (7); i. Vis (Podstraaje) (18); Donja & Gornja Vruoica, i. Koroula (near Vela Luka) (20); i. Koroula (Sv. Antun) (7); i. Koroula (Koroula), i. Suoac (10); i. Lastovo (1); i. Palagruaa (5) (11); i. Mljet, Dubrovnik (7) (10); Rijeka Dubrovaoka (6) (7) (10); Lapad (6) (10); Slano, near Grua, i. Lokrum, i. Lopud (10)
Formicoxenus nitidulus (Nylander, 1846)
Localities in CRO: Mune, Lupoglav (7); Krapina (11)
Leptothorax acervorum (Fabricius, 1793)
Localities in CRO: Leska (18); near Oabar (20); Velika Kapela (19); Kalnik (11)
Leptothorax gredleri Mayr, 1855 Localities in CRO: Velika Kapela (19)
Leptothorax muscorum (Nylander, 1846)
Localities in CRO: near Oabar (20); Velika Kapela (19)
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Manica rubida (Latreille, 1802)
Localities in CRO: Lokve, Zlobin (11); near Oabar (20); Kuaelj, Jasenak (15); Zagreb
(11) (15)
Messor capitatus (Latreille, 1798)
Localities in CRO: Momjan (7); Limski kanal, Pula (20); near Rijeka (1); Rijeka, Veaica
(4); Novi Vinodolski (17); Bakar, Senj, Jablanac (11); i. Krk (between Krk and Baoka,
Treskavac), i. Plavnik, i. Mali Plavnik (9); i. Krk (Baoka) (11); i. Krk (16); i. Rab (Sv.
Matej, Kamenjak), i. Dolin, i. Cres (9); i. Looinj (Sv. Ivan) (7); i. Pag (20); near Zadar
(1); Zemunik, Smiloia, Aegar (7); Benkovac, Drnio (2); Knin (11); Oibenik (19); i. Dugi
otok (Sali, Salsko polje, Dobro polje), i. Tukoooak (12); i. Kornat (20); near Split, near
Makarska (1); Gornje Vinovo (17); Leaevica, Gata, Kotlenice (7); Marjan (8) (10);
Podgora, i. Hvar, i. Sv. Klement, i. Mali Paraan (10); i. Hvar (2); i. Vis (2) (11); i. Vis
(Komiaa), i. Lastovo (Polje) (5); i. Lastovo (2); i. Suoac (7) (10); i. Palagruaa (5) (11); i.
Koroula (Vela Luka) (10) (20); Kuoiote (7); Sv. Ilija, i. Mljet (10); near Dubrovnik (1);
Grua (6); Dubrovnik, Srd, Rijeka Dubrovaoka (10); Dalj (11)
Note: This species is indicated as Messor (or Aphaenogaster) barbarus in older literature.
True M. barbarus (Linnaeus, 1767) is probably absent from the studied territory.
Messor structor (Latreille, 1798)
Localities in CRO: Kanfanar, Vodnjan, Lovran (7); Rovinj (16); Rijeka, Orehovica (4); Grobnioko polje, Bribir, Novi Vinodolski, Senj, i. Krk (Lakmartin) (11); i. Krk (16); i. Krk (between Skrboiai and Malinska, Jurandvor), i. Rab (Rab, Banjol, Sv. Matej, Sv. Ilija, Lopar) (9); i. Cres (near Martinoaica) (1); i. Unije (7); Zadar (1) (7) (11); Privlaka (20); Bokanjac, Diklo, Arbanasi, Bibinje, i. Ugljan, Siveria (7); i. Oiovo (Okrug Gornji) (17); near Split (1); Split (7) (17) (19); Kaotel Suaurac, Kaotel Stari, Solin, Gata (7); Marjan (10) (17); Klis, Podgora (10); Podaopilje, Trnbusi (17); i. Brao (2); i. Hvar, i. Vis (7); i. Bioevo (7) (10); i. Svetac (10); near Dubrovnik (1); Dubrovnik, Lapad, Grua, Oibaoa, i. Lokrum, i. Oipan (10); Dalj (11)
Messor wasmanni Krausse, 1910
Localities in CRO: Markovac, Vodnjan (7); Limski kanal (20); Pula (16) (20); i. Rab (7); i. Rab (Rab, Banjol, Lopar, Jamina), i. Goli, i. Cres (9); i. Cres (Vrana) (20); i. Unije, i. Grujica (7); i. Pag (Novalja, Lun) (15); Zadar, Babindub (7); Privlaka (20); i. Dugi otok (Sali, Salsko polje) (12); Veliki Drvenik (10); Kaotela, Solin (7); Marjan, Klis, between Klis and Solin, Dugopolje (10); i. Brao (Bol, Supetar, Milna) (7); i. Brao (Milna) (17); i. Hvar (Jelsa, Hvar), i. Sv. Klement (10); i. Vis (7) (15); i. Vis (Brguljac) (18); i. Vis (Vis, Komiaa), i. Svetac, i. Suoac, i. Lastovo, i. Palagruaa (10); i. Koroula, i. Mljet (7) (15); Kuoiote (7); Orebia, Trpanj, i. Olipa, i. Jakljan, i. Lopud, i. Kolooep, Grua, near Oibaoa, i. Lokrum (10); Lapad (7) (10)
Monomorium monomorium Bolton, 1987
Localities in CRO: i. Unije, Solin (7); Klis (8); Split (19); Grua, Cavtat (8)
Monomorium pharaonis (Linnaeus, 1758) Localities in CRO: Pula (7); Rijeka (4); Zagreb (11)
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Myrmecina graminicola (Latreille, 1802)
Localities in CRO: Savudrija, Momjan, Tar, Pula, Oijana (7); near Poreo (20); Draga, Orehovica (4); Trsat (15); i. Krk (Ponikva, between Krk and Malinska), i. Cres (9); i. Looinj (Veli Looinj), Zadar, Zemunik, Kistanje, Kosore, Kaotela, i. Hvar (Vrbanj, Jelsa) (7); i. Hvar (Brusje) (2); Makarska (8); Dubrovnik (10); Sljeme (18); Krapina (11)
Myrmica gallienii Bondroit, 1920
Localities in CRO: Plitvice (15); Krbava (17); Narta (20); Osijek (19)
Myrmica hirsuta Elmes, 1978 Localities in CRO: Velika Kapela (19)
Myrmica lobicornis Nylander, 1846
Localities in CRO: Leska (18); Velika Kapela, Oibenik (19)
Myrmica lonae Finzi, 1926
Localities in CRO: Leska (18); Plitvice (15)
Note: this species had been treated as a subspecies or variety of M. sabuleti Meinert,
1861. Seifert (1993, 1996, 2000) raised it to species. On the basis of the checked material,
M. lonae seems to be much rarer than M. sabuleti in Croatia.
Myrmica rubra (Linnaeus, 1758)
Localities in CRO: near Rijeka (4); near Oabar (20); Delnice, Fuaine (11); Leska (18); Dretulja spring (15); Zagreb, Podsused, Kraljioin Zdenac, Pregrada, Krapina, Krapinske Toplice, Trnovec, Jagnjedovac (11); Hlebine, Narta, Milanovac (20)
Myrmica ruginodis Nylander, 1846
Localities in CRO: Mune (7); Rijeka (4); Senj (11); Leska (18); Plitvice (15); Velebit,
Paklenica (7); Medvednica (18); Krapina (11); Javorovac, Milanovac (20)
Myrmica rugulosa Nylander, 1849 Localities in CRO: Krapina (11)
Myrmica sabuleti Meinert, 1861
Localities in CRO: Uoka (7); Lopaoa (4 - indicated as M. scabrinodis); Bakar (9); near
Oabar (20); Leska (18); Velika Kapela (19); i. Cres (9); i. Cres (Vodice) (20); Kosore,
Poljica (7); Makarska (8); Gornja & Donja Vruoica, Kuna (20); Dubrovnik, Grua, Lapad
(10); Sljeme (18); Javorovac, Hlebine, near Naoice, near Donji Miholjac, near Slavonski
Brod (20)
Note: See comment under M. lonae.
Myrmica salina Ruzsky, 1905
Localities in CRO: Struaec, near Naoice (20)
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Myrmica scabrinodis Nylander, 1846
Localities in CRO: Velika Kapela (19); i. Cres (Vodice) (20); Krapina (11); Struaec,
Narta, Javorovac (20)
Myrmica schencki Viereck, 1903 Localities in CRO: Leska (18)
Myrmica specioides Bondroit, 1918
Localities in CRO: Momjan (13); Rijeka (15); Zemunik (7)
Myrmica sulcinodis Nylander, 1846
Localities in CRO: ? Senj (11); Velebit (7); ? i. Hvar, Sljeme, ? Otrigova (11)
Note: Localities Senj, i. Hvar and Otrigova (Vogrin 1955) are doubtful. M. sulcinodis can
be found in subalpine zone in Southern Europe and above 800 m in Central Europe
(Seifert 1988), what contradicts the records from mentioned localities.
Myrmoxenus corsicus (Emery, 1915)
Localities in CRO: i. Krk (near Baoka), Ljubao, Jasenice, near Biograd (Buschinger &
Winter 1985)
Myrmoxenus gordiagini Ruzsky, 1902
Localities in CRO: Momjan (13); near Rovinj, i. Krk (near Baoka), i. Rab (Suha Punta)
(Buschinger et al. 1983)
Myrmoxenus kraussei (Emery, 1915)
Localities in CRO: i. Krk, i. Pag, Paklenica (Buschinger et al. 1986)
Myrmoxenus ravouxi (André, 1896)
Localities in CRO: Dalmacija (Buschinger 1997)
Pheidole pallidula (Nylander, 1849)
Localities in CRO: Savudrija (20); Umag, Tar, Rovinj, Piaan (7); Rijeka (13); Orehovica (4); Prezid, Crikvenica, Novi Vinodolski, Senj (11); i. Krk (16); Bakar, i. Krk (Krk, Jurandvor, near Sv. Roko, Ponikva), i. Rab (Rab, Sv. Ilija, Kamenjak, near Lopar, Rt Zidine, Rt Stojan), i. Goli, i. Dolin, i. Cres (9); i. Cres (Vrana) (20); Velika Kapela (19); near Zadar (1); i. Dugi otok, i. Tukoooak (12); i. Dugi otok (Zaglav), i. Vir (Torovi) (20); Bjelina, Suoci (17); Skradin (7); Lioane Ostrovioke, Ovrljevo, Gornje Vinovo (17); Split (2) (19); near Split (8); Kaotela, Klis, Konjsko, Gata (7); Dugopolje, Marjan, between Klis and Solin, Podgora (10); i. Hvar (2) (7); i. Brao (Milna) (17) (Petrov & Collingwood 1992 - misidentified as P. megacephala); i. Brao (Zlatni rat), i. Vis, i. Jabuka (7); i. Vis (Brguljac) (18); i. Vis (Komiaa), i. Kamenjak, i. Kamik, i. Bioevo, i. Svetac (10); i. Koroula (near Raoioae) (20); i. Koroula (Lumbarda, Kuoiote), i. Suoac, i. Kopiote (7); i. Bijelac, i. Mroara, i. Veli & Mali Lastovljan (10); i. Mljet (7) (10); Gornja & Donja Vruoica (20); near Dubrovnik (1) (8); Grua (6) (10); i. Olipa, Slano, Dubrovnik, Lapad, i. Lopud, Trsteno, Rijeka Dubrovaoka, Srd, Oibaoa, i. Lokrum (10)
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Pyramica argiola (Emery, 1869) Localities in CRO: Zemunik (7)
Pyramica baudueri (Emery, 1875)
Localities in CRO: Zemunik, Kistanje, Kaotela (7); ? Krapina (11)
Note: since this is a typical Mediterranean species, the record from Krapina (Vogrin
1955) is very doubtful.
Solenopsis fugax (Latreille, 1798)
Localities in CRO: Savudrija, Mune - Aejane, Momjan, i. Tovarjea, Veruda (7); Pula (20); Draga, Orehovica (4); Trsat (15); Senj (11); Velika Kapela (19); i. Krk (Krk, near Sv. Roko), i. Plavnik, i. Rab (Rab, Banjol, Rt Zidine), i. Cres (9); Zemunik, Zadar, Nin, i. Rava (7); i. Vir (Vir) (20); Rogoznica (7); Lioane Ostrovioke, Split, Gornje Vinovo, Marjan (17); Kaotela, i. Hvar (Vrbanj, Jelsa), i. Jabuka (7); i. Vis (Vis), i. Mali Paraan, i. Bijelac, Rijeka Dubrovaoka, Lapad (10); Pregrada (4); Krapina, Otrigova (11); Javorovac, Hlebine, near Donji Miholjac, near Naoice, near Slavonski Brod (20)
Solenopsis wolfi Emery, 1915
Localities in CRO: near flevrske, Kaotela, i. Hvar (Privorac cave) (7)
Stenamma debile (Förster, 1850)
Localities in CRO: Rakalj, Mune, Oijana, Uoka, Velebit, Paklenica (7); Medvednica (18);
Krapina (11)
Note: Until the revision by DuBois (1993), this species had been treated as S. westwoodii
Westwood, 1839. The latter is restricted to Western Europe while S. debile has wider
range.
Stenamma striatulum Emery, 1895
Localities in CRO: i. Tovarjea, Oijana (7); i. Krk (near Ponikva, between Krk and
Malinska), i. Cres (9)
Strongylognathus testaceus (Schenck, 1852) Localities in CRO: Krapina (11)
Temnothorax affinis (Mayr, 1855)
Localities in CRO: Lupoglav (7); Rijeka (13); i. Krk, near Dubrovnik (14); Zagreb (15);
near Naoice, near Slavonski Brod (20)
Temnothorax algiricus (Forel, 1894) Localities in CRO: i. Vir (Torovi) (20)
Temnothorax angustulus (Nylander, 1856)
Localities in CRO: Rijeka Dubrovaoka, Dubrovnik (10); Rijeka (13)
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Temnothorax clypeatus (Mayr, 1853) Localities in CRO: Savudrija (13)
Temnothorax crassispinus (Karavaiev, 1926)
Localities in CRO: Mune (7); near Rijeka (4); Orehovica, Plitvice (15); i. Cres (9); i. Hvar
(Brusje) (2); near Dubrovnik (1); Podsused (15); Sljeme (18); Krapina (11); Hlebine,
Javorovac, Virovitica, Kolac (20)
Note:  Seifert (1995,  1996)  showed that Leptothorax (=  Temnothorax) nylanderi
(Foerster, 1850) consists of two different parapatric species: L. nylanderi, which is
distributed in Western Europe, and L. slavonicus, later defined as junior synonym of L.
crassispinus (Radchenko 2000), which is present in Eastern part of Europe. The general
distribution pattern of the two sibling species, the data from neighbouring countries, and
the fact that all examined material belongs to Temnothorax crassispinus, indicate that T.
nylanderi is most probably absent from the territory of Croatia.
Temnothorax exilis (Emery, 1869)
Localities in CRO: Velika Kapela (19); i. Krk (14); i. Krk (near Baoka) (Buschinger & Winter 1985); i. Palacol, i. Trasorka (7); i. Rab (between Rab and Sv. Ilija) (9); Ljubao (14) (Buschinger & Winter 1985); Jasenice, near Biograd (Buschinger & Winter 1985); Podgora, i. Vis, i. Veli Barjak, i. Greben, i. Mali Paraan, i. Kamik, i. Svetac, i. Bioevo (10); i. Jabuka (7); i. Bisaoe, i. Kopiote, Dubrovnik (10)
Temnothorax flavicornis (Emery, 1870)
Localities in CRO: i. Krk (14) (16); Ljubao (14); i. Hvar (Jelsa) (7); i. Vis (Brguljac) (18);
i. Koroula (near Koroula) (20); near Dubrovnik (14)
Temnothorax graecus (Forel, 1911) Localities in CRO: i. Jabuka (7)
Temnothorax interruptus (Schenck, 1852)
Localities in CRO: Limski kanal (20); i. Rab (Rab) (9); i. Vir (Vir) (20); Slano, Rijeka
Dubrovaoka (7); Lapad, near Grua (10)
Temnothorax italicus (Consani, 1952) Localities in CRO: near Poreo (20)
Temnothorax lichtensteini (Bondroit, 1918)
Localities in CRO: Savudrija, i. Tovarjea, Mooaeniaka Draga (7); Limski kanal (20); near Rovinj, i. Krk (near Baoka) (Buschinger et al. 1983); i. Krk (14); i. Rab (Rab, Banjol) (9); Ljubao (14); Kistanje, Kaotela (7); Dubrovnik (10)
Temnothorax nigriceps (Mayr, 1855)
Localities in CRO: Velika Kapela (19); Biokovo (10)
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Temnothorax parvulus (Schenck, 1852)
Localities in CRO: Cicilinovo, Veaica, Bakar (4 - indicated as Leptothorax tuberum); Sukooan, Ovrljevo (17); i. Koroula (near Koroula), Gornja Vruoica (20); Lapad (10); Krapina (11); near Slavonski Brod (20)
Temnothorax pelagosanus (Müller, 1923) Localities in CRO: i. Palagruaa (7)
Temnothorax recedens (Nylander, 1856)
Localities in CRO: i. Rab (Kamenjak) (7); i. Rab (Rab) (9); i. Krk, i. Pag, Paklenica (Buschinger et al. 1986); i. Krk (near Baoka), i. Pag (Kolan), Ljubao (14); i. Vir (Torovi), i. Hvar (near Hvar) (20); Split (2); i. Jabuka (7); Lapad (10)
Temnothorax rottenbergii (Emery, 1870)
Localities in CRO: Pula (13); i. Cres (Vrana) (20); i. Rab (Dundo) (9); Kozjak (7); Lapad
(8)
Temnothorax sordidulus (Müller, 1923) Localities in CRO: Grua (8)
Temnothorax tuberum (Fabricius, 1775)
Localities in CRO: Uoka (7); Trsat (11); Rijeka (13); Rijeka Dubrovaoka (6); Podsused,
Krapina (11)
Temnothorax unifasciatus (Latreille, 1798)
Localities in CRO: Tar, Rovinj (7); Momjan (13); Cicilinovo, Trsat (4); i. Krk (near
Baoka) (14); i. Looinj, Vrana (7); i. Vir (Vir) (20); Marjan, Dubrovnik (10)
Tetramorium biskrense Forel, 1904
Localities in CRO: i. Bijelac, near Grua, Lapad (10)
Tetramorium caespitum (Linnaeus, 1758)
Localities in CRO: Umag, Mune - Aejane (7); Pula (11) (16) (20); Orehovica, Veaica (4); Bakar, Fuaine (11); Leska (18); near Oabar (20); Velika Kapela (19); Senj, Plitvice (15); i. Krk (16); i. Krk (Krk, between Malinska and Skrboiai, Jurandvor), i. Rab (Kamenjak, Lopar) (9); i. Rab, i. Cres, i. Unije, i. Susak, i. Looinj, Velebit, Mali Halan, Zemunik (7); Zadar (2) (7); i. Ugljan, Benkovac (7); Tisno (17); Kaotel Suaurac, Solin, Mosor (7); Split (2); Trogir, Biokovo (10); Ovrljevo, Seoca (17); i. Hvar (Hvar) (20); i. Palagruaa (5) (11); Orebia (10); Dubrovnik (2); Zagreb, Podsused, Krapina (11); Struaec, Narta, Hlebine, Javorovac, Virovitica, Kolac, near Donji Miholjac, near Naoice, near Slavonski Brod (20)
Tetramorium diomedeum Emery, 1908
Localities in CRO: Marjan (17); i. Bioevo, ? Lapad, ? Rijeka Dubrovaoka (10)
Note: Zimmermann (1934) mentioned that the material from Lapad and Rijeka
Dubrovaoka showed some differences from typical T. diomedeum.
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Tetramorium ferox Ruzsky, 1903 Localities in CRO: Lapad (8)
Tetramorium impurum (Förster, 1850) Localities in CRO: Velika Kapela (19)
Tetramorium moravicum Kratochvil, 1941 Localities in CRO: Podaopilje (17)
Tetramorium semilaeve Andre, 1883
Localities in CRO: Pula (7) (20); Oijana (7); i. Krk (16); i. Kormati, i. Rab (Sv. Ilija, near Lopar), i. Dolin (9); i. Cres, i. Galijola, i. Okoljia, i. Kozjak, i. Sv. Petar (7); Modria (17); Bokanjac, Diklo, i. Svilan (7); i. Dugi otok (12); Kaotela (7); Marjan (10) (17); Trogir, Podgora, i. Brao (Puoioaa), i. Hvar (10); i. Hvar (Brusje) (2); i. Vis (Komiaa), i. Bioevo, i. Brusnik (7); i. Vis (Brguljac, Milna) (18); i. Kamenjak, i. Greben, i. Mali Paraan, i. Kamik (10); i. Suoac (7); i. Koroula (near Koroula, near Vela Luka) (20); i. Koroula, i. Mljet (Sobra) (10); i. Mljet (7); Dubrovnik (7) (10); Grua, Lapad, Rijeka Dubrovaoka, near Trsteno, near Oibaoa, i. Lokrum (10)
Tetramorium cf. splendens Ruzsky, 1902
Localities in CRO: Marjan, between Klis and Solin, Podgora, i. Sv. Klement, i. Vis, i.
Brusnik, i. Jabuka, i. Mroara, Lapad (10)
Note: The status of Tetramorium splendens is quite problematic. It was described from
Caucasus by Ruzsky, based on a single worker, but the holotype is lost. It is an enigmatic
taxon and is better to consider it as incertae sedis (Radchenko pers. comm.). It could be
any other Tetramorium species with dilute  sculpture.  Even Zimmermann (1934)
mentioned that some samples were close to T. semilaeve and some to T. diomedeum. It is
also possible that some of the records refer to T. hungaricum Röszler, 1935, a species
more widely distributed as thought before (Csosz & Radchenko pers. comm.).
Subfamily DOLICHODERINAE Forel, 1878
Bothriomyrmex adriacus Santschi, 1922
Localities in CRO: Sv. Stjepan - Livade, Rovinj (7); Momjan, Rijeka (13); i. Rab (near Sv. Ilija, Kamenjak), i. Cres (9); Modria (17); i. Pag, near Zadar, Bokanjac, Bibinje (7); i. Dugi otok (Sali) (12); Lioane Ostrovioke (17); i. Svilan, between Klis and Solin (7); Ovrljevo (17); Solin, Marjan, i. Sv. Klement (10); i. Hvar (10) (15); i. Vis, i. Mljet (7); i. Koroula, i. Otok, Sv. Ilija, i. Oipan, i. Olipa, Rijeka Dubrovaoka, i. Daksa (10); Lapad, Srebreno, Cavtat (8)
Note: B. adriacus is a sibling species of B. meridionalis (Roger, 1836). Müller (1923) and Soudek (1925) regard all their material as B. meridionalis, while Zimmermann (1934) says that most of the material shows characteristics typical for B. adriacus and eventually lists all the records, including those of Müller and Soudek, under B. adriacus. I therefore regard all the records of those three authors as B. adriacus, beeing aware that some could actually be B. meridionalis.
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Bothriomyrmex meridionalis (Roger, 1863)
Localities in CRO: Orehovica (4); i. Krk (16); i. Vir (Vir) (20); i. Brao (Milna) (Petrov
1993 - misidentified as B. syrius)
Note: See comment under B. adriacus.
Dolichoderus quadripunctatus (Linnaeus, 1771)
Localities in CRO: Orehovica (4); Rijeka (15); Zagreb, Krapina (11); Struaec, Slatina,
near Donji Miholjac, near Slavonski Brod (20)
Liometopum microcephalum (Panzer, 1798)
Localities in CRO: Uoka (7); Rijeka (4) (13); Gorski kotar (11); near Skradin, near Split
(1); Kaotela, Solin, Stobreo, Gata (7); Dugopolje (10); Oibaoa (7) (10); Zagreb (11)
Tapinoma ambiguum Emery, 1925
Localities in CRO: Leska (18); Split (17); Javorovac (20)
Note: Of two closely related species, T. ambiguum and T. erraticum (Latreille, 1798), the
first one was generally ignored by most authors. At first only males were known to show
distinct differences between the two species, but Seifert (1984) demonstrated a reliable
method to separate female castes as well. All the literature studied here mention only T.
erraticum, so that T. ambiguum is certainly more common than shown by these results.
Tapinoma erraticum (Latreille, 1798)
Localities in CRO: Mune, Piaan (7); Pula (7) (11); near Poreo, near Oabar (20); Rijeka (1) (7); near Draga (1); Orehovica, Trsat, Veaica (4); Prezid, Bakar, Senj (11); Velika Kapela (19); i. Krk (16); i. Krk (Krk, near Sv. Roko), i. Rab (Rab, near Lopar, near Banjol), i. Cres (9); i. Galijola, i. Looinj (Osorooica), i. Pag (7); i. Pag (Saline) (15); Pridraga, Koaino, Diklo (7); near Zadar (1); Privlaka, i. Vir (Torovi) (20); i. Dugi otok (Dugo polje) (12); Lioane Ostrovioke, Ovrljevo, Arnovnica (17); Split (2) (17) (19); Kaotela, Kozjak (7); between Klis and Solin, Podgora (10); Podaopilje (17); i. Hvar (Brusje) (2); i. Hvar (Vrboska) (7); i. Hvar (Hvar), i. Sv. Klement (10); i. Bioevo (7) (10); i. Koroula (7); i. Suoac, Orebia, i. Oipan, Rijeka Dubrovaoka (10); Grua (7); Konavle (10); Zagreb, Krapina (11); Slatina, near Naoice, near Slavonski Brod (20); Pleternica, Valpovo - Jugovaoa (11) Note: See comment under T. ambiguum.
Tapinoma nigerrimum (Nylander, 1856)
Localities in CRO: i. Rab (near Lopar) (9); Paklenica, Promina, Bokanjac (7); Marjan,
between Klis and Solin, i. Oipan, i. Lopud, Dubrovnik (10); Lapad (7) (10); Oibaoa (7)
Subfamily FORMICINAE Latreille, 1809
Camponotus aethiops (Latreille, 1798)
Localities in CRO: Savudrija, Pula (20); Mune - Aejane, Rovinj, Mooaeniaka Draga, Opatija (7); Rijeka (13); Orehovica (4); Draga, Bakar, Crikvenica, Senj (11); i. Krk (16); i. Krk (Jurandvor) (15); i. Krk (Krk, between Krk and Malinska ), i. Rab (Rab, near Sv.
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Ilija, Dundo, Lopar, Rt Stojan, Rt Zidine) (9); i. Rab, i. Cres (Vrana) (20); i. Cres (7) (9); i. Unije, i. Susak, i. Male Srakane, Podprag (7); Zadar (7) (11); Sukooan (17); i. Piokera (20); i. Dugi otok (Sali) (7) (12); Promina (7); Oibenik (19); Tisno, Lioane Ostrovioke, Vinioae (17); Labin, Split (2); Kaotel Stari, Klis (7); near Klis, Marjan, near Solin, Podgora, Biokovo, i. Brao (Puoioae) (10); i. Brao (Milna) (17); i. Hvar (10) (15); i. Hvar (Suauraj) (2); i. Hvar (Brusje) (2) (3); i. Hvar (Hvar) (20); i. Vis (7); i. Vis (Komiaa, Hum) (5); i. Vis (Brguljac, Podstraaje) (18); i. Kamenjak, i. Svetac (10); i. Bioevo (7) (10); i. Bisaoe, i. Otok (10); i. Koroula (15); i. Koroula (near Koroula, near Vela Luka) (20); i. Koroula (Raoioae), i. Kopiote, i. Suoac (7); i. Lastovo (1); i. Palagruaa (5) (11); Donja Vruoica (20); i. Mljet (7); i. Mljet (Sobra), i. Oipan, Trsteno, Rijeka Dubrovaoka, Oibaoa, i. Lokrum, i. Daksa (10); Dubrovnik (8) (10); Grua (7) (10); Lapad (6) (10); Zagreb (15); Krapina (11); near Slavonski Brod (20); Osijek (19) Note: Sometimes this species is defined as C. sylvaticus (Olivier, 1792) in older literature.
Camponotus dalmaticus (Nylander, 1849)
Localities in CRO: Rovinj, Krapan (7); near Flengi (20); Orehovica, Veaica, Cicilinovo (4); Rijeka, Suoak, Draga, Senj (11); i. Krk (7); i. Rab (9); i. Looinj (Osorooica) (7); i. Dugi otok (12); Oibenik (19); Gornje Ogorje, Vinioae, Trnbusi (17); i. Brao (Puoioaa) (10); i. Hvar (near Hvar) (20); i. Vis (Milna) (18); i. Bioevo, i. Koroula (Pupnat, Arnovo) (7); i. Koroula (near Vela Luka) (20); Grua (7) (10); Dubrovnik, Rijeka Dubrovaoka, Lapad (10)
Camponotus fallax (Nylander, 1856)
Localities in CRO: Poreo (20); Orehovica (15); Kuna (20); Srebreno (8); Krapina,
Otrigova (11)
Camponotus herculeanus (Linnaeus, 1758)
Localities in CRO: Rijeoina, Skrad (11); Fuaine, Mrzla vodica (4); Kuaelj (15); near
Oabar (20); ? Zagreb, Sljeme, ? Krapina, Ivanooica (11)
Note: This is a boreo-montane species which is very rarely found below 300 m altitude
(Seifert 1989, 1996), so that the records from Zagreb and Krapina (Vogrin 1955) are
questionable.
Camponotus lateralis (Olivier, 1792)
Localities in CRO: Umag, Pula, Oijana, Lovran (7); Rijeka (13); Orehovica (4); Suoak, Veaica, Senj (15); Bakar, Novi Vinodolski (11); Fuaine (4); i. Krk (Jurandvor) (11); i. Rab (7); i. Rab (Rab, Lopar), i. Cres (9); i. Pag (Lun) (15); i. Looinj, Bokanjac (7); Zadar (1) (11); i. Dugi otok (Sali) (12); i. Dugi otok (Zaglav) (20); Krka (11); Split (2) (7); Kaotel Suaurac (7); Dugopolje (10); Marjan (10) (17); Makarska (15); Podgora (10); i. Brao (Kneza Ravan, Nereaioaa, Bol, Supetar) (7); i. Brao (Milna) (17); i. Hvar (near Hvar) (20); i. Hvar (Brusje, Suauraj) (2); i. Vis (Komiaa) (7); i. Vis (Milna) (18); i. Bioevo (10); i. Svetac, i. Suoac, i. Koroula (Pupnat, Arnovo) (7); i. Koroula (Koroula) (10) (20); i. Koroula (near Raoioae, near Vela Luka) (20); i. Lastovo (1); i. Veli Lastovljan (7);
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near Dubrovnik (1); i. Oipan, near Trsteno, Rijeka Dubrovaoka, i. Lokrum, i. Daksa (10); Grua (7) (10)
Camponotus ligniperda (Latreille, 1802)
Localities in CRO: Uoka (7); Orehovica, Draga (4); Rijeoina (15); Senj (11); Fuaine, Mrzla vodica (4); near Oabar (20); Ravna Gora, Lokve, Lioko polje (11); Plitvice, near Veliki Zaviaan (15); Mali Halan, Paklenica (7); Dalmacija (6); Oibenik, Dubrovnik (19); Stojdraga, Samobor, Zagreb, Kraljioin Zdenac, Sljeme (11); Medvednica (18); Krapina, Otrigova (11)
Camponotus piceus (Leach, 1825)
Localities in CRO: Savudrija (20); Umag, Kroan, Pula, Oijana, Lovran (7); Draga, Orehovica, Veaica, Senj, i. Krk (Jurandvor) (15); i. Krk (16); i. Rab (Lopar) (9) (20); i. Rab (Rab, Dundo, near Sv. Ilija, Rt Zidine), i. Dolin, i. Cres (9); i. Cres (Vodice) (20); i. Oruda, i. Looinj, i. Kozjak, i. Orjule, i. Trasorka, i. Unije (7); Paklenica (15); Zadar (7); i. Dugi otok (12); i. Dugi otok (Zaglav), i. Vir (Torovi) (20); between Klis and Solin (7); Klis, Kotlenice, Marjan (10); Seoca (17); Podgora, Biokovo, i. Mali Paraan, i. Bioevo, i. Svetac (10); i. Koroula (near Koroula) (20); Slano (7); Lapad, Grua (10); near Slavonski Brod (20)
Camponotus truncatus (Spinola, 1808)
Localities in CRO: Livade (7); near Poreo (20); Orehovica, Cicilinovo (4); Senj (11); i. Dugi otok (Zaglav) (20); Knin (3); Dubrovnik, Grua (10); Zagreb (15); Krapina (11); near Slavonski Brod (20)
Camponotus vagus (Scopoli, 1763)
Localities in CRO: Umag, Lovran - Mooaeniaka Draga (7); Novigrad (11); Voziliai (15); near Poreo (20); Rijeka (13); Orehovica (4); Ponikve, Bakar, Novi Vinodolski, Lokve (11); Velika Kapela (19); i. Krk (16); i. Krk (Lakmartin, Njivice) (11); i. Cres (9); i. Rab, i. Cres (Martinoaica) (11); Plitvice (15); Paklenica, Bibinje (7); Split (1); Ilica Planina (11); Dubrovnik (7); Karlovac, Zagreb, Graoani, Sljeme, Pljeoevica, Krapina, Otrigova, Naoice, Kroenik (11); Slatina (20)
Cataglyphis aenescens (Nylander, 1849)
Localities in CRO: Diklo (7); Lioane Ostrovioke (17); Leaevica (7); between Klis and
Solin (7) (10); i. Brao (Bol) (10)
Cataglyphis nodus (Brullé, 1832)
Localities in CRO: Dalmacija (1); i. Hvar (Brusje) (2); Metkovia (8)
Formica cinerea Mayr, 1853
Localities in CRO: Fuaine (11); i. Krk (Krk, near Malinska) (9); Zagreb (11); Pregrada
(4); Kriaovec (15)
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Formica cunicularia Latreille, 1798
Localities in CRO: Savudrija (20); Umag, Mune, Lovran, Opatija (7); Rijeka (13); Velika Kapela (19); i. Krk (Krk), i. Rab (Rab, Sv. Ilija, Lopar, Rt Stojan) (9); i. Rab, i. Unije (7); i. Cres (Vodice), i. Dugi otok (Zaglav) (20); Buljma (15); Vrana (7); Oibenik (19); Gornje Ogorje (17); Mosor (7); i. Vis (Hum) (5 - indicated as F. cinerea); Trnovec (15); Kalnik (11); Hlebine, Javorovac, Kolac, near Donji Miholjac, near Naoice (20); Poaega (15)
Formica foreli Emery, 1909 Localities in CRO: Mune (7)
Formica fusca Linnaeus, 1758
Localities in CRO: Mune (7); Lokve, Fuaine, Klek (11); near Oabar (20); Velika Kapela (19); i. Cres (9); Velebit (11); Plitvice (15); Sv. Ilija (10); Zagreb, Sljeme (11); Medvednica (18); Narta, Virovitica (20); Poaega, Naoice (11)
Formica gagates Latreille, 1798
Localities in CRO: Orehovica (4); Graoac (11); Oibenik (19); Ovrljevo (17); Labin (2); i. Hvar (10) (11); i. Koroula (11); i. Koroula (Koroula), Oskoruono, Kuna (20); Oibaoa, Konavle (10); Sljeme (18); Krapina (11); Virovitica (20)
Formica lemani Bondroit, 1917
Localities in CRO: Uoka (15); Velika Kapela (19); Buljma (15); Dinara, Biokovo (7)
Formica lugubris Zetterstedt, 1838 Localities in CRO: Sedalce (15)
Formica lusatica Seifert, 1997 Localities in CRO: Mooilo (15)
Formica polyctena Förster, 1850
Localities in CRO: Uoka (7); Fuaine, Mrzla vodica, Plitvice (15); Medvednica (18)
Formica pratensis Retzius, 1783
Localities in CRO: Mune, Uoka (7); Kuaelj, Lokve (15); Rijeoina spring (4); Senj (11); Velika Kapela (19); Plitvice (15); Strmica (7); Split (19); Gera, Sooice, Sljeme, Krapina (11); Javorovac, near Naoice (20)
Formica rufa Linnaeus, 1761
Localities in CRO: Mune (7); Uoka (15); Rijeka, Rijeoina (11); Fuaine (4); Plitvice, Buljma (15); Paklenica (7); Oibenik (19); Dalmacija (6); Zagreb (15); Krapina, Otrigova, Kalnik (11)
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Formica rufibarbis Fabricius, 1793
Localities in CRO: Umag (7); Pula (20); near Rijeka, Veaica, Fuaine (4); Lokve, Novi Vinodolski (11); Velika Kapela (19); near Latin, Plitvice (15); i. Cres (9); Prekriaje (15); Pregrada (4); Krapina, Kalnik (11); Javorovac, Narta, Struaec (20); Opioia Bukovica (11)
Formica sanguinea Latreille, 1798
Localities in CRO: Mune, Uoka (7); near Oabar (20); Gorski kotar, Fuaine (4); Leska (18); Velika Kapela (19); Graoac (11); Dalmacija (6); Sljeme, Krapina (11); near Domaoinec (15); Struaec, near Naoice (20)
Formica truncorum Fabricius, 1804
Localities in CRO: near Rijeka, Gorski kotar (4); Sljeme (15); Krapina (11)
Lasius alienus (Förster, 1850)
Localities in CRO: Lupoglav, Pula, Uoka, Lovran (7); Rijeka, Senj, Delnice (11); i. Krk
(Krk, near Baoka), i. Rab (Rab, Sv. Ilija, Banjol, Dundo, Lopar, Rt Zidine, Rt. Stojan) (9);
i. Rab (Kamenjak), i. Unije (7); Gospia (11); Obrovac, Karin, Babindub, Zemunik (7);
near Zadar (1); i. Dugi otok (12); Benkovac, Kistanje, Kaotela (7); Marjan, Kotlenice,
Mosor, Podgora (10); i. Vis (7); Orebia, Slano, Grua, Lapad (10); Dubrovnik, Trsteno, i.
Lokrum, Kupari, Cavtat (8); Zagreb, Krapina, flurfievac (11)
Note: In the revision of the subgenus Lasius s. str. of Palaearctic, Seifert (1992) described
two new species which previously belonged to L. alienus, i.e. L. paralienus and L.
psammophilus. All the specimens from the L. alienus complex that were checked here
were either L. paralienus or L. psammophilus. It is very likely that many or even all of
the literature data for L. alienus refer to one of those two species. Although it was not
found among the checked material, the presence of L. alienus in Croatia is most probably
not questionable, since it is distributed also in the neighbouring countries.
Lasius bicornis (Förster, 1850)
Localities in CRO: Lokve, Kraljioin Zdenac, Otrigova, Bregi (11)
Lasius brunneus (Latreille, 1798)
Localities in CRO: near Oabar (20); near Rijeka (1); Veaica (4); Plitvice (15); near Split
(1); i. Hvar (Brusje) (2); Krapina, Kalnik (11); Hlebine, Javorovac, Kolac (20)
Lasius carniolicus Mayr, 1861 Localities in CRO: Mosor (7)
Lasius distinguendus (Emery, 1916)
Localities in CRO: Sv. Stjepan - Livade, Gradole, i. Brijuni (7); near Rijeka (1); Orehovica (4 - indicated as L. bicornis var. affinis); i. Krk (Krk), i. Rab (Rab, between Rab and Banjol, Lopar, Stanioae) (9); i. Cres (Martinoaica) (1); i. Unije (7); Peruoia (15); Zemunik, Kaotela, Solin, Lovrea (7); Podaopilje (17); Prekriaje, Pregrada, Konjooina (15); Javorovac, Narta (20)
Lasius emarginatus (Olivier, 1792)
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Localities in CRO: Savudrija, Umag, Mune, i. Tovarjea, Uoka, Opatija, Lovran (7); Rovinj (13); Limski kanal (20); Rijeka (7); Cicilinovo, Orehovica, Veaica, Fuaine (4); Novi Vinodolski (11) (17); Alan (11); Senj (15); Velika Kapela (19); i. Rab (Rab), i. Cres (9); i. Susak, Zadar (7); Sukooan (17); Privlaka, i. Piokera (20); i. Dugi otok (12); Kaotela, Split (7); Marjan, Klis, Podgora, Biokovo, i. Brao (Milna), i. Hvar, i. Vis, i. Bioevo, i. Svetac (10); i. Koroula (Samograd) (7); i. Suoac, i. Mroara, Orebia, Sv. Ilija, Metkovia, Dubrovnik, Grua, Lapad, Rijeka Dubrovaoka, Srd, Oibaoa, i. Lokrum, Konavle (10); Medvednica (18); Zagreb, Krapina, Galdovo (11); Otrigova (15); Javorovac, near Slavonski Brod (20); Jasenovaoa (11)
Lasius flavus (Fabricius, 1781)
Localities in CRO: Uoka (7); Rijeka (13); Draga, Orehovica (4); Zlobin, Delnice (11); Leska (18); Velika Kapela (19); i. Plavnik, i. Rab (Rab, near Sv. Ilija), i. Male Srakane (9); Plitvice (15); Mali Halan (7); Dalmacija (1) (6); near Zadar (2); Oibenik (19); between Klis and Solin (10); Zapreoia (15); Sljeme, Krapina, Kalnik (11); Hlebine, Javorovac (20)
Lasius fuliginosus (Latreille, 1798)
Localities in CRO: Pula, Oijana (7); Orehovica, Draga (4); Novi Vinodolski, Senj, Fuaine, Bjelolasica (11); Leska (18); i. Krk (Ponikva, between Krk and Malinska), i. Rab (Rab, Lopar) (9); i. Rab (11); Paklenica (7) (11); Split (10); Gata (7); Ozalj (11); Prekriaje, Zapreoia (15); Zagreb, Krapina (11); Pregrada (4); Otrigova, Bregi (11); Hlebine (20); Novigrad Podravski (15); Kutjevo (11); near Naoice, near Donji Miholjac, near Slavonski Brod (20)
Lasius lasioides (Emery, 1869) Localities in CRO: Sukooan (17)
Lasius mixtus (Nylander, 1846)
Localities in CRO: Zlobin (11); Velika Kapela, Split (19); Krapina (11)
Lasius myops Forel, 1894
Localities in CRO: Orehovica (15); i. Rab (Banjol, near Lopar) (9); i. Unije, i. Male Srakane, i. Pag (7); i. Pag (Lun) (15); Dubrovnik (8); Lapad, Rijeka Dubrovaoka (10); near Slavonski Brod (20)
Lasius niger (Linnaeus, 1758)
Localities in CRO: Mune (7); near Oabar (20); Leska (18); Velika Kapela (19);
Crikvenica (11); i. Male Srakane (7); near Latin, Plitvice (15); Dalmacija (1) (6);
Obrovac, Koljane, Solin, i. Brao (Milna) (7); i. Hvar (3); Struaec (20); Zagreb (15);
Sljeme, Krapina (11); Pregrada (4); Otrigova (11); Javorovac, Hlebine, Kolac, Narta, near
Donji Miholjac (20); Jasenovaoa (11); near Naoice (20)
Note: Seifert (1991) separated a new species, L. platythorax, from closely related L.
niger. Literature sources mention only L. niger, but L. platythorax was confirmed among
the checked material.
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Lasius nitidigaster Seifert, 1997
Localities in CRO: i. Krk (16); near Slavonski Brod (20)
Lasius paralienus Seifert, 1992
Localities in CRO: Savudrija, Pula (20); Orehovica, Bakar (4 - indicated as L. alienus);
i. Cres (Vodice), i. Rab (Lopar), i. Pag, i. Dugi otok (Zaglav) (20); Ovrljevo, Seoca,
Podaopilje (17); Donja Vruoica, Hlebine, Javorovac, near Donji Miholjac, near Slavonski
Brod (20)
Note: See comment under L. alienus.
Lasius platythorax Seifert, 1991
Localities in CRO: near Oabar (20); Leska (18); Plitvice, Podsused (15); Struaec, Narta,
Milanovac (20)
Note: See comment under L. niger.
Lasius psammophilus Seifert, 1992
Localities in CRO: near Oabar (20); Leska (18); i. Krk (16); Gornje Ogorje (17); Koroula
(near Raoioae) (20)
Note: See comment under L. alienus.
Lasius umbratus (Nylander, 1846)
Localities in CRO: Momjan, Poreo (13); Leska (18); Krapina (11)
Lasius viehmeyeri Emery, 1922
Localities in CRO: Knin, Dalmacija (Boer 2003)
Lepisiota frauenfeldi (Mayr, 1855)
Localities in CRO: Dalmacija (11); Zadar (15); Privlaka, i. Piokera (20); near Sinj (1); Split (2); Kaotela, Kaotel Suaurac, Klis (7); Marjan (8) (10); between Klis and Solin (10); i. Hvar (Brusje) (2); i. Hvar (near Hvar), i. Adrilca (20); i. Brao (Milna) (17) (Petrov & Collingwood 1992 - misidentified as L. splendens); i. Vis (Vis) (10); i. Brusnik, i. Palagruaa (5); Mljet (10); Dubrovnik (10) (19); Srd, Lapad, i. Daksa (10); Cavtat (8)
Lepisiota nigra (Dalla Torre, 1893)
Localities in CRO: Pula (16); Duga uvala (7); i. Rab (Rab, Lopar, Rt Zidine) (9); i. Oruda, i. Unije, i. Sv. Petar, Smiloia, i. Molat (7); i. Dugi otok (12); Oibenik (7); Veliki Drvenik, Trogir (10); Kaotel Suaurac, Solin (7); Split (7) (8); Marjan, Podgora (10); i. Brao (Milna) (7); i. Hvar (8) (10); i. Vis (7); i. Vis (Komiaa), i. Mali & Veli Barjak, i. Kamenjak, i. Kamik, i. Greben, i. Mali & Veli Paraan, i. Bioevo, i. Svetac, i. Brusnik, i. Suoac, i. Bisaoe, i. Koroula (Koroula) (10); i. Koroula (Pupnat), i. Crnac (7); i. Lastovo (near Skrivena Luka), i. Kopiote, i. Podkopiote, i. Mroara, i. Mali & Veli Lastovljan, i. Palagruaa, i. Mljet (Sobra), Orebia (10); Ston (7); i. Olipa (10); Dubrovnik (7) (8) (10) (19); Trsteno (8); Grua, Lapad, i. Lokrum, i. Lopud (10)
Paratrechina vividula (Nylander, 1846) Localities in CRO: Zagreb (11)
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Plagiolepis pygmaea (Latreille, 1798)
Localities in CRO: Savudrija (20); Pula (7) (20); Rovinj, Lovran (7); near Rijeka (1); Orehovica, Trsat, Bakar (4); Crikvenica (11); i. Krk (16); i. Krk (Krk, near Sv. Roko), i. Kormati, i. Rab (Rab, near Sv. Ilija, Banjol, Dundo, Lopar, near Stanioae, Visilica), i. Dolin, i. Cres (9); i. Cres (near Martinoaica) (1); i. Cres (Vodice) (20); i. Looinj, i. Sv. Petar, i. Susak (7); Peruoia (11); Modria (17); near Zadar (1); Bibinje (7); Sukooan (17); i. Vir (Torovi) (20); i. Dugi otok (Dugo polje) (12); i. Dugi otok (Sali), Knin (7); Vrpolje, Lioane Ostrovioke, Ovrljevo, Arnovnica (17); Kaotela, Klis, Split (7); Trogir, between Klis and Solin, Podgora (10); Marjan (10) (17); Podaopilje (17); i. Adrilca (20); i. Hvar (3) (10); i. Hvar (Vrbanj, Jelsa, Vrboska) (7); i. Brao (Bol) (10); i. Brao (Milna) (17); i. Vis (7); i. Vis (Brguljac) (18); i. Vis (Komiaa), i. Mali & Veli Barjak, i. Greben, i. Mali Paraan, i. Svetac, i. Jabuka, i. Bioevo (10); i. Koroula (10) (11); i. Koroula (near Koroula) (20); i. Otok, i. Bisaoe, Orebiè, i. Kopiote, i. Bijelac, i. Mali Lastovljan, i. Mljet (Sobra) (10); near Dubrovnik (1); i. a ipan, Rijeka Dubrovaoka, Dubrovnik, near Grua, Lapad, i. Lokrum, Oibaoa (10); Krapina (11); near Slavonski Brod (20)
Plagiolepis vindobonensis Lomnicki, 1925
Localities in CRO: Velika Kapela (19); i. Vis (Milna, Podstraaje) (18)
Plagiolepis xene Stärcke, 1936 Localities in CRO: Podaopilje (17)
Polyergus rufescens (Latreille, 1798)
Localities in CRO: Kortivi (7); Rijeka (13); Zagreb, Krapina (11); Pregrada (4)
Prenolepis nitens (Mayr, 1853)
Localities in CRO: Rovinj (7) (16); Momjan, Lupoglav, Raoka, Opatija (7); Pula (16); i. Veliki Brijuni (20); Rijeka (13); Orehovica, Veaica, Bakar (4); Draga (15); i. Cres (9); i. Looinj (Veli Looinj), i. Sv. Petar, i. Ilovik (7); Dalmacija (2); i. Dugi otok (Sali) (12); Kaotel Lukoia, Kaotel Suaurac (7); Split, i. Hvar (3); Gornja Vruoica, Oskoruono (20); Dubrovnik, Grua, Lapad (10); Zagreb (11); Oestine (15); Sljeme (18); Krapina (11)
Results and discussion
For the territory of Croatia, 140 ant species are listed. The presence of Lasius alienus was not directly confirmed, but it most probably occurs here, while the situation with Tetramorium splendens is rather unclear (see comments for the two species in the list). Since a large part of Croatia lies in the Mediterranean region which has the richest ant fauna in Europe, the number of recorded ant species is expectedly higher than in neighbouring Central European countries. Thus, there are 118 species known for Slovenia (Braoko 2003) and 101 for Hungary (Gallé et al. 1998). If compared to nearby Mediterranean countries, for instance Italy with 226 species (Poldi et al. 1995), this number is much lower. In part this is certainly a consequence of lesser known ant fauna of Croatia, therefore many new species are expected with further investigations.
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Most species were found in the Mediterranean part of the country. The ant fauna of this region of Croatia has approximately half of the species that are otherwise widely distributed in Europe. The other half are those with more or less Mediterranean distribution, namely the species generally found in the Mediterranean as well as on thermophilous sites of Central Europe and the species mainly limited to the Mediterranean region. Among the latter, the representative species recorded in Croatia are Proceratium algiricum, most Aphaenogaster species, Messor wasmanni, Myrmoxenus corsicus, M. kraussei, Pyramica spp., Temnothorax algiricus, T. angustulus, T. exilis, T. graecus, T. italicus, T. rottenbergii, Tetramorium diomedeum, Tapinoma nigerrimum, Lasius lasioides, Lepisiota spp. The composition of the ant fauna of the Croatian interior resembles the fauna of Central European countries. Crematogaster ionia, C. schmidti, Messor capitatus, M. structor, found in the eastern Pannonian Croatia, are somewhat more exceptional. The mountain area of Gorski Kotar, Velika Kapela and Velebit represents a refuge for species otherwise more common in Northern Europe and mostly higher altitudes of Central Europe (i.e. Leptothorax acervorum, L. muscorum, Myrmica lobicornis, M. sulcinodis, Camponotus herculeanus, Formica lemani, F. lugubris). Two of the listed species, Monomorium pharaonis and Paratrechina vividula, are introduced to the native European ant fauna and are normally established in heated premises. M. pharaonis is a very common pest species, while P. vividula is rarer. The latter was reported to be found in a glasshouse in Zagreb (Vogrin 1955).
Ant sampling has not been evenly performed over the territory of Croatia. Most of the literature data and unpublished collected material originate from the Mediterranean part of the country. Although the ant fauna of this region is best known, many new species are still expected from there, as the Mediterranean territory is very rich in ant species in general. Besides, some genera that are very frequently represented there, i.e. Messor, Aphaenogaster, Temnothorax, Tetramorium, need to be revised what could contribute to additional species. Other parts of Croatia are much less investigated, therefore the picture of ant fauna of these regions will also change with future sampling.
In this study, 117 ant species were obtained from published sources and 100 from unpublished material, where 23 species were previously unknown for Croatia. This suggests that further field work could be very productive. The present review can also be a stimulus for other myrmecologists to supplement the list, for it is supposed that there is still much ant material from Croatia dispersed in various institutional and personal collections.
Acknowledgements
I wish to thank Al Vrezec (National Institute of Biology, Ljubljana) for the collected ants. Franjo Perovia (Croatian Natural History Museum, Zagreb) was very generous to give me access to the ant collection of the Museum. Many thanks to Tonai Rafia (Split), Paula Durbeoia and Lucija Oeria Jelaska (both from the Department of Zoology, Faculty of Science, Zagreb) for letting me to examine their collected ant material. I would like to thank Cedric Collingwood (Skipton, UK) who kindly sent me his list of Croatian ants. I am very grateful to Andreas Schulz (Leverkusen) for help in the identification of
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certain Aphaenogaster and Temnothorax specimens. I also kindly thank Alexander Radchenko (Institute of Zoology of the Ukrainian National Academy of Science, Kiev), Sándor Csosz (Hungarian Natural History Museum, Budapest), Ivan Petrov (Faculty of Biology, Institute of Zoology, Belgrade), and Xavier Espadaler (Universitat Autonoma de Barcelona) for useful information.
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Received / Prejeto: 27. 9. 2006
Appendix
Position of less known localities (abbreviations: i. - island, n. - near): Alan - n. Senj, Arbanasi - n. Zadar, Babindub - n. Zadar, Baovice - Split, Bjelina - n. Benkovac, Bokanjac - n. Zadar, Bregi - n. Koprivnica, Bribir - n. Crikvenica, Buljma -S Velebit, Cicilinovo - n. Rijeka, Oibaoa - n. Dubrovnik, Ovrljevo - n. Split, Diklo - n. Zadar, Donja Vruoica - Peljeoac, Draga - n. Rijeka, Dretulja spring - n. Plaoki, Duga uvala - Istra, Dugopolje - n. Split, flevrske - n. Benkovac, Flengi - n. Poreo, Fuaine - n. Delnice, Galdovo - Sisak, Gata - n. Omio, Gera - Aumberaoka gora, Gornja Vruoica -Peljeoac, Gornje Ogorje - n. Sinj, Gornje Vinovo - n. Split, Graoani - Zagreb, Gradole -Istra, Grua - Dubrovnik, Hlebine - n. Koprivnica, i. Bijelac - n. i. Lastovo, i. Bisaoe - n. i. Koroula, i. Brusnik - n. i. Vis, i. Crnac - n. i. Lastovo, i. Daksa - n. Dubrovnik, i. Dolin
- n. i. Rab, i. Galijola - n. i. Cres, i. Goli - n. i. Rab, i. Greben - n. i. Premuda, i. Grujica
- n. i. Looinj, i. Ilovik - n. i. Looinj, i. Jabuka - n. i. Vis, i. Jakljan - n. i. Oipan, i. Kamik
- n. i. Vis, i. Kolooep - n. Dubrovnik, i. Kopiote - n. i. Lastovo, i. Kormati - n. i. Krk, i.
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Kozjak - n. i. Looinj, i. Lokrum - n. Dubrovnik, i. Lopud - n. Dubrovnik, i. Male Srakane
- n. i. Looinj, i. Mali Barjak - n. i. Vis, i. Mali Lastovljan - n. i. Lastovo, i. Mali Paraan
- n. i. Vis, i. Mali Plavnik - n. i. Krk, i. Mroara - n. i. Lastovo, i. Olipa - n. i. Oipan, i. Orjule - n. i. Looinj, i. Oruda - n. i. Cres, i. Otok - n. i. Koroula, i. Palacol - n. i. Looinj, i. Piokera - n. i. Kornat, i. Plavnik - n. i. Krk, i. Podkopiote - n. i. Lastovo, i. Rava - n. i. Dugi otok, i. Sv. Klement - n. i. Hvar, i. Sv. Petar - n. i. Looinj, i. Svetac - n. i. Vis, i. Svilan - n. Oibenik, i. Okoljia - n. i. Unije, i. Tovarjea - n. Vrsar, i. Trasorka - n. i. Looinj, i. Tukoooak - n. i. Dugi otok, i. Veli Barjak - n. i. Vis, i. Veli Lastovljan - n. i. Lastovo, i. Veli Paraan - n. i. Vis, i. Zabodarski - n. i. Looinj, i. Adrilca - n. i. Hvar, Ilica Planina
- n. Plooe, Jagnjedovac - n. Koprivnica, Jasenak - Velika Kapela, Jasenice - n. Obrovac, Jasenovaoa - n. Donji Miholjac, Javorovac - n. Koprivnica, Kalnik - n. Koprivnica, Kanfanar - Istra, Karin - n. Obrovac, Kistanje - n. Knin, Klek - n. Ogulin, Klis - n. Split, Kolac - n. Virovitica, Koljane - n. Knin, Konavoski Dvori - n. Dubrovnik, Konjsko - n. Split, Kortivi - Istra, Kosore - n. Knin, Kotlenice - n. Split, Kozjak - n. Split, Koaino -n. Zadar, Kraljioin Zdenac - Medvednica, Krapan - Istra, Kroenik - n. Donji Miholjac, Kriaovec - Mefiimurje, Krka - n. Oibenik, Krmoina - n. Zadar, Kuoiote - Peljeoac, Kuna
- Peljeoac, Kupari - n. Dubrovnik, Kutjevo - n. Slav. Poaega, Kuaelj - n. Brod na Kupi, Labin - n. Trogir, Lapad - Dubrovnik, Latin - n. Plaoki, Leaevica - n. Split, Leska -Gorski kotar, Lioane Ostrovioke - n. Oibenik, Livade - Istra, Ljubao - n. Zadar, Lopaoa -n. Rijeka, Lovran - n. Opatija, Lovrea - n. Makarska, Lupoglav - Istra, Mali Halan -Velebit, Marjan - n. Split, Markovac - Istra, Milanovac - n. Virovitica, Mooilo -Krbavsko polje, Modria - n. Obrovac, Momjan - Istra, Mooaeniaka Draga - n. Opatija, Mrzla vodica - Gorski kotar, Mune - Istra, Narta - n. Bjelovar, Orehovica - n. Rijeka, Oskoruono - Peljeoac, Peruoia - n. Gospia, Piaan - n. Pazin, Pljeoevica - Medvednica, Podaopilje - n. Omio, Podgora - n. Makarska, Podprag - Velebit, Podsused - n. Zagreb, Poljica - n. Trogir, Ponikve - n. Rijeka, Prekriaje - n. Zagreb, Pridraga - n. Benkovac, Privlaka - n. Zadar, Privorac cave - n. Brusje, Rakalj - Istra, Raoka - Istra, Ravna Gora
-  Gorski kotar, Rijeoina - n. Rijeka, Rogoznica - n. Oibenik, Rt Stojan - n. Lopar, Rt Zidine - n. Lopar, Samograd - n. Raoioae, Sedalce - n. Brod na Kupi, Seoca - n. Omio, Siveria - n. Drnio, Skrad - Gorski kotar, Skradin - n. Oibenik, Sljeme - Medvednica, Smiloia - n. Benkovac, Sooice - Aumberaoka gora, Srd - n. Dubrovnik, Srebreno - n. Dubrovnik, Stanioae - n. Lopar, Stobreo - n. Split, Stojdraga - Aumberaoka gora, Ston -Peljeoac, Strmica - n. Knin, Struaec - n. Sisak, Suoci - n. Sinj, Sv. Ilija - Peljeoac, Sv. Stjepan - Istra, Oestine - n. Zagreb, Oijana - n. Pula, Opioia Bukovica - n. Virovitica, Otrigova - n. Oakovec, Tar - Istra, Teradige - n. Pula, Tisno - n. Vodice, Trnbusi - n. Omio, Trnovec - n. Varaadin, Trpanj - Peljeoac, Trsat - n. Rijeka, Trsteno - n. Dubrovnik, Veliki Drvenik - n. Oibenik, Veliki Zaviaan - N Velebit, Veruda - n. Pula, Veaica - n. Rijeka, Vinioae - n. Trogir, Vodnjan - Istra, Volosko - n. Opatija, Voziliai -Istra, Vrana - n. Biograd, Vrpolje - n. Oibenik, Zemunik - n. Zadar, Zlobin - n. Kraljevica, Aegar - n. Obrovac, Aejane - Istra, Arnovnica - n. Split
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