FOLIA BIOLOGICA ET GEOLOGICA Ex: Razprave razreda za naravoslovne vede Dissertationes classis IV (Historia naturalis) 56/1 2015 SLOVENSKA AKADEMIJA ZNANOSTI IN UMETNOSTI ACADEMIA SCIENTIARUM ET ARTIUM SLOVENICA Razred za naravoslovne vede – Classis IV: Historia naturalis LJUBLJANA 2015 FOLIA BIOLOGICA ET GEOLOGICA ISSN 1855-7996 Uredniški odbor / Editorial Board Matjaž Gogala, Špela Goričan, Milan Herak (Hrvaška), Ivan Kreft, Ljudevit Ilijanič (Hrvaška), Mario Pleničar, Livio Poldini (Italija) in Branko Vreš Glavni in odgovorni urednik / Editor Mitja Zupančič Tehnični urednik / Technical Editor Janez Kikelj Oblikovanje / Design Milojka Žalik Huzjan Prelom / Layout Medija grafično oblikovanje Sprejeto na seji razreda za naravoslovne vede SAZU dne 2. oktobra 2014 in na seji predsedstva dne 11. novembra 2014. 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Noben del te izdaje ne sme biti reproduciran, shranjen ali prepisan v kateri koli obliki oz. na kateri koli način, bodisi elek­tronsko, mehansko, s fotokopiranjem, snemanjem ali kako drugače, brez predhodnega pisnega dovoljenja last­nikov avtorskih pravic. / All rights reserved. No part of this publication may be reproduced, stored in a retrieval system or transmit­ted, in any form or by any means, electronic, mechanical, photocopying, recording or otherwise, without the prior permission of the publisher. Naslovnica: Saxifraga cuneifolia. (Foto: I. Dakskobler) Cover photo: Saxifraga cuneifolia. (Photo: I. Dakskobler) Folia biologica et geologica · Volume / Letnik 56 · Number / Številka 1 · 2015 VSEBINA CONTENTS RAZPRAVE / ESSAYS Igor Dakskobler 5 Phytosociological analysis of montane beech forests on steep shady slopes on mixed geological bedrock in western Slovenia 5 Fitocenološka analiza montanskih bukovih gozdov na strmih osojnih pobočjih na mešani geološki podlagi v zahodni Sloveniji Igor Dakskobler, Branko Dolinar, Branko Zupan, Florijan Poljšak & Peter Strgar 105 Fitocenološka oznaka rastišč vrste Nigritella archiducis-joannis v Dolini Triglavskih jezer (Julijske Alpe, Slovenija) 105 Phytosociological characteristics of the sites of Nigritella archiducis-joannis in the Triglav Lakes Valley (The Julian Alps, Slovenia) Igor Paušič 115 Confirmation of the Austrian vanilla orchid, Nigritella austriaca (Teppner & E. Klein) P. Delforge (Orchidaceae) a new species in the Slovenian flora 115 Potrditev avstrijske murke, Nigritella austriaca (Teppner & E. Klein) P. Delforge (Orchidaceae), nove vrste v flori Slovenije Mitja Zupančič & Jože Skumavec 125 Flora and vegetation in Pokljuka Gorge (Julian Alps, NW Slovenia) 125 Flora in vegetacija Pokljuške soteske (Julijske Alpe, SZ Slovenija) Božidar Drovenik & Branko Vreš 151 Novosti v favni rodu Otiorhynchus s. lat. (Coleoptera: Curculionidea) v Sloveniji 151 Novelties in the fauna of Otiorhynchus s. lat. (Coleoptera: Curculionidea) in Slovenia IGOR DAKSKOBLER: PHYTOSOCIOLOGICAL ANALYSIS OF MONTANE BEECH FORESTS ON STEEP SHADY SLOPES IGOR DAKSKOBLER: PHYTOSOCIOLOGICAL ANALYSIS OF MONTANE BEECH FORESTS ON STEEP SHADY SLOPES 6 7 FOLIA BIOLOGICA ET GEOLOGICA 56/1, 5–104, LJUBLJANA 2015 FOLIA BIOLOGICA ET GEOLOGICA 56/1 – 2015 FOLIA BIOLOGICA ET GEOLOGICA 56/1 – 2015 PHYTOSOCIOLOGICAL ANALYSIS OF MONTANE BEECH FORESTS ON STEEP SHADY SLOPES ON MIXED GEOLOGICAL BEDROCK IN WESTERN SLOVENIA FITOCENOLOŠKA ANALIZA MONTANSKIH BUKOVIH GOZDOV NA STRMIH OSOJNIH POBOČJIH NA MEŠANI GEOLOŠKI PODLAGI V ZAHODNI SLOVENIJI Igor Dakskobler1 ABSTRACT Phytosociological analysis of montane beech forests on steep shady slopes on mixed geological bedrock in western Slovenia We have conducted phytosociological research into beech forests in western Slovenia that occur on steep shady slopes on dolomite or limestone with a frequent admixture of silicate. We classified the stands on dolomite and partly on platy limestone into the association Arunco-Fagetum and described many of its new subassociations (-caricetosum ferruginei, -vincetosum minoris, -tilietosum). In terms of phytogeography they belong to the geographical variant var. geogr. Anemone trifolia and to two new geographical sub­variants: subvar. geogr. Omphalodes verna and subvar. geogr. Aconitum angustifolium. Beech stands on very steep shady slopes in the montane belt of the southern Julian Alps (from 500 m do 1400 m a.s.l.), which occur on sites character­ised by an admixture of chert, marlstone and (or) claystone on calcareous bedrock and shallow, stony and slightly acid soil are because of significant floristic differences classified into a new, intrazonal association Saxifrago cuneifolii-Fa­getum (alliance Aremonio-Fagion), dependent on land con­figuration and edaphic conditions. We described the follow­ing subassociations: -typicum, -fraxinetosum orni, -rhodo­dendretosum hirsuti,-aruncetosum and -gymnocarpieto­sum dryopteridis. In terms of floristic composition the new association most resembles some forms of the associations Homogyno sylvestris-Fagetum and Rhododendro hisuti­-Fagetum. Key words: phytosociology, synsystematics, phytoge­ography, Arunco-Fagetum, Saxifrago cuneifolii-Fage­tum, Aremonio-Fagion, Julian Alps, Trnovski Gozd pla­teau, Slovenia IZVLEČEK Fitocenološka analiza montanskih bukovih gozdov na strmih osojnih pobočjih na mešani geološki pod­lagi v zahodni Sloveniji V zahodni Sloveniji smo fitocenološko preučili bukove gozdove, ki uspevajo na strmih osojnih pobočjih na dolo­mitu ali apnencu s pogosto silikatno primesjo. Sestoje na dolomitu in deloma ploščastem apnencu smo uvrstili v aso­ciacijo Arunco-Fagetum in opisali več njenih novih sub­asociacij (-caricetosum ferruginei, -vincetosum minoris, -tilietosum). V fitogeografskem smislu pripadajo geografski varianti var. geogr. Anemone trifolia in dvema novima geo­grafskima subvariantama: subvar. geogr. Omphalodes verna in subvar. geogr. Aconitum angustifolium. V to aso­ciacijo zaradi prevelikih florističnih razlik nismo mogli uvrstiti sestojev na zelo strmih osojnih pobočjih v montan­skem pasu južnih Julijskih Alp (od 500 m do 1400 m nm. v.). Zanje je značilna primes roženca, laporovca in (ali) glinavca v karbonatni geološki podlagi in plitva, kamnita in nekoliko zakisana tla. Zato jih uvrščamo v novo edafsko in reliefno pogojeno intraconalno asociacijo Saxifrago cuneifolii-Fa­getum iz zveze Aremonio-Fagion.Opisali smo naslednje njene subasociacije: -typicum, -fraxinetosum orni, -rhodo­dendretosum hirsuti,-aruncetosum in -gymnocarpieto­sum dryopteridis. Po floristični sestavi so novi asociaciji najbolj podobne nekatere oblike asociacij Homogyno sylve­stris-Fagetum in Rhododendro hisuti-Fagetum. Ključne besede: fitocenologija, sinsitematika, fitogeo­grafija, Arunco-Fagetum, Saxifrago cuneifolii-Fagetum, Aremonio-Fagion, Julijske Alpe, Trnovski gozd, Slovenija 1 INTRODUCTION Beech forests on step shady dolomite slopes with shal­low soil (rendzina) in the northwestern part of the Il­lyrian floral province (Marinček 1995) are classified into the association Arunco-Fagetum (Košir 1962, 1979, Accetto 2007). Košir (1979) and Accetto (2007, 2015) published analytical tables for this associ­ation, but descriptions of its species composition, stand structure and ecological conditions are available also in other publications (Marinček 1987: 118–119, Košir 2010: 174–177). Its sites are marked within many poly­gons on two synoptic vegetation maps of Slovenia (Čarni et al. 2002, Košir et al. 2003) as well as on larg­er scale maps (Marinček et al. 2003, 2006), mainly in the Alpine, pre-Alpine, Dinaric and pre-Dinaric phy­togeographical regions. They have also been mapped in western Slovenia, in the Upper Soča Valley, in the Cerkno and the Idrija region. Our inventory of beech stands on steep shady dolomite and limestone slopes began in 1983 in the Cerkno region. They were found and inventoried in large areas in the Bača Valley (1986–1990) and later in the vicinity of Tolmin and the Zadlaščica valley; they were found to be less frequent in the Kobarid region and very rare in the Bovec area, but more frequent in the valleys of the Idrijca and Trebušica. In comparison with sites in other parts of Slovenia the sites in the Upper Soča Valley are clearly different in terms of geological bedrock and conse­quently also in terms of soil conditions. Dolomite is usually mixed with chert; in some places the geological bedrock consists of platy limestone mixed with chert, marlstone and (or) claystone. Admixture of silicate is reflected in skeleton, depth and acidity of the soil and in turn in the species composition. In nearly 30 years we have made more than 200 relevés. By processing them we will try to answer the question of whether the studied beech forests on mixed calcareous-silicate rocks can still be classified within the association Arunco-Fagetum. 2 METHODS Relevés of beech forests were made according to the standard Central-European phytosociological method (Braun-Blanquet 1964) and entered into the FloVeg­Si database (Seliškar & al. 2003). Combined cover-abundance values were transformed into ordinal val­ues 1– 9 (van der Maarel 1979). Numerical compari­sons were conducted with the software package SYN-TAX (Podani 2001). Relevés were arranged into ana­lytic tables based on hierarchical classification. We used the (unweighted) “average linkage” method – UPGMA. Comparison of communities in the synthet­ic table was made with both the UPGMA method and with the “incremental sum of squares” (MISSQ) meth­od and Wishart’s similarity ratio was applied in both. The columns in the synthetic table were compared also with the principal coordinates analysis (PCoA) and Wishart’s coefficient as the similarity ratio as well as with the non-metric multidimensional scaling (NMDS) method with Goodman-Kruskal’S . coeffi­cient. As not all of the authors keep moss species in their tables we only considered vascular plants in these comparisons. Results of numerical methods were com­bined with classic arrangement based on diagnostic species. The nomenclature source for the names of vas­cular plants is Martinčič & al. (2007), Martinčič (2003, 2011) for the names of mosses, and Suppan, Prügger & Mayrhofer (2000) for the names of li­chens. Marinček et al. (1993), is nomenclature source for the names of syntaxa from the alliance Aremonio­-Fagion and Šilc & Čarni (2012) for the names of other syntaxa, except for the name of the class Querco­-Fagetea Braun-Blanquet et Vlieger in Vlieger 1937. Urbančič et al. (2005) is the source for the nomencla­ture of soil types. 2.1 Short ecological description of the study area The Upper Soča Valley is a part of the southern or southwestern Julian Alps. Its geological bedrock is ex­tremely diverse (Buser 1986, 1987), especially in the valleys of the Tolminka, Zadlaščica, Knežica and Bača. The bedrock in these valleys is dominated by Triassic, Jurassic and Cretaceous layers, especially by Bača dolo­mite with cherts and platy limestone mixed with cherts, marlstone and claystone. The Idrijca and Trebušica Valleys also have diverse geological bedrock (Mlakar & Čar 2009, Čar 2010), but Triassic dolo­mite there is not usually mixed with chert. The climate is moist, with mean annual precipitation exceeding 2000 mm (B. Zupančič, 1998); it is relatively warm, with the mean annual temperature of 6°C – 10°C (Ce­gnar 1998). We inventoried the shady slopes that have a considerably colder local climate than the average. Forests in the study area have been managed under forest management plans for a very long time (Perko et al. 2014). Their structure and quality today are large­ly the result of heavy cutting and charcoal production in the first third of the 20th century. In late winter 2014 large areas of these forests were heavily affected by sleet. 3 RESULTS AND DISCUSSION 3.1 Numerical comparison and assessment of diagnostic species Phytosociological relevés of beech forests on steep shady dolomite and limestone slopes in the Upper Soča Valley, on the northern edge of the Trnovski Gozd pla­teau and in the Cerkno and Idrija Hills were arranged based on the results of hierarchical classification into several analytic tables (1 to 10). Most of them were mu­tually compared with relevé material from similar for­est communities (Arunco-Fagetum, Homogyno sylve­stris-Fagetum, Rhododendro hirsuti-Fagetum, Ranun­culo platanifolii-Fagetum) in western and partly also southeastern Slovenia. The synthetic table (Table 11) comprises 18 columns. The compared syntaxa grouped as shown in Figures 1, 2, 3, and 4. The results point to the following conclusions. The larger group of relevés that were arranged into two vegetation tables (1 and 2, see also columns 5 and 6 in Table 11) can, according to their floristic composi­tion, undoubtedly still be classified into the association Arunco-Fagetum. Other relevés are more similar to the relevés of the associations Homogyno sylvestris-Fage­tum and Rhododendro hirsuti-Fagetum, which we stud­ied in western Slovenia and published the results sev­eral years ago (Dakskobler 2002, 2003). If the com­parison includes also the analysis of diagnostic species (Table 12) and the analysis of percentages by groups of diagnostic species (Table 13) we find that in all com­pared beech forests there are very few species associ­ated with only one vegetation type. Having conducted these comparisons we classify into the association Arunco-Fagetum in western Slovenia the beech stands in the (sub)montane belt (200 to 1000 m a.s.l.) on shady dolomite slopes on the northern edge of the Trnovski Gozd plateau, in the Idrija and Cerkno Hills as well as some beech stands on similar slopes and elevation in the southern Julian Alps with their foothills, where the geological bedrock is frequently dolomite with chert, in places also platy limestone with chert. The stands in the montane-altimontane belt on mixed geological bedrock (dolomite with chert, marl­stone or claystone, platy limestone with chert, marl­stone or claystone) are floristically very different. Some of them were mapped as the association Homogyno sylvestris-Fagetum already around 1990 (the map was published much later, Dakskobler 2007a), while oth­ers were marked on field maps and in the database under the provisional names Calamagrostio-Fagetum and Saxifrago cuneifolii-Fagetum. Since 1993 (Marin­ček et al. 1993) the association Homogyno sylvestris­-Fagetum comprises the pre-Alpine fir-beech commu­nity that had previously been known as Abieti-Fage­tum prealpinum.This community is distributed in the larger part of northern Slovenia on calcareous and mixed calcareous-silicate bedrock (Marinček & Čarni 2007). The species composition of its geograph­ical variant from the Kneža, Zadlaščica and Tolminka valleys (Homogyno sylvestris-Fagetum var. geogr. Sesle­ria autumnalis) is in fact very similar to the species composition of some forms of the studied communi­ties, which is partly attributed to the same distribution area. This similarity would be much smaller if the comparison included other geographical variants of the association Homogyno sylvestris-Fagetum. The principal diagnostic species of this community is silver fir (Abies alba), which is very rare in our relevés; it oc­curs only individually and mainly in the shrub layer. This could be partly attributed to the past manage­ment practice and degradation processes. In this fir-beech community it is Adenostyles glabra and Homo­gyne sylvestris that have a higher frequency and medi­an cover. The beech stands that are more indicative of a floristic similarity with fir-beech forests were grouped in Tables 6 and 10 (see also columns 8 and 9 in Table 11) and we allow for the possibility for them to be classified also into the association Homogyno sylve­stris-Fagetum. Phytocoenoses in Tables 7 and 9 (see also columns 10, 11 and 12 in Table 11) on the other hand are floristically well differentiated from pre-Al­pine fir-beech forests. Some of the studied stands (Table 8, relevés 1–8, see also column 13 in Table 11) are, in terms of species composition, slightly similar also to the stands of the association Rhododendro hir­suti-Fagetum.These usually occur on even more ex­treme sites and have a physiognomy of a subalpine beech forest with abundant hairy alpenrose (Rhodo­dendron hirsutum) in the undergrowth. In our stands, hairy alpenrose occurs only individually and therefore serves merely as a differential species of the subasso­ciation. The special type of beech forest from the alliance Aremonio-Fagion in Tables 6, 7, 8, 9 and 10, which is associated with special edaphic factors and land con­figuration, is therefore classified into the new associa­tion Saxifrago cuneifolii-Fagetum.The stands of this association are characterised by dominating Calama­grostis arundinacea and the occurrence of frigophilous and acidophilous species, especially character species of spruce forests that grow together with thermophil­ous species from the order Quercetalia pubescenti-pe­traeae and class Trifolio-Geranietea, as well as with the species of dry and subalpine grasslands, screes and rock crevices in the herb layer. With regard to the dominant species in the herb layer these stands could also be classified into the association Calamagrostio arundinaceae-Fagetum Cerovečki 2009, which Cero­večki (2009) described in the altimontane belt of west­ern Croatia, on calcareous bedrock and shallow soil. This name is a subsequent homonym, because the same name was used to describe the association Cala­magrostio arundinaceae-Fagetum Sýkora 1972 (Sýkora 1972, Boublík et al. 2007) that had validly been de­scribed before, in the Carpathians. The stands of the Carpathian association as well as the stands of the as­sociation Calamagriostidi (Abieti)-Fagetum (for which Hartmann & Jahn (1967) published a table based on Hartmann’s relevés from Central-European mountain ranges north of the Alps) are considerably different from our relevés. We have established a certain simi­larity with the relevés published by Cerovečki (ibid.). Common to both associations are some species from the alliance Aremonio-Fagion, order Fagetalia sylvatica and classes Erico-Pinetea, Vaccinio-Piceetea and Mul­gedio-Aconitetea. The stands of both compared syn­taxa comprise some of the species that characterise cold, stony, slightly acid sites, such as Calamagrostis arundinacea, Veronica urticifolia, Adenostyles glabra and Homogyne sylvestris. The association Saxifrago cu­neifolii-Fagetum is differentiated against the associa­tion Calamagrostio-Fagetum sensu Cerovečki by sev­eral diagnostic species: Tanacetum corymbosum, Lu­zula luzuloides, Saxifraga cuneifolia and Huperzia sela­go; phytogeographical differential species: Aconitum angustifolium, Larix decidua, Anemone trifolia,Carduus crassifolius; some thermophilous species: Sesleria autu­mnalis, Ostrya carpinifolia, Fraxinus ornus, Melittis melissophyllum, Arabis turrita, as well as by chasmo­phytic species and species of subalpine grasslands: Asplenium viride, Sesleria caerulea subsp. calcaria, Be­tonica alopecuros, Laserpitium peucedanoides. The spe­cies that differentiate the association Calamagrostio­-Fagetum sensu Cerovečki against the association Saxifrago cuneifolii-Fagetum include Calamintha gran­diflora, Ribes petraeum and Rosa pimpinellifolia. The differences are confirmed also by the calculation of floristic similarity according to Sorensen (1948). Flo­ristic similarity between the syntaxa Calamagrostio arundinaceae-Fagetum sensu Cerovečki and Saxifrago cuneifolii-Fagetum typicum is only 43 %, which means they cannot be classified into the same association. In addition to identifying the stands that indicate a contact or similarity between associations Homogyno sylvestris-Fagetum and Saxifrago cuneifolii-Fagetum (Table 10), and the stands that indicate a contact or similarity between the stands of associations Rhodo­dendro hirsuti-Fagetum and Saxifrago cuneifolii-Fage­tum (Table 8), we joined in Table 6 (see also column 8 in Table 11) the relevés that are, according to numerical comparisons, floristically more or less similar to the relevés of the association Saxifrago cuneifolii-Fagetum and to the relevés of the association Homogyno sylve­stris-Fagetum, but are, on account of the abundance of diagnostic species of the association Arunco-Fagetum, treated as a special subassociation Saxifrago cuneifolii­-Fagetum aruncetosum. 3.2 Overview of the newly described syntax The determined syntaxa are described below with con­sideration of the above-mentioned reservations. Class: Querco-Fagetea Br.-Bl. et Vlieger in Vlieger 1937 Order: Fagetalia sylvaticae Walas 1933 Alliance: Aremonio-Fagion (Ht. 1938) Borhidi in Török, Podani & Borhidi 1989 Association: Arunco-Fagetum Košir 1962 - anemonetosum trifoliae prov. var. Cardamine pentaphyllos var. Gymnocarpium robertianum - caricetosum ferrugineae subss. nov. - vincetosum minoris subass. nov. - tilietosum subass. nov. var. Luzula nivea var. Hacquetia epipactis Association: Saxifrago cuneifolii-Fagetum ass. nov. -typicum subass. nov. var. Calamagrostis varia var. Primula columnae -fraxinetosum orni subass. nov. -aruncetosum dioicae subass. nov. var. Veratrum album var. Hedera helix -gymnocarpietosum dryopteridis subass. nov. -rhododendretosum hirsuti subass. nov. -caricetosum albae prov. -caricetosum humilis prov. The division of the new forms of the association Arun­co-Fagetum in terms of phytogeography is as follows: Arunco-Fagetum Košir 1962 var. geogr. Anemone trifo­lia Košir ex Dakskobler var. geogr. nov. subvar. geogr. Omphalodes verna subvar. geogr. nov. subvar. geogr. Aconitum angustifolium subvar. geogr. nov. 3.3 Association Arunco-Fagetum in the Soča Val­ley 3.31 Phytogeographical division of the association Arunco-Fagetum in Slovenia Approximate localities of relevés of the association Arunco-Fagetum in the Soča Valley are presented in Figures 5 and 6. These relevés were arranged in three tables (Tables 1, 2 and 3). Several relevés from other parts of Slovenia were arranged in Tables 4 and 5. The association Arunco-Fagetum from the Soča Valley is classified into the geographical variant var. geogr. Anemone trifolia var. geogr. nova that was first decribed by Košir (1979: 164), but only with a few sen­tences and without supporting tabular material. The nomenclature type (holotypus) of the new geographical variant is relevé No. 24 in Table 2. This is also the holo­type of the new geographical subvariant, subvar. geogr. Aconitum angustifolium subvar. geogr. nov. that com­prises the relevés from both the southern and western foothills of the Julian Alps. We also described a new geographical subvariant, subvar. geogr. Omphalodes verna subvar. geogr. nov. Its nomenclature type, holo­typus, is relevé No. 18 in Table 1. Its stands dominate on the slopes above the Idrijca and Trebušica valleys, at the contact of the pre-Alpine and Dinaric phytogeo­graphical regions. The stands of the association Arun­co-Fagetum are distributed also in other parts of the pre-Alpine region of Slovenia, so the geographical var­iant Anemone trifolia is likely to comprise other geo­graphical subvariants. Stands of this geographical variant may occur also in the Sava Valley. Relevé No. 1 in Table 4, which was made on the slopes of Kum, could be classified either within this variant or into a special geographical subvariant var. geogr. Ruscus hypoglossum Košir 1979 subvar. geogr. Anemone trifo­lia prov. The stands of the association Arunco-Fagetum may also be occurring in the sub-Pannonian region of Slovenia. Table 5 comprises three relevés from the shady slopes of Donačka Gora, whose ecology and in part also species composition indicate a certain simi­larity with the stands of the association Arunco-Fage­tum. These relevés stood out from all other compared relevés with their entire floristic composition and grouped into a completely separate cluster. They stand out also in terms of geological bedrock, which is quartz conglomerate. The slopes are very steep. The stand structure and relief conditions are very similar to those in the stands of the association Arunco-Fagetum. These three relevés are temporarily classified into the new geographical variant Arunco-Fagetum var. geogr. Hie­racium rotundatum. Its nomenclature type, holotypus, is relevé No. 1 in Table 5. Phytogeographical differen­tial species include Poa stiriaca, Sesleria sadlerana and Galium sylvaticum. Cimperšek (2004: 441) classifies such stands on Donačka Gora into the syntaxon Lamio orvalae-Fagetum var. geogr. Dentaria polyphylla luna­rietosum redivivae var. Festuca altissima subvar. Arun­cus dioicus and mentions their physiognomic, floristic and ecological similarity with the stands of the asso­ciation Arunco-Fagetum. Marinček and Marinšek (in litt.) classified these stands into the association Festuco drymejae-Fagetum. The existing phytogeographical division of the as­sociation Arunco-Fagetum is therefore as follows: Arunco-Fagetum Košir 1962 var. geogr. Ruscus hypo­glossum Košir 1979 (Dolenjska/Lower Carniola) Arunco-Fagetum Košir 1962 var. geogr. Acer obtusatum Accetto 2007 (Upper Kolpa Valley) Arunco-Fagetum Košir 1962 var. geogr. Calamintha grandiflora Accetto 2015 (Iški vintgar) Arunco-Fagetum Košir 1962 var. geogr. Anemone trifo­lia Košir ex Dakskobler 2015 (Alpine, pre-Alpine and partly Dinaric region of Slovenia) subvar. geogr. Omphalodes verna (the Idrijca Val­ley with its tributaries Trebušica, Belca) subvar. geogr. Aconitum angustifolium (Upper Soča Valley) Arunco-Fagetum Košir 1962 var. geogr. Hieracium ro­tundatum (Donačka Gora) 3.32 Arunco-Fagetum var. geogr. Anemone trifolia sub­var geogr. Omphalodes verna (Table 1) The stands of this geographical subvariant were re­corded mainly on the northern edge of the Trnovski Gozd plateau above the Trebušica and Belca valleys, in the gorges between Vojsko and Trebuša, in the Idrija and Cerkno Hills. The elevation of the relevés ranges between 200 and 1000 m; the predominantly dolomite geological bedrock is sporadically mixed with chert or marlstone and the soil is mainly rendzina. Slopes range between 25° and 45°; the aspect is mainly shady, only occasionally sunny (southwestern). The diagnostic species of the association as selected by Košir (1962, 1979) are represented in different proportions in our relevés. The only relatively frequent character species in the relevés of this geographical subvariant are Arun­cus dioicus and Helleborus niger, but their abundance is far below that in the sample plots from the region where this association was originally described. Arun­cus dioicus is definitely not a species that would spe­cifically characterise these stands or stand out in any way, and the same applies to most of the herb species that give their name beech associations from the alli­ance Aremonio-Fagion. The differential species Carda­mine enneaphyllos and Mercurialis perennis are gener­ally distributed in Illyrian beech forests and are not presented separately in the tables. The same can be said for Prenanthes purpurea, which we kept in the di­agnostic combination. Based on the comparative table (Tables 11, 12) we additionally classified Tamus com­munis and Veratrum nigrum as differential species of the association Arunco-Fagetum. Both these species are otherwise more characteristic for stands of the as­sociation Ostryo-Fagetum and well-characterise forests on dolomite bedrock and shallow soil. Geographical differential specis are Anemone trifolia, Omphalodes verna, Phyteuma spicatum subsp. coeruleum, Tephrose­ris longifolia, Rhamnus fallax, Scopolia carniolica, Po­tentilla carniolica and Lathyrus vernus subsp. flaccidus. Anemone trifolia characterises mainly beech forests in the pre-Alpine-Alpine part of Slovenia. Omphalodes verna is most frequent in beech forests of the Dinaric part of Slovenia. The taxon Phyteuma spicatum subsp. coeruleum is frequent in the Idrijca and Trebuša val­leys and in the Trnovski Gozd plateau, but is not known in the Bača Valley and in valleys to the north of Tolmin. Rhamnus fallax, Scopolia carniolica and Tephroseris longifolia have a predominantly eastern-Alpine–Illyrian distribution. The northernmost sites of Potentilla carniolica in the Soča Valley are (beside newly discovered locality in Trenta – Trnkoczy, mscr.) in the valley of the Idrijca at Slap ob Idrijci. Its distri­bution is Dinaric (Čušin & Dakskobler 2001: 77). The taxon Lathyrus vernus subsp. flaccidus is frequent in beech forests in the Soča Valley (Dakskobler 1991: 20). The listed species well characterise the geographi­cal area between Most na Soči, Cerkno, Idrija and Gorenja Trebuša. Two subassociations are distinguished. The relevés of the subassociation -caricetosum ferruginei were made on the slopes above Črna Draga above the valley of the Belca in Zgornja Idrijca Landscape Park and in Mali Govci above the headwaters of the Trebušica. The differential species of the subassociation are Labur­num alpinum, Carex ferruginea and Adenostyles glabra. These three species characterise beech forests in the montane belt, at elevations between 800 m and 1000 m, in areas otherwise dominated by fir-beech forest (Omphalodo-Fagetum s. lat.). Surina (2002) and Suri­na & Dakskobler (2013) classified similar beech stands into the syntaxon Omphalodo-Fagetum var. geogr. Saxifraga cuneifolia calamagrostietosum variae. We allow for the possibility that the absence of fir, or its expressly insignificant occurrence (individual spec­imens within different stand layers), are partly associ­ated with past management. In any case, stands of this subassociation indicate a contact between the associa­tions Arunco-Fagetum and Omphalodo-Fagetum. The nomenclature type, holotypus, of the new subassocia­tion Arunco-Fagetum caricetosum ferruginei is relevé No. 7 in Table 1. Relevés of the subassociation -vincetosum minoris were made mainly in the Cerkno Hills and in the Trebuša Valley, where we made several relevés also in Govci on the northern edge of the Trnovski Gozd pla­teau. The elevation is slightly lower with most relevés made at between 500 m and 600 m and quite a few made even lower, in the submontane belt. The differen­tial species of the subassociation are Vinca minor, Fraxinus ornus, Ostrya carpinifolia, Vincetoxicum hi­rundinaria, Veratrum nigrum (diagnostic species for the association) and Campanula rapunculoides. The differential species indicate relatively favourable ther­mal conditions that allow for forest growth despite the steep slopes and a shady aspect. The nomenclature type, holotypus, of the new subassociation is relevé No. 18 in Table 1. 3.33 Arunco-Fagetum var. geogr. Anemone trifolia sub­var geogr. Aconitum angustifolium (Table 2) Localities of the stands of this geographical subvariant are mainly in the vicinity of Tolmin (Bučenica, Kozlov Rob, Godiča), some ot the relevés are from the lower part of the Idrijca Valley, i.e. from the slopes of lateral ravines in the valley; many of the relevés come from the steep slopes of Mts. Matajur and Mija above the Nadiža gorge. One relevé is from the shady slopes of the Stol ridge above Srpenica in the Bovec region and one from the Čepovan Valley. The diagnostic species of the association, especially Helleborus niger, are rela­tively rare. This is probably because the geological bedrock is frequently platy limestone with chert or Bača dolomite with chert. On such bedrock, Hellebo­rus niger occurs much more rarely than on pure dolo­mite or on dolomitised limestone. Geographical dif­ferential species Anemone trifolia, Geranium nodosum, Sesleria autumnalis, Luzula nivea, Aconitum angustifo­lium, Tephroseris pseudocrispa and Lathyrus vernus subsp. flaccidus very well characterise forests in the wider vicinity of Tolmin, i.e. in that part of the Upper Soča Valley where the sub-Mediterranean influence is still reflected in both the climate and vegetation. The stands of the association Arunco-Fagetum are very rare in the more Alpine part of the Upper Soča Valley (Bovec region) and are replaced by different forms of the association Anemono trifoliae-Fagetum. Similar applies to other regions of the Slovenian Alps (Košir 1979: 164). The stands of this geographical subvariant were divided into two subassociations. All relevés in Table No. 2 are classified into the new subassociation Arun­co-Fagetum tilietosum. Its differential species are Tilia cordata, T. platyphyllos and Phillitis scolopendrium. The listed species indicate relatively moist, stony sites and slightly colluvial soil; indirectly they also indicate geological bedrock that is not characteristic for the as­sociation Arunco-Fagetum, namely platy limestone with cherts. Freqeuntly occuring in this part of the Upper Soča Valley, on talus slopes and rockfall at the foot of shady slopes, are lime stands classified into the associations Saxifrago petraeae-Tilietum and Veratro nigri-Fraxinetum (Dakskobler 2007b), and at least in some places these lime stands are in contact also with the stands of the association Arunco-Fagetum. The no­menclature type, holotypus, of the new subassociation Arunco-Fagetum tilietosum is relevé No. 24 in Table 2. Two variants are distinguished. The variant Luzula nivea, whose differential species include Cardamine pentaphyllos, Moehringia muscosa and Vincetoxicum hirundinaria, characterises the sites where the geologi­cal bedrock predominantly consists of limestone with chert. Stoniness is considerable and the soil in places more developed – brown rendzina with transitions to chromic cambisol. The dominant stands are those at slightly higher elevations and in a slightly colder cli­mate. The stands of this variant indicate a certain flo­ristic similarity with the stands of the geographical subvariant Lamio orvalae-Fagetum var. geogr. Denta­ria pentaphyllos subvar. geogr. Luzula nivea. Stands of the variant with Hacquetia epipactis and other differ­ential species such as Allium ursinum, Euphorbia amygdaloides, Pulmonaria officinalis and Ruscus acu­leatus dominate at lower elevations, in the submontane belt, in a slightly warmer climate, on dolomite with chert, on less stony and relatively fresh soil. The dif­ferential species of the variant indicate a certain simi­larity or contact with the stands of the syntaxon Ha­cquetio-Fagetum var. geogr. Anemone trifolia. In Table 3 we collected the relevés that grouped separately from all other relevés of the association Arunco-Fagetum in the numerical comparison; these relevés are quite special and differ also from each other. They were made in different parts of the Soča Valley. Relevés Nos. 1 to 10 are temporarily classified into the provisional subassociation Arunco-Fagetum anemonetosum trifoliae prov. Two variants are distin­guished. Var. Cardamine pentaphyllos with differential subspecies Aconitum degenii subsp. paniculatum char­acterises fresher sites, while the var. Gymnocarpium robertianum (its differential species include Sesleria caerulea subsp. calcaria, Campanula rapunculoides, Rubus saxatilis, Vincetoxicum hirundinaria, Luzula nivea, Aconitum angustifolium and Vinca minor) indi­cates more initial, shallow and skeletal soil. In two rel­evés at the foot of Mt. Mija we recorded also Epimedi­um alpinum. For the time being, relevés Nos. 11 to 13 in Table 3 remain indeterminate in terms of their syn­taxonomical status. Relevé No. 12 shows considerable similarity with the stands of the association Saxifrago cuneifolii-Fagetum that will be presented below. 3.4 Description of the new association Saxifrago cuneifolii-Fagetum 3.41 Localities and ecological conditions Approximate localities of the relevés of the association Saxifrago cuneifolii-Fagetum are shown in Figures 7 and 8. These relevés were arranged in five tables (Ta­bles 6, 7, 8, 9 and 10). So far, its stands have been found only in the upper Bača Valley, primarily on the steep slopes of Babji Zob, Šoštar and Kobla above the streams of Linderpoh and Kacenpoh, in Trtnikarski Gozd for­est at Šprickovbl Falls above Kacenpoh, under Robar­jev Grič above Batava at Podbrdo, under Mt. Kojca, on the peak of Bizle under Mt. Hohkovbl (Matajurski Vrh) above Rut, under the Jehle– Koblar–Znojilski Vrh ridge above Bideržuna or the Huda Grapa gorge and above the Koritnica valley, on steep slopes under the Čumik–Luken ridge above the Žventarska Grapa gorge at the village of Grant, on steep shady slopes of Jalovnik above the Kneža Gorge (Temna Brda, Temnak above the Mohor homestead), under Tisovec, Krikov Vrh and the mountain pasture Kuk above the headwaters of the Lipovšček, also in the Kneža (Knežica) basin, and under Vrh nad Sopotom above the Zadlaščica valley. There is also an individual locality in the Kobarid re­gion under the peak Črnik above Drežnica in the Krn Mountains and one in the Bovec region on Strmi Breg on the shady slopes of the Stol ridge above Srpenica. The localities of the stands of this association are large­ly situated where geologists have mapped layers of Ju­rassic micrite and calcarenite limestone with chert sheets and subordinate clayey marlstone, as marked by Buser (1986, 1987) on the geological map and in the accompanying explanatory notes. On every relevé, limestone or dolomite is mixed with chert, claystone or marlstone, with chert completely dominating in some places. Mixed geological bedrock is therefore one of the key factors that condition the special (unique) spe­cies composition of this association. Another decisive factor is the considerable slope (usually between 30° and 45°) and the third is the predominantly shady as­pect. The vertical range of the association is consider­able. The lower line of occurrence is at the elevation of around (400) 500 m. The highest elevation at which we found these stands was under Mt. Črna Prst and Mt. Hohkovbl (Matajurski Vrh) at nearly 1400 m, but most of the stands are situated at the elevations between 900 m and 1250 m. Whereas the stands at lower elevations always occur on shady slopes, the stands in the alti­montane belt can be situated also on sunny slopes. The soil conditions were not analysed in much detail. The soil is always shallow, skeletal. The dominating soils are rendzina-type soils with transitions to cambisols, in places dystric cambisols. The climate is montane and very humid, with mean annual precipitation ex­ceeding 2000 mm, in places even 3000 mm; due to the predominating shady aspect it is also cold. The forest growth conditions are rather extreme as a result of the slope, stoniness and wind exposure, and the growth of these stands is correspondingly inhibited. Only the stands at lower elevations occasionally have larger growing stocks and better growth rates and beech can reach the heights of more than 25 m. Stands in the alti­montane belt are low, forming clusters or coppice, with the tree height reaching between 15 m and 20 m, some­times less. 3.42 Species composition, diagnostic species and no­menclature type of the new association The tree layer is dominated by beech (Fagus sylvatica). The most common companion species are Sorbus aria and Laburnum alpinum, in places also Acer pseudopla­tanus, Picea abies, Ostrya carpinifolia, Fraxinus ornus, Sorbus aucuparia, Larix decidua (differential against the association Arunco-Fagetum), rarely Abies alba, and other. The dominating shrubs are Daphne mezere­um, Lonicera alpigena, Rosa pendulina and in places Rubus hirtus. The most common species in the herb layer in­clude Cyclamen purpurascens, Cardamine enneaphyl­los, Cardamine trifolia, Primula vulgaris, Mercurialis perennis, Prenanthes purpurea, Dryopteris filix-mas, Salvia glutinosa, Symphytum tuberosum, Galium laevi­gatum, Anemone nemorosa, Cirsium erisithales, Solida­go virgaurea, Gentiana asclepiadea, Oxalis acetosella, Homogyne sylvestris, Veronica urticifolia, Calamagro­stis arundinacea, Luzula luzuloides, Hieracium muro­rum, Phyteuma ovatum, Senecio ovatus, Athyrium filix­-femina, Adenostyles glabra and some other species. The moss layer is abundant and can cover more than 20% of the sample plots. The most common spe­cies in the moss layer are Ctenidium molluscum, Necke­ra crispa, Polytrichum formosum, Tortella tortuosa, Atrichum undulatum, Isothecium alopecuroides, Met­zgeria furcata and many others. The moss layer has been only partly inventoried and our determination is not always reliable. The composition of the community by groups of diagnostic species is shown in Table 13, columns 8 to 13. Compared to the stands of the association Arunco­-Fagetum its stands comprise considerably fewer meso­philous species characteristic for the alliance Tilio­-Acerion and beech species from the alliance Aremo­nio-Fagion, but contain substantially more acidophil­ous species from the class Vaccinio-Piceetea. In com­parison with the stands of the association Ranunculo platanifolii-Fagetum the stands of the new association comprise significantly more species of the order Quer­cetalia pubescenti-petraeae and of the class Elyno-Sesle­rietea, but considerably fewer tall herb species from the class Mulgedio-Aconitetea. The diagnostic species of the new association are Veronica urticifolia, Calama­grostis arundinacea, Luzula luzuloides, Huperzia sela­go, Saxifraga cuneifolia, Tanacetum corymbosum s. lat. (inc. T. clusii), Campanula witasekiana, Festuca hete­rophylla, Clinopodium vulgare and Carex humilis. Geo­graphical differential species are Anemone trifolia, Aconitum angustifolium, Lathyrus vernus subsp. flacci­dus, Larix decidua and Sesleria autumnalis. The listed species reflect the special ecology of these sites – very steep slopes in the (alti)montane belt, mixed geological bedrock, shallow, but slightly acid soil in the southern Julian Alps. The nomenclature type of the new associ­ation Saxifrago cuneifolii-Fagetum ass. nov., holotypus, is relevé No. 4 in Table 7. It is classified into the alliance Aremonio-Fagion,order Fagetalia sylvaticae and class Querco-Fagetea. The new association was named after Saxifraga cuneifolia, a southern-European montane species, a character species of the alliance Piceion excelsae and its suballiance Abieti-Piceenion. It usually grows in the montane and subalpine belt on acid, ni­trogen-rich soil. It is distributed from the Pyrenees across the Alps to the eastern Carpathians, as well as in the Apennines and in the Dinaric Mountains (Ae­schimann et al. 2004: 692, Accetto 1993, 1995, 2002, 2006, Zupančič & Accetto 1994, Zupančič 1999). Distribution in Slovenia presented according to the data collected in the FloVegSi database (Seliškar et al. 2003) is shown in Figure 9. It is very frequent in the Alpine and pre-Alpine regions of Slovenia, but consid­erably less frequent in its Dinaric and pre-Dinaric parts. It occurs in many forest communities, especially in beech, fir-beech, spruce and larch communities. It is the most frequent in the stands of the associations Anemono trifoliae-Fagetum, Rhododendro hirsuti-Fage­tum, Ranunculo platanifolii-Fagetum, Polysticho lon­chitis-Fagetum, Homogyno sylvestris-Fagetum, Ade­nostylo glabrae-Piceetum,Laburno alpini-Piceetum, Asplenio-Piceetum and Rhodothamno-Laricetum, sometimes it occurs also in the stands of the syntaxa Saxifrago petraeae-Tilietum,Veratro nigri-Fraxinetum excelsioris, Arunco-Fagetum, Lamio orvalae-Fagetum, Omphalodo-Fagetum var. geogr. Saxifraga cuneifolia, Tanaceto clusii-Fagetum, Luzulo sylvaticae-Piceetum var. geogr. Luzula nivea, Campanulo justinianae-Picee­tum, Ribeso alpini-Piceetum, Neckero crispae-Campa­nuletum justinianae and several others. It characterises cold stony sites with moder rendzina or acid soil reac­tion in the montane belt and with its ecology it is a very suitable species to describe the sites of the new associa­tion. 3.43 Lower syntaxonomic units of the association Saxi­frago cuneifolii-Fagetum Most of the relevés of the stands of the subassociation Saxifrago cuneifolii-Fagetum aruncetosum were made on the steep slopes above the valley of the Kneža or its right tributary Lipovšček, and only two relevés were made on the slopes above the Bača Valley at Hudajužna or Zakojca. Most of the relevés are from the montane belt and were made at elevations of between 600 m and 1200 m, the geological bedrock is mainly dolomite with chert. Except for Carex humilis and Campanula witasekiana the diagnostic species of the association are relatively frequent. Also present are the geographi­cal differential species Anemone trifolia, Sesleria autu­mnalis, Lathyrus vernus subsp. flaccidus and Aconitum angustifolium. Differential species of the subassocia­tion are Festuca altissima, Aruncus dioicus, Veratrum nigrum and Laserpitium krapfii s. str. The latter is an Illyrian-Carpathian taxon and a character species of the association Arunco-Fagetum. The northern border of its distribution area and its only localities in the Alps are in the slopes above the gorges of the Lipovšček and the Zadlaščica and above the Bača Valley. The sub­species L. krapfii subsp. gaudinii occurs elsewhere in the Alps (Aeschimann et al. 2004: 1144, Bačič et al. 2015). The listed species indicate a similarity with the stands of the association Arunco-Fagetum. Based on diagnostic species it could also be classified into this association. In terms of their floristic composition these stands are very similar to the stands of the syn­taxon Homogyno sylvestris-Fagetum var. geogr. Sesleria autumnalis typicum, which is one of the forms of pre-Alpine fir-beech forests described in the same area. At present, these stands cannot be classified within this syntaxon as the fir very rarely occurs within them and even then only with individual specimens. With the name of the subassociation and by selecting their dif­ferential species we demonstrated the contact or simi­larity with the stands of another two associations, Arunco-Fagetum and Homogyno sylvestris-Fagetum. The nomenclature type, holotypus, of the new subas­sociation Saxifrago cuneifolii-Fagetum aruncetosum di­oici is relevé No. 3 in Table 6. Two variants are distin­guished. The variant with Veratrum album character­ises forests in the upper part of the montane belt, on fresh sites. Its differential species are Veratrum album and Aconitum degenii subsp. paniculatum. The variant with Hedera helix (its differential species include Asa­rum europaeum subsp. caucasicum, Galium odoratum and Helleborus odorus) characterises the sites and stands in the lower montane belt that are floristically slightly similar also to the stands of the associations Lamio orvalae-Fagetum s. lat. and Hacquetio-Fagetum s. lat. The nomenclature type of the typical subassocia­tion Saxifrago cuneifolii-Fagetum typicum is the same as the nomenclature type of the new association, i.e. relevé No. 4 in Table 7. It incorporates the stands of the altimontane belt at the elevations between 900 m and 1370 m. All of the relevés were made on very steep slopes above the Bača Valley on shady aspects, apart from the few that were made on sunny aspects. They occur in the belt otherwise dominated by altimontane beech stands of the association Ranunculo platanifolii­-Fagetum. The comparison with two forms of this as­sociation as described by Marinček & Čarni (2010) in the same phytogeographical region demonstrates notable differences (compare Tables 11 and 13, col­umns 11 and 12, and 17 and 18). The species that dif­ferentiate the typical form of the association Saxifrago cuneifolii-Fagetum against the association Ranunculo platanifolii-Fagetum are the species of the order Quer­cetalia pubescenti-petraeae and classes Trifolio-Gerani­etea and Elyno-Seslerietea. All diagnostic species of the association are frequent in the stands of the typical subassociation. In addition to the typical variant that comprises primarily the stands on very steep, shady slopes, we differentiate two other variants. The variant with Calamagrostis varia, its differential species in­clude Carex digitata, Carduus crassifolius and Laserpi­tium peucedanoides, characterises open stands on very steep, sunny and shady slopes with shallow soil on limestone or dolomite with chert, claystone or marl­stone in the upper Bača Valley. The variant with the taxon Primula veris subsp. columnae characterises ex­treme sites, mainly sunny slopes on limestone with chert or almost pure chert on slopes above the central and lower Bača Valley (above Rut and at the mountain pasture Kuk). Its differential species are also Sesleria autumnalis, Lathyrus vernus subsp. flaccidus and Ane­mone trifolia. The role of the forests of this subassocia­tion is exlusively protective as they represent extreme sites for forest growth. The nomenclature type, holotypus, of the subasso­ciation Saxifrago cuneifolii-Fagetum rhododendretosum hirsuti, is relevé No. 1 in Table 8. The differential spe­cies of the subassociation are Rhododendron hirsutum and Clematis alpina. Stands of this subassociation were recorded in the altimontane belt, at the elevations be­tween 1100 m and 1200 m, mostly on both sides of the ridge between the valleys of the Zadlaščica and the Lipovšček. With their entire floristic composition these stands very much resemble the stands of the local form of the association Rhododendro hirsuti-Fagetum (Dakskobler 2003). A detailed comparison of phyto­sociological tables of both syntaxa, especially of abun­dance (frequency and cover value) of diagnostic spe­cies, demonstrates notable differences. In the stands of the association Rhododendro hirsuti-Fagetum hairy al­penrose (Rhododendron hirsutum) usually has a cover value of 1, 2 or 3, but in our case only “r” or “+”. These relevés might indicate a certain similarity, in terms of increasingly extreme sites, between the stands of two intrazonal associations dependent on land configura­tion and edaphic conditions. Table 8 also encompasses several relevés that cannot yet be synsystematically classified in more detail. They formed completely sep­arate groups from other relevés in numerical compari­sons. Relevés 13 to 15 in this table might characterise the most thermophilous form of the association Saxi­frago cuneifolii-Fagetum, which occurs on steep sunny slopes and open rocky ridges. They are temporarily treated as the provisional subassociation Saxifrago cu­neifolii-Fagetum caricetosum humilis prov. Given the poorly represented diagnostic species of the studied as­sociation these relevés may even belong to a new asso­ciation provisionally named Carici humilis-Fagetum prov. The nomenclature type, holotypus, of the subasso­ciation Saxifrago cuneifolii-Fagetum fraxinetosum orni, is relevé No. 26 in Table 9. The differential species of the subassociation is Fraxinus ornus that characterises the relevés at slightly lower elevations, in the montane belt between 500 m and 1000 m. Its species composi­tion comprises also the species that are more typical for the submontane belt and were not recorded in the stands of the typical subassociation, such as Hedera helix, Lonicera xylosteum, Acer campestre, Vinca minor. These are slightly less estreme sites compared to the typical subassociation, on almost consistently shady aspect; the soil is occasionally more developed, with transitions to cambisols, which slightly improves the forest growth rate. These are mainly more or less man­aged forests and have a strong protective role. Some sections were heavily cut in the past. The largest areas of stands of this subassociation are on shady slopes above the Kacenpoh gorge at Podbrdo and on shady slopes above Žventarska Grapa gorge at Grant. Slightly rarer among the diagnostic species of the association are those associated with shallow soil and moder rendzina, such as Campanula witasekiana, Carex hu­milis and Clinopodium vulgare. Also the stands of this subassociation are slightly similar to the stands of the association Arunco-Fagetum both in their physiogno­my and entire floristic composition, but the numerical comparison nevertheless clearly demonstrates they cannot be classified therein. The nomenclature type, holotypus, of the new sub­association Saxifrago cuneifolii-Fagetum gymnocarpie­tosum dryopteridis, is relevé No. 2 in Table 10. The dif­ferential species of the subassociation are Gymnocarpi­um dryopteris, Acer pseudoplatanus, Abies alba, Phyte­uma ovatum, Actaea spicata, Polystichum aculeatum, Dryopteris affinis and Adoxa moschatellina. They indi­cate very steep and stony slopes on dominating chert with shallow, very acid and moist soil in the montane belt, at elevations between 600 m and 1000 m. Most of the relevés were made on the shady slopes of Jalovnik above the Kneža valley, some of them also on the shady slopes of the Koriška Gora–Obloško Brdo ridge above the Koritnica valley. This is the most acidophilous form of the association Saxifrago cuneifolii-Fagetum. Despite occasionally dystric soil these relevés with their entire species composition cannot be classified into the association Luzulo-Fagetum. In terms of their floristic composition they slightly resemble the stands of the syntaxon Homogyno sylvestris-Fagetum var. geogr. Sesleria autumnalis typicum. With a larger pro­portion of fir also in the tree layer similar stands could be classified also into this syntaxon. The current status of some of the stands of this subassociation may indi­cate a form of degradation of a former fir-beech forest. Table 9 comprises several other relevés that cannot yet be syntaxonomically classified and which grouped separately from other relevés. Relevés 14 and 15 may indicate a special subassociation, -caricetosum albae prov. The differential species are Carex alba and Cor­nus mas. In addition to beech the tree layer consists of mainly hop hornbeam and these two relevés could therefore also indicate a contact with the stands of the association Ostryo-Fagetum, which dominate on the sunny slopes in the study area. 4 CONCLUSIONS Beech forests on steep shady slopes with mixed geo­logical bedrock (dolomite with chert, platy limestone with cherts, marlstone or claystone) and shallow, stony soil in the montane belt of the Upper Soča Valley have so far been classified into the associations Arunco-Fa­getum and Homogyno sylvestris-Fagetum. Having pro­cessed extensive material (more than 200 relevés) we were able to better identify the characteristics of the association Arunco-Fagetum also in western Slovenia. Based on these relevés we were able to typify the new geographical variant Arunco-Fagetum var. geogr. Ane­mone trifolia, which has so far been described with no more than a few sentences and has not been supported with any tabular material. It was divided into two geo­graphical subvariants, the northern-Dinaric (subvar. geogr. Omphalodes verna) and the southern-Alpine (subvar. geogr. Aconitum angustifolium). We described several new subassociations: -caricetosum ferrugineae (the northern edge of the Trnovski Gozd plateau, con­tact with the stands of the association Omphalodo-Fa­getum), -vincetosum minoris (Idrija and Cerkno Hills, the Trebuša Valley, submontane and lower montane belt) and -tilietosum (Tolmin region, submontane belt, dolomite or limestone with cherts, fresh, slightly col­luvial soil). The combination of diagnostic species of the association Arunco-Fagetum is supplemented with Veratrum nigrum and Tamus communis. The stands of this association are managed, frequently with a strong protective role. They are important also as biotopes, sites of some rare species of conservation concern and (or) protected species (Anonymous 2002, 2004) such as Helleborus niger, H. odorus, Erythronium dens-canis, Galanthus nivalis, Lilium martagon, L. carniolicum, Li­stera ovata, Platanthera bifolia, Cephalanthera damaso­nium, C. longifolia, C. rubra, Neottia nidus-avis, Leuco­jum vernum, Epipactis helleborine, E. leptochila, E. mu­elleri, Dactylorhiza fuchsii, Convallaria majalis, Orchis mascula subsp. speciosa, Ruscus aculeatus, Ilex aquifoli­um, Veratrum nigrum, Taxus baccata, Aconitum angu­stifolium, Hemerocallis lilioasphodelus,Primula x terno­vania, Primula carniolica and Leucobryum glaucum. In the numerical processing of our phytosociologi­cal material a large group of relevés clearly grouped separately from the relevés of the association Arunco­-Fagetum. Some of them are floristically quite similar to the stands of the associations Homogyno sylvestris-Fage­tum and Rhododendro hirsuti-Fagetum, whereas some are distinctly different. They indicate a montane beech forest in the elevation belt between (400) 500 and 1300 (1400) m a.s.l., on rather extreme sites, very steep shady slopes on mixed geological bedrock. The bedrock is never pure dolomite or limestone, but always consists also of cherts, claystone or marlstone. The soil is conse­quently shallow, skeletal, but also rather moist and acid. The tree species that frequently accompanies beech on such sites is Sorbus aria, as well as Laburnum alpinum, less frequently also Acer pseudoplatanus, Ostrya carpini­folia, Fraxinus ornus, Sorbus aucuparia, Picea abies, Larix decidua and rarely Abies alba. In the herb layer, Calamagrostis arundinacea stands out with the frequen­cy and the area it covers, and along with it also many other species of beech and spruce forests as well as the species characteristic for thermophilous oak forests of the order Quercetalia pubescenti-petraea, for thermo­philous forest edges of the class Trifolio-Geranietea and for subalpine grasslands of the class Elyno-Seslerietea. Such beech stands are classified into the new associa­tion Saxifrago cuneifolii-Fagetum sylvaticae which we describe as an intrazonal forest community of the southern Julian Alps subject to edaphic factors and land configuration. We described several subassocitions, be­side the typical (-typicum) also subassociations -arunce­tosum (montane form on dolomite with cherts, similar to the stands of the associations Homogyno sylvestris­-Fagetum and Arunco-Fagetum), -fraxinetosum orni, which indicates stands in the elevation belt between 500 m and 1000 m a.s.l., -rhododendretosum hirsuti, which indicates a syndynamic relationship with the stands of the association Rhododendro hirsuti-Fagetum and sub­association -gymnocarpietosum dryopteridis, which is the most acidophilous form of the new association on chert-dominated slopes. Some stands of the association Saxifrago cuneifolii-Fagetum are managed and have a strong protective role, those on very steep slopes in the altimontane belt are exclusively protective and are also important as biotopes, sites for some species of conser­vation concern, of rare or protected vascular plants (Anonymous 2002, 2004) such as Leucobryum glaucum, Cephalanthera damasonium, C. longifolia, C. rubra, Co­rallorhiza trifida, Neottia nidus-avis, Epipactis hellebori­ne, E. leptochila, E. atrorubens, Platanthera bifolia, Dac­tylorhiza fuchsii, Listera ovata, Huperzia selago, Lilium carniolicum, L. martagon, Iris graminea, Asphodelus albus, Arabis pauciflora, Aconitum angustifolium, Taxus baccata, Veratrum nigrum, Primula auricula and Hiera­cium pospichalii. According to the typology of Sloveni­an forest sites (Kutnar et al. 2012) the association Saxi­frago cuneifolii-Fagetum is classified into the group of montane–upper-montane beech stands on calcareous and mixed bedrock as a new forest site type Beech stands with Saxifraga cuneifolia. The areas reported so far are estimated at about 500 hectares. 5 POVZETEK 5.1 Uvod Bukove gozdove na strmih osojnih dolomitnih pobočjih s plitvimi tlemi (rendzino) v severozahodnem delu ilirske florne province (Marinček 1995) uvrščamo v asociacijo Arunco-Fagetum (Košir 1962, 1979, Accetto 2007). Analitski preglednici o tej aso­ciaciji sta objavila Košir (1979) in Accetto (2007, 2015), opise njene vrstne sestave in sestojne zgradbe ter ekoloških razmer pa najdemo tudi v nekaterih knjigah (Marinček 1987: 118–119, Košir 2010: 174–177). Njena rastišča so na veliko poligonih označena na dveh preglednih vegetacijskih kartah Slovenije (Čarni et al. 2002, Košir et al. 2003), prav tako na kartah podrobnejših meril (Marinček et al. 2003, 2006), predvsem v alpskem, predalpskem, dinarskem in pred­dinarskem fitogeografskem območju. Kartirani so tudi v zahodni Sloveniji, v Zgornjem Posočju, na Cerk­ljanskem in Idrijskem. V teh pokrajinah smo bukove sestoje na strmih osojnih dolomitnih in apnenčastih pobočjih začeli popisovati leta 1983 na Cerkljanskem. Na precejšnjih površinah smo jih našli in popisali v Baški dolini (1986–1990), kasneje v okolici Tolmina in v dolini Zadlaščice, redkeje na Kobariškem, zelo redko na Bovškem in pogosto v dolinah Idrijce in Trebušice. V primerjavi z opisi teh rastišč v drugih delih Slovenije je v Zgornjem Posočju očitna razlika v geološki podla­gi in posledično talnih razmerah. Dolomitu je navad­no primešan roženec, ponekod je geološka podlaga ploščasti apnenec s primesjo roženca, laporovca in (ali) glinavca. Silikatna primes se kaže v skeletnosti, globini in kislosti tal in posledično v vrstni sestavi. V skoraj 30 letih smo naredili več kot 200 vegetacijskih popisov in z njihovo obdelavo bomo poskušali odgovoriti tudi na vprašanje, ali preučene bukove gozdove na mešanih karbonatno-silikatnih kamninah še lahko uvrstimo v asociacijo Arunco-Fagetum. 5.2 Metode Fitocenološke popise bukovih gozdov smo naredili po srednjeevropski metodi (Braun-Blanquet 1964). Popise smo vnesli v bazo FloVegSi (Seliškar et al. 2003). Kombinirane ocene zastiranja in pogostnosti smo pretvorili v ordinalne vrednosti od 1 do 9 (van der Maarel 1979). Numerične primerjave smo opravili s programom SYN-TAX 2000 (Podani 2001). Fitocenološke popise smo v analitske preglednice ure­dili na podlagi hierarhične klasifikacije. Uporabili smo metodo kopičenja na podlagi povezovanja (netehtanih) srednjih razdalj – »(Unweighted) average linkage method – UPGMA«. Primerjavo združb v sintezni preglednici smo naredili s to metodo in z metodo min­imalnega porasta vsote kvadratov ostanka – »Incre­mental sum of squarres – MISSQ« – pri obeh smo upo­rabili Wishartov koeficient podobnosti (similarity ratio). Stolpce v sintezni preglednici smo primerjali tudi z ordinacijsko metodo glavnih koordinat (PCoA) in Wishartovim koeficientom podobnosti (similarity ratio) ter z nemetrično ordinacijsko metodo – Non-metric Multidimensional Scaling (NMDS) in količnikom Goodman-Kruskal’s .. Pri teh primerjavah smo upoštevali le praprotnice in semenke, saj vsi av­torji v svojih tabelah nimajo tudi mahovnih vrst. Re­zultate numeričnih metod smo kombinirali s klasično ureditvijo na podlagi diagnostičnih vrst. Nomen­klaturni viri za imena praprotnic in semenk so Mar­tinčič & al. (2007), za imena mahov Martinčič (2003, 2011), za imena lišajev Suppan, Prügger & Mayrhofer (2000), za imena sintaksonov iz zveze Aremonio-Fagion Marinček et al. (1993), za ostale sin­taksone pa Šilc & Čarni (2012), razen za ime razreda Querco-Fagetea Braun-Blanquet et Vlieger in Vlieger 1937, in za imena talnih tipov Urbančič et al. (2005). 5. 2.1 Kratka ekološka oznaka raziskovanega območja Zgornje Posočje je del južnih oz. jugozahodnih Juli­jskih Alp. Geološka podlaga je zelo pisana (Buser 1986, 1987), kar velja še posebej za doline Tolminke, Zadlaščice, Knežice in Bače. V njih prevladujejo tri­asne, jurske in kredne plasti, predvsem baški dolomit z roženci in ploščasti apnenec s primesjo rožencev, laporovca in glinavcev. Tudi v dolinah Idrijce in Trebušice je geološka podlaga pisana (Mlakar & Čar 2009, Čar 2010), vendar je tamkajšnji triasni dolomit navadno brez primesi rožencev. Podnebje je vlažno, s povprečno letno množino padavin več kot 2000 mm (B. Zupančič 1998) in razmeroma toplo, s povprečno letno temperaturo 6 °C–10 °C (Cegnar 1998). Popis­ovali smo osojna pobočja, ki imajo v primerjavi s splošnim precej bolj hladno krajevno podnebje. Goz­dovi v raziskovanem območju so že zelo dolgo načrtno gospodarjeni (Perko et al. 2014). Njihova zdajšnja zas­nova in kakovost je v glavnem posledica močnih sečenj in oglarjenja v prvi tretjini 20. stoletja. Februarja 2014 jih je na precejšnji površini močno poškodoval žled. 5.3 Rezultati in razprava 5.3.1 Numerična primerjava in presoja diagnostičnih vrst Fitocenološke popise bukovih gozdov na strmih oso­jnih dolomitnih in apnenčastih pobočjih v Zgornjem Posočju, na severnem robi Trnovskega gozda in v Cerkljanskem in Idrijskem hribovju smo na podlagi rezultatov hierarhične klasifikacije uredili v več analit­skih preglednic (1 do 10) in večino od njih med seboj primerjali s popisnim gradivom podobnih gozdnih združb (Arunco-Fagetum, Homogyno sylvestris-Fage­tum, Rhododendro hirsuti-Fagetum, Ranunculo plata­nifolii-Fagetum) v zahodni in deloma jugovzhodni Sloveniji. Sintezna preglednica (preglednica 11, glej tudi legendo k slikam 1–4) vsebuje 18 stolpcev. Primer­jani sintaksoni so se združevali, kot kažejo slike 1–4. Rezultati teh primerjav so prinesli naslednje zaključke. Večjo skupina popisov, ki smo jih uredili v dve vegetacijski preglednici (1 in 2, glej tudi stolpce 5 in 6 v preglednici 11) po floristični sestavi nedvomno lahko uvrstimo še v asociacijo Arunco-Fagetum. Ostali popisi so bolj podobni popisom asociacij Homogyno sylve­stris-Fagetum in Rhododendro hirsuti-Fagetum, ki smo jih v zahodni Sloveniji preučili in rezultate objavili pred leti (Dakskobler 2002, 2003). Če v primerjavo vključimo še analizo diagnostičnih vrst (preglednica 12) in analizo deležev po skupinah diagnostičnih vrst (preglednica 13) ugotovimo, da v vseh primerjanih bu­kovih gozdovih uspeva zelo malo vrst, ki bi bile vezane samo na en vegetacijski tip. Po teh primerjavah v aso­ciacijo Arunco-Fagetum v zahodni Sloveniji uvrščamo bukove sestoje v (sub)montanskem pasu (od 200 m do 1000 m nm. v.) na osojnih dolomitnih pobočjih na sev­ernem robu Trnovskega gozda, v Idrijskem in Cerkl­janskem hribovju ter nekatere bukove sestoje na po­dobnih pobočjih in na podobni nadmorski višini v južnih Julijskih Alpah s prigorjem, kjer je geološka podlaga pogosto dolomit z rožencem in ponekod ploščasti apnenec z rožencem. Sestoji v montansko-altimontanskem pasu na mešani geološki podlagi (dolomit z rožencem, lapor­ovcem ali glinavcem, ploščasti apnenec z rožencem, laporovcem ali glinavcem) so floristično precej drugačni. Nekatere od njih smo že okoli leta 1990 kar­tirali kot asociacijo Homogyno sylvestris-Fagetum (karta je bila natisnjena šele veliko pozneje, Daksko­bler 2007a), druge pa na terenskih kartah in v podat­kovni bazi provizorno označevali z imeni Calamagro­stio-Fagetum in Saxifrago cuneifolii-Fagetum. V aso­ciacijo Homogyno sylvestris-Fagetum od leta 1993 (Ma­rinček et al. 1993) uvrščamo predalpsko združbo bukve in jelke, za katero smo prej uporabljali ime Abi­eti-Fagetum prealpinum. Ta združba je razširjena v večjem delu severne Slovenije na karbonatni in mešani karbonatno-silikatni podlagi (Marinček & Čarni 2007). Vrstna sestava njene geografske variante iz dolin Kneže, Zadlaščice in Tolminke (Homogyno sylve­stris-Fagetum var. geogr. Sesleria autumnalis) je dejan­sko zelo podobna vrstni sestavi nekaterih oblik preučenih združb, kar je deloma povezano z istim območjem razširjenosti. Če bi v primerjavo vključili druge geografske variante asociacije Homogyno sylve­stris-Fagetum, bi bila ta podobnost precej manjša. Bist­vena diagnostična vrsta te združbe je jelka (Abies alba), ki pa je v naših popisih zelo redka, prisotna le posamično in v glavnem le v grmovni plasti. Deloma bi to lahko bila posledica preteklega gospodarjenja in degradacijskih procesov. V tej jelovo-bukovi združbi sta tudi večinoma številčnejši in imata večje srednje zastiranje vrsti Adenostyles glabra in Homogyne sylve­stris. Bukove sestoje, ki bolj nakazujejo floristično po­dobnost z jelovim bukovjem, smo vzdružili v pregled­nicah 6 in 10 (glej tudi stolpca 8 in 9 v preglednici 11) in te sestoje bi morda lahko uvrstili tudi v asociacijo Homogyno sylvestris-Fagetum. To ni mogoče za fitioce­noze v preglednicah 7 in 9 (glej tudi stolpce 10, 11 in 12 v preglednici 11), ki se floristično dobro razlikujejo od predalpskega jelovo-bukovja. Nekateri od preučenih sestojev (preglednica 8, popisi 1 do 8, glej tudi stolpec 13 v preglednici 11) so po vrstni sestavi nekoliko po­dobni tudi sestojem asociacije Rhododendro hirsuti-Fa­getum, ki pa navadno uspevajo na še bolj skrajnih rastiščih in imajo fiziognomsko podobo subalpinskega bukovega gozda z obilnim dlakavim slečem (Rhodo­dendron hirsutum) v podrasti. V naših popisih se dlakavi sleč pojavlja le posamično in je zato zgolj raz­likovalnica subasociacije. Poseben edafsko in reliefno pogojen tip bukovega gozda iz zveze Aremonio-Fagion s posebno floristično sestavo (preglednice 6, 7, 8, 9 in 10) zato uvrščamo v novo asociacijo Saxifrago cuneifolii-Fagetum.V njenih sestojih je značilna prevlada gozdne šašulice (Calama­grostis arundinacea) ter prisotnost hladnoljubnih in kisloljubnih vrst, predvsem značilnic smrekovih goz­dov, ob hkratnem uspevanju toploljubnih vrst iz reda Quercetalia pubescenti-petraeae in razreda Trifolio-Ge­ranietea kot tudi vrst suhih in subalpinskih travišč, melišč ter skalnih razpok v zeliščni plasti. Po domi­nantni vrsti zeliščne plasti bi lahko te sestoje uvrstili tudi v asociacijo Calamagrostio arundinaceae-Fagetum Cerovečki 2009, ki jo je Cerovečki (2009) opisal v al­timontanskem pasu zahodne Hrvaške, na karbonatni podlagi in plitvih tleh. To ime je poznejši homonim, saj je bila z enakim imenom v Karpatih prej veljavno opisana asociacija Calamagrostio arundinaceae-Fage­tum Sýkora 1972 (Sýkora 1972, Boublík et al. 2007). Sestoji asociacije iz Karpatov, prav tako sestoji aso­ciacije Calamagriostidi (Abieti)-Fagetum, katere tabelo sta na podlagi Hartmannovih popisov iz v srednjeev­ropskih gorovjih severno od Alp objavila Hartmann & Jahn (1967), so z našimi popisi zelo različni. Določeno podobnost ugotavljamo s fitocenološkimi popisi, ki jih je objavil Cerovečki (ibid.). Obema aso­ciacijama so skupne nekatere vrste zveze Aremonio-Fa­gion, reda Fagetalia sylvatica in razredov Erico-Pinetea, Vaccinio-Piceetea in Mulgedio-Aconitetea. V sestojih obeh primerjanih sintaksonov uspevajo nekatere vrste, ki označujejo hladna, kamnita, nekoliko zakisana rastišča, kot so Calamagrostis arundinacea, Veronica urticifolia, Adenostyles glabra in Homogyne sylvestris. Asociacijo Saxifrago cuneifolii-Fagetum nasproti aso­ciacije Calamagrostio-Fagetum sensu Cerovečki razli­kujejo nekatere diagnostične vrste: Tanacetum corym­bosum, Luzula luzuloides, Saxifraga cuneifolia in Hu­perzia selago, fitogeografske razlikovalnice: Aconitum angustifolium, Larix decidua, Anemone trifolia,Carduus crassifolius, nekatere toploljubne vrste: Sesleria autu­mnalis, Ostrya carpinifolia, Fraxinus ornus, Melittis melissophyllum, Arabis turrita, prav tako vrste skalnih razpok in subalpinskih travišč: Asplenium viride, Se­sleria caerulea subsp. calcaria, Betonica alopecuros, La­serpitium peucedanoides. Med vrstami, ki razlikujejo asociacijo Calamagrostio-Fagetum sensu Cerovečki nasproti asociacije Saxifrago cuneifolii-Fagetum so Ca­lamintha grandiflora, Ribes petraeum in Rosa pimpinel­lifolia. Razlike potrjuje tudi izračun floristične podob­nosti po Sorensenu (1948). Ta je med sintaksonoma Calamagrostio arundinaceae-Fagetum sensu Cerovečki in Saxifrago cuneifolii-Fagetum typicum le 43 %, kar ne dopušča uvrstitve v isto asociacijo. Tako kot smo našli sestoje, ki kažejo na stik oz. podobnost asociacij Homogyno sylvestris-Fagetum in Saxifrago cuneifolii-Fagetum (preglednica 10), in ses­toje, ki kažejo na stik oz. podobnost med sestoji aso­ciacij Rhododendro hirsuti-Fagetum in Saxifrago cune­ifolii-Fagetum (preglednica 8), smo v preglednici 6 (stolpec 8 v preglednici 11) združili popise, ki so po numeričnih primerjavah floristično bolj ali manj po­dobni popisom asociacije Saxifrago cuneifolii-Fagetum in popisom asociacije Homogyno sylvestris-Fagetum, a so v njih dobro zastopane tudi diagnostične vrste aso­ciacije Arunco-Fagetum.Zato jih uvrščamo v posebno subsociacijo Saxifrago cuneifolii-Fagetum arunceto­sum. 5.3.2 Pregled novo opisanih sintaksonov Na podlagi zgornjih premislekov bomo v nadaljevanju opisali ugotovljene sintaksone: Razred: Querco-Fagetea Br.-Bl. et Vlieger in Vlie­ger 1937 Red: Fagetalia sylvaticae Walas 1933 Zveza: Aremonio-Fagion (Ht. 1938) Borhidi in Török, Podani & Borhidi 1989 Asociacija: Arunco-Fagetum Košir 1962 - anemonetosum trifoliae prov. var. Cardamine pentaphyllos var. Gymnocarpium robertianum - caricetosum ferrugineae subss. nov. - vincetosum minoris subass. nov. - tilietosum subass. nov. var. Luzula nivea var. Hacquetia epipactis Asociacija: Saxifrago cuneifolii-Fagetum ass. nov. -typicum subass. nov. var. Calamagrostis varia var. Primula columnae -fraxinetosum orni subass. nov. -aruncetosum dioicae subass. nov. var. Veratrum album var. Hedera helix -gymnocarpietosum dryopteridis subass. nov. -rhododendretosum hirsuti subass. nov. -caricetosum albae prov. -caricetosum humilis prov. Členitev novih oblik asociacije Arunco-Fagetum v fitogeografskem smislu pa je naslednja: Arunco-Fagetum Košir 1962 var. geogr. Anemone trifo­lia Košir ex Dakskobler var. geogr. nov. subvar. geogr. Omphalodes verna subvar. geogr. nov. subvar. geogr. Aconitum angustifolium subvar. geogr. nov. 5.3.3 Asociacija Arunco-Fagetum v Posočju 5.3.31 Fitogeografska členitev asociacije Arunco-Fage­tum v Sloveniji Približne lokacije popisov asociacije Arunco-Fagetum v Posočju so na slikah 5 in 6. Te popise smo uredili v treh preglednicah (preglednice 1, 2 in 3). Nekaj popi­sov iz drugih delov Slovenije pa smo uredili v pregled­nicah 4 in 5. Asociacijo Arunco-Fagetum v Posočju uvrščamo v geografsko varianto var. geogr. Anemone trifolia var. geogr. nova, ki jo je le z nekaj stavki in brez tabelarnega gradiva prvi opisal Košir (1979: 164). No­menklaturni tip (holotypus) nove geografske variante je popis št. 24 v preglednici 2. To je tudi holotip nove geografske subvariante, subvar. geogr. Aconitum an­gustifolium subvar. geogr. nova, v katero združujemo popise iz južnega in zahodnega prigorja Julijskih Alp. Opisali smo še eno novo geografsko subvarianto, sub­var. geogr. Omphalodes verna subvar. geogr. nova. Njen nomenklaturni tip, holotypus, je popis št. 18 v preglednici 1. Njeni sestoji prevladujejo na pobočjih nad dolinama Idrijce in Trebušice, na stiku predalp­skega in dinarskega fitogeografskega območja. Ker so sestoji asociacije Arunco-Fagetum razširjeni tudi v drugih delih predalpskega dela Slovenije, je znotraj geografske variante Anemone trifolia verjetna še kakšna geografska subvarianta. Morda sestoji te geo­grafske variante uspevajo tudi v Zasavju. Popis št. 1 v preglednici 4, narejen na pobočjih Kuma, bi že lahko uvrstili vanjo, ali pač v posebno geografsko subvari­anto var. geogr. Ruscus hypoglossum Košir 1979 sub­var. geogr. Anemone trifolia prov. Morda uspevajo ses­toji asociacije Arunco-Fagetum tudi v subpanonskem delu Slovenije. V preglednici 5 so trije popisi iz oso­jnih pobočij Donačke gore, ki po ekologiji in deloma vrstni sestavi kažejo določeno podobnost s sestoji aso­ciacije Arunco-Fagetum. Po celotni floristični sestavi so v naši primerjavi odstopali od vseh ostalih primer­janih popisov, torej so se združevali v čisto poseben sklop. Odstopajo tudi glede geološke podlage, ki je kremenov konglomerat. Pobočja so zelo strma in zgradba sestojev in reliefne razmere so zelo podobne tistim, ki jih poznamo v sestojih asociacije Arunco-Fa­getum. Za zdaj te tri popise uvrščamo v novo geograf­sko varianto Arunco-Fagetum var. geogr. Hieracium rotundatum. Njen nomenklaturni tip, holotypus, je popis št. 1 v preglednici 5. Fitogeografske razlikoval­nice so tudi vrste Poa stiriaca, Sesleria sadlerana in Galium sylvaticum. Cimperšek (2004: 441) takšne ses­toje na Donački gori uvršča v sintakson Lamio orva­lae-Fagetum var. geogr. Dentaria polyphylla lunarieto­sum redivivae var. Festuca altissima subvar. Aruncus dioicus in omenja fiziognomsko, floristično in ekološko podobnost s sestoji asociacije Arunco-Fage­tum. Marinček in Marinšek (in litt.) sta te sestoje uvrstila v asociacijo Festuco drymejae-Fagetum. Zdaj znana fitogeografska členitev asociacije Arunco-Fagetum je torej naslednja: Arunco-Fagetum Košir 1962 var. geogr. Ruscus hypo­glossum Košir 1979 (Dolenjska) Arunco-Fagetum Košir 1962 var. geogr. Acer obtusatum Accetto 2007 (zgornja Kolpska dolina) Arunco-Fagetum Košir 1962 var. geogr. Calamintha grandiflora Accetto 2015 (Iški vintgar) Arunco-Fagetum Košir 1962 var. geogr. Anemone trifo­lia Košir ex Dakskobler 2015 (alpski, predalpski in deloma dinarski del Slovenije) subvar. geogr. Omphalodes verna (dolina Idrijce s pritokoma Trebušica, Belca) subvar. geogr. Aconitum angustifolium (Zgornje Posočje) Arunco-Fagetum Košir 1962 var. geogr. Hieracium ro­tundatum (Donačka gora) 5.3.32 Arunco-Fagetum var. geogr. Anemone trifolia subvar geogr. Omphalodes verna (preglednica 1) Sestoje te geografske subvariante smo popisali predvs­em na severnem robu Trnovskega gozda nad dolinama Trebušice in Belce, v grapah med Vojskim in Trebušo, v Idrijskem in Cerkljanskem hribovju. Nadmorska višina popisov je od 200 m do 1000 m, geološka podla­ga največkrat dolomit, le ponekod s primesjo roženca ali laporovca, tla v glavnem rendzina. Strmina je od 25° do 45°, lega pa v glavnem osojna, le ponekod tudi prisojna (jugozahodna). Diagnostične vrste asociacije, kot jih je izbral Košir (1962, 1979), so v naših popisih zastopane različno. Med značilnicami sta v popisih te geografske subvariante razmeroma pogosti le vrsti Aruncus dioicus in Helleborus niger, a še zdaleč ne s tako obilnostjo kot na popisnih ploskvah iz območja izvornega opisa asociacije. Kresničje (Aruncus dioicus) vsekakor ni vrsta, ki bi te sestoje posebej označevala oz. v njih posebej izstopala, a podobno je z večino zeliščnih vrst, po katerih poimenujemo bukovja iz zveze Aremonio-Fagion. Med razlikovalnicami sta vrsti Cardamine enneaphyllos in Mercurialis perennis v ilirskih bukovjih splošno razširjeni in ju v tabelah ne izpostavljamo posebej. Podobno lahko zapišemo za vrsto Prenanthes purpurea, ki smo jo v diagnostični kombinaciji obdržali. Na podlagi primerjalne pregled­nice (preglednici 11 in 12) smo med razlikovalnice aso­ciacije Arunco-Fagetum dodatno uvrstili vrsti Tamus communis in Veratrum nigrum. Ti dve vrsti sta sicer bolj značilni za sestoje asociacije Ostryo-Fagetum in dobro označujeta gozdove na dolomitni podlagi in plitvih tleh. Geografske razlikovalne vrste so Anemone trifolia, Omphalodes verna, Phyteuma spicatum subsp. coeruleum, Tephroseris longifolia, Rhamnus fallax, Sco­polia carniolica, Potentilla carniolica in Lathyrus ver­nus subsp. flaccidus. Vrsta Anemone trifolia označuje predvsem bukove gozdove v predalpsko-alpskem delu Slovenije. Vrsta Omphalodes verna je najbolj pogosta v bukovih gozdovih dinarskega dela Slovenije. Takson Phyteuma spicatum subsp. coeruleum je pogost v doli­nah Idrijce in Trebuše in v Trnovskem gozdu, ne poznamo pa ga v Baški dolini in v dolinah severno od Tolmina. Vrste Rhamnus fallax, Scopolia carniolica in Tephroseris longifolia imajo predvsem vzhodnoalpsko-ilirsko razširjenost. Vrsta Potentilla carniolica ima (razen osamljenega nahajališča v Trenti – Trnkoczy, mscr.) najbolj severna nahajališča v Posočju v dolini Idrijce pri Slapu ob Idrijci, njena razširjenost je dinar­ska (Čušin & Dakskobler 2001: 77). Takson Lathyrus vernus subsp. flaccidus je pogost v bukovih gozdovih v Posočju (Dakskobler 1991: 20). Naštete vrste dobro označujejo geografski prostor med Mostom na Soči, Cerknim, Idrijo in Gorenjo Trebušo. Razlikujemo dve subasociaciji. Popise subasociaci­je -caricetosum ferruginei smo naredili na strminah nad Črno drago nad dolino Belce v krajinskem parku Zgornja Idrijca in v Malih Govcih nad povirjem Trebušice. Razlikovalnice subasociacije so vrste Labur­num alpinum, Carex ferruginea in Adenostyles glabra. Te tri vrste označujejo bukove gozdove v montanskem pasu, 800 m do 1000 m nadmorske višine, v območjih, kjer sicer prevladuje jelovo-bukov gozd (Omphalodo­-Fagetum s. lat.). Surina (2002) in Surina & Daksko­bler (2013) sta podobne bukove sestoje uvrstila v sin­takson Omphalodo-Fagetum var. geogr. Saxifraga cu­neifolia calamagrostietosum variae. Dopuščamo možnost, da je odsotnost jelke oz. njena izrazito ma­jhna primes (posamični primerki v različnih sestojnih plasteh) deloma povezana s preteklim gospodarjenjem. Vsekakor sestoji te subasociacije kažejo na stik med asociacijama Arunco-Fagetum in Omphalodo-Fagetum. Nomenklaturni tip, holotypus, nove subasociacije Arunco-Fagetum caricetosum ferruginei je popis št. 7 v preglednici 1. Popise subasociacije -vincetosum minoris smo večinoma naredili v Cerkljanskem hribovju in v dolini Trebuše – tam nekaj popisov tudi v Govcih na sev­ernem robu Trnovskega gozda. Nadmorska višina je nekoliko nižja, prevladujejo popisi na višini med 500 m in 600 m, kar precej smo jih naredili še nižje, v podgorskem pasu. Razlikovalnice subasociacije so vrste Vinca minor, Fraxinus ornus, Ostrya carpinifolia, Vincetoxicum hirundinaria, Veratrum nigrum (tudi diagnostična vrsta za asociacijo) in Campanula rapun­culoides. Razlikovalnice kažejo na razmeroma ugodne toplotne razmere, v katerih kljub strmini in osojni legi uspevajo ti gozdovi. Nomenklaturni tip, holotypus, nove subasociacije je fitocenološki popis št. 18 v preglednici 1. 5.3.33 Arunco-Fagetum var. geogr. Anemone trifolia subvar geogr. Aconitum angustifolium (preglednica 2) Nahajališča sestojev te geografske subvariante so pred­vsem v okolici Tolmina (Bučenica, Kozlov rob, Godiča), nekaj popisov je iz spodnjega dela doline Idrijce oz. iz pobočij nad stranskimi grapami v tej dolini, precej popisov je tudi iz strmih pobočij Matajurja in Mije nad sotesko Nadiže, po en popis pa sta iz osojnih pobočij Stolovega grebena nad Srpenico na Bovškem in iz Čepovanske doline. Diagnostične vrste asociacije so razmeroma redke, kar še posebej velja za vrsto Helle­borus niger. Najbrž je to posledica dejstva, da je geološka podlaga pogosto ploščasti apnenec z rožencem ali baški dolomit z rožencem. Na takih podlagah se črni teloh pojavlja precej redkeje kot na čistem dolo­mitu ali na dolomitiziranem apnencu. Geografske raz­likovalnice Anemone trifolia, Geranium nodosum, Se­sleria autumnalis, Luzula nivea, Aconitum angustifoli­um, Tephroseris pseudocrispa in Lathyrus vernus subsp. flaccidus zelo dobro označujejo gozdove v širši okolici Tolmina oz. v tistem delu Zgornjega Posočja, kjer je v podnebju in rastju še opazen submediteranski vpliv. V bolj alpskem delu Zgornjega Posočja (Bovško) so ses­toji asociacije Arunco-Fagetum redkost, nadomeščajo jih različne oblike asociacije Anemono trifoliae-Fage­tum. Podobno velja tudi za druge dele naših Alp (Košir 1979: 164). Sestoje te geografske subvariante smo členili v dve subasociaciji. Vse popise v preglednici št. 2 uvrščamo v novo subasociacijo Arunco-Fagetum tilietosum. Njene razlikovalnice so vrste Tilia cordata, T. platyphyllos in Phillitis scolopendrium. Naštete vrste kažejo na razm­eroma vlažna, kamnita rastišča, nekoliko koluvialna tla, posredno tudi na geološko podlago, ki za asociacijo Arunco-Fagetum ni tako običajna, namreč ploščasti apnenec z roženci. V tem delu Zgornjega Posočja so na pobočnem grušču in podornem skalovju na vznožju osojnih pobočij pogosta lipovja, ki jih uvrščamo v aso­ciaciji Saxifrago petraeae-Tilietum in Veratro nigri­-Fraxinetum (Dakskobler 2007b) in vsaj ponekod so ta lipovja stična tudi s sestoji asociacije Arunco-Fage­tum. Nomenklaturni tip, holotypus, nove subasociacije Arunco-Fagetum tilietosum je popis št. 24 v preglednici 2. Razlikujemo dve varianti. Varianta Luzula nivea, ra­zlikovalnice so tudi vrste Cardamine pentaphyllos, Mo­ehringia muscosa in Vincetoxicum hirundinaria, označuje rastišča, kjer je pogosto geološka podlaga ap­nenec z rožencem. kamnitost je velika, tla so ponekod bolj razvita – rjava rendzina s prehodi v rjava pokar­bonatna tla. Prevladujejo sestoji na nekoliki višji nad­morski višini in v nekoliko hladnejšem podnebju. Ses­toji te variante kažejo določeno floristično podobnost s sestoji geografske subvariante Lamio orvalae-Fagetum var. geogr. Dentaria pentaphyllos subvar. geogr. Luzula nivea. Sestoji variante z vrsto Hacquetia epipactis in drugimi razlikovalnicami Allium ursinum, Euphorbia amygdaloides, Pulmonaria officinalis in Ruscus aculea­tus prevladujejo na nižji nadmorski višini, v podgor­skem pasu, v nekoliko toplejšem podnebju, na dolo­mitu z rožencem, na manj kamnitih in razmeroma svežih tleh. Razlikovalnice variante kažejo na določeno podobnost oz. stik s sestoji sintaksona Hacquetio-Fage­tum var. geogr. Anemone trifolia. V preglednici 3 smo zbrali popise, ki so se pri numerični primerjavi združevali ločeno od vseh os­talih popisov asociacije Arunco-Fagetum in so floristično nekoliko posebni in tudi med seboj so si precej različni. Naredili smo jih v različnih delih Posočja. Popise št. 1 do 10 začasno uvrščamo v provi­zorno subsociacijo Arunco-Fagetum anemonetosum trifoliae prov. Razlikujemo dve varianti. Var. Cardami­ne pentaphyllos,razlikovalnica je tudi podvrsta Aconi­tum degenii subsp. paniculatum, označuje bolj sveža rastišča, var. Gymnocarpium robertianum, razlikoval­nice so tudi vrste Sesleria caerulea subsp. calcaria, Campanula rapunculoides, Rubus saxatilis, Vincetoxi­cum hirundinaria, Luzula nivea, Aconitum angustifoli­um in Vinca minor pa bolj inicalna, plitva in skeletna tla. V dveh popisih na vznožju Mije smo popisali tudi vrsto Epimedium alpinum. Popisi št. 11 do 13 v pregled­nici 3 ostajajo za zdaj sintaksonomsko neopredeljeni. Popis št. 12 kaže precejšnjo podobnost s sestoji aso­ciacije Saxifrago cuneifolii-Fagetum, ki jo bomo pred­stavili v nadaljevanju. 5.3.4 Opis nove asociacije Saxifrago cuneifolii-Fagetum 5.3.41 Nahajališča in ekološke razmere Približna nahajališča popisov asociacije Saxifrago cu­neifolii-Fagetum so na slikah 7 in 8. Te popise smo ure­dili v petih preglednicah (preglednice 6, 7, 8, 9 in 10, glej tudi stolpce 8 do 13 v preglednici 11). Za zdaj smo njene sestoje našli le v zgornji Baški dolini, predvsem na zelo strmih pobočjih Babjega zoba, Šoštarja in Koble nad potokoma Linderpoh in Kacenpoh, v Trtni­karskem gozdu pri slapu Šprickovbl nad Kacenpohom, pod Robarjevim gričem nad Batavo pri Podbrdu, pod Kojco, na vzpetini Bizle pod Hohkovblom (Matajur­skim vrhom) nad Rutom, pod grebenom Jehle–Kob­lar–Znojilski vrh nad Bideržuno oz. Hudo grapo in nad dolino Koritnice, na strmih pobočjih pod gre­benom Čumik–Luken nad Žventarsko grapo pri Gran­tu, na strmih osojnih pobočjih Jalovnika nad Kneško grapo (Temna brda, Temnak nad domačijo Mohor), pod Tisovcem, pl. Kuk in Krikovim vrhom nad povir­jem Lipovščka prav tako v povodju Kneže (Knežice) in pod Vrhom nad Sopotom nad dolino Zadlaščice. Osa­mljeno nahajališče poznamo na Kobariškem – pod Črnikom nad Drežnico v Krnskem pogorju in eno na Bovškem – Strmi breg na osojnih pobočjih Stolovega grebena nad Srpenico. V precejšnji meri so nahajališča sestojev te asociacije tam, kjer imajo geologi kartirane plasti jurskega mikritnega in kalkarenitnega apnenca s polami roženca in podrejeno glinastega laporovca, kot jih na geološki karti in v Tolmaču označuje Buser (1986, 1987). Prav na vseh popisih so apnencu ali dolo­mitu primešani roženec, glinavec ali laporovec, roženec ponekod povsem prevladuje. Zato je mešana geološka podlaga eden izmed ključnih dejavnikov, ki pogojujejo posebno (svojsko) vrstno sestavo te aso­ciacije. Drugi odločilni dejavnik je velika strmina pobočij (navadno med 30° in 45°) in tretji prevladujoča osojna lega. Višinski razpon uspevanja asociacije je precej velik. Spodnja meja uspevanja je na nadmorski višini okoli (400) 500 m, najvišje pa smo te sestoje našli pod Črno prstjo in Hohkovblom (Matajurskim vrhom) na nadmorski višini skoraj 1400 m, pri čemer prev­ladujejo sestoji na nadmorski višini med 900 m in 1250 m. Sestoji na nižji nadmorski višini so vedno na oso­jnih pobočjih, sestoji v altimontanskem pasu so lahko tudi na prisojnih pobočjih. Talnih razmer podrobneje nismo raziskali. Tla so vedno plitva, skeletna. Prev­ladujejo različne oblike rendzine s prehodi v rjava tla, ki so ponekod distrična. Podnebje je gorsko, zelo hu­midno, s povprečno množino padavin več kot 2000 mm, ponekod celo 3000 mm, in zaradi prevladujoče osojne lege hladno. Razmere za uspevanje gozda so zaradi strmine, kamnitosti in izpostavljenosti vetro­vom precej skrajne, temu primerna je tudi rastnost ses­tojev. Le sestoji v nižjih nadmorskih višinah imajo ponekod večje lesno zaloge in boljšo rastnost. Tam bukev doseže zgornjo drevesno višino več kot 25 m. Sestoji v altimontanskem pasu so nizki, pogosto šopaste in panjevske rasti, z zgornjo drevesno višino med 15 m in 20 m, ponekod tudi manj. 5.3.42 Vrstna sestava, diagnostične vrste in nomen­klaturni tip nove asociacije V drevesni plasti prevladuje bukev (Fagus sylvatica). Najbolj pogosti spremljevalni vrsti sta Sorbus aria in Laburnum alpinum, ponekod tudi Acer pseudoplata­nus, Picea abies, Ostrya carpinifolia, Fraxinus ornus, Sorbus aucuparia, Larix decidua (razlikovalnica nasproti asociaciji Arunco-Fagetum), redko Abies alba in še nekatere druge. Prevladujoče grmovnice so vrste Daphne mezereum, Lonicera alpigena, Rosa pendulina in ponekod Rubus hirtus. Najbolj pogoste vrste zeliščne plasti so Cyclamen purpurascens, Cardamine enneaphyllos, Cardamine tri­folia, Primula vulgaris, Mercurialis perennis, Pre­nanthes purpurea, Dryopteris filix-mas, Salvia glutino­sa, Symphytum tuberosum, Galium laevigatum, Ane­mone nemorosa, Cirsium erisithales, Solidago virgaurea, Gentiana asclepiadea, Oxalis acetosella, Homogyne syl­vestris, Veronica urticifolia, Calamagrostis arundina­cea, Luzula luzuloides, Hieracium murorum, Phyteuma ovatum, Senecio ovatus, Athyrium filix-femina, Ade­nostyles glabra in še nekatere druge. Bogato razvita je mahovna plast, ki ponekod za­stira tudi več kot 20 % popisne površine. Najbolj po­goste vrste mahovne plasti so Ctenidium molluscum, Neckera crispa, Polytrichum formosum, Tortella tortuo­sa, Atrichum undulatum, Isothecium alopecuroides, Me­tzgeria furcata in številne druge. Mahovna plast je pop­isana le delno in naše določitve niso vedno zanesljive. Sestava združbe po skupinah diagnostičnih vrst je v preglednici 13, v stolpcih 8 do 13. V primerjavi s ses­toji asociacije Arunco-Fagetum je v njenih sestojih bist­veno manj mezofilnih vrst značilnih za zvezo Tilio­-Acerion in bukovih vrst iz zveze Aremonio-Fagion, a precej več kisloljubnih vrst iz razreda Vaccinio-Picee­tea. V primerjavi s sestoji asociacije Ranunculo platani­folii-Fagetum je v sestojih nove asociacije bistveno več vrst reda Quercetalia pubescenti-petraeae in vrst raz­reda Elyno-Seslerietea in precej manj vrst visokih steb­lik iz razreda Mulgedio-Aconitetea. Diagnostične vrste nove asociacije so Veronica urticifolia, Calamagrostis arundinacea, Luzula luzuloides, Huperzia selago, Saxi­fraga cuneifolia, Tanacetum corymbosum s. lat. (inc. T. clusii), Campanula witasekiana, Festuca heterophylla, Clinopodium vulgare in Carex humilis. Geografske raz­likovalne vrste so Anemone trifolia, Aconitum angusti­folium, Lathyrus vernus subsp. flaccidus, Larix decidua in Sesleria autumnalis. Naštete vrste označujejo poseb­no ekologijo teh rastišč – zelo strma pobočja v (alti)montanskem pasu, mešano geološko podlago, plitva, a nekoliko zakisana tla v južnih Julijskih Alpah. No­menklaturni tip nove asociacije Saxifrago cuneifolii­-Fagetum ass. nov., holotypus, je popis št. 4 v pregledn­ici 7. Uvrščamo jo v zvezo Aremonio-Fagion, red Fage­talia sylvaticae in razred Querco-Fagetea. Novo aso­ciacijo smo imenovali po vrsti Saxifraga cuneifolia. To je južnoevropska montanska vrsta, značilnica zveze Piceion excelsae in njene podzveze Abieti-Piceenion. Navadno uspeva v montanskem in subalpinskem pasu na kislih, z dušikom bogatih tleh. Razširjena je od Pirenejev preko Alp do vzhodnih Karpatov, tudi v Ap­eninih in v Dinarskem gorstvu (Aeschimann et al. 2004: 692, Accetto 1993, 1995, 2002, 2006, Zupančič & Accetto 1994, Zupančič 1999). Razširjenost v Sloveniji po podatkih v bazi FloVegSi (Seliškar et al. 2003) prikazuje slika 9. Zelo pogosta je v alpskem in predalpskem delu Slovenije, precej redkejša pa v njen­em dinarskem in preddinarskem delu. Raste v številnih gozdnih združbah, predvem bukovih, jelovo-bukovih, smrekovih in macesnovih. Najbolj pogosta je v sestojih asociacij Anemono trifoliae-Fagetum, Rhododendro hir­suti-Fagetum, Ranunculo platanifolii-Fagetum, Polysti­cho lonchitis-Fagetum, Homogyno sylvestris-Fagetum, Adenostylo glabrae-Piceetum, Laburno alpini-Piceetum, Asplenio-Piceetum in Rhodothamno-Laricetum, pone­kod raste tudi v sestojih sintaksonov Saxifrago petrae­ae-Tilietum, Veratro nigri-Fraxinetum excelsioris, Arunco-Fagetum, Lamio orvalae-Fagetum, Omphalo­do-Fagetum var. geogr. Saxifraga cuneifolia, Tanaceto clusii-Fagetum, Luzulo sylvaticae-Piceetum var. geogr. Luzula nivea, Campanulo justinianae-Piceetum, Ribeso alpini-Piceetum, Neckero crispae-Campanuletum justi­nianae in še nekaterih drugih. Označuje hladna, kam­nita rastišča s prhninasto rendzino oz. kislo reakcijo tal v montanskem pasu in je po svoji ekologiji zelo primerna za oznako rastišč nove asociacije. 5.3.43 Nižje sintaksonomske enote asociacije Saxifrago cuneifolii-Fagetum Večino popisov sestojev subasociacije Saxifrago cune­ifolii-Fagetum aruncetosum smo naredili na strminah nad dolino Kneže oz. njenega desnega pritoka Lipovščka, le dva popisa smo naredili na pobočjih nad dolino Bače pri Hudajužni oz. Zakojci. Večino popisov je iz gorskega pasu, na nadmorski višini od 600 m do 1200 m, geološka podlaga pa je v glavnem dolomit z rožencem. Diagnostične vrste asociacije so razmeroma pogoste, z izjemo vrst Carex humilis in Campanula wi­tasekiana. Prisotne so tudi geografske razlikovalne vrste, Anemone trifolia, Sesleria autumnalis, Lathyrus vernus subsp. flaccidus in Aconitum angustifolium. Ra­zlikovalnice subasociacije so vrste Festuca altissima, Aruncus dioicus, Veratrum nigrum in Laserpitium kra­pfii s. str. Še posebej zanimiv je takson Laserpitium krapfii subsp. krapfii, sicer značilen za asociacijo Arun­co-Fagetum. Ima ilirsko-karpatsko razširjenost in v strminah nad grapama Lipovščka in Zadlaščice in nad dolino Bače ima severno mejo svojega celotnega areala in edina nahajališča v Alpah. Drugod v Alpah je pris­otna podvrsta L. krapfii subsp. gaudinii (Aeschimann et al. 2004: 1144, Bačič et al. 2015). Naštete razliko­valne vrste torej kažejo na podobnost s sestoji asociaci­je Arunco-Fagetum. Če bi upoštevali zgolj diagnostične vrste, bi te sestoje lahko uvrstili tudi v to asociacijo. Po celotni floristični sestavi so precej podobni tudi sesto­jem sintaksona Homogyno sylvestris-Fagetum var. geogr. Sesleria autumnalis typicum, torej eni izmed oblik predalpskega jelovega-bukovja, opisani v istem območju. V ta sintakson teh sestojev za zdaj ne more­mo uvrstiti, saj se v njih jelka pojavlja redko in zgolj posamično. Z imenom subasociacije in izborom raz­likovalnih vrst smo pokazali na stik oz. podobnost ses­tojev asociacije Saxifrago cuneifolii-Fagetum s sestoji dveh drugih asociacij, Arunco-Fagetum in Homogyno sylvestris-Fagetum.Nomenklaturni tip, holotypus, nove subasociacije Saxifrago cuneifolii-Fagetum arunceto­sum dioici je popis št. 3 v preglednici 6. Razlikujemo dve varianti. Varianta z vrsto Veratrum album označuje gozdove v zgornjem delu montanskega pasu, na svežih rastiščih. Njeni razlikovalnici sta vrsti Veratrum album in Aconitum degenii subsp. paniculatum. Varianta z vrsto Hedera helix, razlikovalnice so tudi vrste Asarum europaeum subsp. caucasicum, Galium odoratum in Helleborus odorus, označuje rastišča in sestoje v spod­njem montanskem pasu, ki so floristično nekoliko po­dobni tudi sestojem asociacij Lamio orvalae-Fagetum s. lat. in Hacquetio-Fagetum s. lat. Nomenklaturni tip tipične subasociacije Saxifrago cuneifolii-Fagetum typicum je isti, kot je nomenklatur­ni tip nove asociacije, to je popis št. 4 v preglednici 7. V njej združujemo sestoje altimontanskega pasu, na nad­morski višini med 900 m in 1370 m. Vse popise smo naredili na zelo strmih pobočjih nad Baško dolino, v osojni, a nekatare tudi v prisojni legi. Uspevajo v pasu, kjer sicer prevladuje altimontansko bukovje iz aso­ciacije Ranunculo platanifolii-Fagetum. Primerjava z dvema oblikama te asociacije, kot sta ju opisala Ma­rinček & Čarni (2010) v istem fitogeografskem območju, kaže na očitne razlike (primerjaj preglednice 11 in 13, stolpce 11 in 12 ter 17 in 18). Za tipično obliko asociacije Saxifrago cuneifolii-Fagetum so nasproti aso­ciaciji Ranunculo platanifolii-Fagetum razlikovalne vrste reda Quercetalia pubescenti-petraeae in razredov Trifolio-Geranietea in Elyno-Seslerietea. Vse dia­gnos­tične vrste asociacije so v sestojih tipične subasociacije pogoste. Poleg tipične variante, v katero uvrščamo predvsem sestoje na zelo strmih osojnih pobočjih, raz­likujemo še dve varianti. Varianta z vrsto Calamagro­stis varia, razlikovalnice so tudi vrste Carex digitata, Carduus crassifolius in Laserpitium peucedanoides, označuje svetle sestoje na zelo strmih prisojnih in oso­jnih pobočjih s plitvimi tlemi na apnencu ali dolomitu z rožencem, glinavcem ali laporovcem v zgornji Baški dolini. Varianta s taksonom Primula veris subsp. colu­mnae označuje skrajna rastišča, v glavnem prisojna pobočja na apnencu z rožencem ali skoraj čistem rožencu na pobočjih nad srednjo in spodnjo Baško dolino (nad Rutom in pri pl. Kuk). Njene razlikoval­nice so tudi vrste Sesleria autumnalis, Lathyrus vernus subsp. flaccidus in Anemone trifolia. Gozdovi te sub­asociacije so izključno varovalni, saj so to skrajna rastišča za uspevanje gozda. Nomenklaturni tip, holotypus, subasociacije Saxi­frago cuneifolii-Fagetum rhododendretosum hirsuti je popis št. 1 v preglednici 8. Razlikovalnici subasociacije sta vrsti Rhododendron hirsutum in Clematis alpina. Sestoje te subasociacije smo popisali v altimontanskem pasu, na nadmorski višini med 1100 m in 1200 m, večinoma na obeh straneh grebena med dolinama Zadlaščice in grapo Lipovščka. Po celotni floristični sestavi so ti sestoji precej podobni sestojem tukajšnje oblike asociacije Rhododendro hirsuti-Fagetum (Dakskobler 2003). Podrobna primerjava fito­ceno­loških preglednic obeh sintaksonov, še posebej abun­dance (pogostnosti in srednjega zastiranja) diagnos­tičnih vrst, pokaže na očitne razlike. V sestojih aso­ciacije Rhododendro hirsuti-Fagetum ima dlakavi sleč (Rhododendron hirsutum) navadno oceno srednjega zastiranja 1, 2 ali 3, v našem primeru pa le “r” ali “+”. Morda ti popisi kažejo na določeno podobnost med sestoji dveh edafsko in reliefno pogojenih intraconal­nih asociacij, v smislu stopnjevanja skrajnosti rastišč. V preglednico 8 smo uvrstili še nekaj popisov, ki jih sinsistematsko za zdaj ne moremo podrobneje opre­deliti. V numeričnih primerjavah so se združevali povsem ločeno od ostalih popisov. Popisi 13 do 15 v tej preglednici morda označujejo najbolj toploljubno ob­liko asociacije Saxifrago cuneifolii-Fagetum, na strmih prisojnih pobočjih in odprtih grebenih. Začasno jih vrednotimo kot provizorno subasociacijo Saxifrago cu­neifolii-Fagetum caricetosum humilis prov. Glede na skromno zastopanost diagnostičnih vrst obravnavane asociacije ti popisi lahko pripadajo celo novi asociaciji s provizornim imenom Carici humilis-Fagetum prov. Nomenklaturni tip, holotypus, subasociacije Saxi­frago cuneifolii-Fagetum fraxinetosum orni je popis št. 26 v preglednici 9. Razlikovalnica subasociacije je vrsta Fraxinus ornus, ki označuje popise na nekoliko nižji nadmorski višini, v montanskem pasu od 500 m do 1000 m. V njeni vrstni sestavi so prisotne tudi vrste, bolj značilne za podgorski pas, ki jih v sestojih tipične subasociacije nismo popisali, na primer vrste Hedera helix, Lonicera xylosteum, Acer campestre, Vinca minor. V primerjavi z njo so to nekoliko manj skrajna rastišča, lega je skoraj vedno osojna, tla so ponekod bolj razvita, s prehodi v rjava tla. Rastnost gozdov je zato nekoliko boljša in večinoma so to bolj ali manj gospodarski goz­dovi, s poudarjeno varovalno vlogo. V preteklosti so jih ponekod močno sekali. Največje površine sestojev te subasociacije so na osojnih pobočjih nad grapo Ka­cenpoha pri Podbrdu in na osojnih pobočjih nad Žventarsko grapo pri Grantu. Med diagnostičnimi vrstami asociacije so nekoliko redkejše tiste, ki označujejo plitva tla in prhninasto rendzino, na primer vrste Campanula witasekiana, Carex humilis in Clino­podium vulgare. Tudi sestoji te subasociacije so po vid­ezu in celotni floristični sestavi nekoliko podobni ses­tojem asociacije Arunco-Fagetum, a numerična prim­erjava očitno pokaže, da jih vanjo ne moremo uvrstiti. Nomenklaturni tip, holotypus, nove subasociacije Saxifrago cuneifolii-Fagetum gymnocarpietosum dryop­teridis je popis št. 2 v preglednici 10. Razlikovalnice subasociacije so vrste Gymnocarpium dryopteris, Acer pseudoplatanus, Abies alba, Phyteuma ovatum, Actaea spicata, Polystichum aculeatum, Dryopteris affinis in Adoxa moschatellina. Označujejo zelo strma in kam­nita pobočja na prevladujočem rožencu s plitvimi, precej zakisanimi in vlažnimi tlemi v montanskem pasu, na nadmorski višini med 600 m in 1000 m. Večino popisov smo naredili na osojnih pobočjih Jal­ovnika nad dolino Kneže, nekatere tudi na osojnih pobočjih grebena Koriška gora–Obloško brdo nad dolino Koritnice. To je najbolj kisloljubna oblika aso­ciacije Saxifrago cuneifolii-Fagetum.Kljub temu, da so tla ponekod distrična, teh popisov po celotni vrstni sestavi nikakor ne moremo uvrstiti v asociacijo Lu­zulo-Fagetum. Po floristični sestavi so nekoliko podob­ni sestojem sintaksona Homogyno sylvestris-Fagetum var. geogr. Sesleria autumnalis typicum. Ob večjem deležu jelke tudi v drevesni plasti bi bilo mogoče po­dobne sestoje uvrstiti tudi v ta sintakson. Zdajšnje stanje nekaterih sestojev te subasociacije morda kaže na degradacijsko obliko nekdanjega jelovo-bukovega gozda. V preglednici 9 je še nekaj popisov, ki jih za zdaj sintaksonomsko ne moremo opredeliti in so se združevali ločeno od ostalih popisov. Popisa 14 in 15 morda nakazujeta posebno subasociacijo, -caricetosum albae prov. Razlikovalni vrsti sta Carex alba in Cornus mas. V drevesni plasti je ob bukvi precej črnega gabra in morda ta dva popisa kažeta tudi na stik s sestoji aso­ciacije Ostryo-Fagetum, ki v raziskovanem območju prevladujejo na prisojnih pobočjih. 5.4 Zaključki Bukove gozdove na strmih osojnih pobočjih z mešano geološko podlago (dolomit z roženci, ploščasti apnenec z roženci, laporovcem ali glinavcem) in plitvimi, kam­nitimi tlemi v gorskem pasu Zgornjega Posočja smo do zdaj uvrščali v asociaciji Arunco-Fagetum in Homo­gyno sylvestris-Fagetum. Obdelava obsežnega gradiva (več kot 200 fitocenoloških popisov) nam je omogočila boljše prepoznavanje značilnosti asociacije Arunco-Fa­getum tudi v zahodni Sloveniji. Na podlagi teh popisov smo lahko tipizirali novo geografsko varinato Arunco­-Fagetum var. geogr. Anemone trifolia, ki je bila do zdaj opisana zgolj z nekaj stavki in brez podpore tabelarnega gradiva. Členili smo jo na dve geografski subvarianti, severnodinarsko (subvar. geogr. Omphalodes verna) in južnoalpsko (subvar. geogr. Aconitum angustifolium). Opisali smo več novih subasociacij: -caricetosum ferru­gineae (severni rob Trnovskega gozda, stik s sestoji asociacije Omphalodo-Fagetum), -vincetosum minoris (Idrijsko in Cerkljansko hribovje, dolina Trebuše, podgorski in spodnji gorski pas) in -tilietosum (Tol­minsko, podgorski pas, dolomit ali apnenec z roženci, sveža, nekoliko koluvialna tla). Kombinacijo diagnos­tičnih vrst asociacije Arunco-Fagetum dopolnjujemo z vrstama Veratrum nigrum in Tamus communis. Sestoji te asociacije so gospodarski, a pogosto s poudarjeno varovalno vlogo. Imajo tudi biotopski pomen, kot rastišča nekaterih redkih, varstveno pomembnih in (ali) zavarovanih vrst (Anonymous 2002, 2004) kot so Helleborus niger, H. odorus, Erythronium dens-canis, Galanthus nivalis, Lilium martagon, L. carniolicum, Li­stera ovata, Platanthera bifolia, Cephalanthera damaso­nium, C. longifolia, C. rubra, Neottia nidus-avis, Leuco­jum vernum, Epipactis helleborine, E. leptochila, E. mu­elleri, Dactylorhiza fuchsii, Convallaria majalis, Orchis mascula subsp. speciosa, Ruscus aculeatus, Ilex aquifo­lium, Veratrum nigrum, Taxus baccata, Aconitum an­gustifolium, Hemerocallis lilioasphodelus, Primula x ternovania, Primula carniolica in Leucobryum glau­cum. Pri numerični obdelavi našega fitocenološkega gradiva se je večja skupina popisov nedvomno zdru­ževala ločeno od popisov asociacije Arunco-Fagetum. Nekateri med njimi so floristično precej podobni ses­tojem asociacij Homogyno sylvestris-Fagetum in Rho­dodendro hirsuti-Fagetum, nekateri pa od njih očitno različni. Označujejo montanski bukov gozd v višin­skem pasu med (400) 500 m in 1300 (1400) m nm. v., precej skrajnih rastišč, zelo strmih osojnih pobočij na mešani geološki podlagi. Ta ni nikoli čisti dolomit ali apnenec, temveč je v podlagi vedno primes rožencev, glinavca ali laporovca. Posledično so tla plitva, skelet­na, a tudi bolj vlažna in ponekod precej kisla. Drevesna vrsta, ki na takih rastiščih skoraj vedno spremlja bukev, je mokovec (Sorbus aria), prav tako alpski neg­noj (Laburnum alpinum), manj pogosto tudi gorski javor (Acer pseudoplatanus), črni gaber (Ostrya carpi­nifolia), mali jesen (Fraxinus ornus), jerebika (Sorbus aucuparia), smreka (Picea abies), macesen (Larix deci­dua) in redko jelka (Abies alba). V zelišni plasti po po­gostnosti in površini, ki jo zastira, izstopa vrsta Cala­magrostis arundinacea, ob njej pa še številne druge vrste bukovih in smrekovih gozdov ter tudi vrste, značilne za toploljubne hrastove gozdove iz reda Quer­cetalia pubescenti-petraea, za toploljubne gozdne robove iz razreda Trifolio-Geranietea in za subalpinska travišča iz razreda Elyno-Seslerietea.Take bukove ses­toje uvrščamo v novo asociacijo Saxifrago cuneifolii­-Fagetum sylvaticae in jo označujemo kot edafsko in reliefno pogojeno intraconalno gozdno združbo južnih Julijskih Alp. Opisali smo več subasociacij, poleg tipične (-typicum) še subasociacije -aruncetosum (mon­tanska oblika na dolomitu z roženci, nekoliko podobna tudi sestojem asociacij Homogyno sylvestris-Fagetum in Arunco-Fagetum), -fraxinetosum orni, ki označuje sestoje v višinskem pasu med 500 m in 1000 m nm. v., -rhododendretosum hirsuti, ki kaže na sindinamsko povezanost s sestoji asociacije Rhododendro hirsuti-Fa­getum in subasociacijo -gymnocarpietosum dryopteri­dis, ki je najbolj kisloljubna oblika nove asociacije, na pobočjih, kjer prevladuje roženec. Nekateri sestoji aso­ciacije Saxifrago cuneifolii-Fagetum so gospodarski in imajo poudarjeno varovalno vlogo, tisti na zelo strmih pobočjih v altimontanskem pasu pa so izključno var­ovalni, tudi s pomembno biotopsko vlogo, kot življenjski prostor nekaterih varstveno pomembnih, redkih ali zavarovanih praprotnic in semenk (Anonymous 2002, 2004) kot so Leucobryum glaucum, Cephalanthera damasonium, C. longifolia, C. rubra, Corallorhiza trifida, Neottia nidus-avis, Epipactis helle­borine, E. leptochila, E. atrorubens, Platanthera bifolia, Dactylorhiza fuchsii, Listera ovata, Huperzia selago, Li­lium carniolicum, L. martagon, Iris graminea, Asphode­lus albus, Arabis pauciflora, Aconitum angustifolium, Taxus baccata, Veratrum nigrum, Primula auricula in Hieracium pospichalii. Po tipologiji gozdnih rastišč Slovenije (Kutnar et al. 2012) asociacijo Saxifrago cu­neifolii-Fagetum uvrščamo v skupino gorsko-zgornje­gorskih bukovij na karbonatnih in mešanih kamninah kot nov gozdni rastiščni tip Bukovje s klinolistnim kamnokrečem. Njegove zdaj znane površine ocenju­jemo na okoli 500 hektarov. ACKNOWLEDGEMENTS – ZAHVALA Iztok Sajko prepared Figures 6 and 8 for print. Mag. Boško Čušin, Dr. Andrej Rozman and Dr. Branko Vreš helped with the relevés of the stands of the association Arunco-Fagetum in the Dolenjska region and on Donačka Gora. Dr. Aleksander Marinšek provided the so far unpublished phytosociological table of forests on Donačka Gora. I am very grateful to Academician Dr. Mitja Zupančič and Dr. Lado Kutnar for their insights and peer review. Sincere thanks also to Prof. Dr. Jean-Paul Theurillat and Prof. Dr. Dan Gafta for their valu­able help with the validity and use of the names of the syntaxa Calamagrostio arundinacea-Fagetum and Saxifrago cuneifolii-Fagetum and to Dr. Branko Vreš who helped me with the distribution of the taxa Saxi­fraga cuneifolia and Laserpitium krapfii in Slovenia. English translation by Andreja Šalamon Verbič. 6. REFERENCES – LITERATURA Accetto, M., 1993: Mraziščna smrečja (Asplenio-Piceetum R. Kuoch 1954 var geogr. Omphalodes verna var. geogr. nova) v koliševkah Kočevske. Gozdarski vestnik (Ljubljana) 51 (10): 426–445. Accetto, M., 1995: Neckero crispae-Campanuletum justinianae ass. nova v Sloveniji. Razprave 4. razreda SAZU (Ljubljana) 36 (2): 31–48 + 2 preglednice. Accetto, M., 2002: Nova spoznanja o rastlinstvu in rastju Gorjancev. Gozdarski vestnik, (Ljubljana) 60 (4): 192–205. Accetto, M., 2006: Campanulo justinianae-Piceetum abietis var. ass. nova v Dinarskem gorstvu južne Slovenije. 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Razprave 4. razreda SAZU (Ljubljana) 35 (9): 151–175. 1 Institute of Biology, Scientific Research Centre of the Slovenian Academy of Sciences and Arts, Regional unit Tolmin, Brunov drevored 13, SI-5220 Tolmin and Biotechnical Faculty of the University in Ljubljana, Department of Forestry and Renewable Forest Resources, Večna pot 83, 1000 Ljubljana, igor.dakskobler@zrc-sazu.si Figure 1: Dendrogram of (sub)montane-altimontane beech and fir-beech forests of western and southeastern Slovenia (UPGMA, similarity ratio) Slika 1: Dendrogram (sub)montansko-altimontanskih bukovih in jelovo-bukovih gozdov zahodne in jugovzhodne Slovenije (UPGMA, similarity ratio) Figure 2: Dendrogram of (sub)montane-altimontane beech and fir-beech forests of western and southeastern Slovenia (MISSQ, similarity ratio) Slika 2: Dendrogram (sub)montansko-altimontanskih bukovih in jelovo-bukovih gozdov zahodne in jugovzhodne Slovenije (MISSQ, similarity ratio) Figure 3: Two-dimensional scatter diagram of (sub)montane-alti­montane beech and fir-beech forests of western and southeast­ern Slovenia (PCoA, similarity ratio) Slika 3: Dvorazsežni ordinacijski diagram (sub)montansko-altimon­tanskih bukovih in jelovo-bukovih gozdov zahodne in jugovzhodne Slovenije (PCoA, similarity ratio) Figure 4: Two-dimensional scatter diagram of (sub)montane-altimontane beech and fir-beech forests of western and south­eastern Slovenia (NMDS, Goodman-Kruskal’s .) Slika 4: Dvorazsežni ordinacijski diagram (sub)montansko-altimontanskih bukovih in jelovo-bukovih gozdov zahodne in jugovzhodne Slovenije (NMDS, Goodman-Kruskal’s .) Legend to Figures 1–4: ArFov Arunco-Fagetum var. geogr. Anemone trifolia subvar. geogr. Omphalodes verna, this article, Table 1; ArFtil Arunco-Fagetum var. geogr. Anemone trifolia tilietosum, this article, Table 2; ArF Arunco-Fagetum, Dolenjska, this article, Table 4; ScFar Saxifrago cuneifolii-Fagetum aruncetosum, this article, Table 6; ScFluz Saxifrago cuneifolii-Fagetum gymnocarpietosum dryopteridis, this article, Table 10, columns 1–12; ScFfo Saxifrago cuneifolii-Fagetum fraxinetosum orni, this article, Table 9; ScFca Saxifrago cuneifolii-Fagetum typicum var. Calamagrostis varia, this article, Table 7, relevés 1–18; ScFpc Saxifrago cuneifolii-Fagetum typicum var. Primula columnae, this article, Table 7, relevés 19–26; ScFrh Saxifrago cuneifolii-Fagetum rhododendretosum hirsuti, this article, Table 8, relevés 1–8. ArFty Arunco-Fagetum var. geogr Ruscus hypoglossum typicum, Košir 1979, Table 8, columns 1–18; ArFho Arunco-Fagetum var. geogr. Ruscus hypoglossum homogynetosum sylvestris, Košir (1979, Table 8, columns 19–26); ArFao Arunco-Fagetum var. geogr. Acer obtusatum, Accetto (2007, Table 1); HoFty Homogynos sylvestris-Fagetum var. geogr. Sesleria autumnalis typicum, Dakskobler (2002, Phytosocio­logical table 2, relevés 1–31); RF-At Rhododendro hirsuti-Fagetum var. geogr. Anemone trifolia subvar. geogr. Aconitum angustifolium forma Pinus mugo, Dakskobler (2003, Phytosociological table 3); RF-Sa Rhododendro hirsuti-Fagetum var. geogr. Anemone trifolia subvar. geogr. Aconitum angustifolium forma Sesleria autumnalis, Dakskobler (2003, Phytosociological table 4); HoFrh Homogyno sylvestris-Fagetum var. geogr. Sesleria autumnalis rhododendretosum hirsuti, Dakskobler (2002, Phytosociological table 1); RpFho Ranunculo platanifolii-Fagetum var. geogr. typica homogynetosum sylvestris, Marinček & Čarni (2010), Table 14, relevés 1–10); RpFty Ranunculo platanifolii-Fagetum var. geogr. typica typicum, Marinček & Čarni (2010, Table 1, relevés 1–12); Figure 5: Distribution of the association Arunco-Fagetum in the Soča Valley Slika 5: Razširjenost asociacije Arunco-Fagetum v Posočju Figure 6: Approximate localities of relevés of the association Arunco-Fagetum in western Slovenia Slika 6: Približna lokacija popisov asociacije Arunco-Fagetum v zahodni Sloveniji Figure 7: Known distribution of the association Saxifrago cuniefolii-Fagetum Slika 7: Zdaj znana razširjenost asociacije Saxifrago cuniefolii-Fagetum Figure 8: Approximate localities of relevés of the association Saxifrago cuneifolii-Fagetum Slika 8: Približna lokacija popisov asociacije Saxifrago cuniefolii-Fagetum Figure 9: Distribution of Saxifraga cuneifolia in Slovenia Slika 9: Razširjenost vrste Saxifraga cuneifolia v Sloveniji 1 Stand of the syntaxon Arunco-Fagetum vincetosum minoris in the Trebuša Valley 1 Sestoj sintaksona Arunco-Fagetum vincetosum minoris v dolini Trebuše 2 Stand of the syntaxon Arunco-Fagetum tilietosum, Kozlov Rob near Tolmin 2 Sestoj sintaksona Arunco-Fagetum tilietosum, Kozlov rob pri Tolminu 3 Aruncus dioicus 4 Saxifraga cuneifolia 6a Slopes of Šoštar above the Linderpoh gorge, stands of the syntaxon Saxifrago cuneifolii-Fagetum typicum 6a Pobočja Šoštarja nad Linderpohom, sestoji sintaksona Saxifrago cuneifolii-Fagetum typicum 5 Stand of the syntaxon Saxifrago cuneifolii-Fagetum typi­cum, Krikov Vrh 5 Sestoj sintaksona Saxifrago cuneifolii-Fagetum typicum, Krikov vrh 6b Bizle above Rut, stands of the syntaxon Saxifrago cuneifolii-Fagetum typicum 6b Bizle nad Rutom, sestoji sintaksona Saxifrago cuneifolii-Fagetum typicum 7 Stand of the syntaxon Saxifrago cuneifolii-Fagetum fraxinetosum orni, Trtnikarski Gozd near Šprickovbl 7 Sestoj sintaksona Saxifrago cuneifolii-Fagetum fraxinetosum orni, Trtnikarski gozd pri Šprickovblu 8 Stand of the syntaxon Saxifrago cuneifolii-Fagetum aruncetosum, slopes of Krikov Vrh above the Mirna Grapa gorge 8 Sestoj sintaksona Saxifrago cuneifolii-Fagetum aruncetosum, pobočja Krikovega vrha nad Mirno grapo Table 1 (Preglednica 1): Arunco-Fagetum var. geogr. Anemone trifolia subvar geogr. Omphalodes verna Number of relevé (Zaporedna številka popisa) 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 31 32 33 34 35 36 37 38 39 40 41 Database number of relevé (Delovna številka popisa) 217992 217993 217994 217995 217996 217997 241562 241566 218035 218036 218038 218037 229402 247019 247020 247021 229406 234559 234793 246306 238012 246296 246303 246302 235036 235037 235038 247035 247084 255569 238021 246408 245853 246960 246961 217875 234789 246405 246409 246406 246407 Elevation in m (Nadmorska višina v m) 870 900 940 960 980 980 790 810 570 610 580 650 360 540 550 600 530 540 220 260 540 430 400 430 340 400 490 490 500 290 370 360 890 900 930 290 190 290 360 330 350 Aspect (Lega) SW SW SW SW SW SE N NNE NE NE N NE N NE W E N NE N NNW SE NW NNE NE NNW NNW NNW NE E NE N E E NW E NE N W W NE SE Slope in degrees (Nagib v stopinjah) 30 35 40 35 40 35 45 35 35 30 35 40 35 30 35 30 45 35 35 25 25 30 40 35 35 35 35 30 35 30 30 40 45 35 35 35 35 35 35 30 35 Parent material (Matična podlaga) D D D D D D D D D D D D D D D D D D D DR D D D D D D D DL DR D D D D DA DA D Ko D D D D Soil (Tla) Re Re Re Re Re Re Re Re Re Re Re Re Re Re Re Re Re Re Re Re Re Re Re Re Re Re Re Re Re Re Re Re Re Re Re Re Re Re Re Re Re Stoniness in % (Kamnitost v %) 20 30 40 20 20 30 20 20 10 20 20 30 10 20 30 30 10 10 5 10 20 15 20 10 5 5 5 10 10 5 10 20 10 30 10 10 10 10 10 20 20 Cover in % (Zastiranje v %): Upper tree layer (Zgornja drevesna plast) E3b 70 80 80 90 80 90 80 80 80 80 90 80 90 90 80 80 80 90 90 90 80 90 80 90 70 80 80 90 90 90 80 90 80 90 90 70 70 70 80 80 70 Lower tree layer (Spodnja drevesna plast) E3a 20 5 10 5 5 5 . 10 10 20 10 10 10 5 5 10 10 10 5 5 10 5 10 10 20 10 10 10 10 10 10 5 . 10 10 20 30 10 10 10 10 Shrub layer (Grmovna plast) E2 20 30 30 20 20 10 5 5 10 10 10 10 5 5 5 5 5 5 10 20 10 5 5 5 10 5 5 10 10 70 30 10 5 5 5 20 30 60 30 30 30 Herb layer (Zeliščna plast) E1 50 60 60 60 60 70 50 70 30 50 40 50 30 50 60 40 40 40 40 70 50 50 70 50 50 50 40 40 60 20 50 50 20 40 20 60 50 70 80 70 70 Moss layer (Mahovna plast) E0 5 5 5 5 5 5 5 5 5 10 10 10 5 5 5 10 5 5 2 5 5 5 10 10 5 10 5 5 10 5 5 5 5 10 5 10 10 10 5 10 10 Maximum diameter of trees (Največji prsni premer dreves) cm 80 50 70 75 70 50 50 40 45 35 35 40 50 45 45 50 35 50 40 40 35 45 40 40 45 45 40 40 50 35 40 40 35 40 30 35 40 50 45 45 40 Maximum height of trees (Največja drevesna višina) m 32 29 30 30 27 22 27 19 25 25 25 22 28 30 25 34 20 24 28 30 24 25 24 26 26 26 24 22 26 22 25 27 20 20 20 22 28 30 28 28 26 Number of species (Število vrst) 70 65 61 48 41 46 58 54 61 39 41 36 46 48 51 49 52 51 64 61 64 72 61 60 73 49 49 62 53 66 41 54 18 45 32 54 82 63 74 81 76 Relevé area (Velikost popisne ploskve) m2 900 900 900 900 400 400 400 200 400 400 400 200 400 400 400 400 400 400 400 400 400 400 400 400 400 400 400 400 200 400 400 400 100 400 400 200 400 400 400 400 400 Date of taking relevé (Datum popisa) 5/8/2007 5/8/2007 5/8/2007 5/8/2007 5/8/2007 5/8/2007 5/5/2011 5/5/2011 4/20/2007 4/20/2007 4/20/2007 4/20/2007 5/8/2009 5/16/1997 5/16/1997 9/27/1996 5/8/2009 5/19/2004 4/7/2010 6/17/2011 6/28/2010 6/17/2011 6/17/2011 6/17/2011 4/9/2010 4/9/2010 4/9/2010 4/22/2005 4/22/2005 5/1/1993 10/29/2010 4/22/2011 4/18/2012 7/26/2002 7/26/2002 4/19/2007 4/7/2010 4/22/2011 4/22/2011 4/22/2011 4/22/2011 Locality (Nahajališče) Črna draga Črna draga Črna draga Črna draga Črna draga Črna draga Mali Govci Mali Govci Govci - Velika grapa Govci-Velika grapa Govci-Velika grapa Govci-Velika grapa Kamrtna skala Gorenja Trebuša - Govci Gorenja Trebuša - Govci Gorenja Trebuša - Govci Kamrtna skala Govci-Govška grapa Dolenja Trebuša - Na lazu Stopnik - Pri Vojsku Šebrelje - Sv. Ivan Šebrelje - Reka Šebrelje-Vojsk Šebrelje - Kanačnk Reka - Vidršek Reka - Vidršek Reka - Vidršek Police-Snoviška grapa Police-Snoviška grapa Trebuša - Krtovše Reka - Sv. Ivan Sjavnica-Šebrelje Velike Vrše Drnova Drnova Pršjak Dolenja Trebuša - Ravnica Sjavnica - Jelje Sjavnica- Jelje Sjavnica-Šebrelje Sjavnica - Šebrelje Quadrant (Kvadrant) 0049/2 0049/2 0049/2 0049/2 0049/2 0049/2 9949/3 9949/3 9949/3 9949/3 9949/3 9949/3 9849/3 9949/3 9949/3 9949/3 9849/3 9948/4 9849/3 9849/3 9849/4 9849/3 9849/3 9849/3 9849/4 9849/4 9849/4 9849/3 9849/3 9948/2 9849/4 9949/2 9948/2 9850/1 9850/1 9949/1 9949/1 9849/4 9949/2 9849/4 9949/2 Coordinate GK Y (D-48) m 416192 416162 416125 416114 416123 416105 413677 413779 411905 411923 411811 411739 415609 409813 409914 409995 415720 409282 412023 415776 416428 416144 415841 415946 418463 418611 418862 415592 415457 409474 416710 417211 407961 424009 424242 409837 411292 417141 417254 417128 417160 Coordinate GK X (D-48) m 5093506 5093578 5093683 5093731 5093764 5093767 5095806 5095726 5096853 5096818 5096841 5096762 5107010 5098038 5098058 5098001 5106777 5098836 5106855 5108296 5107978 5108152 5108045 5108023 5107705 5107635 5108078 5109015 5109298 5101474 5108235 5106503 5105386 5112366 5112428 5101852 5106631 5106996 5106544 5106822 5106692 Diagnostic species of the association (Diagnostične vrste asociacije) Pr. Fr. AF Helleborus niger E1 . . + 1 . . 1 . + + + . . . . 1 2 1 1 1 2 + 1 1 2 2 1 2 2 + 1 . + . . . 1 + 1 . 26 63 TA Aruncus dioicus E1 + . . . . . + + + . . . . + + + + + . 1 + + . + + . . . . . + . . . . . + . + . + 18 44 FS Prenanthes purpurea E1 1 1 1 1 1 1 + + + . . . . + . + . + . + + . . . . . . . . . . . . + + . . . . . 16 39 Fs Epipactis helleborine E1 . + + + + + . . . . . . . . . + . . . . + + . . . . . . . . . . . . . . . . . . 8 20 QF Spiraea chamaedryfolia E2a . . . . . . . . . . . . . . . . . + . . . . . . 1 + + . . . . . . . . + + . . + + 8 20 VP Rosa pendulina E2a + . + . . . . 1 . . + . . . . . . . . . . . . . . . . . . . . . . r + . . . . + 7 17 QP Tamus communis E1 . . . . . . . . . . . . . . . . . r . . + + . + . . + . . . . . . . . . . . . . 5 12 MuA Phyteuma ovatum E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . . 1 2 Geographical differential species (Geografske razlikovalne vrste) AF Anemone trifolia E1 . . . . . . + . 1 1 1 1 1 1 1 1 1 + 1 1 1 1 + + + + 1 1 1 1 . 2 . . . 1 1 1 2 1 2 30 73 AF Omphalodes verna E1 . + + + + 1 r 1 1 1 1 1 + + + + . + . . . 1 1 . . . . + + 1 . + . + . 1 + + + 2 + 29 71 FS Phyteuma spicatum subsp. coeruleum E1 + + 1 1 1 1 + + . + + + . . . . . . + + + . + + . . . . . + . . . . . . . + + . . 19 46 TA Tephroseris longifolia E1 . . . . . . . . . . . . + . . . . . . . + + + . + + + + + . . + . . . . . . + 1 + 13 32 AF Rhamnus fallax E2 + + + . . + + + . . . + . r . + . + . . . . . . . . . . . . . . . . . . . . . . . 10 24 AF Potentilla carniolica E1 . . . . . . . . . . . . . . . . . . . . . + . . + . + . . . . + . . . . . . r . + 6 15 AF Scopolia carniolica E1 1 + . . . . . . . . . . . . . . . . . . . . . 1 . . . . . . . . . . . . . . . . . 3 7 FS Lathyrus vernus subsp. flaccidus E1 . . . . . . . . . . . . . . . . r . . . . . . . . . . . . . . . . . . . . . . . . 1 2 Differential species of the subassociations (Razlikovalnice subasociacij) FS Laburnum alpinum E3a + . . . + + . + . + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 5 12 FS Laburnum alpinum E2b . + + . . + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + 4 10 FS Laburnum alpinum E2a . . . . . . . + . . + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 2 5 FS Laburnum alpinum E1 + + + + + + + . . + . . . . . . . r . . + . . . . . . . . . . . . . . . . . . . . 10 24 ES Carex ferruginea E1 + + + r . + 1 + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 7 17 TR Adenostyles glabra E1 1 1 1 1 . . 1 + . . . . . . . 1 . . . . . . . . . . . . . . . . . . . . . . . . . 7 17 QF Vinca minor E1 . . . . . . . . + . + . 1 1 . . . 1 1 3 1 . . + 1 1 1 . 1 1 + 2 . 1 1 . 1 2 2 1 2 23 56 QP Fraxinus ornus E3b . . . . . . . + . . 1 . . . . . . . . . . . . . . . . . . . . . . . . 1 + . . . . 4 10 QP Fraxinus ornus E3a . . . . . . . . + . . + r r r . + . . . . . . . . . r . . . . + . . . 2 + . . + + 12 29 QP Fraxinus ornus E2b . . . . . . . + . . . + . . . . . + . r . . . . . . . . . . + + . + . 1 . . . + 1 10 24 QP Fraxinus ornus E2a . . . . . . . . . . . . r . + . . + . + . + + . . . . . . + + . . . . . . + + . + 11 27 QP Fraxinus ornus E1 + + + . . . + + . + . . + + + r + + . . + . + r . . . . . . + + . + + . . . . . . 19 46 QP Ostrya carpinifolia E3b . . . . . . . . + + 2 1 . . + 1 2 . . . + + . r . r r r . . . + . + . 1 r . . . + 18 44 QP Ostrya carpinifolia E3a . . . . . . . . + 1 . + r . . + . . . . + + + + + . . + . + . + . + + + . . . + + 18 44 QP Ostrya carpinifolia E2b r . . . . . . . . . . . . . . . . . . . . . r . . . . . . . . . . . . . . . . . . 2 5 QP Ostrya carpinifolia E2a . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . 1 2 QF Veratrum nigrum E1 . . . . . . . . r 1 1 . + + . . . r + 1 1 1 + + 1 . . + r . . . . . . . + . . . 1 17 42 TG Vincetoxicum hirundinaria E1 . r . . . . . . . . . . r . + r + + . + . + + + . . . + . . . + . . + . . . . . + 14 34 TG Campanula rapunculoides E1 . . . . . . . . . . . . r . . . . . . . + + . . + . . + + . . 1 . r + . . . . . + 10 24 AF Aremonio-Fagion Cyclamen purpurascens E1 1 + + . + + + 1 1 1 1 1 + 1 + 1 1 + 1 + + + + + 1 1 1 1 1 1 1 1 + 1 1 1 + + 1 1 1 40 98 Cardamine enneaphyllos E1 2 1 1 1 1 1 2 2 1 1 1 1 1 1 1 1 1 1 + 1 + 1 1 1 2 2 2 + . + + . 1 1 1 r + + + + 1 39 95 Cardamine trifolia E1 + + + . . . + + 1 + + . + + + + + . + + + + 1 1 + + . . . . . . . . . + + + + + . 26 63 Euphorbia carniolica E1 + + + + . . . . . . . . . . + . . + + + + + + . + . . + + . . . . r . + . 1 1 + + 20 49 Lamium orvala E1 + . + 1 . . . . . . . . 1 + + 1 + . r . + + 2 1 + . . + + + . . . . . . . + 1 . + 20 49 Hacquetia epipactis E1 . . 1 1 . . . . . . . . . . + . . . + 1 1 + + . + . . 1 + + . . . . . . + + + + . 16 39 Knautia drymeia E1 . . . . + + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + + . + . 5 12 Geranium nodosum E1 . . . . . . . . . . . . + . . . . . 1 1 . . . . . . . . . . . . . . . . 1 . . . . 4 10 Daphne laureola E2a . . . . . . . . + + . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . 3 7 Anemone x pittonii E1 . . . . . . + . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . . . . 2 5 Aremonia agrimonoides E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . 1 2 EC Erythronio-Carpinion Primula vulgaris E1 r 1 1 1 1 1 . . 1 1 1 1 + + + + + 1 + . + + . . + 1 1 1 + + + + + + + . + + + + + 35 85 Helleborus odorus E1 . . . . . . . . . . . . . . . . . . 1 . . . . . . . . + . . . . . . . . + + 1 . . 5 12 Erythronium dens-canis E1 . . . . . . . . . . . . . . . . . . . . . + . . 1 1 . 1 . . . . . . . . . . . . . 4 10 Lonicera caprifolium E2a . . . . . . . . . . . . . . . . . . . + . . . . . . . + . . . . . . . . + . . . + 4 10 Lonicera caprifolium E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . 1 2 Galanthus nivalis E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . 1 2 TA Tilio-Acerion Polystichum aculeatum E1 1 + . + . . + + 1 + 1 . + 1 + + 1 1 1 + + + 1 1 1 + . + + + + 1 . + . 2 1 + + 1 1 34 83 Acer pseudoplatanus E3b + 1 + 1 + + . + 1 1 + 1 . . + . 1 + . + . + + . 1 + r 1 1 . . . . + . . r + + + + 28 68 Acer pseudoplatanus E3a r . . + . + . + + + + + . . r + + 1 + . + + + 1 . + r . . . . . r . . . . 1 . . . 21 51 Acer pseudoplatanus E2b . . . . . . . + . . . . . . . + + + . . . . . + . . . . . . . . . + . + + 1 . . . 9 22 Acer pseudoplatanus E2a . . . . . . . + . . . . . . . . + . . . . . + . . . . . . . . . . . . + + 1 + + 1 9 22 Acer pseudoplatanus E1 + 1 1 + 1 1 1 1 + 1 . + . + 1 + + 1 + . + 1 . + + + + . 1 + + + . + + . + + . 1 + 33 81 Ulmus glabra E3b . . . . . . . . . . + . . . . . . . . . r r . 1 + . . . . r . + . . . . . . . + . 8 20 Ulmus glabra E3a . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . . . 1 2 Ulmus glabra E2b . . . . . . . . . . + . . . . . . + . r . . . . . . . . . . + . . . . . . + . 1 . 6 15 Ulmus glabra E2a . . . . . . . . . . . . . . . . . . . . . . . . + + . . . . . . . . . . . 1 . 1 1 5 12 Ulmus glabra E1 + . + . . . r . . . . . . . . . . . . + + + . + + + . . . + . + . . . . . . . 1 . 12 29 Acer platanoides E3b . . . . . . . . . . . . . . . . . . . r r r . . . . . r . . . . . . . . . . . . . 4 10 Acer platanoides E3a . . . . . . . . . . . . . . . . . . . . + . r . . . . . . . . . . . . . . . . . r 3 7 Acer platanoides E2b . . . . . . . . . . . . . . . . . . . . + . . . . . . . . . . + . . . . . + . . + 4 10 Acer platanoides E2a . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + + . . 2 5 Acer platanoides E1 . . . . . . . . . . + . . . . . . . . + + + + . . . . . . . . . . . . . + + . + + 9 22 Euonymus latifolia E2b . . . . . . . + . . . . . . . . . + . . . . . . . . . . . + . . . . . + . . . . . 4 10 Juglans regia E3a . . . . . . . . . . . . . . . . . . . . . . . . . . r . . . . . . . . . . . . . . 1 2 Juglans regia E2b . . . . . . . . . . . . . . . . . . . . . . r . . . . . . . . . . . . . . . . . . 1 2 Juglans regia E2a . . . . . . . . . . . . . . . . . . . . . + . r . . . . . . . + . . . . . . . . + 4 10 Juglans regia E1 . . . . . . . . . . . . . . . . r . . r . . . . . . . . . . + . . . . . . . . . . 3 7 Dryopteris affinis E1 . . . . . . . . . . . . . . . . . . . + . . . . + + . . . . . . . . . . . . + . . 4 10 Phyllitis scolopendrium E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + + + + . 4 10 Thalictrum aquilegiifolium E1 . . . . . . . . . . . . . . . . . . . . . + . + . . . . . . . . . . . . . . . . . 2 5 Geranium robertianum E1 . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . . . . . + . . . 2 5 Polystichum setiferum E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 + . . 2 5 Lunaria rediviva E1 r . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 2 Staphylea pinnata E2a . . . . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . . . . . . . . 1 2 Tephroseris pseudocrispa E1 . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . 1 2 Polystichum braunii E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . 1 2 FS Fagetalia sylvaticae Fagus sylvatica E3b 4 4 5 5 4 4 5 5 4 5 4 4 5 5 5 5 4 5 5 5 5 5 5 5 4 5 5 5 5 5 5 5 5 5 5 4 3 4 5 4 4 41 100 Fagus sylvatica E3a 2 1 + 1 1 1 + 1 1 . 1 1 + + + + + 1 . + 1 + + + 2 + + 1 + 1 + 1 . 1 1 . 1 1 + 1 + 37 90 Fagus sylvatica E2b 2 2 3 2 2 1 . + 1 1 + 1 + . . . + + . + 1 + + . 1 . + 1 . 1 1 + . . + . + 1 1 1 1 30 73 Fagus sylvatica E2a 1 1 1 + + . + + . . . . . . + + . . . 2 . + . . 1 . . . 1 . + . . . . + + 3 2 2 2 20 49 Fagus sylvatica E1 + 1 1 1 1 1 1 + + + + + . + + + + 1 + 1 1 + + . . + + 1 1 1 1 1 . . + . . . 2 1 . 32 78 Mercurialis perennis E1 1 1 1 1 1 1 1 1 1 1 1 1 + + + 1 . + 2 1 1 1 2 1 1 1 1 + + 1 + + 1 2 1 1 . 1 1 1 + 39 95 Daphne mezereum E2a + 1 1 1 1 . 1 + + . + + + + 1 + + 1 + + + + + + + + + + + 1 . + r + + 1 + + + + + 38 93 Galium laevigatum E1 1 1 + 1 1 1 + + + + 1 + + + + + + + . 1 1 1 1 + 1 + . + + + . + . + + 1 + + 1 1 + 37 90 Galeobdolon flavidum E1 1 1 1 1 + . + + 1 1 1 + + + + 1 + . + 1 + 1 1 1 1 . . + + + 1 + . . . 1 + 1 + 1 1 34 83 Salvia glutinosa E1 + 1 1 1 + . 1 . 1 + 1 . + + + 1 + + 1 1 + . + . . + . + + . + 1 r + r . + + + + + 32 78 Asarum europaeum subsp. caucasicum E1 . . . . . . . . 1 1 + . + 1 1 1 1 + + 1 1 1 1 1 + + . 1 . . 1 1 . . + 1 + 1 1 1 1 27 66 Dryopteris filix-mas E1 + + + + r . . . + . . . + + + + + + . + + + + 1 + + + . . + + . . + . . . + + + + 27 66 Euphorbia amygdaloides E1 . + 1 1 . . + + + + + + + . + . + + + . + + . . 1 + + + . + + 1 . . . + + . . . + 26 63 Actaea spicata E1 + + . . . . 1 + + . + . . . . . + + + + + + + 1 + 1 . . . . . . . + . . . . . + + 19 46 Lathyrus vernus subsp. vernus E1 . . . . 1 + . . . . . . . . . . + . + + + + + + + 1 + . . . . . + . 1 + + 1 1 + . 19 46 Neottia nidus-avis E1 . . + + + + . . . . . + + + + + + + . . . + + + . . . . . . + + . . + . . . . . + 18 44 Symphytum tuberosum E1 . + + . . . . . . . . . + + + . . . + . + + + + + + . . r + . . . . . + + . 1 + . 18 44 Mycelis muralis E1 + + + . . . 1 . . . . + + . + + + + . . . + + + . . + . . . + . . . + . . . + . . 17 41 Lonicera alpigena E2 1 + + + + 1 + + . + . + . . 1 + 2 . . . . . . . . . . . . . . . . + + . . . . . . 15 37 Melica nutans E1 + + . + . . . . . . + + . . . . . . . . + . + . . . . + + + . . . . . . + + + + + 15 37 Polygonatum multiflorum E1 + + + . + + . + . . . . . . . . + . + . + . . + . . . . . . . . + + + + . . . . . 14 34 Fraxinus excelsior E3b . . r r . . . . . . . . . . . . . . r + . . . . . . . r + . . . . . . . r . . . . 7 17 Fraxinus excelsior E3a r . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 2 Fraxinus excelsior E2b + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . 2 5 Fraxinus excelsior E2a . + . . + . . . . . . . . . . . . . . . . . + . + . . . . + . + . . . . . + . . . 7 17 Fraxinus excelsior E1 1 + 1 1 1 1 + . . . . . . . . . . . . . . . . . + . . + + . . . . . . . . . + + + 13 32 Lilium martagon E1 . . . + . . + + . . + + . + . . . . + . r + . . . . . + + . . + . . . . . . . . . 12 29 Paris quadrifolia E1 + . . . . . . + . . . . . . . . . . + + 1 + + + . . . . . . . . . + . . + . . + . 11 27 Viola reichenbachiana E1 . . . . . . . . . + . . . . . . . . + . . . . . + + 1 . . . . . . . . . + + + + . 9 22 Pulmonaria officinalis E1 . . . . . . . . . . . . . . . . . . . . . . . . + . . . . + . . . . . . + + + + + 7 17 Campanula trachelium E1 . . . . . . . . . . . . . . . . . . + . . . . . + + . + . . . . . . + . . . + . . 6 15 Heracleum sphondylium E1 + . . . . . . . . . . . . . . . . . + . + . . . + . . . r + . . . . . . . . . . . 6 15 Prunus avium E3b . . . . . . . . . . . . . . . . . . . . r r . . . . r r . . . . . . . . + . + . . 6 15 Prunus avium E3a . . . . . . . . . . . . . . . . . . + . . . . . . . . + . . . . . . . . . . . . . 2 5 Prunus avium E2b . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + + . . . . . . . . . 2 5 Prunus avium E2a . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . . . . + . . . . 2 5 Prunus avium E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . . 1 2 Carpinus betulus E3b . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 2 . . . . 1 2 Carpinus betulus E3a . . . . . . . . . . . . . . . . . . . . . . . . . . . . r . . . . . . . 1 + + . + 5 12 Carpinus betulus E2b . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . . . . 1 2 Carpinus betulus E2a . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . 1 2 Carpinus betulus E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + 1 2 Sambucus nigra E3a . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . . 1 2 Sambucus nigra E2b . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . 1 . 2 5 Sambucus nigra E2a + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . + . + + . 5 12 Brachypodium sylvaticum E1 . . . . . . . . + . . . . . . . . . . + . . . . + . . . . . + . . . . . . . . . . 4 10 Galium odoratum E1 + . . . . . . . . . . . . . . . . . + + . . . . . . . . . . . . . . . . . 1 . . . 4 10 Euphorbia dulcis E1 . . . . . . . . . . . . . . . . . . + . . . . . + . . . . . . . . . . . + . . . . 3 7 Petasites albus E1 + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 2 Cardamine bulbifera E1 + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 2 Sanicula europaea E1 . . + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 2 Festuca altissima E1 . . . . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . . . . . . . . 1 2 Allium ursinum E1 . . . . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . . . . . . . . 1 2 Cardamine pentaphyllos E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . r . . 1 2 QP Quercetalia pubescenti-petraeae Melittis melissophyllum E1 + + . + + + . . . . . . . . . . . + . . + + . . . . + . . . . + . + . . . . . . + 12 29 Euonymus verrucosa E2a r . . . . . . . + . . . . . . . . . . . . + . . . . . . . + . . . . . + + . + + . 8 20 Cornus mas E2b . . . . . . . . . . . . r . . . r . . . . . . . . . . . . . 1 . . . . + + . . . + 6 15 Cornus mas E2a . . . . . . . . . . . . . . . . . . . . . . . . . . r . . + + + . . . . . . + . + 6 15 Sorbus aria E3b . . . . . . . + . . . . . . . . . . . . . . . . . . . r . . . . . . . . . . . . . 2 5 Sorbus aria E3a . . . . . + . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . . 2 5 Sorbus aria E2b . . . . . . . . . . . . . . . . . . . . . r . . . . . . . . + . . . . . . . . . r 3 7 Sorbus aria E2a + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 2 Sorbus aria E1 + . + . + + . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . . . 5 12 Mercurialis ovata E1 . . . . . . . . . . . . . . . . . . . . . + . . . . . + . . . r . . . . . . . . + 4 10 Aristolochia lutea E1 . . . . . . . . . . . . . . . . . . . . . + . . + . . . . . . . . . . . . + . . . 3 7 Asparagus tenuifolius E1 . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . + . . . . . . . . . . 2 5 Orchis mascula subsp. speciosa E1 . . . . . . . . . . . . . . . . . . . . . . . . . . r . . . . . . . . . . . . . . 1 2 Sesleria autumnalis E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . . . . 1 2 Hypericum montanum E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . 1 2 Calamintha sylvatica E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + 1 2 QR Quercetalia roboris Rubus hirtus E2a . . . . . . . . + . . . + . . . + . + + . . + . . . . . . . . . . . . 1 . 1 + + + 11 27 Pteridium aquilinum E1 . . . . . . . . . . . . . . . . . 1 . . + + . . . . . . . . + 1 . . . . . . + . . 6 15 Quercus petraea E3b . . . . . . . . . . . . . . . . . . r . . . . . . . . + . . . . . . . . . . . . . 2 5 QF Querco-Fagetea Hepatica nobilis E1 + + . . . . + 1 + . + . . + 1 + . . . . + . . + . . 1 1 1 + . . . 1 + 1 1 + 1 + . 22 54 Carex digitata E1 . + + . + + + + . . . + . . . . . . + . . . . . + . . 1 + + + 1 . . . + + + 1 + . 19 46 Clematis vitalba E2 + + . . . . . . . . . + . . + . . . . . + + + + . + . . . . + + . . . + . + + + + 16 39 Lonicera xylosteum E2a . . . . . . . . . . . . + . . . . . + + . . + + + . + + + + + . . . . + + . + + + 16 39 Anemone nemorosa E1 + + + + . . + + . . . . . . . . . . + + . . . + . . . . + + . . . 1 . . + + . . . 14 34 Aegopodium podagraria E1 . . . . . . . . . . . . . . . . . . + . . . . + 1 . 1 + + + . . . . . + 1 1 + . . 11 27 Hedera helix E3a . . . . . . . . + . . . . . . . . . . . . . . . . + . + + + . . . . . . . + + . . 7 17 Hedera helix E1 . . . . . . . . + . + . . . . . . . + . . . . . . . . + + . . 1 . . . . 1 + 1 + . 10 24 Corylus avellana E3a . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 . . . . 1 2 Corylus avellana E2b + + . . . . . . + . . . . . . . + . . . . . . . . . . . . . . + . . . . 1 + . + 2 9 22 Corylus avellana E2a + . . . . . . . . . + . . . . . . . . . . . . . . . . . . + . . . . . + . + . . . 5 12 Corylus avellana E1 . . . . . . . . . . . . . r . . . . . . + + . . . . . . . . . . . . . . . . . . . 3 7 Ilex aquifolium E3a . . . . . . . . . 1 . . . . . . . . . . . . . + . . . . . . . . . . . . . . . . . 2 5 Ilex aquifolium E2 . . . . . . . . r + . . . . . . . + . r + . . + . . . . . . 1 . . . . . + . . . . 8 20 Acer campestre E3b . . . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . . . + . . + . . 3 7 Acer campestre E3a . . . . . . . . . . . . . . . . . . . . . . . . . . r + + . . + . . . . . + 1 + . 7 17 Acer campestre E2b . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . 1 + . . 3 7 Acer campestre E2a . . . . . . . . . . . . . . . . . . . . . . . . . . + . . + + . . . . . . 1 1 . + 6 15 Acer campestre E1 . . . . . . . . . . . . . . . . . . + . . . . . . . . . . . . + . . . . . + 1 . + 5 12 Rosa arvensis E2a . . . . . . . . . . . . . . . . . . . . . . + . + . + . + . . + . . . . . + . . . 6 15 Listera ovata E1 . . . . . . . . . . . . . . . . . . + . . . . . + . . + . . . . . . . . + . . . + 5 12 Taxus baccata E3b . . . . . . . . . . + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 2 Taxus baccata E3a . . . . . r . . . . 1 . . r . . . . . . . . . . . . . . . . . . . . . . . . r . + 5 12 Taxus baccata E2b . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 1 2 Taxus baccata E2a . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . . . . 1 2 Taxus baccata E1 . . . . . . . . r . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 2 Viburnum opulus E2a . . . . . . . . . . . . . . . . . . . . . . . . . . . . + + . . . . . . + . . + . 4 10 Cephalanthera longifolia E1 . + . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . . . . . . . . . . 2 5 Viola riviniana E1 . . . . . . . . + . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . 2 5 Viola mirabilis E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . r . . . . . . . . . . . + 2 5 Rubus caesius E2a . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . + 2 5 Platanthera bifolia E1 r . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 2 Carex umbrosa E1 . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . . . 1 2 Malus sylvestris E3a . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + 1 2 Malus sylvestris E2b . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . 1 2 Malus sylvestris E2a . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . 1 2 Anemone ranunculoides E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . . . . . 1 2 Moehringia trinervia E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . . 1 2 Equisetum hyemale E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 3 . . . . 1 2 Scilla bifolia E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 . . . 1 2 EP Erico-Pinetea Carex alba E1 . + 1 . . 2 . 2 1 1 + 2 3 + + + 1 . 1 + 1 1 + + + + 1 1 1 1 + 2 . 1 + 3 1 + 1 2 2 35 85 Cirsium erisithales E1 + 1 1 + 1 1 . + + . + + . . 1 + + + + + + + 1 + + + . . . . . + . + + + + . . + + 29 71 Calamagrostis varia E1 + 1 1 + 1 + + . r . . + . + r + + . + . . + + . . . + . . . . 1 . . + + . . . + + 22 54 Rubus saxatilis E1 . . . . . . . + . . . . . . + + . . . . . . + . . . . . + + . . . . . . . . . + + 8 20 Aquilegia nigricans E1 . . . . . . + . . . . . . . . + + . . . . + . + + + . . . . . . . . . . . . . . . 7 17 Buphthalmum salicifolium E1 . + + . . + . + . . . . . . . . . r . . . . . . . . . . . . . . . . . . . . . . . 5 12 Erica carnea E1 . . . . . . . r . . . . . . . . . . . . . . . . . . . . . . . . . . . + r . . + + 5 12 Rhododendron hirsutum E2a . . . . . . . r . . . . r . . . + . . . . . . . . . . . . . . . . . . + . . . . . 4 10 Epipactis atrorubens E1 . . . . . . . . . . . . . . r . . . . . . . . . . . . . . . . . . . . . . . . . . 1 2 Molinia caerulea subsp. arundinacea E1 . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . . . . . . 1 2 Peucedanum austriacum E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + 1 2 VP Vaccinio-Piceetea Aposeris foetida E1 + 1 1 1 1 1 . . . . . + + + + . . . + + + + + . + + + + 1 1 + . . + . . + 1 1 + + 28 68 Solidago virgaurea E1 + + + + + + . . + + . . . r + . + + + + . . + + + . . + + + + + . . . . . + + + + 26 63 Valeriana tripteris E1 + . . + + 1 + 1 + . . + + + r + . . + . . . 1 . + . + + . + . . + + . + + . + 1 + 25 61 Veronica urticifolia E1 + . + 1 1 1 + . + + . . + + + . + + . + + . + + + . . . . + . . . + + . + . . . . 22 54 Homogyne sylvestris E1 . . . . + + + + 1 1 1 + 1 + + + + 1 . . . . . . . . . . . + . . + 1 . 1 . . . + . 19 46 Oxalis acetosella E1 . . . . . . + . + 1 + . + 1 . . . . + + + . . + + . . . . . + . . . . 1 1 + + + . 17 42 Gentiana asclepiadea E1 . + + + . + . . . . . . . . . . + + . . + . + . . . . . . . . . . . . . . . . . . 8 20 Hieracium murorum E1 . + + . . + + + . . . . . . . . + . . . . . . . . . . . . . . . + . . . . . . . . 7 17 Picea abies E3b . r . . . . . . 1 + 1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . 5 12 Picea abies E3a . . . . . . . . 1 . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . . 2 5 Picea abies E2b . . . . . . . + . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . + . 3 7 Picea abies E2a r r . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . . r . . 4 10 Picea abies E1 . . . . . + r + . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . + . 5 12 Abies alba E3b . r r . . . . . + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 3 7 Abies alba E3a r . . r . . . . . r . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 3 7 Abies alba E2b . . . . . . . . r . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 2 Abies alba E1 . + + . . . r . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 3 7 Vaccinium myrtillus E1 . . . . . + . + . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . 3 7 Maianthemum bifolium E1 . . . . . . . + + . . . . + . . . . . . . . . . . . . . . . . . . . . . . . . . . 3 7 Saxifraga cuneifolia E1 . . . . . . . . . . . . . . . . + . . + . . . . . . . . . . . . . . . . . . . + . 3 7 Clematis alpina E2 . . . . . . r . . . . . . . . r . . . . . . . . . . . . . . . . . . . . . . . . . 2 5 Gymnocarpium dryopteris E1 . . . . . . . + + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 2 5 Lonicera nigra E2a r . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 2 Calamagrostis arundinacea E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . r . . . . . . . 1 2 Phegopteris connectilis E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . 1 2 SSC Sambuco-Salicion capreae Sorbus aucuparia E1 + . . . . + . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . 3 7 RP Rhamno-Prunetea Crataegus monogyna E3a . . . . . . . . . . . . . . . . . . . + . . . . . . . . . + . . . . . . . . . . . 2 5 Crataegus monogyna E2a . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . 1 2 Crataegus monogyna E2b . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . + + 3 7 Crataegus monogyna E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . . . . . 1 2 Berberis vulgaris E2b . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . 1 2 Berberis vulgaris E2a . . . . . . . . . . . . . . . . . . . . . . . . + . r . . . . . . . . . + . . . . 3 7 Cornus sanguinea E2b . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + + . . . 2 5 Cornus sanguinea E2a . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . + . . + . 3 7 Cornus sanguinea E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + 1 2 Euonymus europaea E2a . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . . + . . 2 5 Viburnum lantana E2a . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . + 2 5 Ligustrum vulgare E2a . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . 1 2 TG Trifolio-Geranietea Digitalis grandiflora E1 . . . . . . . . . . . . . . . . + . . . . . . . + + . . . . . + . . . + . . . . . 5 12 Lilium carniolicum E1 . . . . . . . . . . . . . . . . . . . . . + . . . . + . . . . . . . . . . . . . + 3 7 Clinopodium vulgare E1 . + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 2 Viola hirta E1 . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . . 1 2 Laserpitium latifolium E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . . 1 2 Verbascum lanatum E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . 1 2 EA Epilobietea angustifolii Eupatorium cannabinum E1 + + + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . 4 10 Fragaria vesca E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . . . . . 1 2 BA Betulo-Alnetea viridis Salix glabra E2a . . . . . . . + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 2 MuA Mulgedio-Aconitetea Senecio ovatus E1 + . 1 1 1 . 1 . + + . . + + + + + + + 1 + 1 1 + + . + + + + . + . + . . 1 + + + + 31 76 Aconitum lycoctonum s. lat. E1 + 1 . 1 1 . . . . . . + . + 1 2 + + 2 + 1 1 + + + + . + r . . . . . + . + . . . . 22 58 Athyrium filix-femina E1 + + . . . . . . + . + . . + r . . + . + . + . + . . . + . + + . . . . . . . + + . 15 37 Aconitum degenii subsp. paniculatum E1 . + . . . . . . . . . . . . . . . . + . + + 1 1 1 + . + . . . . . . . . . . . . . 9 22 Polygonatum verticillatum E1 1 + + + + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 5 12 Veratrum album s. lat. E1 . . . . . . . . . . . . . . . . . . . . + + 1 . . . . . . . + . . . . . . . . . . 4 10 Centaurea montana E1 . + + . . + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 3 7 Chaerophyllum hirsutum E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + 1 2 ES Elyno-Seslerietea Betonica alopecuros E1 . . . . . . . + . . . . . . r . . . . . . + . . . . . . . . . . . . . . . . . . . 3 7 Sesleria caerulea subsp. calcaria E1 . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . . . 1 + 3 7 Aster bellidiastrum E1 . . r . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 2 Laserpitium peucedanoides E1 . . . . . . . + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 2 FB Festuco-Brometea Carex humilis E1 . + . . . . r . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 2 5 MA Molinio-Arrhenatheretea Angelica sylvestris E1 + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . + 3 7 Cirsium oleraceum E1 . . . . . . + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 2 TR Thlaspietea rotundifolii Gymnocarpium robertianum E1 + + . . . . + + . . . . . + + 1 + r . . + . + + . . . . . + . . . . . + . . . + + 16 39 Astrantia carniolica E1 . . . . . . + . . . . . . . . . . . . . . . . . r . . . . . . . . . . . . . . + . 3 7 Hieracium bifidum E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . r . . . . 1 2 AT Asplenietea trichomanis Asplenium viride E1 + + r . . . + + + . . + . + r + + . . + . + + + + + + + + . . . . . . + . . . . . 21 51 Asplenium trichomanes E1 + + . + . . + . + . . + + . . + . . + . . . + + . + + + . . . . . . . . + + + + . 18 44 Asplenium ruta-muraria E1 . . + . . + . . + . . . . . . + . . + + . + . + . . . + + . . . . . . . . . + + . 12 29 Moehringia muscosa E1 . . . . . . . . . . . . + . . . . . . + . . + + . + . + . . . . . . . + + . + + + 11 27 Paederota lutea E1 . r r + + 1 + + . . . . . r . r . . . . . . . + . . . . . . . . . . . . . . . . . 10 24 Polypodium vulgare E1 . . . . . . . . + . . . . . . . . . + + . . + . . . + . . . . + . . . . r . + + . 9 22 Phyteuma scheuchzeri subsp. columnae E1 . . + . . . + . . . . . . + . + . . . . . . . . . . . . . . . . r . . . . . . + . 6 15 Cystopteris fragilis E1 . . . + . . + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 2 5 Carex brachystachys E1 . . . . . . . . r . . . . . r . . . . . . . . . . . . . . . . . . . . . . . . . . 2 5 Primula carniolica E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . 1 2 ML Mosses and lichens (Mahovi in lišaji) Ctenidium molluscum E0 . 1 + . + + + 1 1 1 1 1 1 1 1 1 1 1 + + + 1 + 1 + + + 1 + 1 + 1 + + . 2 1 1 1 1 1 38 93 Neckera crispa E0 + . + . + + . . + 1 . 1 1 + + 1 + + + + + + + 1 + + + 1 + + + + + + . + 1 + + 1 + 35 85 Fissidens dubius E0 + + + + + + + . + + . . + . + 1 . . + . + + . . + + + + . + . . . + . . + . . + . 23 56 Polytrichum formosum E0 . . . + . + . + + + . . . . . . + + + . . + . . + + + . + + + + . + + + . . . . . 19 46 Plagiochila porelloides E0 . . . . . . + . + + . . + + . . . 1 . . . . . . + + . + . + . . . + . . + . . + . 13 32 Tortella tortuosa E0 . . . + . + + + . . . + . . . . . . . . . . . . + + + . . + . . . + . + . . . + . 12 29 Isothecium alopecuroides E0 . . . . . . . . . . + . 1 . . . . . + + . + . + + . + . + + . . . . . . . . . . . 10 24 Schistidium apocarpum E0 + + . + + . . . . + . . . . + . . . . . + + . . . . . . . . . . . . . . . . . . . 8 20 Encalypta streptocarpa E0 . . + . + . . . . . . . . . . . . . . . . . . . . + . + . . . . . . . . . . . . . 4 10 Peltigera canina E0 . . . . . . . . + . . . . . . . . . . . . . . . + + . . . . . . . . . . . . . . . 3 7 Bartramia halleriana E0 . . . . . . . . . . . . . . . . + + . . . . . . . . . . . 1 . . . . . . . . . . . 3 7 Marchantia polymorpha E0 . . . . . . . . . . . . . . . . . . . . . . + . + . . . . . . . . . . . + . . . . 3 7 Eurhynchium angustirete E0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . + . . . + . 3 7 Rhytidiadelphus triquetrus E0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + + . . + . 3 7 Mnium thomsonii E0 . . . . . . + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . 2 5 Metzgeria furcata E0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . . + . . . . 2 5 Thuidium tamariscinum E0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . + . . 2 5 Brachythecium rutabulum E0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . + 2 5 Cladonia pyxidata E0 . . . . . . . + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 2 Plagiothecium sp. E0 . . . . . . . . + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 2 Homalothecium philippeanum E0 . . . . . . . . . . . + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 2 Hypnum cupressiforme E0 . . . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . . . . . . . . . 1 2 Atrichum undulatum E0 . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . 1 2 Thuidium delicatulum E0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 . . . . . . . . . . . 1 2 Bryum capillare E0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . . . . 1 2 Fissidens taxifolius E0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . . . . 1 2 Dicranum scoparium E0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . 1 2 Mnium marginatum E0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . 1 2 Conocephalum conicum E0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . 1 2 Anomodon viticulosus E0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . 1 2 Legend - Legenda D Dolomite - dolomit DR Dolomite with chert - dolomit z rožencem DL Dolomite with marlstone - dolomit z laporovcem DA Dolmite limestone - dolomitiziran apnenec Ko Conglomerate - konglomerat Re Rendzina - rendzina Number of relevé (Zaporedna številka popisa) 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 31 32 33 34 35 36 37 38 39 40 41 Pr. Fr. Number of relevé (Zaporedna številka popisa) 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 31 32 33 34 35 36 37 38 39 40 41 Pr. Fr. Number of relevé (Zaporedna številka popisa) 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 31 32 33 34 35 36 37 38 39 40 41 Pr. Fr. Number of relevé (Zaporedna številka popisa) 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 31 32 33 34 35 36 37 38 39 40 41 Pr. Fr. Number of relevé (Zaporedna številka popisa) 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 31 32 33 34 35 36 37 38 39 40 41 Pr. Fr. Number of relevé (Zaporedna številka popisa) 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 31 32 33 34 35 36 37 38 39 40 41 Pr. Fr. Table 2 (Preglednica 2): Arunco-Fagetum tilietosum Number of relevé (Zaporedna številka popisa) 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 31 32 Database number of relevé (Delovna številka popisa) 221126 228755 222979 222970 223038 222973 222975 255546 255547 255556 255557 245848 246726 246728 255542 255543 246727 223031 221237 221238 221240 233729 221239 255558 255562 255560 255559 255561 246304 255564 255565 255566 Elevation in m (Nadmorska višina v m) 890 270 800 410 260 440 600 370 340 440 270 715 440 380 520 540 440 620 300 300 290 320 340 270 390 370 290 390 330 520 370 400 Aspect (Lega) NE NE NNW W E NW NNW N NE N NE NW NNE N NNE NNW NNE SE E E E NE NE NNE NE NE N N NNW NNW NW N Slope in degrees (Nagib v stopinjah) 45 45 35 30 15 35 35 35 35 35 35 30 40 30 35 30 45 35 35 30 35 35 35 40 35 30 35 35 35 35 35 30 Parent material (Matična podlaga) AR D D D Gr D D AR AR AR AR D A AR AR AR AR DA DR DR DR DR DR DR DR DR DR DR D DR DRG DRG Soil (Tla) Re Re Re Re Re Re Re Re Re Re Re Re Re Re Re Re Re Re Re Re Re Re Re Re Re Re Re Re Re Re Re Re Stoniness in % (Kamnitost v %) 30 30 40 10 20 10 30 20 20 30 30 50 30 10 70 60 30 10 20 10 10 10 10 10 10 10 20 10 40 10 5 10 Cover in % (Zastiranje v %): Upper tree layer (Zgornja drevesna plast) E3b 80 80 80 70 90 60 80 60 80 70 70 90 90 90 80 90 90 70 70 80 70 90 80 100 80 80 90 70 90 100 90 100 Lower tree layer (Spodnja drevesna plast) E3a . 20 10 30 . 30 10 40 20 20 20 10 10 10 10 10 5 20 30 10 20 10 10 10 20 20 10 30 10 10 10 10 Shrub layer (Grmovna plast) E2 10 10 5 20 5 5 2 10 10 10 20 10 10 5 5 5 5 2 40 30 40 30 10 10 10 5 10 20 10 10 10 10 Herb layer (Zeliščna plast) E1 50 70 50 60 70 50 40 60 50 40 50 60 60 70 50 50 60 40 70 70 70 70 70 60 70 70 70 80 70 70 90 80 Moss layer (Mahovna plast) E0 10 20 30 10 10 5 15 10 10 10 10 20 10 5 20 10 10 5 15 5 20 10 10 10 10 10 20 10 10 5 5 5 Maximum diameter of trees (Največji prsni premer dreves) cm 35 35 40 35 35 30 35 35 30 30 35 40 35 40 40 40 40 35 45 50 40 40 40 45 40 40 40 40 40 50 45 60 Maximum height of trees (Največja drevesna višina) m 16 19 22 20 18 20 20 18 20 20 20 25 20 24 25 2 22 22 28 30 25 24 27 24 22 24 24 22 25 24 22 35 Number of species (Število vrst) 68 70 61 69 60 69 71 68 68 68 71 69 47 44 60 54 49 59 86 68 65 77 56 69 70 59 79 86 66 49 45 36 Relevé area (Velikost popisne ploskve) m2 200 400 200 400 400 400 400 400 400 400 400 400 400 400 400 400 400 200 400 400 400 400 400 400 400 400 200 400 400 400 400 400 Date of taking relevé (Datum popisa) 6/30/2004 5/21/2002 7/26/1990 5/25/1991 7/27/1990 7/26/1990 7/26/1990 24.5,1990 6/11/1990 24.5,1990 6/11/1990 5/3/2012 4/26/2011 4/26/2011 4/15/2003 4/15/2003 4/26/2011 7/27/1990 4/17/2008 4/17/2008 4/17/2008 4/11/2009 4/17/2008 5/16/2003 5/23/1990 5/23/1990 4/17/2001 5/23/1990 6/17/2011 5/6/2005 5/6/2005 5/6/2005 Locality (Nahajališče) Srpenica - Strmi breg Dolenja Trebuša - Prvejk Matajur - Nadiža Matajur-Nadiža Mija Matajur - Nadiža Matajur - Nadiža Bučenica Bučenica Bučenica Bučenica Čepovan-Vrše Bučenica Bučenica Godiča - Bukovo Godiča - Bukovo Bučenica Mija Kozlov rob Kozlov rob Kozlov rob Kozlov rob Kozlov rob Kozlov rob Kozlov rob Kozlov rob Kozlov rob Kozlov rob Stopnik-Pri Vojsku Idrija pri Bači - Mlaka Idrija pri Bači - Mlaka Idrija pri Bači - Mlaka Quadrant (Kvadrant) 9747/1 9949/1 9747/3 9747/3 9747/3 9747/3 9747/3 9848/1 9848/1 9848/1 9848/1 9948/2 9848/1 9848/1 9848/2 9848/2 9848/1 9746/4 9848/1 9848/1 9848/1 9848/1 9848/1 9848/1 9848/1 9848/1 9848/1 9848/1 9849/3 9848/4 9848/4 9848/4 Coordinate GK Y (D-48) m 384707 410303 385597 385009 384478 385093 385337 402812 403283 402511 403423 407654 402823 402064 404833 404943 402647 383785 402162 402199 402199 402102 402157 402109 402101 402089 402059 402023 415624 407062 406912 406750 Coordinate GK X (D-48) m 5127062 5106569 5121282 5121302 5121086 5121383 5121354 5114840 5114638 5114798 5114622 5105516 5114751 5115038 5115670 5115680 5114770 5121161 5117138 5117061 5117094 5117144 5117010 5117006 5116999 5117040 5117227 5117040 5108170 5111432 5111636 5111562 Diagnostic species of the association (Diagnostične vrste asociacije) Pr. Fr. QF Veratrum nigrum E1 + 1 1 1 1 + r 1 + + 1 2 2 2 1 1 16 50 TA Aruncus dioicus E1 + 1 + + 1 + + + + + 1 1 + 1 14 44 FS Prenanthes purpurea E1 + + + + + + + 1 + + 10 31 Fs Epipactis helleborine E1 + + + + + + 6 19 QF Spiraea chamaedryfolia E2a + + + + 4 13 QP Tamus communis E1 + + + 3 9 AF Helleborus niger E1 + 1 3 MuA Phyteuma ovatum E1 + 1 3 Geographical differential species (Geografske razlikovalne vrste) AF Anemone trifolia E1 1 1 1 1 1 + 1 1 + 1 1 1 1 1 1 + + 1 1 1 1 1 1 1 1 1 1 1 + + . 1 31 97 AF Geranium nodosum E1 . . . . . . . 1 . . . . + + + 1 + . 1 1 1 1 . 1 1 1 + 1 1 . . . 16 50 QP Sesleria autumnalis E1 . . . . . . . + . + + . + . . . . . r . . + + r + . . + . . . . 10 31 MuA Aconitum angustifolium E1 . . . + 1 . . . + . 1 . . . . . . . . . + . . r . . . + . . . . 7 22 FS Lathyrus vernus subsp. flaccidus E1 . . . . . . . . + + . . + . . . + . . . . . . . . . . . . . . . 4 13 TA Tephroseris pseudocrispa E1 . . . . . . . . . . . . . . . . . . . . . . . + 1 . + + . . . . 4 13 Differential species of the subassociaton (Razlikovalnice subasociacije) TA Phyllitis scolopendrium E1 + 2 1 + + + + 1 + 1 1 + + + 1 1 1 + + + + 1 + 1 1 1 1 2 1 . + . 30 94 FS Tilia cordata E3b . . . + . + . . . . . r . . 1 . . 1 3 3 3 1 + 1 1 1 + 2 + . . + 17 53 FS Tilia cordata E3a . + + . . + + . . . . . . + + . . . 1 1 . . . + 1 + . . . . . . 11 34 FS Tilia cordata E2b . . . . . . . . . . . . . . . . . . + . . . . + . + + . + . . . 5 16 FS Tilia cordata E2a . . . . . . . . . . . + . . . . . . . . . + . . + . + 1 . + . . 6 19 FS Tilia cordata E1 . . . + . . . . . . . + . . . . . . . . . . . + + + . + . . . + 7 22 TA Tilia platyphyllos E3b . + 2 . . . . . + . . + . . . . . + . . . + . . . . . . . + . . 7 22 TA Tilia platyphyllos E3a . + 1 . . . + . . . . + . . . . + + . + + + . . . . . . . . . . 9 28 TA Tilia platyphyllos E2b . . . . . . . + . . . . . . . . . . . . . . . . . . . . . . . . 1 3 TA Tilia platyphyllos E2a . . . . . . . . . . . + . . . . . . . . . + . . . . . . . . . . 2 6 TA Tilia platyphyllos E1 . . + . . . . . . . + . . . . . + . . . . . . . . . . . . + . . 4 13 Differential species of variants (Razlikovalnice variant) FS Luzula nivea E1 2 . + + 1 . + 1 + 1 + . 1 + . . + + . . . . . . + . . + . . . . 15 47 AT Moehringia muscosa E1 . . + + + + . + + + + + + . + + . . . . . . . . . . + . 1 . . . 14 44 FS Cardamine pentaphyllos E1 1 . 1 + . + 1 + . 1 . + . . . . 2 . . . . . . . . . + . . . . . 10 31 TG Vincetoxicum hirundinaria E1 . + + + 1 1 + + . . . . . . . . . + . . . . . . . . . . . . . . 8 25 AF Hacquetia epipactis E1 . + . . . . . . . . . . . + . . . . 1 1 1 1 + + + . 1 1 + 1 1 . 14 44 FS Allium ursinum E1 . . . . . . . . . . . . . . . . . . r + + + . + . . 2 2 + + 4 4 11 34 FS Euphorbia amygdaloides E1 . . . . + . . . . . . . . . . . . + + + 1 + + + . . + + . . . . 10 31 FS Pulmonaria officinalis E1 . . . . . . . . . . . . . . . + . . 1 + + + . . . . + + 1 . + . 9 28 QP Ruscus aculeatus E1 . . . . . . . . . . . . . . . . . . + + + + + . . + . . . . + . 7 22 AF Aremonio-Fagion Cardamine enneaphyllos E1 1 1 + 2 2 1 2 2 2 + 2 2 2 2 2 3 2 2 + 1 1 2 1 2 2 2 1 2 1 1 2 1 32 100 Cyclamen purpurascens E1 1 1 1 + 1 1 . 1 1 1 + 1 + + + 1 + 1 2 1 1 1 1 1 1 1 1 1 + + + + 31 97 Lamium orvala E1 1 1 . 1 . + + + + + . 1 1 + . . 1 1 . . . + . + + . . + 2 1 + + 21 66 Cardamine trifolia E1 . . + . + + 1 + 1 1 1 + + 1 + . . . . . . . . . . . + . 1 . + + 16 50 Euphorbia carniolica E1 . . . . + . . . . . . . . . . . . 1 . . . . . . . . . . . + + . 4 13 Epimedium alpinum E1 . . . . . 2 . . . . . . . . . . . . . . . . . . . . . . . . . . 1 3 Daphne laureola E2a . . . . . . . . . . . + . . . . . . . . . . . . . . . . . . . . 1 3 Omphalodes verna E1 . . . . . . . . . . . r . . . . . . . . . . . . . . . . . . . . 1 3 Rhamnus fallax E2 . . . . . . . . . . . + . . . . . . . . . . . . . . . . . . . . 1 3 Vicia oroboides E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . 1 3 EC Erythronio-Carpinion Helleborus odorus E1 . + . 1 + 1 1 + 1 + + 1 . 1 + + + + 1 1 2 . 2 1 . 1 + . 1 1 + . 25 78 Primula vulgaris E1 . + . . . + 1 . + . . + . + . . . 1 1 + 1 + + + + + + 1 . + . . 18 56 Galanthus nivalis E1 . . . . + + . . . . . . . 2 . . . . 1 1 1 2 1 2 2 1 1 2 . . 1 . 14 44 Ornithogalum pyrenaicum E1 . . . . . . . . . . . . . . . . . . + + + + . . . . . . . + . . 5 16 Crocus vernus subsp. vernus E1 . . . . . . . . . . . . . . . . . . + . . + . . . . . . . . . . 2 6 TA Tilio-Acerion Acer pseudoplatanus E3b + + 1 . . 1 1 . . . + . + . . + + 2 + . . . . r + + + . 1 1 1 + 19 59 Acer pseudoplatanus E3a 1 . + . . + + . . + + . . . . . . . . . . . . r . + . 1 . + + . 11 34 Acer pseudoplatanus E2b . + + . . . + . . . . r + . . . . + + . . . . + . . . . . . . . 8 25 Acer pseudoplatanus E2a + . . . . . . . . . . . . . . . . + + . 1 + . . . + 1 1 . . + . 9 28 Acer pseudoplatanus E1 + . . . . . 1 + . . 1 . + . . + . . 1 1 1 + 1 + . . + . + 1 1 + 17 53 Polystichum aculeatum E1 + + + + + . + . + + . + . . + + + . + + . . . . . . + 1 1 + . + 19 59 Acer platanoides E3b . + . . . . . . . . . . . . . . + + . . . . . + + . . . . . . . 5 16 Acer platanoides E3a . . + . . . + . . . . . . . . . + . . . . . . . . . . . . . . . 3 9 Acer platanoides E2b . + . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . 2 6 Acer platanoides E2a . . . . . . . . . . . + . . . . + . . + . + . . + . . . . . . . 5 16 Acer platanoides E1 . + . . . + . . . . . 1 1 + . . 1 . . . . 1 + + + + + . + + . . 14 44 Ulmus glabra E3b . . . 1 . . + . . . . . . . + + + + r . . + . r + . . . . 1 + + 13 41 Ulmus glabra E3a . . . . . . . . . . . . . . + + + . + + . . . . . . . + . + . . 7 22 Ulmus glabra E2b . . . + . . . . . . . . . + . + . . 1 + + + . . . . . . . . + . 8 25 Ulmus glabra E2a . . . . . . . . . . + . . . . . . + 1 + 1 + + . . . + + . . . . 9 28 Ulmus glabra E1 . . . . . . . . . . + . + + + + . . + . . . . . . . . . + . + + 9 28 Geranium robertianum E1 + . . . . . . . . . . . . . + + 1 + r . . . . . . . . . + . . . 7 22 Polystichum setiferum E1 . 1 . . . . . . . . . . . . . + . . . . . . . . . . . + 1 . 2 1 6 19 Adoxa moschatellina E1 . . . . . . . . . + . . . + 1 1 . . . . . . . + . . . + . . . . 6 19 Dryopteris affinis E1 . . . . . . . . . + . . . . . . . . . . . + . . . . + + + . . + 6 19 Staphylea pinnata E2a . + . 1 + + . . . . . . . . . . . . . . . . . . . . . . . . . . 4 13 Staphylea pinnata E1 . + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 3 Euonymus latifolia E2b + . . . . . . . . . . + . . . . . . . . . . . . . . . . . + . . 3 9 Arum maculatum E1 . . . . . . . . . . . . . . + . . . . . . . . . . . . . + . . . 2 6 Polystichum x bicknellii E1 . + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 3 Polystichum x illyricum E1 . . . . . . r . . . . . . . . . . . . . . . . . . . . . . . . . 1 3 Juglans regia E1 . . . . . . . . . . . . + . . . . . . . . . . . . . . . . . . . 1 3 Corydalis solida E1 . . . . . . . . . . . . . . 2 . . . . . . . . . . . . . . . . . 1 3 Polystichum braunii E1 . . . . . . . . . . . . . . r . . . . . . . . . . . . . . . . . 1 3 FS Fagetalia sylvaticae Fagus sylvatica E3b 4 4 3 3 5 4 4 4 5 5 4 5 4 5 4 5 4 3 3 3 3 4 5 5 4 4 5 4 3 5 5 5 32 100 Fagus sylvatica E3a 1 . + + . . . . 1 + 1 1 + . + + . . . + . + . 1 . 1 1 + 1 + 1 1 20 63 Fagus sylvatica E2b 1 1 + 1 + + . 1 1 + . . 1 + + + 1 . . 1 . 1 . + . + 1 . 1 1 1 1 23 72 Fagus sylvatica E2a . + . . . . . . . . . . 1 + . . . . 1 2 2 2 + + + . 1 1 . + + + 15 47 Fagus sylvatica E1 1 1 . . + . . . + . + 1 1 + . . . . + 1 1 1 1 + + . . . . 1 . 1 17 53 Asarum europaeum subsp. caucasicum E1 + + 1 1 1 1 1 1 1 1 1 1 1 + + 1 + + 1 1 1 . 1 1 1 1 1 1 1 + + + 31 97 Mercurialis perennis E1 1 1 . + 1 + + 1 1 1 1 2 1 + + + + 1 2 1 2 1 2 2 1 . 1 1 1 3 + . 29 91 Lathyrus vernus subsp. vernus E1 . 1 . + 1 + + 1 1 1 + 1 + + + + + . 1 1 1 1 1 1 . 1 1 1 + + + + 28 88 Actaea spicata E1 1 . 1 + + + 1 + 1 1 + + + + + . + 1 . . + + . 1 1 + + 1 + 1 + + 27 84 Dryopteris filix-mas E1 1 . 1 + + + 1 1 1 1 1 1 1 1 + 1 1 1 . . . + . + + + + 1 1 1 + 1 27 84 Salvia glutinosa E1 1 + + + + + + + + + + + + . + . + . 1 1 + + + . + + + + + + . . 26 81 Galium laevigatum E1 1 1 . + . . 1 1 1 + 1 . + . + + + + 1 . + + + 1 + + + + + + + . 25 78 Galeobdolon flavidum E1 1 + . + 1 . + 1 1 + 1 + . . + . . . 1 1 + . + 1 + 1 + 1 1 + + + 24 75 Daphne mezereum E2a + + + . . + + + + + + + + . . + . + + + . . + + + + + . + + . . 22 69 Polygonatum multiflorum E1 + + . . + . + . . + + + + + . . . + 1 1 + . 1 + 1 + + 1 . + . . 20 63 Carpinus betulus E3b . . . . . . . . . . . . . . + + . . + . . . . . . . . . + + 2 . 6 19 Carpinus betulus E3a . . . + . . + . . . + . . + + . . 2 1 1 + + . . . + + + + + + . 16 50 Carpinus betulus E2b . + . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . 2 6 Carpinus betulus E2a . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . 1 3 Euphorbia dulcis E1 . . . + . + . + + + + . . . . . . . 1 + + + . + + + + 1 . + . . 16 50 Mycelis muralis E1 + + . + + . . + + + + + + . + + . + . . . . + + + . . . . . . . 16 50 Symphytum tuberosum E1 + . . . . . . . + . + . . . + . . . 1 1 + 1 . + + + + . 1 + + + 16 50 Paris quadrifolia E1 . . . . . . . . . . . + + + . . + . + + . . + + . + + + + + + + 15 47 Cardamine bulbifera E1 . . . . . . . . . . . . . 1 + 1 . 1 . + + 1 1 + . . 1 + . 1 + 1 14 44 Campanula trachelium E1 + + . + . + . . + + . + + . . . . + . . . . . + + . + . . . . . 12 38 Neottia nidus-avis E1 . . . . . + . + + 1 + . . . . . . + . + . . + 1 + + . . + . . . 12 38 Lilium martagon E1 + . . . . . . . . . . . + + . . . . 1 1 . + + . . + + 1 . + . . 11 34 Fraxinus excelsior E3b . + . . . . . . . . . + . + r . . . r . + . . . . . 1 + 3 . . . 9 28 Fraxinus excelsior E2b . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . 1 3 Fraxinus excelsior E2a . . . . . . . . . . . . . . . . . . . 2 + + . . . . + . . + . . 5 16 Fraxinus excelsior E1 . . . . . . . . . . . + . + . . . . . . . . + . . . 1 . . . . + 5 16 Prunus avium E3b . . . . . . . . . . . . . r . . . . + + . + . r + . . . + . r + 9 28 Prunus avium E3a . . . . . . . + . . . . . . . . . . . . . . . . . . . . . . . . 1 3 Prunus avium E2b . . . . . . . 1 . . . . . . . . . . + . . . . . . . . . . . . . 2 6 Prunus avium E2a . . . . . . . . . . . . . . . . . . + . . . . + . . . . . . . . 2 6 Prunus avium E1 . . . . . . . + . . . . . + . . . . . . . . . . . . . . . . . + 3 9 Viola reichenbachiana E1 . . . . . + + . . . . . . . + . . . + . . + + . + . + . . . . . 8 25 Heracleum sphondylium E1 . . . + . . . + . . . . . . . . . . . . . . . + + + + + . . . . 7 22 Sambucus nigra E2b . . . . . . . . . . + . . . . . . . . . . . . . . + . . . . . . 2 6 Sambucus nigra E2a . . . . . . . . . + . + . . . . 1 . . . . . . . . . . + + . + + 7 22 Sambucus nigra E1 . . . . . . . . . . . . . . + . . . . . . . . . . . . . . + + + 4 13 Scrophularia nodosa E1 + . + + . . . . . + . . . . . . . + . . . . . . . . . . + . . . 6 19 Melica nutans E1 . . . + . . . . . . + . . . . . . . + + . + . . + . . . . . . . 6 19 Galium odoratum E1 . . . . . . . . . . . . . + + + . . . . . . . . . . . . . . + + 5 16 Brachypodium sylvaticum E1 . . . . . . . . . . . . . . . . + . 1 + . + . . . . . . + . . . 5 16 Petasites albus E1 . . . . . . . . . . . . . . . . . . . . . . . + + . + + . 1 . . 5 16 Orchis pallens E1 . . . . . . . . . . . . . . . . . . 1 + + + . . . . . . . . . . 4 13 Laburnum alpinum E3b + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 3 Laburnum alpinum E2b . + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 3 Laburnum alpinum E1 + . + . . . + . . . . . . . . . . . . . . . . . . . . . . . . . 3 9 Cephalanthera damasonium E1 . . . . . . . + + + . . . . . . . . . . . . . . . . . . . . . . 3 9 Epilobium montanum E1 + . . . . . . . . . . . . . + . . . . . . . . . . . . . . . . . 2 6 Cardamine impatiens E1 . . . + . . . . . . . . . . + . . . . . . . . . . . . . . . . . 2 6 Myosotis sylvatica E1 . . . . . . . . . . . . . . . . . . . . . . . + + . . . . . . . 2 6 Lonicera alpigena E2 1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 3 Festuca altissima E1 + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 3 Ranunculus lanuginosus E1 . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . 1 3 Phyteuma spicatum subsp. coeruleum E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . 1 3 QP Quercetalia pubescenti-petraeae Ostrya carpinifolia E3b 1 1 . 1 . . . . . . + 1 1 + 1 1 2 . 1 + . + . r . . . . + + + . 17 53 Ostrya carpinifolia E3a . + 1 1 . + 1 1 + + 1 . 1 . . + 1 + . . + + . + 1 + . 1 . . . . 19 59 Ostrya carpinifolia E2a . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . 1 3 Ostrya carpinifolia E1 . . . . + . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 3 Fraxinus ornus E3b . . . . . . . . . . . . . . r . . . + . . . . . . . . . . . + . 3 9 Fraxinus ornus E3a . . . 1 + 1 + + . . . . . . . r . + + . + . . . . . . + . . . . 10 31 Fraxinus ornus E2b + + . . . . . . . . . . . . . . . . 1 . . . . . . . . . . . . . 3 9 Fraxinus ornus E2a . . . + . . . . . . . . . . . . . . 1 + . + . . . . . . . . . . 4 13 Fraxinus ornus E1 + . . + + . + . . . 1 + + + . . + . 1 . . 1 + + . . . . + . . . 14 44 Melittis melissophyllum E1 + + + + 1 + + + + + . . . . . . . . . . . . . . . + . . . . + . 12 38 Sorbus aria E3b r . . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . 2 6 Sorbus aria E3a + + + . . 1 + . + + . . . . . . . . + . . . . . . . . + . . . . 9 28 Sorbus aria E2b . . . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . 1 3 Sorbus aria E2a . . . . . . . . + . . . . . . . . . + . . . . . . . . . . . . . 2 6 Tanacetum corymbosum E1 . . . + . . . . . . . . . . . . . . + + + + + . + + . . . . . . 8 25 Euonymus verrucosa E2a . . . . + . . . . . . . . . . . . . + + + + . . . . + + . . . . 7 22 Arabis turrita E1 + . + . . . . . + . + . . . . . . . . . . . . . . . . + . . . . 5 16 Cornus mas E2b . + . + . . . + . . . . . . . . . . 1 . . . . . . . . . + . . . 5 16 Cornus mas E2a . . . . . . . . + . . . . . . . . . . . . . . . . . . . . . . . 1 3 Hypericum montanum E1 + . + . . . + . . . . . . . . . + . . . . . . . . . . . . . . . 4 13 Convallaria majalis E1 . . . . + . . . . . + . . . . . . . . + . . . . . . . . . . . . 3 9 Asparagus tenuifolius E1 . . . . + . . . . . . . . . . . . . + . . . . . . . . . . . . . 2 6 Primula x ternovania E1 . . . 1 . . + . . . . . . . . . . . . . . . . . . . . . . . . . 2 6 Calamintha sylvatica E1 . . . + . . . . . . . . . . . . . + . . . . . . . . . . . . . . 2 6 Primula veris subsp. columnae E1 + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 3 Quercus cerris E1 . . . . . . . . . . + . . . . . . . . . . . . . . . . . . . . . 1 3 QR Quercetalia roboris . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Quercus robur E3b . . . . . . . . . . . . . . . . . . + + . + . . . . . + . . . . 4 13 Betula pendula E3b . . . . . . . . . . . r . . . . . . . . . . . . . . . . . . . . 1 3 Rubus hirtus E2a . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . 1 3 OF Querco-Fagetea Carex digitata E1 . + + + 1 + 1 + + . + + + + + + + 1 1 1 1 + . + + + + + . + . . 26 81 Hedera helix E3a . . . 1 + . . . . . . + + + . . . . 1 1 + + + + 1 + + . . . . . 14 44 Hedera helix E2b . . . . . . . 1 . + 1 . . . . . . . . . . . . + . . . . . . . . 4 13 Hedera helix E1 . + . . + . + + + . . . + + + + + . 1 1 2 2 + 1 1 + 1 1 . . + + 22 69 Vinca minor E1 . + . 1 2 1 1 . + + . 1 + + . + . 1 2 1 2 . 1 . . + + . 1 1 + + 22 69 Lonicera xylosteum E2a . + . + . . + 1 + . + r + . . . + + 1 1 1 + + . . . . + + + + . 19 59 Hepatica nobilis E1 + . . . . . + . + . 1 + . . + + . . 1 + 1 1 + 1 1 1 1 1 . . . . 17 53 Clematis vitalba E3a . . . . . . . . . . . . . . . . . . + . . . . . . + . . . . . . 2 6 Clematis vitalba E2 . . . + . . . . + + + + . . . . + + + . + . + r . . + . + . . . 13 41 Acer campestre E3b . . . . . . . . . . . . . . . . r . . . . . . . . . . . . . + . 2 6 Acer campestre E3a . . . + . . . + . . . . . . . r + . + . + + . . + . . + + . + . 11 34 Acer campestre E2b . . . + . . . . . . . . . . . . + . + . . + . . . . . . + . . . 5 16 Acer campestre E2a . + . . . . . . . . . . . . . . r . + + 1 + + . . . . . . . . . 7 22 Acer campestre E1 . . . . . . . . . . . . . . . . + . . . . . . . . . + . + . + . 4 13 Listera ovata E1 . . . + . + . . . . . . . . . . . + 1 . + . + + + + . + + . . . 11 34 Corylus avellana E2b + + . 1 . . . + . . . . . . . . + . . . . . . . . . . . 1 + . . 7 22 Corylus avellana E2a . . . . . . . + . . . . . . . . . . + . . . . . . . + . . . . . 3 9 Aegopodium podagraria E1 . . . . . . . . . . . . . . . + . . . + . . . r . . + . 1 + 1 . 7 22 Moehringia trinervia E1 + . . . . . . . . + . . . . + . . . . . . . . . . . + + . . . . 5 16 Viola mirabilis E1 . . . + . . . . . . . . . . . . . . + + . . . . . . + + . . . . 5 16 Rosa arvensis E2a . . . . . . . . + . . . . . . . . . 1 + + + . . . . . . . . . . 5 16 Viburnum opulus E2a . . . . . + . . . . + . . . . . . . . . . . . . . . . + . . . . 3 9 Cerastium sylvaticum E1 . . . . . . . . . . . . . . . . . . . . . . . + + . . + . . . . 3 9 Ilex aquifolium E3a . . . . . . . . . . . + . . . . . . . . . . . . . . . . . . . . 1 3 Ilex aquifolium E2 . . . + . . . . . . . + . . . . . . . . . . . . . . . . . . . . 2 6 Gagea lutea E1 . . . . . . . . . . . . . . + + . . . . . . . . . . . . . . . . 2 6 Galium schultesii E1 . . . . . . . . . . . . . . . . . . . . . . . . + . . + . . . . 2 6 Melica uniflora E1 . . . . + . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 3 Lathraea squamaria E1 . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . . . 1 3 Viola riviniana E1 . . . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . 1 3 EP Erico-Pinetea Carex alba E1 . . . + 2 1 + 1 + . + + . . . . . . 2 + 1 2 + r + + . + + . . . 18 56 Calamagrostis varia E1 + + . + . . . + . . + . + . . + . + . . + + + . . . . . . . . . 11 34 Aquilegia atrata E1 . . . . . . . . + . + . . . . . . . . . . . . . . + . . . . . . 3 9 Aquilegia nigricans E1 . . . . . + 1 . . . . . . . . . . + . . . . . . . . . . . . . . 3 9 Cirsium erisithales E1 . . . . . + + . . . . . . . . . . . + . . . . . . . . . . . . . 3 9 Rubus saxatilis E1 . . + . + . . . . . . . . . . . . . . . . . . . . . . . . . . . 2 6 Buphthalmum salicifolium E1 . . . . . + . . . . . . . . . . . . . . . . . . . . . . . . . . 1 3 VP Vaccinio-Piceetea Oxalis acetosella E1 . . + . + . . + + + . + + + + + + . . . . . . + + . + . + + . + 17 53 Solidago virgaurea E1 + + . . . + . + + + . + . + . . . . 1 + + + + + + . + . . . . . 16 50 Veronica urticifolia E1 + 1 + . . + + . + + + + . . . . + . . . . . . . + . + + . . . . 13 41 Aposeris foetida E1 . . . + + + . . . . . . . . . . . . + + . + . . . + . . . . . . 7 22 Saxifraga cuneifolia E1 + 1 + . . . . . . + . . + . . + . . . . . . . . . . . . + . . . 7 22 Valeriana tripteris E1 + + + . . . . + . . + r . . . . . . . . . . . . . . . . . . . . 6 19 Maianthemum bifolium E1 . . . . + . . . . . . . . + . . . . . . . + . . . . + + . . . . 5 16 Picea abies E3b . . . . . . . . . . . + . . . . . . . . . + r . . . . . . . + . 4 13 Picea abies E3a . . . . . . . . . . . + . . . . . . + . . . . . . + . + . . . . 4 13 Picea abies E2b . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . 1 3 Picea abies E2a . . . . . . . . . . . . . . . . . . . . . . . r . . . + . . . . 2 6 Abies alba E3b . . . . . . . . . . . . . . . . . . r . . + . . . . . . . . . . 2 6 Abies alba E3a . . . . . . . . . . . . . . . . . . . . r . . . . . . . . . . . 1 3 Abies alba E2b . . . . . . . . . . . . . . . . . . + + . . . . . . . . . . . . 2 6 Abies alba E2a . . . . . . . . . . . . . . . . . . . . + 1 . . . . + . . . . . 3 9 Abies alba E1 . . . . . . . . . . . . . . . . . . + . . 1 + . . . . + . . . . 4 13 Hieracium murorum E1 . + . . . . . + + . . . . . . . . . . . . . . . . . . . . . . . 3 9 Luzula luzuloides E1 . . . . . . . . . + . . . . . . . . . . . + . . + . . . . . . . 3 9 Homogyne sylvestris E1 . . + . . + . . . . . . . . . . . . . . . . . . . . . . . . . . 2 6 Luzula pilosa E1 . . . . . . . . . . . . . . . . . . . . . . . . . . + + . . . . 2 6 Dryopteris expansa E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . + 2 6 Polystichum lonchitis E1 . . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . 1 3 RP Rhamno-Prunetea Euonymus europaea E2a . . + . . . . . . . . . . . . . . . + + + . . . . . . + . . . . 5 16 Viburnum lantana E2a . . . . . . . . + . . . . . . . . . + + . . . . . . . . . . . . 3 9 Berberis vulgaris E2a . . . . . . . . . . + . . . . . . . + . . + . . . . . . . . . . 3 9 Crataegus monogyna E2b . . . . . . . . . . . . . . . . . . + . + . . . . . . . . . . . 2 6 Crataegus monogyna E2a . . . . . . . . . . . . . . . . . . + . + . . . . . . . . . . . 2 6 Cornus sanguinea E2b . + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 3 Ligustrum vulgare E2a . . . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . 1 3 Rubus fruticosus agg. E2a . . . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . 1 3 TG Trifolio-Geranietea Campanula rapunculoides E1 . . . + + + . + + + + . + . . . . 1 1 . + + + + . + . + . . . . 16 50 Digitalis grandiflora E1 . + . . . . . . . . . r . . . . . + . . . . . . . . . . . . . . 3 9 Verbascum lanatum E1 + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 3 Vicia sylvatica E1 . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . 1 3 EA Epilobietea angustifolii Hypericum hirsutum E1 + + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 2 6 Fragaria vesca E1 . . . . . . + . . . . . . . . . . . + . . . . . . . . . . . . . 2 6 Eupatorium cannabinum E1 + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 3 Rubus idaeus E2a + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 3 Solanum dulcamara E1 . . . . . . . . . . + . . . . . . . . . . . . . . . . . . . . . 1 3 Urtica dioica E1 . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . . 1 3 Stachys sylvatica E1 . . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . 1 3 MuA Mulgedio-Aconitetea Aconitum lycoctonum s. lat. E1 . + . 1 + 1 . . . . + . . + . . . . + 1 1 + 2 1 1 2 + 1 . . . . 16 50 Senecio ovatus E1 1 1 + . . + + . 1 + 1 + . . . . + . . . . . . . . . . . 1 + + . 13 41 Milium effusum E1 . + . . . . . . . . . . . . . . + . . . . . . + 1 . + 1 . . . . 6 19 Aconitum degenii subsp. paniculatum E1 + + . + . 1 1 . . . . . . . . . . . . . . . . . . . . . . . . . 5 16 Athyrium filix-femina E1 . . . . + . + . . + . . . . . . . . . . . . . . . . . + + . . . 5 16 Silene dioica E1 + + . . . . . . . . . . . . . + . 1 . . . . . . . . . . . . . . 4 13 Saxifraga rotundifolia E1 . + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 3 Veratrum album s. lat. E1 . . . + . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 3 Doronicum austriacum E1 . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . 1 3 ES Elyno-Seslerietea Sesleria caerulea subsp. calcaria E1 . . + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 3 MA Molinio-Arrhenatheretea Ajuga reptans E1 . . . . + . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 3 Veronica chamaedrys E1 . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . 1 3 Angelica sylvestris E1 . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . 1 3 TR Thlaspietea rotundifolii Adenostyles glabra E1 + . 1 . . . 1 + . + . . . . . . . . . . . . . . . . . . . . . . 5 16 Gymnocarpium robertianum E1 . . . . . . . + . . . . + . . . . . . . . . . . . . . . + . . . 3 9 Cerastium subtriflorum E1 + + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 2 6 AT Asplenietea trichomanis Asplenium trichomanes E1 . 1 + + + + + + + + + + + + + + + + + + + . + 1 + + + + + . . . 27 84 Polypodium vulgare E1 . . + . + + + + . . + + + + + + . + + + 1 1 . + + + + + + . . . 22 69 Asplenium ruta-muraria E1 . . . + + + + . . . . + . . . . . + + . . + . . . . . . + . . . 9 28 Asplenium viride E1 + . + . . + + . . + . . . . . . . . . . . . . . . . . . + . . . 6 19 Cystopteris fragilis E1 + . + . . . . . . + . r . . . . . . . . . . . . . . . . . . . . 4 13 Saxifraga petraea E1 . . + . . . . . . . . r . . . . . . . . . . . . . . . . . . . . 2 6 Cardaminopsis arenosa E1 . 1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 3 Phyteuma scheuchzeri subsp. columnae E1 . . . . . . . . . . . r . . . . . . . . . . . . . . . . . . . . 1 3 Asplenium adiantum-nigrum E1 . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . . . 1 3 ML Mosses and lichens (Mahovi in lišaji) Isothecium alopecuroides E0 + 1 1 1 2 1 + 1 1 1 1 1 1 1 2 2 1 1 + . . 2 . 1 1 1 1 1 1 . . . 26 81 Ctenidium molluscum E0 1 1 1 1 + 1 . . + . 1 2 1 . 1 . . + 2 + 1 1 1 + . + 1 1 1 + + + 25 78 Fissidens dubius E0 . . + . + 1 + + + 1 + + . . + . . + + + + . . + 1 + . . . + + + 20 63 Neckera crispa E0 . 1 2 + + 1 1 . + . 1 2 . . + + . . 2 + 1 1 + . . 1 + . 1 . + . 20 63 Thamnobryum alopecurum E0 + . + . . + + . . . . + + + + 1 + . . . + 1 . + 1 + 1 . . 1 1 + 19 59 Plagiothecium denticulatum E0 + + + . + + + 2 1 1 1 . . . + . . . . . . . . . + + + . . . . . 14 44 Brachythecium rutabulum E0 . . + . . . . 1 . 1 + . . + + + . . . . + . . + + 1 . . . . . . 11 34 Eurhynchium angustirete E0 . . . . . + + 1 + + + . . . . . . . + + + + . . . . . . . . . . 10 31 Thuidium tamariscinum E0 . . . . . + + + + + . . . . . . . . . . + 1 + . . . + . . . . + 10 31 Anomodon attenuatus E0 . . . 1 1 + + . . . . . . . + . + + . . . 1 . . . . . . + . . . 9 28 Plagiochila porelloides E0 . + + . + . 1 + . + . . . . + + . . . . . . . . . . . 1 . . . . 9 28 Polytrichum formosum E0 . . + . . . + . . + + + . . . . . . + . . + . . . . + . . . . + 9 28 Anomodon viticulosus E0 . . . + + . . . . . . + . . . + . 1 1 . . + + . . . . . . . . . 8 25 Mnium thomsonii E0 + . + . . . . . . + + + . . + . . . . . . + . . . . . + . . . . 8 25 Plagiomnium undulatum E0 + . . . . . . . + + + . . . . + . . . . . . . + . . . + + . . . 8 25 Schistidium apocarpum E0 + + + . 1 . . . . . . + . . 1 + . + . . . . . . . . . . . . . . 8 25 Brachythecium velutinum E0 . . . + . . . + . + + . . . . . . + . . . . . . + . . + . . . . 7 22 Hypnum cupressiforme E0 . . + . + . . + . . . . . . . . . . . . . . . . + + . 1 . . . + 7 22 Porella platyphylla E0 . . . 1 + . . . + . . . + . + . + + . . . . . . . . . . . . . . 7 22 Peltigera canina E0 . + + . . . . . . . . . . . + + + . . . . . . + . . . . . . . . 6 19 Bryum capillare E0 + . . . . . . . + . . . . . . + . . . . . . + . + . . . . . . . 5 16 Metzgeria furcata E0 . . + . . . . . + . . . . . . + . . . . . . . . + + . . . . . . 5 16 Homalothecium philippeanum E0 . . . . . . . . 1 . + . . . 1 . . . . . . . + . . . . . . . . . 4 13 Isothecium myosuroides E0 . . 1 . . + + . . . . . . . . . . . . . . . . . . . . . . . . . 3 9 Homalothecium lutescens E0 . . . + . . . + . . . . . . . + . . . . . . . . . . . . . . . . 3 9 Tortella tortuosa E0 . . . . . . + . . . . . . . . . . . . . . . + . + . . . . . . . 3 9 Neckera complanata E0 . . . . . . . . . . + . . . + + . . . . . . . . . . . . . . . . 3 9 Atrichum undulatum E0 + . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . 2 6 Conocephalum conicum E0 . . . . . . + . . . . . . . . . . . . . . . . . . . . + . . . . 2 6 Bartramia halleriana E0 . . . . . . . + . + . . . . . . . . . . . . . . . . . . . . . . 2 6 Rhytidiadelphus triquetrus E0 . . + . . . . . . . . . . . . . . . . . . + . . . . . . . . . . 2 6 Thuidium delicatulum E0 . . . . + + . . . . . . . . . . . . . . . . . . . . . . . . . . 2 6 Hylocomium splendens E0 . . + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 3 Hypnum cupressiforme var. filiforme E0 . . . + . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 3 Ulota crispa E0 . . . . . + . . . . . . . . . . . . . . . . . . . . . . . . . . 1 3 Plagiothecium undulatum E0 . . . . . . . + . . . . . . . . . . . . . . . . . . . . . . . . 1 3 Collema cristatum E0 . . . . . . . + . . . . . . . . . . . . . . . . . . . . . . . . 1 3 Plagiomnium cuspidatum E0 . . . . . . . . . . + . . . . . . . . . . . . . . . . . . . . . 1 3 Dicranum scoparium E0 . . . . . . . . . . . + . . . . . . . . . . . . . . . . . . . . 1 3 Scleropodium purum E0 . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . 1 3 Legend - Legenda A Limestone - apnenec AR Limestone with chert - apnenec z rožencem D Dolomite - dolomit DR Dolomite with chert - dolomit z rožencem DRG Dolomite with chert and claystone - dolomit z rožencem in glinavcem Gr Gravel - grušč Re Rendzina - rendzina Number of relevé (Zaporedna številka popisa) 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 31 32 Pr. Fr. Number of relevé (Zaporedna številka popisa) 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 31 32 Pr. Fr. Number of relevé (Zaporedna številka popisa) 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 31 32 Pr. Fr. Number of relevé (Zaporedna številka popisa) 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 31 32 Pr. Fr. Number of relevé (Zaporedna številka popisa) 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 31 32 Pr. Fr. Number of relevé (Zaporedna številka popisa) 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 31 32 Pr. Fr. Table 3 (Preglednica 3): Arunco-Fagetum var. geogr. Anemone trifolia Number of relevé (Zaporedna številka popisa) 1 2 3 4 6 5 7 8 9 10 11 12 13 Database number of relevé (Delovna številka popisa) 221163 221164 228715 241500 249666 222812 255544 223033 223035 223037 255509 223067 246272 Elevation in m (Nadmorska višina v m) 900 880 260 830 570 580 540 420 420 350 560 1050 590 Aspect (Lega) N NNW NNE E NE N NE NE N SE SEE NE NNE Slope in degrees (Nagib v stopinjah) 45 40 45 40 35 35 35 30 35 35 30 40 30 Parent material (Matična podlaga) AR AR D D D D DR DA D D AR DR D Soil (Tla) Re Re Re Re Re Re Re Re Re Re Re Re Re Stoniness in % (Kamnitost v %) 30 20 20 10 5 10 15 25 5 10 40 50 0 Cover in % (Zastiranje v %): Upper tree layer (Zgornja drevesna plast) E3b 80 90 80 80 90 80 70 80 70 60 90 90 80 Lower tree layer (Spodnja drevesna plast) E3a 20 5 10 10 10 10 20 20 20 20 5 10 10 Shrub layer (Grmovna plast) E2 10 5 20 10 30 10 20 10 5 20 5 5 20 Herb layer (Zeliščna plast) E1 60 70 60 60 60 60 80 80 70 70 30 30 40 Moss layer (Mahovna plast) E0 20 10 5 5 5 20 20 10 10 10 5 10 5 Maximum diameter of trees (Največji prsni premer dreves) cm 30 45 40 35 45 45 30 30 30 30 45 40 40 Maximum height of trees (Največja drevesna višina) m 20 26 18 20 27 22 15 18 18 16 20 24 25 Number of species (Število vrst) 58 49 62 35 64 64 89 80 75 62 43 41 77 Relevé area (Velikost popisne ploskve) m2 400 400 200 200 400 200 400 200 200 200 400 400 400 Date of taking relevé (Datum popisa) 5/22/2008 5/22/2008 4/22/2004 4/12/2011 8/29/2013 6/6/1996 8/13/1991 7/27/1990 7/27/1990 7/27/1990 7/5/1991 8/27/1998 3/21/2012 Locality (Nahajališče) Črvov vrh Črvov vrh Straža Skopica Idrijska Bela-Kucelj Učja - Žila Godiča Mija Mija Mija Ljubinj - Pilanec Lom - Struje Godovič Quadrant (Kvadrant) 9849/3 9849/3 9849/4 9948/2 0049/2 9646/4 9848/2 9746/4 9746/4 9746/4 9848/2 9748/4 0050/1 Coordinate GK Y (D-48) m 412424 412550 418432 408310 421265 381736 405207 384095 384125 384274 406684 405884 428942 Coordinate GK X (D-48) m 5111682 5111742 5106906 5106307 5090282 5130772 5115854 5121050 5121170 5121077 5115360 5118164 5090688 Diagnostic species of the association (Diagnostične vrste asociacije) Pr. Fr. TA Aruncus dioicus E1 . + 1 + . . . + + . . . + 6 46 MuA Phyteuma ovatum E1 + + . . . . + 1 . . . + . 5 38 VP Rosa pendulina E2a + . . . . . + + + . . + . 5 38 QF Veratrum nigrum E1 . . 1 . . . . 1 2 + . . . 4 31 FS Prenanthes purpurea E1 . . . . + . + . . . . + + 4 31 FS Epipactis helleborine E1 . . . . + + + . . . . . + 4 31 AF Helleborus niger E1 . . . 1 + . . . . . . . 1 3 23 QF Tamus communis E1 . . . . . . . . . + . . + 2 15 QF Spiraea chamaedryfolia E2a . . 1 . . . . . . . . . . 1 8 VP Laserpitium krapfii E1 . . . . . . . . . . . + . 1 8 Geographical differential species (Geografske razlikovalne vrste) AF Anemone trifolia E1 . . 1 + . 1 . 1 1 + 1 . . 7 54 AF Omphalodes verna E1 . . + . 1 . . . . . . . 1 3 23 QF Lathyrus vernus subsp. flaccidus E1 . . . . . . + . . . . . . 1 8 Differential species of lower units (Razlikovalne vrste nižjih enot) FS Cardamine pentaphyllos E1 + 2 . 1 . . . . . . . . . 3 23 MuA Aconitum degenii subsp. paniculatum E1 + + . + . . . . . . . . . 3 23 TR Gymnocarpium robertianum E1 . . . . + + + . + + . . + 6 46 TG Campanula rapunculoides E1 . . . . . + 1 1 + 1 + . . 6 46 ES Sesleria caerulea subsp. calcaria E1 . . + . . . 2 1 1 + . + . 6 46 TG Vincetoxicum hirundinaria E1 . . . . . + + + + + . . . 5 38 QP Convallaria majalis E1 . . . . + . 1 + 1 2 . . . 5 38 EP Rubus saxatilis E1 . . . . 1 + . + + . . . . 4 31 FS Luzula nivea E1 . . . . . 1 + 2 + . . . . 4 31 MuA Aconitum angustifolium E1 . . . . . . 1 + + 1 . . . 4 31 QF Vinca minor E1 . . . . . + 1 2 . 1 . . . 4 31 AF Epimedium alpinum E1 . . . . . . . . 3 1 . . . 2 15 AF Aremonio-Fagion Cyclamen purpurascens E1 + . 1 + + 1 1 1 1 1 . 1 1 11 85 Cardamine enneaphyllos E1 1 1 2 1 1 . + . 1 . 1 . . 8 62 Euphorbia carniolica E1 . . + . + 1 . + 1 + . . + 7 54 Knautia drymeia E1 . . + . + + . . . . . . . 3 23 Geranium nodosum E1 . . + . . . 1 . . . . . . 2 15 Lamium orvala E1 . . . + . + . . . . . . . 2 15 Rhamnus fallax E2 . . . + . . . . . . . . + 2 15 Daphne laureola E2a . . . . + . . . . . . . + 2 15 Hemerocallis lilio-asphodelus E1 . . + . . . . . . . . . . 1 8 Potentilla carniolica E1 . . + . . . . . . . . . . 1 8 Cardamine trifolia E1 . . . . . . + . . . . . . 1 8 Hacquetia epipactis E1 . . . . . . . . . . . . 1 1 8 EC Erythronio-Carpinion Primula vulgaris E1 + . . + + 1 + 1 1 . + . + 9 69 Helleborus odorus E1 . + . . . . 1 . 1 . 1 . . 4 31 Galanthus nivalis E1 1 1 . . . . . . . . 1 . . 3 23 Erythronium dens-canis E1 . . . . . . . . . . . . 1 1 8 TA Tilio-Acerion Acer pseudoplatanus E3b + 1 2 2 + . . . . . . . 2 6 46 Acer pseudoplatanus E3a + . 1 . + . . + . . . . . 4 31 Acer pseudoplatanus E2b . . . . . . . . . . . + . 1 8 Acer pseudoplatanus E2a . . + . . . . . + . . . . 2 15 Acer pseudoplatanus E1 + + + . 1 + . + + . . . . 7 54 Polystichum aculeatum E1 1 1 + . + . . + + . . . + 7 54 Ulmus glabra E3b . . . . . . . . . . . . + 1 8 Ulmus glabra E2a . . . . . . . + + . . . . 2 15 Ulmus glabra E1 . . . . . . . + . . . r + 3 23 Phyllitis scolopendrium E1 . + . . . . + + . . . . . 3 23 Acer platanoides E3b + + . . . . . . . . . . . 2 15 Acer platanoides E3a + . . . . . . + . . . . . 2 15 Acer platanoides E2b + . . . . . . . . . . . . 1 8 Acer platanoides E2a . . . . + . + . . . . . . 2 15 Acer platanoides E1 . + . . + . . . . . . . . 2 15 Tilia platyphyllos E3b . . . + . . + . . . . . . 2 15 Tilia platyphyllos E3a + . . . . . . . . . . . . 1 8 Tilia platyphyllos E1 + . . . . . . . . . . . . 1 8 Thalictrum aquilegiifolium E1 . . + . . . . . . . . . + 2 15 Tephroseris pseudocrispa E1 . . . . r . . + . . . . . 2 15 Lunaria rediviva E1 . + . . . . . . . . . . . 1 8 Scrophularia vernalis E1 . + . . . . . . . . . . . 1 8 Adoxa moschatellina E1 . + . . . . . . . . . . . 1 8 Euonymus latifolia E2b . . . . . . + . . . . . . 1 8 Juglans regia E2a . . . . . . . . . . . . + 1 8 FS Fagetalia sylvaticae Fagus sylvatica E3b 4 4 3 3 5 5 4 4 4 4 5 5 3 13 100 Fagus sylvatica E3a + + 1 . + 1 1 . . . 1 1 1 9 69 Fagus sylvatica E2b + . + + . 1 + 1 + . 1 1 1 10 77 Fagus sylvatica E2a . . . . 1 . + . . . 1 . + 4 31 Fagus sylvatica E1 . + . . . + . . . . + . . 3 23 Dryopteris filix-mas E1 + 1 + + + + + + + . + 1 + 12 92 Mercurialis perennis E1 1 1 + 1 1 + 1 + + . . + 1 11 85 Lathyrus vernus subsp. vernus E1 . + 1 + . + + + 1 + + . + 10 77 Salvia glutinosa E1 + + 1 + 1 + + . + . + . + 10 77 Asarum europaeum subsp. caucasicum E1 1 + . . 1 . 1 1 + 1 2 . + 9 69 Galium laevigatum E1 + . 1 . 1 . 1 . + + 1 + 1 9 69 Daphne mezereum E2a + . + + 1 . + + . . . + + 8 62 Melica nutans E1 + . + . . + + + . . . . + 6 46 Actaea spicata E1 + + . . . . + 1 + . . . . 5 38 Lonicera alpigena E2 + . . + 2 . + . . . . + . 5 38 Mycelis muralis E1 + + . + . + . . . . + . . 5 38 Neottia nidus-avis E1 . . + + . . . . + + + . . 5 38 Galeobdolon flavidum E1 . + . . + + . . . . . . + 4 31 Laburnum alpinum E3b . . . . . . . + . . . . . 1 8 Laburnum alpinum E3a r . . . . . + . . + . r . 4 31 Laburnum alpinum E2b . . . . + . . . . . . . . 1 8 Laburnum alpinum E2a . . . . 1 + + . . . . + . 4 31 Laburnum alpinum E1 . . . . + + . . . + . + . 4 31 Polygonatum multiflorum E1 + . . . + . . . + . . . + 4 31 Viola reichenbachiana E1 . . . . . + + + . . + . . 4 31 Cardamine bulbifera E1 + 1 . . . . . . . . 2 . . 3 23 Sambucus nigra E2b . . + . . . . . . . . . . 1 8 Sambucus nigra E2a + + . . . . . . . . + . . 3 23 Tilia cordata E3b . + 1 1 . . . . . . . . . 3 23 Tilia cordata E3a . . + . . . . + . . . . . 2 15 Tilia cordata E2b . . + . . . . + . . . . . 2 15 Tilia cordata E2a . . . . . . + . + . . . . 2 15 Tilia cordata E1 . . . . . . . . . . + . . 1 8 Euphorbia amygdaloides E1 . . 1 . . . . . . + . . + 3 23 Campanula trachelium E1 . . . . . . + . + . + . . 3 23 Fraxinus excelsior E3b . + . . . . . . . . . . + 2 15 Fraxinus excelsior E2a . . . . . . . . . . . . + 1 8 Fraxinus excelsior E1 + + . . . . . . . . . . . 2 15 Poa nemoralis E1 + . . . . . + . . . . . . 2 15 Scrophularia nodosa E1 . + . . . . . . . . + . . 2 15 Carpinus betulus E3a . r . . . . . . . . + . . 2 15 Euphorbia dulcis E1 . . . . . . + . . . 1 . . 2 15 Epilobium montanum E1 . + . . . . . . . . . . . 1 8 Festuca altissima E1 . + . . . . . . . . . . . 1 8 Galium odoratum E1 . + . . . . . . . . . . . 1 8 Paris quadrifolia E1 . + . . . . . . . . . . . 1 8 Prunus avium E2a . . + . . . . . . . . . . 1 8 Symphytum tuberosum E1 . . . + . . . . . . . . . 1 8 Lilium martagon E1 . . . . . . . . . . . . 1 1 8 Pulmonaria officinalis E1 . . . . . . . . . . . . 1 1 8 Allium ursinum E1 . . . . . . . . . . . . + 1 8 Carex sylvatica E1 . . . . . . . . . . . . + 1 8 Cephalanthera damasonium E1 . . . . . . . . . . . . + 1 8 Heracleum sphondylium E1 . . . . . . . . . . . . + 1 8 QP Quercetalia pubescenti-petraeae Fraxinus ornus E3a . . + + + + 1 + 1 1 . . + 9 69 Fraxinus ornus E2b . . + . 2 . + . . 1 . . . 4 31 Fraxinus ornus E2a . . + . 2 + + . . . . . . 4 31 Fraxinus ornus E1 + . . . + + . + . . . . . 4 31 Ostrya carpinifolia E3b 2 + 2 2 . + 2 . . . . + . 7 54 Ostrya carpinifolia E3a . . . . . + . 1 2 1 + . . 5 38 Ostrya carpinifolia E2b . . + . . . . . . . . . . 1 8 Sorbus aria E3b + + . . . . 1 . . . . . . 3 23 Sorbus aria E3a . . . . + . 1 + 1 + . + + 7 54 Sorbus aria E2b + . . . + + + . . . . + . 5 38 Sorbus aria E2a . . . . r + . + . . . . . 3 23 Sorbus aria E1 . . . . + + . . . + . . . 3 23 Melittis melissophyllum E1 . . . . + + + + 1 1 . . 1 7 54 Euonymus verrucosa E2a . . + . . . + + . + . . . 4 31 Tanacetum corymbosum E1 . . . . . + . + . + . . . 3 23 Primula x ternovania E1 . . . . . . . + 1 + . . . 3 23 Cephalanthera rubra E1 . . . . . + . . . . 1 . . 2 15 Campanula persicifolia E1 . . . . . . . + . + . . . 2 15 Cornus mas E3a . . . . . . . . . 1 . . . 1 8 Cornus mas E2b . . . . . . . . . 1 . . + 2 15 Cnidium silaifolium E1 . . . . . . + . . . . . . 1 8 Epipactis muelleri E1 . . . . . . . + . . . . . 1 8 Hypericum montanum E1 . . . . . . . + . . . . . 1 8 Sesleria autumnalis E1 . . . . . . . + . . . . . 1 8 Calamintha sylvatica E1 . . . . . . . . . + . . . 1 8 Piptatherum virescens E1 . . . . . . . . . . . . + 1 8 Asparagus tenuifolius E1 . . . . . . . . . . . . + 1 8 QR Quercetalia roboris Pteridium aquilinum E1 . . . . 1 . . + . + . . + 4 31 Serratula tinctoria E1 . . . . . . . + . + . . . 2 15 Quercus petraea E1 . . . . . . + . . . . . . 1 8 Hieracium racemosum E1 . . . . . . . . . . + . . 1 8 Betula pendula E3b . . . . . . . . . . . r . 1 8 Betonica officinalis E1 . . . . . . . . . . . . + 1 8 QF Querco-Fagetea Carex digitata E1 + . + . + + 1 + + + + + + 11 85 Hepatica nobilis E1 . . . + . + + 1 . 1 . . + 6 46 Hedera helix E3a . . . . . . . 1 . + . . . 2 15 Hedera helix E2b . . . . . . . . . . + . . 1 8 Hedera helix E1 . . . . . . + + + + . . + 5 38 Corylus avellana E2b . . . . . . . . . + . . + 2 15 Corylus avellana E2a . . r . + . . + + . . . . 4 31 Lonicera xylosteum E2a . . + . + . . + . + . . . 4 31 Festuca heterophylla E1 + . . . . . + + . . . . . 3 23 Clematis vitalba E2 . . . . + + . . . . . . + 3 23 Viburnum opulus E2a . . . . . . . + + + . . . 3 23 Anemone nemorosa E1 1 1 . . . . . . . . . . . 2 15 Viola mirabilis E1 . . . . . . . + . + . . . 2 15 Rubus caesius E2a . . 1 . . . . . . . . . . 1 8 Viola riviniana E1 . . . . + . . . . . . . . 1 8 Acer campestre E1 . . . . . . + . . . . . . 1 8 Cephalanthera longifolia E1 . . . . . . . . . + . . . 1 8 Taxus baccata E3a . . . . . . . . . . . . 2 1 8 Taxus baccata E2b . . . . . . . . . . . . 2 1 8 Taxus baccata E1 . . . . . . . . . . . . + 1 8 EP Erico-Pinetea Carex alba E1 . . + . + 1 1 2 2 2 . . 1 8 62 Calamagrostis varia E1 . . + . 2 + 2 + + 2 . . . 7 54 Cirsium erisithales E1 . . + . 1 + + . + . . r + 7 54 Aquilegia nigricans E1 . . + . . . + + 1 + . . . 5 38 Rhododendron hirsutum E2a . . + . + + . . + . . . . 4 31 Buphthalmum salicifolium E1 . . . . + . . . + + . . . 3 23 Peucedanum austriacum E1 . . . . + . 1 . . . . . . 2 15 Erica carnea E1 . . . . . . . . + + . . . 2 15 Polygala chamaebuxus E1 . . . . + . . . . . . . . 1 8 Pinus nigra E3a . . . . . . . . + . . . . 1 8 VP Vaccinio-Piceetea Veronica urticifolia E1 1 + + . 1 + 1 + + . . 1 + 10 77 Hieracium murorum E1 + . . . + + + + + + + . . 8 62 Solidago virgaurea E1 . . . . 1 + . + + + 1 + 1 8 62 Valeriana tripteris E1 . . 1 . + + + . + . . + + 7 54 Gentiana asclepiadea E1 . . . . + + . + + . . . 1 5 38 Homogyne sylvestris E1 . . . . 1 1 . + 1 . . . 1 5 38 Oxalis acetosella E1 + + . . . . + + . . . . + 5 38 Saxifraga cuneifolia E1 1 + . . . + 1 1 . . . . . 5 38 Aposeris foetida E1 . . . . . 1 + + . . . . + 4 31 Luzula luzuloides E1 + + . . . . 1 . . . . 1 . 4 31 Picea abies E3b r r . . r . . . . . . . 3 4 31 Picea abies E3a r r . . r . . . . . . . . 3 23 Picea abies E2b . . r . . . . . + . . . . 2 15 Picea abies E2a . . . . + . . . . . . . . 1 8 Calamagrostis arundinacea E1 + . . . . . + . . . . 1 . 3 23 Maianthemum bifolium E1 + . . . . . + . . . . . + 3 23 Vaccinium myrtillus E1 . . . . + . . . . . . . . 1 8 Huperzia selago E1 . . . . . . . . + . . . . 1 8 Abies alba E2b . . . . . . . . . . . r . 1 8 Abies alba E1 . . . . . . . . . . . r . 1 8 SSc Sambuco-Salicion capreae Sorbus aucuparia E2a . . . . . . . . . . . . + 1 8 RP Rhamno-Prunetea Cornus sanguinea E2b . . + . . . . . . + . . . 2 15 Crataegus monogyna E2b . . . . . . . . . + . . . 1 8 Euonymus europaea E2a . . + . . . . . . . . . . 1 8 Rubus fruticosus agg. E2a . . . . + . . . . . . . . 1 8 TG Trifolio-Geranietea Digitalis grandiflora E1 . . . . + + + + . . . . + 5 38 Verbascum lanatum E1 . . . + . . . . . . + . . 2 15 Anthericum ramosum E1 . . . . . . . . + + . . . 2 15 Thalictrum minus E1 . . + . . . . . . . . . . 1 8 Lilium carniolicum E1 . . . . . . . . + . . . . 1 8 Origanum vulgare E1 . . . . . . . . . + . . . 1 8 Vicia sylvatica E1 . . . . . . . . . . + . . 1 8 EA Epilobietea angustifolii Urtica dioica E1 . + . . . . . . . . . . . 1 8 Fragaria vesca E1 . . . . . + . . . . . . . 1 8 Galeopsis speciosa E1 . . . . . . . . . . + . . 1 8 BA Betulo-Alnetea viridis Salix appendiculata E2a . . + . . . . . . . . . . 1 8 Salix glabra E2a . . . . . . . . + . . . . 1 8 MuA Mulgedio-Aconitetea Senecio ovatus E1 1 1 + + + . + . . . + + . 8 62 Athyrium filix-femina E1 . 1 + + . . + . . . . . + 5 38 Aconitum lycoctonum s. lat. E1 . . + . . . + . . + . . . 3 23 Centaurea montana E1 . . + . . . . . . . . . + 2 15 Milium effusum E1 . + . . . . . . . . . . . 1 8 Saxifraga rotundifolia E1 . r . . . . . . . . . . . 1 8 Veratrum album s. lat. E1 . . . . . . . . . . . . + 1 8 ES Elyno-Seslerietea Phyteuma orbiculare E1 . . . . . + . . . . . . . 1 8 FB Festuco-Brometea Peucedanum oreoselinum E1 . . . . . . . . . + . . . 1 8 MA Molinio-Arrhenatheretea Ajuga reptans E1 . . . . . . . . . . . . + 1 8 Taraxacum officinale E1 . . . . . . . . . . + . . 1 8 Veronica chamaedrys E1 . . . . . . . . . . + . . 1 8 Cirsium oleraceum E1 . . . . . . . . . . . . r 1 8 TR Thlaspietea rotundifolii Adenostyles glabra E1 . . . . . . + . . . . 1 . 2 15 Cerastium subtriflorum E1 2 . . . . . . . . . . . . 1 8 AT Asplenietea trichomanis Asplenium trichomanes E1 + . + + + + 1 + . . + + . 9 69 Asplenium viride E1 + . + + + + . . + . . 1 . 7 54 Asplenium ruta-muraria E1 . . . + + + . + . + . + . 6 46 Polypodium vulgare E1 . . + . . + 1 . + + . . . 5 38 Moehringia muscosa E1 . . + + . . + . + . . . . 4 31 Cystopteris fragilis E1 + . . + . . . . . . . . . 2 15 Campanula carnica E1 . . . . . . + . . . . + . 2 15 Cardaminopsis arenosa E1 . . + . . . . . . . . . . 1 8 ML Mosses and lichens (Mahovi in lišaji) Ctenidium molluscum E0 1 + + + + 1 + . 1 1 . + . 10 77 Fissidens dubius E0 1 . + . . + + + + + . 1 + 9 69 Neckera crispa E0 + . . . + + + 2 1 + . + + 9 69 Polytrichum formosum E0 + . . . + + + . . + . 1 + 7 54 Isothecium alopecuroides E0 1 . . . + . . 1 + 1 + . . 6 46 Metzgeria furcata E0 . . . . . + + + + + . . . 5 38 Plagiothecium denticulatum E0 . . . . . + 1 + 1 . + . . 5 38 Schistidium apocarpum E0 . + + . . + . + . . + . . 5 38 Hypnum cupressiforme E0 + . . . . + 1 . . . . . + 4 31 Tortella tortuosa E0 . . . . . + . + + 1 . . . 4 31 Thuidium tamariscinum E0 . . . . + . 1 . . . . . + 3 23 Eurhynchium angustirete E0 . . . . . . + + + . . . . 3 23 Plagiochila porelloides E0 . . . . . . + + . . . 1 . 3 23 Dicranum scoparium E0 + . . . . . + . . . . . . 2 15 Peltigera canina E0 + . . . . . . + . . . . . 2 15 Atrichum undulatum E0 + . . . . . . . . . . + . 2 15 Bartramia halleriana E0 . . . . + . . . + . . . . 2 15 Plagiothecium nemorale E0 . . . . . + . . . . + . . 2 15 Neckera complanata E0 . . . . . . + . . . + . . 2 15 Leucobryum glaucum E0 . . . . . . . . . . . + + 2 15 Bryum capillare E0 + . . . . . . . . . . . . 1 8 Brachythecium rutabulum E0 . + . . . . . . . . . . . 1 8 Marchantia polymorpha E0 . . + . . . . . . . . . . 1 8 Mnium thomsonii E0 . . + . . . . . . . . . . 1 8 Isothecium myosuroides E0 . . . . . + . . . . . . . 1 8 Plagiomnium undulatum E0 . . . . . + . . . . . . . 1 8 Homalothecium sericeum E0 . . . . . . + . . . . . . 1 8 Cladonia furcata E0 . . . . . . + . . . . . . 1 8 Collema cristatum E0 . . . . . . . + . . . . . 1 8 Thuidium delicatulum E0 . . . . . . . + . . . . . 1 8 Hypnum cupressiforme var. filiforme E0 . . . . . . . . + . . . . 1 8 Thamnobryum alopecurum E0 . . . . . . . . + . . . . 1 8 Brachythecium velutinum E0 . . . . . . . . . . + . . 1 8 Porella platyphylla E0 . . . . . . . . . . + . . 1 8 Conocephalum conicum E0 . . . . . . . . . . . + . 1 8 Rhizomnium punctatum E0 . . . . . . . . . . . + . 1 8 Rhytidiadelphus triquetrus E0 . . . . . . . . . . . . + 1 8 Legend - Legenda AR Limestone with chert - apnenec z rožencem D Dolomite - dolomit DR Dolomite with chert - dolomit z rožencem DA Dolomite limestone - dolomitni apnenec Re Rendzina - rendzina Number of relevé (Zaporedna številka popisa) 1 2 3 4 6 5 7 8 9 10 11 12 13 Pr. Fr. Number of relevé (Zaporedna številka popisa) 1 2 3 4 6 5 7 8 9 10 11 12 13 Pr. Fr. Number of relevé (Zaporedna številka popisa) 1 2 3 4 6 5 7 8 9 10 11 12 13 Pr. Fr. Number of relevé (Zaporedna številka popisa) 1 2 3 4 6 5 7 8 9 10 11 12 13 Pr. Fr. Table 4 (Preglednica 4): Arunco-Fagetum, Dolenjska Number of relevé (Zaporedna številka popisa) 1 2 3 4 5 6 Database number of relevé (Delovna številka popisa) 235270 249876 248921 252685 249873 248462 Elevation in m (Nadmorska višina v m) 270 660 630 820 820 920 Aspect (Lega) N NW NE NW NW NE Slope in degrees (Nagib v stopinjah) 25 30 30 45 30 40 Parent material (Matična podlaga) D D D D D D Soil (Tla) Re Re Re Re Re Re Stoniness in % (Kamnitost v %) 5 5 5 5 5 20 Cover in % (Zastiranje v %): Upper tree layer (Zgornja drevesna plast) E3b 80 70 90 80 80 80 Lower tree layer (Spodnja drevesna plast) E3a 5 1 5 20 2 10 Shrub layer (Grmovna plast) E2 30 30 10 10 10 20 Herb layer (Zeliščna plast) E1 30 70 70 70 80 60 Moss layer (Mahovna plast) E0 5 5 10 5 5 10 Maximum diameter of trees (Največji prsni premer dreves) cm 60 26 55 60 60 35 Maximum height of trees (Največja drevesna višina) m 30 30 30 25 28 18 Number of species (Število vrst) 75 49 80 69 73 53 Relevé area (Velikost popisne ploskve) m2 900 400 600 400 400 200 Date of taking relevé (Datum popisa) 5/12/2010 5/22/2013 6/12/2013 5/22/2014 5/22/2013 5/28/2013 Locality (Nahajališče) Zasavska reber (Kum) Iška Kobile Kobile - Veliki Tisovec Iška Pšeničev vrh Quadrant (Kvadrant) 9856/3 0153/3 0258/1 0258/1 0153/3 0253/4 Coordinate GK Y (D-48) m 503755 462372 528931 529352 463030 471329 Coordinate GK X (D-48) m 5109052 5077329 5071347 5071392 5077042 5066911 Diagnostic species of the association (Diagnostične vrste asociacije) Pr. Fr. AF Helleborus niger E1 + 1 + + 1 1 6 100 QP Tamus communis E1 + + . + 1 . 4 67 FS Prenanthes purpurea E1 . . 1 + + 1 4 67 VP Rosa pendulina E2a . . + + + 1 4 67 MuA Phyteuma ovatum E1 . . 1 + + . 3 50 TA Aruncus dioicus E1 + . . . . . 1 17 QF Veratrum nigrum E1 + . . . . . 1 17 FS Epipactis helleborine E1 . . + . . . 1 17 QF Spiraea chamaedryfolia E2a . . + . . . 1 17 Geographical differential species (Geografske razlikovalne vrste) AF Ruscus hypoglossum E2a . . 1 . r . 2 33 AF Anemone trifolia E1 + . . . . . 1 17 Differential species of lower units (Razlikovalnice nižjih enot) VP Homogyne sylvestris E1 . . 1 + + 1 4 67 VP Gentiana asclepiadea E1 . . + + + + 4 67 AF Aremonio-Fagion Cyclamen purpurascens E1 + + + + + + 6 100 Cardamine enneaphyllos E1 1 1 1 1 1 . 5 83 Hacquetia epipactis E1 + 1 + + . . 4 67 Cardamine trifolia E1 1 . . . + + 3 50 Euphorbia carniolica E1 + . + . + . 3 50 Lamium orvala E1 1 1 . . + . 3 50 Epimedium alpinum E1 + . + . . . 2 33 Daphne laureola E2a . 1 . . + . 2 33 Potentilla carniolica E1 . . + 1 . . 2 33 Aremonia agrimonoides E1 . . + . + . 2 33 Omphalodes verna E1 . 1 . . . . 1 17 Calamintha grandiflora E1 . . . . 1 . 1 17 Vicia oroboides E1 . . . . + . 1 17 Knautia drymeia E1 . . . . . + 1 17 EC Erythronio-Carpinion Primula vulgaris E1 + + . . + . 3 50 Lonicera caprifolium E2a + . . . . . 1 17 TA Tilio-Acerion Acer pseudoplatanus E3b . + + 1 + . 4 67 Acer pseudoplatanus E3a 1 . + + + + 5 83 Acer pseudoplatanus E2b 1 . . + . . 2 33 Acer pseudoplatanus E2a . 1 . . + . 2 33 Acer pseudoplatanus E1 + 2 + 1 1 + 6 100 Polystichum aculeatum E1 1 1 2 1 + + 6 100 Ulmus glabra E3b . + . . + . 2 33 Ulmus glabra E3a + . . . . . 1 17 Ulmus glabra E2b . . + . . . 1 17 Ulmus glabra E2a . + . . . . 1 17 Ulmus glabra E1 + + + . + . 4 67 Tephroseris longifolia E1 + . . + + . 3 50 Acer platanoides E3b . . + + . . 2 33 Acer platanoides E3a + . . . . . 1 17 Acer platanoides E2b + . . . . . 1 17 Acer platanoides E1 + . + . . . 2 33 Arum maculatum E1 + + . . . . 2 33 Euonymus latifolia E2a . . + . + . 2 33 Staphylea pinnata E2 2 . . . . . 1 17 Lunaria rediviva E1 + . . . . . 1 17 Tilia platyphyllos E3b + . . . . . 1 17 Tilia platyphyllos E1 + . . . . . 1 17 Phyllitis scolopendrium E1 + . . . . . 1 17 Dryopteris affinis E1 . . + . . . 1 17 Polystichum x bicknellii E1 . . + . . . 1 17 Thalictrum aquilegiifolium E1 . . . + . . 1 17 FS Fagetalia sylvaticae Fagus sylvatica E3b 5 4 5 4 4 4 6 100 Fagus sylvatica E3a 1 . 1 + + + 5 83 Fagus sylvatica E2b . + 1 + + . 4 67 Fagus sylvatica E2a . 2 + 1 . + 4 67 Fagus sylvatica E1 . . . 1 + . 2 33 Mercurialis perennis E1 1 . 2 2 2 1 5 83 Galeobdolon flavidum E1 1 + + + + . 5 83 Polygonatum multiflorum E1 + + + + + . 5 83 Dryopteris filix-mas E1 + + + + + . 5 83 Lonicera alpigena E2a . 1 1 1 2 + 5 83 Symphytum tuberosum E1 + + . + 1 . 4 67 Neottia nidus-avis E1 + . + + + . 4 67 Daphne mezereum E2a . . + + + + 4 67 Sambucus nigra E2 1 + . + . . 3 50 Asarum europaeum subsp. europaeum E1 + . 1 . + . 3 50 Mycelis muralis E1 + . + . + . 3 50 Melica nutans E1 + . . . + + 3 50 Actaea spicata E1 . + + + . . 3 50 Lilium martagon E1 . + + . + . 3 50 Galium laevigatum E1 . + . . + + 3 50 Paris quadrifolia E1 . + + + . . 3 50 Cephalanthera damasonium E1 . . + + + . 3 50 Petasites albus E1 . . + + + . 3 50 Pulmonaria officinalis E1 1 . + . . . 2 33 Cardamine bulbifera E1 1 . . . + . 2 33 Scrophularia nodosa E1 + + . . . . 2 33 Heracleum sphondylium E1 + . . . + . 2 33 Allium ursinum E1 + . . + . . 2 33 Salvia glutinosa E1 . + . . 1 . 2 33 Sanicula europaea E1 . + . . 1 . 2 33 Euphorbia amygdaloides E1 . . + . + . 2 33 Ranunculus lanuginosus E1 + . . . . . 1 17 Campanula trachelium E1 + . . . . . 1 17 Leucojum vernum E1 . 1 . . . . 1 17 Asarum europaeum subsp. caucasicum E1 . 1 . . . . 1 17 Euphorbia dulcis E1 . + . . . . 1 17 Brachypodium sylvaticum E1 . . . + . . 1 17 Fraxinus excelsior E2a . . . + . . 1 17 Lathyrus vernus E1 . . . . + . 1 17 Cardamine pentaphyllos E1 . . . . r . 1 17 Phyteuma spicatum E1 . . . . . 1 1 17 QP Quercetalia pubescenti-petraeae Fraxinus ornus E3a + . . . . . 1 17 Fraxinus ornus E2b + . . + . . 2 33 Fraxinus ornus E2a . 1 + 1 . . 3 50 Fraxinus ornus E1 + . + . 1 . 3 50 Ostrya carpinifolia E3b + . + 1 . . 3 50 Sorbus aria E3b . . . + . . 1 17 Sorbus aria E3a . . . + . + 2 33 Sorbus aria E2b . . . . . + 1 17 Sorbus aria E2a . . + + . + 3 50 Sorbus aria E1 . . . . + . 1 17 Hypericum montanum E1 . . + + . . 2 33 Melittis melissophyllum E1 . . . + + . 2 33 Carex flacca E1 . . . + + . 2 33 Convallaria majalis E1 . . . + . + 2 33 Cornus mas E2b + . . . . . 1 17 Arabis turrita E1 + . . . . . 1 17 Cephalanthera rubra E1 . . + . . . 1 17 Clematis recta E1 . . + . . . 1 17 Tanacetum corymbosum E1 . . . + . . 1 17 Euonymus verrucosa E2a . . . . . + 1 17 QR Quercetalia roboris Pteridium aquilinum E1 . . . + + . 2 33 Quercus robur E3b + . . . . . 1 17 QF Querco-Fagetea Clematis vitalba E2a + . + + + . 4 67 Hepatica nobilis E1 . + 1 . + + 4 67 Platanthera bifolia E1 . . + + . + 3 50 Hedera helix E3a 1 . + . . . 2 33 Hedera helix E1 + . 1 . . . 2 33 Anemone nemorosa E1 . 1 . . 1 . 2 33 Lonicera xylosteum E2a . + + . . . 2 33 Rosa arvensis E2a . + . . + . 2 33 Galium sylvaticum E1 . . 1 1 . . 2 33 Cephalanthera longifolia E1 . . + + . . 2 33 Acer campestre E2b + . . . . . 1 17 Acer campestre E2a + . . . . . 1 17 Acer campestre E1 + . . . . . 1 17 Moehringia trinervia E1 + . . . . . 1 17 Viburnum opulus E2a + . . . . . 1 17 Listera ovata E1 r . . . . . 1 17 Carex digitata E1 . . 1 . . . 1 17 Ilex aquifolium E3a . . + . . . 1 17 Ilex aquifolium E2 . . 1 . . . 1 17 Corylus avellana E2 . . . + . . 1 17 Taxus baccata E3a . . . . . + 1 17 Taxus baccata E2b . . . . . + 1 17 EP Erico-Pinetea Carex alba E1 + + + . + 1 5 83 Cirsium erisithales E1 . + + + 1 + 5 83 Calamagrostis varia E1 . . + + . 1 3 50 Buphthalmum salicifolium E1 . . . + + + 3 50 Aquilegia nigricans E1 + . . . . . 1 17 Epipactis atrorubens E1 . . . + . . 1 17 Peucedanum austriacum E1 . . . + . . 1 17 Rubus saxatilis E1 . . . + . . 1 17 Erica carnea E1 . . . . . 2 1 17 VP Vaccinio-Piceetea Solidago virgaurea E1 + . + + + + 5 83 Aposeris foetida E1 . 1 . 1 + . 3 50 Valeriana tripteris E1 . . + . + 1 3 50 Saxifraga cuneifolia E1 + . r . . . 2 33 Dryopteris dilatata E1 + . r . . . 2 33 Picea abies E3b . . . . 1 1 2 33 Picea abies E3a . + . . + . 2 33 Picea abies E2b . + . . . + 2 33 Picea abies E2a . 1 . . . + 2 33 Picea abies E1 . + . . . . 1 17 Abies alba E3a . + . . . r 2 33 Hieracium murorum E1 . . + . . + 2 33 Luzula sylvatica E1 . . . . + 1 2 33 Oxalis acetosella E1 . + . . . . 1 17 Gymnocarpium dryopteris E1 . + . . . . 1 17 Laserpitium krapfii E1 . . 1 . . . 1 17 Vaccinium myrtillus E1 . . . . . 1 1 17 Veronica urticifolia E1 . . . . . 1 1 17 SSc Sambuco-Salicion capreae Sorbus aucuparia E2a . . . . . + 1 17 Sorbus aucuparia E1 . . . + + + 3 50 RP Rhamno-Prunetea Crataegus monogyna E2a + . . . . . 1 17 Ligustrum vulgare E2a + . . . . . 1 17 Viburnum lantana E2a . . . . . + 1 17 TG Trifolio-Geranietea Digitalis grandiflora E1 . + + . + . 3 50 Vincetoxicum hirundinaria E1 . . . + + . 2 33 Campanula rapunculoides E1 + . . . . . 1 17 Iris graminea E1 . . . + . . 1 17 Lilium carniolicum E1 . . . + . . 1 17 EA Epilobietea angustifolii Eupatorium cannabinum E1 . + + + . . 3 50 Stachys alpina E1 . + + . . . 2 33 Chelidonium majus E1 + . . . . . 1 17 Parietaria officinalis E1 1 . . . . . 1 17 Rubus idaeus E1 . . + . . . 1 17 Atropa bella-donna E1 . . + . . . 1 17 Bromopsis benekenii E1 . . . + . . 1 17 MuA Mulgedio-Aconitetea Senecio ovatus E1 + 1 . + + . 4 67 Aconitum degenii subsp. paniculatum E1 + . + + . . 3 50 Aconitum lycoctonum E1 + . + . + . 3 50 Athyrium filix-femina E1 + . . + . . 2 33 Veratrum album s. lat. E1 . 1 + . . . 2 33 Centaurea montana E1 . . + + . + 3 50 Silene dioica E1 + . . . . . 1 17 Crepis paludosa E1 . . . + . . 1 17 ES Elyno-Seslerietea Aster bellidiastrum E1 . . + . . + 2 33 Sesleria tenuifolia subsp. kalnikensis E1 . . . + . . 1 17 Phyteuma orbiculare E1 . . . . . + 1 17 MA Molinio-Arrhenatheretea Angelica sylvestris E1 + . . . . . 1 17 Ajuga reptans E1 . . . . + . 1 17 TR Thlaspietea rotundifolii Gymnocarpium robertianum E1 . . . + . + 2 33 AT Asplenietea trichomanis Moehringia muscosa E1 + . . . . + 2 33 Asplenium trichomanes E1 + . + . . . 2 33 Cardaminopsis arenosa E1 + . . . . . 1 17 Asplenium ruta-muraria E1 . . . . . + 1 17 Asplenium viride E1 . . . . . + 1 17 ML Mosses and lichens (Mahovi in lišaji) Ctenidium molluscum E0 . + + . + 2 4 67 Polytrichum formosum E0 . . + + + 1 4 67 Neckera crispa E0 . + . . . 2 2 33 Dicranum scoparium E0 . . + . . + 2 33 Thuidium tamariscinum E0 . . . . + + 2 33 Conocephalum conicum E0 + . . . . . 1 17 Homalothecium philippeanum E0 + . . . . . 1 17 Schistidium apocarpum E0 + . . . . . 1 17 Atrichum undulatum E0 . . + . . . 1 17 Fissidens dubius E0 . . + . . . 1 17 Plagiomnium undulatum E0 . . . . + . 1 17 Tortella tortuosa E0 . . . . . + 1 17 Peltigera leucophlebia E0 . . . . . + 1 17 Bartramia halleriana E0 . . . . . + 1 17 Legend - Legenda D Dolomite - dolomit Re Rendzina - rendzina Number of relevé (Zaporedna številka popisa) 1 2 3 4 5 6 Pr. Fr. Number of relevé (Zaporedna številka popisa) 1 2 3 4 5 6 Pr. Fr. Number of relevé (Zaporedna številka popisa) 1 2 3 4 5 6 Pr. Fr. Table 5 (Preglednica 5): Arunco-Fagetum var. geogr. Hiracium rotundatum Number of relevé (Zaporedna številka popisa) 1 2 3 Database number of relevé (Delovna številka popisa) 248419 248421 248420 Elevation in m (Nadmorska višina v m) 815 860 850 Aspect (Lega) NW N NW Slope in degrees (Nagib v stopinjah) 40 40 40 Parent material (Matična podlaga) Co Co Co Soil (Tla) Re Re Re Stoniness in % (Kamnitost v %) 20 5 20 Cover in % (Zastiranje v %): Upper tree layer (Zgornja drevesna plast) E3b 80 70 80 Lower tree layer (Spodnja drevesna plast) E3a 5 10 Shrub layer (Grmovna plast) E2 5 5 5 Herb layer (Zeliščna plast) E1 70 70 70 Moss layer (Mahovna plast) E0 5 5 10 Maximum diameter of trees (Največji prsni premer dreves) cm 100 70 90 Maximum height of trees (Največja drevesna višina) m 24 24 22 Number of species (Število vrst) 54 44 37 Relevé area (Velikost popisne ploskve) m2 400 400 400 Date of taking relevé (Datum popisa) 5/21/2013 5/21/2013 5/21/2013 Locality (Nahajališče) Donačka gora Donačka gora Donačka gora Quadrant (Kvadrant) 9760/1 9760/1 9760/1 Coordinate GK Y (D-48) m 557508 557683 557530 Coordinate GK X (D-48) m 5124572 5124605 5124545 Diagnostic species of the association (Diagnostične vrste asociacije) Pr. Fr. FS Mercurialis perennis E1 3 2 2 3 100 FS Festuca altissima E1 3 3 1 3 100 AF Cardamine enneaphyllos E1 2 1 2 3 100 FS Prenanthes purpurea E1 + 1 . 2 67 TA Aruncus dioicus E1 + . . 1 33 AF Helleborus niger E1 + . . 1 33 Geographical differential species (Geografske razlikovalne vrste) QR Hieracium rotundatum E1 + + . 2 67 QF Galium sylvaticum E1 . + + 2 67 FS Poa stiriaca E1 . + . 1 33 FB Sesleria sadlerana E1 . . 1 1 33 Differential species of lower units (Razlikovalnice nižjih enot) VP Homogyne sylvestris E1 r . . 1 33 VP Luzula luzuloides E1 + 3 1 3 100 FB Festuca pallens E1 . + + 2 67 VP Calamagrostis arundinacea E1 . . 1 1 33 AF Aremonio-Fagion Cyclamen purpurascens E1 + + + 3 100 Vicia oroboides E1 + . . 1 33 TA Tilio-Acerion Acer platanoides E1 + + . 2 67 Acer pseudoplatanus E1 + + . 2 67 Scrophularia vernalis E1 r . . 1 33 Tilia platyphyllos E3b + . . 1 33 Tilia platyphyllos E3a + . . 1 33 Tilia platyphyllos E1 + . . 1 33 Dryopteris affinis E1 + . . 1 33 Phyllitis scolopendrium E1 + . . 1 33 Geranium robertianum E1 . + . 1 33 FS Fagetalia sylvaticae Fagus sylvatica E3b 4 4 4 3 100 Fagus sylvatica E3a . . + 1 33 Fagus sylvatica E2a + . . 1 33 Fagus sylvatica E1 + + . 2 67 Phyteuma spicatum E1 1 1 1 3 100 Euphorbia dulcis E1 + + + 3 100 Dentaria bulbifera E1 1 . + 2 67 Galium odoratum E1 1 + . 2 67 Melica nutans E1 + . 1 2 67 Mycelis muralis E1 + + . 2 67 Dryopteris filix-mas E1 1 + . 2 67 Actaea spicata E1 1 . . 1 33 Salvia glutinosa E1 1 . . 1 33 Epilobium montanum E1 + . . 1 33 Galeobdolon flavidum E1 + . . 1 33 Myosotis sylvatica E1 + . . 1 33 Paris quadrifolia E1 + . . 1 33 Petasites albus E1 + . . 1 33 Polygonatum multiflorum E1 + . . 1 33 Sambucus nigra E2a + . . 1 33 Sanicula europaea E1 + . . 1 33 Symphytum tuberosum E1 + . . 1 33 Tilia cordata E3a . + . 1 33 Tilia cordata E2a + . . 1 33 Tilia cordata E1 . + . 1 33 Daphne mezereum E2a . + . 1 33 Heracleum sphondylium E1 . . + 1 33 Lathyrus vernus E1 . . + 1 33 Neottia nidus-avis E1 . . + 1 33 QP Quercetalia pubescenti-petraeae Campanula persicifolia E1 . 1 + 2 67 Tanacetum corymbosum E1 . + + 2 67 Arabis turrita E1 . + . 1 33 Ostrya carpinifolia E2a . . + 1 33 Ostrya carpinifolia E1 . + . 1 33 Sorbus aria E3a . r . 1 33 Sorbus aria E2b . . + 1 33 Hierochloë australis E1 . . + 1 33 Fraxinus ornus E1 . . r 1 33 QR Quercetalia roboris Hieracium praecurrens E1 + . . 1 33 Quercus petraea E1 . + . 1 33 QF Querco-Fagetea Hepatica nobilis E1 + . + 2 67 Festuca heterophylla E1 . + 1 2 67 Clematis vitalba E2a + . . 1 33 Moehringia trinervia E1 + . . 1 33 Anemone nemorosa E1 + . . 1 33 Stellaria holostea E1 . + . 1 33 Carex digitata E1 . . 1 1 33 Viola riviniana E1 . . + 1 33 EP Erico-Pinetea Calamagrostis varia E1 . . 1 1 33 VP Vaccinio-Piceetea Gentiana asclepiadea E1 + + . 2 67 Aposeris foetida E1 + . + 2 67 Hieracium murorum E1 + . . 1 33 Oxalis acetosella E1 + . . 1 33 Maianthemum bifolium E1 . + . 1 33 Vaccinium myrtillus E1 . + . 1 33 Avenella flexuosa (Deschampsia flexuosa) E1 . . + 1 33 TG Trifolio-Geranietea Anthericum ramosum E1 . . + 1 33 Iris graminea E1 . . + 1 33 EA Epilobietea angustifolii Rubus idaeus E1 + . + 2 67 Urtica dioica E1 + . . 1 33 MuA Mulgedio-Aconitetea Senecio ovatus E1 + + . 2 67 Doronicum austriacum E1 1 . . 1 33 Silene dioica E1 + . . 1 33 Athyrium filix-femina E1 + . . 1 33 Milium effusum E1 . + . 1 33 ES Elyno-Seslerietea Aster bellidiastrum E1 . + . 1 33 FB Festuco-Brometea Allium oleraceum E1 . + . 1 33 MA Molinio-Arrhenatheretea Taraxacum officinale E1 . + . 1 33 TR Thlaspietea rotundifolii Hieracium bifidum E1 . + + 2 67 AT Asplenieta trichomanis Asplenium trichomanes E1 + + . 2 67 Campanula rotundifolia agg. E1 . + . 1 33 Cardaminopsis arenosa E1 . + . 1 33 ML Mosses and lichens (Mahovi in lišaji) Isothecium alopecuroides E0 + . . 1 33 Bryum capillare E0 . + . 1 33 Hypnum cupressiforme E0 . + . 1 33 Polytrichum formosum E0 . + . 1 33 Ctenidium molluscum E0 . . + 1 33 Schistidium apocarpum E0 . . + 1 33 Tortella tortuosa E0 . . + 1 33 Legend - Legenda Co Quartz conglomerate - kremenov konglomerat Re Rendzina - rendzina Number of relevé (Zaporedna številka popisa) 1 2 3 Pr. Fr. Number of relevé (Zaporedna številka popisa) 1 2 3 Pr. Fr. Table 6 (Preglednica 6): Saxifrago cuneifolii-Fagetum aruncetosum Number of relevé (Zaporedna številka popisa) 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 Database number of relevé (Delovna številka popisa) 255377 255385 255378 255384 255387 255380 255381 255383 255379 255386 255504 255388 255499 255403 255500 255502 255446 Elevation in m (Nadmorska višina v m) 1090 970 1050 960 940 1170 1210 1160 1030 870 740 580 740 600 750 420 560 Aspect (Lega) E NE NE NE NE NE NE NNE SE SE N NE NW NNE N NW N Slope in degrees (Nagib v stopinjah) 40 35 40 35 40 35 35 35 40 35 35 35 35 35 35 35 30 Parent material (Matična podlaga) DR DR DR DR DR DR DR DR DR DR DR DR DR ALR DR DR DR Soil (Tla) Re Re Re Re Re Re Re Re Re Re Re Re Re Re Re Re Re Stoniness in % (Kamnitost v %) 20 20 25 30 10 30 10 30 20 20 5 10 20 40 50 5 30 Cover in % (Zastiranje v %): Upper tree layer (Zgornja drevesna plast) E3b 90 80 90 90 90 80 85 90 90 90 90 80 90 90 90 70 90 Lower tree layer (Spodnja drevesna plast) E3a 20 10 10 10 5 10 10 20 Shrub layer (Grmovna plast) E2 1 5 2 10 5 10 5 10 2 5 20 5 5 10 5 20 5 Herb layer (Zeliščna plast) E1 70 60 70 60 60 80 75 70 60 50 70 40 40 50 50 60 60 Moss layer (Mahovna plast) E0 5 10 5 10 10 10 5 20 5 5 20 5 5 10 10 10 20 Maximum diameter of trees (Največji prsni premer dreves) cm 35 35 45 40 35 35 40 35 45 50 35 40 70 50 45 40 35 Maximum height of trees (Največja drevesna višina) m 20 20 20 20 22 18 16 15 20 24 20 24 28 26 25 24 22 Number of species (Število vrst) 70 61 79 74 68 74 70 78 66 72 67 53 74 75 60 88 47 Relevé area (Velikost popisne ploskve) m2 400 400 400 400 400 400 400 400 400 400 400 400 400 400 400 400 400 Date of taking relevé (Datum popisa) 5. 7. 1990 7/5/1990 5. 7. 1990 7/5/1990 7/21/1989 7/5/1990 7/5/1990 7/5/1990 7/5/1990 7/7/1990 7/23/1991 5/24/1988 6/1/2000 5/18/1999 6/1/2000 5/22/1989 5/6/1987 Locality (Nahajališče) Krikov vrh- Mirna grapa Krikov vrh - Mirna grapa Krikov vrh - Mirna grapa Krikov vrh - Mirna grapa Krikov vrh - Mirna grapa Krikov vrh Krikov vrh Krikov vrh Krikov vrh - Mirna grapa Krikov vrh - Huda grapa Kojca Loje - Osoje Temljine - Kotel Temna brda Temljine - Kotel Podleskovca Hudajužna - Kobilica Quadrant (Kvadrant) 9748/4 9748/4 9748/4 9748/4 9748/4 9748/4 9748/4 9748/4 9748/4 9748/4 9849/2 9849/1 9849/1 9748/4 9849/1 9849/1 9849/2 Coordinate GK Y (D-48) m 406718 406921 406832 406961 407048 406583 406541 406703 406865 407045 416594 410306 411056 408891 411065 411359 416361 Coordinate GK X (D-48) m 5118877 5118896 5118878 5118835 5118820 5118895 5118859 5118724 5118918 5118490 5113572 5116771 5114342 5118055 5114709 5115942 5114114 Diagnostic species of the association (Diagnostične vrste asociacije) Pr. Fr. VP Veronica urticifolia E1 + . + + + + + + . + 1 + r + + + + 15 88 VP Calamagrostis arundinacea E1 . . + + + 1 1 1 + + 2 + 1 + + . . 13 76 VP Saxifraga cuneifolia E1 + . + + + . + . . . 1 + . r 1 . . 9 53 VP Luzula luzuloides E1 + . . + . + 1 . . . 1 + 1 + + . . 9 53 QP Tanacetum corymbosum E1 + . + + . + . . . . . + + . + . . 7 41 TG Clinopodium vulgare E1 . . . . . + + + + . . . . . . . . 4 24 QF Festuca heterophylla E1 . . . . . + + . . . + . . . . . . 3 18 VP Huperzia selago E1 . . . . . . . . . . + . . . . . . 1 6 Geographical differential species (Geografske razlikovalne vrste) AF Anemone trifolia E1 1 1 + . 1 + . . + 1 . 1 1 1 + 1 1 13 76 QP Sesleria autumnalis E1 + + + + + + + . 1 1 . + . r . 1 . 12 71 FS Lathyrus vernus subsp. flaccidus E1 . . . + . + . . + + . . . . . . . 4 24 MuA Aconitum angustifolium E1 . . . . + . + . . + . . . . . + . 4 24 Differential species of the subassociation (Razlikovalnice subasociacije) FS Festuca altissima E1 1 1 1 1 1 1 1 1 + . 1 . . . + . + 12 71 TA Aruncus dioicus E1 1 . + + . + + + . . + + + + + + . 12 71 VP Laserpitium krapfii E1 + + + + + . . + 1 + . . . . . . . 8 47 QF Veratrum nigrum E1 . + + + 1 . . + + + . + . . . . . 8 47 Differential species of lower units (Razlikovalnice nižjih enot) MuA Veratrum album s. lat. E1 1 1 1 + . 2 2 2 . . . . . . . . . 7 41 MuA Aconitum degenii subsp. paniculatum E1 . + + . . + + . + r . . . . . . . 6 35 FS Asarum europaeum subsp. caucasicum E1 . . . . . . . . . + . 1 + 1 . 1 1 6 35 FS Galium odoratum E1 . . . . . . . . . . + + + 1 + . . 5 29 QF Hedera helix E3a . . . . . . . . . . . . . . . 1 . 1 6 QF Hedera helix E2b . . . . . . . . . . . + . . . . . 1 6 QF Hedera helix E1 . . . . . . . . . . + . + r + . + 5 29 EC Helleborus odorus E1 . . . . . . . . . . . + 1 . . 1 . 3 18 AF Aremonio-Fagion Cardamine enneaphyllos E1 2 2 1 2 2 1 2 1 1 2 2 1 1 2 1 2 3 17 100 Cyclamen purpurascens E1 1 1 1 + 1 1 . 1 1 1 1 1 1 1 . 1 1 15 88 Cardamine trifolia E1 1 1 + 1 + 1 1 1 . . 1 1 . + 1 + 2 14 82 Lamium orvala E1 + + + . . . . . . 1 . + 1 2 + + + 10 59 Anemone x pittonii E1 . . . . . . . . . + . . . . . . . 1 6 Hacquetia epipactis E1 . . . . . . . . . . . . . . . + . 1 6 EC Erythronio-Carpinion Primula vulgaris E1 . + . + + + . . + + + + . . . + . 9 53 TA Tilio-Acerion Acer pseudoplatanus E3b . + + + 1 . . 1 . . . . . . . . . 5 29 Acer pseudoplatanus E3a + . . + . + . + + . . . . . . . . 5 29 Acer pseudoplatanus E2b . . . + . . . . + . . . . . . + . 3 18 Acer pseudoplatanus E2a . . . . + + . . . . . . . . . . . 2 12 Acer pseudoplatanus E1 1 1 + 1 1 1 . + 1 + + . + + + . + 14 82 Polystichum aculeatum E1 + + + 1 + + + + + + . . . + + + + 14 82 Geranium robertianum E1 + . + . . . . . + + . . . . . . . 4 24 Thalictrum aquilegiifolium E1 + . . . . + + + . . . . . . . . . 4 24 Dryopteris affinis E1 . . . . . . . . . . . + . + . + . 3 18 Polystichum setiferum E1 . . . . 1 . . . . . . . . + . . . 2 12 Polystichum x illyricum E1 . . . . . . . . . + . . r . . . . 2 12 Adoxa moschatellina E1 . . . . . . . . . . . . . 1 + . . 2 12 Stellaria montana E1 . . . . . . . . . . . . . 1 + . . 2 12 Phyllitis scolopendrium E1 . . . . . . . . . . . . . + + . . 2 12 Polystichum braunii E1 . . . . . . . . . . . . . . + . + 2 12 Acer platanoides E3b . . . . . . . . . . . . . r . . . 1 6 Acer platanoides E3a . . . . . . . . . . . . . . . + . 1 6 Acer platanoides E2a . . . . . . . . . . . + . . . . . 1 6 Acer platanoides E1 . . . . . . . . . . . 1 . . . . . 1 6 Ulmus glabra E3b . . . . . . . . . . . . + . . . . 1 6 Juglans regia E1 . . . . . . . . . . . . r . . . . 1 6 Arum maculatum E1 . . . . . . . . . . . . . + . . . 1 6 Impatiens noli-tangere E1 . . . . . . . . . . . . . + . . . 1 6 Tephroseris pseudocrispa E1 . . . . . . . . . . . . . . . + . 1 6 FS Fagetalia sylvaticae Fagus sylvatica E3b 5 5 5 5 5 5 5 5 5 5 5 5 5 5 5 5 5 17 100 Fagus sylvatica E3a 1 1 1 1 . 1 1 1 + + 1 . + 1 + 1 . 14 82 Fagus sylvatica E2b . . + 1 1 1 1 . + + + . + 1 + 1 + 13 76 Fagus sylvatica E2a + + . . . . . . . . + 1 . + . . . 5 29 Fagus sylvatica E1 . + . . . . . . . . 1 + 1 + + 1 + 8 47 Actaea spicata E1 1 1 1 1 1 1 + 1 1 + + + + 1 + + + 17 100 Lathyrus vernus subsp. vernus E1 1 1 1 1 1 1 1 1 1 1 1 + 1 + + 1 + 17 100 Galium laevigatum E1 1 1 1 1 1 1 + 1 1 + 1 + + + + 1 . 16 94 Dryopteris filix-mas E1 1 1 1 + 1 2 1 2 1 + 1 + 1 1 1 + . 16 94 Prenanthes purpurea E1 1 1 1 1 1 1 1 + 1 + 1 1 1 1 + + . 16 94 Mercurialis perennis E1 2 2 2 2 1 1 1 2 3 1 + + . 1 . + + 15 88 Daphne mezereum E2a + + + 1 1 1 + + + + + + . + . + . 14 82 Campanula trachelium E1 + + + + + + + + 1 . + + + . . + . 13 76 Pulmonaria officinalis E1 + + + + + + . + . + + . + + + 1 . 13 76 Symphytum tuberosum E1 + . + + + + . . + + 1 + + + + + . 13 76 Euphorbia amygdaloides E1 + + 1 + . 1 . 1 1 + . + + . . + + 12 71 Galeobdolon flavidum E1 + . + . + + 1 1 1 + . . 1 + 1 . + 12 71 Heracleum sphondylium E1 + + + + + + + + + + . . + . . + . 12 71 Neottia nidus-avis E1 + + + + . . + + + + . + + . + . . 11 65 Salvia glutinosa E1 . . + . + + . + + + + + + + . + . 11 65 Lilium martagon E1 + + 1 + 1 1 + 1 . . . . r . . . . 9 53 Paris quadrifolia E1 + . + + + . + . + . . . . + + . + 9 53 Sanicula europaea E1 + + 1 + + 1 . + . . . . . + . + . 9 53 Mycelis muralis E1 + . + + . . + . + + . . + . + . . 8 47 Epipactis helleborine E1 + 1 + + + + + . 1 . . . . . . . . 8 47 Scrophularia nodosa E1 + . . . . + + . + + . . + + + . . 8 47 Lonicera alpigena E2 . . + + 1 . + + + . + . . + . . . 8 47 Polygonatum multiflorum E1 + . . + . . . + + + + . . + . . . 7 41 Cardamine bulbifera E1 . . . . . . . . + + . . 1 + + + . 6 35 Petasites albus E1 . . + . . . . . . . + . . + + . . 4 24 Ranunculus lanuginosus E1 . . + . . 1 + . . . . . . . + . . 4 24 Sambucus nigra E2b . . . + . . . . . . . . . . . + + 3 18 Sambucus nigra E2a . . . . . . . . . + . + . + + . . 4 24 Cephalanthera damasonium E1 . . . . + . . . . + . . . . . + . 3 18 Carex sylvatica E1 . . . . . + + + . . . . . . . . . 3 18 Carpinus betulus E3b . . . . . . . . . . . . + + . . . 2 12 Carpinus betulus E3a . . . . . . . . . . . . r + . 1 . 3 18 Laburnum alpinum E3a . . . . . + + . . . . . . . . . . 2 12 Laburnum alpinum E2b . . . . . + . . . + . . . . . . . 2 12 Laburnum alpinum E1 . . . + + . . . . . . . . . . . . 2 12 Epilobium montanum E1 . . . . . . + . . . . . . . + . . 2 12 Cardamine impatiens E1 . . . . . . . . . . . . + . + . . 2 12 Melica nutans E1 . . . . . . . . . . . . + . . . + 2 12 Tilia cordata E3a . . . . . . . . . . . . . + . . . 1 6 Tilia cordata E2a . . . . . . . . . . . . . . . + . 1 6 Tilia cordata E1 . . . . . . . . . . . . r . . + . 2 12 Myosotis sylvatica E1 + . . . . . . . . . . . . . . . . 1 6 Cardamine pentaphyllos E1 . . + . . . . . . . . . . . . . . 1 6 Epipactis leptochila E1 . . . . + . . . . . . . . . . . . 1 6 Viola reichenbachiana E1 . . . . . . . . . . . . + . . . . 1 6 Corydalis cava E1 . . . . . . . . . . . . . + . . . 1 6 Prunus avium E3b . . . . . . . . . . . . . . . . + 1 6 Prunus avium E1 . . . . . . . . . . . . . . . + . 1 6 Euphorbia dulcis E1 . . . . . . . . . . . . . . . + . 1 6 QP Quercetalia pubescenti-petraeae Melittis melissophyllum E1 . . + . . . . + . . + + + . . + . 6 35 Ostrya carpinifolia E3b . . . . + . . . . . . . + + r . + 5 29 Ostrya carpinifolia E3a . . . . . + . + . . . + r + . + . 6 35 Arabis turrita E1 + . . . . . . . + + . . r . . . . 4 24 Fraxinus ornus E3a . . . . . . . . . . . . . . . . + 1 6 Fraxinus ornus E2b . . . . . . . . . . + . . . . + + 3 18 Fraxinus ornus E2a . . . . . . . . . + + . . . . . . 2 12 Fraxinus ornus E1 . . . . . . . . . . . . + . . . + 2 12 Sorbus aria E3b . + . . + . . . . . . . . . . . . 2 12 Sorbus aria E2b . . . . + . . . . . . . . . . . . 1 6 Sorbus aria E2a . . . . . + + . . . + . . . . . . 3 18 Hypericum montanum E1 . + . . . . . . + . . . . . . . . 2 12 Epipactis muelleri E1 . + . . . . . . . . . . . . . . . 1 6 Calamintha sylvatica E1 . . . . . . . . + . . . . . . . . 1 6 Cornus mas E2b . . . . . . . . . . . . . . . + . 1 6 Euonymus verrucosa E2a . . . . . . . . . . . . . . . + . 1 6 Ruscus aculeatus E1 . . . . . . . . . . . . . . . + . 1 6 QR Quercetalia roboris Hieracium racemosum E1 . . . . . . . . . . + . + . . . . 2 12 Rubus hirtus E2a . . . . . . . . . . . . + . . . . 1 6 Pteridium aquilinum E1 . . . . . . . . . . . . + . . . . 1 6 QF Querco-Fagetea Carex digitata E1 + + + + 1 + + + + + + . + + . + . 14 82 Anemone nemorosa E1 1 1 1 1 1 1 1 1 . . . . . 1 + . . 10 59 Clematis vitalba E2 . . . . . . . . + + . . + . . + + 5 29 Moehringia trinervia E1 . . . . . . . . . + . . + . + . + 4 24 Aegopodium podagraria E1 . . . . . . . . . . . + r + . 1 . 4 24 Corylus avellana E2b . . . . . . . . . . . . r + . . . 2 12 Corylus avellana E2a . . . . . . . . . + + . . . . + . 3 18 Corylus avellana E1 . . . . . . . . . . . . . . . + . 1 6 Platanthera bifolia E1 . . + . . . . + . . + . . . . . . 3 18 Hepatica nobilis E1 . . + . . . . + . . . . . . . . . 2 12 Dactylorhiza fuchsii E1 . . . . . + + . . . . . . . . . . 2 12 Rosa arvensis E2a . . . . . . . . . . . . r . . + . 2 12 Anemone ranunculoides E1 + . . . . . . . . . . . . . . . . 1 6 Lonicera xylosteum E2a . . . . . . . . . . + . . . . . . 1 6 Taxus baccata E3a . . . . . . . . . . . . . . . 1 . 1 6 Taxus baccata E2b . . . . . . . . . . . . . . . 1 . 1 6 Vinca minor E1 . . . . . . . . . . . . . . . 1 . 1 6 Acer campestre E2a . . . . . . . . . . . . . . . + . 1 6 Acer campestre E1 . . . . . . . . . . . . . . . + . 1 6 EP Erico-Pinetea Cirsium erisithales E1 + + + + + + + + + + + . . + . + . 13 76 Aquilegia nigricans E1 . . + . . + + + . + . . . . . . . 5 29 Calamagrostis varia E1 . 1 + . . . . . + + . . . . . + . 5 29 Carex alba E1 + + . + . . . . + . . . + . . . . 5 29 Buphthalmum salicifolium E1 . . . . . . . . + . . . . . . . . 1 6 Carex ornithopoda E1 . . . . . . . . + . . . . . . . . 1 6 VP Vaccinio-Piceetea Oxalis acetosella E1 1 1 1 1 + 1 1 1 + . 1 + . 1 1 + + 15 88 Solidago virgaurea E1 + 1 + . . + 1 + . + + + + . . + . 11 65 Gentiana asclepiadea E1 + + + . 1 + 1 + . . + + . . + . . 10 59 Hieracium murorum E1 + + + + . + + . . . + . + . . . . 8 47 Gymnocarpium dryopteris E1 . . . . + + + + . . + . . + + . + 8 47 Homogyne sylvestris E1 . + + + 1 . . + . . 1 . . . . . + 7 41 Picea abies E3a . . . . . + . . . . . . r . . . + 3 18 Picea abies E2b . . . . . . . . . . . . . . . . + 1 6 Picea abies E2a + . . . . + + + . . 2 . . . . + . 6 35 Picea abies E1 . . . . + . . . . . 1 . . . . . . 2 12 Maianthemum bifolium E1 + + . . . + . . . . 1 . . + . . . 5 29 Valeriana tripteris E1 + . . . + . . + . + 1 . . . . . . 5 29 Rosa pendulina E2a . + + . + . + + . . . . . . . . . 5 29 Aposeris foetida E1 . . . . . . . . . . . 1 1 . . + . 3 18 Abies alba E3a . . + + . . . . . . . . . . . . . 2 12 Abies alba E2b . . . . . . . . . . . . . r . . . 1 6 Abies alba E2a . . . + . . . . . + . . . . . . . 2 12 Dryopteris dilatata E1 . . . . . . . . . . . . . r . . + 2 12 SSC Sambuco-Salicion capreae Salix caprea E3a . . . . . . + + . . . . . . . . . 2 12 RP Rhamno-Prunetea Crataegus monogyna E2b . . . . . . . . . . . . . . . + . 1 6 TG Trifolio-Geranietea Campanula rapunculoides E1 + . . + . . . . + + . + r + . + . 8 47 Verbascum lanatum E1 . . . . . + + + + + . . . . . . . 5 29 Digitalis grandiflora E1 . + + + . . . . . . + . . . . . . 4 24 Origanum vulgare E1 . . . . . . + . + . . . . . . . . 2 12 Vincetoxicum hirundinaria E1 . . . . . . . . . . . . r . . . . 1 6 Lilium carniolicum E1 . . . . . . . . . . . . . . . + . 1 6 EA Epilobietea angustifolii Rubus idaeus E2a + . + + . 1 + + . + . . . . . . . 7 41 Hypericum hirsutum E1 . . + + . + . + + . . . . . . . . 5 29 Stachys sylvatica E1 . . + . . + . . + . . . . . . . . 3 18 Urtica dioica E1 . . . . . . . . . . . . . + + . . 2 12 Galeopsis speciosa E1 . . . + . . . . . . . . . . . . . 1 6 Bromopsis benekenii E1 . . . . . . . + . . . . . . . . . 1 6 Arctium nemorosum E1 . . . . . . . . + . . . . . . . . 1 6 MuA Mulgedio-Aconitetea Senecio ovatus E1 1 + 1 + 1 1 1 1 1 1 + . + 1 1 + + 16 94 Athyrium filix-femina E1 . . + 1 + 1 1 1 + . + + 1 + 1 + + 14 82 Phyteuma ovatum E1 1 + 1 + + . + + + . 1 + . . . . . 10 59 Aconitum lycoctonum s. lat. E1 . . . . . 1 + + . . . + . . . + . 5 29 Silene dioica E1 + . . . . . . . + + . . . . . . . 3 18 Polygonatum verticillatum E1 . . + . . . + + . . . . . . . . . 3 18 Milium effusum E1 . . . . . + . . . . . . . . . . . 1 6 Ranunculus platanifolius E1 . . . . . . + . . . . . . . . . . 1 6 Centaurea montana E1 . . . . . . . + . . . . . . . . . 1 6 Doronicum austriacum E1 . . . . . . . . . . . . . + . . . 1 6 FB Festuco-Brometea Pimpinella saxifraga E1 . . . . . . . . . . . . . . . + . 1 6 MA Molinio-Arrhenatheretea Veronica chamaedrys E1 . . . . . . . . . . . . + . . + . 2 12 Dactylis glomerata s.str. E1 . . . . . . . + . . . . . . . . . 1 6 TR Thlaspietea rotundifolii Adenostyles glabra E1 1 1 + + + 1 1 1 + r 1 . . . + . . 12 71 Gymnocarpium robertianum E1 . . . . + . . . . . . . . . . . . 1 6 AT Asplenietea trichomanis Asplenium trichomanes E1 + + . + . . . + + + . + + + + + . 11 65 Polypodium vulgare E1 . . . + + . . . . . + + + + + + . 8 47 Asplenium viride E1 + . . . + + . + . + . . . . + . + 7 41 Asplenium ruta-muraria E1 . + . . . . . + . + . . . . . + . 4 24 Carex brachystachys E1 . . . . . . . + . . . . . . . . . 1 6 Asplenium adiantum-nigrum E1 . . . . . . . . . . . . r . . . . 1 6 Moehringia muscosa E1 . . . . . . . . . . . . . . . . + 1 6 ML Mosses and lichens (Mahovi in lišaji) Polytrichum formosum E0 + + + + + + + + + + 1 + + . + + 1 16 94 Plagiothecium denticulatum E0 + 1 + + + + + + + + + . . + + 1 1 15 88 Ctenidium molluscum E0 1 . 1 1 + 1 1 1 1 . 1 + . + 1 1 1 14 82 Fissidens dubius E0 + + + 1 + 1 + + + + 1 . . . . . . 11 65 Neckera crispa E0 + 1 . + + . . + . . 1 + . + + + + 11 65 Schistidium apocarpum E0 + 1 . 1 + . + + 1 + . + . + . + . 11 65 Isothecium alopecuroides E0 . + + + + . . . . + + . + + 1 + . 10 59 Tortella tortuosa E0 + + + + + + + . + + . . . . . . . 9 53 Hypnum cupressiforme E0 . . + + + + . + . . 1 . + . + . . 8 47 Metzgeria furcata E0 . . + + + . . + . . 1 . . . + + . 7 41 Plagiochila porelloides E0 + . . . . + . . . . . . . . . + . 3 18 Brachythecium velutinum E0 . + . . + . . . . . . . . . . + . 3 18 Peltigera canina E0 . + . . . . . + . . . . + . . . . 3 18 Isothecium myosuroides E0 . . + + . . + . . . . . . . . . . 3 18 Dicranum scoparium E0 . . + . . . . . . . 2 . . . . . + 3 18 Collema cristatum E0 . . . . . . . . . + . . . + . + . 3 18 Anomodon attenuatus E0 . . . . . . . . . + . . + . . + . 3 18 Thuidium tamariscinum E0 . . . . . . . . . . 2 . . . . + + 3 18 Bartramia halleriana E0 . . . . . . . . . . + . . . . + + 3 18 Atrichum undulatum E0 . . . . . . . . . . . + + . . + . 3 18 Brachythecium rutabulum E0 . . . . . . . . . . . . . + . + + 3 18 Thamnobryum alopecurum E0 . . . . . . . . . . . . . + . + + 3 18 Cladonia furcata E0 . + . + . . . . . . . . . . . . . 2 12 Mnium thomsonii E0 . . . + . . . . . . . . . . + . . 2 12 Plagiomnium undulatum E0 . . . . . . + . . . . . . + . . . 2 12 Bryum capillare E0 . . . . . . . . . + . . . . . . + 2 12 Plagiothecium nemorale E0 . . . . . . . . . . . + . . . . + 2 12 Platygyrium repens E0 . . . + . . . . . . . . . . . . . 1 6 Encalypta streptocarpa E0 . . . . . + . . . . . . . . . . . 1 6 Plagiomnium cuspidatum E0 . . . . . . . + . . . . . . . . . 1 6 Brachythecium oxycladum E0 . . . . . . . + . . . . . . . . . 1 6 Ulota crispa E0 . . . . . . . . . + . . . . . . . 1 6 Dicranodontium denudatum E0 . . . . . . . . . . + . . . . . . 1 6 Cladonia rangiferina E0 . . . . . . . . . . + . . . . . . 1 6 Porella platyphylla E0 . . . . . . . . . . . . + . . . . 1 6 Neckera complanata E0 . . . . . . . . . . . . + . . . . 1 6 Homalothecium lutescens E0 . . . . . . . . . . . . . + . . . 1 6 Conocephalum conicum E0 . . . . . . . . . . . . . . + . . 1 6 Fissidens taxifolius E0 . . . . . . . . . . . . . . . . + 1 6 Legend - Legenda ALR Limestone with chert and marlstone - apnenec z rožencem in laporovcem DR Dolomite with chert - dolomit z rožencem Re Rendzina - rendzina Number of relevé (Zaporedna številka popisa) 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 Pr. Fr. Number of relevé (Zaporedna številka popisa) 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 Pr. Fr. Number of relevé (Zaporedna številka popisa) 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 Pr. Fr. Number of relevé (Zaporedna številka popisa) 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 Pr. Fr. Table 7 (Preglednica 7): Saxifrago cuneifolii-Fagetum typicum Number of relevé (Zaporedna številka popisa) 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 Database number of relevé (Delovna številka popisa) 247261 247624 247264 255353 247628 255354 247629 247631 247632 255503 247622 247623 255323 247630 255355 247626 255356 247633 247636 247637 255516 211738 212216 255391 255392 255390 Elevation in m (Nadmorska višina v m) 1240 1170 1190 1220 1100 1220 1250 1310 1070 900 1270 1220 1150 1140 1072 1070 1150 1060 1340 1370 1230 1351 1260 1050 1150 1210 Aspect (Lega) S NE S NEE SE S SW SSW SW NW NE E NNE NW NW E SE SW E SE NE SE SSW SE E S Slope in degrees (Nagib v stopinjah) 40 35 35 40 35 35 40 35 40 35 40 35 40 35 35 35 35 35 30 40 35 50 45 35 30 35 Parent material (Matična podlaga) AR AR DRG DR AR DR DR AR AR ALR AR AR DR AR AR DR Gl AR DR R DR AL R R ARG AR Soil (Tla) Re Re Re Re Re Re Re Re Re Re Re Re Re Re Re Re Ra Re Re Dy Re Re Re Dy Dy Re Stoniness in % (Kamnitost v %) 20 30 60 10 30 30 10 20 20 20 10 10 5 10 10 40 70 10 . 20 10 20 . . . 1 Cover in % (Zastiranje v %): Upper tree layer (Zgornja drevesna plast) E3b 80 90 90 100 90 95 80 80 90 80 80 90 80 90 95 90 80 90 90 80 100 70 80 90 95 90 Lower tree layer (Spodnja drevesna plast) E3a 10 5 5 . 5 . 10 10 10 20 10 . 10 5 5 10 . 5 10 . 10 20 . . . Shrub layer (Grmovna plast) E2 5 10 5 10 10 5 5 5 10 10 15 10 20 5 70 2 50 5 10 10 5 5 5 15 5 5 Herb layer (Zeliščna plast) E1 70 70 60 70 70 50 80 70 70 70 70 80 80 80 20 50 40 30 90 70 80 70 50 70 70 100 Moss layer (Mahovna plast) E0 10 10 5 10 5 5 10 10 10 10 10 5 5 10 . 10 . 5 5 10 5 5 10 10 10 . Maximum diameter of trees (Največji prsni premer dreves) cm 35 40 40 30 45 40 40 40 35 35 40 20 30 30 30 35 40 30 40 45 35 60 50 30 30 35 Maximum height of trees (Največja drevesna višina) m 10 10 14 12 14 20 16 14 10 15 10 7 10 15 18 14 16 16 15 12 18 14 18 16 18 16 Number of species (Število vrst) 50 65 68 67 69 62 63 71 62 60 59 48 64 74 69 69 71 53 46 55 66 53 53 36 45 48 Relevé area (Velikost popisne ploskve) m2 200 200 200 400 200 400 200 200 200 400 200 200 400 200 400 200 400 200 200 200 400 400 200 400 400 400 Date of taking relevé (Datum popisa) 7/11/1991 7/11/1991 8/17/1992 9/14/1987 8/4/1991 9/14/1987 7/2/1991 7/2/1991 7/2/1991 7/3/1991 7/11/1991 7/17/1991 8/17/1989 7/2/1991 9/15/1987 7/11/1991 9/16/1987 7/2/1991 6/29/1991 8/29/1991 9/5/1986 6/21/2006 6/21/2006 6/26/1988 6/26/1988 6/26/1988 Locality (Nahajališče) Blesar-Žbentre Kacenpoh - Šoštar Šoštar-Blesar Kacenpoh Kacenpoh-Šoštar Kacenpoh - Žbentre Kacenpoh-Kobla Kacenpoh-Kobla-Krevle Kacenpoh-Kobla Jehle - Gladka skala Kacenpoh - Šoštar Kacenpoh-Šoštar Kacenpoh-Fergajnar Kacenpoh-Kobla Kacenpoh-Kobla Kacenpoh-Šoštar Bača - Prbola Kacenpoh-Kobla Kacenpoh - Babji zob Črna prst-Kor Pl. Kuk - Lasene Bizle Bizle Pl. Kuk - Rovje Pl. Kuk - Rovje Pl. Kuk - Lasene Quadrant (Kvadrant) 9749/4 9749/4 9749/4 9749/4 9749/4 9749/4 9749/4 9749/4 9749/4 9749/4 9749/4 9749/4 9749/4 9749/4 9749/4 9749/4 9749/4 9749/4 9749/4 9749/4 9748/4 9749/3 9749/3 9748/4 9748/4 9748/4 Coordinate GK Y (D-48) m 419383 419143 419309 419086 419341 419381 420166 420221 419980 417014 419044 419111 419642 420067 419966 419279 420638 420077 418710 418930 407821 415472 415324 408010 407905 407828 Coordinate GK X (D-48) m 5120991 5120822 5120955 5120818 5120884 5121006 5121604 5121656 5121494 5118128 5120788 5120805 5121245 5121522 5121521 5120734 5121096 5121370 5120708 5120950 5120073 5120103 5120014 5119780 5119893 5119946 Diagnostic species of the association (Diagnostične vrste asociacije) Pr. Fr. VP Calamagrostis arundinacea E1 3 2 1 3 1 1 + 1 1 2 2 4 4 2 3 + 2 + 4 3 3 2 2 4 4 4 26 100 VP Veronica urticifolia E1 1 1 1 1 1 1 1 1 1 1 . 1 + 1 1 1 1 + + 1 1 + 1 1 1 + 25 96 QP Tanacetum corymbosum s. lat. (inc. T. clusii) E1 1 1 + + + + + . . . + + + . + + + + + + 1 + + 1 1 1 22 85 VP Luzula luzuloides E1 1 + + . + + + . . + + 1 . . + + 1 + 1 1 + + + 1 1 + 21 81 ES Campanula witasekiana E1 + + 1 + + + + + . + . . + . . + 1 . . + . . . + + . 15 58 QF Festuca heterophylla E1 1 . + + 1 + . . + + . . . . . + . + . + . + + + + 1 15 58 FB Carex humilis E1 + . 1 + + + 2 1 + . + . . + . . . . . . . 1 + . . . 12 46 VP Saxifraga cuneifolia E1 . + . + . . . . . . . + . + + + 1 . + . + . + . + . 11 42 VP Huperzia selago E1 . + + + . . . . . . . + 1 + 1 . + . . . + . . . . . 9 35 TG Clinopodium vulgare E1 . . + + + + . + . . . . . . . . . . . . + + . . . . 7 27 Geographical differential species (Geografski razlikovalni vrsti) MuA Aconitum angustifolium E1 . + 1 + + + + 1 . . + . . . . . . . . . . . . . . + 9 35 VP Larix decidua E3b . . . . . . . . . + . . . . . . . . . . . . r . . . 2 8 Differential species of variants (Razlikovalnice variant) QF Carex digitata E1 . + + . + + + + + . + + . + 1 . . + . . . . . . . . 12 46 EP Calamagrostis varia E1 + . + + 2 + 1 + . + + . . . . . . . . . . . . . . . 9 35 ES Carduus crassifolius E1 . . + + . + + + + . + . . . . . . . . . . . . . . . 7 27 ES Laserpitium peucedanoides E1 + + + . + . + . . . . + . . . + . . . . . . . . . . 7 27 QP Primula veris subsp. columnae E1 . . . . . . . . . . . . . . . . . . . . . + + . + 1 4 15 QP Sesleria autumnalis E1 . . . . . . . . . . . . . . . . . . . . . . . 1 + + 3 12 FS Lathyrus vernus subsp. flaccidus E1 . . . . . . . . . . . . . . . . . . . . . . r . . + 2 8 AF Anemone trifolia E1 . . . . . . . . . . . . . . . . . . . . . . . 1 + . 2 8 AF Aremonio-Fagion AF Cyclamen purpurascens E1 1 1 1 1 1 1 1 1 + 1 1 1 1 + 1 + + + + + 1 . + . . 1 23 88 AF Dentaria enneaphyllos E1 + 1 1 1 1 + 1 1 + + 1 . . 1 1 1 . + + . . . . . . . 16 62 AF Cardamine trifolia E1 . . . . . . . . . . + . . + + . . . 1 + 1 . . . + . 7 27 AF Knautia drymeia E1 . . + . . . + . + . . . . . . . + . . + . + . . . . 6 23 AF Lamium orvala E1 . . . . . 1 . . . . . . . . + . . + . . . . . . . . 3 12 AF Hacquetia epipactis E1 . . . 1 . . . . . . . . . . . . . . . . . . . . . . 1 4 EC Erythronio-Carpinion Primula vulgaris E1 1 1 1 + 1 + + + + + + + . . . 1 + + + . . + + . + . 19 73 Ornithogalum pyrenaicum E1 . . + . . . . . . . . . . . . . . + . . . . . . . . 2 8 Helleborus odorus E1 . . . . . . . . . . . . . . . . . . . . + . . . . . 1 4 TA Tilio-Acerion Acer pseudoplatanus E3b . . . . . . . . . + . . . + . . . + . . . . . . . . 3 12 Acer pseudoplatanus E3a . . . . + . . . . . . . . . . . . . . . . . . . . . 1 4 Acer pseudoplatanus E2b . . . . . . . . + . + . . + . . . + . . . . . . . . 4 15 Acer pseudoplatanus E2a . . . . . . . . + . + . + . + . . + . . . . . . . . 5 19 Acer pseudoplatanus E1 . . . . + . . . . + + . . . . . . + . . . . + . . . 5 19 Polystichum aculeatum E1 . . . . . . . . . . . . . + + . . . . . + . . . . . 3 12 Aruncus dioicus E1 . . . . . . . . . . + . . . . . . . + . . . . . . . 2 8 Thalictrum aquilegiifolium E1 . . . + . . . . . . . . . . . . . . . . . + . . . . 2 8 FS Fagetalia sylvaticae Fagus sylvatica E3b 5 5 5 5 5 5 5 5 5 5 5 5 5 5 5 5 4 5 5 5 5 4 5 5 5 5 26 100 Fagus sylvatica E3a + 1 1 . 1 . + 1 . . 1 . . + . + . . 1 + . . + . . . 12 46 Fagus sylvatica E2b + 1 + 1 + + . + + + + 1 . . . . . + + + + . + 1 1 + 19 73 Fagus sylvatica E2a . + + . . . . . . . . . . . . + . + + . . + . . . . 6 23 Fagus sylvatica E1 . . . + + . . . . . . . . . . . . . . . . . . + + . 4 15 Galium laevigatum E1 1 1 1 1 1 + 1 1 + 1 1 1 1 1 1 + 1 + 1 1 1 1 1 + . + 25 96 Mercurialis perennis E1 1 1 2 1 1 1 2 2 2 1 1 + 1 1 1 1 + 1 . + + 1 + . . . 22 85 Prenanthes purpurea E1 1 1 + 1 1 . + . 1 1 1 . 1 1 1 1 1 + + 1 1 + 2 1 1 . 22 85 Lonicera alpigena E2a + 1 + 1 . + + + + 1 + . 1 1 1 . + . + + . + . . + . 18 69 Symphytum tuberosum E1 + + + . + . . + . + . + + + . + . + + + + 1 1 . + 1 18 69 Daphne mezereum E2a . + . + + . + + . 1 . . + + 1 + + . . . . + . + . . 13 50 Melica nutans E1 + . . + + + . + 1 1 + + . + + + + . . . . . . . . . 13 50 Euphorbia amygdaloides E1 + + . . + . + . . + + + . + + . . + + . 1 . . . . . 12 46 Salvia glutinosa E1 + + + . 1 1 . + + . . . + + 1 . . + . . + . . . . . 12 46 Dryopteris filix-mas E1 . . . . . . . . + + + . . + + + + + + + 1 . + . . . 12 46 Campanula trachelium E1 . + + 1 1 1 . + . . . . . . . + . + . . 1 . . + . 1 11 42 Laburnum alpinum E3b . + . + . . . . . . . + . . . . . . . . . . . . . . 3 12 Laburnum alpinum E3a . 1 . . + . . . + 1 . . + . . + 1 . . + . . . . . . 8 31 Laburnum alpinum E2b . + + + . . . . . . . + + + + . . . . + . . . . + . 9 35 Laburnum alpinum E2a . . + . . . . + + . + . + + 1 + + + . . . . . . + . 11 42 Laburnum alpinum E1 . + . + + . . . . + + . . . . + + + . . . . . + . . 9 35 Epipactis helleborine E1 . . + + + + 1 + + . . . . . . . . + . . + . . . . . 9 35 Lilium martagon E1 + . + . . . + . . + + . . . . . . . . . + + + . . + 9 35 Festuca altissima E1 . + . . . . . . + . 1 + . + . + . + . . . . . . . . 7 27 Heracleum sphondylium E1 . + . . . . + + . . + . . + . . . + . . + . . . . . 7 27 Lathyrus vernus subsp. vernus E1 . . . . 1 . . . . . . . . . . . . . + . 1 + . . . + 5 19 Galeobdolon flavidum E1 . . . . . 1 + + . . . . . . . . . + . . . . . . . . 4 15 Mycelis muralis E1 . . . . + 1 . . . + . . . . . . . + . . . . . . . . 4 15 Pulmonaria officinalis E1 . . . . + . . . . . . . + . + . . . . . + . . . . . 4 15 Viola reichenbachiana E1 . . . . + . . . . + . . . . . . . . . . . . . + . . 3 12 Neottia nidus-avis E1 . . . . . . + . . . . . . . . . . . . . . . . + . + 3 12 Actaea spicata E1 . . . . . . . . . . . . . + + . . . . . + . . . . . 3 12 Epilobium montanum E1 . . . . . . . . . . . . . . . + . . . + + . . . . . 3 12 Poa nemoralis E1 . . . . . . . . . . . . . . . + . . . + . . . . . + 3 12 Petasites albus E1 . . . + . . . . . . . . . . . . . . . . + . . . . . 2 8 Scrophularia nodosa E1 . . . . . . . . . . . . . . . + . . . . + . . . . . 2 8 Polygonatum multiflorum E1 . . . . . . . . + . . . . . . . . . . . . . . . . . 1 4 Euphorbia dulcis E1 . . . . . . . . . . . . . + . . . . . . . . . . . . 1 4 Cardamine bulbifera E1 . . . . . . . . . . . . . . . . . + . . . . . . . . 1 4 Ranunculus lanuginosus E1 . . . . . . . . . . . . . . . . . . . . + . . . . . 1 4 QP Quercetalia pubescenti-petraeae Melittis melissophyllum E1 + + + + + + 1 . + 1 . + . + + . + + . + . + + . + . 18 69 Sorbus aria E3b . . . + . . . . . . . . . . + . . + . . . + + + + . 7 27 Sorbus aria E3a + + + . + . + + + + . + 1 . . + 1 . . . . . 1 . . . 13 50 Sorbus aria E2b + 1 + 1 . + . + + . + . . + . . 1 . + + + + + 1 . + 17 65 Sorbus aria E2a . + + . . . . . . . + + 1 . . . . . + . . . . . . . 6 23 Sorbus aria E1 . . . + . . . . . . . . . . . . . . . . + . . . . + 3 12 Hypericum montanum E1 + . . . + . . . . + . . . + . . . . . . . . . . . . 4 15 Arabis turrita E1 . . . . + + . + . . . . . . . + . . . . . . . . . . 4 15 Convallaria majalis E1 . . . . . . . . . . + . . + . . + . . . . + . . . . 4 15 Peucedanum schottii E1 + . . . . . . . . . . . . . . . . . . . . . . . . + 2 8 Ostrya carpinifolia E3b . . . . . . . . . . . . . . . . . r . . . . . . . . 1 4 Ostrya carpinifolia E3a . . . . . . . . r + . . . . . . . . . . . . . . . . 2 8 Fraxinus ornus E3a . . . . . . . . r . . . . . . . . . . . . . . . . . 1 4 Fraxinus ornus E1 . . . . . . . . . + . . . . . . . . . . . . . . . . 1 4 QR Quercetalia roboris Hieracium lachenalii E1 . . . . . . . . . . . . + . . . . . . . . + . . . . 2 8 Rubus hirtus E2a . . . . . . . . . . . . . . . . + . . . + . . . . . 2 8 Hieracium racemosum E1 . . . . . . . . . . . . . . . . . . . . . . + . . . 1 4 QF Querco-Fagetea Anemone nemorosa E1 1 1 . 1 1 + 1 1 + 1 1 1 1 1 1 + . 1 + 1 + . + . . + 21 81 Hepatica nobilis E1 . 1 + . + . 1 1 1 . + + + 1 1 1 + . + . 1 . . . . . 15 58 Dactylorhiza fuchsii E1 . + + . . . . . . . . . . + . . . . + . . . . + . . 5 19 Corylus avellana E2 . . + . . . . . . . . + + . . . + + . . . . . . . . 5 19 Platanthera bifolia E1 . . . . . . . . . . . . + . . . . . . . . . . . . . 1 4 Viola riviniana E1 . . . . . . . . . . . . + . . . . . . . . . . . . . 1 4 Clematis vitalba E2a . . . . . . . . . . . . . . . + . . . . . . . . . . 1 4 Moehringia trinervia E1 . . . . . . . . . . . . . . . . . + . . . . . . . . 1 4 Listera ovata E1 . . . . . . . . . . . . . . . . . . . . . . . . + . 1 4 Cephalanthera longifolia E1 . . . . . . . . . . . . . . . . . . . . . . . . . + 1 4 EP Erico-Pinetea Cirsium erisithales E1 + + 1 1 1 + 1 1 1 + . . + + 1 + + + . + 1 1 + . + 1 22 85 Rubus saxatilis E1 + + + 2 . + + + + 1 + 1 + + 1 . + . . . . . + . . . 16 62 Aquilegia nigricans E1 . + + + . . + + + + + . . + + + . . . . + . . . . . 12 46 Buphthalmum salicifolium E1 + + 1 . 1 + + 1 + . + . . . . + . . . . . + . . . + 12 46 Peucedanum austriacum E1 . . . . + + + + + . . . . . . . . . . . . . . . . . 5 19 Erica carnea E1 . . . . . . . . . + + + + . . . . . . . . . . . . . 4 15 Rhododendron hirsutum E2a . + . . . . . . . . . + + . . . . . . . . . . . . . 3 12 Carex alba E1 . . . + . + . . . . . . . . + . . . . . . . . . . . 3 12 Carex ornithopoda E1 . . . . . . . . . . . . . . . + . . . . . . . . . . 1 4 Allium ericetorum E1 . . . . . . . . . . . . . . . . + . . . . . . . . . 1 4 Epipactis atrorubens E1 . . . . . . . . . . . . . . . . . . . . . . . . . + 1 4 VP Vaccinio-Piceetea Solidago virgaurea E1 + 1 1 1 1 1 1 + 1 + + + + 1 1 + 1 + + 1 . + + + + 1 25 96 Hieracium murorum E1 + 1 1 1 + + 1 + 1 + . 1 + + 1 1 1 . 1 + 1 + + 1 1 1 24 92 Valeriana tripteris E1 + 1 1 + 1 + 1 1 1 1 1 1 1 + 1 1 1 . 1 1 + 1 + . + . 23 88 Rosa pendulina E2a + + r 1 + . + + 1 1 1 1 + 1 1 . 1 + + + + . . + . + 21 81 Aposeris foetida E1 + 1 1 1 + 1 . . + . . + 1 1 1 + . + 1 1 1 1 1 . + 2 20 77 Vaccinium myrtillus E1 + + . + . . . . + . + + 1 + + . 1 . + . + . . 1 . . 13 50 Gentiana asclepiadea E1 . . . 1 . . + . + + . . + + 1 . + . + . + . + . . . 11 42 Homogyne sylvestris E1 . 1 . 1 . . . . + . + 1 1 1 1 1 + . . . + . . . . . 11 42 Maianthemum bifolium E1 . + . + . . . . . . + 1 + + + + . . + + + . . . . . 11 42 Clematis alpina E2a . . . . . . + + + . + . + + + + + . . . . . + . . . 10 38 Oxalis acetosella E1 . . . 1 . . . . . + . . + + 1 . . . 1 1 1 . . . 1 . 9 35 Picea abies E3b . . . . . . + + . . . . . + 1 . 1 r . . . + + . . . 8 31 Picea abies E3a . . . . . . . . . + . . . . . . . . . r . . . . . . 2 8 Picea abies E2b . . . . + . . . . . . . . . . . + . . . . . . . . . 2 8 Picea abies E2a . . . . . . . . . . . . + . + . . . . + . . . . . . 3 12 Picea abies E1 . . . + . . . . . . + . . . . . . . . . . . . . . . 2 8 Polystichum lonchitis E1 . . . . . . . . . . + . . . . r . . . . + . . . . . 3 12 Phegopteris connectilis E1 . . . . . . . . . . . . 1 + + . . . . . . . . . . . 3 12 Abies alba E3b . . . . . . . . . . . . r . . . . . . . . . . . . . 1 4 Abies alba E3a . . . . . . . . + . . . . + . . . . . . . . . . . . 2 8 Abies alba E2b . . . . . . . . . . . . . . + . . . . . . . . . . . 1 4 Dryopteris dilatata E1 . . . . . . . . . . . . . + . . . . . + . . . . . . 2 8 Vaccinium vitis-idaea E1 . . . . . . . . . . . . + . . . . . . . . . . . . . 1 4 Luzula sylvatica E1 . . . . . . . . . . . . . . . . . . + . . . . . . . 1 4 Avenella flexuosa (Deschampsia flexuosa) E1 . . . . . . . . . . . . . . . . . . . . + . . . . . 1 4 Gymnocarpium dryopteris E1 . . . . . . . . . . . . . . . . . . . . + . . . . . 1 4 Orthilia secunda E1 . . . . . . . . . . . . . . . . . . . . . . . . + . 1 4 SSc Sambuco-Salicion capreae Sorbus aucuparia E3b . . . . . . . . . . . . . . . . . . . . . . r . . . 1 4 Sorbus aucuparia E3a . . . . . . . . . . . . + . . . . . . . . . . . . . 1 4 Sorbus aucuparia E2b . . . . . . . . . . . + . . . . + . . + . . . . . . 3 12 Sorbus aucuparia E2a . . . . . . . . . + . + + . . . . . . + . . . . . . 4 15 Sorbus aucuparia E1 . . . . . . . . . . . . . + . . + . . . . . + . . . 3 12 TG Trifolio-Geranietea Verbascum lanatum E1 . . . . . + . . . . . . . . . + . + . + + . + + . + 8 31 Iris graminea E1 + . . . . . + + . . . . + + . . . . . . . + + . . . 7 27 Digitalis grandiflora E1 . + . + + + . . . . . . . . . . . . . + + . . . . . 6 23 Laserpitium latifolium E1 . . . . . + + + + . . . . . . . . . . . . + r . . . 6 23 Thalictrum minus E1 . . . . . + . + . . . . . . . . . . . . . + r . . . 4 15 Vicia sylvatica E1 . . . . . . . . . . . . . . . . . . . . . + + . + + 4 15 Lilium carniolicum E1 . + + . . + . . . . . . . . . . . . . . . . . . . . 3 12 Libanotis sibirica s. lat. E1 . . . . . . + + . . . . . . . . . . . . . . . . . + 3 12 Anthericum ramosum E1 . . + . . . . . . . . . . . . + . . . . . . . . . . 2 8 Arabis pauciflora E1 . . + . . . . + . . . . . . . . . . . . . . . . . . 2 8 Vincetoxicum hirundinaria E1 . . . . . + . + . . . . . . . . . . . . . . . . . . 2 8 Verbascum lychnitis E1 . . + . . . . . . . . . . . . . . . . . . . . . . . 1 4 Calamintha einseleana E1 . . . . . + . . . . . . . . . . . . . . . . . . . . 1 4 Origanum vulgare E1 . . . . . . . + . . . . . . . . . . . . . . . . . . 1 4 Trifolium rubens E1 . . . . . . . . . . . . . . . . + . . . . . . . . . 1 4 Campanula rapunculoides E1 . . . . . . . . . . . . . . . . . . . . . . + . . . 1 4 EA Epilobietea angustifolii Fragaria vesca E1 . + + + + + + + + + . . . . . . + . . . + . . . + + 13 50 Hypericum hirsutum E1 . . . . + + . + . . + . . . . + . . . . + . . . . . 6 23 Rubus idaeus E2a . + . . + . . . . . . . + + . . . . . + . . . . + . 6 23 Galeopsis speciosa E1 . . . . . . . . . . . . . . . . . . . + + . . . . . 2 8 BA Betulo-Alnetea viridis Alnus viridis E2b . . . . . . . . . . . . . . . . . . . . . . + . . . 1 4 MuA Mulgedio-Aconitetea Phyteuma ovatum E1 + + + . 1 . + + 1 + . + + 1 + 1 + + 1 1 + 1 1 . 1 . 21 81 Senecio ovatus E1 . . + . + 1 + + + + . + + + . + + + + 1 1 1 + + 1 + 21 81 Aconitum lycoctonum s. lat. E1 . + + + . + + + . . + . . 1 1 . . 1 . . + . . . . + 12 46 Athyrium filix-femina E1 . + . + . . . . . . + . + + + . . . 1 + 1 . . + + . 11 42 Polygonatum verticillatum E1 . . . + . . 1 + . . . . . + + . + + . + + + . . . . 10 38 Veratrum album E1 . . . . . . . . . . + . . + . . . + + + + . . . . 1 7 27 Pleurospermum austriacum E1 . . . + . . . . . . . . . . . . + . . + . + . . . . 4 15 Ranunculus platanifolius E1 . . . . . . . . . . . . . . . . . . . + + . . . . . 2 8 Silene vulgaris subsp. antelopum E1 . . . . . . + . . . . . . . . . . . . . . . . . . . 1 4 Silene dioica E1 . . . . . . . . . . . . . . . . . + . . . . . . . . 1 4 Saxifraga rotundifolia E1 . . . . . . . . . . . . . . . . . . + . . . . . . . 1 4 Chaerophyllum villarsii E1 . . . . . . . . . . . . . . . . . . . + . . . . . . 1 4 Centaurea montana E1 . . . . . . . . . . . . . . . . . . . . . . . . . + 1 4 ES Elyno-Seslerietea Sesleria caerulea subsp. calcaria E1 + + 3 + + + 1 1 2 1 + + . . + + + . . + . 1 + . . . 18 69 Betonica alopecuros E1 + + + 1 + + 1 1 . 1 + . . . 1 + + . . . . + . . . 1 15 58 Phyteuma orbiculare E1 + 1 + . + . + + . . . . . + . . . . . . . + + . . . 9 35 Centaurea haynaldii subsp. julica E1 + . + + + + . . . . . . . . . . . . . . . + . . . . 6 23 Serratula macrocephala E1 . + . . . . . . . . + + . . . . . . . . . . . . . . 3 12 Carex mucronata E1 . . . . . . + . . . . . . . . . . . . . . . . . . . 1 4 Festuca calva E1 . . . . . . . + . . . . . . . . . . . . . . . . . . 1 4 Astrantia bavarica E1 . . . . . . . . . . . . . + . . . . . . . . . . . . 1 4 FB Festuco-Brometea Carlina acaulis E1 . . + + . . . . . . . . . . . . . . . . . + . . . + 4 15 Bromopsis transsilvanica E1 . . . . . . . + . . . . . . . . . . . . . + . . . . 2 8 Brachypodium rupestre E1 . . . . . . . . . . . . . . . . . . . . . 1 . . . + 2 8 Linum viscosum E1 . . + . . . . . . . . . . . . . . . . . . . . . . . 1 4 Allium carinatum subsp. carinatum E1 . . . . . + . . . . . . . . . . . . . . . . . . . . 1 4 Euphorbia cyparissias E1 . . . . . + . . . . . . . . . . . . . . . . . . . . 1 4 Dianthus monspessulanus E1 . . . . . . . . . . . . + . . . . . . . . . . . . . 1 4 Koeleria pyramidata E1 . . . . . . . . . . . . . . . . . . . . . + . . . . 1 4 Festuca valesiaca agg. E1 . . . . . . . . . . . . . . . . . . . . . . . . . + 1 4 MA Molinio-Arrhenatheretea Galium album E1 . . . . + + + + . . . . . . . . . . . . . . . . . . 4 15 Angelica sylvestris E1 . . + . . . . . . . . . . . . + . . . . . . . . . . 2 8 Dactylis glomerata E1 . . . . . . . . . . . . . . . . . . . . . . . . + + 2 8 Taraxacum officinale E1 . . . . . . . + . . . . . . . . . . . . . . . . . . 1 4 CU Calluno-Ulicetea Phyteuma zahlbruckneri E1 . . . . . . . . . . . . . . . . + . . . . . . + . . 2 8 Festuca ovina agg. E1 . . . . . . . . . . . . . . . . + . . . . . . . . . 1 4 Festuca nigrescens E1 . . . . . . . . . . . . . . . . . . . . . + . . . . 1 4 Festuca sp. E1 . . . . . . . . . . . . . . . . . . . . . . . . . + 1 4 Potentilla erecta E1 . . . . . . . . . . . . . . . . . . . . . . . . . + 1 4 TR Thlaspietea rotundifolii Adenostyles glabra E1 + + 1 + + 1 + . 1 . + 1 + 1 2 1 . . + . + + + . + . 19 73 Ligusticum seguieri E1 + . . . . . . . . . . . . . . . . . . . . + . . . . 2 8 Campanula cochleariifolia E1 . . . . . . . . + . . . . . . + . . . . . . . . . . 2 8 Heracleum sphondylium subsp. pollinianum E1 . . + . . . . . . . . . . . . . . . . . . . . . . . 1 4 Arabis alpina E1 . . . . . . . . . . . . . . . r . . . . . . . . . . 1 4 Gymnocarpium robertianum E1 . . . . . . . . . . . . . . . + . . . . . . . . . . 1 4 AT Asplenietea trichomanis Asplenium viride E1 . + + + + + . + + + + . + + 1 1 . . . . . . . . . . 13 50 Asplenium ruta-muraria E1 . . . . + + . + + . . . . . + + + . . . . . + . . + 9 35 Asplenium trichomanes E1 . . . . + . . . . . . . . . . + 1 + . + . . + . + . 7 27 Cystopteris fragilis E1 . . . . . . . + . . + . . . . + . + . + . . . . . . 5 19 Campanula carnica E1 . . . . . . . . + + . . . . . . . . . + . + + . . . 5 19 Polypodium vulgare E1 . . . . . . . . . . . . . . . + + . . + . . . + . . 4 15 Festuca stenantha E1 . . . . . . . + . . . . . . . . + . . . . . . . . . 2 8 Moehringia muscosa E1 . . . . . . . . . + . . . . . . 1 . . . . . . . . . 2 8 Carex brachystachys E1 . . . . . . . . . . + . . . . 1 . . . . . . . . . . 2 8 Saxifraga crustata E1 . . . . . . . . . . . . r . . . + . . . . . . . . . 2 8 Sedum album E1 . . . . . . . . . . . . . . . . + . . . . . . . . . 1 4 Paederota lutea E1 . . . . . . . . . . . . . . . . + . . . . . . . . . 1 4 Primula auricula E1 . . . . . . . . . . . . . . . . . . . . . + . . . . 1 4 Campanula spicata E1 . . . . . . . . . . . . . . . . . . . . . . r . . . 1 4 ML Mosses and lichens (Mahovi in lišaji) Tortella tortuosa E0 + + + + + + + + + 1 + 1 1 + 2 . + . + . . . . + + + 20 77 Polytrichum formosum E0 . . + + + . + . + + 1 1 1 1 1 + + + + + + . + 1 2 . 20 77 Ctenidium molluscum E0 + 1 + + + + + + + 1 1 + + 1 + + . . + . + . . . . + 19 73 Schistidium apocarpum E0 + + + + + . . + + + . . . . . . + . . + . . . + + + 13 50 Fissidens dubius E0 . . . + + + . . + . + + 1 + 1 + . . . . . . . . . . 10 38 Atrichum undulatum E0 . . . . . . . . . + . 1 1 + . . . . + + . . + 1 1 . 9 35 Plagiothecium denticulatum E0 . + + + . . . . . . . . . . 1 . . + + . . . . + . . 7 27 Dicranella heteromalla E0 . . . . . . . . . . . . 1 + 1 . 1 . . . . . . 1 + . 6 23 Cladonia fimbriata E0 + + . + . . . . . . . + . + . . . . . . . . . . . . 5 19 Dicranum scoparium E0 . + . . + . . . . . . . . . + . 1 . . . . . . + . . 5 19 Mnium thomsonii E0 . . . + . . . . . + . . . . 1 . + . . . . . . + . . 5 19 Mnium marginatum E0 . . . . . . . . + . . . . + . + . + + . . . . . . . 5 19 Neckera crispa E0 . . . . . . . . + 1 . . . . . + + . . . . . + . . . 5 19 Brachythecium velutinum E0 + . + . . . . . . . . . . . . . . + . . . . . . + . 4 15 Bryum capillare E0 + + . . . . . . . . . . + . . + . . . . . . . . . . 4 15 Metzgeria furcata E0 . . . . . . . . + . . + . . + . . . . . . . . . + . 4 15 Peltigera canina E0 . . . + + . . . . . . . . . + . . . . . . . . . . . 3 12 Cladonia furcata E0 . . . . . . . . . + . . . . . . . . . . . . . + + . 3 12 Leucobryum glaucum E0 . . . . . . . . . . . . + + + . . . . . . . . . . . 3 12 Isothecium alopecuroides E0 . . . . . . . . . . . . + . . . + . . . . + . . . . 3 12 Hypnum cupressiforme var. filiforme E0 . . . . . . + + . . . . . . . . . . . . . . . . . . 2 8 Encalypta streptocarpa E0 . . . . . . . + . . . . . . . + . . . . . . . . . . 2 8 Plagiochila porelloides E0 . . . . . . . . . + . . . . . . + . . . . . . . . . 2 8 Isothecium myosuroides E0 . . . . . . . . . . + + . . . . . . . . . . . . . . 2 8 Cladonia rangiferina E0 . . . . . . . . . . . . + . . . + . . . . . . . . . 2 8 Bryum argenteum E0 . . . . . . . . . . . . . . + 1 . . . . . . . . . . 2 8 Hypnum cupressiforme E0 . . . . . . . . . . . . . . . . 2 . . . 1 . . . . . 2 8 Polytrichum juniperinum E0 . . . . . . . . . . . . . . . . + . . . 1 . . . . . 2 8 Homalothecium philippeanum E0 . + . . . . . . . . . . . . . . . . . . . . . . . . 1 4 Ulota crispa E0 . . . . . . + . . . . . . . . . . . . . . . . . . . 1 4 Homalothecium sericeum E0 . . . . . . . + . . . . . . . . . . . . . . . . . . 1 4 Dermatocarpon miniatum E0 . . . . . . . + . . . . . . . . . . . . . . . . . . 1 4 Marchantia polymorpha E0 . . . . . . . . . . . . + . . . . . . . . . . . . . 1 4 Conocephalum conicum E0 . . . . . . . . . . . . . . . + . . . . . . . . . . 1 4 Dicranodontium denudatum E0 . . . . . . . . . . . . . . . . 2 . . . . . . . . . 1 4 Thuidium tamariscinum E0 . . . . . . . . . . . . . . . . + . . . . . . . . . 1 4 Rhizocarpon geographicum E0 . . . . . . . . . . . . . . . . + . . . . . . . . . 1 4 Fissidens taxifolius E0 . . . . . . . . . . . . . . . . . . + . . . . . . . 1 4 Paraleucobryum sauteri E0 . . . . . . . . . . . . . . . . . . + . . . . . . . 1 4 Homalothecium lutescens E0 . . . . . . . . . . . . . . . . . . . + . . . . . . 1 4 Pseudoleskeella catenulata E0 . . . . . . . . . . . . . . . . . . . + . . . . . . 1 4 Bartramia halleriana E0 . . . . . . . . . . . . . . . . . . . . + . . . . . 1 4 Plagiothecium nemorale E0 . . . . . . . . . . . . . . . . . . . . . . . . + . 1 4 Legend - Legenda AR Limestone with chert - apnenec z rožencem ARG Limestone with chert and claystone - apnenec z rožencem in glinavcem DR Dolomite with chert - dolomit z rožencem DRG Dolomite with chert and claystone - dolomit z rožencem in glinavcem AL Limestone with marlstone - apnenec z laporovcem ALR Limestone with chert and marlstone R Chert - roženec Gl Claystone - glinavec Re Rendzina - rendzina Ra Ranker - ranker Dy Dystric brown soil - distrična rjava tla Number of relevé (Zaporedna številka popisa) 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 Pr. Fr. Number of relevé (Zaporedna številka popisa) 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 Pr. Fr. Number of relevé (Zaporedna številka popisa) 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 Pr. Fr. Number of relevé (Zaporedna številka popisa) 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 Pr. Fr. Number of relevé (Zaporedna številka popisa) 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 Pr. Fr. Number of relevé (Zaporedna številka popisa) 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 Pr. Fr. Table 8 (Preglednica 8): Saxifrago cuneifolii-Fagetum rhododendretosum hirsuti, typicum, caricetosum humilis Number of relevé (Zaporedna številka popisa) 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 Database number of relevé (Delovna številka popisa) 217672 217673 255517 247260 221359 255357 255358 255359 221130 221131 255362 255363 212221 255321 255479 247627 Elevation in m (Nadmorska višina v m) 1180 1190 1210 1100 1210 1080 1130 1200 1010 1100 1100 1000 1170 1050 1030 1100 Aspect (Lega) NE NE NE W NW SW N N N NNE N N SE S SE E Slope in degrees (Nagib v stopinjah) 45 35 40 35 45 35 45 40 40 35 25 35 40 45 35 30 Parent material (Matična podlaga) DRG DRG AR AR AL AR ALR ALR AR AR AR AR DR DR AL AR Soil (Tla) Rj Rj Re Re Rj Re Re Re Re Re Rj Re Re Re Re Re Stoniness in % (Kamnitost v %) 30 50 60 20 30 70 30 30 30 30 40 70 10 5 10 80 Cover in % (Zastiranje v %): Upper tree layer (Zgornja drevesna plast) E3b 80 80 90 80 80 70 80 80 90 90 90 90 80 90 70 70 Lower tree layer (Spodnja drevesna plast) E3a 20 10 . 20 20 30 20 10 . 10 20 20 . . 10 . Shrub layer (Grmovna plast) E2 10 5 5 5 10 10 20 10 5 5 5 5 5 10 5 20 Herb layer (Zeliščna plast) E1 70 60 80 80 60 60 80 80 60 50 40 20 80 60 80 50 Moss layer (Mahovna plast) E0 30 30 15 10 20 10 20 20 30 10 20 10 5 5 5 10 Maximum diameter of trees (Največji prsni premer dreves) cm 50 60 40 45 45 40 35 35 40 35 45 35 30 40 35 20 Maximum height of trees (Največja drevesna višina) m 17 17 10 14 18 12 12 10 16 16 18 17 10 18 14 5 Number of species (Število vrst) 74 50 57 64 47 75 68 65 62 55 48 51 49 63 67 61 Relevé area (Velikost popisne ploskve) m2 200 200 200 200 400 400 400 400 400 400 400 400 200 400 400 200 Date of taking relevé (Datum popisa) 6/19/2007 6/19/2007 9/2/1986 7/30/1992 6/5/2001 7/29/1992 7/29/1992 7/29/1992 6/30/2004 6/30/2004 7/30/1992 7/31/1992 6/5/2006 8/17/1989 7/1/1991 7/11/1991 Locality (Nahajališče) Tisovec Tisovec Tisovec Zadlaščica - Sopota Črnik - Kladje Vrh nad Sopotom Vrh nad Sopotom Vrh nad Sopotom Srpenica - Strmi breg Srpenica - Strmi breg Vrh nad Sopotom Vrh nad Sopotom Bizle Kacenpoh - Fergajnar Robarjev grič Kacenpoh-Šoštar Quadrant (Kvadrant) 9748/4 9748/4 9748/4 9748/4 9747/2 9748/4 9748/4 9748/4 9747/1 9747/1 9748/4 9748/4 9749/3 9749/4 9749/4 9749/4 Coordinate GK Y (D-48) m 407495 407474 407473 406648 395790 406792 406915 406924 384715 384526 406588 406729 415333 419813 421957 419207 Coordinate GK X (D-48) m 5119608 5119599 5119574 5119827 5125336 5120184 5120110 5120066 5126925 5126934 5119767 5120313 5119910 5121059 5118553 5120868 Diagnostic species of the association (Diagnostične vrste asociacije) Pr. Fr. VP Veronica urticifolia E1 + 1 . + + 1 + + 1 1 + + 1 1 + . 14 88 VP Calamagrostis arundinacea E1 3 2 3 3 2 1 3 3 3 1 1 + . + . . 13 81 QP Tanacetum corymbosum s. lat. (inc. T. clusii) E1 + + 1 + + . . . . . + . 1 + . . 8 50 VP Luzula luzuloides E1 + 1 . . + + + + . . 1 + . . . . 8 50 ES Campanula witasekiana E1 . . + + + + + + . + . . . + . . 8 50 VP Huperzia selago E1 + . + . . . + + 1 + . . . + . . 7 44 VP Saxifraga cuneifolia E1 + + . . + . + . 1 . . + . . . . 6 38 QF Festuca heterophylla E1 1 + . + . 1 . . . . . . + . . . 5 31 FB Carex humilis E1 + . . + . . . . . . . . 2 2 2 . 5 31 TG Clinopodium vulgare E1 . . . + . + . . . . . . + + 1 . 5 31 Geographical differential species (Geografske razlikovalne vrste) MuA Aconitum angustifolium E1 . . + + . + + 1 . . . . . . . + 6 38 FS Lathyrus vernus subsp. flaccidus E1 + . . + . + + + . . . . . . . . 5 31 AF Anemone trifolia E1 . . . . 1 . . . 1 1 . . . . . . 3 19 VP Larix decidua E3b . . . . . . r . . . . . . . + . 2 13 QP Sesleria autumnalis E1 . . . . . + . . . . . . . . . . 1 6 Differential species of subassociation (Razlikovalni vrsti subasociacije) VP Clematis alpina E2a . + + + r + 1 1 1 . . . . . . . 8 50 EP Rhododendron hirsutum E2a + . + + . r + + . . . . . . . . 6 38 AF Aremonio-Fagion Cyclamen purpurascens E1 + + + 1 1 1 1 1 . 1 + 1 + 1 + 1 15 94 Cardamine enneaphyllos E1 . . . . . + . + . 1 . 1 . . . 2 5 31 Cardamine trifolia E1 . . . . . . + + . . 1 + . . . . 4 25 Lamium orvala E1 . . . . . + . . . . . . . . . + 2 13 Knautia drymeia E1 . . . . . . . . . . . . . + . . 1 6 Aremonia agrimonoides E1 . . . . . . . . . . . . . . + . 1 6 EC Erythronio-Carpinion Primula vulgaris E1 . . . . . + . . . . . . + + + + 5 31 Ornithogalum pyrenaicum E1 . . . . . . . . . . . . + . . + 2 13 TA Tilio-Acrion Acer pseudoplatanus E3b . . . . . . . . . 1 + . . . . . 2 13 Acer pseudoplatanus E3a . . . + . . . . . . . . . . . . 1 6 Acer pseudoplatanus E2b . . . . r + . . . . . . . . . . 2 13 Acer pseudoplatanus E2a . . . + + . . . . . . . . . . . 2 13 Acer pseudoplatanus E1 . . . 1 . . . + 1 1 1 r . . + . 7 44 Polystichum aculeatum E1 . . . + . . . . + + . + . . . 1 5 31 Thalictrum aquilegiifolium E1 . . . r . . . + . . . . . . . . 2 13 Acer platanoides E1 . . . . . . . . + . . . . . + . 2 13 Polystichum braunii E1 . . . . . . r . . . . . . . . . 1 6 Adoxa moschatellina E1 . . . . . . . . . + . . . . . . 1 6 Polystichum x illyricum E1 . . . . . . . . . . . + . . . . 1 6 Geranium robertianum E1 . . . . . . . . . . . . . . . + 1 6 Phyllitis scolopendrium E1 . . . . . . . . . . . . . . . + 1 6 FS Fagetalia sylvaticae Fagus sylvatica E3b 5 5 5 5 5 5 5 5 5 5 5 5 4 5 4 5 16 100 Fagus sylvatica E3a + 1 . 1 + + 1 1 1 1 1 1 . . . . 11 69 Fagus sylvatica E2b + + + . + + + + + + + r . . . . 11 69 Fagus sylvatica E2a + . . + . . . . . . . r + . . + 5 31 Fagus sylvatica E1 + 1 . . . + . + 1 1 + 1 . . + . 9 56 Daphne mezereum E2a + + + + + 1 1 + 1 + 1 + + + + . 15 94 Galium laevigatum E1 + . . + 1 + + + 1 1 + + + 1 + 1 14 88 Mercurialis perennis E1 . . . + + 1 1 1 1 . 1 1 2 1 + 1 12 75 Lonicera alpigena E2a + . + + + + + + + . . . + + . 1 11 69 Prenanthes purpurea E1 . . . + + . . . 1 + + + . 1 + . 8 50 Salvia glutinosa E1 . . . + . + . + . + + + + . + . 8 50 Symphytum tuberosum E1 1 1 . + . + . + . . + . + . . . 7 44 Dryopteris filix-mas E1 + 1 . . . . + + 1 . . + . . . 1 7 44 Epilobium montanum E1 + 1 1 . . . + + + . . . . . . . 6 38 Viola reichenbachiana E1 + + + + . . . . . . + . . . + . 6 38 Laburnum alpinum E3b . . . . . . . . . . . . r . + + 3 19 Laburnum alpinum E3a . . . . + 1 + r . . . . . 1 + . 6 38 Laburnum alpinum E2b . . . + + . + + . . . . . . . . 4 25 Laburnum alpinum E2a . . . . . + + . + . . . . . . . 3 19 Laburnum alpinum E1 . . . . . + . . 1 + . . . . + . 4 25 Asarum europaeum subsp. caucasicum E1 . . . + . + . . . + + . . . . + 5 31 Lathyrus vernus subsp. vernus E1 . . . . . + + + . + . + . . . . 5 31 Actaea spicata E1 . . . . . . . + + + . + . . . 1 5 31 Campanula trachelium E1 + . . . . + . . . . . . . 1 1 . 4 25 Lilium martagon E1 . . . . + . . . . . + + . . . + 4 25 Galeobdolon flavidum E1 . . . . . . . . . 1 + + . . . + 4 25 Poa nemoralis E1 1 1 . . . . . . . . . . . . + . 3 19 Luzula nivea E1 . . . . + . . . 2 2 . . . . . . 3 19 Paris quadrifolia E1 . . . . . . . . . + . + . . . + 3 19 Festuca altissima E1 . . . . . . . . . . + + . . . + 3 19 Epipactis helleborine E1 . . . . . . . . . . . . . 1 1 + 3 19 Tilia cordata E3b r . . . . . . . . . . . . . . + 2 13 Mycelis muralis E1 . 1 . . . + . . . . . . . . . . 2 13 Melica nutans E1 . . . + . . . . . . . . . . . 1 2 13 Scrophularia nodosa E1 . . . . . . . . . + . . . . . + 2 13 Petasites albus E1 . . . . . . . . . . . . . + + . 2 13 Pulmonaria officinalis E1 + . . . . . . . . . . . . . . . 1 6 Epipactis leptochila E1 . . . + . . . . . . . . . . . . 1 6 Euphorbia amygdaloides E1 . . . . . . . . . . . . . + . . 1 6 Neottia nidus-avis E1 . . . . . . . . . . . . . + . . 1 6 Cardamine bulbifera E1 . . . . . . . . . . . . . . . + 1 6 QP Quercetalia pubescenti-petraeae Sorbus aria E3 + + + + + 2 + + + + . . 2 1 + + 14 87 Sorbus aria E2b + + + + + 1 1 + + . + . 1 . . . 11 69 Sorbus aria E2a . . . . . . . . . . . . . 1 + + 3 19 Sorbus aria E1 . + . . . . . . + . . . + . . . 3 19 Ostrya carpinifolia E3b . . . . . . . . . . . . . . . . 2 13 Ostrya carpinifolia E3a r . + + r r . . . r . + 2 . + . 9 56 Ostrya carpinifolia E2b . . . . . . . . . . . . + . . . 1 6 Ostrya carpinifolia E1 . . . . . . . . + . . . . . . . 1 6 Primula veris subsp. columnae E1 1 + 1 + + + . . . . . . 1 . . . 7 44 Hypericum montanum E1 + . . + . + . . . . . . . + + . 5 31 Arabis turrita E1 . . . . . + + + . . . . . . + + 5 31 Convallaria majalis E1 + . + . . . . . . . . . 1 . 1 . 4 25 Melittis melissophyllum E1 . . . . . . . . . . . . 1 + + . 3 19 Peucedanum schottii E1 . . . . . . . . . . . . + . . + 2 13 Calamintha sylvatica E1 . . + . . . . . . . . . . . . . 1 6 Potentilla micrantha E1 . . . + . . . . . . . . . . . . 1 6 Fraxinus ornus E3b . . . . . . . . . . . . . . . + 1 6 Fraxinus ornus E3a . . . . . + . . . . . . . . . . 1 6 Fraxinus ornus E2b . . . . . . . . . . . . . . + . 1 6 Fraxinus ornus E2a . . . . . . . . + . . . . . . . 1 6 Carex flacca E1 . . . . . . . . . . . . + . . . 1 6 Cephalanthera rubra E1 . . . . . . . . . . . . . + . . 1 6 Peucedanum schottii var. petraeum E1 . . . . . . . . . . . . . + . . 1 6 QR Quercetalia roboris Betula pendula E3a . . . . . . . . . . . r . . . . 1 6 Rubus hirtus E2a . . + . . . . . . . . . . . . . 1 6 Populus tremula E3a . . . . . . . . . . . r . . . . 1 6 Hieracium lachenalii E1 . . . . . . . . . . . . . . + . 1 6 Hieracium sabaudum E1 . . . . . . . . . . . . . . + . 1 6 Quercus petraea E3b . . . . . . . . . . . . . . + . 1 6 QF Querco-Fagetea Hepatica nobilis E1 + + + 1 . 1 + 1 . . . . . . . 1 8 50 Carex digitata E1 + . + . . . . . . . + . . + + . 5 31 Anemone nemorosa E1 . + . 1 . . + . . . 1 + . . . . 5 31 Veratrum nigrum E1 . . . . + r . . . . . . . . . . 2 13 Cephalanthera longifolia E1 . . . . . . . . . . . . + + . . 2 13 Cruciata glabra E1 . . . + . . . . . . . . . . . . 1 6 Platanthera bifolia E1 . . . . . . . . . r . . . . . . 1 6 Moehringia trinervia E1 . . . . . . . . . . . + . . . . 1 6 Viola riviniana E1 . . . . . . . . . . . . . + . . 1 6 Clematis vitalba E2a . . . . . . . . . . . . . . + . 1 6 Lonicera xylosteum E2a . . . . . . . . . . . . . . . + 1 6 Stellaria holostea E1 . . . . . . . . . . . . . . . + 1 6 EP Erico-Pinetea Cirsium erisithales E1 + + + + + 1 1 . . . . . + 1 + + 11 69 Rubus saxatilis E1 + . . . + . 1 + + . . . . + . . 6 38 Buphthalmum salicifolium E1 . . + . . + + . . . . . . 1 1 . 5 31 Calamagrostis varia E1 . . + . . . . . . . . . 2 2 2 + 5 31 Epipactis atrorubens E1 . . . . . + . . . . . . . + + . 3 19 Erica carnea E1 . . + . r . . . . . . . . . . . 2 13 Peucedanum austriacum E1 . . . . . . . . . . . . . + 1 . 2 13 Polygala chamaebuxus E1 . . . . . . . . . . . . . + 1 . 2 13 Allium ericetorum E1 . . + . . . . . . . . . . . . . 1 6 Aquilegia nigricans E1 . . . . . . . . . . . . . + . . 1 6 VP Vaccinio-Piceetea Rosa pendulina E2a 1 1 1 1 1 + + + + . + . + . + 1 13 81 Valeriana tripteris E1 + 1 1 + 1 + 1 1 + . . . . 1 . . 10 63 Solidago virgaurea E1 + + 1 + + . . . + + . . . 1 1 . 9 56 Gentiana asclepiadea E1 + . . + . . + + . + + . . + . + 8 50 Oxalis acetosella E1 . + . + . . + + 1 1 1 1 . . . . 8 50 Maianthemum bifolium E1 . . . . r . + + + . + + . . 1 . 7 44 Hieracium murorum E1 + . + . + + . . . . . . . 1 1 . 6 38 Homogyne sylvestris E1 . . . . + . 1 1 1 + . . . . . . 5 31 Vaccinium myrtillus E1 + . + + . . . . + . . . . . . . 4 25 Dryopteris dilatata E1 . . . . . . . . + + + . . . . . 3 19 Phegopteris connectilis E1 + . + . . . . . . . . . . . . . 2 13 Gymnocarpium dryopteris E1 + . . . . . . . . . . + . . . . 2 13 Luzula luzulina E1 . . + + . . . . . . . . . . . . 2 13 Picea abies E3b . . . . r . . . . . . . . + . . 2 13 Picea abies E3a . . . . . . . . . . . + . . . . 1 6 Picea abies E2b . . . . . . . + . r . . . . . . 2 13 Picea abies E1 . . . . . . . . r . . . . . . . 1 6 Abies alba E3b . . . . . r . . . . . . . . . . 1 6 Abies alba E3a . . . . . . r . . r . . . . . . 2 13 Abies alba E2b . . . . . . . + . . . . . . . . 1 6 Laserpitium krapfii E1 . . . . . . + + . . . . . . . . 2 13 Luzula sylvatica E1 . . . . . . . . . + . . . . . + 2 13 Aposeris foetida E1 . . . . . . . . . . . . . 1 . + 2 13 Polystichum lonchitis E1 . r . . . . . . . . . . . . . . 1 6 Lonicera nigra E2a . . + . . . . . . . . . . . . . 1 6 SSc Sambuco-Salicion capreae Sorbus aucuparia E3b . . . . . . . + . r . . . . . . 2 13 Sorbus aucuparia E3a + . . . . . . . r . + . . . . . 3 19 Sorbus aucuparia E2b . . . . . . + . . . . . . . + . 2 13 Sorbus aucuparia E2a . . . + . . . . . . . . . . . . 1 6 Sorbus aucuparia E1 . + . . r . . . 1 + + . . . . . 5 31 Sambucus racemosa E2a . . . . . . . . + + . . . . . . 2 13 RP Rubus dumetorum E2a . . . + . . . . . . . . . . . . 1 6 TG Trifolio-Geranietea Achillea distans E1 + . + . . + . . . . . . + + + . 6 38 Campanula rapunculoides E1 1 + . . . + . . . . + . . . . . 4 25 Digitalis grandiflora E1 . . . . . + . . . . . . . + + + 4 25 Vicia sylvatica E1 + . . . . . . . . . . . . + + . 3 19 Verbascum lanatum E1 . . . . . . . . + . . . . + 1 . 3 19 Vincetoxicum hirundinaria E1 . . . . . . . . . . . . + + + . 3 19 Iris graminea E1 . . . . . . . . . . . . 1 . . + 2 13 Polygonatum odoratum E1 . . . . . . . . . . . . + . + . 2 13 Verbascum sp. E1 . . . . . + . . . . . . . . . . 1 6 Laserpitium latifolium E1 . . . . . . . . . . . . 1 . . . 1 6 Arabis pauciflora E1 . . . . . . . . . . . . + . . . 1 6 Geranium sanguineum E1 . . . . . . . . . . . . + . . . 1 6 Laserpitium siler E1 . . . . . . . . . . . . + . . . 1 6 Lilium carniolicum E1 . . . . . . . . . . . . + . . . 1 6 Thalictrum minus E1 . . . . . . . . . . . . + . . . 1 6 Trifolium rubens E1 . . . . . . . . . . . . + . . . 1 6 Valeriana collina (V. wallrothii) E1 . . . . . . . . . . . . + . . . 1 6 Viola hirta E1 . . . . . . . . . . . . + . . . 1 6 Calamintha einseleana E1 . . . . . . . . . . . . . + . . 1 6 Hieracium umbellatum E1 . . . . . . . . . . . . . + . . 1 6 Hypericum perforatum E1 . . . . . . . . . . . . . . + . 1 6 Origanum vulgare E1 . . . . . . . . . . . . . . . + 1 6 EA Epilobietea angustifolii Rubus idaeus E2a + + . + . + + + 1 + + . . . + + 11 69 Fragaria vesca E1 + . + + . + + . + . . . . 1 + . 8 50 Urtica dioica E1 + . . . . . . . . + . . . . . . 2 13 Solanum dulcamara E1 . + . . . . . . . . . . . . . + 2 13 Galeopsis speciosa E1 . . . . . . . . . . . + . . . . 1 6 Hypericum hirsutum E1 . . . . . . . . . . . . . . . + 1 6 BA Betulo-Alnetea viridis Salix appendiculata E2a + . + . . . . + . . . . . . . + 4 25 MuA Mulgedio-Aconitetea Senecio ovatus E1 1 1 + + + 1 + + 1 1 1 + . + + . 14 88 Athyrium filix-femina E1 + + . + . . . . + + + . . . . . 6 38 Phyteuma ovatum E1 . . . + . . . . + + . + . . . . 4 25 Veratrum album E1 . . . . . . + + . . + + . . . . 4 25 Hypericum maculatum E1 + + . . . . . . + . . . . . . . 3 19 Aconitum degenii subsp. paniculatum E1 . . . . . . 1 + . . . + . . . . 3 19 Ranunculus platanifolius E1 . . . . . . + . + + . . . . . . 3 19 Polygonatum verticillatum E1 . . . . . . . + . + + . . . . . 3 19 Pleurospermum austriacum E1 . . . . . . . . . . . . + . . + 2 13 Aconitum lycoctonum s. lat. E1 . . . . . . . . . . . . . + . . 1 6 Centaurea montana E1 . . . . . . . . . . . . . . . + 1 6 Lathyrus occidentalis var. montanus E1 . . . . . . . . . . . . . . . + 1 6 ES Elyno-Seslerietea Sesleria caerulea subsp. calcaria E1 + + + 1 1 2 2 1 . . + + 2 1 + . 13 81 Betonica alopecuros E1 . . + . . + + . . . . . 1 1 . . 5 31 Centaurea haynaldii subsp. julica E1 . . + . . . . . . . . . . + . . 2 13 Laserpitium peucedanoides E1 . . . . + . . . . . . . . . . + 2 13 Festuca calva E1 . . + . . . . . . . . . . . . . 1 6 Phyteuma orbiculare E1 . . . . . . . . . . . . + . . . 1 6 FB Festuco-Brometea Brachypodium rupestre E1 + . + + . . . . . . . . . . 2 . 4 25 Thymus praecox E1 . . . . . 1 . . . . . . + . + . 3 19 Bromopsis transsilvanica E1 + . . . . . . . . . . . . . + . 2 13 Koeleria pyramidata E1 r . . . . . . . . . . . . . . . 1 6 Asphodelus albus E1 . . . . . . . . . . . . + . . . 1 6 Euphorbia cyparissias E1 . . . . . . . . . . . . + . . . 1 6 Linum viscosum E1 . . . . . . . . . . . . . + . . 1 6 Carlina acaulis E1 . . . . . . . . . . . . . . + . 1 6 Dianthus monspessulanus E1 . . . . . . . . . . . . . . + . 1 6 Pimpinella saxifraga E1 . . . . . . . . . . . . . . + . 1 6 MA Molinio-Arrhenatheretea Galium album E1 . . . r . . . . . . . . . . . + 2 13 Vicia cracca E1 . . . . . + . . . . + . . . . . 2 13 Angelica sylvestris E1 . . . . . . . . . . . . . + . . 1 6 Lotus corniculatus E1 . . . . . . . . . . . . . + . . 1 6 CU Calluno-Ulicetea Selaginella helvetica E1 1 + + . . . . . . . . . . . . . 3 19 Phyteuma zahlbruckneri E1 . . . . . . . . . . . . . . + . 1 6 TR Thlaspietea rotundifolii Adenostyles glabra E1 . 1 . + . + + . . + 1 1 . + . + 9 56 Ligusticum seguieri E1 . . . . . . . . . . . . + . + . 2 13 Arabis alpina E1 . . . . . . + . . . . . . . . . 1 6 Astrantia carniolica E1 . . . . . . . + . . . . . . . . 1 6 Gymnocarpium robertianum E1 . . . . . . . + . . . . . . . . 1 6 Cerastium subtriflorum E1 . . . . . . . . + . . . . . . . 1 6 Heracleum sphondylium subsp. pollinianum E1 . . . . . . . . . . . . . . . + 1 6 AT Asplenietea trichomanis Asplenium viride E1 + + + . . + 1 + 1 + . + . . . . 9 56 Campanula carnica E1 + + . . . + + + + . + . . . . . 7 44 Moehringia muscosa E1 + + . . . 1 + + + . . 1 . . . . 7 44 Asplenium trichomanes E1 + + . . . + . . 1 1 . + . . . + 7 44 Cystopteris fragilis E1 + . . . . . 1 + + 1 . + . . . + 7 44 Asplenium ruta-muraria E1 + + . + . + . . . . . . . . + . 5 31 Primula auricula E1 . . + . . + + . . . r . . . . . 4 25 Saxifraga crustata E1 . . + . . + + . . . . . . . . . 3 19 Kernera saxatilis E1 . . . . . + + . . . . . . . . . 2 13 Micromeria thymifolia E1 . . . . . + r . . . . . . . . . 2 13 Sedum maximum E1 . . . . . r . . . . . . + . . . 2 13 Festuca stenantha E1 . . . . . + . . . . . . . . . . 1 6 Hieracium pospichalii E1 . . . . . + . . . . . . . . . . 1 6 Paederota lutea E1 . . . . . + . . . . . . . . . . 1 6 Sedum album E1 . . . . . r . . . . . . . . . . 1 6 Carex brachystachys E1 . . . . . . + . . . . . . . . . 1 6 Polypodium vulgare E1 . . . . . . . . + . . . . . . . 1 6 ML Mosses and lichens (Mahovi in lišaji) Ctenidium molluscum E0 1 2 + 1 + 2 1 2 1 2 1 1 . . . 1 13 81 Tortella tortuosa E0 + 1 + + 1 1 1 1 + + . . . + . . 11 69 Isothecium alopecuroides E0 + + + + 1 . . + + 1 . 1 . . . 1 10 63 Neckera crispa E0 + + + . 1 1 1 1 1 1 . + . . . . 10 63 Schistidium apocarpum E0 + + . + . 1 . + + + + 1 . . + . 10 63 Polytrichum formosum E0 1 2 + + + . . + 2 1 . . . + . . 9 56 Peltigera canina E0 . + + . + . + + + + . + . . . + 9 56 Fissidens dubius E0 1 . . . 1 . + . + . + + . . . . 6 38 Atrichum undulatum E0 1 1 + . . . . . . . + . . . . . 4 25 Conocephalum conicum E0 . . . . . . + + + . . + . . . . 4 25 Homalothecium philippeanum E0 . + . . . 1 . + . . . . . . . . 3 19 Hypnum cupressiforme E0 . . + + . + . . . . . . . . . . 3 19 Plagiothecium denticulatum E0 . . . + . . . . . . + + . . . . 3 19 Dicranella heteromalla E0 . . . + . . . . . . + . . . + . 3 19 Mnium thomsonii E0 . . . . . . + 1 . . + . . . . . 3 19 Bartramia halleriana E0 . . . . + . . . 1 . . . . . . . 2 13 Metzgeria furcata E0 . . . . . . + 1 . . . . . . . . 2 13 Plagiochila porelloides E0 . . . . . . . + . . + . . . . . 2 13 Cladonia pyxidata E0 . . . . . . + . + . . . . . . . 2 13 Homalothecium lutescens E0 . . . . . . . . . . . . . . + 1 2 13 Distichium capillaceum E0 + . . . . . . . . . . . . . . . 1 6 Marchantia polymorpha E0 + . . . . . . . . . . . . . . . 1 6 Rhytidiadelphus triquetrus E0 + . . . . . . . . . . . . . . . 1 6 Peltigera leucophlebia E0 + . . . . . . . . . . . . . . . 1 6 Plagiothecium undulatum E0 . . + . . . . . . . . . . . . . 1 6 Cladonia furcata E0 . . + . . . . . . . . . . . . . 1 6 Bryum capillare E0 . . . . + . . . . . . . . . . . 1 6 Hylocomium splendens E0 . . . . + . . . . . . . . . . . 1 6 Bryum argenteum E0 . . . . . . + . . . . . . . . . 1 6 Dicranum scoparium E0 . . . . . . . . 1 . . . . . . . 1 6 Mnium marginatum E0 . . . . . . . . . 1 . . . . . . 1 6 Brachythecium velutinum E0 . . . . . . . . . . + . . . . . 1 6 Anomodon attenuatus E0 . . . . . . . . . . . . . . . + 1 6 Brachythecium rutabulum E0 . . . . . . . . . . . . . . . + 1 6 Legend - Legenda AR Limestone with chert - apnenec z rožencem ARG Limestone with chert and claystone - apnenec z rožencem in glinavcem DR Dolomite with chert - dolomit z rožencem DRG Dolomite with chert and claystone - dolomit z rožencem in glinavcem AL Limestone with marlstone - apnenec z laporovcem ALR Limestone with chert and marlstone Re Rendzina - rendzina Rj Brown calcareous soil - rjava pokarbonatna tla Number of relevé (Zaporedna številka popisa) 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 Pr. Fr. Number of relevé (Zaporedna številka popisa) 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 Pr. Fr. Number of relevé (Zaporedna številka popisa) 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 Pr. Fr. Number of relevé (Zaporedna številka popisa) 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 Pr. Fr. Table 9 (Preglednica 9): Saxifrago cuneifolii-Fagetum fraxinetosum orni Number of relevé (Zaporedna številka popisa) 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 31 32 33 34 35 36 37 38 39 40 41 42 Database number of relevé (Delovna številka popisa) 247607 255462 255305 255455 255454 247634 255320 255319 247635 255304 255306 255322 255318 255329 255312 255313 255316 255314 255495 255463 255472 255473 255465 255486 255484 255471 255470 255315 255485 255487 255449 255469 255483 255466 255468 255467 255496 255335 255441 255410 255411 255412 Elevation in m (Nadmorska višina v m) 890 760 880 1050 810 1040 950 950 980 900 910 1100 930 890 740 740 710 730 630 1030 980 1110 540 630 720 500 660 670 520 520 500 780 780 700 760 750 630 760 760 990 980 920 Aspect (Lega) NE NNW N NNW N W E SE SWW N N NE NE NE NE NE NE NE NE N NE NE NE NE NE NE NE NE NE NE NE NE NE N NE NE NE NE NNW N N N Slope in degrees (Nagib v stopinjah) 40 35 40 40 35 35 35 40 35 35 30 40 40 40 35 35 30 40 35 30 35 35 30 40 35 35 30 30 30 35 35 30 35 30 30 30 30 30 35 40 35 40 Parent material (Matična podlaga) AR AGR ALR ALR ALR DR DR DR DR ALR ALR DR AR AGP AL AL AL AL AL AGP AGR AR AR AGR AGR AGR AGR AL AL AGR ALR ALR ALR AGR ALR AGR AL AL AL AL ALR ALR Soil (Tla) Re Re Re Re Re Re Re Re Re Re Rj Re Re Rj Re Re Re Re Re Rj Re Re Re Re Re Re Re Re Rj Re Dy Dy Dy Re Dy Dy Rj Re Rj Re Re Re Stoniness in % (Kamnitost v %) 30 60 50 60 60 20 5 10 10 40 30 60 10 10 30 40 40 70 40 10 30 40 30 30 70 10 70 30 20 15 10 20 20 40 30 30 15 20 20 70 90 70 Cover in % (Zastiranje v %): Upper tree layer (Zgornja drevesna plast) E3b 90 100 95 95 85 90 90 90 70 90 95 90 80 90 90 95 100 95 90 100 95 95 100 90 90 90 100 95 90 90 95 100 80 95 95 95 90 70 70 90 90 90 Lower tree layer (Spodnja drevesna plast) E3a 20 . . . . 10 . . 20 . . . 20 . . . . . . . . . . 5 10 5 . . 10 5 . 5 20 5 . . 10 30 30 . 5 10 Shrub layer (Grmovna plast) E2 20 10 20 20 30 5 10 5 5 20 30 5 10 5 5 5 5 5 20 3 10 5 5 10 10 5 5 5 15 10 5 5 10 5 5 5 5 20 10 20 5 5 Herb layer (Zeliščna plast) E1 70 70 60 70 40 80 80 80 50 60 60 50 80 60 60 60 60 50 60 90 80 70 70 80 60 70 60 60 60 70 60 70 70 70 60 70 70 60 60 45 40 70 Moss layer (Mahovna plast) E0 5 20 30 30 30 10 5 5 10 15 20 15 10 10 15 20 10 20 20 10 10 20 20 20 20 30 20 5 20 20 40 10 15 30 30 30 30 5 10 25 20 20 Maximum diameter of trees (Največji prsni premer dreves) cm 40 30 30 30 30 40 30 35 40 40 40 35 40 40 40 35 35 30 30 35 30 35 35 30 40 40 40 25 70 45 30 35 50 35 40 40 50 35 40 35 50 40 Maximum height of trees (Največja drevesna višina) m 16 16 18 10 18 18 16 18 18 20 22 18 18 20 24 20 22 16 18 18 18 16 19 16 18 20 26 22 28 22 20 22 22 20 24 26 34 18 22 16 17 16 Number of species (Število vrst) 68 69 70 73 61 71 87 87 54 64 68 63 83 63 79 84 90 73 77 59 54 68 64 73 79 83 68 95 91 87 66 49 55 73 62 53 89 83 79 72 67 80 Relevé area (Velikost popisne ploskve) m2 200 400 400 400 400 200 400 400 200 400 400 400 400 400 400 400 400 400 400 400 400 400 400 400 400 400 400 400 400 400 400 400 400 400 400 400 400 400 400 400 400 400 Date of taking relevé (Datum popisa) 6/6/1993 5/29/1987 5/23/1988 5/29/1987 5/27/1987 7/2/1991 8/17/1989 8/17/1989 7/2/1991 5/23/1988 5/23/1988 8/17/1989 8/17/1989 8/17/1989 5/6/1989 5/6/1989 8/9/1987 5/6/1989 8/28/1988 6/17/1987 19..6.1987 19..6.1987 6/22/1987 7/12/1991 7/12/1991 6/22/1987 6/22/1987 5/6/1989 7/12/1991 7/12/1991 5/23/1987 6/22/1987 7/12/1991 6/21/1987 6/21/1987 6/21/1987 8/26/1988 5/6/1989 6/24/1990 6/11/1988 7/6/1991 7/6/1991 Locality (Nahajališče) Kacenpoh - Šprickovbl Koblar - Bideržuna Trtnikarski gozd Koblar Koritnica - Zajtl Kacenpoh -Kobla Kacenpoh-Fergajnar Kacenpoh - Fergajnar Kacenpoh-Kobla Trtnikarski gozd Trtnikarski gozd Kacenpoh - Fergajnar Kacenpoh - Fergajnar Kacenpoh - Fergajnar Šprickovbl -Trtnikarski gozd Šprickovbl -Trtnikarski gozd Šprickovbl -Trtnikarski gozd Šprickovbl -Trtnikarski gozd Prodarjeva grapa Grant - Temerce Grant - Luken Grant - Luken Grant - Čumik Grant - Čumik Grant - Čumik Grant - Čumik Grant - Čumik Šprickovbl -Trtnikarski gozd Grant - Čumik Grant - Čumik Koritnica - Jelenca Grant - Čumik Grant - Čumik Grant - Čumik Grant - Čumik Grant - Čumik Porezen - Lanek Kacenpoh Batava Koblar Koblar - Bideržuna Koblar - Bider~una Quadrant (Kvadrant) 9749/4 9849/2 9749/4 9849/2 9849/1 9749/4 9749/4 9749/4 9749/4 9749/4 9749/4 9749/4 9749/4 9749/4 9749/4 9749/4 9749/4 9749/4 9849/2 9749/3 9749/3 9749/3 9849/1 9849/1 9849/1 9849/1 9849/1 9749/4 9849/1 9849/1 9849/1 9849/1 9849/1 9849/1 9849/1 9849/1 9849/2 9749/4 9749/4 9849/2 9849/2 9849/2 Coordinate GK Y (D-48) m 419719 416438 419801 416659 416045 420173 419914 419905 420386 419806 419680 419771 419990 419996 419933 419912 419947 419903 418738 413146 412416 412387 414899 414574 414492 414938 414457 420027 414920 414943 414971 414264 414352 413826 413640 413527 418903 420100 421587 416623 416711 416726 Coordinate GK X (D-48) m 5119956 5117826 5119743 5117576 5117376 5121281 5121128 5121048 5121008 5119710 5119946 5121180 5121062 5121084 5119892 5119936 5119962 5120001 5117254 5119002 5118494 5118188 5117442 5117646 5117552 5117474 5117668 5119925 5117462 5117428 5116930 5117582 5117540 5117734 5117724 5117846 5116795 5120820 5118574 5117722 5117706 5117786 Diagnostic species of the association (Diagnostične vrste asociacije) Pr. Fr. VP Veronica urticifolia E1 1 + + 1 1 1 1 1 + + + 1 + 1 1 1 1 1 1 2 1 1 2 + 1 1 1 1 + . + 1 1 1 1 1 1 + 1 1 1 1 41 98 VP Calamagrostis arundinacea E1 1 . + 2 1 1 2 + + + 1 1 3 2 1 1 2 . 1 3 2 2 1 + 2 + 1 2 2 2 4 3 3 3 2 3 2 1 + 2 + 1 40 95 VP Saxifraga cuneifolia E1 + . + + + . + . . + 1 1 1 1 1 1 1 1 + 1 + 1 1 + + 1 + 1 + 1 + + . + 1 1 1 + + . 1 1 36 86 VP Luzula luzuloides E1 . . + . + + + . + . + + + + + + + . . 1 1 + . + + + + + . + 1 . + + + 1 . + . . + + 29 69 QP Tanacetum corymbosum E1 + . 1 1 . + . . + + . . . . + . + . . . 1 . + + . + + + + + + . . + . . . 1 + . . . 20 48 VP Huperzia selago E1 . . + 1 . . + + . + + + + + + . . . . 1 + . . . . + . . + . 1 + . . . + . . . . . + 18 43 QF Festuca heterophylla E1 . . . . . . . + + . . . + . + . + . + + . . . . . . . + + + . . + . + . . + + . . . 14 33 ES Campanula witasekiana E1 . . . + . + + + . . . . + . . . . . . + . + . . . . . . . . . . . . . . . + . . . + 9 21 FB Carex humilis E1 + . . . . + + 1 + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + + 7 17 TG Clinopodium vulgare E1 . . . . . . + + . . . . + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 3 7 Geographical differential species (Geografske razlikovalne vrste) QP Sesleria autumnalis E1 . + . . + . . . . . . . . . . . . . + . + . . . + + . . + + + . . + + + . . . . . . 12 29 FS Lathyrus vernus subsp. flaccidus E1 . + . . . . . . . . . . . . . . . . . . . + + + . + . . + + . . . + . . . . . . . + 9 21 AF Anemone trifolia E1 . . 1 . . . + . + 1 + . . . . + . . . . . . . . . . . . . . + . . . . . + 1 . . . . 9 21 VP Larix decidua E3b . . . . 2 . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . 1 . + + . + 6 14 VP Larix decidua E3a . . . . . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . . . . . . . . 1 2 MuA Aconitum angustifolium E1 . . . . . . . . . . . . . . + . . . . . . . . . + . . . . . . . + . . . 1 . . . . 4 10 Differential species of the subassociation (Razlikovalnica subasociacije) QP Fraxinus ornus E3b . . + . . . . . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . . . . . 2 5 QP Fraxinus ornus E3a + + . . . . . . . . . . . . . . . + 1 . . . + . . + . + + . + . . + . . . + + . . . 12 29 QP Fraxinus ornus E2b + 1 . . . . . . . . . . . . + . . . . . . . + + . . . + . + + . + . . . . . . . . . 9 21 QP Fraxinus ornus E2a 1 + . . 1 . . + . + + . + . . + + . + . . . . . + + . + 1 + . + + . + + . 1 + . . . 21 50 QP Fraxinus ornus E1 . . . . . . + . . . + . . + + . . . . . . . . . + + + . . . . . . + . . + . + . . . 10 24 AF Aremonio-Fagion Cyclamen purpurascens E1 + 1 + 1 1 + 1 1 1 + 1 + 1 1 + 1 1 1 1 1 1 1 + 1 1 1 1 + . . + + . 1 1 . + 1 1 . + + 37 88 Cardamine trifolia E1 . . + . . + + . . + + + + + 1 1 . 1 1 1 1 + + 1 . + + . . + . . . 1 + . 1 1 + 1 1 1 28 67 Cardamine enneaphyllos E1 r . . . . 1 1 1 1 1 1 + 2 + 1 2 + 1 . + 1 + 1 1 + 1 1 1 . 1 . . . . . . . . . . . . 24 57 Aremonia agrimonoides E1 . . . . . . . . . . . . + . + + . . . . . . . . . . + . . . . . . . . . . . . . . 4 10 Hacquetia epipactis E1 . . . . . 1 . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . . . . 2 5 Anemone x pittonii E1 . . . . . . + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . 2 5 Lamium orvala E1 . . . . . . . . + . . . . + . . . . . . . . . . . . . . . . . . . . . . . . . . . . 2 5 Scopolia carniolica E1 . . . . . . . . . + . + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 2 5 Knautia drymeia E1 . . . . . . . + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 2 Omphalodes verna E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 2 . . . 1 2 EC Erythronio-Carpinion Primula vulgaris E1 . + . . + + + + + . . . + + 1 1 1 + 1 . + . + + + 1 1 1 + 1 + + + 1 . + + 1 1 . . . 30 71 TA Tilio-Acerion Acer pseudoplatanus E3b . . . . 1 . . . . . . . . . . . . . . . . . . . . . . . + . . + + . . . + . . . . r 6 14 Acer pseudoplatanus E3a . . . . . . . . + . . . . . . . . . . . . . . . . . . . + . . . . . . . . + + . + . 5 12 Acer pseudoplatanus E2b . . . + 1 . . . + + . + . . . . . . . . . . . . . . . . + . . . . . . . . . . + + . 8 19 Acer pseudoplatanus E2a . + . . . . . . . . . . . . . . . . . . . . . . . + . . . + . + + + . . + . + . . . 8 19 Acer pseudoplatanus E1 . + . + + + + . + . + . . + + . . . . . . . + + . + + + . + 1 + . + . 1 + 1 . + . 22 52 Aruncus dioicus E1 . . . + . + + + + + + . + . . + . + + . . + . . . . . . + . . . . + + . + + 1 . + + 20 48 Polystichum aculeatum E1 . . + . . . . . . 1 . + 1 1 . . + + . . . . . . . . . r . . . . . . . . . . 1 . . . 9 21 Tephroseris pseudocrispa E1 . . . . . . . . . . . . . . + . + + . . . . . . . . . + . . + . . . . . + + . . . . 7 17 Euonymus latifolia E2 + . . . . . . . . + . . + + . + . . . . . . . . . . . . . . . . . . . . . + 1 . . . 7 17 Acer platanoides E3b . + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 2 Acer platanoides E2b . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . . . . . 1 2 Acer platanoides E2a . . . . + . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . . . 2 5 Acer platanoides E1 . . . . . . . + . . . . . . . + . . . . . . . . . . + + . . . . . . . . + . + . . . 6 14 Dryopteris affinis E1 . . . . . . . . . . . . + + . . . . . . . . . . . . . . . . . . . . + . . . . . . . 3 7 Ulmus glabra E3a . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . 1 2 Ulmus glabra E2a . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . . . . . . 1 2 Ulmus glabra E1 . . . . . . . . . . . . . . . . . . . . . . . . . . 1 . + . + . . . . . . . . . . . 3 7 Polystichum braunii E1 . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . 1 . + 3 7 Geranium robertianum E1 . . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . . . . . . . . . + . 2 5 Polystichum setiferum E1 . . . . . . . . . . . . . + . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 2 Thalictrum aquilegiifolium E1 . . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . . . . . . . . . . . 1 2 Juglans regia E2b . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . . 1 2 Juglans regia E2a . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . 1 2 Juglans regia E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . 1 2 Polystichum x illyricum E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . 1 2 Tilia platyphyllos E3a . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + 1 2 FS Fagetalia sylvaticae Fagus sylvatica E3b 5 5 5 5 3 5 5 5 5 5 5 5 5 5 5 5 5 4 5 5 5 5 5 5 5 5 5 5 5 5 5 5 5 5 5 5 4 4 4 5 5 5 42 100 Fagus sylvatica E3a 1 . . . . + . . 1 . . . 1 . . . + . . . . . . 1 1 . . . 1 + . . 1 . . . . . + . + 1 13 31 Fagus sylvatica E2b . + + 1 1 + + . 1 1 1 + . 1 1 1 + + 1 + . + + + 1 + . + + 1 1 + 1 . . + 1 1 . 2 + 1 34 81 Fagus sylvatica E2a . . . . . . . . . . . . + . . . . . . . + . . . . . . . 1 1 . . . . . 1 . . + . . . 6 14 Fagus sylvatica E1 + . . . . . . . . . + . . . + . + . . + . . . . . + + + 1 1 . . . . 1 . 1 . . . + . 13 31 Galium laevigatum E1 1 1 1 1 1 1 1 1 1 1 1 + 1 + 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 + 1 1 . 1 + + 1 1 1 1 + 41 98 Daphne mezereum E2a + 1 1 1 1 + + + + + + . + . + + + + 1 1 1 1 + 1 1 1 + 1 1 1 + + 1 + + . + 1 1 + + + 39 93 Mercurialis perennis E1 1 + 1 + 1 1 1 1 2 1 1 1 1 + . 1 1 + + 1 1 1 1 1 1 1 1 1 + + + + + 1 + . . 1 1 1 1 + 39 93 Prenanthes purpurea E1 + + . + + 1 1 1 1 + + 1 + 1 1 + 1 . + + . + 1 + 1 + 1 + + + 1 1 1 1 1 1 1 + 1 . + + 38 90 Dryopteris filix-mas E1 . + + + 1 + + . + 1 + + + + + + + 1 + + + + + . 1 + 1 + 1 1 + + + 1 1 . + + + + + 1 38 90 Actaea spicata E1 . . 1 + + + + + + 1 1 1 + + + + + + + . + + + + + + . + 1 1 . + + + + + 1 + 1 + + + 37 88 Salvia glutinosa E1 + + + . . + . . 1 + + + 1 + + + 1 + 1 . + + 1 + 1 + + 1 1 + + 1 + + 1 1 1 + 1 + . . 35 83 Lonicera alpigena E2a + 1 1 2 1 + + . . 1 2 + 1 1 + + . + . . . 1 . . + . + + . . + 1 1 + . 1 + + 1 + . + 29 69 Asarum europaeum subsp. caucasicum E1 + 1 + . 1 . . . . + . . + + + 1 1 1 . . . + + + . 1 1 1 1 1 . . . + . . 1 + + + . + 25 60 Lilium martagon E1 + + + + + . . . . + + + . . . + + . . . + + 1 + + + 1 + + 1 . . + . . . . + . . . . 22 52 Symphytum tuberosum E1 . . . . . + . + + . . . . . + . + . . + . + 1 + + + + + . + + + + + . + 1 . + . . . 21 50 Lathyrus vernus subsp. vernus E1 . + . 1 1 . . . . . 1 . . . . . . . + 1 1 1 1 1 1 1 1 + 1 1 . . . + . . . . . . . + 18 43 Pulmonaria officinalis E1 . . + . + + . . + + . + . + + + 1 . . . . + . . . . . + + + . . . + . . + + + . . . 18 43 Viola reichenbachiana E1 + + . + . . . . . . . . + + + . + . + . . . . . + . . + + + + . + . + . . . . + . + 17 40 Sanicula europaea E1 . . . . . . + . . . + + 1 1 + . + . . + + . . . . . + 1 + + . . . . + + + . + . . . 17 40 Campanula trachelium E1 + + . . . + 1 1 + . + . 1 . + + 1 + + . . . . . . . . + . . + . . . . . + + . . . . 17 40 Euphorbia amygdaloides E1 . . . . . . . + + . . . . . + . + . + + . . . . + . + + + + . + + + . . . . + . . . 15 36 Mycelis muralis E1 . . . . . + . . + + . + + . + . . + + . . . . . . . . + . . . . . . . + 1 . + . 1 + 14 33 Laburnum alpinum E3a + + + 1 + . . + + . . + . . . . . . + . . . . . . . . . . . . . . + . . . . + + . . 12 29 Laburnum alpinum E2b . . + . . . + + . . . . . + + . + + . 1 . . . . + . . + . . . . . . . + . . . . . . 11 26 Laburnum alpinum E2a + . . . . 1 . + . . . + + . . + . . . . . . . . . . . . . . . . . . . . . . 1 + . . 8 19 Laburnum alpinum E1 + . . . . . . . + . . . . . . . + . + . . . . . . . . + . . . . . . . . + + . . + + 9 21 Melica nutans E1 . + . . . + + . . . + . . . . . . + . . . . . . + + . + . . . . . . . . . . + + . + 11 26 Epipactis helleborine E1 . . . . . + 1 1 + . . . + + . . . . + . . . . . . . . . . + . . . . . . . + + . . . 10 24 Neottia nidus-avis E1 . . . . . . . + . + . . . . + . . . . . . . . . . . . + + . . + . . + + + + . . . 10 24 Paris quadrifolia E1 . . + . + + . . . + + + . . . . . . . . . . . . . . + . . . . . . . . . . . . + + . 9 21 Polygonatum multiflorum E1 . . . . . + + . + + + . + . . . . . . . . . . . . + . . + . . . . . . . . + . . . . 9 21 Festuca altissima E1 . . . . . . . . + . + . + 1 . . . . . . . + . . . . + . . . . . . . . . . . + . 1 1 9 21 Galeobdolon flavidum E1 . . . . . + . . 1 . . . . . . . . + . . . + . . . + . . . . . . . . . . . . . + 1 + 8 19 Petasites albus E1 . . . . . . + + . . . + . 1 . . . . . . . . . . . . . . . . . + . + . . + . . . . . 7 17 Scrophularia nodosa E1 . . . . . . . . . + . + + . . . + + . . . . . . . . + . . . . . . . + . . . . . . . 7 17 Epilobium montanum E1 . . . . . . . . . . . + . + . . . + . . . . . . . . . . . . . . . . . . . . . + + + 6 14 Cephalanthera damasonium E1 + . . . . . . . . . . . . . . . + . + . . . . + . . . . + . . . . . . . . . . . . . 5 12 Euphorbia dulcis E1 . . . . . . . . . . . . . . . . . . + . . . . . + . . . . . . . . . . . + + + . . . 5 12 Sambucus nigra E2 . . . . . . . . . . . . . + . . . + . . . . + . . . . . . . . + . . . . . . . . . . 4 10 Poa nemoralis E1 . . . . . . . + . . . . . . . . . . . . . + . . . . . . . . . . . . . . . . . . . . 2 5 Prunus avium E3b . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . 1 2 Prunus avium E3a . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . 1 2 Prunus avium E2a . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . . . . . 1 2 Prunus avium E1 . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . . + . . . . . 2 5 Heracleum sphondylium E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . + + . . . . . . . . . . . . 2 5 Orobanche salviae E1 . . . . . . . . . . . . + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 2 Hordelymus europaeus E1 . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . . . . . . . 1 2 Carpinus betulus E3a . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . 1 2 Carpinus betulus E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . 1 2 Tilia cordata E2b . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . . . . . 1 2 Tilia cordata E2a . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . 1 2 Galium odoratum E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . 1 2 QP Quercetalia pubescenti-petraeae Melittis melissophyllum E1 . 1 . . . . + + + . . . . . 1 + 1 . + + + . 1 1 1 1 + + 1 1 . + . + . . + 1 . . . . 22 52 Sorbus aria E3b . . . . . . . . . . . . . . + . . . . . . . . + + . . . . . . + + 1 . . . . . . . . 6 14 Sorbus aria E3a + + . + + . . + + . . . . . . . . . + . . . . . . . . . . + + . . . 1 . . + + + . . 13 31 Sorbus aria E2b 1 1 + 1 1 . . 1 . . + . . . . + + + 1 . . + + + + . . . . . + . . + . . . . . . . + 18 43 Sorbus aria E2a 1 . . . . . 1 . . . . . 1 . . . . . . . 1 + . + . 1 . + + + . . + . . . . + + 1 . . 14 33 Sorbus aria E1 . . . . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . + . . . . . . . . 2 5 Ostrya carpinifolia E3b . . . . . . . . . . . . . . + + . 2 . . . . . + + . . . . . . . . + . . . . . . + r 8 19 Ostrya carpinifolia E3a + + 1 . 1 . . . . + . . . . . . + . 1 . . . . . . + . + r + . . . . + . . 2 1 + . . 15 36 Ostrya carpinifolia E2b . . . . 1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 2 Ostrya carpinifolia E2a . + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . 2 5 Hypericum montanum E1 + . + . . . . . + . + . + + . . . . + . . . . . . . . + . + . . . . . . + + + . . . 12 29 Arabis turrita E1 + . . . . . . . + + . . . . . . . + . . . . . . + . + . . . . . . . . . . + . + . . 8 19 Convallaria majalis E1 . + + + . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . . . + . . . . 5 12 Cephalanthera rubra E1 . . . . . + . + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 2 5 Epipactis muelleri E1 . . . . . . + + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 2 5 Sorbus x thuringiaca E2b + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 2 Primula veris subsp. columnae E1 . . . + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 2 Calamintha sylvatica E1 . . . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . . . . . . . . . . 1 2 Euonymus verrucosa E2 . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . 1 2 Orchis mascula subsp. speciosa E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . 1 2 QR Quercetalia roboris Hieracium racemosum E1 . . . . . . + + . . . . + + + + 1 . + . . . . + . . . + + . + . + . . . + . . . . . 14 33 Rubus hirtus E2a . . . . . . . . . + . + . . . . . . . . . . . . . . . . + + . . + . . . . . . . . . 5 12 Betonica officinalis E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 + + . . . . . . . . . . . 3 7 Hieracium sabaudum E1 . . . . . . . . . . . . . . . . + . . . . . . + . . . . . . . . . + . . . . . . . . 3 7 Quercus petraea E3b . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . 1 2 Quercus petraea E1 . . . . . . . . . . . . . . . . . . . . . . . . . . + + . + . . . . . . . . . . . . 3 7 Betula pendula E2b . . . . + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 2 Hieracium lachenalii E1 . . . . . . . + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 2 Pteridium aquilinum E1 . . . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . . . . . . . . . . 1 2 Castanea sativa E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . 1 2 QF Querco-Fagetea Carex digitata E1 + + + 1 + + + + + . + + + + + + + . + + + + + 1 + + + + + . + + . + 1 + + 1 + . + . 36 86 Lonicera xylosteum E2a + + + . + . + . . + + . + + + + + + + . . + + + + + + + + + + + + . . . 1 + + . . . 29 69 Hepatica nobilis E1 . 1 1 1 1 1 1 + + + . + + + + 1 1 1 1 . . . + 1 1 1 . 1 + 1 . . . . . . . 1 1 . . . 26 62 Anemone nemorosa E1 . . . + . + 1 . + + + 1 + . . . . . . + . + 1 . + 1 1 . 1 + 1 1 1 1 . . + . . . + . 22 52 Corylus avellana E2a . . . . . . . . . + + + . . + . . . + . . . . + + + . . + + + . . . . . + . . . . . 12 29 Vinca minor E1 . . . . + . . . . . . . . . + + . . . . . . . . . + + 2 1 1 + . . . . . + 1 . . . . 11 26 Dactylorhiza fuchsii E1 . . + + . + + + . . + . + . + + . . . . . . . . . . + . . . . . . . + . . . . . . 11 26 Hedera helix E3a . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 1 . . . . . . . 2 5 Hedera helix E1 . . . . . . . . . . + . . . r . . + . . . + + . + + . . . + 1 + . . . . . . . . . 10 24 Rosa arvensis E2a . . . . . . . . . . . . + . + + . + . . . . . . . + + + + . . . . . . . . + . . . 9 21 Viola riviniana E1 . . . . . . + . . . + . . . + + + + . . . . . . . . . + . . . . . . . . . . . . . . 7 17 Aegopodium podagraria E1 . . . . . . . . . . . . . . + . + . . . . . . . . . . + 1 + . . . + . . . . . . . . 6 14 Clematis vitalba E2a . . . . . . . + . . . . . . + . . + . . . . . . . + . + . . . . . . . . . . . . . 5 12 Acer campestre E3a . . . . . . . . . . . . . . . . . . . . . . . . . + . . . + . . . . . . . . . . . . 2 5 Acer campestre E2b . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . . . . . 1 2 Acer campestre E2a . . . . . . . . . . . . . . . . . . . . . . . . . + . . + . . . . . . . . . . . . . 2 5 Acer campestre E1 . + . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . + . . . + . . . . 4 10 Luzula nivea E1 . . . . . . . . . . . . . . . . . . . . . . . + + . . . . . . . . . . . . . . . + + 4 10 Cephalanthera longifolia E1 . . . . . . . + . . . . + + . . . . . . . . . . . . . . . . . . . . . . . . . . . . 3 7 Veratrum nigrum E1 . . . . . . . . . . . . . . . . . . . . . . . r + . . . . . . . + . . . . . . . . . 3 7 Cruciata glabra E1 . . . . . . . . . . . . . . . . . . . + + . . . . . . . . . . . . . . . . . . . . . 2 5 Platanthera bifolia E1 r . . . . . . . . . . . . . r . . . . . . . . . . . . . . . . . . . . . . . . . . . 2 5 Taxus baccata E2b . . . . . . . . . . . . . + . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 2 Moehringia trinervia E1 . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . . . . 1 2 Pyrus pyraster E2a . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . 1 2 Malus sylvestris E2a . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . 1 2 Galium schultesii E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . 1 2 Listera ovata E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . 1 2 EP Erico-Pinetea Cirsium erisithales E1 1 + + + + + + + + . . . + . + + 1 + + + + + + + + + . + + + . . + . . + . + + + . . 30 71 Rubus saxatilis E1 . 1 + 1 1 + 1 + . . . . . . . . . . . . . . 1 1 + + . . + + . + + . . . . + . . . + 17 40 Carex alba E1 . . . + . . 1 1 . . . . . . . . . . . . + . 2 3 + 3 . . + . + . . + . . . . . + . . 12 29 Peucedanum austriacum E1 + 1 1 . . . + 1 . . . . . . + + . . . . . . . . 1 . + . + . . . . . . . 1 . . . . 11 26 Calamagrostis varia E1 + 2 . . . + + 3 + . . . . . + + + . . . . . . . . . . . . . . . . . . . . 1 . . . 10 24 Aquilegia nigricans E1 . . . + . + + + . . . . . . r + . . . . . . . . . + + + . . . . . . . . + . . . . 10 24 Buphthalmum salicifolium E1 + . . . . . 1 1 . . . . + . . . + . + . . . . + . . . + . . . . . . . . . . + . . . 9 21 Erica carnea E1 . + . + . . + . . . . . . . . . . . . . . . . . . + . . . . + . . . . . . . . . . . 5 12 Rhododendron hirsutum E2a . . . + . . + + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 3 7 Carex ornithopoda E1 . . . . . . + + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 2 5 Cotoneaster tomentosus E2a + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 2 Epipactis atrorubens E1 . . . . . . . . . . . . + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 2 Polygala chamaebuxus E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . 1 2 VP Vaccinio-Piceetea Solidago virgaurea E1 1 1 1 1 1 1 1 1 + + + + 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 + 1 1 1 + 1 . 1 + + 1 + + 41 98 Hieracium murorum E1 + + + 1 . 1 1 1 . . + + 1 + + 1 1 + 1 1 1 + + + + + + 1 + 1 + 1 + 1 1 1 1 + + . + 1 38 90 Homogyne sylvestris E1 1 1 1 2 1 1 1 1 . 1 1 + 1 1 1 1 1 + 2 2 1 2 2 1 1 1 + 1 . 1 + 1 + 1 . . 1 + 2 1 + 2 38 90 Oxalis acetosella E1 . 1 . 1 1 + + . . + 1 1 + . 1 + . . . 1 + + 1 + 1 1 1 1 1 1 + 1 1 1 1 1 1 + + 1 1 1 34 81 Gentiana asclepiadea E1 + 1 + 1 . + . . . + + . . + . + + + + + 1 + + + + + . + + + + + . 1 1 + 1 + 1 + . + 32 76 Valeriana tripteris E1 + 1 1 2 . . . 1 + 1 1 + 1 . + 1 + 1 1 + 1 1 + + + + . 1 . . . . . + + . + + 1 1 . 1 30 71 Rosa pendulina E2a + 1 1 1 1 + + . . 1 + . . . + . . . . 1 + 1 + + + + . + . + . 1 + + + . + . + 1 1 1 28 67 Maianthemum bifolium E1 . 1 1 . . + + + . + 1 + . . + + + . . 1 + + 1 + + . + + + . . 1 + + + . 1 + + . . + 28 67 Aposeris foetida E1 . . . + . 1 1 + + . . + . . . . . . . + . . + + + + + . 1 1 + . . . . . . . . . . . 15 36 Picea abies E3b . + . . + . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . 1 . + + . . 6 14 Picea abies E3a . . . . . . . . . . . . + . . . . . + . + . . . . . . . . . . . . . . . . . + . . . 4 10 Picea abies E2b . . + . . . . + . + + . . . . . . . . + . . . + . + . . . . . . . . . . . . . . . . 7 17 Picea abies E2a + . . . 1 + + . . . . . . . + + + + + . . + + . . . . + + . . . + + . . . . . . . . 15 36 Picea abies E1 . + . + . . . + . . . . . + . . + . . + . . + + + . . . . . . . . . . + + . . + . . 12 29 Vaccinium myrtillus E1 . + . + . . + + . . + . . . . . . . . + . . . . . . . + . . + . . . . . + + . . . . 10 24 Phegopteris connectilis E1 . . . . . . . . . . . . . + . . . . . + . . . . . . . . . . 2 . . + + + + . . + . . 8 19 Gymnocarpium dryopteris E1 . . . . . . . . . . . . . . . . . . . . . . + . + . + . . . . . . 1 1 + 1 . . . . . 7 17 Clematis alpina E2a . . 1 2 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + 1 + 1 6 14 Orthilia secunda E1 . . . . . . + + . . + . + + . . . . . . . . . . . . . . . . . . . . . . . . . . . . 5 12 Dryopteris dilatata E1 . . . . . . . . . . . + . . . . . . . + . . . . . . . . . . . . . . . . . . . . . . 2 5 Luzula pilosa E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . + . . . . . . . . . . . 2 5 Lonicera nigra E2a . . + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 2 Vaccinium vitis-idaea E1 . . . + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 2 Abies alba E2b . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . 1 2 Abies alba E2a . . . . . . + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 2 Corallorhiza trifida E1 . . . . . . . . . . . . . . r . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 2 Avenella flexuosa (Deschampsia flexuosa) E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . 1 2 Polystichum lonchitis E1 . . . . . . . . . + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 2 Luzula luzulina E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . 1 2 Laserpitium krapfii E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . 1 2 SSc Sambuco-Salicion capreae Sorbus aucuparia E3a . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . . . . . . . 1 2 Sorbus aucuparia E2b + + 1 1 1 . . . . + + . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 . + 9 21 Sorbus aucuparia E2a . + . . . . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . . . . . . . 2 5 Sorbus aucuparia E1 + . . + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . 3 7 Salix caprea E3a . . . . . . . . . . . r . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 2 RP Rhamno-Prunetea Crataegus monogyna E2 . . . . . . . . . . . . . . . . . . . . . . . . . . . + . + . . . . . . . . . . . . 2 5 Rosa canina E2a . + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 2 Viburnum lantana E2a . . . . . . . + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 2 TG Trifolio-Geranietea Campanula rapunculoides E1 1 . . . . . . . . . . . . . + + . + 1 . . . + 1 + 1 + + 1 1 . . . + + + . + . . . . 17 40 Digitalis grandiflora E1 + . . . . . + + . . . . + . + 1 . . . . . + . . + . + . + . . . . . . . + . . . . 11 26 Vicia sylvatica E1 . + . + + . . . . . + . . . . . . . . . . . . . . . . + . . . . . . . . . . + . . . 6 14 Vincetoxicum hirundinaria E1 . . . . . . . . . . . . . . . . + . . . . . . . + + . . . + . . . . . . . + . . . . 5 12 Achillea distans E1 . . . . . . . + . . . . . . . . . . . + . . . . . . . . . . . . . . . . . + . . . . 3 7 Verbascum lanatum E1 . . . . . . . + + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 2 5 Polygonatum odoratum E1 + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 2 Lilium carniolicum E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . 1 2 EA Epilobietea angustifolii Fragaria vesca E1 + . + . . + 1 + . . + + + . + 1 + . + + . . + . + + + . + . . . . . . . + . . . . 19 45 Rubus idaeus E2a . . . . . . . . . . . + . . . . . + . . . + . . + . . . . . . . + . . . . . . + . . 6 14 Hypericum hirsutum E1 . . . . . . . . . . . + . . . . + + . . . + . . . . + . . . . . . . . . . . . . . . 5 12 Tussilago farfara E1 . . . . . . . + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . 2 5 Galeopsis speciosa E1 . . . . . . . . . . . + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 2 Bromopsis benekenii E1 . . . . . . . . . . . . + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 2 Solanum dulcamara E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . 1 2 BA Betulo-Alnetea viridis Salix appendiculata E2b . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . 1 2 MuA Mulgedio-Aconitetea Phyteuma ovatum E1 1 1 1 1 + 1 1 + 1 + 1 1 + + 1 1 + + + 1 1 1 1 1 1 1 1 1 + 1 + 1 1 1 1 + + 1 1 . 1 1 41 98 Senecio ovatus E1 . + . . + + + + 1 . . + + 1 + . 1 + + + . + . . + . + + 1 + . 1 + 1 1 1 1 + + + 1 + 31 74 Athyrium filix-femina E1 . . . . + + + . . + . 1 + 1 + . . + . + . + . . + + + + + + + + + + 1 + 1 . + + + + 28 67 Aconitum lycoctonum s.lat. E1 . . . . . + + + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 3 7 Polygonatum verticillatum E1 . . . . . + + . . . . . . . . . . . . . . + . . . . . . . . . . . . . . . . . . . . 3 7 Silene dioica E1 . . . . . . . . + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 2 Ranunculus platanifolius E1 . . + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 2 Doronicum austriacum E1 . . . . . . . . . . . . . 1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 2 ES Elyno-Seslerietea Sesleria caerulea subsp. calcaria E1 2 3 + 2 . + . + . . + . . . . . . + . . . + . . . . . . . . . . . . . . . . . + . 2 11 26 Betonica alopecuros E1 . . . 1 . . + + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 3 7 Laserpitium peucedanoides E1 . . . + . + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + 3 7 Aster bellidiastrum E1 . . . + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 2 Centaurea haynaldii subsp. julica E1 . . . . . . . + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 2 FB Festuco-Brometea Pimpinella saxifraga E1 . . . . . . . . . . . . . . . + 1 . . . . . + . . . . + . . . . . . . . . . . . . . 4 10 Allium carinatum subsp. pulchellum E1 . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . . . . . . . + . . . . 2 5 Festuca valesiaca agg. E1 . . . . . . + . . . . . . . . . . . + . . . . . . . . . . . . . . . . . . . . . . . 2 5 Carlina acaulis E1 . . . . . . . + . . . . . . . . . . . . + . . . . . . . . . . . . . . . . . . . . . 2 5 Brachypodium rupestre E1 . . . + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 2 MA Molinio-Arrhenatheretea Angelica sylvestris E1 . . . . . . . + . . . . . . + . . + . . . . . . . + . . + + + . . . . . . . . . . . 7 17 Galium album E1 . . . . . . . + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 2 Lotus corniculatus E1 . . . . . . . + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 2 JT Juncetea trifidi Selaginella helvetica E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . . 1 2 TR Thlaspietea rotundifolii Adenostyles glabra E1 . . . . . 1 + + + . . + . + + + + 1 . 1 + 2 . . + + + . . . . . + 1 + + + . + 1 1 1 25 60 Gymnocarpium robertianum E1 . . + + . . . . . . . . . . + . 1 + . + + + + + 1 . . . + . . . . . . . . + + + + 16 38 Hieracium bifidum E1 + . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . 2 5 Astrantia carniolica E1 . . . . . . + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 2 Campanula cespitosa E1 . . . . . . . + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 2 AT Asplenieta trichomanis Asplenium viride E1 + + + 1 + + + + . + + + + + + 1 + 1 + + + 1 + 1 + 1 . + . + . . . . + + . . . + 1 1 32 76 Moehringia muscosa E1 + . + . + . . . . + . . + . + + + + + . + + + + + + . + . + . . . . . . + . . + . + 21 50 Asplenium trichomanes E1 + + . . . + . . + . . . . + + + + + . . + + + + . + + + + . . . + + . . + . . . . 20 48 Polypodium vulgare E1 + + . . + . . . . . . . . . + . + . . + . + + . + + . + + + + . . + + + + . . . + . 19 45 Asplenium ruta-muraria E1 + . + . . . . + . . . . + . + + . + . . . . . . . + + . + . . . . . . . . . . + . 11 26 Cystopteris fragilis E1 . . . . . . + . . . . . . . . . . + . . . + . . . . + . . . . . . . . . . . . + + + 7 17 Saxifraga crustata E1 + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . + 3 7 Campanula carnica E1 + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + 2 5 Carex brachystachys E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + + 2 5 Saxifraga hostii E1 + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 2 Epilobium collinum E1 . . . . . . . . . . . . + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 2 Asplenium adiantum-nigrum E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . 1 2 ML Mosses and lichens (Mahovi in lišaji) Polytrichum formosum E0 + 1 1 + . . + + + + + + + + + 1 1 + + 1 + . 1 + + 1 + 2 1 1 2 1 1 1 1 1 1 1 1 + + + 39 93 Ctenidium molluscum E0 + 2 . 1 . + 1 + . 2 2 + 2 . 1 1 1 2 2 2 1 1 2 2 2 . 1 1 1 1 1 1 + 1 2 1 2 . 1 2 2 2 36 86 Neckera crispa E0 + 1 1 1 + . . + . + . . + + + + + 1 + + + 1 1 2 . 1 + 1 1 1 + 1 1 1 + + + + 1 2 1 35 83 Tortella tortuosa E0 + 1 . + 1 + + + . + 1 + + . + + . . 1 2 + 1 . . + 1 . . + + 1 1 1 1 2 1 + . . 1 + + 31 74 Plagiothecium denticulatum E0 . . + . 1 1 + . + + + + 1 + 1 . 1 + + . . . . + + + + 1 1 1 . . . 1 1 + + + + + + 1 30 71 Fissidens dubius E0 + + + 1 1 + . . + + 1 + . . + + . + + . + + . + + . + + + . + + 1 1 + 1 + + 1 . . 30 71 Metzgeria furcata E0 . . . 2 . + . + . + 1 + + . + 1 2 . + . . 1 1 + . . + 1 + 1 + . + . + 1 + + . . + 1 26 62 Hypnum cupressiforme E0 . + + . 1 . . . . . . . . . 1 + 1 1 1 . 1 + . 1 . 1 + . + . . + + + 1 2 2 + 1 + + 1 25 60 Peltigera canina E0 + . + + + . . . . . . . + . + + + + + + + + . + + . . + . . . . . + + + + . . + + + 23 54 Isothecium alopecuroides E0 . 1 + + + + + . . . . . + . 1 1 1 2 . . + 1 . + 1 . . 1 + . . . . + 1 + + . . . . 1 22 52 Schistidium apocarpum E0 . . . . . . + + + . + + + . + 1 . + . . . . . + + . . + + . . + + . . . + . + 1 1 + 20 48 Dicranum scoparium E0 + + 2 1 1 . + + . + + . + . . . . . + + . . . . . + . . . . . . . . . + + . + . + + 18 43 Plagiochila porelloides E0 . . . . . . + + . 1 . . . . + + + 1 + . . . . . + + . . + . . . . 1 1 . 1 + 1 1 + . 18 43 Thuidium tamariscinum E0 . + . + + . . . . . . . . . . + 1 + 1 . . . + . . . . + . + 1 . . + + + 1 . . . + + 17 40 Mnium thomsonii E0 . . . + + . . . . . + . + + + . + + + . . . . + . . . + . . . . . . . . + 1 . 1 + + 16 38 Atrichum undulatum E0 . . . . . . . . . . + 1 + 1 . . + . . . . . . . . + + . + . + 2 1 . + . 1 + . . . . 14 33 Cladonia furcata E0 . . . . . . . . . . . . . . . . + . + 1 + . 1 + . + . . + + 1 . + . . . + . . . + + 14 33 Bartramia halleriana E0 . . . . . . . . . . . . . . + + + . + . . . + + . + . + + . 1 + . + + + . . . . . . 14 33 Hylocomium splendens E0 . . + + . . . . . . . . . . . . . . + . . + . + . + . + . . . . . . . . + . . + + + 11 26 Homalothecium lutescens E0 + . . + . + . . . . . . . . . + . + . . . . . . + . . + . . . . . . . . . . . + + + 10 24 Dicranella heteromalla E0 . . . + . . + . . . + 1 . + . . . . . . . . . . . . + . + . 1 . + . . . + . . . . . 10 24 Collema cristatum E0 . . . . . . . . . . . . . . . + . . + . . . . . . . . . + + . . . . . . + + + + + + 10 24 Plagiothecium nemorale E0 . + + . 1 . . . . + . . . . + . . . . . . . 1 . . + . . . . . . . 1 + . . . . . . . 9 21 Mnium marginatum E0 . . + . . 1 . . . 1 . . . . + 1 . 1 . . . . . . . . . . . . . . . . . . . . . 1 . . 7 17 Eurhynchium angustirete E0 . . + . . . . . . . . . . . . . + . + . . . . . . + . . . + . . . . . . + + . . . . 7 17 Brachythecium velutinum E0 . . . . . + . . + . . + . . + . + . . . . . . . . . . . + . . . . . . . . . . + . . 7 17 Rhizomnium punctatum E0 . . . . . . . . . . . . . . . . + . . . + . . . . + . . . . . . . . . 1 . . + + + . 7 17 Plagiomnium undulatum E0 . + . . + . . . . . . . . . . . . . . . . . . . . + . . . . . . . . . + . . + + . . 6 14 Cladonia rangiferina E0 . . + . . . . . . . . . . . . . + . + . . . . + . . . . . . + . . . . . + . . . . . 6 14 Bryum capillare E0 . 1 . . + . . . . . . . . . + . . . . . . . . . . . . . . . . . . . . . + + . . . . 5 12 Rhytidiadelphus triquetrus E0 . . + + . . . . . . . . . . . . . . . . . + . . . + . . . . . . . . . + . . . . . . 5 12 Plagiomnium affine E0 . . . + . . . . . . . . . . . . . . . + . . . . . . + . . . . . . . + . . . . + . . 5 12 Porella platyphylla E0 . . . . . . . . . . . . . + + + . + . . . . . . . . . . . . . . . . . . . . . . + . 5 12 Isothecium myosuroides E0 . . . . . . . . . . . . . . . . . . . . . . . + . . . . + . . . + . . . . . . . + + 5 12 Homalothecium philippeanum E0 + . . . . . . . . . . . . . + . . . . . . . . . . . . . . . . . . . . . . . . . + + 4 10 Peltigera leucophlebia E0 . + + + . . . . . . + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 4 10 Brachythecium rutabulum E0 . . . . . + . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . + . . . . + 4 10 Cladonia pyxidata E0 . . . . . . + + . . . . + . . . . . . . . . . . + . . . . . . . . . . . . . . . . . 4 10 Fissidens taxifolius E0 . . . . . . . . . . . + + + . . . . . . . . . . . . . . . . + . . . . . . . . . . . 4 10 Conocephalum conicum E0 . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . . . . . + . . . . 1 + . 4 10 Hypnum cupressiforme var. filiforme E0 . + . . . . . . . . . . . . . . . . . . . . . . . . . . + + . . . . . . . . . . . . 3 7 Cladonia sp. E0 . . + . . . . . . . . . . . . + . . . . . . . . . . . . . . . . . . . . . . . + . . 3 7 Leucobryum glaucum E0 . . . . . . + . . . . . + . . . . . . . . . . . . . . . . . + . . . . . . . . . . . 3 7 Thamnobryum alopecurum E0 . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . + . . . + . . . . . . 3 7 Radula complanata E0 . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . + . . . . . . . . . . . 2 5 Scapania nemorea E0 . . . . . . . . . . . . . . . . . . . + . . . . . . . . + . . . . . . . . . . . . . 2 5 Homalothecium sericeum E0 . . . . . . . . . . . . . . . . . . . . . . . . + . . . . + . . . . . . . . . . . . 2 5 Bryum argenteum E0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . + 2 5 Bryum sp. E0 + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 2 Dicranum majus E0 . . . . + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 2 Paraleucobryum sauteri E0 . . . . . . . + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 2 Riccardia sp. E0 . . . . . . . . . . + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 2 Platygyrium repens E0 . . . . . . . . . . . + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 2 Neckera complanata E0 . . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . . . . . . . . . . . 1 2 Brachythecium sp. E0 . . . . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . . . . . . . . . 1 2 Scleropodium purum E0 . . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . . . . . 1 2 Anomodon attenuatus E0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . 1 2 Porella arboris-vitae E0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . 1 2 Orthotrichum anomalum E0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . 1 2 Brachythecium oxycladum E0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . 1 2 Legend - Legenda AR Limestone with chert - apnenec z rožencem AGR Limestone with chert and claystone - apnenec z rožencem in glinavcem DR Dolomite with chert - dolomit z rožencem DRG Dolomite with chert and claystone - dolomit z rožencem in glinavcem AL Limestone with marlstone - apnenec z laporovcem ALR Limestone with chert and marlstone AGP Limestone with claystone and sandstone - apnenec z glinavcem in peščenjakom Re Rendzina - rendzina Rj Brown calcareous soil - rjava pokarbonatna tla Dy Dystric brown soil - distrična rjava tla Number of relevé (Zaporedna številka popisa) 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 31 32 33 34 35 36 37 38 39 40 41 42 Pr. Fr. Number of relevé (Zaporedna številka popisa) 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 31 32 33 34 35 36 37 38 39 40 41 42 Pr. Fr. Number of relevé (Zaporedna številka popisa) 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 31 32 33 34 35 36 37 38 39 40 41 42 Pr. Fr. Number of relevé (Zaporedna številka popisa) 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 31 32 33 34 35 36 37 38 39 40 41 42 Pr. Fr. Number of relevé (Zaporedna številka popisa) 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 31 32 33 34 35 36 37 38 39 40 41 42 Pr. Fr. Number of relevé (Zaporedna številka popisa) 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 31 32 33 34 35 36 37 38 39 40 41 42 Pr. Fr. Number of relevé (Zaporedna številka popisa) 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 31 32 33 34 35 36 37 38 39 40 41 42 Pr. Fr. Table 10 (Preglednica 10): Saxifrago cuneifolii-Fagetum gymnocarpietosum dryopteridis Number of relevé (Zaporedna številka popisa) 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 Database number of relevé (Delovna številka popisa) 254399 255533 255534 255532 255536 255537 255447 255448 254400 255339 255541 255491 255328 255450 255464 Elevation in m (Nadmorska višina v m) 650 710 680 540 850 960 660 660 710 830 970 580 870 520 570 Aspect (Lega) NE NE N NE NE N N NE NNE NE NE NE SE N NE Slope in degrees (Nagib v stopinjah) 35 35 35 30 30 30 35 35 40 40 35 35 40 40 30 Parent material (Matična podlaga) AR R AR AR AR AR DR DR ALR AGR ALR ALR AGR AGR AR Soil (Tla) Ra Dy Re Re Rj Rj Re Re Dy Re Re Re Re Re Re Stoniness in % (Kamnitost v %) 70 80 80 80 70 60 60 20 50 10 40 60 20 30 80 Cover in % (Zastiranje v %): Upper tree layer (Zgornja drevesna plast) E3b 90 100 100 100 100 100 90 95 80 80 90 90 90 95 100 Lower tree layer (Spodnja drevesna plast) E3a 10 5 5 10 Shrub layer (Grmovna plast) E2 5 1 5 1 5 3 5 5 5 5 5 5 20 30 5 Herb layer (Zeliščna plast) E1 15 15 15 30 30 30 40 70 50 50 50 50 60 50 40 Moss layer (Mahovna plast) E0 40 20 10 10 10 5 20 10 30 5 20 20 30 50 30 Maximum diameter of trees (Največji prsni premer dreves) cm 40 30 30 30 35 35 35 35 40 40 45 50 30 30 35 Maximum height of trees (Največja drevesna višina) m 18 20 18 20 22 18 22 20 22 22 19 28 16 16 20 Number of species (Število vrst) 58 49 46 44 51 36 61 44 60 66 58 66 83 75 72 Relevé area (Velikost popisne ploskve) m2 400 400 400 400 400 400 400 400 400 400 400 400 400 400 400 Date of taking relevé (Datum popisa) 9/19/2014 5/2/1987 5/2/1987 5/2/1987 5/2/1987 5/2/1987 5/9/1987 5/9/1987 9/19/2014 8/18/1989 5/31/2002 7/12/1991 8/17/1989 5/9/1987 7/13/1987 Locality (Nahajališče) Mohor-Temnak Mohor-Temnak Mohor-Temnak Mohor-Temnak Mohor-Temnak Mohor-Temnak Koritnica - Jelenca Koritnica - Jelenca Mohor-Temnak Kacenpoh Grant-Luken Grant - Čumik Kacenpoh - Fergajnar Koritnica - Jelenca Grant - Čumik Quadrant (Kvadrant) 9848/2 9848/2 9848/2 9848/2 9848/2 9848/2 9849/1 9849/1 9848/2 9749/4 9749/3 9849/1 9749/4 9849/1 9849/1 Coordinate GK Y (D-48) m 409083 409083 408792 409279 408705 408549 414910 415268 408917 419966 412945 414724 420022 415052 414520 Coordinate GK X (D-48) m 5117500 5117496 5117482 5117587 5117248 5117118 5116586 5116283 5117461 5120916 5118127 5117600 5121008 5116996 5117162 Diagnostic species of the association (Diagnostične vrste asociacije) Pr. Fr. VP Calamagrostis arundinacea E1 + 1 1 1 . + + 1 3 1 + + 3 1 + 14 93 VP Luzula luzuloides E1 + 1 1 + 1 1 + 1 1 1 1 + + + . 14 93 VP Saxifraga cuneifolia E1 + + 1 + + 1 . . . + 1 + + + 1 12 80 VP Veronica urticifolia E1 + + + 1 + . 1 + . + 1 . + + + 12 80 VP Huperzia selago E1 + + . . . . + . . . + . + . . 5 33 QP Tanacetum corymbosum E1 . . . . . + . . . . . . + . . 2 13 TG Clinopodium vulgare E1 . . . . . . . . . . . . + . . 1 7 QF Festuca heterophylla E1 . . . . . . . . . . . . + . . 1 7 ES Campanula witasekiana E1 . . . . . . . . . . . . + . . 1 7 Geographical differential species (Geografske razlikovalne vrste) AF Anemone trifolia E1 r 1 1 . 1 . 1 . + . . . + . + 8 53 QP Sesleria autumnalis E1 r + + . + + . . . . . + . + + 8 53 FS Lathyrus vernus subsp. flaccidus E1 . . . . + . . . . . 1 . . + + 4 27 VP Larix decidua E3b . . . . . . . 1 . . . . . . . 1 7 Differential species of the subassociations (Razlikovalne vrste subasociacij) TA Acer pseudoplatanus E3b . + . . . . . + . . . + . . . 3 20 TA Acer pseudoplatanus E3a . . . . + + + . r . . . . . . 4 27 TA Acer pseudoplatanus E2b . . + . . . + . . . . . . + . 3 20 TA Acer pseudoplatanus E1 + + + + + + . + + . + + . . . 10 67 MuA Phyteuma ovatum E1 + + + + . + + 1 . + 1 + . . . 10 67 FS Actaea spicata E1 . . + 1 + + 1 1 + + + + . . . 10 67 VP Gymnocarpium dryopteris E1 + + + + . . 1 + 1 + + + . . . 10 67 TA Polystichum aculeatum E1 r . . . + + 1 + . + + + . . + 9 60 VP Abies alba E3a r . . . . . . . . . . . . . . 1 7 VP Abies alba E2b r + + + + + . . + . . . . . . 7 47 TA Dryopteris affinis E1 . . . . . + . . 1 + . . . . . 3 20 TA Adoxa moschatellina E1 . . . . . . . . + + + . . . . 3 20 EP Carex alba E1 . . . . . . 1 2 . . . . . 2 2 4 27 QP Cornus mas E2b . . . . . . . . . . . . . + + 2 13 AF Aremonio-Fagion Cardamine trifolia E1 1 1 1 1 1 1 + . 1 + + 1 . 1 1 13 87 Cyclamen purpurascens E1 1 1 . 1 1 1 1 1 1 + . + 1 1 1 13 87 Cardamine enneaphyllos E1 . . + 1 2 2 1 2 3 2 1 2 . . . 10 67 Lamium orvala E1 . . . . + . . . + 1 . 1 . . . 4 27 Aremonia agrimonoides E1 . . . . . . . . . . . . + . . 1 7 Knautia drymeia E1 . . . . . . . . . . . . + . . 1 7 EC Erythronio-Carpinion Primula vulgaris E1 . . . + . . + 1 . . . + + + + 7 47 TA Tilo-Acerion Aruncus dioicus E1 + + . . + . . . + + . . . . . 5 33 Phyllitis scolopendrium E1 . . . . . + . . + . . . . + . 3 20 Polystichum braunii E1 . . + . . . . . 1 . . . . . + 3 20 Acer pseudoplatanus E2a . . . . + . . . . . . + . . . 2 13 Polystichum setiferum E1 . . . . . . . . . 1 . . . . + 2 13 Euonymus latifolia E2 . . . . . . + . . . . . + . . 2 13 Acer platanoides E2a . . . . . . . . . . . . . + . 1 7 Acer platanoides E1 . . . . . . + . . . . . . . . 1 7 Ulmus glabra E3b . . . . . . . . . . . . . . + 1 7 Ulmus glabra E3a . . . . . . + . . . . . . . . 1 7 Ulmus glabra E2b . . . . . . . . . . . + . . . 1 7 Stellaria montana E1 . . . . . . . . + . . . . . . 1 7 Lunaria rediviva E1 . . . . . . . . . . . + . . . 1 7 Tephroseris pseudocrispa E1 . . . . . . . . . . . . . + . 1 7 Tilia platyphyllos E2a . . . . . . . . . . . . . + . 1 7 FS Fagetalia sylvaticae Fagus sylvatica E3b 5 5 5 4 5 5 5 4 4 5 5 5 5 4 4 15 100 Fagus sylvatica E3a 1 . . . . . . . + . + 1 . . . 4 27 Fagus sylvatica E2b 1 + + + + + + 1 + . + 1 . 1 . 12 80 Fagus sylvatica E2a + . . . . . . . + . + + . . . 4 27 Fagus sylvatica E1 1 . . + + . . + 1 . 1 . + . . 7 47 Mercurialis perennis E1 + 1 . 1 1 1 1 1 1 1 + + + . + 13 87 Galium laevigatum E1 + 1 + 1 . + 1 1 . . 1 + 1 1 + 12 80 Dryopteris filix-mas E1 + + . 1 + + + 1 . 1 1 1 . + + 12 80 Daphne mezereum E2a . + + + + + 1 + . . + . + 1 . 10 67 Prenanthes purpurea E1 1 + + . . . 1 . 1 + 1 + + + . 10 67 Asarum europaeum subsp. caucasicum E1 + 1 1 1 . . 1 . + . . + . + 1 9 60 Lonicera alpigena E2a 1 1 . . . . 1 1 . + + . + 1 + 9 60 Salvia glutinosa E1 . . . + + . . . + 1 . . + + . 6 40 Galeobdolon flavidum E1 . . . . + + . . 1 + + . . . 1 6 40 Pulmonaria officinalis E1 . . . + . . + . . + . . . + + 5 33 Symphytum tuberosum E1 . . . + . . . . . + . + . 1 + 5 33 Mycelis muralis E1 . . . . . . . + + . + + . . + 5 33 Petasites albus E1 + + . . . . . . . + . . . + . 4 27 Polygonatum multiflorum E1 . . + + . . . + . . + . . . . 4 27 Lilium martagon E1 . . + . . + + . . + . . . . . 4 27 Paris quadrifolia E1 . . + . . + . . . + + . . . . 4 27 Lathyrus vernus subsp. vernus E1 . . . . . + . . . . + + . + . 4 27 Festuca altissima E1 . . . . . . . . + + + . . . . 3 20 Galium odoratum E1 . . . . . . . . 1 + 1 . . . . 3 20 Sambucus nigra E2 . . . . . . . . + + . + . . . 3 20 Scrophularia nodosa E1 . . . . . . . . + + . + . . . 3 20 Campanula trachelium E1 . . . . . . . . . + + . + . . 3 20 Laburnum alpinum E3b r . . . . . . . . . . . + . . 2 13 Laburnum alpinum E3a . . . . . + . . . . . . + . . 2 13 Laburnum alpinum E2b . . . . . . . . . + . . . . . 1 7 Laburnum alpinum E1 . . . . . . + . . . . . . . . 1 7 Carpinus betulus E3b . . . r . . . . . . . + . . . 2 13 Carpinus betulus E3a . . + . . . . . . . . . . . . 1 7 Carpinus betulus E2b . . . . . . + . . . . . . . . 1 7 Carpinus betulus E2a . . . + . . . . . . . . . . . 1 7 Viola reichenbachiana E1 . . . . . . + . . . . + . . . 2 13 Sanicula europaea E1 . . . . . . . . . + . + . . . 2 13 Euphorbia amygdaloides E1 . . . . . . . . . . . + + . . 2 13 Prunus avium E3b . . . . + . . . . . . . . . . 1 7 Prunus avium E2a . . . . . . . . . . . + . . . 1 7 Tilia cordata E3a . . . . + . . . . . . . . . . 1 7 Circaea lutetiana E1 . . . . . . . . . + . . . . . 1 7 Dentaria bulbifera E1 . . . . . . . . . + . . . . . 1 7 Epilobium montanum E1 . . . . . . . . . + . . . . . 1 7 Neottia nidus-avis E1 . . . . . . . . . . + . . . . 1 7 Heracleum sphondylium E1 . . . . . . . . . . . + . . . 1 7 Epipactis helleborine E1 . . . . . . . . . . . . + . . 1 7 Poa nemoralis E1 . . . . . . . . . . . . + . . 1 7 Melica nutans E1 . . . . . . . . . . . . . . + 1 7 QP Quercetalia pubescenti-petraeae Ostrya carpinifolia E3b 1 . . . . . . . + . . + . 3 2 5 33 Ostrya carpinifolia E3a + + + 2 + + 1 1 . + . . . . . 9 60 Ostrya carpinifolia E2a . . . . . . . . + . . . . . . 1 7 Ostrya carpinifolia E1 . + . . . . + . . . . . . . . 2 13 Sorbus aria E3b + . . . . . . . . . . . . 1 . 2 13 Sorbus aria E3a + + + + + . . + . . . . + . . 7 47 Sorbus aria E2b . . . . . . . + . . . . . . + 2 13 Sorbus aria E2a . . . . . . . . . . . . 1 1 . 2 13 Fraxinus ornus E3b . . . . . . . . . . . . . 1 . 1 7 Fraxinus ornus E3a r . + . . . . . . + . . . . + 4 27 Fraxinus ornus E2b . . . . . . . . . . . . . . 1 1 7 Fraxinus ornus E2a . . . . . . . . . . . . 1 + . 2 13 Fraxinus ornus E1 . . . . . . . . . . . + . . . 1 7 Melittis melissophyllum E1 . . . . . . + . . . . . + . + 3 20 Arabis turrita E1 . . . . + . . . . . . + . . . 2 13 Hypericum montanum E1 . . . . . . . . . . . . + + . 2 13 Euonymus verrucosa E2a . . . . . . . . . . . . . + . 1 7 QR Quercetalia roboris Rubus hirtus E2a + + . . . . . . . + . . + . . 4 27 Populus tremula E3b r . + . . . . . + . . . . . . 3 20 Populus tremula E1 . . . . . . . . + . . . . . . 1 7 Hieracium racemosum E1 + . . . . . . . . . . . 1 . . 2 13 Hieracium sabaudum E1 . . . . . . . . . . . . + . . 1 7 QF Querco-Fagetea Carex digitata E1 r + + + + + + . . . . . + + 1 10 67 Anemone nemorosa E1 + + . . + . . 2 . . + + . . 1 7 47 Corylus avellana E2a . . . . + . . . . . . + + . . 3 20 Hedera helix E3a . . . . . . . . . . . . . . 1 1 7 Hedera helix E1 . . . . . . + . . . . + . . + 3 20 Moehringia trinervia E1 . . . . + . . . r . . + . . . 3 20 Vinca minor E1 . . . . . . . . . . . + . 1 + 3 20 Acer campestre E3b . . . . . . . . . . . . . . 1 1 7 Acer campestre E2a . . . . . . . . . . . + . . + 2 13 Acer campestre E1 . . . . . . + . . . . . . . + 2 13 Clematis vitalba E2a . . . . . . . . . + . + . . . 2 13 Rosa arvensis E2a . . . . . . . . . . . . + . + 2 13 Hepatica nobilis E1 . . . . . . . . . . . . . 1 1 2 13 Lonicera xylosteum E2a . . . . . . . . . . . . . 1 + 2 13 Aegopodium podagraria E1 . . . . . . . . . . . + . . . 1 7 Cephalanthera longifolia E1 . . . . . . . . . . . . + . . 1 7 Viola riviniana E1 . . . . . . . . . . . . + . . 1 7 Viburnum opulus E2a . . . . . . . . . . . . . + . 1 7 EP Erico-Pinetea Erica carnea E1 r . . . . . . . . . . . + + . 3 20 Cirsium erisithales E1 . . . + . . + . . . . . + . . 3 20 Rubus saxatilis E1 . . . . . . + . . . . . . + . 2 13 Calamagrostis varia E1 . . . . . . . + . . . . + . . 2 13 Molinia caerulea subsp. arundinacea E1 + . . . . . . . . . . . . . . 1 7 Buphthalmum salicifolium E1 . . . . . . . . . . . . + . . 1 7 Polygala chamaebuxus E1 . . . . . . . . . . . . + . . 1 7 VP Vaccinio-Piceetea Oxalis acetosella E1 + 1 1 + 1 1 + 1 + 1 1 + . 1 1 14 93 Solidago virgaurea E1 + + + . . . . . . + + . 1 + + 8 53 Phegopteris connectilis E1 + + 1 . . . + . 1 + . . + . . 7 47 Picea abies E3b . . . . . . . 1 . . . . . + + 3 20 Picea abies E3a . . . . . . . . . . . . + . . 1 7 Picea abies E2b . . . . . + + + . + . r . + . 6 40 Picea abies E2a . . . . . . . . . . . . + . . 1 7 Picea abies E1 + . . . + . . + . . . . . . . 3 20 Rosa pendulina E2a + + . . . . + 1 . . . . . 2 . 5 33 Gentiana asclepiadea E1 . . . . . . + + . + . . + . + 5 33 Homogyne sylvestris E1 . . . . . . 1 1 . . + . . 2 + 5 33 Hieracium murorum E1 + . + . . . . . . . . . 1 . . 3 20 Dryopteris dilatata E1 . . . . + . . . + . + . . . . 3 20 Valeriana tripteris E1 . . . . . . . . r . . . + 1 . 3 20 Aposeris foetida E1 . . . . . . . . . . . + . . + 2 13 Thelypteris limbosperma E1 . . . . . . . . + . . . . . . 1 7 Maianthemum bifolium E1 . . . . . . . . . . + . . . . 1 7 Vaccinium myrtillus E1 . . . . . . . . . . . . 2 . . 1 7 Dryopteris expansa E1 . . . . . . . . . . . . . + . 1 7 SSC Sambuco-Salicion capreae Sorbus aucuparia E2a . . . . . . + . . . . . . . 1 7 Sorbus aucuparia E1 r + + . + . . + . . . . + . . 6 40 RP Rhamno-Prunetea Crataegus monogyna E2 . . . . . . . . . . . . . + . 1 7 Berberis vulgaris E2a . . . . . . . . . . . . . . + 1 7 TG Trifolio-Geranietea Campanula rapunculoides E1 . . . + . . . . . . . + . . + 3 20 Digitalis grandiflora E1 . . . . . . . . . + . . + . . 2 13 Verbascum lanatum E1 . . . . . . . + . . . . . . . 1 7 Achillea distans E1 . . . . . . . . . . . . + . . 1 7 Lilium carniolicum E1 . . . . . . . . . . . . . . + 1 7 EA Epilobietea angustifolii Rubus idaeus E2a + + . . . . . . + . . . . . . 3 20 Urtica dioica E1 . . + . . . . . r . . . . . . 2 13 Hypericum hirsutum E1 . . . . . . + . . . . . . . . 1 7 Galeopsis speciosa E1 . . . . . . . . . 1 . . . . . 1 7 Stachys alpina E1 . . . . . . . . . + . . . . . 1 7 Fragaria vesca E1 . . . . . . . . . . . . + . . 1 7 MuA Mulgedio-Aconitetea Senecio ovatus E1 + + + + + . 1 + + 1 1 + . + + 13 87 Athyrium filix-femina E1 + . + 1 + . 1 1 1 1 . + + . + 11 73 ES Elyno-Seslerietea Sesleria caerulea subsp. calcaria E1 + + . + . . . . . . . . + . + 5 33 Betonica alopecuros E1 . . . . . . . . . . . . + . . 1 7 Phyteuma orbiculare E1 . . . . . . . . . . . . + . . 1 7 FB Festuco-Brometea Brachypodium rupestre E1 . . . . . . . . . . . . + . . 1 7 Carlina acaulis E1 . . . . . . . . . . . . + . . 1 7 Dianthus monspessulanus E1 . . . . . . . . . . . . + . . 1 7 Festuca valesiaca agg. E1 . . . . . . . . . . . . + . . 1 7 Pimpinella saxifraga E1 . . . . . . . . . . . . . . + 1 7 MA Molinio-Arrhenatheretea Angelica sylvestris E1 . . . . . . . . . . . . . + . 1 7 TR Thlaspietea rotundifolii Adenostyles glabra E1 . . . . + . . . . 1 1 + . . . 4 27 Gymnocarpium robertianum E1 . . . . . . + + . . . . . + + 4 27 Cerastium subtriflorum E1 . . . . . + . . . . . . . . . 1 7 Campanula cespitosa E1 + 1 7 AT Asplenieta trichomanis Asplenium viride E1 + + + + + + . + + + + + . 1 + 13 87 Polypodium vulgare E1 + + + . . . . . + . + + + + + 9 60 Asplenium trichomanes E1 . . . + + . . . + + + + + . + 8 53 Asplenium ruta-muraria E1 . . . . + . . . . . . . + . + 3 20 Moehringia muscosa E1 . . . . . . . . . . . . + + + 3 20 Cystopteris fragilis E1 . . . . . . . . . + + . . . . 2 13 Epilobium collinum E1 . . . . . . . . . . . . + . . 1 7 ML Mosses and lichens (Mahovi in lišaji) Polytrichum formosum E0 2 2 1 + 1 + 1 + 2 . 2 . 2 1 + 13 87 Neckera crispa E0 + 1 + 2 + + + . + + + . + 1 2 13 87 Isothecium alopecuroides E0 + + . 1 + + + + + + + 2 + . 1 13 87 Ctenidium molluscum E0 . + + 1 + + 1 1 + . + 1 + 1 2 13 87 Hypnum cupressiforme E0 1 + . + . . + + + + . 1 1 1 + 11 73 Plagiothecium denticulatum E0 . + . 1 1 . + + + . + . + 1 + 10 67 Atrichum undulatum E0 . 1 1 + 1 . + . + + + . + . . 9 60 Fissidens dubius E0 . . + + + + + . + . . + . 1 1 9 60 Thuidium tamariscinum E0 2 . . . . . + . 1 . . . . 1 + 5 33 Metzgeria furcata E0 . . . . . . + . . + + . + 1 . 5 33 Leucobryum glaucum E0 1 . 1 . . . . . . . . . + + . 4 27 Dicranum scoparium E0 1 + . . . . . . + . . . . 1 . 4 27 Tortella tortuosa E0 + . . . . . . . + . . . + 1 . 4 27 Dicranella heteromalla E0 . + 1 . 1 . . . . . . . + . . 4 27 Plagiochila porelloides E0 . . . . + . + + . . . . . 2 . 4 27 Bryum capillare E0 . . . . + . + + + . . . . . . 4 27 Peltigera canina E0 . . . . . . . . + . + + . + . 4 27 Mnium thomsonii E0 . . . . . . . . . + + . . + + 4 27 Fissidens taxifolius E0 + + . . . . . . . . . . . + . 3 20 Bartramia halleriana E0 . . . + . . . + . . . . . 1 . 3 20 Eurhynchium angustirete E0 . . . . + . . . . . . . . 1 + 3 20 Thamnobryum alopecurum E0 . . . . . . + . . + + . . . . 3 20 Conocephalum conicum E0 . . . . . . . . + + + . . . . 3 20 Porella platyphylla E0 . . . . . . . . + . . + + . . 3 20 Cladonia rangiferina E0 1 . + . . . . . . . . . . . . 2 13 Cladonia furcata E0 . + . . . . . . . . . . . + . 2 13 Hylocomium splendens E0 . . . . . . . . + . . . . 1 . 2 13 Plagiothecium nemorale E0 . . . . . . . . . + . . . . + 2 13 Brachythecium velutinum E0 . . . . . . . . . + + . . . . 2 13 Anomodon attenuatus E0 . . . . . . . . . . + + . . . 2 13 Schistidium apocarpum E0 . . . . . . . . . . . + + . . 2 13 Homalothecium lutescens E0 . . . . . . . . . . . . + . + 2 13 Bazzania trilobata E0 + . . . . . . . . . . . . . . 1 7 Marchantia polymorpha E0 . . . . . . . . 1 . . . . . . 1 7 Homalothecium philippeanum E0 . . . . . . . . . . + . . . . 1 7 Mnium sp. E0 . . . . . . . . . . + . . . . 1 7 Anomodon viticulosus E0 . . . . . . . . . . + . . . . 1 7 Hypnum cupressiforme var. filiforme E0 . . . . . . . . . . . 1 . . . 1 7 Brachythecium oxycladum E0 . . . . . . . . . . . + . . . 1 7 Neckera complanata E0 . . . . . . . . . . . + . . . 1 7 Collema cristatum E0 . . . . . . . . . . . + . . . 1 7 Plagiomnium undulatum E0 . . . . . . . . . . . . . 1 . 1 7 Rhytidiadelphus triquetrus E0 . . . . . . . . . . . . . 1 . 1 7 Mnium marginatum E0 . . . . . . . . . . . . . . 1 1 7 Plagiomnium affine E0 . . . . . . . . . . . . . . + 1 7 Plagiothecium undulatum E0 . . . . . . . . . . . . . . + 1 7 Legend - Legenda AR Limestone with chert - apnenec z rožencem AGR Limestone with chert and claystone - apnenec z rožencem in glinavcem DR Dolomite with chert - dolomit z rožencem AL Limestone with marlstone - apnenec z laporovcem ALR Limestone with chert and marlstone - apnenec z rožencem in laporovcem R Chert - roženec Re Rendzina - rendzina Dy Dystric brown soil - distrična rjava tla Number of relevé (Zaporedna številka popisa) 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 Pr. Fr. Number of relevé (Zaporedna številka popisa) 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 Pr. Fr. Number of relevé (Zaporedna številka popisa) 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 Pr. Fr. Number of relevé (Zaporedna številka popisa) 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 Pr. Fr. Table 11: Synoptic table of syntaxa Arunco-Fagetum, Homogyno-Fagetum, Saxifrago cuneifolii-Fagetum, Rhododendro hirsuti-Fagetum and Ranunculo platanifolii-Fagetum Preglednica 11: Sintezna tabela sintaksonov Arunco-Fagetum, Homogyno-Fagetum, Saxifrago cuneifolii-Fagetum, Rhododendro hirsuti-Fagetum in Ranunculo platanifolii-Fagetum Successsive number (Zaporedna številka) 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 Sign for syntaxa (Oznaka sintaksonov) ArFty ArFho ArFao ArF ArFov ArFtil HoFty ScFar ScFluz ScFfo ScFca ScFpc ScFrh RF-At RF-Sa HoFrh RpFho RpFty Author (Avtor popisov) ŽK ŽK MA ID ID ID ID ID ID ID ID ID ID ID ID ID LM LM Number of relevés (Število popisov) 18 8 12 6 41 32 31 17 12 42 18 8 8 18 20 18 10 12 AF Aremonio-Fagion Cyclamen purpurascens E1 94 100 75 100 98 97 90 88 83 88 100 63 100 100 95 94 70 25 Cardamine enneaphyllos E1 100 100 75 83 95 100 55 100 83 57 83 13 25 56 10 6 70 67 Helleborus niger E1 67 63 92 100 63 3 16 . . . . . . . 5 6 80 17 Ruscus hypoglossum E1 39 63 42 33 . . . . . . . . . . . . . . Cardamine trifolia E1 22 13 . 50 63 50 61 82 92 67 17 50 25 28 20 11 70 92 Aremonia agrimonoides E1 17 . . 33 2 3 . . 10 . . . . . . 30 42 Vicia oroboides E1 17 13 17 17 . 3 . . . . . . . . . . 20 17 Lamium orvala E1 11 . . 50 49 66 26 59 33 5 17 . 13 . . 6 . 33 Knautia drymeia E1 6 . 8 17 12 . . . . 2 22 25 . . . . . . Hacquetia epipactis E1 . 13 42 67 39 44 . 6 . 5 6 . . . . . 20 8 Omphalodes verna E1 . . 92 17 71 3 . . . 2 . . . . . . 10 8 Euphorbia carniolica E1 . . 57 50 49 13 . . . . . . . . . . . . Potentilla carniolica E1 . . 42 33 15 . . . . . . . . . . . . . Rhamnus fallax E2 . . 17 . 24 3 6 . . . . . . 39 . . . . Daphne laureola E2a . . 17 33 7 3 . . . . . . . . . . . . Scopolia carniolica E1 . . 8 . 7 . . . 5 . . . . . . . . Epimedium alpinum E1 . . 8 33 . 3 . . . . . . . . . . . . Calamintha grandiflora E1 . . 8 17 . . . . . . . . . . . . . 8 Anemone trifolia E1 . . . 17 73 97 68 76 50 21 . 25 13 83 40 61 . . Geranium nodosum E1 . . . . 10 50 . . . . . . . . . . . . Anemone x pittonii E1 . . . . 5 . 6 . 5 . . . 6 . . . . EC Erythronio-Carpinion Primula vulgaris E1 . . 42 50 85 56 29 53 33 71 83 50 13 28 15 44 20 17 Erythronium dens-canis E1 . . 33 17 10 . . . . . . . . . . . . . Lonicera caprifolium E2a . . . 17 10 . . . . . . . . . . . . . Helleborus odorus E1 . . . . 12 78 . 18 . . . 13 . . . . . 8 Galanthus nivalis E1 . . . . 2 44 . . . . . . . . . . . . Ornithogalum pyrenaicum E1 . . . . . 16 . . . . 11 . . . . . . . Crocus vernus subsp. vernus E1 . . . . . 6 . . . . . . . . . . . . TA Tilio-Acerion Acer pseudoplatanus E3 72 75 83 100 80 66 58 47 58 24 22 . 13 6 25 44 40 33 Acer pseudoplatanus E2 55 38 42 50 32 47 52 29 33 38 33 . 38 . 30 50 30 50 Acer pseudoplatanus E1 78 38 33 100 80 53 77 82 83 52 22 13 25 6 20 11 20 25 Polystichum aculeatum E1 67 100 33 100 83 59 48 82 67 21 11 13 13 11 10 11 40 33 Aruncus dioicus E1 61 75 100 17 44 44 6 71 42 48 6 13 . 17 10 11 30 25 Euonymus latifolia E2 50 50 . 33 10 9 6 . 8 17 . . . 6 5 17 . . Acer platanoides E3 22 25 17 50 15 22 3 12 . 2 . . . . 5 . . . Acer platanoides E2 44 13 . . 12 22 . 6 . 7 . . . . . . . . Acer platanoides E1 28 . 17 33 22 44 3 6 8 14 . . . . 5 . . . Ulmus glabra E3 22 25 . 50 20 47 . 6 8 2 . . . . . . . 8 Ulmus glabra E2 28 25 . 33 22 41 . . 8 2 . . . . . . 10 8 Ulmus glabra E1 11 13 8 67 29 28 . . . 7 . . . . . . . . Phyllitis scolopendrium E1 11 13 . 33 10 94 . 12 17 . . . . . . . . . Thalictrum aquilegiifolium E1 11 . . 17 5 . . 24 . 2 6 13 25 6 . . 40 . Arum maculatum E1 11 . . 33 . 6 . 6 . . . . . . . . . . Tephroseris longifolia E1 . . 25 50 32 . . . . . . . . . . . . . Isopyrum thalictroides E1 . . 17 . . . . . . . . . . . . . . . Dryopteris affinis E1 . . . 17 10 19 3 18 25 7 . . . . . 6 . . Lunaria rediviva E1 . . . 17 2 . . . 8 . . . . . . . . . Staphylea pinnata E2 . . . 17 2 13 . . . . . . . . . . . . Staphylea pinnata E1 . . . 17 . 3 . . . . . . . . . . . . Tilia platyphyllos E3 . . . 17 . 34 6 . . 2 . . . . 5 . . . Tilia platyphyllos E2 . . . . . 9 . . . . . . . . 10 6 . . Tilia platyphyllos E1 . . . 17 . 13 . . . . . . . . . 6 . . Polystichum x bicknellii E1 . . . 17 . 3 3 . . . . . . . . . . . Juglans regia E3 . . . . 2 . . . . . . . . . . . . . Juglans regia E2 . . . . 12 . . . . 5 . . . . . . . . Juglans regia E1 . . . . 7 3 . 6 . 2 . . . . . . . . Geranium robertianum E1 . . . . 5 22 13 24 . 5 . . . . . 11 . 17 Polystichum setiferum E1 . . . . 5 19 . 12 8 2 . . . . . . . . Tephroseris pseudocrispa E1 . . . . 2 13 . 6 . 17 . . . 11 5 . . . Polystichum braunii E1 . . . . 2 3 6 12 17 7 . . 13 . . 28 . . Adoxa moschatellina E1 . . . . . 19 10 12 25 . . . . . . . . 58 Corydalis solida E1 . . . . . 3 . . . . . . . . . . . . Polystichum x illyricum E1 . . . . . 3 6 12 . 2 . . . 6 10 . . . Stellaria montana E1 . . . . . . 3 12 8 . . . . . . . . . Impatiens noli-tangere E1 . . . . . . . 6 . . . . . . . . . 8 Chrysosplenium alternifolium E1 . . . . . . . . . . . . . . . . . 25 FS Fagetalia sylvaticae Fagus sylvatica E3 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 Fagus sylvatica E2 100 100 58 83 93 88 97 100 92 90 72 100 100 78 55 72 90 67 Fagus sylvatica E1 33 . 58 33 78 53 71 47 50 31 11 25 50 . 15 28 30 8 Mercurialis perennis E1 100 100 100 83 95 91 81 88 92 93 100 50 63 100 60 67 90 25 Cardamine bulbifera E1 100 100 8 33 2 44 6 35 8 . 6 . . . . . 20 50 Lonicera alpigena E2 94 50 75 83 37 3 52 47 50 69 78 50 88 100 80 56 50 50 Daphne mezereum E2a 89 100 92 67 93 69 81 82 67 93 61 25 100 83 100 100 60 33 Prenanthes purpurea E1 89 100 75 67 39 31 97 94 67 90 83 88 25 78 50 78 50 33 Polygonatum multiflorum E1 89 88 67 83 34 63 6 41 33 21 6 . . 17 . . 20 17 Dryopteris filix-mas E1 72 100 8 83 66 84 68 94 83 90 44 50 50 39 55 44 50 83 Euphorbia amygdaloides E1 72 75 . . 63 31 10 71 . . 56 25 . 28 . . 70 75 Actaea spicata E1 72 50 17 50 46 84 65 100 83 88 11 13 13 22 20 39 50 25 Paris quadrifolia E1 67 25 17 50 27 47 19 53 33 21 . . . 6 . 6 20 67 Mycelis muralis E1 56 50 17 50 41 50 45 47 33 33 22 . 25 6 15 39 50 50 Lilium martagon E1 50 25 25 50 29 34 16 53 33 52 28 50 13 22 10 11 . 33 Galeobdolon flavidum E1 39 38 33 83 83 75 26 71 42 19 22 . . . 10 . 40 67 Neottia nidus-avis E1 39 25 . 67 44 38 13 65 8 24 6 25 . 11 5 . 10 . Epipactis helleborine E1 39 88 8 17 20 19 42 47 . 24 44 13 . 39 5 11 10 17 Galium odoratum E1 33 13 . . 10 16 . 29 25 2 . . . . . . 20 42 Viola reichenbachiana E1 17 38 8 . 22 25 . 6 17 40 11 13 50 39 40 . 20 17 Pulmonaria officinalis E1 17 25 17 33 17 28 19 76 25 43 17 13 13 . 5 11 . 17 Fraxinus excelsior E3 . . 33 . 20 28 6 . . . . . . . . . . . Fraxinus excelsior E2 11 . . 17 22 19 3 . . . . . . . . . . . Fraxinus excelsior E1 . . . . 32 16 6 . . . . . . . . . . . Sambucus nigra E3 . . . 17 2 . . . . . . . . . . . . . Sambucus nigra E2 11 13 . 17 15 28 10 41 25 10 . . . . . 6 10 . Festuca altissima E1 11 . . . 2 3 23 71 25 21 39 . . . . 6 20 8 Carex sylvatica E1 11 . . . . . . 18 . . . . . . . . . 50 Salvia glutinosa E1 6 13 100 33 78 81 77 65 33 83 61 13 38 39 50 94 10 8 Prunus avium E2 6 . . . 10 9 . . 8 2 . . . . . . . . Epilobium montanum E1 6 25 . . . 6 13 12 8 14 6 25 63 6 5 6 . 25 Asarum europaeum subsp. europaeum E1 . 25 8 50 . . . . . . . . . . . . . . Campanula trachelium E1 . 13 17 17 15 38 32 76 17 40 44 38 25 17 20 6 10 25 Heracleum sphondylium E1 . . 75 33 15 22 . 71 8 5 33 13 . . . . . 17 Symphytum tuberosum E1 . . 33 67 44 50 32 76 25 50 61 88 63 11 30 33 10 75 Melica nutans E1 . . 25 50 37 19 3 12 . 26 72 . 13 67 35 39 . . Laburnum alpinum E3 . . . . 12 3 48 12 17 43 50 13 50 61 85 50 . . Laburnum alpinum E2 . . 25 . 15 3 65 12 8 26 72 25 63 39 85 72 . . Laburnum alpinum E1 . . . . 24 9 . 12 8 19 44 13 13 22 35 . . . Brachypodium sylvaticum E1 . . 17 17 10 16 . . . . . . . . . . . . Petasites albus E1 . . 8 50 2 16 10 24 25 17 6 13 . . . 6 10 8 Prunus avium E3 . . . . 17 31 3 6 8 5 . . . . . . . . Prunus avium E1 . . 8 . 2 9 3 6 . 5 . . . . . . . . Leucojum vernum E1 . . 8 17 . . . . . . . . . . . . . . Euphorbia dulcis E1 . . 8 17 7 50 . 6 . . 6 . . . 5 . . . Galium laevigatum E1 . . . 50 90 78 . 94 75 98 100 88 75 100 95 . . 25 Asarum europaeum subsp. caucasicum E1 . . . 17 66 97 29 35 58 60 . . 25 17 40 79 . . Lathyrus vernus subsp. vernus E1 . . . 17 46 88 16 100 25 43 6 50 38 39 30 11 . 42 Cephalanthera damasonium E1 . . . 50 . 9 . 18 . 12 . . . . . . . . Allium ursinum E1 . . . 33 2 34 . . . . . . . . . . . 8 Sanicula europaea E1 . . . 33 2 . 3 53 17 40 . . . . . . 10 58 Scrophularia nodosa E1 . . . 33 . 19 16 47 25 17 6 13 . . . . 10 50 Cardamine pentaphyllos E1 . . . 17 2 31 3 6 . . . . . . . . . . Sambucus nigra E1 . . . 17 . 13 . . . . . . . . . . . . Ranunculus lanuginosus E1 . . . 17 . 3 . 24 . . . 13 . . . . 30 42 Phyteuma spicatum subsp. spicatum E1 . . . 17 . . . . . . . . . . . . 60 50 Phyteuma spicatum subsp. coeruleum E1 . . . . 46 3 . . . . . . . . . . . . Carpinus betulus E3 . . . . 12 53 3 18 25 2 . . . . 5 . . . Carpinus betulus E2 . . . . 5 9 . . 17 . . . . . . . . . Carpinus betulus E1 . . . . 2 . . . . 2 . . . . . . . . Lathyrus vernus subsp. flaccidus E1 . . . . 2 13 10 24 17 21 . 25 63 6 30 28 . . Tilia cordata E3 . . . . . 66 6 6 8 . . . 13 . 15 17 . . Luzula nivea E1 . . . . . 47 . . . 10 . . 13 17 . . . . Tilia cordata E2 . . . . . 31 3 6 . 5 . . . 6 10 . . . Tilia cordata E1 . . . . . 22 3 12 . . . . . . . 11 . . Orchis pallens E1 . . . . . 13 . . . . . . . . . . . . Cardamine impatiens E1 . . . . . 6 3 12 . . . . . . . . . . Myosotis sylvatica E1 . . . . . 6 . 6 . . . . . . . . . . Poa nemoralis E1 . . . . . . 3 . . 5 6 25 25 6 . . . . Corydalis cava E1 . . . . . . . 6 . . . . . . . . 10 8 Epipactis leptochila E1 . . . . . . . 6 . . . . 13 . . . . . Circaea lutetiana E1 . . . . . . . . 8 . . . . . . . . . Hordelymus europaeus E1 . . . . . . . . . 2 . . . . . . . . Orobanche salviae E1 . . . . . . . . . 2 . . . . . . . . QP Quercetalia pubescenti-petraeae Sorbus aria E3 . . 33 33 10 28 58 12 50 57 83 50 100 89 90 94 . . Sorbus aria E2 33 63 . 50 10 6 48 24 8 71 67 88 100 94 100 89 40 . Sorbus aria E1 . . . 17 12 . . . . 5 6 25 13 . . . . . Tamus communis E1 22 13 50 67 12 9 . . . . . . . . . . . . Cephalanthera rubra E1 22 38 . 17 . . . . . 5 . . . . . . . . Convallaria majalis E1 11 25 50 33 . 9 . . . 12 17 13 25 72 40 . . . Melittis melissophyllum E1 6 . 17 33 29 38 26 35 8 52 78 50 . 44 20 11 . . Ostrya carpinifolia E3 . 13 8 50 59 84 68 53 25 92 17 . 63 61 100 94 . . Ostrya carpinifolia E2 . . . . 7 3 68 . 8 7 . . . 28 70 94 . . Ostrya carpinifolia E1 . . . . . 3 . . 17 . . . . . . . . . Asparagus tenuifolius E1 . . 50 . 5 6 . . . . . . . . . . . . Acer obtusatum E3 . . 17 . . . . . . . . . . . . . . . Acer obtusatum E2 . . 42 . . . . . . . . . . . . . . . Fraxinus ornus E3 . . . 17 34 38 29 6 25 33 6 . 13 33 40 61 . . Fraxinus ornus E2 . . 33 50 41 19 19 24 . 60 . . . 50 55 56 . . Fraxinus ornus E1 . . 25 50 46 44 16 12 8 24 6 . . . 30 28 . . Cornus mas E2 . . 25 17 24 19 . 6 . . . . . . . . . . Carex flacca E1 . . 25 33 . . . . . . . . . 6 . . . . Hypericum montanum E1 . . . 33 2 13 6 12 . 29 22 . 38 11 5 6 . . Euonymus verrucosa E2 . . . 17 20 22 3 6 . 2 . . . 17 15 . . . Tanacetum corymbosum E1 . . . 17 . 25 19 41 8 48 78 100 63 17 15 6 . . Arabis turrita E1 . . . 17 . 16 19 24 8 36 22 . 38 . 10 11 . . Clematis recta E1 . . . 17 . . . . . . . . . . . . . . Mercurialis ovata E1 . . . . 10 . . . . . . . . . . . . . Aristolochia lutea E1 . . . . 7 . . . . . . . . . . . . . Sesleria autumnalis E1 . . . . 2 31 29 71 50 29 . 38 13 . 30 44 . . Calamintha sylvatica E1 . . . . 2 6 . 6 . 2 . . 13 . . . . . Orchis mascula subsp. speciosa E1 . . . . 2 . . . . . . . . . . . . . Ruscus aculeatus E1 . . . . . 22 . 6 . . . . . . . . . . Primula x ternovania E1 . . . . . 6 . . . . . . . . . . . . Primula veris subsp. columnae E1 . . . . . 3 6 . . 2 . 50 75 . . . . . Quercus cerris E1 . . . . . 3 . . . . . . . . . . . . Sorbus mougeoti agg. (inc. S. austriaca) E2 . . . . . . 3 . . . . . . . 5 6 . . Epipactis muelleri E1 . . . . . . . 6 . 5 . . . . . . . . Sorbus x thuringiaca E2 . . . . . . . . . 2 . . . . . . . . Peucedanum schottii E1 . . . . . . . . . . 6 13 . . . . . . Potentilla micrantha E1 . . . . . . . . . . . . 13 . . . . . QR Quercetalia roboris Rubus hirtus E2a 50 38 8 . 27 3 3 6 25 12 6 13 13 6 . . . . Pteridium aquilinum E1 11 . 25 33 15 . . 6 . 2 . . . 6 . . . . Quercus robur E3 . . . 17 . 13 . . . . . . . . . . . . Quercus petraea E3 . . . . 5 . . . . 2 . . . . . . . . Quercus petraea E1 . . . . . . . . . 7 . . . . . . . . Betula pendula E3 . . . . . 3 3 . . . . . . 6 5 6 . . Betula pendula E2 . . . . . . . . . 2 . . . . . . . . Populus tremula E3 . . . . . . 3 . 25 . . . . . . . . . Populus tremula E1 . . . . . . . . 8 . . . . . . . . . Hieracium racemosum E1 . . . . . . . 12 8 33 . 13 . . 5 . . . Hieracium sabaudum E1 . . . . . . . . . 7 . . . . . . . . Betonica officinalis E1 . . . . . . . . . 7 . . . 6 . . . . Castanea sativa E1 . . . . . . . . . 2 . . . . . . . . Hieracium lachenalii E1 . . . . . . . . . 2 6 13 . . . . . . QF Querco-Fagetea Anemone nemorosa E1 78 25 33 33 34 . . 59 50 52 89 63 38 17 20 11 70 83 Lonicera xylosteum E2 61 75 . 33 39 59 3 6 . 69 . . . 11 20 17 40 . Hedera helix E3a 50 13 8 33 17 44 . 6 . 5 . . . . . . . . Hedera helix E1 . . 25 33 24 75 . 35 17 24 . . . . . . . . Galium sylvaticum E1 50 75 83 33 . . . . . . . . . . . . . . Carex digitata E1 39 63 17 17 46 81 . 82 58 86 67 . 25 44 85 . 50 . Platanthera bifolia E1 33 13 8 . 2 . . 18 . 5 6 . . 22 5 6 10 . Corylus avellana E3a . . . . 2 . . . . . . . . . . . . . Corylus avellana E2 17 25 . 17 37 28 42 29 17 29 28 . . 22 25 17 30 17 Ilex aquifolium E3a . . . 17 5 3 . . . . . . . . . . . . Ilex aquifolium E2 17 25 17 17 20 6 3 . . . . . . . . . . . Spiraea chamaedryfolia E2a 17 13 . 17 20 13 . . . . . . . 6 . . . . Clematis vitalba E2 6 25 25 67 39 41 10 29 17 10 6 . . . . . . . Hepatica nobilis E1 . 75 75 67 54 53 16 12 . 62 72 25 88 61 20 33 50 25 Pulmonaria stiriaca E1 . . 25 . . . . . . . . . . . . . . . Taxus baccata E3 . . 17 17 12 . . 6 . . . . . . . . . . Taxus baccata E2 . . . 17 7 . 10 6 . 2 . . . . 11 22 . . Cephalanthera longifolia E1 . . 17 33 5 . . . . 7 . 13 . . . . . . Rosa arvensis E2a . . . 33 15 16 . 12 . 26 . . . 11 . . . . Veratrum nigrum E1 . . . 17 41 50 13 47 . 7 . . 25 11 5 11 . . Acer campestre E3 . . . 17 22 34 . . . 5 . . . . . . . . Acer campestre E2 . . . 17 17 28 . 6 8 7 . . . . . . . . Acer campestre E1 . . . 17 12 13 . 6 8 10 . . . . . . . . Listera ovata E1 . . . 17 12 34 . . . 2 . 13 . . . . 20 . Viburnum opulus E2 . . . 17 10 9 . . . . . . . . 5 . . . Moehringia trinervia E1 . . . 17 2 16 3 24 25 2 6 . . . . . . 8 Vinca minor E1 . . . . 56 69 . 6 8 26 . . . . . . . . Aegopodium podagraria E1 . . . . 27 22 3 24 8 14 . . . . . . . . Malus sylvestris E3 . . . . 2 . . . . . . . . . . . . . Malus sylvestris E2 . . . . 5 . . . . 2 . . . . . . . . Rubus caesius E2a . . . . 5 . . . . . . . . . . . . . Viola mirabilis E1 . . . . 5 16 . . . . . . . . . . . . Viola riviniana E1 . . . . 5 3 . . . 17 6 . . . . . . . Anemone ranunculoides E1 . . . . 2 . . 6 . . . . . . . . . 8 Carex umbrosa E1 . . . . 2 . . . . . . . . . . . . . Scilla bifolia E1 . . . . 2 . . . . . . . . . . . . . Equisetum hyemale E1 . . . . 2 . . . . . . . . . . . . . Cerastium sylvaticum E1 . . . . . 9 . . . . . . . . . . . . Gagea lutea E1 . . . . . 6 . . . . . . . . . . . 8 Galium schultesii E1 . . . . . 6 . . . 2 . . . . . . 20 . Lathraea squamaria E1 . . . . . 3 . . . . . . . . . . . . Melica uniflora E1 . . . . . 3 . . . . . . . . . . . . Festuca heterophylla E1 . . . . . . 10 18 . 33 50 75 50 6 10 . 10 . Dactylorhiza fuchsii E1 . . . . . . . 12 . 26 17 25 . 17 . . . . Cruciata glabra E1 . . . . . . . . . 5 . . 13 . . . . . Pyrus pyraster E2 . . . . . . . . . 2 . . . . . . . . Viscum album E3 . . . . . . . . . . . . . . 5 . . . Veronica montana E1 . . . . . . . . . . . . . . . . . 25 EP Erico-Pinetea Calamagrostis varia E1 11 38 . 50 54 34 23 29 8 24 50 . 13 89 70 67 60 33 Cirsium erisithales E1 . 25 92 83 71 9 55 76 17 71 89 75 88 83 75 67 10 . Carex alba E1 . . 33 83 85 56 26 29 17 29 17 . . 78 30 44 20 . Erica carnea E1 . . 33 . 12 . 6 . 8 12 22 . 25 100 80 61 10 . Rubus saxatilis E1 . . 25 17 20 6 6 . 8 40 83 13 50 89 65 22 20 . Aquilegia nigricans E1 . . 8 17 17 9 . 29 8 24 61 13 . 33 10 . . . Buphthalmum salicifolium E1 . . . 50 12 3 19 6 . 21 56 25 38 56 45 17 . . Peucedanum austriacum E1 . . . 17 2 . . . . 26 28 . . 22 10 . . . Epipactis atrorubens E1 . . . 17 2 . 3 . . 2 . 13 13 17 5 . . . Rhododendron hirsutum E2a . . . . 10 . 23 . . 7 17 . 75 100 100 100 . . Molinia caerulea subsp. arundinacea E1 . . . . 2 . . . 8 . . . . . 5 . . . Aquilegia atrata E1 . . . . . 9 . . . . . . . 6 5 . . . Carex ornithopoda E1 . . . . . . . 6 . 5 6 . . 28 15 . . . Polygala chamaebuxus E1 . . . . . . 3 . . 2 . . . 44 25 6 . . Centaurea jacea agg. E1 . . . . . . 3 . . . . . . 33 5 . . . Cotoneaster tomentosus E2 . . . . . . . . . 2 . . . 17 15 . . . Allium ericetorum E1 . . . . . . . . . . 6 . 13 6 . . . . Rhodothamnus chamaecistus E2 . . . . . . . . . . . . . 50 10 . . . Pinus mugo E2 . . . . . . . . . . . . . 22 . . . . Amelanchier ovalis E2 . . . . . . . . . . . . . 17 . . . . Chamaecytisus hirsutus E1 . . . . . . . . . . . . . 17 . . . . Gymnadenia odoratissima E1 . . . . . . . . . . . . . 11 . . . . VP Vaccinio-Piceetea Laserpitium krapfii E1 100 88 92 17 . . 3 47 . 2 . . 25 11 5 11 . . Rosa pendulina E2a 89 88 67 67 17 . 35 29 33 67 89 63 100 100 90 78 50 17 Gentiana asclepiadea E1 33 50 83 67 20 . 52 59 25 76 44 38 50 61 45 33 60 58 Solidago virgaurea E1 17 38 42 83 63 50 71 65 42 98 100 88 63 72 95 83 10 8 Homogyne sylvestris E1 11 100 50 67 46 6 68 41 25 90 56 13 38 78 85 94 100 . Oxalis acetosella E1 11 38 . 17 41 53 71 88 100 81 28 50 50 17 25 78 50 83 Luzula sylvatica E1 6 13 8 33 . . . . . . . 13 . . . . 30 8 Polystichum lonchitis E1 . 25 . . . 3 . . . 2 11 13 13 17 5 . 20 8 Aposeris foetida E1 . . 75 50 68 22 6 18 8 36 72 88 . 33 . . 40 75 Valeriana tripteris E1 . . 58 50 61 19 52 29 8 71 94 75 100 89 80 83 100 17 Abies alba E3 . . 25 33 15 9 100 12 8 . 11 . 13 11 15 94 . 17 Abies alba E2 . . 17 . 2 16 87 18 58 5 6 . 13 22 45 94 10 0 Abies alba E1 . . . . 7 13 48 . . . . . . 6 30 28 . 25 Veronica urticifolia E1 . . 17 17 54 41 71 88 75 98 94 100 88 44 70 44 70 50 Hieracium murorum E1 . . . 33 17 9 48 47 17 90 89 100 50 50 45 28 60 8 Picea abies E3 . . . 50 15 22 45 18 8 21 39 38 13 33 55 22 80 33 Picea abies E2 . . . 33 17 9 10 41 42 52 22 13 13 61 40 28 40 33 Picea abies E1 . . . 17 12 . 13 12 25 29 11 . . . . . . . Saxifraga cuneifolia E1 . . . 33 7 22 61 53 75 86 39 50 50 22 20 6 20 25 Dryopteris dilatata E1 . . . 33 . . 19 12 25 5 6 13 . 11 10 6 . . Vaccinium myrtillus E1 . . . 17 7 . 19 . . 24 56 38 38 72 45 33 10 8 Gymnocarpium dryopteris E1 . . . 17 5 . 35 47 83 17 . 13 13 28 65 28 30 17 Maianthemum bifolium E1 . . . . 7 16 10 29 8 67 44 38 38 50 5 6 20 58 Clematis alpina E1 . . . . 5 . 23 . . 14 50 13 88 67 65 56 40 8 Calamagrostis arundinacea E1 . . . . 2 . 87 76 92 95 100 100 100 50 95 83 10 . Lonicera nigra E2a . . . . 2 . . . . 2 . . 13 11 . 6 . . Phegopteris connectilis E1 . . . . 2 . 19 . 50 19 17 . 25 11 20 33 . . Luzula luzuloides E1 . . . . . 9 52 53 100 69 72 100 75 11 30 17 . 8 Luzula pilosa E1 . . . . . 6 . . . 5 . . . . . . . . Dryopteris expansa E1 . . . . . 6 . . . . . . . . . . 20 33 Huperzia selago E1 . . . . . . 45 6 33 43 44 13 50 83 60 72 . . Orthilia secunda E1 . . . . . . 13 . . 12 . . . 6 5 . . . Larix decidua E3 . . . . . . 13 . . 17 6 13 13 22 35 28 10 . Larix decidua E2 . . . . . . . . . . . . . 6 5 . . . Vaccinium vitis-idaea E1 . . . . . . 3 . . 2 6 . . 17 25 6 . . Dryopteris carthusiana E1 . . . . . . 3 . . . . . . . . 6 . . Thelypteris limbosperma E1 . . . . . . . . 8 . . . . . . . . . Avenella flexuosa E1 . . . . . . . . . 2 . 13 . . . . . 8 Luzula luzulina E1 . . . . . . . . . 2 . . 25 . . . . . Corallorhiza trifida E1 . . . . . . . . . 2 . . . . . . . . Calamagrostis villosa E1 . . . . . . . . . . . . . . . 11 . . SSc Sambuco-Salicion capreae Sorbus aucuparia E3 . . . . . . 19 . . . 6 13 25 17 65 39 . . Sorbus aucuparia E2 33 38 . 17 . . 13 . . 21 17 13 15 72 45 61 60 25 Sorbus aucuparia E1 . . . 50 7 . 23 . 42 7 22 13 25 22 30 22 . . Salix caprea E3 . . . . . . . 12 . 2 . . . . . . . . Sambucus racemosa E2 . . . . . . . . . . . . . . . 6 . . RP Rhamno-Prunetea Crataegus monogyna E2a . . . 17 10 6 . 6 . 5 . . . 6 . . . . Crataegus monogyna E3a . . . . 5 . . . . . . . . . . . . . Viburnum lantana E2 . . . 17 5 9 . . . 2 . . . 11 . . . . Ligustrum vulgare E2a . . . 17 2 3 . . . . . . . . . . . . Cornus sanguinea E2b . . . . 12 3 . . . . . . . . . . . . Berberis vulgaris E2 . . . . 10 9 . . . . . . . . . . . . Euonymus europaea E2 . . . . 5 16 . . . . . . . . . . . . Rubus fruticosus agg. E2a . . . . . 3 . . . . . . . . . . . . Rosa canina E2a . . . . . . . . . 2 . . . . . . . . Rubus dumetorum E2a . . . . . . . . . . . . 13 . . . . . Rhamnus catharticus E2 . . . . . . . . . . . . . 6 . . . . Juniperus communis E2 . . . . . . . . . . . . . . 15 . . . Cotoneaster integerrimus E2 . . . . . . . . . . . . . . 5 . . . TG Trifolio-Geranietea Melampyrum velebiticum E1 39 88 42 . . . . . . . . . . . . . . . Digitalis grandiflora E1 . 25 42 50 12 9 6 24 8 26 22 25 13 . 5 11 20 8 Vincetoxicum hirundinaria E1 . . 8 33 34 25 13 6 . 12 11 . . 28 10 33 . . Campanula rapunculoides E1 . . . 17 24 50 . 47 17 40 . 13 38 11 45 . . . Lilium carniolicum E1 . . . 17 7 . . 6 . 2 17 . . 6 5 . . . Iris graminea E1 . . . 17 . . . . . . 28 25 . 6 5 . . . Clinopodium vulgare E1 . . . . 2 . . 24 . 7 28 25 25 . . 6 . . Laserpitium latifolium E1 . . . . 2 . . . . . 22 25 . . 5 . . . Viola hirta E1 . . . . 2 . . . . . . . . . . . . . Vicia sylvatica E1 . . . . . 3 10 . . 14 . 50 13 . 5 . . . Origanum vulgare E1 . . . . . . . 12 . . 6 . . 6 . . . . Achillea distans E1 . . . . . . 10 . . 7 . . 38 . 10 6 . . Polygonatum odoratum E1 . . . . . . . . . 2 . . . 22 . . . . Anthericum ramosum E1 . . . . . . . . . . 11 . . . 5 . . . Thalictrum minus E1 . . . . . . . . . . 11 25 . 11 5 . . . Arabis pauciflora E1 . . . . . . . . . . 11 . . 11 . . . . Libanotis sibirica s. lat. E1 . . . . . . . . . . 11 13 . 11 25 . . . Verbascum lychnitis (inc. Verbascum sp.) E1 . . . . . . . . . . 6 . 13 . . . . . Calamintha einseleana E1 . . . . . . . . . . 6 . . . . . . . Trifolium rubens E1 . . . . . . . . . . 6 . . . . . . . Laserpitium siler E1 . . . . . . . . . . . . . 17 5 . . . Valeriana collina (V. wallrothii) E1 . . . . . . . . . . . . . 6 . . . . Hypericum perforatum E1 . . . . . . . . . . . . . . . . 10 . Fragaria moschata E1 . . . . . . . . . . . . . . . . 10 . EA Epilobietea angustifolii Verbascum nigrum E1 39 50 . . . . . . . . . . . . . . . . Fragaria vesca E1 11 38 . . 2 6 10 . . 45 56 38 63 . 15 6 . . Astragalus glycyphillos E1 6 . . . . . . . . . . . . . . . . . Eupatorium cannabinum E1 . . . 50 10 3 3 . . . . . . . . . . . Stachys alpina E1 . . . 33 . . . . 8 . . . . . . . . . Rubus idaeus E2a . . . 17 . 3 42 41 25 14 22 25 75 . 55 61 . 25 Bromopsis benekenii E1 . . . 17 . . . 6 . 2 . . . . . . . 17 Atropa bella-donna E1 . . . 17 . . 13 . . . . . . . . . . . Verbascum lanatum E1 . . . . 2 3 10 29 8 5 17 63 . . . 6 . . Hypericum hirsutum E1 . . . . . 6 16 29 8 12 28 13 . . . 6 . . Solanum dulcamara E1 . . . . . 3 6 . . 2 . . 13 . . 6 . . Stachys sylvatica E1 . . . . . 3 . 18 . . . . . . . . . 8 Galeopsis speciosa E1 . . . . . . 6 6 8 2 . 25 . . . . . 8 Galeopsis pubescens E1 . . . . . . 3 . . . . . . . . . . . Arctium nemorosum E1 . . . . . . . 6 . . . . . . . . . . Tussilago farfara E1 . . . . . . . . . 5 . . . . . . . . BA Betulo-Alnetea Salix glabra E2 . . . . 2 . . . . . . . . 22 . . . . Salix appendiculata E2 . . . . . . . . . 2 . . 38 39 60 39 . . Alnus viridis E2b . . . . . . . . . . . 13 . . 5 17 . . Sorbus chamaemespilus E2 . . . . . . . . . . . . . 6 . . . . MuA Mulgedio-Aconitetea Phyteuma ovatum E1 72 88 67 50 2 3 68 59 83 98 83 75 13 39 30 11 . . Senecio ovatus E1 56 88 75 67 76 41 97 94 92 74 72 100 100 6 55 94 20 100 Aconitum lycoctonum s. lat. E1 22 50 75 50 54 50 . 29 . 7 56 25 . . . . . . Athyrium filix-femina E1 11 . 8 33 37 16 52 82 75 67 33 63 38 39 25 28 30 92 Veratrum album s. lat. E1 11 . 33 33 10 3 10 41 . . 17 50 25 6 . . 50 67 Polygonatum verticillatum E1 . . 25 . 12 . 13 18 . 7 39 38 13 11 . 6 80 92 Centaurea montana E1 . . 42 50 7 . . 6 . . . 13 . 11 . . . . Lathyrus laevigatus E1 . . 8 . . . . . . . . . . . . . . . Aconitum degenii subsp. paniculatum E1 . . . 50 22 16 13 35 . . . . 25 11 . 6 . 8 Silene dioica E1 . . . 17 . 13 . 18 . 2 6 . . . . . . . Crepis paludosa E1 . . . 17 . . . . . . . . . . . . . . Chaerophyllum hirsutum E1 . . . . 2 . . . . . . . . . . . 10 8 Aconitum angustifolium E1 . . . . . 22 19 24 . 10 44 13 63 56 25 28 . . Milium effusum E1 . . . . . 19 6 6 . . . . . . . . 10 8 Doronicum austriacum E1 . . . . . 3 . 6 . 2 . . . . . . . 8 Saxifraga rotundifolia E1 . . . . . 3 . . . . . 13 . . . . 40 33 Ranunculus platanifolius E1 . . . . . . 10 6 . 2 . 25 13 6 . . 60 42 Hypericum maculatum E1 . . . . . . 3 . . . . . . . . . . . Pleurospermum austriacum E1 . . . . . . . . . . 11 25 . . . . . . Silene vulgaris subsp. antelopum E1 . . . . . . . . . . 6 . . . . . . . Chaerophyllum villarsii E1 . . . . . . . . . . . 13 . . . . . . Cicerbita alpina E1 . . . . . . . . . . . . . . . . 10 . Geum rivale E1 . . . . . . . . . . . . . . . . 10 . Anthriscus nitidus E1 . . . . . . . . . . . . . . . . . 33 ES Elyno-Seslerietea Betonica alopecuros E1 . . 25 . 7 . 6 . . 7 72 25 38 61 25 11 . . Aster bellidiastrum E1 . . . 33 2 . . . . 2 . . . 17 30 17 . . Phyteuma orbiculare E1 . . . 17 . . . . . . 39 25 . 6 . . . . Sesleria tenuifolia subsp. kalnikensis E1 . . . 17 . . . . . . . . . . . . . . Carex ferruginea E1 . . . . 17 . . . . . . . . . . . . . Sesleria caerulea subsp. calcaria E1 . . . . 7 3 39 . 25 26 83 38 100 94 100 94 . . Laserpitium peucedanoides E1 . . . . 2 . . . . 7 39 . 13 44 20 . . . Campanula witasekiana E1 . . . . . . 6 . . 21 67 38 75 56 40 11 . . Centaurea haynaldii subsp. julica E1 . . . . . . . . . 2 28 13 13 . . . . . Carduus crassifolius E1 . . . . . . . . . . 39 . . 17 . . . . Serratula macrocephala E1 . . . . . . . . . . 17 . . . . . . . Astrantia bavarica E1 . . . . . . . . . . 6 . . . . . . . Carex mucronata E1 . . . . . . . . . . 6 . . 6 . . . . Globularia cordifolia E1 . . . . . . . . . . . . . 11 . . . . Senecio abrotanifolius E1 . . . . . . . . . . . . . 6 . . . . Gentiana lutea subsp. symphyandra E1 . . . . . . . . . . . . . . 25 . . . FB Festuco-Brometea Pimpinella saxifraga E1 . . . 50 . . . 6 . 10 . . . . . . . . Carex humilis E1 . . . . 5 . . . . 17 56 25 25 33 45 . . . Brachypodium rupestre E1 . . . . . . 3 . . 2 . 25 38 6 20 . . . Allium carinatum subsp. pulchellum E1 . . . . . . . . . 5 6 . . . . . . . Carlina acaulis E1 . . . . . . . . . 5 11 25 . 6 . . . . Bromopsis transsilvanica E1 . . . . . . . . . . 6 13 13 . 15 . . . Dianthus monspessulanus E1 . . . . . . . . . . 6 . . . . . . . Euphorbia cyparissias E1 . . . . . . . . . . 6 . . . . . . . Koeleria pyramidata E1 . . . . . . . . . . . 13 13 . . . . . Thymus praecox E1 . . . . . . . . . . . . 13 . . . . . Campanula rotundifolia agg. E1 . . . . . . . . . . . . . . . . . 20 MA Molinio-Arrhenatheretea, Molinion caeruleae Ajuga reptans E1 11 . . 17 . 3 . . . . . . . . . . . 17 Adenophora liliifolia E1 . . 25 . . . . . . . . . . . . . . . Narcisus poeticus E1 . . 8 . . . . . . . . . . . . . . . Angelica sylvestris E1 . . . 17 7 3 . . . 17 11 . . 6 . . . . Cirsium oleraceum E1 . . . . 2 . . . . . . . . . . . . . Veronica chamaedrys E1 . . . . . 3 . 12 . . . . . . . . . . Dactylis glomerata s.str. E1 . . . . . . . 6 . . . 25 . . . . . . Galium album E1 . . . . . . . 6 . 2 22 . 13 . . . . . Lotus corniculatus E1 . . . . . . . . . 2 . . . 6 . . . . Taraxacum officinale E1 . . . . . . . . . . 6 . . . 5 . . . Vicia cracca E1 . . . . . . . . . . . . 13 . 5 . . . CU Calluno-Ulicetea JT Selaginella helvetica E1 . . . . . . 3 . . 2 . . 38 . 10 17 . . Festuca ovina agg. (inc. Festuca sp.) E1 . . . . . . . . . . 6 13 . . . . . . Phyteuma zahlbruckneri E1 . . . . . . . . . . 6 13 . . . . . . Potentilla erecta E1 . . . . . . . . . . . 13 . . . . . . Festuca nigrescens E1 . . . . . . . . . . . 13 . 6 . . . . GU Galio-Urticetea Parietaria officinalis E1 . . . 17 . . . . . . . . . . . . . . SM Chelidonium majus E1 . . . 17 . . . . . . . . . . . . . . Urtica dioica E1 . . . . . 3 3 12 17 . . . 13 . . 6 . 8 TR Thlaspietea rotundifolii Gymnocarpium robertianum E1 . . . 33 39 9 13 6 17 38 6 . 13 72 50 56 . . Adenostyles glabra E1 . . . . 17 16 77 71 33 60 78 63 50 67 65 67 60 67 Astrantia carniolica E1 . . . . 7 . 3 . . 2 . . 13 33 5 6 30 . Hieracium bifidum E1 . . . . 2 . . . . 5 . . . . . . . . Cerastium subtriflorum E1 . . . . . 6 . . 8 . . . . . 5 . . . Campanula cochleariifolia E1 . . . . . . 6 . . . 11 . . 6 5 6 . . Campanula cespitosa E1 . . . . . . . . . 2 . . . 22 . 6 . . Epilobium collinum E1 . . . . . . . . . 2 . . . . . . . . Arabis alpina E1 . . . . . . . . . . 6 . 13 . . . . . Heracleum pollinianum E1 . . . . . . . . . . 6 . . . . . . . Ligusticum seguieri E1 . . . . . . . . . . 6 13 . . . . . . Aquilegia einseleana E1 . . . . . . . . . . . . . 6 . . . . Petasites paradoxus E1 . . . . . . . . . . . . . 6 15 . . . Scrophularia juratensis E1 . . . . . . . . . . . . . 6 . . . . Gypsophila repens E1 . . . . . . . . . . . . . . 10 . . . Saxifraga aizoides E1 . . . . . . . . . . . . . . . 6 . . AT Asplenietea trichomanis Asplenium viride E1 . 13 8 17 51 19 58 41 92 76 72 . 75 72 75 78 10 8 Asplenium trichomanes E1 . . . 33 44 84 55 65 50 48 22 38 38 22 50 56 . 17 Asplenium ruta-muraria E1 . . . 17 29 28 29 24 8 26 39 25 50 50 85 61 10 . Moehringia muscosa E1 . . . 33 27 44 13 6 . 50 11 . 63 . 40 6 10 . Cardaminopsis arenosa E1 . . . 17 . 3 . . . . . . . . 5 . . . Paederota lutea E1 . . . . 24 . . . . . 6 . 13 22 . 6 10 . Polypodium vulgare E1 . . . . 22 69 26 47 50 45 11 25 . 11 40 56 . 33 Phyteuma scheuchzeri subsp. columnae E1 . . . . 15 3 . . . . . . . 17 . . . . Carex brachystachys E1 . . . . 5 . 10 6 . 5 11 . 13 33 55 22 . . Cystopteris fragilis E1 . . . . 5 13 16 . 17 17 22 13 38 11 5 17 . 17 Primula carniolica E1 . . . . 2 . . . . . . . . . . . . . Saxifraga petraea E1 . . . . . 6 . . . . . . . . 5 . . . Asplenium adiantum-nigrum E1 . . . . . 3 . 6 . 2 . . . . . . . . Campanula carnica E1 . . . . . . 6 . . 5 11 38 63 11 40 44 . . Hieracium glaucum E1 . . . . . . 3 . . . . . . 6 . . . . Ceterach officinarum s. lat. E1 . . . . . . 3 . . . . . . . . . . . Saxifraga crustata E1 . . . . . . . . . 7 11 . 38 . 50 11 . . Saxifraga hostii E1 . . . . . . . . . 2 . . . . 5 . . . Festuca stenantha (inc. F. valesica agg.) E1 . . . . . . . . . . 11 13 13 33 10 6 . . Sedum album E1 . . . . . . . . . . 6 . 13 . . . . . Campanula spicata E1 . . . . . . . . . . . 13 . . . . . . Primula auricula E1 . . . . . . . . . . . 13 38 17 25 . . . Micromeria thymifolia E1 . . . . . . . . . . . . 25 6 5 6 . . Kernera saxatilis E1 . . . . . . . . . . . . 25 . 10 . . . Hieracium pospichalii E1 . . . . . . . . . . . . 13 . . . . . Sedum maximum E1 . . . . . . . . . . . . 13 . . . . . Valeriana saxatilis E1 . . . . . . . . . . . . . 50 30 17 . . Potentilla caulescens E1 . . . . . . . . . . . . . 11 5 . . . Athamanta turbith E1 . . . . . . . . . . . . . 6 5 . . . Hieracium villosum E1 . . . . . . . . . . . . . . 5 . . . Moehringia villosa E1 . . . . . . . . . . . . . . 5 . . . Sedum hispanicum E1 . . . . . . . . . . . . . . 5 . . . ML Mosses and lichens (Mahovi in lišaji) Ctenidium molluscum E0 . . 25 67 93 78 10 82 83 86 89 38 100 94 95 28 70 50 Neckera crispa E0 . . 25 33 85 63 74 65 83 83 22 13 88 72 95 94 20 17 Polytrichum formosum E0 . . 8 67 46 28 81 94 83 93 78 75 75 78 90 83 . 8 Thuidium tamariscinum E0 . . . 33 5 31 3 18 25 40 6 . . . 25 11 10 . Dicranum scoparium E0 . . . 33 2 3 48 18 25 43 22 13 . 33 70 72 . . Plagiomnium undulatum E0 . . . 33 . 25 13 12 . 14 . . . . . 6 . . Fissidens dubius E0 . . . 17 56 63 65 65 58 71 56 . 38 67 75 72 . . Tortella tortuosa E0 . . . 17 29 9 71 53 17 74 89 50 100 94 90 94 . 17 Schistidium apocarpum E0 . . . 17 20 25 65 65 8 48 50 50 63 39 45 61 . . Bartramia halleriana E0 . . . 17 7 6 13 18 17 33 . 13 13 6 40 28 10 . Atrichum undulatum E0 . . . 17 2 6 19 18 67 33 22 63 38 11 20 17 . . Conocephalum conicum E0 . . . 17 2 6 10 6 25 10 6 . 25 11 10 28 . . Homalothecium philippeanum E0 . . . 17 2 13 6 . 8 10 6 . 38 6 . . . . Peltigera leucophlebia E0 . . . 17 . . 3 . . 10 . . 13 . 20 6 . . Plagiochila porelloides E0 . . . . 32 28 35 18 25 43 11 . 13 33 70 78 50 33 Isothecium alopecuroides E0 . . . . 24 81 55 59 92 52 11 13 75 22 35 28 30 42 Encalypta streptocarpa E0 . . . . 10 . . 6 . . 11 . . . 5 . . . Eurhynchium angustirete E0 . . . . 7 31 10 . 8 17 . . . 17 15 6 . . Marchantia polymorpha E0 . . . . 7 . 6 . 8 . 6 . 13 . 5 6 . . Rhytidiadelphus triquetrus E0 . . . . 7 6 3 . . 12 . . 13 17 50 44 10 . Peltigera canina E0 . . . . 7 19 35 18 25 54 17 . 63 6 5 28 . . Brachythecium rutabulum E0 . . . . 5 34 . 18 . 10 . . . . . . 20 . Metzgeria furcata E0 . . . . 5 16 35 41 25 62 17 13 25 22 40 39 30 . Mnium thomsonii E0 . . . . 5 25 . 12 17 38 22 13 25 . . . . . Anomodon viticulosus E0 . . . . 2 25 . . 8 . . . . . . . . . Bryum capillare E0 . . . . 2 16 6 12 33 12 22 . 13 . 40 6 . . Fissidens taxifolius E0 . . . . 2 . 6 6 17 10 . 13 . . . . 30 42 Hypnum cupressiforme E0 . . . . 2 22 65 47 67 60 6 13 38 44 70 67 10 . Mnium marginatum E0 . . . . 2 . 32 . 7 17 22 13 . 11 15 44 . . Plagiothecium sp. E0 . . . . 2 . . . . . . . . . . . . . Thuidium delicatulum E0 . . . . 2 6 . . . . . . . . . . . . Cladonia pyxidata E0 . . . . 2 . . . . 10 . . 13 44 60 . . . Thamnobryum alopecurum E0 . . . . . 59 . 18 25 7 . . . . . . . . Plagiothecium denticulatum E0 . . . . . 44 39 88 58 71 28 25 13 28 50 28 10 . Anomodon attenuatus E0 . . . . . 28 . 18 17 2 . . . . . . . . Brachythecium velutinum E0 . . . . . 22 6 18 17 17 17 13 . . . . . . Porella platyphylla E0 . . . . . 22 . 6 17 12 . . . . . . . . Homalothecium lutescens E0 . . . . . 9 3 6 . 24 . 13 . 6 10 . . . Isothecium myosuroides E0 . . . . . 9 35 18 . 12 11 . . 11 45 39 . . Neckera complanata E0 . . . . . 9 6 6 8 2 . . . . . 6 . . Hylocomium splendens E0 . . . . . 3 6 . 8 26 . . 13 . 45 39 . . Hypnum cupressiforme var. filiforme E0 . . . . . 3 19 . 8 7 11 . . . 5 44 . . Plagiomnium cuspidatum E0 . . . . . 3 3 6 . . . . . . . 3 . . Plagiothecium undulatum E0 . . . . . 3 10 . . . . . 13 . . 22 . . Scleropodium purum E0 . . . . . 3 . . . 2 . . . 6 10 . . . Ulota crispa E0 . . . . . 3 . 6 . . 6 . . . . . . . Collema cristatum E0 . . . . . 3 . 18 8 24 . . . . 15 6 . . Dicranella heteromalla E0 . . . . . . 29 . 25 24 22 25 13 11 20 11 . . Cladonia sp. E0 . . . . . . 23 . . 7 . . . . . 22 . . Leucobryum glaucum E0 . . . . . . 10 . 17 7 17 . . 17 30 28 . . Bryum argenteum E0 . . . . . . 10 . . 5 11 . 13 . 10 11 . . Dicranum majus E0 . . . . . . 10 . . 2 . . . 11 15 17 . . Plagiothecium nemorale (P. sylvaticum) E0 . . . . . . 6 12 8 21 . 13 . . 5 28 . . Rhizomnium punctatum E0 . . . . . . 3 . . 17 . . . . 5 6 . . Bazzania trilobata E0 . . . . . . 3 . 8 . . . . . . . . . Hookeria lucens E0 . . . . . . 3 . . . . . . . . 17 . . Plagiomnium affine E0 . . . . . . 3 . . 12 . . . . . . . . Mnium sp. E0 . . . . . . 3 . 8 . . . . . 10 17 . . Orthothecium rufescens E0 . . . . . . 3 . . . . . . 17 5 . . . Cladonia furcata E0 . . . . . . . 12 8 33 6 25 13 . . . . . Cladonia rangiferina E0 . . . . . . . 6 17 14 11 . . . . . . . Brachythecium oxycladum E0 . . . . . . . 6 8 2 . . . . . . . . Platygyrium repens E0 . . . . . . . 6 . 2 . . . . . . . . Dicranodontium denudatum E0 . . . . . . . 6 . . 6 . . . . . . . Homalothecium sericeum E0 . . . . . . . . . 5 6 . . . . . . . Paraleucobryum sauteri E0 . . . . . . . . . 2 . 13 . . . . . . Bryum sp. E0 . . . . . . . . . 2 . . . . 5 6 . . Brachythecium sp. E0 . . . . . . . . . 2 . . . . . . . . Orthotrichum anomalum E0 . . . . . . . . . 2 . . . . . . . . Porella arboris-vitae E0 . . . . . . . . . 2 . . . . 5 . . . Radula complanata E0 . . . . . . . . . 5 . . . . . . . . Riccardia sp. E0 . . . . . . . . . 2 . . . . . . . . Scapania nemorea E0 . . . . . . . . . 5 . . . . . . . . Rhizocarpon geographicum E0 . . . . . . . . . . 6 . . . . . . . Distichium capillaceum E0 . . . . . . . . . . . . 13 . . . . . Rhytidiadelphus loreus E0 . . . . . . . . . . . . . . . 12 . . 1 Arunco-Fagetum var. geogr Ruscus hypoglossum typicum, Košir 1979, Table 8, columns 1-18 2 Arunco-Fagetum var. geogr. Ruscus hypoglossum homogynetosum sylvestris, Košir 1979, Table 8, columns 19-26 3 Arunco-Fagetum var. geogr. Acer obtusatum, Accetto 2007, Table 1 4 Arunco-Fagetum, Dolenjska, this paper, Table 4 5 Arunco-Fagetum var. geogr. Anemone trifolia subvar. geogr. Omphalodes verna, this paper, Table 1 6 Arunco-Fagetum var. geogr. Anemone trifolia tilietosum, this paper, Table 2 7 Homogynos sylvestris-Fagetum var. geogr. Sesleria autumnalis typicum, Dakskobler 2002, Phyt. Table 2, relevés 1-31 8 Saxifrago cuneifolii-Fagetum aruncetosum, this paper, Table 6 9 Saxifrago cuneifolii-Fagetum gymnocarpietosum dryopteridis, this paper, Table 10, columns 1-12 10 Saxifrago cuneifolii-Fagetum fraxinetosum orni, this paper,Table 9 11 Saxifrago cuneifolii-Fagetum typicum var. Calamagrostis varia, this paper, Table 7, relevés 1-18 12 Saxifrago cuneifolii-Fagetum typicum var. Primula columnae, this paper, Table 7, relevés 19-26 13 Saxifrago cuneifolii-Fagetum rhododendretosum hirsuti, this paper, Table 8, relevés 1-8 14 Rhododendro hirsuti-Fagetum var. geogr. Anemone trifolia subvar. geogr. Aconitum angustifolium forma Pinus mugo, Dakskob­ler 2003, Phyt.Table 3 15 Rhododendro hirsuti-Fagetum var. geogr. Anemone trifolia subvar. geogr. Aconitum angustifolium forma Sesleria autumnalis, Dakskobler 2003, Phyt. Table 4 16 Homogyno sylvestris-Fagetum var. geogr. Sesleria autumnalis rhododendretosum hirsuti Dakskobler, 2002, Phyt. Table 1 17 Ranunculo platanifolii-Fagetum var. geogr. typica homogynetosum sylvestris, Marinček & Čarni 2010, Table 14, relevés 1-10. 18 Ranunculo platanifolii-Fagetum var. geogr. typica typicum, Marinček & Čarni 2010, Table 1, relevés 1-12 Successsive number (Zaporedna številka) 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 Successsive number (Zaporedna številka) 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 Successsive number (Zaporedna številka) 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 Successsive number (Zaporedna številka) 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 Successsive number (Zaporedna številka) 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 Successsive number (Zaporedna številka) 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 Successsive number (Zaporedna številka) 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 Table 12: Diagnostic species of syntaxa Arunco-Fagetum, Homogyno-Fagetum, Saxifrago cuneifolii-Fagetum, Rhododendro hirsuti-Fa­getum and Ranunculo platanifolii-Fagetum Preglednica 12: Diagnostične vrste sintaksonov Arunco-Fagetum, Homogyno-Fagetum, Saxifrago cuneifolii-Fagetum, Rhododendro hirsuti-Fagetum in Ranunculo platanifolii-Fagetum Successsive number (Zaporedna številka) 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 Sign for syntaxa (Oznaka sintaksonov) ArFty ArFho ArFao ArF ArFov ArFtil HoFty ScFar ScFluz ScFfo ScFca ScFpc ScFrh RF-At RF-Sa HoFrh RpFho RpFty Author (Avtor popisov) ŽK ŽK MA ID ID ID ID ID ID ID ID ID ID ID ID ID LM LM Number of relevés (Število popisov) 18 8 12 6 41 32 31 17 12 42 18 8 8 18 20 18 10 12 Diagnostic species of the association Arunco-Fagetum (Diagnostične vrste asociacije Arunco-Fagetum) VP Laserpitium krapfii E1 100 88 92 17 . . 3 47 . 2 . . 25 11 5 11 . . AF Helleborus niger E1 67 63 92 100 63 3 16 . . . . . . . 5 6 80 17 TA Aruncus dioicus E1 61 75 100 17 44 44 6 71 42 48 6 13 . 17 10 11 30 25 AF Ruscus hypoglossum E1 39 63 42 33 . . . . . . . . . . . . . . TG Melampyrum velebiticum E1 39 88 42 . . . . . . . . . . . . . . . QP Tamus communis E1 22 13 50 67 12 9 . . . . . . . . . . . . QF Spiraea chamaedryfolia E2a 17 13 . 17 20 13 . . . . . . . 6 . . . . QF Veratrum nigrum E1 . . . 17 41 50 13 47 . 7 . . 25 11 5 11 . . Diagnostic species of the association Homogyno sylvestris-Fagetum (Diagnostične vrste asociacije Homogyno sylvestris-Fagetum) VP Abies alba E3 . . 25 33 15 9 100 12 8 . 11 . 13 11 15 94 . 17 VP Abies alba E2 . . 17 . 2 16 87 18 58 5 6 . 13 22 45 94 10 . VP Abies alba E1 . . . . 7 13 48 . . . . . . 6 30 28 . 25 VP Veronica urticifolia E1 . . 17 17 54 41 71 88 75 98 94 100 88 44 70 44 70 50 TR Adenostyles glabra E1 . . . . 17 16 77 71 33 60 78 63 50 67 65 67 60 67 VP Homogyne sylvestris E1 11 100 50 67 46 6 68 41 25 90 56 13 38 78 85 94 100 . VP Saxifraga cuneifolia E1 . . . 33 7 22 61 53 75 86 39 50 50 22 20 6 20 25 AT Asplenium viride E1 . 13 8 17 51 19 58 41 92 74 72 0 75 72 75 78 10 8 Diagnostic species of the association Saxifrago cuneifolii-Fagetum (Diagnostične vrste asociacije Saxifrago cuneifolii-Fagetum) VP Veronica urticifolia E1 . . 17 17 54 41 71 88 75 98 94 100 88 44 70 44 70 50 VP Calamagrostis arundinacea E1 . . . . 2 . 87 76 92 95 100 100 100 50 95 83 10 . VP Saxifraga cuneifolia E1 . 33 7 22 61 53 75 86 39 50 50 22 20 6 20 25 VP Luzula luzuloides E1 . . . . . 9 52 53 100 69 72 100 75 11 30 17 . 8 VP Huperzia selago E1 . . . . . . 45 6 33 43 44 13 50 83 60 72 . . QP Tanacetum corymbosum E1 . . . 17 . 25 19 41 8 48 78 100 63 17 15 6 . . QF Festuca heterophylla E1 . . . . . . 10 18 . 33 50 75 50 6 10 . 10 TG Clinopodium vulgare E1 . . . . 2 . . 24 . 7 28 25 25 . . 6 . . ES Campanula witasekiana E1 . . . . . . 6 . . 21 67 38 75 56 40 11 . . FB Carex humilis E1 . . . . 5 . . . . 17 56 25 25 33 45 . . . Diagnostic species of the association Rhododendro hirsuti-Fagetum (Diagnostične vrste asociacije Rhododendro hirsuti-Fagetum) EP Rhododendron hirsutum E2a . . . . 10 . 23 . . 7 17 . 75 100 100 100 . . EP Rhodothamnus chamaecistus E2 . . . . . . . . . . . . . 50 10 . . . VP Homogyne sylvestris E1 11 100 50 67 46 6 68 41 25 90 56 13 38 78 85 94 100 . VP Clematis alpina E1 . . . . 5 . 23 . . 14 50 13 88 67 65 56 40 8 BA Salix appendiculata E2 . . . . . . . . . 2 . . 38 39 60 39 . . AT Carex brachystachys E1 . . . . 5 . 10 6 . 5 11 . 13 33 55 22 . . FS Laburnum alpinum E3 . . . . 12 3 48 12 17 43 50 13 50 61 85 50 . . FS Laburnum alpinum E2 . . 25 . 15 3 65 12 8 26 72 25 63 39 85 72 . . FS Laburnum alpinum E1 . . . . 24 9 . 12 8 19 44 13 13 22 35 . . . QP Ostrya carpinifolia E3 13 8 50 59 84 68 53 25 92 17 . 63 61 100 94 . . QP Ostrya carpinifolia E2 . . . . 7 3 68 . 8 7 . . . 28 70 94 . . QP Ostrya carpinifolia E1 . . . . . 3 . . 17 . . . . . . . . . QP Fraxinus ornus E3 . . 17 34 38 29 6 25 33 6 . 13 33 40 61 . . QP Fraxinus ornus E2 . . 33 50 41 19 19 24 . 60 . . . 50 55 56 . . QP Fraxinus ornus E1 . . 25 50 46 44 16 12 8 24 6 . . . 30 28 . . QP Melittis melissophyllum E1 6 . 17 33 29 38 26 35 8 50 78 50 . 44 20 11 . . QP Convallaria majalis E1 11 25 50 33 . 9 . . . 12 17 13 25 72 40 . . . Diagnostic species of the association Ranunculo platanifolii-Fagetum (Diagnostične vrste asociacije Ranunculo platanifolii-Fagetum) TR Adenostyles glabra E1 . . . . 17 16 77 71 33 60 78 63 50 67 65 67 60 67 MuA Polygonatum verticillatum E1 . . 8 . 12 . 13 18 . 7 39 38 13 11 . 6 80 92 MuA Ranunculus platanifolius E1 . . . . . . 10 6 . 2 . 25 13 6 . . 60 42 Table 13: Phytosciological groups in syntaxa Arunco-Fagetum, Homogyno-Fagetum, Saxifrago cuneifolii-Fagetum, Rhododendro hirsuti-Fagetum and Ranunculo platanifolii-Fagetum (relative frequencies) Preglednica 13: Fitosociološke skupine sintaksonov Arunco-Fagetum, Homogyno sylvestris-Fagetum, Saxifrago cuneifolii-Fagetum, Rhododendro hirsuti-Fagetum in Ranunculo platanifolii-Fagetum (relativne frekvence) Successsive number (Zaporedna številka) 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 Sign for syntaxa (Oznaka sintaksonov) ArFty ArFho ArFao ArF ArFov ArFtil HoFty ScFar ScFluz ScFfo ScFca ScFpc ScFrh RF-At RF-Sa HoFrh RpFho RpFty Author (Avtor popisov) ŽK ŽK MA ID ID ID ID ID ID ID ID ID ID ID ID ID LM LM Number of relevés (Število popisov) 18 8 12 6 41 32 31 17 12 42 18 8 8 18 20 18 10 12 Aremonio-Fagion 10,7 9,53 14,4 10,9 12,3 8,9 6,82 6,73 7,68 4,29 4,19 3,72 3,14 5,21 2,95 3,65 10,5 8,84 Erythronio-Carpinion 0 0 1,8 1,21 2,15 3,33 0,61 1,14 0,75 1,18 1,62 1,33 0,22 0,47 0,26 0,87 0,57 0,7 Tilio-Acerion 16,4 12,8 8,99 12,8 9,85 12,7 6,36 8,05 9,54 4,67 1,71 1,06 2,26 1,15 2,43 3,98 5,95 8,08 Fagetalia sylvaticae 41, 369 27,5 26,3 30,6 36,1 28,3 37,6 32,2 28 23,8 23,1 22,9 20,48 20,8 22,3 29,5 38,3 Quercetalia pubescenti-petraeae 2,7 3,97 8,99 8,21 6,06 7,55 8,75 5,51 4,86 9,34 6,96 9,04 10,1 8,71 10,8 11,9 1,13 0 Quercetalia roboris 1,75 0,99 0,79 0,72 0,84 0,31 0,19 0,38 1,5 1,24 0,19 0,8 0,22 0,4 0,17 0,12 0 0 Querco-Fagetea 10,6 11,1 8,39 8,7 110 12,3 2,37 7,19 4,87 8,57 5,91 4,52 4,26 3,81 3,66 2,32 8,5 4,85 Erico-Pinetea 0,32 1,64 4,58 4,83 5,19 2,13 3,51 2,84 1,69 4,5 7,42 2,9 5,59 15,3 9,88 7,61 3,4 0,92 Vaccinio-Piceetea 7,66 11,5 12,8 10,6 8,94 5,51 24,8 14,3 21,3 21,4 20,7 23,2 20,6 19,6 21,7 23,7 24,9 16,9 Sambuco-Salicion capreae 0,95 0,99 0 0,97 0,13 0 1,15 0,19 0,94 0,51 0,76 0,81 1,17 1,85 2,43 2,54 1,7 0,7 Rhamno-Prunetea 0 0 0 0,72 0,88 0,83 0 0,1 0 0,15 0 0 0,22 0,38 0,35 0 0 0 Trifolio-Geranietea 1,12 2,95 2,21 1,93 1,54 1,46 0,82 1,9 0,56 1,84 3,33 4,25 2,46 2,25 2,25 1,11 1,13 0,22 Epilobietea angustifolii 1,61 2,3 0 1,93 0,26 0,47 2,29 2,18 1,31 1,46 2,1 3,46 2,69 0 1,21 1,69 0 1,62 Betulo-Alnetea 0 0 0 0 0,04 0 0 0 0 0,04 0 0,27 0,67 1,12 1,13 1,11 0 0 Mulgedio-Aconitetea 4,94 5,9 7,98 5,31 4 3,17 6,11 6,82 5,62 4,41 6,29 9,57 5,15 3,09 2,34 3,43 9,07 13,7 Elyno-Seslerietea 0 0 0,6 0,97 0,66 0,05 1,07 0 0,56 1,1 6,76 2,92 4,26 5,31 4,16 2,64 0 0 Festuco-Brometea 0 0 0 0,72 0,09 0 0,06 0,09 0 0,63 1,52 2,13 1,79 0,75 1,39 0 0 0,56 Molinio-Arrhenatheretea 0,32 0 0,79 0,48 0,18 0,16 0 0,38 0 0,35 0,67 0,53 0,45 0,2 0,17 0 0 0,47 Calluno-Ulicetea 0 0 0 0 0 0 0,06 0 0 0,04 0,19 1,06 0,67 0,1 0,17 0,34 0 0 Galio-Urticetea 0 0 0 0,48 0 0,05 0,06 0,19 0,37 0 0 0 0,22 0 0 0,12 0 0,22 Thlaspietea rotundifolii 0 0 0 0,48 1,23 0,52 2,08 1,23 1,31 1,72 1,91 1,6 1,57 3,64 2,69 2,91 2,55 1,87 Asplenietea trichomanis 0 0,34 0,19 1,69 4,05 4,53 4,6 3,13 4,87 4,51 4 3,72 9,41 6,31 9,7 7,65 1,13 2,09 Skupaj (Total) 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 I. DAKSKOBLER, B. DOLINAR, B. ZUPAN, F. POLJŠAK & P. STRGAR: FITOCENOLOŠKA OZNAKA RASTIŠČ VRSTE NIGRITELLA I. DAKSKOBLER, B. DOLINAR, B. ZUPAN, F. POLJŠAK & P. STRGAR: FITOCENOLOŠKA OZNAKA RASTIŠČ VRSTE NIGRITELLA 106 107 FOLIA BIOLOGICA ET GEOLOGICA 56/1, 105–114, LJUBLJANA 2015 FOLIA BIOLOGICA ET GEOLOGICA 56/1 – 2015 FOLIA BIOLOGICA ET GEOLOGICA 56/1 – 2015 FITOCENOLOŠKA OZNAKA RASTIŠČ VRSTE NIGRITELLA ARCHIDUCIS-JOANNIS V DOLINI TRIGLAVSKIH JEZER (JULIJSKE ALPE, SLOVENIJA)* PHYTOSOCIOLOGICAL CHARACTERISTICS OF THE SITES OF NIGRITELLA ARCHIDUCIS-JOANNIS IN THE TRIGLAV LAKES VALLEY (THE JULIAN ALPS, SLOVENIA) Igor DAKSKOBLER1, Branko DOLINAR2, Branko ZUPAN3, Florijan POLJŠAK4 & Peter STRGAR5 *Posvečeno prof. dr. Vladu Ravniku ob njegovi 90. letnici IZVLEČEK Fitocenološka oznaka rastišč vrste Nigritella archiducis-joannis v Dolini Triglavskih jezer (Julijske Alpe, Slovenija) V članku podajamo kronologijo odkrivanja vrste Nigri­tella archiducis-joannis v Sloveniji in s fitocenološko tabelo opišemo njena rastišča v Dolini Triglavskih jezer. Raste na alpinskih traviščih, ki jih uvrščamo v asociacijo Ranunculo hybridi-Caricetum sempervirentis in v jugovzhodnoalpsko zvezo Caricion austroalpinae. Ugotavljamo, da vrsti Nigritel­la archiducis-joannis in N. widderi zaradi njunega uspevanja v Julijskih Alpah nista najboljši značilnici severnoalpske zveze Seslerion variae, saj s svojo prisotnostjo povezujeta po­dobna travišča tako v Severovzhodnih kot v Jugovzhodnih Alpah. Ključne besede: Nigritella archiducis-joannis, Ranuncu­lo hybridi-Caricetum sempervirentis, Julijske Alpe,Triglavsko pogorje, Triglavski narodni park, Slovenija ABSTRACT Phytosociological characteristics of the sites of Nigritella archiducis-joannis in the Triglav Lakes Valley (The Julian Alps, Slovenia) The article provides a chronology of the discovery of Nigritella archiducis-joannis in Slovenia and a phytosocio­logical table that offers a description of its sites in the Triglav Lakes Valley. It grows on alpine grasslands classified into the association Ranunculo hybridi-Caricetum sempervirentis and into the southeastern-Alpine alliance Caricion austroal­pinae. We have determined that Nigritella archiducis-joannis and N. widderi cannot be considered good character species of the northern-Alpine alliance Seslerion variae, as they are native to the Julian Alps and as such represent a link with similar grasslands both in the Northeastern and the South­eastern Alps. Key words: Nigritella archiducis-joannis, Ranunculo hy­bridi-Caricetum sempervirentis, Julian Alps, Triglav Moun­tains, Triglav National Park, Slovenia 1 UVOD Nigritella archiducis-joannis je apomiktični, tetraploi­dni takson iz skupine rdeče murke (Nigritella rubra = N. miniata s. lat.), ki sta ga kot novega opisala Teppner & Klein (1985). Njegovo klasično nahajališče je v po­gorju Totes Gebirge, na prisojnih alpinskih karbona­tnih traviščih. Je endemit avstrijske dežele Štajerske oz. širše endemit Avstrije (Teppner 2008, Staudinger & al. 2009). Aeschimann & al. (2004: 1118) ga ozna­čujejo kot vzhodnoalpski takson, ki ga poznajo le na Štajerskem v Avstriji in je značilen za subalpinsko-al­pinska travišča na karbonatni podlagi iz podzveze Se­slerienion variae. Domnevno uspeva tudi v Karavan­kah, na Ojstrcu (Hochobir) na avstrijskem Koroškem (Foelsche 2011). V Dolini Triglavskih jezer je Florijan Poljšak 31.7. 2004 fotografiral murko, ki se je razliko­vala tako od črne (Nigritella rhellicani) kot od rdeče (Nigritella rubra = N. miniata) in je ni znal določiti. Helmuth Zelesny in Hans-Peter Welle sta 12. julija 2008 na alpinskem travišču v osrednjem delu Trigla­vskega narodnega parka, na nadmorski višini okoli 1850 m, opazila devet cvetočih primerkov murke, ki je po videzu povsem ustrezala taksonu N. archiducis-jo­annis, kot sta ga poznala s Štajerske (Zelesny 2008: 589). Bolj natančno smo položaj tega nahajališča ugo­tovili iz fotografij, ki nam jih je prijazno poslal Zelesny. Iz njih smo lahko sklepali na dolino Triglavskih jezer med Dvojnim jezerom in Jezerom v Ledvicah (9648/4) in posledično tudi ugotovili, da gre za isto nahajališče, kot ga je Florijan Poljšak našel že leta 2004. Njegov po­snetek njemu takrat neznane murke odgovarja posnet­kom, ki jih je objavil Zelesny (ibid.) in nedvomno pripadajo taksonu N. archiducis-joannis. Torej je F. Poljšak prvi, ki je opazil in fotografiral to murko tudi v Sloveniji. O tej najdbi, oziroma o uspevanju taksona Nigritella archiducis-joannis v Julijskih Alpah v Slove­niji, je kasneje poročal tudi Foelsche (2011: 83). Med murkami, ki uspevajo v Sloveniji, smo ga omenili tudi mi, ko smo poročali o najdbi Widderjeve murke (Nigri­tella widderi) v Julijskih Alpah (Dakskobler & al. 2012). Kljub intenzivnemu iskanju v letih 2012 in 2013 (iskali so ga F. Poljšak in B. Dolinar ter P. Strgar, B. Zupan in I. Veber) nam nahajališča v Dolini Trigla­vskih jezer ni uspelo potrditi. Pač pa je poleti 2013 Branko Dolinar našel vrsto Nigritella archiducis-joan­nis, za katero predlagamo slovensko ime Janezova murka (glej tudi Dolinar 2015), pod vrhom Viševni­ka, prav tako v Triglavskem pogorju, kjer je naštel šest rastlin. Posamezne primerke, ki morfološko ustrezajo Janezovi murki, je sam ali v družbi s F. Poljšakom opa­zil tudi na širšem območju Tegoške gore v grebenu Ko­šute v Karavankah, a ti podatki so še v obdelavi in do­ločitev za zdaj ni preverjena. Poleti 2014 so Polona in Peter Strgar ter Branko Zupan Janezovo murko ponov­no opazili in fotografirali na znanem nahajališču v Dolini Triglavskih jezer. Branko Zupan je cvetoče pri­merke opazil tudi 9. 8. 2014 in nato skupaj z I. Daksko­blerjem še 11. 8. 2014. Takrat sta na tem nahajališču naštela 12 cvetočih Janezovih murk. Istega dne sta našla še eno nahajališče nekoliko nižje pri Utah, v ko­tanji pod Kopico, kjer je cvetelo šest primerkov te vrste. Skupno število opaženih Janezovih murk v Dolini Tri­glavskih jezer v letu 2014 je 18. Na obeh nahajališčih smo naredili fitocenološke popise homogenih sestojev, skupno pet, in v tem članku bomo kratko opisali rasti­šča in združbene razmere tega za zdaj izjemno redkega taksona v flori Slovenije. Menimo, da je prav zaradi njegove redkosti takšna obravnava smiselna. Branko Dolinar je pripravil tudi njegov kratek morfološki opis in si pomagal z ustreznimi strokovnimi deli, ki obravnavajo to in druge kukavičevke (Redl 1999, Ravnik 2003, Delforge 2006, Kretzschmar 2008, Foelsche 2011): Nigritella archiducis-joannis Teppner & E. Klein Sin.: Gymnadenia archiducis-joannis (Teppner & E. Klein) Teppner & E. Klein, Nigritella rubra subsp. ar­chiducis-joannis (Teppner & E. Klein) H. Kretzschmar comb. nov. Janezova murka ima do deset pritličnih žlebastih listov strnjenih v rozeto. Eden med njimi je izrazito krajši in širši od ostalih. Običajno je nekaj listov pole­glih, ostali se dvigajo proti socvetju nekoliko odma­knjeno od stebla. Do osem svetlo zelenih stebelnih li­stov, ki sežejo do socvetja, se tesno prilega steblu. Po robovih so srebrno obarvani. Socvetje je enobarvno, polokroglo, na vrhu nekoliko potlačeno (topo). Barva cvetov je temnorožnata (mesnordeča) proti koncu cve­tenja postane svetlejša. Podporni listi so proti vrhu vi­jolični. Cvet ima oba stranska zunanja cvetna lista po­udarjeno štrleča navzven. Zunanji srednji in oba no­tranja cvetna lista se tesno prilegajo k medeni ustni, ki je po dolžini cevasto zvita in obrnjena navzgor. Dolga je do osem in široka do štiri milimetre, malo pod sre­dino ima zažetek. Ostroga je dolga največ dva milime­tra, plodnica ni zasukana. 2 METODE Travišča v Dolini Triglavskih jezer smo preučili po standardni srednjeevropski metodi (Braun-Blanquet 1964). Fitocenološke popise smo vnesli v bazo FloVegSi (Seliškar & al. 2003). S pomočjo te podatkovne baze smo izdelali tudi arealno karto (slika 1). Iz fitocenolo­ških popisov smo naredili preglednico 1, ki smo jo ure­dili s pomočjo programskega paketa SYN-TAX (Poda­ni 2001). Najprej smo kombinirane vrednosti zastira­nja in pogostnosti pretvorili v numerične vrednosti (1– 9) po van der Maarel (1979) in popise primerjali z metodo hierarhične klasifikacije “(unweighted) ave­rage linkage method” – UPGMA. Pri tem smo upora­bljali Wishartov koeficient podobnosti (Wishart’s si­milarity ratio). Nomenklaturni vir za imena prapro­tnic in semenk je Mala flora Slovenije (Martinčič & al. 2007), razen za vrste Nigritella widderi Teppner & E. Klein, Nigritella bicolor W. Foelsche in Nigritella ar­chiducis-joannis Teppner & E. Klein, za mahove pa Martinčič (2003). Pri določanju višjih rastlin smo si pomagali tudi z avstrijsko ekskurzijsko floro (Fischer & al. 2008). Nomenklaturna vira za imena rastlinskih sintaksonov sta Theurillat (2004) in Grabherr & Mucina (1993). Podatke o podnebnih razmerah in ge­ološki zgradbi povzemamo po Erhartič (2012). 2.1 Ekološka oznaka raziskovanega območja Geološka podlaga na nahajališču Janezove murke v Dolini Triglavskih jezer južno od Jezera v Ledvicah je jurski apnenec, ponekod s primesjo laporovca, na dru­gem nahajališču pod Kopico pa prevladuje ledeniško gradivo in grušč iz zgornjetriasnih apnencev. Podnebje je gorsko, s povprečno letno temperaturo okoli 2 °C in s povprečno letno množino padavin okoli 3000 mm, vsaj polovica jih pade v obliki snega, ki tla pokriva 150 do 200 dni. Gozd je v Dolini segal vse do nadmorske višine okoli 1900 m in vrzelasto macesnovje (Rhodot­hamno-Laricetum) s starimi drevesi macesna najdemo na tej nadmorski višini še zdaj, nad Jezerom v Led­vicah. 3 REZULTATI IN RAZPRAVA 3.1 Pregled zdaj znanih nahajališč vrste Nigri­tella archiducis-joannis v Sloveniji 9648/4 (UTM 33TVM03) Slovenija: Gorenjska, Ju­lijske Alpe, Triglavsko pogorje, Dolina Triglavskih jezer, južno od Jezera v Ledvicah, nad potjo proti Koči pri Sedmerih jezerih, 1830 m do 1835 m nm. v., alpin­sko travišče, popisi št. 2 do 5 v preglednici 1. Leg. F. Poljšak, 31. 7. 2004, avtorjeve fotografije; leg. & det. B. Zupan, Peter in Polona Strgar, 16. 7. 2014 in B. Zupan in I. Dakskobler, 11. 8. 2014, herbarij LJS in fotografije avtorjev. 9648/4 (UTM 33TVM03) Slovenija: Gorenjska, Ju­lijske Alpe, Triglavsko pogorje, Dolina Triglavskih jezer, pri Utah, kotanja pod Kopico, 1810 m nm. v., al­pinsko travišče na ledeniškem gradivu, popis št. 1 v preglednici 1. Leg. & det. B. Zupan in I. Dakskobler, 11. 8. 2014, herbarij LJS in fotografije avtorjev. 9649/1 (UTM 33TVM13) Slovenija: Gorenjska, Ju­lijske Alpe, Triglavsko pogorje, Viševnik nad Pokljuko, 1965 m nm. v., alpinsko travišča na vzhodnem vršnem delu gore. Leg. & det. B. Dolinar, 14. 7. 2013, herbarij LJS in avtorjeve fotografije. V zgodnjem poletju 2015 smo Branko Zupan, Brane Anderle, Blaž Blažič in Igor Dakskobler potrdili in popisali nahajališče pod Viševnikom, Peter in Polo­na Strgar pa sta pri Utah v Dolini Triglavskih jezer našla še eno novo nahajališče. 3.2 Fitocenološka oznaka rastišč vrste Nigritella archiducis-joannis v Dolini Triglavskih jezer Prevladujoči vrsti v alpinskih traviščih, v katerih v Dolini Triglavskih jezer uspeva vrsta Nigritella archidu­cis-joannis, sta Sesleria caerulea subsp. calcaria in Carex sempervirens. Zato te sestoje uvrščamo v asociacijo Ra­nunculo hybridi-Caricetum sempervirentis in v jugo­vzhodnoalpsko zvezo Caricion austroalpinae. Sestoj pod Kopico je bolj inicialen, na plitvih kamnitih tleh, kar nakazujejo nekatere vrste, na primer Anemone bal­densis, Hieracium bifidum, Valeriana saxatilis, z večjo pokrovnostjo kot na drugih ploskvah tudi vrsta Dryas octopetala. Sestoji južno od Jezera v Ledvicah so vrstno bogatejši, v njih je tudi nekaj bolj vlagoljubnih vrst, na primer Carex ferruginea, Soldanella alpina in Tofieldia calyculata in tudi nekaj bolj kisloljubnih vrst, na primer Festuca nigrescens in Meum athamanticum, ki morda kažejo na manjšo primes laporovca v tleh. V splošnem je vrstna sestava na nahajališčih Janezove murke v Dolini Triglavskih jezer precej podobna vrstni sestavi alpin­skih travišč, v katerih v Triglavskem pogorju raste Wid­derjeva murka (Nigritella widderi) – primerjaj Daksko­bler & al. (2012). Med vrstami, ki jih kot najpogostejše spremljevalke Janezove murke navajajo na nahajališčih v Avstriji (Teppner & Klein 1985, Staudinger & al. 2009), so tudi take, ki rastejo na opisanih nahajališčih v Julijskih Alpah: Sesleria caerulea subsp. calcaria (= S. albicans), Carex semprevirens, Achillea clavenae, Anthyl­lis vulneraria subsp. alpestris, Galium anisophyllum, Helianthemum alpestre, Juncus monanthos in Phyteuma orbiculare, nismo pa popisali vrst Festuca norica, F. pu­mila in Carduus defloratus. Grabherr & al. (1993: 414) so vrsti Nigritella widderi in N. archiducis-joannis uvr­stili med značilnice zveze Seslerion coeruleae (= Sesleri­on variae), ki združuje subalpinsko-alpinska travišča na karbonatni podlagi Severnih Alp. Floristično precej podobna travišča v Jugovzhodnih Alpah uvrščamo v zvezo Caricion austroalpinae. Med njenimi značilnica­mi, ki smo jih našli v preučenih sestojih, so vrste Laser­pitium peucedanoides, Heracleum austriacum subsp. si­ifolium in Linum julicum. Nedvomno so rastišča Jane­zove murke v Severovzhodnih Alpah ekološko (in flori­stično) precej podobna nahajališčem te murke v Jugo­vzhodnih Alpah (isti višinski pas, okoli 1800 m do 2000 m nm. v., podobna geološka podlaga, podoben tip tra­višča, podobna vrstna sestava z nekaj fitogeografskimi posebnostmi). Glede na pojavljanje vrst Nigritella wid­deri in N. archiducis-joannis tako v Severovzhodnih kot v Jugovzhodnih Alpah ugotavljamo, da to nista najbolj­ši diagnostični vrsti za razlikovanje združb iz zvez Se­slerion variae in Caricion austroalpinae, temveč s svojo prisotnostjo povezujeta ekološko in floristično precej podobna travišča Severovzhodnih in Jugovzhodnih Alp. Morda je glede na podobnost teh travišč umestna njihova obravnava znotraj iste zveze Seslerion variae in dveh podzvez Caricenion austroalpinae in Seslerienion variae, kot predlaga Theurillat (2004). 4 ZAKLJUČKI Nigritella archiducis-joannis je apomiktični in tetra­ploidni takson iz skupine rdeče murke (Nigritella mi­niata = N. rubra agg.). Do nedavna je to bil endemit Severnih oz. Severovzhodnih Alp. Z odkritjem naha­jališča v Julijskih Alpah (Zelesny 2008), ki smo ga potrdili in našli še nova, mogoča pa so tudi v Kara­vankah (Foelsche 2011, Dolinar, in litt.), se je nje­gova znana razširjenost precej povečala, podobno kot se je pred nekaj leti precej povečala vednost o arealu vrste Nigritella widderi (Dakskobler & al. 2012). Ra­stišča obeh murk so si v Severovzhodnih in Jugovzho­dnih Alpah precej podobna, tako po ekoloških značil­nostih kot po vrstni sestavi, zato obe našteti vrsti ne razlikujeta severnoalpske zveze Seslerion variae na­sproti jugovzhodnoalpski zvezi Caricion austroalpi­nae, temveč njune združbe na nek način povezujeta. Populacija vrste Nigritella archiducis-joannis v Slove­niji je zelo majhna in jo na zdaj znanih nahajališčih v Triglavskem pogorju ocenjujemo na manj kot 30 pri­merkov. Zaradi te redkosti je potencialno zelo ogrože­na (kategorija R), podobno velja tudi za Avstrijo (Sta­udinger & al. 2009: 177). Čeprav so do zdaj znana nahajališča v bližini zelo obiskanih planinskih poti (velja tako za Triglavska jezera kot za Viševnik) meni­mo, da jo turistični obisk za zdaj ne ogroža. Ta murka cveti konec junija in začetek julija, če je s snegom bo­gata zima tudi pozneje, torej v času, ko se planinska sezona šele začenja. 5 SUMMARY Nigritella archiducis-joannis is an apomictic and tetra­ploid taxon from the group of red vanilla orchids (Ni­gritella miniata = N. rubra agg.). Until recently it was considered an endemic species of the Northeastern Alps. The discovery of the locality in the Julian Alps (Zelesny 2008), which we confirmed and also found several new ones while establishing that it had poten­tial localities also in the Karavanke – Foelsche 2011, Dolinar, in litt.), considerably expanded the scope of its known distribution area; a similar situation hap­pened several years ago when we substantially im­proved our knowledge of the distribution area of Nigri­tella widderi (Dakskobler & al. 2012). The sites of both orchids are very similar in the Northeastern and Southeastern Alps, both in terms of their ecological characteristics (altitude, parent material), and species composition (Table 1), so they cannot differentiate the northern-Alpine alliance Seslerion variae against the southeastern-Alpine alliance Caricion austroalpinae; instead, we can say that they in a certain way connect their communities. Given the similarity of these grass­lands they could be discussed within the same alliance Seslerion variae and within two suballiances Cariceni­on austroalpinae and Seslerienion variae, as proposed by Theurillat (2004). The population of Nigritella archiducis-joannis in Slovenia is very small and is currently estimated to comprise fewer than 30 specimens on the localities known so far in the Triglav Mountains. In view of this fact it is potentially highly endangered (category R, rare) and the situation is similar in Austria (Staudin­ger & al. 2009: 177). Although its localities known so far are in the vicinity of highly frequented mountain trails (both around Triglav Lakes and Mt. Viševnik) we do not think it is currently threatened by tourists. This orchid blossoms quite early (at the end of June, the beginning of July, in snow-rich winters even later), at the time when the hiking season is only just begin­ning, so it is normally very inconspicuous outside its peak flowering season. ZAHVALA – ACKNOWLEDGEMENTS Pri pripravi tega članka so nam z nasveti dragoceno pomagali prof. Wolfram Foelsche, dr. Helmuth Zele­sny in Janez Mihael Kocjan. Pri terenskem delu nas je nekajkrat spremljala Polona Strgar. Angleški prevod izvlečka in povzetka Andreja Šalamon Verbič. 6. LITERATURA–REFERENCES Aeschimann, D., Lauber, K, , Moser, D. M. & Theurillat, J-P., 2004: Flora alpina. Bd. 2: Gentianaceae–Orchi­daceae. Haupt Verlag, Bern, Stuttgart, Wien. Braun-Blanquet, J., 1964: Pflanzensoziologie. Grundzüge der Vegetationskunde. 3. Auf., Springer Verlag, Wien–New York. Dakskobler, I., B. Dolinar, B. Zupan, R, Iskra, P. Strgar & A. Trnkoczy, 2012: Nigritella widderi Teppner & E. Klein, a new species in the flora of Slovenia. Folia biologica et geologica (Ljubljana) 53 (1–2): 25–43. Delforge, P., 2006: Orchids of Europe, North Africa and the Middle East. Timber Press London. Dolinar, B., 2015: Kukavičevke v Sloveniji. Pipinova knjiga, Podsmreka. Erhartič, B., 2012: Geomorfološka dediščina v Dolini Triglavskih jezer (Geografija Slovenije, 23). Založba ZRC, Ljubljana. Fischer M. A., W. Adler & K. Oswald, 2008: Exkursionsflora von Österreich, Liechtenstein und Südtirol. Land Oberösterreich, Biologiezentrum der OÖ Landesmuseen, Linz. Foelsche, W., 2011: Das Erzherzog-Johann Kohlröschen, Nigritella archiducis-joannis Teppner & E. Klein, und die Kohlröschen-Arten Österreichs. Joannea Botanik (Graz) 9: 61–95. Grabherr, G. & L. Mucina (eds.), 1993: Die Pflanzengesellschaften Österreichs. Teil II: Natürliche waldfreie Vege­tation. Gustav Fischer Verlag, Jena - Stuttgart - New York. Grabherr, G., J. Greimler & L. Mucina, 1993: Seslerietea albicantis. In: Grabherr, G. & L. Mucina (eds.): Die Pflanzengesellschaften Österreichs. Teil II: Natürliche waldfreie Vegetation, Gustav Fischer Verlag, Jena - Stutt­gart - New York, pp. 402–446. Kretzschmar, H., 2008: Die Orchideen Deutschlands und angrenzender Länder. Quelle & Meyer Verlag GmbH & Co., Wiebelsheim. Maarel, van der E., 1979: Transformation of cover-abundance values in phytosociology and its effects on communi­ty similarity. Vegetatio 39 (2): 97–114. Martinčič, A., 2003: Seznam listnatih mahov (Bryopsida) Slovenije. Hacquetia (Ljubljana) 2 (1): 91–166. Martinčič, A., T. Wraber, N. Jogan, A. Podobnik, B. Turk, B. Vreš, V. Ravnik, B. Frajman, S. Strgulc Krajšek, B. Trčak, T. Bačič, M. A. Fischer, K. Eler & B. Surina, 2007: Mala flora Slovenije. Ključ za dolo­čanje praprotnic in semenk. Četrta, dopolnjena in spremenjena izdaja. Tehniška založba Slovenije, Ljubljana. Podani, J., 2001: SYN-TAX 2000. Computer Programs for Data Analysis in Ecology and Systematics. User's Manual, Budapest. Ravnik, V., 2002: Orhideje Slovenije. Tehniška založba Slovenije, Ljubljana. Redl, K., 1999: Wildwachsende Orchideen in Ostereich, 2.Auflage, Kurt Redel Eigenverlag, Druckerei Theiss GmbH, Wolfsberg. Seliškar, T., B. Vreš & A. Seliškar, 2003: FloVegSi 2.0. Računalniški program za urejanje in analizo bioloških podatkov. Biološki inštitut ZRC SAZU, Ljubljana. Staudinger, M., O. Stöhr, F. Essl, L. Schratt-Ehrendorfer, H. Niklfeld & W. Gutermann, 2009: Gefäs­spflanzen. In: Rabitsch, W. & F. Essl, (eds.): Endemiten – Kostbarkeiten in Österreichs Pflanzen- und Tierwelt. Naturwissenschaftlicher Verein für Kärnten und Umweltbundesamt GmbH, Klagenfurt und Wien, pp. 64–267. Teppner, H., 2008: Kohlröschen / Nigritella. In: Fischer M. et al. (eds.): Exkursionsflora von Österreich, ­Liechtenstein und Südtirol. Land Oberösterreich, Biologiezentrum der OÖ Landesmuseen (Linz), pp. 1058– 1060. Teppner, H. & E. Klein, 1985: Karyologie und Fortpflanzungsmodus von Nigritella (Orchidaceae-Orchideae) inkl. N. archiducis-joannis spec. nov. und zweier Neukombinationen. Phyton (Horn, Austria) 52 (1): 147–176. Theurillat, J.-P., 2004: Pflanzensoziologisches System. In: Aeschimann, D., K. Lauber, D. M. Moser & J.-P. Theurillat: Flora alpina 3: Register. Haupt Verlag, Bern, Stuttgart, Wien, pp. 301–313. Zelesny, H., 2008: Nigritella rubra subsp. archiducis-joannis in Slowenien und Bemerkungen zu Nigritella rubra. Journal Europäischer Orchideen 40 (3): 587–598. 1 Biološki inštitut Jovana Hadžija ZRC SAZU, Regijska raziskovalna enota Tolmin, Brunov drevored 13,5220 Tolmin in Uni­verza v Ljubljani, Biotehniška fakulteta, Oddelek za gozdarstvo in obnovljive gozdne vire, Večna pot 83, 1000 Ljubljana igor.dakskobler@zrc-sazu.si 2 Bizjanova 21, SI-1107 Ljubljana, branko.dolinar@telemach.net, http://www.orhideje.si 3 Savica 6, SI-4264 Bohinjska Bistrica 4 Selce 23, SI-6257 Pivka, florijan.poljsak@siol.com 5 Zoisova 11, SI-4264 Bohinjska Bistrica, peter.strgar@petersfoto.si, http://www.petersfoto.si Slika 1: Razširjenost vrste Nigritella archiducis-joannis v Sloveniji Figure 1: Distribution of Nigritella archiducis-joannis in Slovenia Preglednica 1: (Sub)alpinska travišča z vrsto Nigritella archiducis-joannis v Dolini Triglavskih jezer Table 1: (Sub)alpine grasslands with Nigritella archiducis-joannis in the Triglav Lake Valley Zaporedna številka popisa (Number of relevé) 1 2 3 4 5 Številka popisa v podatkovni bazi (Database number of relevé ) 253930 253922 253923 253925 253924 Nadmorska višina v m (Altitude in m) 1810 1835 1835 1832 1833 Lega (Aspect) S E SE SE SE Nagib v stopinjah (Slope in degrees) 10 20 20 15 20 Matična podlaga (Parent material ) A A A A A Tla (Soil ) R R R R R Zastiranje zeliščne plasti v % (Cover of herb layer in %): E1 85 90 80 90 95 Kamnitost v % (Stoniness in %) 15 10 20 10 5 Velikost popisne ploskve (Relevé area) m2 5 10 10 10 20 Število vrst (Number of species) 29 40 39 32 38 Date of taking relevé (Datum popisa) 8/11/2014 8/11/2014 8/11/2014 8/11/2014 8/11/2014 Nahajališče (Locality) Triglavska jezera - Ute Triglavska jezera -Ledvica Triglavska jezera -Ledvica Triglavska jezera -Ledvica Triglavska jezera -Ledvica Kvadrant (Quadrant) 9648/4 9648/4 9648/4 9648/4 9648/4 Koordinata (Coordinate) GK Y (D-48) m 406584 406570 406581 406584 406588 Koordinata (Coordinate) GK X (D-48) m 5132731 5133355 5133370 5133355 5133367 CA Caricion austroalpinae Pr. Fr. Laserpitium peucedanoides E1 + 1 1 1 . 4 80 Heracleum austriacum subsp. siifolium E1 . 1 . . . 1 20 Linum julicum E1 + . . . . 1 20 CFr Caricion firmae Helianthemum alpestre E1 + . + . . 2 40 CF Caricion ferrugineae Carex ferruginea E1 . + + + . 3 60 ES Elyno-Seslerietea Sesleria caerulea subsp. calcaria E1 3 1 3 4 3 5 100 Carex sempervirens E1 2 3 3 3 4 5 100 Anthyllis vulneraria subsp. alpestris E1 1 3 2 2 1 5 100 Astrantia bavarica E1 1 2 1 1 2 5 100 Bartsia alpina E1 + 1 + + + 5 100 Achillea clavenae E1 + 1 + + + 5 100 Nigritella archiducis-joannis E1 + + + + + 5 100 Thymus praecox subsp. polytrichus E1 + + + 1 . 4 80 Polygonum viviparum E1 . + 1 1 1 4 80 Hieracium villosum E1 . + + + + 4 80 Phyteuma orbiculare E1 + 1 . + . 3 60 Gentiana clusii E1 + . . + + 3 60 Heliosperma alpestre E1 + . + + . 3 60 Lotus alpinus E1 . + + 1 . 3 60 Pulsatilla alpina subsp. austroalpina E1 . + + . + 3 60 Galium anisophyllum E1 . + + . + 3 60 Biscutella laevigata E1 . . + + + 3 60 Hieracium glabratum E1 . . + + + 3 60 Salix alpina E1 . . + + + 3 60 Aster bellidiastrum E1 + . . + . 2 40 Arabis vochinensis E1 . + + . . 2 40 Pedicularis rostrato-capitata E1 . . + + . 2 40 Nigritella bicolor E1 . . + . + 2 40 Gentianella anisodonta E1 . . . + + 2 40 Euphrasia picta E1 + . . . . 1 20 Agrostis alpina E1 + . . . . 1 20 Polygala alpestris E1 + . . . . 1 20 Helianthemum nummularium subsp. grandiflorum E1 . + . . . 1 20 Ranunculus carinthiacus E1 . + . . . 1 20 Juncus monanthos E1 . + . . . 1 20 Betonica alopecuros E1 . . + . . 1 20 Potentilla crantzii E1 . . + . . 1 20 Saussurea pygmaea E1 . . . . + 1 20 Alchemilla exigua E1 . . . . + 1 20 OE Oxytropido-Elynion Dryas octopetala E1 3 2 2 2 2 5 100 Gentiana nivalis E1 + . . . . 1 20 TG Trifolio-Geranietea Laserpitium siler E1 + + + + . 4 80 FB Festuco-Brometea Gymnadenia conopsea E1 . + . + + 3 60 JT Juncetea trifidi Soldanella alpina E1 + + + . + 4 80 Potentilla aurea E1 . + + . + 3 60 Gentiana pannonica E1 . + . . r 2 40 CU Calluno-Ulicetea Meum athamanticum E1 . 1 1 + + 4 80 Coeloglossum viride E1 . + + . + 3 60 Luzula exspectata E1 . . + . + 2 40 PaT Poo alpinae-Trisetalia Campanula scheuchzeri E1 . . + + + 3 60 Festuca nigrescens E1 . + . . + 2 40 Crepis aurea E1 . + . . . 1 20 MA Molinio-Arrhenatheretea Trifolium pratense E1 . + . + + 3 60 Leontodon hispidus E1 . 1 . . + 2 40 CD Caricetalia davallianae Selaginella selaginoides E1 + r r . . 3 60 Parnassia palustris E1 . + + . 1 3 60 Tofieldia calyculata E1 . . + . . 1 20 AC Arabidetalia caeruleae Homogyne discolor E1 + 1 1 1 1 5 100 Trifolium pallescens E1 . + . . + 2 40 TR Thlaspietea rotundifolii Anemone baldensis E1 + . . . . 1 20 Hieracium bifidum E1 + . . . . 1 20 AT Asplenietea trichomanis Valeriana saxatilis E1 + . . . . 1 20 MuA Mulgedio-Aconitetea Peucedanum ostruthium E1 . + . + . 2 40 Veratrum album E1 . + . + r 3 60 Viola biflora E1 . . . . + 1 20 EP Erico-Pinetea Carex ornithopoda E1 + . . . + 2 40 Rhododendron hirsutum E2 . . + . + 2 40 Rhodothamnus chamaecistus E1 + . . . . 1 20 Erica carnea E1 . . + . . 1 20 Pinus mugo E2 . . . + . 1 20 ML Mahovi (Mosses) Tortella sp. E0 . + . . . 1 20 Legenda - Legend A apnenec - Limestone R rendzina - Rendzina Zaporedna številka popisa (Number of relevé) 1 2 3 4 5 Pr. Fr. Slika 2: Nigritella archiducis-joannis v dolini Triglavskih jezer, 31. 7. 2004. Foto F. Poljšak Figure 2: Nigritella archiducis-joannis in the Triglav Lake Valley, 31. 7. 2004. Photo F. Poljšak Slika 3: Nigritella archiducis-joannis v dolini Triglavskih jezer, 16. 7. 2014. Foto P. Strgar Figure 3: Nigritella archiducis-joannis in the Triglav Lake Valley, 16. 7. 2014. Photo P. Strgar Slika 4: Rastišče vrste Nigritella archiducis-joannis v Dolini Triglavskih jezer. Foto I. Dakskobler Figure 4: Site of Nigritella archiducis-joannis in the Triglav Lake Valley. Photo I. Dakskobler IGOR PAUŠIČ: CONFIRMATION OF THE AUSTRIAN VANILLA ORCHID, NIGRITELLA AUSTRIACA (TEPPNER & E. KLEIN) IGOR PAUŠIČ: CONFIRMATION OF THE AUSTRIAN VANILLA ORCHID, NIGRITELLA AUSTRIACA (TEPPNER & E. KLEIN) 116 117 FOLIA BIOLOGICA ET GEOLOGICA 56/1, 115–123, LJUBLJANA 2015 FOLIA BIOLOGICA ET GEOLOGICA 56/1 – 2015 FOLIA BIOLOGICA ET GEOLOGICA 56/1 – 2015 CONFIRMATION OF THE AUSTRIAN VANILLA ORCHID, NIGRITELLA AUSTRIACA (TEPPNER & E. KLEIN) P. DELFORGE (ORCHIDACEAE) A NEW SPECIES IN THE SLOVENIAN FLORA POTRDITEV AVSTRIJSKE MURKE, NIGRITELLA AUSTRIACA (TEPPNER & E. KLEIN) P. DELFORGE (ORCHIDACEAE), NOVE VRSTE V FLORI SLOVENIJE Igor PAUŠIČ1 ABSTRACT A new orchid species, Nigritella austriaca was finally con­firmed for Slovenia. It’s recently discovered locality is on the summit of Mt. Uršlja gora (1699 m), Carinthia, N Slovenia. According to the floral morphometrics and the early flower­ing period of the observed orchids (compared with N. rhelli­cani), the plants examined on Uršlja gora mountain were determined to be Austrian vanilla orchid, Nigritella austria­ca, a new species in the Slovenian flora. Austrian vanilla or­chid, Nigritella austriaca (Teppner & E. Klein) P. Delforge is distributed across the Central and Eastern Alps, Dolomites, Jura, Central Massif and as far as the Pyrenees. Keywords: Nigritella austriaca, floral morphometrics, Mt. Uršlja gora, Slovenia IZVLEČEK V prispevku dokazujemo prisotnost avstrijske murke (Nigri­tella austriaca) v flori Slovenije ter rastišče majhne populaci­je le-te na vrhu Uršlje gore (1966 m) na Koroškem. Opravili smo morfometrične analize cvetnih elementov socvetja z Uršlje gore ter dobljene vrednosti primerjali z vrednostmi morfoloških potez pri podobni vrsti - Rhellikanovi murki (Nigritella rhellicani). Dimenzije cvetnih elementov Rhelli­kanove murke povzemamo po literaturnih virih, ki jih na­vajamo v nadaljevanju. Glede na izmerjene vrednosti in di­menzije analiziranih cvetnih elementov, obliko socvetja ter zgodnji čas cvetenja smo osebke iz populacije na Uršlji gori determinirali kot avstrijsko murko Nigritella austriaca (Te­ppner & E. Klein) P. Delforge, novo vrsto v flori Slovenije. Ključne besede: Nigritella austriaca, morfometrija cvet­nih elementov, Uršlja gora, Slovenija. INTRODUCTION The genus Nigritella L.C.M. Richard (Orchidaceae) in­cludes about a dozen species that are distributed over the mountains of Central and Southern Europe (the Alps, Apennines, Carpathians, Balkans, Pyrenees and the Cantabrian mountains) and Scandinavia (Teppner 1996, Teppner & Klein 1998, Delforge 2001, Del­forge 2006). Only three species from the genus Nigri­tella are mentioned by Ravnik in his survey of Slove­nian vanilla orchids (Ravnik 2002): N. rhellicani Tepp­ner & E. Klein, N. lithopolitanica Ravnik and N. rubra (Wettst.) K. Richt. More recently, occurrences of new taxa of the genus Nigritella in Slovenia were reported (Dakskobler et al. 2012): N. widderi Teppner & E. Klein, N. archiducis-joannis Teppner & E. Klein, N. bi­color W. Foelsche and N. hygrophila W. Foelsche & Heidtke, mostly in the Julian Alps. Nigritella austriaca (synonyms: Gymnadenia au­striaca (Teppner & E.Klein) P. Delforge, Gymnadenia nigra subsp. austriaca (Teppner & E.Klein) Teppner & E. Klein, Nigritella nigra subsp. austriaca Teppner & E. Klein) is distributed across the Central and Eastern Alps, Dolomites Jura, Central Massif in the west as far as the Pyrenees (Delforge 2001 and 2006, Sáez 2004). Mala Flora Slovenije (Jogan et al. 2007) briefly men­tion the species under the synonym Nigritella nigra subsp. austriaca (Teppner & E. Klein) - the authors mention the occurrence of this species as “possible” for Slovenia and that it is distinguished from N. rhellicani by the margins of the bracteae, which don’t bear papil­lae. However, this is not always the case. Other floral morphological characteristics should also be consid­ered. Moreover such specimens are found on Mt. Rod­ica (Julian Alps). Dakskobler et al. (2012) - briefly discuss the occurrence of the species in Slovenia, but again using the same source (Jogan et al. 2007), and claiming it as “possible” for Slovenia. The species is also listed in the official checklist of plant species of Triglav National Park. The photos were made by Špela Novak within the Triglav National Park (mail conver­sation is stored by the author), in the area called Lukn­ja. In fact, those plants were never morphologically or genetically determined to be Nigritella austriaca. The species is included in the web page Slovenske orhideje (Slovene Orchids) (www.orhideje.si, 23.12.2014) by the author Branko Dolinar. According to the Branko Doli­nar the material was never morphologically or geneti­cally determined as Nigritella austriaca. The species (again under the homotypic synonym N. nigra subsp. austriaca) is also mentioned in a recently published monography, Kukavičevke Slovenije (Dolinar 2015) using this, unpublished manuscript as a source. Franc Sušnik (Sušnik 1955) mentiones the presence of the taxon Nigritella nigra in his gradual thesis on the veg­etation of Uršlja gora. Nigritella nigra var. rosea is the former synonym of the species Nigritella lithopolitani­ca, which is actually quite abundant on the summit of Uršlja gora. It’s unclear whether the author was writing about Nigritella lithopolitanica or, maybe, even about Nigritella austriaca in his thesis. According to bota­nists actively involved in research into the genus Nigri­tella (Gymnadenia) in Slovenia, all the material being attributed to Nigritella austriaca that has been mor­phologically (or genetically) analysed in Slovenia to date belongs to the species Nigritella rhellicani. From above, it can be concluded, that no published morpho­logical (or any other) data from Slovenia for the taxon N. austriaca exist, that could prove the determination and existence of this species in any specific locality. So, contrary to expectations, Nigritella austriaca has not yet been confirmed for Slovenia. Mt. Uršlja Gora (1699 m) represents the eastern calcareous outpost of the Karavanke mountain range (Map 1). The origin of the tree line (natural or anthro­pogenic), restricted to the summit area, was already a source of speculations in the past (Gams 1976). The tree line was lowered in the past, owing to pasture ac­tivities in the summit area, but it has been gaining al­titude since the change in traditional alpine extensive agriculture and the abandonment of pastures. Accord­ing to Wraber 1959, the association Adenostylo gla­brae- Piceetum var. geogr. Cardamine trifolia forms the natural vegetation of the summit area above 1500 m. The soil substrate on the summit area of Uršlja gora is acidic; meadows preserved by people for centuries have turned into tall herb overgrown areas. METHODS The area near the summit of Mt. Uršlja gora was sur­veyed during June 2014 in order to study the ecology of Nigritella lithopolitanica, which is abundant in the area. However, two unusually deep red flowering specimens were found within the population of N. lithopolitanica (Fig. 1). After a closer inspection of the summit area, 11 additional plants were found among the numerous flowering Nigritella lithopolitanica specimens. Because of the very low number of deep red flowering speci­mens, only two inflorescences were taken from the lo­cality for further analysis on 27 June 2014. We com­pared the floral morphometrics of the flowers taken from the field with similar taxa- we particularly focused on the differences between the morphologically and physiognomically closely related taxa, N. rhellicani and N. austriaca, which resembled the deep red specimens from Mt. Uršlja gora. Floral traits (macro-morphology) were examined on flowers from the basal 1/3 of the in­florescence and studied under the binocular stereo­scopic microscope Nikon SMZ 1000 (8-80 x). NIS Ele­ments D 4.2 software was used for the measurements. The measured floral dimensions of the Nigritella austriaca from Mt. Uršlja gora (27 flowers analysed) were compared with the macro- morphological traits of N. rhellicani but also with N. austriaca, for both ob­tained from the literature available: Brütsch 2000, Foelsche 2011, 2012 and 2014, Klein & Drescher 1996, Teppner 1990 and 2002, Delforge 2001 and 2006, Wucherpfennig 1999, Timpe & Mrkvicka 1991, Sáez 2004 and Hedrén 2000. In the determina­tion of vascular plants, we followed Martinčič et al. 2007. The occurrence of denticles in the margin of the lowermost bracts of the inflorescence (a characteristic that should be used with caution!), the length of the labellum, sepals, petals and spur, the colour of the flowers and the shape of the inflorescence were used as diagnostic characteristics to distinguish between Ni­gritella austriaca and Nigritella rhellicani, both taxa yet unknown in the area being studied. RESULTS In the close vicinity of the Nigritella austriaca speci­mens, on the summit of Mt. Uršlja gora, the following plant taxa were recorded: Arnica montana, Aster alpi­num, Botrychium lunaria, Briza media, Calluna vulga­ris, Campanula barbata , Carlina acaulis, Coeloglossum viride, Cruciata glabra, Dactylorhiza fuchsii, Daphne mezereum, Geum rivale, Gymnadenia conopsea, Hiera­cium aurantiacum, Listera ovata, Lotus corniculatus, Nigritella lithopolitanica, Phyteuma orbiculare, Pinus mugo, Plantago media, Poa alpina, Polygala alpestris, Polygonum viviparum, Potentilla erecta, Rhododendron hirsutum, Tofieldia calyculata, Traunsteinera globosa, Trifolium montanum, Trollius europaeus, Veratrum album, Vaccinium myrtillus and Vaccinium vitis-idaea. Plant height of the 13 Austrian vanilla orchid spec­imens (measured in the field) varies between 11 and 17 cm (average 14.4 cm). The inflorescence of all speci­mens (observed in the field) was (hemi) spherical, even in the later stages of flowering (Figure 1B), obviously wider than long, with relatively large, wide- open, deep reddish flowers. Floral elements were closely analysed (from the lower 1/3 of the inflorescences, 27 flowers). The labellum was distinctly open, slightly narrow in its basal third, 7.5-8.7 mm long, (average 8.1 mm), 5.0- 5.5 mm wide (average 5.1 mm), sepals 7.5-8.1 mm (average 7.8 mm), petals 6.7-8.0 mm (average 6.8 mm), only slightly shorter and narrower than sepals. The spur was rather short: 0.8-1.2 mm (average 1.1 mm). On av­erage, all the floral traits measured (except the spur) are longer than those mentioned in the literature in the case of N. rhellicani (Table 1). Closer inspection of the bracts revealed no denticles (marginal epidermal cells elongated to cylindrical papillae). The lowermost bracts of the inflorescence were without papillae (which are typical of N. rhellicani), and smooth or sometimes with only some marginal epidermal cells somewhat elongated in the margin of the lowermost bracts. Because of the colour, the dimensions of the measured floral elements (especially the length of the labellum, sepals and petals), which are longer on aver­age (and wider) in the case of N. austriaca (compared to N. rhellicani), and owing to the different shape of the inflorescence, this taxon exhibits characteristics dif­ferent from those of N. rhellicani even in the field. Thus, based on a close comparison of the studied plant material, we conclude that the deep red flowered va­nilla orchid specimens from Mt. Uršlja gora do belong to Nigritella austriaca. Table 1 gives the most typical morphomertic differences (some of them easily observable even in the field) be­tween N. austriaca and N. rhellicani, differences that are crucial in our opinion: Nigritella austriaca Nigritella austriaca, Mt. Uršlja gora Nigritella rhellicani Flowering time VI-VIII, at the same time as Nigritella rubra in the same habitat Specimens were observed in full bloom on 27.6.2014. So the flowering started at least a week earlier. The same flowering phenophase as Nigritella lithopolitanica VI-VIII, at least 7-10 days later than Nigritella austri­aca in the same habitat Inflorescence shape Hemi- spherical, even in later stages of flowering ob­viously wider than long Hemi- spherical, even in later stages of flowering ob­viously wider than long At first conical, becoming sub-cylindrical at the end of flowering (longer than wide) Colour Flowers deep reddish (rarely brown) to brownish purple Flowers deep reddish Flowers dark reddish brown (very rarely red) to brown­ish (almost black), pink, yel­lowish, orange or white Floral dimensions Large flowers, Labellum length: (6.9-)7.5- 9.8 mm Sepals: 6.3- 8.4 (-9.4) mm Petals: 6.0- 7.5 (-8.2) mm Large flowers, Labellum length: 7.5- 8.7 mm Sepals: 7.5- 8.1 mm Petals: 6.7- 8.0 mm Small flowers, Labellum length: (4.5-) 5.0- 7.7 mm Sepals: (3.8-) 4.2 (-7.4) mm Petals: (3.5-) 4.0- 5.6 (-6.1) mm Bracts Bracts glabrous or rarely the 1-5 lower bracts slight­ly papillose (about 10% of them) Bracts without papillae and smooth or sometimes with only some marginal epider­mal cells somewhat elon­gated in the margin of the lowermost bracts Lower bracts about equal­ling flowers, with papil­lae (up to 0.1 mm long) on edges Spur length 0.9-1.3 mm 0.8-1.2 mm 1.1-1.7 mm Reproduction apomictic sexual Ploidy level 2n=4x=80 2n=2x=40 Distribution Alps, Jura, Central Massif and the Pyrenees The summit area of Mt. Uršlja gora Alps, Jura, Balkans Tab. 1. Floral macro- morphometric and other differences between Nigritella austriaca and Nigritella rhellicani. Tabela 1. Makro- morfometrične in druge razlike med vrstama Nigritella austriaca in Nigritella rhellicani. DISCUSSION AND CONCLUSIONS The potential for occurrence of Nigritella austriaca in Slovenia had already been the subject of speculation in the past (Jogan 2007, Dakskobler et al. 2012). The recently discovered locality on the summit of Mt. Uršlja gora in Carinthia represents the first published record for Slovenia, supported with the morphometric measurements. The location of a small population on the summit of Mt. Uršlja gora is possibly one of the south-easternmost in the species’ range. The locality is threatened by the rapid succession of herbaceous per­ennials and Pinus mugo. Because of the poorly known distribution in Slovenia, this taxon definitely belongs to our Red List as a vulnerable taxon (V). The reason for this lack of data for Slovenia is mainly due to mor­phological similarity with the N. rhellicani, which is, especially in the Julian Alps and the western Kara­vanke mountain range, quite abundant, but which be­comes extremely rare in the eastern part of the Kara­vanke mountain range (Mt. Košuta being its eastern­most known limit in Slovenia). Closer inspection of the inflorescence and floral elements, on the other hand, reveals quite sufficient and reliable morphometrical dissimilarities between these two taxa. The occurrence of this taxon in Slovenia is not surprising, and it is likely to have more localities elsewhere in the northern Slovenian territory. Specimens of N. austriaca on Mt. Uršlja gora flower at least 7-10 days before N. rhellicani on its closest known locality on the slopes of Mt. Košuta in Karavanke. The closest known N. austriaca locality lies on the slopes of Mt. Obir (Hochobir, Aus­tria, Carinthia). The determination key for the Slove­nian taxa from the genus Nigritella (Jogan 2007) is not sufficiently sophisticated or up-to-date and must be upgraded in its next edition, especially since new taxa have been described for the Slovenian territory, includ­ing Nigritella austriaca. POVZETEK 5.1 Uvod Rod Nigritella L. C. M. Richard (Orchidaceae) obsega približno ducat vrst in podvrst (Teppner 1996, Te­ppner & Klein 1998). Vsi taksoni se pojavljajo v mon­tanskem in alpinskem pasu v gorovjih Srednje in Južne Evrope (Alpe, Apenini, Karpati, Pireneji, Kantabrija) ter v Skandinaviji (Teppner 1996, Teppner & Klein 1998, Delforge 2001, Delforge 2006). Ravnik (Rav­nik 2002) navaja v svoji monografiji Kukavičevke Slo­venije za območje Slovenije le tri vrste iz roda Nigritel­la: N. rhellicani Teppner & E. Klein, N. lithopolitanica Ravnik in N. rubra (Wettst.) K. Richt. V zadnjih letih dokazujejo avtorji pojavljanje novih vrst iz rodu Nigri­tella na območju Slovenije (Dakskobler s sod. 2012): N. widderi Teppner & E. Klein, N. archiducis-joannis Teppner & E. Klein, N. bicolor W. Foelsche in N. hy­grophila W. Foelsche & Heidtke, večinoma na območju Julijskih Alp. Areal razširjenosti avstrijske murke - Nigritella austriaca (sinonimi: Gymnadenia austriaca (Teppner & E.Klein) P. Delforge, Gymnadenia nigra subsp. au­striaca (Teppner & E.Klein) Teppner & E. Klein, Nigri­tella nigra subsp. austriaca Teppner & E. Klein) - obse­ga Osrednje in Vzhodne Alpe in Dolomite. Vrsto sre­čamo v Juri in Centralnem masivu. Na vzhod pa sega areal razširjenosti vrste do Pirenejev (Delforge 2001 in 2006, Sáez 2004). Mala flora Slovenije (Jogan s sod. 2007) omenja vrsto s sinonimom Nigritella nigra subsp. austriaca (Teppner & E. Klein). Navaja tudi možno oz. verjetno pojavljanje te vrste v Sloveniji. Po Mali flori Slovenije (2007) se avstrijska murka razlikuje od Rhel­likanove murke po robovih spodnjih braktej, ki niso papilozne (upoštevanje tega znaka kot dominantnega kriterija je pomanjkljivo, glede na dejstvo, da lahko opazujemo redke primerke Rhellikanove murke, pri katerih rob braktej ni izrazito papilozen). Tako je po­trebno pri zanesljivi morfološki/morfometrični deter­minaciji avstrijske murke upoštevati še druge znake cvetov in socvetja kot celote. Dakskobler s sod. (2012) prav tako omenja vrsto ter njeno “verjetno” pojavljanje na območju Slovenije, a se ponovno sklicuje na isti vir (Jogan s sod. 2007). Vrsta je navedena v seznamu vrst Triglavskega narodnega parka. Avtorica fotografij je Špela Novak, ki je avstrijski murki podobne rastline fotografirala na območju Luknje v Triglavovi skupini. Za fotografirane primerke pa ni nobenih morfometrič­nih podatkov, tudi genetskih raziskav ne, ki bi lahko potrdile prisotnost avstrijske murke v visokogorju v bližini Triglava oz. Slovenije. Tudi na spletni strani Slovenske orhideje (www.orhideje.si, 23.12.2014) avtor­ja Braneta Dolinarja je vrsta navedena. Avtor tudi teh fotografiranih primerkov v preteklosti ni nikoli mor­fometrično ovrednotil ali poslal delov rastlin na genet­ske raziskave. Vrsta (avtor je uporabil sinonim N. nigra subsp. austriaca) je vključena v monografijo Kukavi­čevke Slovenije (Dolinar 2015), kjer avtor za lokacijo rastišča navaja tudi Uršljo goro, nanašajoč se na ta, ta­krat še ne objavljen rokopis. Franc Sušnik (Sušnik 1955) omenja v svojem diplomskem delu o vegetaciji in flori Uršlje gore tudi takson Nigritella nigra. Nigritella nigra var. rosea je star sinonim za kamniško murko (Nigritella lithopolitanica), ki je danes pogosta na ovr­šnem predelu Uršlje gore. Ni povsem jasno, ali je avtor v svojem delu navajal prisotnost vrste Nigritella litho­politanica ali pa je morda res naletel na avstrijsko murko. Strnemo lahko, da pripadajo primerki, ki so jih nekateri avtorji v preteklosti uvrščali oz. poimenovali avstrijska murka, v resnici vrsti Rhellikanova murka (Nigritella rhellicani), ki je na prvi pogled avstrijski murki sicer res zelo podobna. Tako nimamo nobenih morfometričnih ali pa genetskih dokazov o obstoju av­strijske murke na območju Slovenije do sedaj, torej ob­stoj vrste v Sloveniji še ni bil potrjen. Geološko predstavlja Uršlja gora (1699 m) enega najbolj vzhodno ležečih apnenčastih osamelcev, od­rastkov Karavank (karta 1). O pojavu gozdne meje na samem vrhu gore (naravna ali pa antropogena) je bilo v preteklosti že veliko napisanega (Gams 1976). Goz­dna meja je segala v preteklosti precej nižje kot danes, kar je bilo posledica paše na ovršnem predelu gore. Danes se travišča na vrhu gore spet zaraščajo, pred­vsem zaradi odsotnosti paše ali redne košnje. Na vrhu Uršlje gore se pojavlja združba Adenostylo glabrae - Pi­ceetum var. geogr. Cardamine trifolia (Wraber 1959) in predstavlja naravno obliko vegetacije vrha gore nad sklenjenim območjem gozda (nad 1500 m). Prsti na vrhu so kisle, travnike in pašnike, ki so bili v preteklo­sti aktivni, danes zarašča rušje (Pinus mugo) in zelnate trajnice. 5.2 Metode Na vrhu Uršlje gore smo junija 2014 analizirali ekolo­ške značilnosti rastišča kamniške murke (Nigritella li­thopolitanica), katere populacija je na Uršlji gori šte­vilčna. Pozorni smo postali na dva škrlatno rdeča pri­merka (fotografija 1). Po natančnejšem pregledu ob­močja vrha smo zabeležili še dodatnih 11 škrlatno rdečih primerkov murke, skupno torej trinajst. 27. 6. 2014 smo odvzeli dve kompletni socvetji. Primerjali smo morfometrične vrednosti izbranih cvetnih ele­mentov z vrednostmi, ki jih literatura navaja za vrsti N. rhellicani in N. austriaca. Na oddelku za biologijo FNM UM smo opravili morfometrijo cvetnih elemen­tov 27 cvetov iz spodnje tretjine socvetja. Meritve smo opravili z uporabo stereolupe Nikon SMZ 1000 (8-80 x) ter programskega paketa NIS Elements D 4.2. Primerjali smo nekatere morfometrične vrednosti cvetnih elementov avstrijske murke z Uršlje gore (27 cvetov) z znanimi vrednostmi, ki jih literatura podaja za avstrijsko in Rhellikanovo murko: Brütsch 2000, Foelsche 2011 in 2012, Klein & Drescher 1996, Te­ppner 1990 in 2002, Delforge 2001 in 2006, Wu­cherpfennig 1999, Timpe & Mrkvicka 1991, Sáez 2004 in Hedrén 2000. Prisotnost obeh zgoraj omenje­nih vrst na Uršlji gori do sedaj še ni bila potrjena. No­menklaturo višjih rastlin na rastišču na vrhu Uršlje gore povzemamo po Mali flori Slovenije (Martinčič s sod. 2007). Med pomembnimi diagnostičnimi, razli­kovalnimi znaki med vrstama N. austriaca in N. rhelli­cani smo obravnavali: odsotnost papil na robovih braktej, dolžino medene ustne, dolžino sepalov, dolži­no petalov ter dolžino ostroge, barvo cvetov ter obliko socvetij - pri primerkih v poznih stadijih cvetenja. 5.3 Rezultati V neposredni okolici primerkov avstrijske murke na ovršnem delu Uršlje gore smo zabeležili naslednje ra­stlinske vrste: Arnica montana, Aster alpinum, Bot­rychium lunaria, Briza media, Calluna vulgaris, Cam­panula barbata , Carlina acaulis, Coeloglossum viride, Cruciata glabra, Dactylorhiza fuchsii, Daphne mezere­um, Geum rivale, Gymnadenia conopsea, Hieracium aurantiacum, Listera ovata, Lotus corniculatus, Nigri­tella lithopolitanica, Phyteuma orbiculare, Pinus mugo, Plantago media, Poa alpina, Polygala alpestris, Polygo­num viviparum, Potentilla erecta, Rhododendron hirsu­tum, Tofieldia calyculata, Traunsteinera globosa, Trifo­lium montanum, Trollius europaeus, Veratrum album, Vaccinium myrtillus and Vaccinium vitis-idaea. Na terenu samem smo izmerili višino socvetij vseh 13 primerkov, ki variira med 11 in 17 cm (povprečna višina znaša 14.4 cm). Vsi primerki (opazovani v po­znih stadijih cvetenja) so imeli izrazito poloblasta so­cvetja, širša kot pa visoka (fotografija 1B). Cvetovi vseh primerkov so bili škrlatno rdeče barve s široko razprti­mi cvetnimi elementi. Analizirali smo dolžino cvetnih elementov: sepalov, petalov, medene ustne ter ostroge na spodnji tretjini dveh socvetij (27 cvetov). Lateralna robova medene ustne sta bila široko razprta. Medena ustna je bila rahlo zožena na spodnji (distalni) tretjini, 7.5-8.7 mm dolga (povprečje znaša 8.1 mm) in 5.0-5.5 mm široka (povprečje znaša 5.1 mm). Sepali merijo v dolžino 7.5-8.1 mm (povprečje znaša 7.8 mm), petali 6.7-8.0 mm (povprečje znaša 6.8 mm). Petalni listi so v povprečju le nekoliko krajši in ožji od sepalnih. Ostro­ga analiziranih cvetov je bila kratka, 0.8-1.2 mm dolga (povprečje znaša 1.1 mm). V povprečju so bile izmerje­ne vrednosti vseh opazovanih cvetnih elementov (z izjemo ostroge) večje, kot jih literatura navaja za vrsto Nigritella rhellicani, ter ustrezajo dimenzijam, ki jih li­terature navaja za vrsto Nigritella austriaca (tabela 1). Opazovali smo rob braktej, ki v nobenem primeru ni bil papilozen. Zaradi izmerjenih vrednosti analizira­nih cvetnih elementov, škrlatno rdeče barve cvetov ter poloblaste oblike socvetij v poznih fazah cvetenja in braktej brez izrazitih papil na robovih ugotavljamo, da pripada populacija škrlatno rdečih murk z vrha Uršlje gore vrsti avstrijska murka (Nigritella austriaca). Tabela 1 prikazuje najpomembnejše morfometrič­ne razlike (nekatere od njih lahko brez težav spremlja­no na terenu samem) med vrstama Nigritella austriaca ter Nigritella rhellicani. Prav tako so podane izmerjene vrednosti spremljanih znakov pri dveh socvetjih z Ur­šlje gore. 5.4 Razprava in zaključki Potencialna prisotnost avstrijske murke v slovenski flori je bila v preteklosti že predmet razprav (Jogan 2007, Dakskobler s sod. 2012). Pričujoč prispevek je prvi, ki s pomočjo morfometrije cvetnih elementov dokazuje prisotnost taksona na območju Slovenije, na vrhu Uršlje gore na Koroškem. Majhna populacija avstrijske murke na Uršlji gori se po do sedaj znanem pojavljanju nahaja na skrajno jugovzhodnem delu celotnega areala razšir­jenosti te vrste. Širše območje je, predvsem zaradi odso­tnosti paše, podvrženo hitri sukcesiji, zaraščanju z zel­natimi trajnicami in grmovnimi vrstami, predvsem z rušjem (Pinus mugo). Zaradi majhnega števila opaženih primerkov na Uršlji gori ter predvsem zaradi do sedaj edinega potrjenega rastišča omenjene vrste sodi avstrij­ska murka nedvomno na naš Rdeči seznam kot ranljiva vrsta (V). Razlog za pomanjkanje podatkov o prisotno­sti vrste v našem alpskem svetu je tudi velika morfolo­ška podobnost z Rhellikanovo murko (Nigritella rhelli­cani), ki je zlasti v Julijskih Alpah in zahodnih Kara­vankah relativno pogosta, a redka v vzhodnem delu Karavank (najbolj vzhodno se vrsta po danes znanih podatkih pojavlja na Košuti). Po drugi strani pa ob na­tančnem morfološkem pregledu lahko obe vrsti, ki ju v prispevku primerjamo, relativno enostavno ločimo. Po­javljanje avstrijske murke na območju Slovenije je bilo pričakovano, morda ima vrsta drugod v našem visoko­gorju celo še več rastišč. Primerki avstrijske murke cve­tijo na Uršlji gori približno 7-10 dni pred Rhellikanovo murko v Karavankah. Najbližja znana populacija av­strijske murke pa se pojavlja na pobočjih Obirja (Ho­chobir) na Avstrijskem Koroškem. Ključ za določevanje murk (Jogan 2007) bo v prihodnje potrebno ažurirati, saj je bilo na območju Slovenije že dokazano pojavljanje novih vrst murk, sedaj tudi avstrijske murke. ACKNOWLEDGEMENTS Sincere thanks to prof. Mitja Kaligarič for the correc­tions of the manuscript. Thanks to botanists actively involved in research into the genus Nigritella (Gymna­denia) in Slovenia: Matej Lipovšek, Branko Dolinar and Mihael Kocjan for a constructive debate on the oc­currence of Nigritella austriaca on Mt. Uršlja gora. ZAHVALA Iskreno se zahvaljujem profesorju Mitji Kaligariču za pregled rokopisa. Hvala botanikom, ki se aktivno po­svečajo problematiki pri rodu Nigritella (Gymnadenia): Mateju Lipovšku, Branetu Dolinarju in Mihaelu Ko­cjanu za konstruktivno debato o pojavljanju avstrijske murke na Uršlji gori. REFERENCES-LITERATURA Brütsch, J.P., 2000: Die Gattung Nigritella Rich. Bauhinia 14: 21-32. Dakskobler, I., B. Dolinar, B. Zupan, R. Iskra, P. Strgar & A. Trnkoczy, 2012: Nigritella widderi Teppner & E. Klein, a new species in the flora of Slovenia. Folia Biologica et Geologica (Ljubljana) 53/1-2: 25-43. Delforge, P., 2001: Guide des Orchidées d’Europe, d’Afrique et du Proche Orient. Ed. 2. Delachaux et Niestlé. Lausanne-Paris. Delforge, P., 2006: Orchids of Europe, North Africa and the Middle East. A&C Black Publishers Ltd. (Ljubljana), London: 640 pp. Dolinar, B.: Slovenske orhideje (http://www.orhideje.si/), 23.12.2014. Dolinar, B., 2015: Kukavičevke Slovenije. Pipinova knjiga. Dobrova: 120-121. Foelsche, W., 2011: Das Erzherzog-Johann-Kohlröschen, Nigritella archiducis-joannis Teppner & E. Klein, und die Kohlröschen-Arten Österreichs. Joannea Botanik 9: 61–95. Foelsche, W., 2012: Bestimmungsschlüssel für die Kohlröschen der Ostalpen, Version 10. May 2012. http://www.museum-joanneum.at/upload/file/Bestimmungsschluessel_Nigritella_2012.pdf (7.7.2015) Foelsche, W., 2014: Bestimmungsschlüssel für die Arten und Unterarten der Gattung Nigritella mit Kurz­beschreibungen, Verbreitungsangaben und Abbildungen, Vestion 1, Dezember 2014. http://www.museum-joanneum.at/upload/file/Bestimmungsschluessel_Nigritella_2012.pdf (7.7.2015) Gams, I., 1976: O zgornji gozdni meji na jugovzhodnem Koroškem. Geografski zbornik (Ljubljana) 16: 158-163. Hedrén, M., E. Klein & H. Teppner, 2000: Evolution of Polyploids in the European Orchid genus Nigritella: Evi­dence from Allozyme Data. Phyton 40 (2): 239-275. Jogan, N., 2007: Orchidaceae – kukavičevke. In: Martinčič, A. (ed.): Mala flora Slovenije. Ključ za določanje praprotnic in semenk. Tehniška založba Slovenije, Ljubljana: 756–784. Klein, E. & A. Drescher, 1996: Nigritella nigra (Orchidaceae - Orchideae) im Massif Central (Frankreich). Phyton, 36 (2): 231-250. Martinčič, A. s sod.: 2007. Mala flora Slovenije. Ključ za določanje praprotnic in semenk. Tehniška založba Slovenije, Ljubljana. Ravnik, V., 2002: Orhideje Slovenije. Tehniška založba, Ljubljana. Sáez, L., 2004: The genus Nigritella (Orchidaceae) in the Iberian Peninsula. Anales del Jardín Botánico de Madrid 61(1): 81-90. Sušnik, F., 1955: Vegetacija in flora Uršlje gore. Diplomsko delo. 106 str., Ljubljana. Teppner, H. & E. Klein, 1990: Nigritella rhellicani spec. nova und N. nigra (L.) Rchb. f. s. str. (Orchidaceae - Orchi­deae). Phyton 31 (1): 5-26. Teppner, H. & E. Klein, 1993: Nigritella gabasiana spec. nova, N. nigra subsp. iberica subsp. nova (Orchidaceae­-Orchideae) und deren Embryologie. Phyton 33 (2): 179-322. Teppner, H. & E. Klein, 1998: Etiam atque etiam: Nigritella versus Gymnadenia: Neukombinationen und Gymna­denia dolomitensis spec. nova (Orchidaceae - Orchideae). Phyton 38 (1): 220-225. Teppner, H. & E. Klein, 1990: Nigritella rhellicani spec, nova und N. nigra (L.) RCHB. f. s. str. (Orchidaceae - Or­chideae). Phyton 31 (1): 5-26. Teppner, H. & T. Ster, 1996: Nigritella buschmanniae spec. nova (Orchidaceae-Orchideae) und eine Biographie für Frau Adolfine Buschmann. Phyton 36 (2): 277-294. Teppner, H., 2002: A review of new results in Nigritella (Orchidaceae). Polska Akademia Nauk, Tom XLVI (2), Krakow 2004. Timpe, W. & A. Ch. Mrkvicka, 1991: Zur Unterscheidung von Nigritella nigra (L.) RCHB.fil. subsp. austriaca Tep­pner & Klein und Nigritella rhellicani Teppner & Klein anhand makroskopischer Merkmale. Mitt.Bl. AHO Baden- Württ. 23(3): 449- 466. Wraber, T., 1959: Tipološka razčlenitev gozdne vegetacije zgornje Mežiške doline. Tipkopis, knjižnica Biološkega inštituta J. Hadžija, SAZU, Ljubljana. Wucherpfennig, W., 1999: Die Gattung Nigritella. AHO Bayern articles, Wolfgang Wucherpfennig, Lerchenweg 1, D-85386 Eching. 1 Department of Biology, Faculty of Natural Sciences and Mathematics, University of Maribor, Koroška c. 160, SI-2000 Maribor, Slovenia, igor.pausic@um.si Map 1. Geographic position of Slovenia in Europe and the location of Mt. Uršlja gora, locality of Nigritella austriaca (Teppner & Klein) P. Delforge. Karta 1. Geografski položaj Slovenije v Evropi in lokacija populacije avstrijske murke Nigritella austriaca (Teppner & Klein) P. Delforge na Uršlji gori. B A C Fig. 1. Nigritella austriaca (Teppner & Klein) P. Delforge. A – habitus, B – inflorescence, C– habitat (photos by Igor Paušič, Mt. Uršlja gora, 27.6.2014). Fotografija 1. Avstrijska murka, Nigritella austriaca (Teppner & Klein) P. Delforge. A- habitus, B- socvetje, C-habitat. (Igor Paušič, Uršlja gora, 27.6.2014) MITJA ZUPANČIČ & JOŽE SKUMAVEC: FLORA AND VEGETATION IN POKLJUKA GORGE (JULIAN ALPS, NW SLOVENIA) MITJA ZUPANČIČ & JOŽE SKUMAVEC: FLORA AND VEGETATION IN POKLJUKA GORGE (JULIAN ALPS, NW SLOVENIA) 126 127 FOLIA BIOLOGICA ET GEOLOGICA 56/1, 125–150, LJUBLJANA 2015 FOLIA BIOLOGICA ET GEOLOGICA 56/1 – 2015 FOLIA BIOLOGICA ET GEOLOGICA 56/1 – 2015 FLORA AND VEGETATION IN POKLJUKA GORGE (JULIAN ALPS, NW SLOVENIA) FLORA IN VEGETACIJA POKLJUŠKE SOTESKE (JULIJSKE ALPE, SZ SLOVENIJA) Mitja ZUPANČIČ1 & Jože SKUMAVEC2 ABSTRACT Flora and vegetation in Pokljuka Gorge (Julian Alps, NW Slovenia) The vascular flora and vegetation of Pokljuka Gorge are described. Five forest and one shrub community and 262 taxons of vascular flora were identified. Central European flora predominate, with 158 (60,3%), there are 11 (4,2%) spe­cies of Illyrian flora, 14 (5,3%) protected species and the sub­endemic species Saxifraga burseriana is also present. We classified the forest and shrub associations into Central Eu­ropean phytocenoses, although more widespread southeast European-Illyrian species are present in some. Key words: flora, vegetation, Alpine region of Slovenia, Triglav National Park, Pokljuka Gorge. IZVLEČEK Flora in vegetacija Pokljuške soteske (Julijske Alpe, SZ Slovenija) Opisana je vaskularna flora in vegetacija Pokljuške soteske. Določili smo 5 gozdnih in 1 grmiščno združbo ter 262 taksonov vaskularne flore. Prevladuje srednjeevropska flora s 158 taksoni (60,3 %), ilirske flore je 11 (4,2 %) vrst, zavarovanih vrst je 14 (5,3 %), prisoten je še subendemit Saxifraga burseriana. Gozdne in grmiščno združbo uvrščamo v srednjeevropske fitocenoze, vendar so v nekat­erih prisotne širše razširjene jugovzhodnoevropsko-ilirske vrste. Ključne besede: flora, vegetacija, alpsko območje Slovenije, Triglavski narodni park, Pokljuška soteska. INTRODUCTION Pokljuka Gorge is among the most interesting natural features of Triglav National Park. It is cut into the steep northeast edge of Pokljuka plateau at an altitude of 670 to 800 m. It is the largest fossil gorge in Slovenia, cre­ated many millions of years ago by the waters of Tri­glav glacier. The gorge is for the most part dry today, only during heavy rain and the spring snow melt does water flow in the lower part of the gorge in Ribščica stream, which flows into the Rodovna (Smolej 1982, Ramovš 1986, Skumavec 1995, Skumavec & Skobr­ne 1995). The retreat of Bohinj glacier during the last glaciation in the Würm was important for the today’s form of the area in question. The Radovna river with tributaries had a large amount of water, which had flowed from beneath Radovna glacier and the ice-bound Pokljuka plateau and had great erosive power (ŠIfrer 1983). Flowing along tectonic cracks it also created Pokljuka Gorge, with many interesting natural phenomena, such as Pokljuka Cave, a natural bridge and »vrtci« (garden plots) with flat bottoms in the form of sinkholes. Pokljuka Gorge is traversable from Jela in the northeast of the gorge to the cliff above Srednji vrtec. There is no natural passage between the cliffs; this is only possible in the final, upper part in the fissure be­tween the cliffs. Passage is possible via bridges and steps, called the Galleries. The Galleries were built in 1930 and at that time were known as the Galleries of Prince Andrew (Karadžordžević). The bridges col­lapsed during the Second World War and were reno­vated in 1982. After the renovation of the Galleries, my colleague and friend Jože Skumavec took me to Pokljuka Gorge. We have visited the gorge many times since the nine­teen eighties, where, more in an amateur than profes­sional manner, we observed the nature of the gorge, above all the flora and vegetation. In the second half of the nineteen eighties, I began planned research of the vegetation of Pokljuka. M. Wraber, Tregubov and Pisk­ernik had researched Pokljuka prior to me. Their stud­ies were published in local scientific reports of Bled Forest Management and in two informative publica­tions, M. Wraber (1960) and Tregubov (1957). At the time of my Pokljuka research, my colleague Skumavec had the idea of a planned study of the flora and vegeta­tion of Pokljuka Gorge. We began the research, which with major breaks, lasted until today. The research path led us along the gorge, from Jela, Kobalove rovte, Stranska soteska, Pokljuka cave, Sred­nji vrtec, Galerije, Veliki vrtec, below the path towards Zatrnik above the gorge, across the transitional or cir­cular path towards Stara Pokljuka or below Pustovo polje, through Pokljuka cave and back to Jela. The area covers almost 2 km of the length of the gorge. We wanted to include and inventory the flora as complete­ly as possible, although we probably did not fully suc­ceed in this and we expect that it is or will be possible to find some species that we overlooked or that were not there at the time of our surveys due to growth or other natural reasons. The forest-shrub vegetation is represented by 5 phytocenological relevés and a phyto­cenological table with three relevés. METHODS The floristic research took place according to estab­lished standard methods, in which we used Mala flora Slovenia (Martinčič et al. 2007), Flora alpina (Ae­schimann et al. 2004), Register flore Slovenije (Trpin & Vreš 1995) and the database FloVegSi (T. Seliškar, Vreš & A. Seliškar 2003) for identifying species. We treated the vegetation according to the standard Cen­tral European (Zurich-Montpellie) method (Braun-Blanquet 1964). We inventoried the flora and vegeta­tion along the mountaineering-tourist path right up to the cliffs that confine the gorge, and in the area above the gorge, which geographically sensibly belongs to it. We visited the research area a number of times from spring to autumn. ECOLOGICAL CHARACTERISTICS OF THE RESEARCH AREA Climatic conditions in Pokljuka Gorge are more or less similar to those that prevail in the Alps. Pokljuka Gorge is a frost area, in which temperature inversions appear in spring and autumn, and the area is in gen­eral colder than the surroundings throughout the year. A humid climate prevails in the gorge, with fresh sum­mers and cold winters. Average annual precipitation is from 1500 to 2000 mm and more. The majority of rainfall, around 60% and perhaps even more, falls dur­ing the vegetation period. On the floor of the gorge, in sheltered positions and rough ground, snow lies late into spring. We estimate average annual air tempera­ture to be as in the Alps, between 3° and 6° C, depend­ing on the configuration of the terrain. Pokljuka Gorge has a uniform geological composi­tion, with Upper Triassic limestone predominating. There is some dolomite of the same age at the start of the path into the gorge. Lithologically, there are several variants of limestone, from massive grey limestone, siliconised or almost solid limestone to dense grey limestone, which often contains ‘bulbs’ of greyer or black chert. (Ramovš 1986). The gorge was gauged out to a depth of 50m by glacial outflows along tectonic cracks. Traces of the action of the glacial water are vis­ible on the overhanging cliffs, natural bridges, in smaller tunnels, Pokljuka cave and on the rubbly, col­lapse ground, from which the disappearing stream Ribščica originates, which flows out into the Radovna. (Ramovš 1986, Skumavec & Skobrne 1995). The soils on this carbonate base are basic eutric brown carbonate soils (calcocambisols), shallow to me­dium deep, in places there are rendzinas. In soils in depressions there are mosaic soils between rendzinas, which are scattered with flint-chert, and lithosols on rocks and boulders. The soils scattered with chert are more or less acidic. There is also greater acidity be­neath the lithosols on limestone rocks and blocks, where a thin layer of raw humus accumulates. In the »garden plots« are distric brown soils with an abun­dant admixture of sandy flint created from the cherts, which are moderately acid to neutral. The mezo- and microclimatic, geological-litholog­ical and soil conditions enable the varied flora and vegetation in Pokljuka Gorge. FLORA The flora and vegetation of the research area are de­fined by their phytogeographic position. The majority of Slovenia belongs to the Euro-Siberian-North Amer­ican floral region. A particularity here is the Illyrian floral province, in which Pokljuka Gorge is classified. There are some southeast European-Illyrian species with narrower or wider distribution in this area. These are: Anemone x pitonii, A. trifolia, Aposeris foetida, Cardamine ennaphyllos, C. trifolia, Galium laevigatum, Helleborus niger, Homogyne sylvestris, Knautia drymeia ssp. drymeia in Lamium orvala. Additional southeast European species are more frequent and more wide­spread in large numbers than elsewhere in Europe, some of which are in Pokljuka Gorge: Cardamine pen­taphyllos, Chaerophyllum hirsutum, Peucedanum au­striacum, Primula vulgaris in Stellaria montana. Irre­spective of POLDINI’s (1991) phytogeographic divi­sion of flora, it is more or less justified to classify as southeast European species, with a hint of Illyrian, the species Aruncus dioicus, Fraxinus ornus, Helleborus odorus, Ostrya carpinifolia and Saxifraga rotundifolia, which also denote the Illyrian floral province. Poklju­ka Gorge is located on the edge of the Julian Alps, which are the eastern part of the southeast limestone Alps, so it is placed in the southeast Alpine floral sec­tor and, because it lies below the Alps, in the subalpine floral subsector and in the Julian Alps-West Kara­vanke-Kamnik Alps district (Zupančič et al. 1987). The already mentioned species Anemone trifolia, Hel­leborus niger and Larix decidua are characteristic of this district. We did not find the characteristic–en­demic species Pedicularis elongata ssp. julica in Poklju­ka Gorge. The sub-endemic species Saxifraga burseria­na is important in Pokljuka Gorge for the mentioned district, which is widespread in the southeast Alpine region. The large settlement of the species Saxifraga burseriana in the cliffs below the entrance to Pokljuka cave is interesting. Even recently we thought (Skuma­vec & Zupančič 2014) that it is almost the only pre­served site. The most recent data according to the FloVegSi database of the Biological Institute ZRC SAZU (T. Seliškar et al. 2003) state a number of loca­tions in the Julian and Savinja Alps and Karavanke. There are 13 protected species. These are: Cephalanthe­ra damasonium, C. rubra, Convallaria majalis, Cycla­men purpurascens, Dactylorchiza maculata ssp. fuchsii, Dianthus hyssopifolius, Epipactis helleborine, Hellebo­rus niger, H. odorus, Huperzia selago, Lycopodium an­notinum, Neottia nidus avis in Primula auricula. Forest and shrub communities supplement or confirm the designation of southeast Alpine phytogeographic posi­tion of Pokljuka Gorge in the context of the Illyrian floral province. If we add to the above diagnostically important analysis for the phytogeographic determination of the research area also an analysis of other species accord­ing to Poldini (1991), it can be seen that psychro­philic Circumboreal, Mediterranean-Montane, Euro­siberian, Paleotemperate, Arctic–Alpine and Eastern Alpine geoelements are present in Pokljuka Gorge, with more than two fifths participation. We conceive Poldini’s definition of Mediterranean-Montane ele­ments as mountain elements under specific Mediter­ranean climatic influences, to which the Julian Alps are subject. These are a quarter of all those recorded. The presence of the enumerated geoelements con­firms the cold climatic conditions. The other major group, with slightly over a third share, are European and Euroasian geoelements, which are generally wide­spread in the European temperate zone. A more de­tailed analysis of geoelements in Pokljuka Gorge is shown in Table 1. The biological form of plants according to Raunki­aer indicates how a plant adapts to the environment in which it lives (thrives) or what sort of life capacity it has to survive the most unfavourable seasons (e.g., winter cold or summer drought). The biological spec­trum, which is the relation between the biological forms, shows the ecological conditions in the area in question (habitat) (M. Wraber 1946). The biological spectrum of Pokljuka Gorge confirms that ecological conditions predominate here that are normal in the temperate belt. (Table 2). Below are stated a list of plants of Pokljuka Gorge based on their affiliation to families. The 262 plant taxons belong to 61 families. Lycopodiaceae Huperzia selago (L.) Mart. Lycopodium annotinum L. Equisetaceae Equisetum sylvaticum L. Hypolepidaceae Pteridium aquilinum (L.) Kuhn Thelypteridaceae Phegopteris connectilis (Michx.) Watt Thelypteris limbosperma (AU.) H. P. Fuchs Aspleniaceae Asplenium ruta-muraria L. Asplenium trichomanes L. ssp. quadrivalens (?) Asplenium trichomanes L. s. lat. Asplenium viride Huds. Phyllitis scolopendrium (L.) Newman Athyriaceae Athyrium filix-femina (L.) Roth Cystopteris fragilis (L.) Bernh. Matteucia struthiopteris (L.) Tod. Aspidiaceae Dryopteris affinis (Löve) Fraser-Jenkis Dryopteris expansa (Presl.) Fraser-Jenkis & Jermy Dryopteris filix-mas (L.) Schott Gymnocarpium dryopteris (L.) Newman Polystichum aculeatum (L.) Roth Polystichum braunii (Spenn.) Fee Polystichum lonchitis (L.) Roth Blechnaceae Blechnum spicant (L.) Roth Polypodiaceae Polypodium vulgare L. Pinaceae Alies alba Miller Larix decidua Miller Picea abies (L.) Karsten Pinus sylvestris L. Aristolochiaceae Asarum europaeum L. ssp. caucasicum (Ducharte) Soó Asarum europaeum L. ssp. europaeum Ranunculaceae Aconitum degenii Gayer ssp. paniculatum (Archang.) Mucher Aconitum lycoctonum L. em. Koelle ssp. ranunculifo­lium (Rchb.) Schinz & Keller Actaea spicata L. Anemone nemorosa L. Anemone trifolia L. Caltha palustris L. Clematis alpina (L.) Mill. Clematis vitalba L. Helleborus niger L. Helleborus odorus Waldst. & Kit. Hepatica nobilis Mill. Ranunculus acris L. ssp. acris Ranunculus lanuginosus L. Thalictrum aquilegiifolium L. Trollius europaeus L. Papaveraceae Chelidonium majus L. Fumariaceae Corydalis cava (L.) Schweiger & Koerte Corydalis solida (L.) Clairv. Caryophyllaceae Dianthus hyssopifolius L. Moehringia muscosa L. Silene dioica (L. em. Mill.) Clairv. Silene nutans (L.) Wibel. s. lat. Stellaria montana Perrat Stellaria nemorum L. Polygonaceae Rumex acetosa L. Rumex alpestris Jacq. Fagaceae Fagus sylvatica L. Quercus petraea (Matt.) Liebl. Quercus robur L. Betulaceae Betula pendula Roth Corylaceae Corylus avellana L. Carpinaceae Ostrya carpinifolia Scop. Juglandaceae Juglans regia L. Ulmaceae Ulmus glabra Huds. Urticaceae Parietaria officinalis L. Urtica dioica L. Grossulariaceae Ribes alpinum L. Ribes uva-crispa L. ssp. laciocarpum Gaud. ex. Monn. Crassulaceae Sedum album L. Saxifragaceae Chrysosplenium alternifolium L. Saxifraga burseriana L. Saxifraga cuneifolia L. Saxifraga rotundifolia L. Rosaceae Alchemilla sp. (?) Aremonia agrimonioides (L.) DC. Aruncus dioicus (Walter) Fernald Filipendula ulmaria (L.) Maxim. s. lat. Fragaria vesca L. Geum urbanum L. Potentilla caulescens L. Potentilla erecta (L.) Raeusch. Rosa pendulia L. Rubus idaeus L. Rubus plicatus Weiche & Nees Rubus saxatilis L. Sorbus aria (L.) Crantz. Sorbus aucuparia L. ssp. aucuparia Fabaceae Genista tinctoria L. Laburnum alpinum (Mill.) Presl. Lathyrus vernus (L.) Bernh. ssp. vernus Lotus corniculatus L. s. lat. Trifolium campestre Schreb Vicia cracca L. Vicia oroboides Wulfen Onagraceae Circaea alpina L. Circea x intermedia Ehrh. Circaea lutetiana L. Epilobium montanum L. Aceraceae Acer campestre L. Acer platanoides L. Acer pseudoplatanus L. Oxalidaceae Oxalis acetosella L. Geraniaceae Geranium phaeum L. s. lat. Geranium robertianum L. Balsaminaceae Impatiens noli-tangere. L. Polygalaceae Polygala chamaebuxus L. Rhamnaceae Rhamnus catharticus L. Rhamnus pumilus Turra Santalaceae Thesium bavarum Schrank Euphorbiaceae Euphorbia amygdaloides L. Euphorbia cyparissias L. Euphorbia dulcis L. ssp. incompta (Cesati) Nyman Mercurialis perennis L. Thymeleaceae Daphne mezereum L. Apiaceae Aegopodium podagraria L. Angelica sylvestris L. Astrantia major L. s. lat. Chaerophyllum aureum L. Chaerophyllum hirsutum L. Laserpitium siler L. Myrrhis odorata (L.) Scop. Peucedanum austriacum (Jacq.) Koch ssp. rablense (Wulfen) Schrank Peucedanum schottii Bess. Peucedanum verticillare (L.) Koch Pimpinella saxifraga L. Sanicula europaea L. Hypericaceae Hypericum montanum L. Violaceae Viola biflora L. Viola reichenbachiana Jord. ex. Bureau Cistaceae Helianthemum nummularium (L.) Mill. ssp. grandi­florum (Scop.) Schinz & Thell Brassicaceae Arabis turrita L. Cardamine bulbifera (L.) Crantz Cardamine enneaphyllos (L.) Crantz Cardamine impatiens L. Cardamine pentaphyllos (L.). Crantz Cardamine trifolia L. Kernera saxatilis (L.) Rchb. Lunaria rediviva L. Salicaceae Popolus tremula L. Salix appendiculata L. Salix caprea L. Tiliaceae Tilia cordata Mill. Primulaceae Cyclamen purpurascens Miller Lysimachia vulgaris L. Primula auricula L. Primula vulgaris Hudson Ericaceae Calluna vulgaris (L.) Hull Erica carnea L. Vaccinium myrtillus L. Vaccinium vitis-idaea L. Pyrolaceae [Moneses uniflora (L.) A. Gray] Orthylia secunda (L.) House Monotropaceae Montropa hypophagea Walls. Adoxaceae Adoxa moschatellina L. Sambucaceae Sambucus ebulus L. Sambucus nigra L. Sambucus racemosa L. Caprifoliacae Lonicera alpigena L. Lonicera xylosteum L. Lonicera nigra L. Valerianaceae Valeriana tripteris L. Dipsacaceae Knautia drymeia Heufel ssp. drymeia Scabiosa lucida Vill. s. lat. Oleaceae Fraxinus excelsior L. Fraxinus ornus L. Gentianaceae Gentiana asclepiadea L. Ascplepidiaceae Vincetoxicum hirundinaria Medic. Rubiaceae Galim laevigatum L. Galium mollugo L. Galium odoratum (L.) Scop. Galium L. sp. (?) Solanaceae Atropa bella-donna L. Solanum dulcamara L. Boraginaceae Myosotis sylvatica (Ehrh.) Hoffm. Pulmonaria officinalis L. Symphytum tuberosum L. ssp. tuberosum Scrophulariaceae Digitalis grandiflora Miller Lathraea squamaria L. Melampyrum pratense L. ssp. vulgatum (Pers.) Ronninger Melampyrum sylvaticum L. ssp. sylvaticum Scrophularia nodosa L. Veronica montana L. Veronica officinalis L. Veronica urticifolia Jacq. Plantaginaceae Plantago major L. Lamiaceae Ajuga reptans L. Calamintha menthifolia Host. Galeobdolon flavidum (F. Herm.) Holub Galeopsis speciosa Mill. Galeopsis pubescens Besser Lamium orvala L. Lamium maculatum L. Prunella vulgaris L. Salvia glutinosa L. Thymus praecox Opiz ssp. polytrichus (Berb.) Jalas Campanulaceae Campanula cochlaeriifolia Lam. Campanula patula L. Campanula rapunculoides L. Campanula scheuchzeri Vill. Campanula trachelium L. Phyteuma ovatum Honck Phyteuma spicatum L. s. lat. Asteraceae Adenostyles glabra (Miller) DC Arctium lappa L. Aster bellidiastrum (L.) Scop. Buphthalmum salicifolium L. Carduus personata (L.) Jacq. Cirsium Miller s. lat. Cirsium erisithales (Jacq.) Scop. Cirsium oleraceum (L.) Scop. Cirsium vulgare (Savi)Tenore Doronicum austriacum Jacq. Erigeron annus (L.) Pers. Eupatorium cannabinum L. Hieracium murorum L. Homogyne sylvestris Cass. Petasites albus (L.) Gaertner Petasites paradoxus (Retz.) Baumg. Senecio ovatus (Gaertn., Mey. & Scherb.) Willd. Solidago virgaurea L. ssp. virgaurea Tussilago farfara L. Cichoriaceae Aposeris foetida (L.). Less. Mycelis muralis (L.) Dumort Prenanthes purpurea L. Taraxacum officinale agg. Trilliaceae Paris qudrifolia L. Convallariaceae Convallaria majalis L. Majanthemum bifolium L. Polygonatum verticillatum (L.) All. Melanthiaceae Veratrum album L. s. lat. Orhidaceae Cephalanthera damasonium (Mill.) Druce Cephalanthera rubra (L.) L. C. Rich. Dactylorhiza maculata (L.) Soó ssp. fuchsii (Druce) Hyl. Epipactis atrorubens (Hoffm.ex Bernk.) Besser Epipactis helleborine (L.) Crantz s. lat. Neottia nidus-avis (L.) Rich. Juncaceae Luzula luzuloides (Lam.) Dandy & Wilmott s. lat. Luzula pilosa (L.) Wild. Cyperaceae Carex alba Scop. Carex branchystashys Schrank & Moll. Carex digitata L. Carex sylvatica Huds. Poaceae Brachypodium rupestre (Host.) Roem & Schult. Brachypodium sylvaticum (Huds.) PB. Calamagrostis arundinacea (L.) Roth Calamagrostis varia (Schrad.) Host Calamagrostis villosa (Chaixex ex VilL.) J. F. Gmel. Dactylis glomerata L. Deschampsia flexuosa (L.) Trin. Deschampsia caespitosa (L.) P. Beauv. Festuca altissima All. Festuca gigantea (L.) Vill. Melica nutans L. Milium effusum L. Sesleria caerulea (L.) Ard. ssp. calcaria (Opiz) Čelak ex Hegi Table 1: Geoelements according to Poldini (1991) Geoelement Number % European 59 22,5 Circumboreal 38 14,5 Euroasian 38 14,5 Mediterranean-montane 34 13,0 Eurosiberian 22 8,4 Paleotemperate 15 5,7 Cosmopolitan 11 4,2 Northern Illyrian 7 2,7 Arctic-Alpine 5 1,9 Southern Illyrian 4 1,5 Southeast European 9 3,4 European-Mediterranean 4 1,5 Pontic 5 1,9 Mediterranean-Pontic 3 1,1 Endemic 2 0,8 Eastern Alpine 2 0,8 Mediterranean-Atlantic 2 0,8 Adventitious species 2 0,8 TOTAL 262 100,0 Table 2: Biological forms according to Raunkiaer (M. Wraber 1946) Biological form Number % Phanerophytes 42 16,0 Chamaephytes 19 7,3 Hemicryptophytes 132 50,4 Geophytes 61 23,2 Therophytes 8 3,1 TOTAL 262 100,0 VEGETATION Despite the small area of Pokljuka Gorge, 5 forest and 1 shrub association thrive in it. The largest area is oc­cupied by the associations Anemono trifoliae-Fagetum var. geogr. Helleborus niger; all other associations Ho­mogyno sylvestris-Fagetum, Mastigobryo-Piceetum var. geogr. Anemone trifolia, Corydalido cavae-Aceretum var. geogr. Dentaria enneaphyllos and Fraxino orni­-Ostryetum carpinifoliae grow on smaller areas. The association Rhytidiadelpho lorei-Piceetum is only frag­mentarily developed. The association Anemono-Fage­tum var. geogr. Helleborus niger, Homogyno-Fagetum, Corydalido-Aceretum var. geogr. Dentaria enneaphyl­los and Fraxino-Ostryetum are placed into the class of mesophilous deciduous forests on eutric soils of the class Querco-Fagetea. We classify them variously into orders and alliances, the first three phytocenoses into the order of mesophilous beech forests Fagetalia sylva­ticae and the last Fraxino-Ostryetum into the order of thermophilous oak forests Quercetalia pubescentis. Classification into alliances is more complicated. We classify the associations Anemono-Fagetum var. geogr. Helleborus niger, Homogyno-Fagetum and Corydalido­-Aceretum var. geogr. Dentaria enneaphyllos in the Il­lyrian alliance of beech forests Aremonio-Fagion. Southeast European-Illyrian, southeast European and southeast Alpine species are classified into them. Šilc & Čarni (2012), on the example of some Eu­ropean associations, classify the phytocenosis Coryda­lido-Aceretum into the alliance Tilio-Acerion and order Aceretalia pseudoplatani. We are of the opinion that the mentioned alliance and order are not supported by diagnostically important species for them but only with generally widespread Central European species of beech forests from the order Fagetalia sylvaticae s. lat. (Zupančič 1996). We classify the shrub community Fraxino-Ostryetum into the Illyrian-Balkan alliance of thermophilous continental thermophilous forests of oak and hop hornbeam Fraxino orni-Ostryion carpini­foliae. We classify the spruce associations Rhytidiade­lpho lorei-Piceeetum and Mastigobryo-Piceeetum into the class of Holarctic coniferous forests of the Eurosi­berian-North American region, Vaccinio-Piceetea, the order of Euroasian boreo-montane coniferous forests Vaccinio-Piceetalia (Piceetalia excelsae) and the alli­ance of European boreo-montane coniferous forests Vaccinio-Piceion (Piceion excelsae). ANEMONO TRIFOLIAE-FAGETUM Tre­gu­bov 1962 var. geogr. HELLEBORUS NIGER Marinček, Poldini & Zupančič 1989 The association Anemono trifoliae-Fagetum was first mentioned in print in 1957, without the publication of a phytocenological table (Tregubov 1957 a, b). It was reasoned with a phytocenological table in 1962 (Tregu­bov 1962). In both publications, Tregubov envisaged the following characteristic species for the association Anemono-Fagetum: Anemone trifolia, Cyclamen purpu­rascens, Hepatica nobilis and Helleborus niger. M. Wra­ber (1959) was of similar thinking about the associa­tion, but additionally articulated the phytocenosis into lower sinsystematic units – sub-associations. In a paper on the vegetation of Triglav National Park, the authors (Marinček et al. 1983) doubted the correctness of Tregubov’s (1957 a, b, 1962) choice of characteristic species and designated them relative characteristic spe­cies. Marinček (1983), in his book Bukovi gozdovi na Slovenskem (Beech Forests in Slovenia), speaks in ge­neral about the phytocenosis Anemono-Fagetum but does not mention its possible characteristic or differen­tial species. The findings of the phytocenologists Pol­dini from Italy and Zukrigle from Austria that the as­sociation Anemono-Fagetum or its similar in southern Austria probably thrives in northern Italy, dictated joint research, the result of which was published in Ma­rinček et al. (1989). Comparisons showed that Trebu­gov’s chosen characteristic species do not correspond. We have proposed the species Anemone trifolia as a relative characteristic species of the wider phytogeo­graphic region and added the distinguishing species Picea abies, Larix decidua, Vaccinium myrtillus, V. vitis­-idaea and Carex alba and, as further relative character­istic species, Saxifraga rotundifolia, Ranunculus platani­folius, Adenostyles glabra and Polygonatum verticilla­tum, which are constants in altimontane and subalpine beech forests. In a paper (Marinček et al. 1989), we divided the association Anemono-Fagetum into two geographic variants. In the east of the area of distribu­tion of the association is the geographic variant with the species Helleborus niger, and in the southwest that with the species Luzula nivea. Synthesis of altimontane beech forests of the alliance Aremonio-Fagion (Zupan­čič 2012) showed that the species Polygala chamae­buxus and Orthilia secunda and the distinguishing spe­cies Picea abies and Larix decidua must be considered as characteristics of the association Anemono-Fagetum. A geographic variant Anemono trifoliae-Fagetum var. geogr. Helleborus niger is widespread in Pokljuka Gorge. In the associations are represented the relative characteristic species Anemone trifolia, the distin­guishing species Picea abies and Larix decidua and the relative distinguishing species Saxifraga rotundifolia and Polygonatum verticillatum. The association grows over the majority of steep slopes of the gorge, where there are eutric shallow, skeletal carbonate brown soils or rendzinas on limestone or dolomite. The area on do­lomite is more or less smooth, but on limestone bro­ken, sometimes more rocky, so similar to ecological conditions that have already been described in previ­ously mentioned papers (Tregubov 1957 a, b, 1962, M. Wraber 1960, Marinček et al. 1989). We add a phy­tocenological relevé »in situ«. Relevé 1 Anemono-Fagetum var. geogr. Helleborus niger cephalantheretosum Altitude: 660 m, exposure: N, inclination: 40 °, soil: brown carbonate soils, geological base: dolomite. I: I = 80, II = 10, III = 50, IV = 10 I: Fagus sylvatica 4.3, Picea abies 3.2, Larix decidua +. II: Fagus sylvatica 2.3, Acer campestre +, A. plata­noides +, Acer pseudoplatanus +, Clematis vitalba +, Corylus avellana +, Fraxinus excelsior +, Lonicera nigra +, Rubus idaeus +, Sambucus nigra +, Ulmus glabra +. III: Anemone trifolia 2.2, Oxalis acetosella 2.2, Stel­laria montana 1.2, Acer pseudoplatanus 1.1, Cardamine trifolia 1.1, Galeobdolon flavidum 1.1, Homogyne sylve­stris 1.1, Prenanthes purpurea 1.1, Veronica urticifolia 1.1, Viola reichenbachiana 1.1, Actaea spicata +.2, Asa­rum europaeum +.2, Gymnocarpium dryopteris +.2, He­patica nobilis +.2, Lunaria rediviva +.2, Mercurialis pe­rennis +.2, Saxifraga cuneifolia +.2, S. rotundifolia +.2, Adoxa moschatellina +, Ajuga reptans +, Aremonia agri­monioides +, Aruncus dioicus +, Athyrium filix-femina +, Asplenium trihomanes ssp. trichomanes +, A. viride +, Campanula trachelium +, Cardamine bulbifera +, C. enneaphyllos +, Cephalanthera rubra +, Chaerophyllum hirsutum +, Cyclamen purpurascens +, Cystopteris fra­gilis +, Dactylorhiza maculata ssp. fuchsii +, Dryopteris filix-mas +, Epilobium montanum +, Euphorbia amyg­daloides +, Euphorbia dulcis ssp. incompta +, Fagus syl­vatica +, Geranium robertianum +, Geum urbanum +, Helleborus niger +, Lamium orvala +, Maianthemum bifolium +, Milium effusum +, Myosotis sylvatica +, Paris quadrifolia +, Petasites albus +, Phegopteris con­nectilis +, Phyllitis scolopendrium +, Picea abies +, Polygonatum verticillatum +, Polypodium vulgare +, Polystichum aculeatum +, Primula vulgaris +, Pulmona­ria officinalis +, Ranunculus lanuginosus +, Salvia glut­inosa +, Senecio ovatus +, Scrophularia nodosa +, Ulmus scabra +, Valeriana tripteris +, Vicia oroboides +. IV: Isothecium mysuroides 1.4, Euhrynchium zet­terstedtii 1.3, Neckera crispa +.4, Ctenidium molluscum +.2, Lobaria pulmonaria, Minum undulatum +. The relevé is defined by the lowland thermophil­ous sub-association of cephalanthera of the north-eastern geographic variant with black hellebore of al­pine forest of beech and three-leaved anemone – Ane­mono-Fagetum Tregubov 1956 var. geogr. Helleborus niger Marinček, Poldini & Zupančič 1988 cephalanthe­retosum Marinček, Poldini & Zupančič 1988. HOMOGYNO SYLVESTRIS-FAGETUM Marin­ček et al. 1993 The Dinarid phytocenosis of fir and beech was first de­scribed in Slovenia as the association Abieti-Fagetum dinaricum (Tregubov 1957 c), derived from the asso­ciation of I. Horvat (1938) Fagetum silvaticae croati­cum australe abietetosum. Tregubov (1957 a, b) later observed an individual smaller core of pre-alpine fir-beech forest on limestone in the Karavanke and desig­nated it the phytocenosis Abieti-Fagetum homogyneto­sum sylvestris. M. Wraber (1960) describes a south-eastern Alpine forest of beech and fir – Abieti-Fagetum austroalpinum - in the pre-Alpine/Alpine region as a geographic variant of Central European fir-beech forest (J. & M. Bartsch 1940). Marinček (1987), in his mon­ograph on beech forests, draws attention in the descrip­tion of pre-Alpine forest of beech and fir – Abieti-Fage­tum praealpinum – to »a fairly numerous group of Illyr­ian plant species, which indicate an Illyrian character of pre-Alpine beech forests with fir«. A year later, Ma­rinček & Dakskobler (1988) in a paper on acidophil­ous beech forests of the pre-Alpine world of Slovenia demonstrate with phytocenological tables a new acido­philous fir-beech association Luzulo-Abieti-Fagetum praealpinum with three sub-associations: typicum, gali­etosum rotundifolii and lamietosum orvalae. According to adopted Codices (Barkman et al. 1976, 1986, Weber et al. 2000) the name of the association was invalid. On the basis of phytocenological tables, the authors deter­mined distinguishing species for the association, to wit: Abies alba, Adenostyles glabra, Anemone trifolia, Festu­ca altissima, Polygonatum verticillatum, Ranunculus platanifolius and Veronica urticifolia. In a synthesis paper by Marinček et al. (1992, published 1993), the authors proposed that the pre-Alpine fir-beech forest be called after the southeast European-Illyrian species Homogyne sylvestris, namely Homogyno-Fagetum; for the nomenclature type they took relevé number 16 from Table 3 in the paper by Marinček & Dakskob­ler (1988), which is in accordance with the aforemen­tioned Codices. In research of fir-beech forests of north-western Slovenia in the region of the southern Julian Alps, Dakskobler (2002 a, 2002 b, 2009) deter­mined more exactly characteristic and distinguishing species of the association Homogyno sylvestris-Fagetum, which in the previous paper he had only defined as dis­tinguishing species, although only three of them, i.e., the species Abies alba, Adenostyles glabra and Veronica urticifolia. Instead of the other previous ones, he added the species Asplenium viride, Homogyne sylvestris and Saxifraga cuneifolia, which are more acceptable for rec­ognising the association Homogyno-Fagetum. There are smaller areas of pre-Alpine fir-beech for­est Homogyne-Fagetum in the area of Pokljuka Gorge, on its upper western edge, from whence it spreads to­wards Zatrnik and Stara Pokljuka and onwards to the Pokljuka plateau. Ecological conditions are similar to those described in the publications of Tregubov (1957), M. Wraber (1960), Marinček (1987), Marin­ček & Dakskobler (1988) and Dakskobler (2002 a, 2002 b, 2009). Upper Triassic limestone predominates on the site of pre-Alpine fir-beech forest, where there are shallow to medium deep carbonate brown soils. The relief is broken, partially rocky, here and there level, where the humus horizon is slightly acidic be­cause of the appearance of crumbled chert in the soils. Fir is rarer than in similar, more optimal conditions and a significant amount has been felled. Unfortunate­ly, there is too much management in this part of the forest. In addition to fir, of other distinguishing species are also present Veronica urticifolia, Asplenium viride, Homogyne sylvestris and Saxifraga cuneifolia. On the mixed silicaceous-limestone geological base, where there are acidic brown soils on cherts, an acidophilous variant of the association appears, Homogyno-Fagetum var. Calamagrostis arundinacea var. nova. We envisage the following distinguishing species for the variant: Calamagrostis arundinacea, Lycopodium annotinum and Huperzia selago. The holotype of the variant is the submitted phytocenological relevé 3. The edge of Pokljuka Gorge is part of a natural monument, in which management is restricted or even undesired. For illus­tration of the association Homogyno-Fagetum in Poklju­ka Gorge, we add phytocenological relevés 2 and 3. Relevé 2 Homogyno sylvestris-Fagetum Altitude: 860 m, eksposure: N, inclination: 30 °, soil: brown carbonate soils, geological base: limestone, stoniness: 40 %. I = 80, II = 0, III = 30, IV = 40 I: Fagus sylvatica 2.2, Picea abies 2.2, Abies alba 2.1, Acer pseudoplatanus 1.2, Betula pendula +, Larix decidua +, Populus tremula +. II: Abies alba +, Acer pseudoplatanus +, Corylus avellana +, Daphne mezereum +, Fagus sylvatica +, Fraxinus ornus +, Laburnum alpinum +, Lonicera alpi­gena +, Lonicera nigra +, Populus tremula +, Rosa pen­dulina +, Rubus idaeus +, Rubus saxatilis +, Sorbus aria +, Sorbus aucuparia +, Ulmus glabra +. III: Anemone trifolia 3.2, Homogyne sylvestris 1.2, Oxalis acetosella 1.2, Polygonatum verticillatum 1.2, Abies alba 1.1, Calamagrostis varia 1.1, Veronica urtici­folia 1.2, Cyclamen purpurascens + .2, Hieracium muro­rum + .2, Saxifraga cuneifolia +, Valeriana tripteris +, Acer pseudoplatanus +, Adoxa moschatellina +, Arun­cus dioicus +, Asplenium trihomanes +, A. viride +, Athyrium filix-femina +, Campanula trachelium +, Carex digitata +, Cardamine trifolia +, Cirsium erisi­thales +, Digitalis grandiflora +, Dryopteris filix-mas +, Galium laevigatum +, Gentiana asclepiadea +, Huper­zia selago +, Hypericum montanum +, Laburnum alpi­num +, Maiantemum bifolium +, Mercurialis perennis +, Mycelis muralis +, Phegopteris connectilis +, Phyteuma ovatum +, Picea abies +, Polypodium vulgare +, Pre­nanthes purpurea +, Scrophularia nodosa +, Sorbus aucuparia ssp. aucuparia, +, Veronica montana +, Viola reichenbachiana +. IV: Ctenidium molluscum 5.3, Isothecium mysuroi­des 1.2, Hylocomium splendens +.3, Bazzania trilobata +.2, Euhrynchium zetterstedtii +.2, Fissidens taxifolius +.2, Mnium sp. (?)+.2, Neckera crispa +.2, Peltigera leu­cophlebia +.2, Plagiochila asplenioides +.2, Polytrichum formosum +.2, Rhytidiadelphus triquetrus +.2, Clado­nia pyxydata +, Dicranum scoparium +. Relevé 3 Homogyno sylvestris-Fagetum var. Calamagrostis arundinacea Altitude: 850 m, exposure: N, inclination: 25 °, soil: acid brown soil on chert, geological base: limestone/chert, stoniness: 0 %, I = 60, II = 20, III = 30, IV = 5 I: Fagus sylvatica 2.2, Abies alba 1.1, Picea abies +.2, Laburnum alpinum + II: Fagus sylvatica 2.2, Lonicera nigra +.2, Betula pendula +, Picea abies +.2, Corylus avellana +, Daphne mezereum +, Laburnum alpinum +, Picea abies +, Sor­bus aria +, S. aucuparia ssp. aucuparia +, Rosa pendu­lina +, Rubus saxatilis +. III: Calamagrostis arundinacea 2.2, Vaccinium myr­tillus 1.2, Anemone trifolia + .2, Galium laevigatum + .2, Homogyne sylvestris + .2, Luzula luzuloides + .2, Ly­copodium annotinum + .2, Phegopteris connectilis +, .2, Abies alba +, Acer pseudoplatanus +, Ajuga reptans +, Campanula trachelium +, Cyclamen purpurascens +, Dryopteris filix-mas +, D. expansa (D. assimilis) +, Fagus sylvatica +, Fragaria vesca +, Gentiana asclepia­dea +, Hepatica nobilis +, Hieracium murorum +, Hu­perzia selago +, Oxalis acetosella +, Picea abies +, Polygo­natum verticillatum +, Prenanthes purpurea +, Solidago virgaurea +, Valeriana tripteris +, Veronica urticifolia +, IV: Ctenidium molluscum 1.2, Bazzania trilobata +.3, Hylocomium splendes +.3, Isothecium mysuroides +.3, Plagiochila asplenioides +.2, Polytrichum formosum +.2, Tortella tortuosa +.2, Cladonia pyxydata +, Clado­nia squamosa +, Dicranum scoparium +. CORYDALIDO CAVAE-ACERETUM PSUEDO­PLATANI Moor 1938 var. geogr. DENTARIA ENNEAPHYLLOS Zupančič 1996 The »garden plots« of Pokljuka Gorge are settled by an Illyrian variant of the Central European sycamore maple forest Corydalido cavae-Aceretum pseudoplatani var. geogr. Dentaria enneaphyllos. Habitats are frag­mentary, individual ones cover around 0.5 ha. Of char­acteristic species of Central European phytocenoses, Corydalis cava, C. solida and Lathraea squamaria are represented. Distinguishing species for the Illyrian variant are the south-eastern European-Illyrian spe­cies Anemone trifolia, Cardamine ennephyllos, C. trifo­lia and Saxifraga rotundifolia. A particularity of habi­tats of this geographic variant is the flourishing growth of the circumboreal species Matteucia struthiopteris, because of which we decided to form a sub-association according to it Corydalido cavae-Aceretum psuedopla­tani Moor 1938 var. geogr. Dentaria enneaphyllos mat­teucietosum (Zupančič 1996). Because of ill-consid­ered anthropogenic interventions in the »garden plots« and thus thinning of the tree layer, exposure to sun has affected the abundant growth of the shade loving spe­cies Matteucia struthiopteris and enabled the luxurious growth of ruderal species such as Urtica dioica, Chae­rophyllum hirsutum, Filipendula ulmaria, Lysimachia nemorum, Cirsium sp. etc. (Skumavec & Zupančič 2014). There are favourable conditions in Pokljuka Gorge for the growth of the sycamore maple association, since cold and humid air predominates here and there are slightly acid brown soils on limestone intermixed to a significant extent with chert. Under these ecologi­cal conditions, the semi-psychrophilic association of sycamore maple (Zupančič 1996), has become suc­cessfully established. In a paper by Zupančič (1996), the sycamore maple phytocenosis in two »garden plots« was described when the vegetation was in a more or less optimal state. For orientation, we add the diagnostically important species of the geographical variant Corydalido-Aceretum var. Dentaria ennaphyl­los, which are: Corydalis cava 22, C. solida 22, Lathraea squamaria 1+, Cardamine ennephyllos 2+, C. trifolia 11, Anemone trifolia 1+, Saxifraga rotundifolia 1+, Matteu­cia struthiopteris 22-3. FRAXINO ORNI-OSTRYETUM CARPINIFOLIAE Aichinger 1933 The shrub association Fraxino-Ostryetum was de­scribed by the Austrian phytocenologist Aichinger in the Austrian and partly also Slovene region of the Karavanke (Aichinger 1933). He originally called the association Ostrya carpinifolia-Fraxinus ornus. He clas­sified it phytogeographically in the Illyrian floral prov­ince, although there is no trace in it of more than two southeast European-Illyrian species: Anemone trifolia and Cyclamen purpurascens – and some more wide­spread species that we formerly classified there, i.e., Fraxinus ornus, Ostrya carpinifolia and Pinus nigra. The association occupies extreme habitats on lime­stone, where the soils are skeletal rendzinas. In Slove­nia the association was first recognised by the phytoce­nologists Tregubov (1957) and M. Wraber (1960) and they briefly described it in their contributions. Later there followed a further brief description by the au­thors of the report on the vegetation of Triglav Nation­al Park (Marinček et al. 1983). In the report are first described the characteristic and distinguishing species of the association, to wit Erica carnea, Calamagrostis varia, Polygala chamaebuxus and Sesleria caerulea s. lat. Aichinger (1933) in his monograph on the vegeta­tion of the Karavanke did not explicitly state its charac­teristic and distinguishing species but only gave a wide choice of 18 types of plant combination. Dakskobler (2015) recently performed a revision of associations of hop hornbeam and manna ash in the area of the Julian Alps and northern part of the Di­narid Massif (including the area of northern Italy). He compared them with similar associations in Austria and Croatia. He came to the conclusion that the asso­ciations differ among themselves and, on the basis of synthesis tables, showed that the following are diag­nostic species of the association Fraxino orni-Ostrye­tum Aichinger 1933: Campanula caespitosa, Primula auricula, Hieracium porrifolium, Asperula aristata, Al­lium ericetorum, Paederota lutea, Betonica alopecuros, Rhamnus fallax, Picea abies, Anemone trofolia, Valeri­ana tripteris, Salix glabra, S. appediculata, Rosa pendu­lina, Laburnum alpinum, Phyteuma orbiculare, Cam­panula carnica, Galium purpureum, Euphrasia cuspi­data, Rhododendron hirsutum, Festuca calva, Saxifraga crustata, S. hostii, Potentilla caulescens, Aconitum an­gustifolium, Seslaria caerulea ssp. calcaria. The follow­ing of the diagnostic species are present in our associa­tion: Anemone trifolia, Laburnum alpinum, Pica abies; Sesleria caerulea ssp. calcaria, (Potentilla caulescens). The cause of the impoverishment of diagnostic species is the smallness of the object, since the association is more or less fragmentarily developed. The association Fraxino-Ostryetum appears in Pokljuka Gorge above Pokljuka cave. It occupies the steeply precipitous southeast slope that transitions into the cliff above Pokljuka Gorge. The soils on the lime­stone or dolomite base are shallow rendzinas, which become lithosols in the cliff. Hop hornbeam above manna ash predominates in the shrub layer. There are individual small shrubs of the mentioned shrub spe­cies on the cliff. Phytocenological relevé 4 provides the following image of this shrub vegetation. Relevé 4 Fraxino orni-Ostryetum carpinifoliae Altitude: 750 m, eksposure: E, inclination: 75 - 90 °, soil: rendzina, geological base: dolomitized limestone, stoniness: 10 % I = 20, II = 100, III = 80, IV = 5, I: Ostrya carpinifolia 1.2, Fraxinus ornus 1.1, Betula pendula +, Fagus sylvatica +, Laburnum alpinum +, Picea abies +, Sorbus aria +. II: Ostrya carpinifolia 3.2, Fraxinus ornus 2.2, Quercus petraea 1.1, Sorbus aria 1.1, Acer platanoides +, Daphne mezereum +, Fagus sylvatica +, Laburnum alpinum +, Picea abies +, Populus tremula +, Quercus robur +, Sorbus aucuparia ssp. aucuparia +. III: Melampyrum pratense ssp. vulgatum 3.4, Cala­magrostis varia 2.2, Peucedanum schottii 1.1, Carex alba +, Convallaria majalis +, Carex digitata +, Genista tinc­toria +.2, Peucedanum austriacum ssp. rablense +.-2 , Polygala chamaebuxus +, Polypodium vulgare +.2 +, Se­sleria caerulea ssp. calcaria +.2, Thymus praecox s. lat. +.2, Acer pseudoplatanus +, Anemone trifolia +, Arabis turrita, Asplenium ruta- muraria +, A. trichomanes ssp. trichomanes +, A. viride +, Buphthalmum salicifolium +, Campanula persicifolia +, C. rapuncoloides +, Cirsium erisithales +, Conzya canadensis +, Cyclamen purpura­scens +, Dianthus hyssopifolius +, Digitalis grandiflora +, Epipactis atrorubens +, Erica carnea +, Euphorbia amygdaloides +, E. cyparissias +, Fragaria vesca +, Ga­lium laevigatum +, Hieracium murorum +, Knautia drymeia ssp. drymeia +, Laburnum alpinum +, Lathyrus pratensis +, Lotus corniculatus +, Melittis melysophyl­lum +, Moehringia muscosa +, Oryganum vulgare +, Picea abies +, Pimpinella saxifraga +, Primula vulgaris +, Pteridium aquilinum +, Scabiosa lucida +, Silene nu­tans +, Solidago virgaurea +, Sorbus aucuparia ssp. au­cuparia +, Thesium bavarum +, Veronica urticifolia +, Vicia cracca +, Vincetoxicum hirundinaria +, Viola rei­chenbachiana +, Vaccinium myrtillus +°. IV: Ctenidium molluscum 1.2, Homalothecium phi­lippeanum +.3, Grimmia pulvinata +.2, Isothecium mysuroides +.2, Neckera crispa +.2. RHYTIDIADELPHO LOREI-PICEETUM Zu­pan­čič 1981 em. 1999 M. Wraber (1953) was the first to draw attention to the spruce association in question. His description was general, without evidentiary material – phytocenologi­cal tables. He also behaved similarly later (M. Wraber 1960). He was uncertain in his descriptions whether the spruce phytocenosis with the moss Rhytidiadelp­hus loreus was an independent association or only a sub-association of some other association with the species Luzula sylvatica subsp. sylvatica, which at that time had not been validly described (Luzulo sylvaticae­-Piceetum M. Wraber 1963). Research of spruce forests of Slovenia by Zupančič showed that the spruce phyto­cenosis with the moss Rhytidiadelphus loreus is an in­dependent association. Zupančič (1980) first present­ed it in a synthesis table comparatively with other Eu­ropean spruce association. It was validly presented for a second time in a paper by Culiberg, ŠErcelj & Zu­pančič (1981). Zupančič (1999) finally formed it in a monograph on spruce forests of Slovenia. The charac­teristic species of the association are Dicranum polyse­tum, Rhytidiadelphus loreus and Thelypteris limbosper­ma. In addition to characteristic species, we chose a further group of mosses and lichens that characteristi­cally mark the phytocenosis Rhytidiadelpho-Piceetum, with an average 70 % cover of the habitat. These are: Bazzania trilobata, Cetraria islandica, Cladonia pyxi­data, C. rangiferina, Fissidens taxifilius, Hylocomium splendens, Leucobrium glaucum, Mylia taylori, Plagio­chila aspleniodies var. major, Plagiothecium neglectum, P. undulatum, Pleurozium schreberi, Polytrichum for­mosum, Rhytidiadelphus triquetrus, Scapania nemoro­sa and Tortella tortuosa. The association Rhytidialpho­-Piceetum normally grows on distric acid brown soils on silicate, and on Pokljuka, unagglutinated moraine with chert of Quaternary age. In Pokljuka Gorge there are moderately acid brown soils on a limestone base, intermixed with a consider­able quantity of crumbled chert. The described soils, with boulders and rocks and with temperature inver­sion, are only suitable for the development of spruce forest. These ecological conditions enable the growth of beech, although on small areas of only a few are, or it appears only here or there, visibly feeble. There are two fragments of spruce forest in Pokljuka Gorge in which, of the characteristic species, there is only the moss Rhytidialphus loreus on very small areas. Other mosses achieve greater cover values, as is evident from phytocenological relevé 5, the majority being acidophi­lous which characteristically ecologically mark piceetal habitats. In addition to acidophilous mosses, tubulous flowers from the class Vaccinio-Piceetea are fairly nu­merous. The phytocenological relevé shows the selec­tion of plants. Relevé 5 Rhytidiadelpho lorei-Piceetum Altitude: 680 m, exposure: N, slope: 35 °, stoniness: 80 % fallen boulders and rocks, soils: rendzinas, litho­sols, geological base: limestone with chert I = 60, II = 10, III = 50, IV = 80 I: Picea abies 3.3, Fagus sylvatica +, Larix decidua + II: Lonicera nigra 2.2, Clematis alpina 1.2, Picea abies +.2, Fagus sylvatica +, Fraxinus ornus +, Labur­num alpinum +, Lonicera alpigena +, Ostrya carpinifo­lia +, Rosa pendulina +, Rubus idaeus +, R. saxatilis +, Sorbus aucuparia ssp. aucuparia +, Ulmus glabra +. III: Homogyne sylvestris 2.2, Gymnocaripium dryopteris 1.2, Oxalis acetosela 1.2, Polypodium vulgare 1.2, Valeriana tripteris 1.2, Veronica urticifolia 1.2, Ly­copodium annotinum +.3, Adenostyleds glabra +.2, Asplenium trihomanes ssp. quadrivalens (?) +.2, Cala­magrostis arundinacea +.2, C. villosa +.2, Carex digita­ta +.2, Cystopteris fragilis +.2, Dryopteris filix-mas +.2, Festuca altissima +.2, Luzula luzuloides +.2, Saxifraga cuneifolia +.2, Vaccinium myrtllus +.2, Acer platanoides +, A. pseudoplatanus +, Actea spicata +, Adoxa mo­schatellina +, Anemone trifolia +, Aruncus dioicus +, Athyrium filix-femina +, Campanula cochlaeriifolia +, Cardamine pentaphyllos +, C. trifolia +, Cephalanthera damasonium +, Circaea lutetiana +, Cirsium erisithales +, Cyclamen purpurascens +, Dryopteris expansa, Epilo­bium montanum +, Fagus sylvatica +, Galeobdolon fla­vidum +, Galium leavigatum +, Hieracium murorum +, Melampyrum sylvaticum +, Mercurialis perennis +, Myosotis sylvatica +, Petasites albus +, Phyllitis scolo­pendrium +, Phyteuma ovatum +, Picea abies +, Polysti­chum aculeatum +, Prenanthes purpurea +, Sanicula europaea +, Saxifraga cuneifolia +, Senecio ovatus +, Solidago virgaurea +, Sorbus aucuparia s. lat. +, Symphytum tuberosum +., Ulmus glabra +, Viola rei­chenbachiana. IV: Isothecium mysuroides 3.5, Eurhynchium zetter­stedtii 2.4, Hylocomium splendes 2.4, Minum undula­tum 1.4, Plagiochila asplenioides 1.3, Rhytidiadelphus triquetrus 1.3, Ctenidium molluscum +.3, Fissdens taxi­folius +.3, Mnium spinosum (?)+.3, M. punctatum +.2, Metzgeria furcata +.2, Neckera crispa +.2, Plagiothecium undulatum +.2, Polytrichum formosum +.2, Bazzania trilobata +, Cladonia rangiferina +, Dicranum scopari­um +, Peltigera leucophlebia +, Rhytidiadelphus loreus +. A few years ago, we recorded some specimens of the north-eastern Eurasian-Circumpolar species Mo­neses uniflora on this habitat. We have no longer found this species recently. They were probably picked by visitors to whom we showed them on natural history excursions. MASTIGOBRYO-PICEETUM (Schmidt & Gais­berg 1936) Br.-Bl. & Sissingh in Br.-Bl. et al. 1939 corr. Zupančič 1999 var. geogr. ANEMONE TRIFOLIA var. geogr. nova German phytocenologists have for the most part been involved in research of the Central European acidophi­lous spruce association Mastigobryo-Piceetum (Schmidt, Gaisberg, R. Tüxen, Oberdorfer, J. & M. Bartsch, Jahn, Hartmann). In the vicinity of Slovenia in Koroška (Carinthia), it was already recognised at the start of the nineteen thirties by the Austrian phytoce­nologist Aichinger. His compatriot Smettan presented it in the Tyrol with 10 phytocenological relevés. The association was first described in Slovenia by Persoglio (in Tregubov 1957) under the name Bazzanio-Picee­tum, in the Upper Sava Valley. Zupančič (1999), while studying spruce association checked Persoglio’s re­search of the association Mastigobryo-Piceetum and compared his results with the results of the previously mentioned European phytocenologists; he came to the conclusion that the northwest European phytocenosis differs from ours, so he characterised it as a new geo­graphic variant Mastigobryo-Piceetum var. geogr. Tri­entalis europaea Zupančič 1999, with distinguishing species the Arctic-Nordic boreal element Trientalis eu­ropaea and the acidophilous boreal moss Ptilium cri­sta-castrensis. At the same time, on the basis of exam­ples from 9 analytical tables of the association Masti­gobryo-Piceetum, more or less reliably determined characteristic or distinguishing species of this associa­tion, from those previously very loosely envisaged by Braun-Blanquet (1939): these are Blechnum spicant, Bazzania trilobata and Sphagnum nemoreum. Along the extreme northeast edge of Pokljuka Gorge appears the acidophilous forest association Ma­stigobryo-Piceetum var. geogr. Anemone trifolia. On the basis of the predominately acidophilous vegetation and the prevailing chert in the soils, we conclude that it grows on distric brown soils that are shallow to medi­um deep. Two characteristic species are present on the described surfaces, Bazzania trilobata and Blechnum spicant. In view of the mass appearance of the north­east-Euroasian-Suboceanic species Melampyrum pra­tense L. subsp. vulgatum (Pers.) Ronniger, we classified both described habitats of the association into a new sub-association Mastigobryo-Piceetum melampyreto­sum vulgati subass. nova. The holotype of the sub-as­sociation is relevé 1 (Phytocenological table). The spe­cies Melampyrum pratense subsp. Vulgatum, together with some other species, e.g., Calluna vulgaris, Pteridi­um aquilinum, Potentilla ercta, Carex alba, appears on drier or, rather, less damp habitats. We classified the association Mastigobryo-Picee­tum according to phytogeographic principles as a northwest geographic variant of the pre-Alpine-Alpine region of Slovenia, with the distinguishing species Anemone trifolia L., which for this region is an explic­itly characteristic Alpine-southeast European-Illyrian species. We also ranked in the phytocenological table the relevé described in 1986 from the region of the Karavanke – Sava Caves (Zupančič 1999), which be­longs in this geographic variant and shows similarities to the already known sub-association lueucobrietosum (Persoglio in Tregubov 1957). The phytocenological table shows the vegetation and floristic image more precisely. MASTYGOBRYO-PICEETUM (Schmidt & Gaisberg 1936) Br.-Bl. & Sissingh in Br.-Bl. et al. 1939 corr. Zupančič 1999 var. geogr. ANEMONE TRIFOLIA var. geogr. nova Zaporedna številka popisa (Number of relevé) 1 2 3 Delovna številka popisa (Working number of relevé) 7/14 8/14 38/86 Datum (Date) 18.8.2014 18.8.2014 27.8.1986 Nadmorska višina v m (Altitude in m) 841 862 1250 Sinsistematska pripadnost (Sinsistematical characteristic) Nebesna lega (Aspect) NE E E Nagib v stopinjah (Slope in degrees) 20 20 25 Kamnitost v % (Stoniness in %) 0 0 0 Geološka podlaga (Bedrock) r o ž e, a p n e sili, skri Tla (Soil) kisla distrična rjava tla Acidophilous dystric cambisol Pokrovnost (Cover) %: drevesna plast (Tree layer) I 60 60 70 grmovna plast (Shrub layer) II 5 5 30 zeliščna plast (Herb layer) III 60 90 70 mahovna plast (Moss layer) IV 15 15 30 Velikost popisne ploskve (Relevé) m2 400 400 400 Presence (Prezenca) Kraj popisov (Location) Pokljuška soteska Julijske Alpe Savske jame Karavanke ZNAČILNICE ZA ASOCIACIJO (Characteristic species of association) 1 2 3 VP Bazzania trilobata IV 1.3 +.3 2.3 3+-2 VP Blechnum spicant III + +.3 2.2 3+-2   RAZLIKOVALNICA ZA GEOGRAFSKO VARIANTO (Diferential speecies of geographical variant)   1 2 3 F Anemone trifolia III +.2 +.2 + 3+   RAZLIKOVALNICA ZA SUBASOCIACIJO M.-P. var. geogr. ANEMONE TRIFOLIA MELAMPYRETOSUM subass. nova (Diferential species of subassociation)   1 2 3 RP Melampyrum pratense subsp. vulgatum III 2.3 2.3 . 22   RAZLIKOVALNICA ZA SUBASOCIACIJO M.-P. var. geogr. ANEMONE TRIFOLIA LEUCOBRYETOSUM Persoglio 1957 (Diferential species of subassociation) 1 2 3 VP Leucobryum glaucum IV +.2 1.2 2.3 3+-2   VP VACCINIO-PICEETEA Br.-Bl. in Br.-Bl. et al. 1939 em. Zupančič (1980) 2000 s. lat.   1 2 3   I 3.3 3.3 4.1 33-4 Picea abies II . + 2.3 2+-2 3+-4   III + + . 2+ Vaccinium myrtillus 3.3 3.3 + 0 3+-3 Calamagrostis arundinacea 2.3 1.3 2.4 31-2 Polytrichum formosum IV 1.3 1.2 2.3 31-2 Bazzania trilobata 1.3 +.3 2.3 3+-2 Leucobryum glaucum +.2 1.2 2.3 3+-2 Blechnum spicant III + +.3 2.2 3+-2 Lycopodium annotinum + +.2 2.2 3+-2 Maianthemum bifolium 1.1 + 1.1 3+-1 Dicranum scoparium IV +.2 1.2 +.2 3+-1   I . . + 1+ Abies alba II . . 1.1 11 3+-1   III + + . 2+ Dicranum polysetum IV +.2 +.3 +.2 3+ Plagiothecium undulatum +.3 +.2 +.2 3+ Dryopteris expansa (D. assimilis) III +.2 +.2 + 3+ Phegopteris connectilis + +.2 + 3+ Oxalis acetosella + + +.2 3+ Calluna vulgaris + +.2 + 3+ Hieracium murorum + + + 3+ Hypnum cupressiforme IV 1.2 1.2 . 21 Thelypteris limbosperma III . + 1.3 2+-1 Huperzia selago + . 1.2 2+-1 Luzula luzuloides +.2 +.2 . 2+ Sphagnum girgensohnii IV +.2 . +.2 2+ Luzula pilosa III . + +.2 2+ Gymnocarpium dryopteris + + . 2+ Solidago virgaurea + + . 2+ Gentiana asclepiadea + . + 2+ Hylocomium splendens IV . +.3 . 1+ Peltigera leucophlebia . +.3 . 1+ Dicranella heteromalla +.2 . . 1+ Homogyne alpina III . . +.2 1+ Aposeris foetida . . +.2 1+ Cantharellus cibarius + . . 1+ Larix decidua II . + . 1+ Lonicera nigra + . . 1+ Monotropa hypophegea III . + . 1+ Plagiochila asplenioides var. major IV + . . 1+ Plagiothecium neglectum + . . 1+ Rosa pendulina II . + . 1+ Rubus saxatilis + . . 1+ Vaccinium vitis-idaea III + . . 1+ Veronica urticifolia . + . 1+ Melampyrum sylvaticum . . + 1+ Cetraria islandica IV . . + 1+ Rhytidiadelphus triquetrus . . + 1+ Equisetum sylvaticum III . . + 1+   RP QUERCETALIA ROBORIS-PETRAEAE R. Tx. (1931) 1937 s. lat.   1 2 3 Melampyrum pratense subsp. vulgatum III 2.3 2.3 . 22 Populus tremula I . +.3 . 1+ 2+ II + + . 2+ Pteridium aquilinum III + +.2 . 2+ Betula pendula II + + . 2+ Veronica officinalis III + + . 2+ Carex montana . . +.2 1+ Potentilla erecta . + . 1+   EP ERICO-PINETEA Ht. 1959 s. lat.   1 2 3 Carex alba III 1.2 +.3 . 2+-1 Erica carnea . + . 1+   A ADENOSTYLETALIA G. & J. Br.-Bl. 1931   1 2 3 Polygonatum verticillatum III + + . 2+ Rubus idaeus II + + . 2+ Veratrum album III + . . 1+   F QUERCO-FAGETEA Br.-Bl. & Vlieger in Vlieger 1937 s. lat.   1 2 3 Anemone trifolia III +.2 +.2 + 3+ Laburnum alpinum II + 1.1 . 2+-1 2+-1 III . + . 1+ Carex digitata + +.2 . 2+ Ctenidium molluscum IV + +.2 . 2+ Fagus sylvatica I + + . 2+ 2+ II +.2 + . 2+ Acer pseudoplatanus + + . 2+ Cirsium erisithales III + + . 2+ Prenanthes purpurea + + . 2+ Eurhynchium zetterstedtii IV . +.2 . 1+ Aruncus dioicus III + . . 1+ Corylus avellana II + . . 1+ Isothecium myosuroides IV . + . 1+ Mycelis muralis III . + . 1+ Sambucus racemosa II + . . 1+   Q QUERCETALIA PUBESCENTIS Br.-Bl. (1931 n. nud.) 1932 s. lat.   1 2 3 Sorbus aria II + + . 2+ Fraxinus ornus . + 0 . 1+0   O OSTALE VRSTE (Other Species)   1 2 3 Cladonia pyxidata IV + . +.2 2+ Cladonia rangiferina IV + + . 2+ Sorbus aucuparia subsp. aucuparia II + + . 2+ 2+ III . + . 1+ Tortella tortuosa IV . +.2 . 1+ Ajuga reptans III + . . 1+ Galeopsis pubescens . + . 1+ LEGENDA (Legend) Geološka podlaga (Bedrock) apne Apnenec (Limestone) rože Roženec (Chert) sili Silikat (Silikate) skri Skriljavec (Schist) CONCLUSIONS Because of its natural features, Pokljuka Gorge is enti­tled to special treatment within the framework of Triglav National Park. Because of its botanical and geological features of interest, it should be protected on the same level as the protection of the central part of the park. Today it is protected according to the criteria of the outer zone of Triglav National Park. In the small area of Pokljuka Gorge – around 36 hectares – there are a large number of geological karst phenomena, such as sink­holes, natural bridges, Pokljuka Save, Stranska soteska, narrows, chert inserts in limestone etc., which is a rarity in the world. The flora is even more interesting, with the subendemic Saxifraga burseriana, alpine flora and mountain or high mountain vegetation because of air inversion, which causes plant inversion. Because of the specific colder micro- and mezoclimatic conditions in Pokljuka Gorge, which is located in the montane belt from 670 to 800 m asl, alpine plants appear: Primula au­ricula, Potentilla caulescens, Viola biflora, Saxifraga cu­neifolia, Lycopodium annotinum, Huperzia selago, Vero­nica urticifolia, Lonicera nigra, (Moneses uniflora) and many others. Spruce communities occupies the coldest or acidic soil parts of the gorge, and sycamore maple thrives in the coldest »garden plots«. This region, with a relatively small area, is settled by 6 forest-shrub associa­tions and 262 plant species, which gives the gorge a spe­cial seal because of the botanical and vegetational vari­ety, which is supplemented by the Alpine-Karstic geo­logical and soil variety. With our presentation of the flora and vegetation, we wished to show a part of these interesting features of Pokljuka Gorge. We would like this contribution to encourage zoologists to deal with the fauna of Pokljuka Gorge, because we do not doubt that the variety is even greater than the botanical. The description of the vegetation gave us the op­portunity to show the development path and problems of sintaxonomy, which required lengthy deliberation during the search for more or less final solutions. This seemed to us particularly necessary and important for an area in which there are not optimal but exceptional ecological conditions for the forest associations in ques­tion. Their development tends towards opposing condi­tions, i.e., to low altitudes with specific microclimatic influences. A continuous struggle for the dominance of one or another influence takes place among them, which enables the optimal development of one phyto­cenosis or another. The consequence of these conditions is that the plant cover of the phytocenoses is limited to the most adaptable plant species, which can withstand the daily or annual exchange of temperature influences. This is also reflected in the described forest phyto­cenoses, in which not all characteristic and distinguish­ing species are represented, lacking those that need opti­mal conditions for their development. Only characteris­tic and distinguishing species are present that are adapt­ed to these unquiet (turbulent) ecological conditions. ACKNOWLEDGEMENT We are grateful to colleagues Dr. Igor Dakskobler and Emeritus Professor Dr. Ljudevit Ilijanic, correspond­ing member of SAZU, for reviewing the paper and use­ful advice. We are similarly grateful to Vinko Žagar BA, for computer processing of the phytocenological table. POVZETEK Pokljuška soteska je med najzanimivejšimi naravnimi znamenitostmi Triglavskega narodnega parka. Vreza­na je v strm severovzhodni rob Pokljuške planote na nadmorskih višinah od 670 do 800 m. Je največja fosil­na soteska v Sloveniji, nastala pred mnogimi milijoni let, ko so jo izoblikovale vode Triglavskega ledenika. (Smolej 1982, Ramovš 1986, Skumavec 1995, Skuma­vec & Skobrne 1995). Pokljuško sotesko so mnogokrat obiskovali nara­voslovci, tudi midva. Predvsem sva opazovala floro in vegetacijo. V letu 1996 je bila natisnjena razprava o be­lojavorjevi združbi v Pokljuški soteski (Zupančič 1996). S tem v zvezi se je kolegu Skumavcu porodila zamisel o načrtnem pregledu vaskularne flore in vege­tacije Pokljuške soteske. Rezultat je pričujoča razprava. Raziskovana pot naju je vodila vzdolž soteske, od Jele, Kobalovega rovta, Stranske soteske, Pokljuške lu­knje, Srednjega vrtca, Galerij, Velikega vrtca, po poti proti Zatrniku nad sotesko, prek prehoda ali krožne poti proti Stari Pokljuki oziroma pod Pustovem polju, skozi Pokljuško luknjo in nazaj do Jele. Območje sote­ske obsega skoraj 2 km dolžine. Želela sva zajeti in po­pisati čim več flore, verjetno se nama to ni povsem posrečilo in predvidevava, da je ali bo mogoče najti še kakšno vrsto, ki sva jo spregledala. Gozdne združbe so predstavljene s 5 fito­ceno­loškimi popisi in fitocenološko tabelo. Floristične in vegetacijske raziskave so potekale po standardnih metodah. Klimatske razmere v Pokljuški soteski so bolj ali manj podobne tistim, ki vladajo v alpskem svetu (1500–2000 m padavin, povprečna letna temperatura 3 do 6 oC). Pokljuška soteska je mraziščno območje. Geološko sestavo večinoma predstavlja zgornjetri­adni apnenec in nekaj enako starega dolomita. Na ne­katerih mestih (npr. v »vrtcih«) je primešan roženec. Tla na apnencu in dolomitu so karbonatna bazična, evtrična rjava in rendzine. Tla, presuta z rožencem, so distrična, zmerno kisla rjava. Na balvanih in skalah se pojavlja litosol. Zabeležili smo 262 taksonov vaskularne flore (Ta­bela 2). Posebnost je prisotnost 11 jugovzhodnoevrop­sko-ilirskih vrst: Anemone x pitonii, A. trifolia, Aspose­ris foetida, Cardamine ennaphyllos, C. trifolia, Galium laevigatum, Helleborus niger, Homogyne sylvestris, Knautia drymeia ssp. drymeia in Lamium orvala ter subendemit, Saxifraga burseriana, ki ga še s 5 vrstami: Cardamine pentaphyllos, Chaerophyllum hirsutum, Pe­ucedanum austriacum ssp. rablense, Primula vulgaris in Stellaria montana, uvrščamo med jugovzhodnoe­vropsko floro. Več kot dve petini vrst je hladnoljubnih, to so cirkamborealni, mediteransko-montanski, evro­sibirski, paleotemperatni, arktično-alpski in vzhodno­alpski geoelementi (po Poldiniju 1991 – Tabela 1). Analiza flore kaže, da območje Pokljuške soteske uvr­ščamo v ilirsko florno provinco, jugovzhodnoalpski florni sektor in v julijskoalpski-zahodnokaravanški­-kamniškoalpski distrikt (Zupančič et al. 1987). Zava­rovanih vrst je 14: Cephalanthera damasonium, C. rubra, Convallaria majalis, Cyclamen purpurascens, Dactylorchiza maculata ssp. fuchsii, Dianthus hyssopi­folius, Epipactis helleborine, Helleborus niger, H. odo­rus, Huperzia selago, Hypericum montanum, Lycopodi­um annotinum, Neottia nidus-avis in Primula acaulis. Pomembna je prisotnost subendemita Saxifruga burse­riana. V Pokljuški soteski smo določili 5 gozdnih in 1 gr­miščno združbo. Od listnatih gozdov je najbolj razšir­jena asociacija oziroma geografska varianta Anemono trifoliae-Fagetum var. geogr. Helleborus niger, ki pora­šča evtrična plitva, skeletna karbonatna rjava tla ali rendzine na apnencu in dolomitu. Na apnencu, kjer so plitva do srednje globoka karbonatna rjava tla, uspeva asociacija Homogyno-Fagetum. Kisla varianta asociaci­je Homogyno-Fagetum var. Calamograstis arundinacea var. nova. je na mešani silikatno-apnenčasti geološki podlagi, kjer so kisla rjava tla na rožencih. V »vrtcih« se pojavlja belojavorjeva asociacija oziroma geografska varianta Corydalido cavae-Aceretum pseudoplatani var. geogr. Dentaria enneaphyllos, ker tam prevladuje hla­den in vlažen zrak. Tla so zakisana rjava na apnencu s primesjo roženca. Fragmentarno razvita smrekova združba Rhytidiadelpho lorei-Piceetum naseljuje razgi­ban apnenčast skalovit svet na balvanih, kjer so tla po­mešana z rožencem. Na skalah (balvanih) pa je inicial­ni kisli litosol. Ta rastišča so pod vplivom temperatur­ne inverzije in hladnih tal, ki jih povzroča kroženje zraka med skalami, kjer dolgo časa ležita sneg in led. Drugi smrekov gozd oziroma geografska varianta Ma­stigobryo-Piceetum var. geogr. Anemone trifolia var. geogr. nova je na distričnih kislih rjavih plitvih do sre­dnje globokih tleh z obilico roženca. Strmo apnenča­sto-dolomitno pobočje in stene nad Pokljuško luknjo porašča toploljubno grmišče Fraxino orni-Ostryetum. Tla so rendzine, ki v prepadni steni prehajajo v bazični litosol. Opisana gozdno-grmiščna vegetacija je utemeljena s fitocenološkimi popisi in fitocenološko tabelo »in situ« ter razpravo Zupančiča (1996). Pri opisu vegetacije smo izkoristili priložnost, da prikažemo razvojno pot in probleme sintaksonomije, ki je zahtevala dolgotrajna razmišljanja ob iskanju bolj ali manj dokončnih rešitev. To se nam je zdelo še pose­bej potrebno in pomembno za območje, kjer za obrav­navane gozdne združbe ni optimalnih ekoloških raz­mer, temveč so te izredne. Njihov razvoj poteka v na­sprotujočih si razmerah, to je na nizkih nadmorskih višinah s posebnim mikroklimatskim vplivom. Med njimi se odvija neprestan boj za prevlado enega ali dru­gega vpliva, kar onemogoča optimalni razvoj te ali one fitocenoze. Posledica teh razmer je, da je rastlinska odeja fitocenoz omejena na najbolj prilagodljive ra­stlinske vrste, ki prenesejo vsakodnevno oziroma vsa­koletno menjavanje temperaturnih vplivov. To se odra­ža tudi na opisanih gozdnih fitocenozah, kjer niso za­stopane vse njihove značilnice in razlikovalnice, ki za svoj razvoj potrebujejo optimalne razmere. Prisotne so le tiste značilnice ali razlikovalnice, ki so prilagojene tem nemirnim (turbulentnim) ekološkim razmeram. Pokljuška soteska je zaradi svoje naravoslovne znamenitosti upravičena do posebne obravnave v sklo­pu Triglavskega narodnega parka. Zaradi botaničnih in geoloških zanimivosti bi jo morali zavarovati na ravni varovanja osrednjega dela Triglavskega narodne­ga parka. Danes je zavarovana po kriterijih zunanjega pasu Triglavskega narodnega parka. REFERENCES Aeschimann, D., K. Lauber, D. M. Moser & J.-P. 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Razprave 4. razr. SAZU (Ljubljana) 37 (8): 189–205. Tab. 1. Zupančič, M., 1999: Smrekovi gozdovi Slovenije. Dela SAZU 36 (Ljubljana). Zupančič, M., 2012: Syntaxonomic problems of altimontane beech forests of the alliance Aremonio-Fagion in Slove­nia. Folia biologica et geologica (Ljubljana) 53 (1-2): 83–127. 1 SAZU, Novi trg 5, SI-1000 Ljubljana, Slovenija. 2 Zgornje Laze 21, SI-4247 Zgornje Gorje, Slovenija, email: skumavec@siol.net Figure 1: Location of the research area Figure 2: Notice in Jela at the entrance to Pokljuka Gorge (Photo: J. Skumavec) Figure 3: Burser‘s saxifrage - Saxifraga burseriana L. on the cliff in front of Pokljuka Cave (Photo: J. Skumavec) Figure 4: Pre-Alpine beech forest - Anemono-Fagetum (Photo: J. Skumavec) Figure 5: Ostrich fern - Matteuccia struthiopteris (L.) Tod. in sycamore-maple forest (Photo: J. Skumavec) Figure 6: Sub-Alpine spruce forest - Rhytidiadelpho lorei-Piceetum (Photo: J. Skumavec) Figure 7: Bridge, called the Galleries, in Pokljuka Gorge (Photo: J. Skumavec) Figure 8: Cherts in Triassic limestone (Photo: J. Skumavec) Figure 9: Pokljuka Gorge seen from Mežakla (Photo: J. Skumavec) B. DROVENIK & B. VREŠ: NOVOSTI V FAVNI RODU OTIORHYNCHUS S. LAT. (COLEOPTERA: CURCULIONIDEA) V SLOVENIJI B. DROVENIK & B. VREŠ: NOVOSTI V FAVNI RODU OTIORHYNCHUS S. LAT. (COLEOPTERA: CURCULIONIDEA) V SLOVENIJI 152 153 FOLIA BIOLOGICA ET GEOLOGICA 56/1, 151–162, LJUBLJANA 2015 FOLIA BIOLOGICA ET GEOLOGICA 56/1 – 2015 FOLIA BIOLOGICA ET GEOLOGICA 56/1 – 2015 NOVOSTI V FAVNI RODU OTIORHYNCHUS S. LAT. (COLEOPTERA: CURCULIONIDEA) V SLOVENIJI NOVELTIES IN THE FAUNA OF OTIORHYNCHUS S. LAT. (COLEOPTERA: CURCULIONIDEA) IN SLOVENIA Božidar DROVENIK1 & Branko VREŠ2 Izvleček Novosti v favni rodu Otiorhynchus s. lat. (Coleoptera: Cur­culionidea) v Sloveniji Favna hroščev rilčkarjev iz rodu Otiorhynchus s. lat. v Sloveniji doslej še ni bila zadostno poznana. V preteklosti je razširjenost tega rodu na širšem ozemlju nekdanje Jugoslavi­je doslej predstavil le Kovačević (1971), ki v svojem delu omenja tudi okrog 84 vrst iz Slovenije. Pri raziskavah razšir­jenosti vrst rodu Otiorhynchus s. lat. v Sloveniji, smo v prete­klih dveh desetletjih zabeležili 17 novih vrst: Dodecastichus aurosignatus (Apfelbeck 1889) = Otiorhynchus aurosignatus Apfelbeck 1889, D. brevipes (Apfelbeck 1894) = Otiorhynchus brevipes Apfelbeck 1894, D. dalmatinus (Gyllenhal, 1834) = Otiorhynchus dalmatinus Gyllenhal 1834, D. heydeni (Stier­lin, 1861) = Otiorhynchus heydeni Stierlin 1861, Otiorhynchus alutaceus (Germar 1817), O. azaleae Penecke 1894, O. brusi­nae Stierlin 1888, O. pantherinus Apfelbeck 1898, O. porcatus (Herbst 1795), O. primigenius Apfelbeck 1918 = O. austriacus primigenius Apfelbeck 1918, O. remotegranulatus Stierlin 1891, O. rugifrons (Gyllenhal 1813), O. subdentatus Bach 1834, O. tenebricosus (Herbst 1784) = fuscipes (Olivier 1807), O. truncatus Stierlin 1861, O. turca Boheman 1843 in O. un­cinatus Germar 1824. V prispevku predstavljamo sedanje poznavanje njihove razširjenosti in ekologije. Podatki teme­ljijo predvsem na prepariranih primerkih B. Drovenika in V. Furlana, ki so shranjeni v koleopteroloških zbirkah Biolo­škega inštituta Jovana Hadžija ZRC SAZU in Prirodoslovne­ga muzeja Slovenije. Skupno število sedaj znanih vrst iz rodov Dodecastichus in Otiorhynchus s.str. v Sloveniji je 103. Ključne besede: Curculionidae, Dodecastichus, Oti­orhynchus, rilčkarji, favna, Slovenija Abstract Novelties in the fauna of Otiorhynchus s. lat. (Coleoptera: Curculionidea) in Slovenia Fauna of weevil beetles from genus Otiorhynchus s. lat. was not yet very well known in Slovenia. Distribution of this genus on the territory of former Yugoslava was presented by Kovačević (1971) who mentioned about 80 species from Slo­venia. Our investigation of distribution of Otiorhynchus s. lat. in Slovenia in the last two decades has added 17 addi­tional species for this territory. They are: Dodecastichus aurosignatus (Apfelbeck 1889) = Otiorhynchus aurosignatus Apfelbeck 1889, D. brevipes (Apfelbeck 1894) = Otiorhynchus brevipes Apfelbeck 1894, D. dalmatinus (Gyllenhal, 1834) = Otiorhynchus dalmatinus Gyllenhal 1834, D. heydeni (Stier­lin, 1861) = Otiorhynchus heydeni Stierlin 1861, Otiorhynchus alutaceus (Germar 1817), O. azaleae Penecke 1894, O. brusi­nae Stierlin 1888, O. pantherinus Apfelbeck 1898, O. porcatus (Herbst 1795), O. primigenius Apfelbeck 1918 = O. austriacus primigenius Apfelbeck 1918, O. remotegranulatus Stierlin 1891, O. rugifrons (Gyllenhal 1813), O. subdentatus Bach 1834, O. tenebricosus (Herbst 1784) = fuscipes (Olivier 1807), O. truncatus Stierlin 1861, O. turca Boheman 1843 in O. un­cinatus Germar 1824. Recent knowledge of their distribution and ecology are presented. Data are based on pinned speci­mens from the collections of B. Drovenik and V. Furlan in Jovan Hadži Institute of Biology SRC SASA and Slovenian Museum of Natural History in Ljubljana. 103 species of wee­vil beetles from genera Dodecastichus and Otiorhynchus s. str. are known from the territory of Slovenia at the present time. Keywords: Curculionidae, Dodecastichus, Otiorhynchus, weevil beetles, fauna, Slovenia 1 UVOD Rilčkarji (Curculionidae) Slovenije in med njimi tudi rod Otiorhynchus, Germar 1824, so do sedaj še vedno slabo poznana skupina živali. To je vrstno zelo velika skupina hroščev, ki šteje v Evropi preko 5000 različnih taksonov, v Sloveniji pa je trenutno poznanih preko 900 vrst. Skupina rilčkarjev je dokaj zahtevna za pre­poznavanje (določanje), majhno pa je tudi število razi­skovalcev, ki se z njihovo taksonomijo in razširjenostjo ukvarja tako v Evropi kot v Sloveniji. Rod Otiorhync­hus spada med vrstno najbogatejše rodove rilčkarjev v Evropi, za katerega je poznanih 864 taksonov (vrst in podvrst) rodu Otiorhynchus s. str. oziroma 934 takso­nov rodu Otiorhynchus s. lat. (Alonso-Zarazaga 2004). V Sloveniji je rod Otiorhynchus s. lat. po doslej znanih podatkih zastopan s 103 vrstami in podvrstami (B. Drovenik, neobjavljeno), kar predstavlja 11 % po­znanih evropskih taksonov. V delu Kovačevića (1971) je navedenih za Slovenijo 84, za vso (nekdanjo) Jugo­slavijo pa 228 vrst. Večletne raziskave favne hroščev v Sloveniji so po­kazale, da je poznavanje te velike skupine žuželk še zelo nepopolno in do danes smo za favno Slovenije zbrali številne nove podatke tudi za rilčkarje. Pri rodu Otiorhynchus smo našli 17 novih vrst, ki jih predsta­vljamo v tem prispevku s kartami njihove razširjenosti in kratkimi opisi, 8 med njimi pa tudi slikovno. Ta rod ima številne vrste, ki so vezane na ozko omejeno geo­grafsko območje; številne med njimi so endemiti ali imajo disjunktne areale. Vzrok dokaj veliki številčno­sti endemitov te skupine hroščev je med drugim tudi v tem, da osebki ne letajo, ker so nekrilati oziroma imajo zraščena krila (pokrovke). Mnogi med njimi so mono­fagi, hranijo se izključno le z določeno rastlinsko vrsto, ki je lahko zelo omejeno razširjena. V zadnjih desetletjih so rilčkarje v Sloveniji razi­skovali predvsem S. Brelih, V. Furlan, B. Kofler in M. Zdešar. Del njihovih podatkov smo uporabili tudi v tem prispevku. Nekaj posameznih podatkov so prispe­vali še tuji strokovnjaki E. Holzer, M. Kahlen, H. Ki­ppenberg in H. Freude. Večji del predstavljenih podat­kov za rilčkarje oziroma rod Otiorhynchus v Sloveniji je zbral B. Drovenik. Določanje oziroma revizijo dolo­čitev sta opravila vodilna evropska strokovnjaka tega rodu L. Behne in O. Vořišek. Pri našem delu smo upoštevali prispevek Kovače­vića (1971), ki je objavil razširjenost vseh do takrat znanih vrst rodu Otiorhynchus v Jugoslaviji. Pri svojem delu je zbral tudi vse podatke iz zbirk v Sloveniji in v Trstu. To so zbirke A. Gspana, V. Kodriča, G. Depolia, E. Pretnerja in J. Müllerja; med temi koleopterologi se je z rilčkarji ukvarjal predvsem V. Kodrič. Vsi podatki, ki jih je uporabil Kovačević v svojem delu, so bili zbra­ni pred drugo svetovno vojno; njihova določitev ni popolnoma zanesljiva. Menimo, da so nadaljnje sistematične favnistične raziskave rilčkarjev v Sloveniji zelo potrebne in bodo dale še več novih podatkov o razširjenosti in pojavlja­nju vrst iz te skupine hroščev pri nas. Katalog rilčkar­jev Slovenije je v pripravi. 2 MATERIAL IN METODE DELA Objekt: vrste hroščev rilčkarjev iz rodu Otiorhynchus s. lat. Metode dela: Terensko vzorčenje smo izvajali po UTM metodologiji kartiranja favne Evrope in Sloveni­je (Sivec 1980, Carnelutti, Gogala & Sivec 1982) v okviru temeljnih raziskav naravne dediščine Slovenije (temeljni raziskovalni program Favna, flora in vegeta­cija Slovenije in sosednjih območij). Material: pri terenskem vzorčenju zbrane osebke hroščev smo preparirali in so shranjeni v koleopterolo­ški zbirki Biološkega inštituta Jovana Hadžija. Pri ob­delavi podatkov smo uporabili tudi primerke iz osebne zbirke hroščev V. Furlana, ki je je shranjena v Prirodo­slovnem muzeju Slovenije. Podatki: zbrani terenski podatki in podatki iz zbirk so digitalizirani in vneseni v podatkovno bazo FloVegSi (Favna, flora in vegetacija Slovenije) Biološke­ga inštituta Jovana Hadžija ZRC SAZU. Kartografsko gradivo: arealne karte (po UTM mreži) za Slovenijo so bile narejene s pomočjo aplikacije FloVegSi (Seliškar T., Vreš & A. Seliškar 2003). Obdobje raziskav: zadnji dve desetletji 20. stoletja in do danes. Nomenklatura: Reitter E. 1913, Winkler A. 1927 – 1932, Freude & al. 1981, Lucht 1987, Fauna Europaea Web Service (Alonso-Zarazaga 2004). 3 REZULTATI IN RAZPRAVA V preteklih dveh desetletjih smo zabeležili 17 novih vrst za Slovenijo. To so: Dodecastichus aurosignatus (Apfelbeck 1889) = Otiorhynchus aurosignatus Apfel­beck 1889, D. brevipes (Apfelbeck 1894) = Otiorhynchus brevipes Apfelbeck, 1894, D. dalmatinus (Gyllenhal 1834), D. heydeni (Stierlin 1861) = Otiorhynchus heyde­ni Stierlin 1861, Otiorhynchus alutaceus (Germar 1817), O. azaleae Penecke 1894, O. brusinae Stierlin 1888, O. pantherinus Apfelbeck 1898, O. porcatus (Herbst 1795), O. primigenius Apfelbeck 1918, O. remotegranulatus Stierlin 1891, O. rugifrons (Gyllenhal 1813), O. subden­tatus Bach 1834, O. tenebricosus (Herbst 1784), O. trun­catus Stierlin 1861, O. turca Boheman 1843 in O. unci­natus Germar 1824. Skupno število trenutno znanih vrst iz rodov Dodecastichus in Otiorhynchus s. str. v Sloveniji je 103 (B. Drovenik, neobjavljeno). V pri­spevku je vsaka vrsta predstavljena z arealno karto po UTM sistemu (iz baze FloVegSi) in opisom razširjeno­sti za Slovenijo ter kratkim opisom njenega pojavljanja na Balkanskem polotoku in v Evropi (Kovačević 1971, Alonso-Zarazaga 2004), osem vrst je predstavljenih tudi slikovno (Slika 18 A – H). Pregled vrst: Dodecastichus aurosignatus (Sliki 1 in 18 A): vrsta je razširjena po gorovjih Balkanskega polotoka v Ro­muniji, Bolgariji, Albaniji in po vseh republikah nek­danje Jugoslavije, razen Slovenije, kjer doslej še ni bila poznana (Kovačević 1971, Alonso-Zarazaga 2004). Kovačević (1971: 7-8) jo obravnava kot gorsko vrsto; doslej najbližje nahajališče je Gorski Kotar (Lokve). Iz Slovenije je znanih 5 nahajališč; pojavlja v alpskem in predalpskem svetu z enim nahajališčem celo v subpa­nonskem svetu (Slika 1). Nahajališča: 33T VM63 (9553/4) Slovenija, Štajerska, Savinjske Alpe, Matkov kot. 1049 m n. m. 5.7.1994. Leg. Drovenik B. 33T VM33 (9650/2) Slovenija, Gorenjska, Moste (Jese­nice), Piškotarjev most. 460 m n. m. 17.7.1996. Leg. Drovenik B. 33T VM33 (9650/2) Slovenija, Gorenjska, Bled, pri mostu čez Savo. 436 m n. m. 23.7.1996. Leg. Drove­nik B. 33T VM71 (9853/2) Slovenija, Gorenjska, Volčji Potok, Šmarca. 340 m n. m. 6.4.1996. Leg. Drovenik B. 33T WM76 (9361/2) Slovenija, Štajerska, Police (Gornja Radgona), 290 m n. m. 2.7.2000. Leg. Drovenik B. Dodecastichus brevipes (Sliki 2 in 18 B): vrsta je raz­širjena v osrednjem delu Balkanskega polotoka tako v nižinah kot višjih legah (Kovačević 1971: 8). Iz sever­nega dela Dalmacije in Istre še ni podatkov o razširje­nosti te vrste. V Sloveniji poznamo tri nova nahajališča: 33T VL14 (0449/2) Slovenija, Primorska, Kozina, Tu­blje pri Hrpeljah. 500 m n. m. 13.6.1999. Leg. Hol­zer E., Det. Behne L. 33T VL54 (0452/2) Slovenija, Snežnik, vrh. 1705 m n. m. 18.6.1997. Leg. Drovenik B. 33T VL24 (0450/3) Slovenija, Primorska, Gradišče, Golac, Velika Pleševica. 20.6.1996. Leg. Drovenik B. Vsa tri nahajališča so iz jugozahodne Slovenije, iz česar lahko sklepamo, da je vrsta verjetno razširjena tudi v severni Dalmaciji in Istri. Vrsta ima v Sloveniji montanski karakter – na Krasu (pri Kozini) dosega se­verozahodno mejo svojega areala. Dodecastichus dalmatinus (Slika 3): vrsta je razšir­jena vzdolž jadranske obale (Italija, Hrvaška, Bosna in Hercegovina – Alonso-Zarazaga 2004, Fauna Euro­paea). Za nekdanjo Jugoslavijo jo navaja že Kovačević (1971: 16), ki jo obravnava kot podvrsto O. pulverulen­tus subsp. dalmatinus; doslej najbližje nahajališče na­vaja za hrvaško Istro (Opatija), pojavlja pa se vse od Kvarnerja do Črne gore. Slednja v Favni Evrope ni omenjena (Alonso-Zarazaga 2004). V Sloveniji je bila najdena le v skrajnem jugozahodnem delu, v slo­venski Istri ob meji s Hrvaško. Nahajaličše: 33T VL24 (0450/3) Slovenija, Primorska, Golac, Velika Pleševica. 13.6.1999. Leg. Holzer E., det. L. Behne. Dodecastichus heydeni (Sliki 4 in 18 C): vrsta je razširjena v osrednjem delu Balkanskega polotoka – Dalmacija (obalni del), Bosna in Hercegovina ter na Apeninskem polotoku (Kovačević 1971: 12). Najdemo jo od nižin do montanskega pasu; v Sloveniji živi na krasu, kjer dosega severozahodno mejo svojega areala. Nahajališča: 33T VL07 (0148/4) Slovenija, Kras, Mali Dol (Komen). 220 m n. m. 12..9.1981. Leg. Drovenik B. 33T VL13 (0549/2) Slovenija, Kras, Veliki Badin, Mlini. 370 m n. m. 4.9.1990. Leg. Furlan V. 33T VL25 (0349/2) Slovenija, Kras, Škocjanske jame, Sapendol. 360 m n. m. 22.7.2004. Leg. Drovenik B. Otiorhynchus alutaceus (Sliki 5 in 18 F): Kovače­vič (1971: 22) jo označuje kot “transadriatično” vrsto, ki je razširjena le v Mediteranskem območju Istre in Dalmacije. Obravnava jo kot gospodarsko pomembno vrsto (»škodljivec«) na vinski trti. Glede na razširjenost vrste po Fauna Europaea (Alonso-Zarazaga 2004) se vrsta pojavlja predvsem na Balkanskem (Hrvaška, Gr­čija in Bolgarija) in Apeninskem polotoku (Italija), na­vedena pa je tudi iz Slovaške (verjetno sekundarno po­javljanje v vinogradih). V Sloveniji živi v submedite­ranskem območju predvsem v vinogradniških nasadih. Nahajališča: 33T UL94 (0447/4) Slovenija, Istra, Strunjan/Strugna­no, Ronek. 28 m n. m. 4.9.1998. Leg. Furlan V. 33T VL07 (0148/4) Slovenija, Kras, Mali Dol (Komen). 260 m n. m. 10.5.1981. Leg. Drovenik B. 33T VL25 (0349/2) Slovenija, Kras, Škocjanske jame, Schmidlova dvorana. 313 m n. m. 5.7.2001. Leg. Drovenik B. 33T VL14 (0449/1) Slovenija, Kozina, Petrinjski kras, 428 m n. m. 20.5.1995. Leg. Drovenik B. 33T VL25 (0349/2) Slovenija, Kras, Divača. 452 m n. m. 19.5.1979. Leg. Furlan V. 33T VL16 (0249/4) Slovenija, Primorska, Kras, Divača, Brestovica pri Povirju, 385 m n. m. 16.5.1984. Leg. Drovenik B. Otiorhynchus azaleae (Slika 6): visokogorska vrsta, poznana iz centralnih in vzhodnih Alp (Avstrija, Itali­ja in Švica, Alonso-Zarazaga 2004, Fauna Europa­ea); njena hranilna rastlina je alpska azaleja - Loiseleu­ria procumbens (Holdhaus 1954: 106). Za Slovenijo jo sicer že navaja Kovačević (1971: 24) na nahajališčih Sovatna (Julijske Alpe), Kropa in Tržič (vir njegovih podatkov nam ni poznan). Glede na njen visokogorski karakter sta nahajališči Kropa in Tržič dvomljivi. Do­slej edino zanesljivo (s primerki potrjeno) nahajališče v Sloveniji je na visokogorskih travnikih Mokrice (Ka­mniško-Savinjske Alpe, Drovenik 2000: 124, Drove­nik 2006: 244). Nahajališče: 33T VM62 (9653/3) Slovenija, Savinjske Alpe, Mokrica, vrh. 1778 m n. m. 14.7.1976. Leg. Drovenik B. Otiorhynchus brusinae (Slika 7): gorska do visoko­gorska vrsta; Kovačević (1971: 53) jo najbližje Sloveni­ji navaja za Gorski Kotar (Hrvaška) in Julijske Alpe (Italija; Alonso-Zarazaga 2004). Pri nas je bila doslej najdena le v mrazišču Paradana v Trnovskem gozdu, verjetno pa je, da živi tudi na Javornikih in Snežniku. Nahajališče: 33T VL09 (0049/1) Slovenija, Trnovski gozd, Paradana. 1100 m n. m. 10.6.1978. Leg. Furlan V. Otiorhynchus pantherinus (Sliki 8 in 18 H): je su­balpinska do alpinska vrsta. Kovačević (1971: 62) jo navaja za gorovja Balkanskega polotoka (Vranica, Bje­lašnica, Volujak, Maglič, Koritnik in Ljuboten na Šar­planini). Glede na Fauna Europaea je razširjena v na­slednjih državah: Bosna in Hercegovina, Srbija in Črna gora ter Albanija (Alonso-Zarazaga 2004). Nahaja­lišče v Julijskih Alpah predstavlja skrajno severozaho­dno mejo njenega areala. Nahajališči: 33T VM22 (9750/3) Slovenija, Julijske Alpe, Soriška planina, Možic. 1448 m n. m. 10.6.1979. Leg. Pre­tner E. 33T VL54 (0452/2) Slovenija, Notranjska, Snežnik, vrh. 1780 m n. m. 1.6.2000. Leg. Drovenik B., det. Behne L. Otiorhynchus porcatus (Slika 9): je zahodno, sre­dnje in severnoevropska vrsta (glej Fauna Europaea, Alonso-Zarazaga 2004), ki je Kovačević (1971) ne navaja za nekdanjo Jugoslavijo. Je petrofilna, terikolna in polifagna vrsta, ki je aktivna ponoči in se podnevi skriva pod kamenjem ali rastlinskimi ostanki (Hold­haus 1954: 259). Pri nas ima subalpinski karakter; do­slej je bila najdena le na Soriški planini, kar domnevno predstavlja jugovzhodno mejo njenega areala. Nahajališče: 33T VM22 (9750/3) Slovenija, Julijske Alpe, Soriška planina. 1423 m n. m. 8.7.1982. Leg. Furlan V. Otiorhynchus primigenius (Slika 10): južnobalkan­ska vrsta s subalpinskim karakterjem, ki je bila opisa­na z Bogičevice (Črna Gora) na nadmorski višini 2530 m kot podvrsta O. austriacus subsp. primigenius Apfel­beck 1918 (Kovačević 1971: 52). Drovenik jo je našel tudi na nahajališču Dobri do na Durmitorju (julij 1980). Fauna Europaea za razširjenost vrste prikazuje samo Albanijo (Alonso-Zarazaga 2004). Furlanova najdba te vrste v osrednji Sloveniji (Zasavje) je presene­tljiva in morda le naključna. Nahajališče: 33T VM80 (9955/1) Slovenija, Dolenjska, Breg pri Litiji. 245 m n. m. 21.8.1991. Leg. Furlan V. Otiorhynchus remotegranulatus (Slika 11): Kova­čević (1971) vrste ne navaja za nekdanjo Jugoslavijo. V Sloveniji ima montanski do subalpinski karakter. Glede na razširjenost vrste po Fauna Europaea (Alon­so-Zarazaga 2004) je doslej poznana iz Madžarske in Romunije. Nahajališča: 33T VL19 (0049/1) Slovenija, Trnovski gozd, Paradana. 1143 m n. m. 10.6.1978. Leg. Drovenik B. 33T VL95 (0355/2) Slovenija, Kočevski Rog. 596 m n. m. 4.5.1979. Leg. Drovenik B. 33T VM41 (9851/2) Slovenija, Škofja Loka, Lubnik. 850 m n. m. 29.6.1978. Leg. Furlan V. Otiorhynchus rugifrons (Slika 12): Glede na prika­zano razširjenost v Fauna Europaea (Alonso-Zara­zaga 2004) je vrsta razširjena v zahodni, srednji in severni Evropi. Kovačević (1971) je ne navaja za nek­danjo Jugoslavijo. V Sloveniji ima subalpinski karak­ter; doslej je bila najdena le v Kamniško-Savinjskih Alpah (Drovenik 2006: 245). Drugod po Evropi je po­znana tudi kot gospodarsko pomembna vrsta (»ško­dljivec«) v jagodovih nasadih. Nahajališče: 33T VM63 (9653/1) Slovenija, Savinjske Alpe, Okrešelj. 1563 m n. m. 1.7.1992. Leg. Drovenik B. Otiorhynchus subdentatus (Slika 13 in 18 E): Glede na prikazano razširjenost v Fauna Europaea je vrsta razširjena v zahodni in srednji Evropi, južno od Slo­venije pa je znana le na Hrvaškem (Alonso-Zaraza­ga 2004). Kovačević (1971) je ne navaja za nekdanjo Jugoslavijo. V Sloveniji ima montanski do subalpinski karakter. Doslej je bila najdena v Kamniško-Savinj­skih (Drovenik 2006: 245) in vzhodnih Julijskih Alpah. Nahajališča: 33T VM93 (9655/2) Slovenija, Savinjske Alpe, Šmihel nad Mozirjem. 940 m n. m. 12.7.1996. Leg. Drove­nik B. 33T VM63 (9653/1) Slovenija, Savinjske Alpe, Logarska Dolina, Okrešelj. 1530 m n. m. 1.7.1992. Leg. Fur­lan V. 33T VM31 (9750/4) Slovenija, Gorenjska, Julijske Alpe, Ratitovec, Prtovč. 940 m n. m. 10.6.1979. Leg. Fur­lan V. Otiorhynchus tenebricosus (Slika 14): Glede na pri­kazano razširjenost v Fauna Europaea (Alonso-Za­razaga 2004) je vrsta razširjena v večjem delu osre­dnje Evrope. Kovačević (1971: 27) jo navaja samo za Hrvaško (Štirovaća na Velebitu) na osnovi primerka iz Prirodoslovnega muzeja Slovenije v Ljubljani. Edino doslej znano nahajališče v Sloveniji je s Pohorja, kjer je bila vrsta najdena v smrekovem gozdu. 33T WM24 (9557/2) Slovenija, Pohorje, Jezerski vrh. 1530 m n. m. 29.6.1988. Leg. Kippenberg H. 33T WM24 (9557/2) Slovenija, Pohorje, Jezerski vrh. 1500 m n. m. 28.6.1988. Leg. Kahlen M. Otiorhynchus truncatus (Slika 15): Glede na prika­zano razširjenost v Fauna Europaea (Alonso-Zara­zaga 2004) je vrsta razširjena v srednji Evropi (Češka in Slovaška) in na Balkanskem polotoku (Albanija, Hr­vaška, Bosna in Hercegovina). Kovačević (1971: 38) jo obravnava kot mediteransko montansko vrsto, razšir­jeno od Snježnika (Hrvaška) do Orjena (Črna gora). Novi nahajališči v južni Sloveniji se navezujeta na bal­kanska. Nahajališči: 33T VL75 (0354/1) Slovenija, Velika gora, Turn. 1220 m n. m. 2.6.1978. Leg. Drovenik B. 33T VL54 (0452/1) Slovenija, Snežnik: Sviščaki. 1240 m n. m. 14.5.1996. Leg. Freude H. Otiorhynchus turca (Sliki 16 in 18 G): Vrsta je bila opisana iz Anatolije (locus typicus). Glede na prikaza­no razširjenost v Fauna Europaea (Alonso-Zarazaga 2004) je poznana iz Italije, Hrvaške ter jugovzhodnih delov Balkanskega polotoka (Grčija, Bolgarija, Turčija) in Kavkaza (Winkler 1927-1932). Mihajlova in sode­lavci (Mihajlova et al. 2008) jo naštevajo med primer­ki shranjenimi v Sarajevskem muzeju, vendar brez na­hajališč. Kovačević (1971) je za nekdanjo Jugoslavijo ne navaja. Njeni nahajališči pri nas sta verjetno na­ključni (prinešena vrsta) ali pa se je pojavila kot gospo­darsko pomembna vrsta (»škodljivec« ) v vinogradih. Nahajališči: 33T VL14 (0449/2) Slovenija, Primorska, Kozina: Tu­blje pri Hrpeljah. 500 m n. m. 13.6.1999. Leg. Hol­zer E., det. Behne L. 33T VL04 (0448/1) Slovenija, Istra, Ankaran /Ancara­no. 30 m n. m. 20.7.1997. Leg. Drovenik B. Otiorhynchus uncinatus (Sliki 17 in 18 D): Vrsta je razširjena predvsem v osrednji in severni Evropi (Alonso-Zarazaga 2004). Kovačević (1971) je za nekdanjo Jugoslavijo ne omenja. V Sloveniji ima mon­tanski karakter; najdena je bila na območju Julijskih in Kamniško-Savinjskih Alp (Drovenik 2006: 246), kar glede na sedanje poznavanje njene razširjenosti pred­stavlja jugovzhodno mejo njenega areala. Nahajališči: 33T VM73 (9654/1) Slovenija, Savinjske Alpe: Solčava, Rogovilec. 674 m n. m. 12.7.1983. Leg. Furlan V. 33T UM92 (9747/4) Slovenija, Julijske Alpe: Kobarid, Ladra, Ladrski vrh. 546 m n. m. 14.5.2002. Leg. Drovenik B. 4 ZAKLJUČKI Favna rilčkarjev Slovenije je dopolnjena s seznamom 17 vrst iz rodov Dodecastichus in Otiorhynchus s.str. Vseh doslej znanih vrst rodu Otiorhynchus s.lat. za Slo­venijo je 103. Razširjenost novo ugotovljenih vrst rodu Oti­orhynchus za Slovenijo dobro dopolnjuje areal teh vrst v Evropi; njihovo pojavljanje predstavlja bodisi skrajno južno mejo areala za (severno)evropske vrste, skrajno severozahodno mejo areala za balkanske vrste ali osre­dnji – vezni del areala za jugovzhodno evropske vrste. 5 SUMMARY 5.1 Introduction Weevils (Curculionidae) of Slovenia, including the genus Otiorhynchus, Germar 1824, are still a poorly known group of animals. This is a very species rich group of beetles that numbers more than 5000 differ­ent taxons in Europe, and more than 900 species have been identified in Slovenia. The group of weevils is fairly demanding for recognition (identification) and there are also few researchers who deal with their tax­onomy and distribution, both in Europe and in Slove­nia. The genus Otiorhynchus belongs among the most species rich genuses of weevils in Europe, where 864 taxons (species and sub-species) of the genus Otiorhyn­chus s.str., or 934 taxons of the genus Otiorhynchus s.lat. (Alonso-Zarazaga 2004) are currently known. In Slovenia, according to date known to date the genus Otiorhynchus s.lat. is represented by 103 species and sub-species (B. Drovenik, unpublished), which is 11 % of known European taxons. In a work by Kovače­vić (1971), 84 are stated for Slovenia, and 228 species for the entire (former) Yugoslavia. Research over several years of beetle fauna in Slo­venia has shown that recognition of this large group of insects is still very incomplete and we have also col­lected to date a lot of new data for weevils. We found 17 new species of the genus Otiorhynchus, which we pre­sent in this contribution with maps of their distribu­tion and short descriptions, 8 of them also with pic­tures. This genus has numerous species that are bound to a strictly limited geographic area: many of them are endemics or have disjunct areas of distribution. One cause of the fairly large number of endemics in this group of beetles, among others, is that individuals of these species do not fly because they are wingless spe­cies or have fused wings (coverings). Many of them are monophagous, feeding exclusively on a specific plant species, which may have a very limited distribution. In recent decades, weevils in Slovenia have been researched above all by S. Brelih, V. Furlan, B. Kofler and M. Zdešar. We also used some of their data in this contribution. Some individual data were also contrib­uted by foreign experts, such as E. Holzer, M. Kahlen, H. Kippenberg and H. Freude. B. Drovenik collected the majority of the presented data for weevils, or the genus Otiorhynchus in Slovenia. Identification or re­view of identification of these animals has been per­formed by leading European experts who study these beetles, such as L. Behne and O. Vořišek. We took into account in our work the contribution of Kovačević (1971), who published the distribution of all species of the genus Otiorhynchus in (former) Yu­goslavia known at that time. He also collected in his work all data from collections in Slovenia and in Tri­este. These are the collections of A. Gspan, V. Kodrič, G. Depoli, E. Pretner and J. Müller; among these co­leopterologists, V. Kodrič dealt with weevils. All data that Kovačević used in his work were collected before the Second World War and their identification may be dubious. We believe that further systematic faunistic re­search of weevils in Slovenia is very necessary and will provide even more new data on the distribution and occurrence here of species from this group of beetles. A catalogue of weevils is in preparation. 5.2 Materials and Methods Object: species of weevil beetles from the genus Oti­orhynchus s. lat. Methods of work: Field sampling was performed by the methodology of mapping the fauna of Europe and Slovenia (Sivec 1980) within the framework of basic research of the natural heritage of Slovenia (basic program of research: Fauna, flora and vegetation of Slovenia and neighbouring regions). Material: in field sampling, the collected material (beetle individuals) is prepared and stored in the co­leopterological collection of the Jovan Hadži Biological Institute. In processing the data, we also used the data of V. Furlan in the basic material of his private collec­tion of beetles (the collection is kept at the Natural History Museum of Slovenia). Data: the collected field data and data from collec­tions have been digitalised and entered into the data­base FloVegSi (Fauna, flora and vegetation of Slovenia) of the Jovan Hadži Biological Institute of ZRC SAZU. Cartographic material: the distribution maps for Slo­venia were made with the help of the application FloVegSi (Seliškar & al. 2003). Period of research: last two decades of the 20th century and to the present. Nomenclature: Reitter E. 1913, Winkler A. 1927 – 1932, Freude & al. 1981, Lucht 1987, Fauna Euro­paea Web Service (Alonso-Zarazaga 2004). 5.3 Results and Discussion In the past two decades we have recorded 17 new spe­cies for Slovenia. These are: Dodecastichus aurosigna­tus (Apfelbeck 1889) = Otiorhynchus aurosignatus Ap­felbeck 1889 (Figures 1 and 18 A), D. brevipes (Apfel­beck 1894) = Otiorhynchus brevipes Apfelbeck (Figures 2 and 18 B), 1894, D. dalmatinus (Gyllenhal 1834) (Fig­ure 3), D. heydeni (Stierlin 1861) = Otiorhynchus heyde­ni Stierlin 1861 (Figures 4 and 18 C), Otiorhynchus alu­taceus (Germar 1817) (Figures 5 and 18 F), O. azaleae Penecke 1894 (Figure 6), O. brusinae Stierlin 1888 (Fig­ure 7), O. pantherinus Apfelbeck 1898 (Figures 8 and 18 H), O. porcatus (Herbst 1795) (Figure 9), O. primigenius Apfelbeck 1918 (Figure 10), O. remotegranulatus Stier­lin 1891 (Figure 11), O. rugifrons (Gyllenhal 1813) (Fig­ure 12), O. subdentatus Bach 1834 (Figures 13 and 18 E), O. tenebricosus (Herbst 1784) (Figure 14), O. trunca­tus Stierlin 1861 (Figure 15), O. turca Boheman 1843 (Figures 16 and 18 G) and O. uncinatus Germar 1824 (Figures 17 and 18 D). The total number of currently known species from the genuses Dodecastichus and Otiorhynchus s. str. In Slovenia is 103 (B. Drovenik, unpublished). Each species is presented in the contri­bution with a map of distribution from the FloVegSi database (Figures 1-17) and a description of its distri­bution for Slovenia and a brief description of its occur­rence in the Balkan Peninsular and in Europe (Kova­čević 1971, Alonso-Zarazaga 2004); eight species are presented with a picture (Figure 18 A – H). 5.4 Conclusions Weevil fauna in Slovenia has been supplemented with a list of 17 species from the genuses Dodecastichus and Otiorhynchus s.str. There are 103 species of the genus Otiorhynchus s.lat. known to date for Slovenia. The distribution of the newly established species of the genus Otiorhynchus for Slovenia is well included in (supplements) the area of distribution of these spe­cies in Europe; their occurrence represents either the extreme southern boundary of the area of distribution of (northern) European species, the extreme north-westerly boundary of the area of distribution of Balkan species or the central – connected part of the area of distribution of south-eastern European species. ZAHVALA – ACKNOWLEDGEMENTS Za uporabo posameznih podatkov se zahvaljujeva V. Furlanu, E. Holzerju, M. Kahlenu, H. Kippenbergu in H. Freudeju, za določitev izbranih vrst pa L. Behneju. Dr. Matjažu Kuntnerju se zahvaljujeva za pomoč pri fotografiji osebkov iz zbirke, dr. Matjažu Gregoriču pa za pripravo slike s fotografijami izbranih vrst hroščev. Raziskavo je sofinancirala ARRS v sklopu temeljnih raziskav Biološkega inštituta Jovana Hadžija ZRC SAZU v Ljubljani (P1-0236). 6 LITERATURA – REFERENCES Alonso-Zarazaga, M. A., 2004: Fauna Europaea: Coleoptera, Curculionidea. Fauna Europaea version 1.1, http://www.faunaeur.org Carnelutti, J., M. Gogala & I. Sivec, 1982: Kartiranje Favne Slovenije – navodila. Prirodoslovni muzej Sloveni­je, Ljubljana, 9 str. Dieckmann, L., 1980: Beiträge zur Insektenfauna der DDR: Coleoptera – Curculionidae (Brachycerinae, Otiorhyn­chinae, Brachyderinae). Beitr. Ent. (Berlin) 30 (1): 145-310. Drovenik, B., 2000: Novosti v favni hroščev rilčkarjev (Curculionidae, Coleoptera) na Kamniškem. Kamniški zbornik XV, Kamnik, str. 123-128. Drovenik, B., 2006: Hrošči rilčkarji (Coleoptera, Curculionidea) Kamniško-Savinjskih Alp. Kamniški zbornik XVIII, Kamnik, str. 239-253. Fauna Europaea Web Service, 2004: Fauna Europaea version 1.1, Available online at http://www.faunaeur.org. Freude, H., K. W. Harde & G. A. Lohse, 1981: Die Käfer Mitteleuropas. Band 10. Goecke & Evers Verlag, Krefeld. 310 str. Heath, J., 1971: Instructions for Recorders Europaean Invertebrate Surwey. Biological Records Centre, Abbots Rip­ton, 23 str. Heath, J. & F. Perring, 1975: Biological recording in Europe. Endeavour 34: 103-108. Holdhaus, K., 1954: Die Spuren der Eiszeit in der Tierwelt Europas. Universitätsverlag Wagner, Insbruck. 1 karta, 493 str. Koch, K., 1992: Die Käfer Mitteleuropas. Ökologie. Band 3. Goecke & Evers Verlag, Krefeld. 389 str. Kovačević, Ž., 1971: Otiorrhynchus vrste i njihovo rasprostranjenje u Jugoslaviji (Coleoptera – Curculionidae). Acta Instituti protectionis plantarum Facultatis agronomicae / Rad Instituta za zaštitu bilja poljoprivrednog fakulteta, Zagreb, 103 str. Lucht, W., 1987: Die Käfer Mitteleuropas. Katalog. Goecke & Evers Verlag, Krefeld. 342 str. Mihajlova, B., S. Pešić & D. Kotrošan, 2008: Otiorhynchini (Coleoptera: Curculionidae, entiminae) in the collec­tion of the National Museum of Bosnia and Herzegovina. Arch. Biol. Sci. (Beograd) 60(4): 713-725. Reitter, E., 1913: Bestimungstabellen der Otiorrhynchus-Arten mit gezähnten Schenkeln aus der palaearktischen Fauna. Genus Dorymerus und Tourineria. Verlag von Edmund Reitter, Brünn, str. 1-123. Reitter, E., 1913: Bestimungstabellen der Otiorrhynchus-Arten mit ungezähnten Schenkeln aus der palaearktischen Fauna. Genus Otiorrhynchus Germ. Verlag von Edmund Reitter, Paskau, str. 25-118. Seliškar, T., B. Vreš & A. Seliškar, 2003: FloVegSi 2.0. Računalniški program za urejanje in analizo bioloških podatkov. Biološki inštitut ZRC SAZU, Ljubljana. Sivec, I., 1980: Kartiranje nevretenčarjev Evrope. Predstavitev in razlaga. Biol. vestnik (Ljubljana) 28 (2): 169-194. Winkler, A., 1927 – 1932: Catalogus Coleopterorum regionis palaearcticae II. Albert Winkler, Wien, str. 1137-1698. 1 Dr., Tunjiška cesta 2, SI-1240 Kamnik 2 Dr., Biološki inštitut Jovana Hadžija ZRC SAZU, Novi trg 2, SI-1001 Ljubljana, Slovenija, e-mail: branevr@zrc-sazu.si Slika 1: Razširjenost Dodecastichus aurosignatus v Sloveniji. Figure 1: Distribution of Dodecastichus aurosignatus in Slovenia. Slika 2: Razširjenost Dodecastichus brevipes v Sloveniji. Figure 2: Distribution of Dodecastichus brevipes in Slovenia. Slika 3: Razširjenost Dodecastichus dalmatinus v Sloveniji. Figure 3: Distribution of Dodecastichus dalmatinus in Slovenia. Slika 5: Razširjenost Otiorhynchus alutaceus v Sloveniji. Figure 5: Distribution of Otiorhynchus alutaceus in Slovenia. Slika 4: Razširjenost Dodecastichus heydeni v Sloveniji. Figure 4: Distribution of Dodecastichus heydeni in Slovenia. Slika 6: Razširjenost Otiorhynchus azaleae v Sloveniji. Figure 6: Distribution of Otiorhynchus azaleae in Slovenia. Slika 8: Razširjenost Otiorhynchus pantherinus v Sloveniji. Figure 8: Distribution of Otiorhynchus pantherinus in Slovenia. Slika 7: Razširjenost Otiorhynchus brusinae v Sloveniji. Figure 7: Distribution of Otiorhynchus brusinae in Slovenia. Slika 9: Razširjenost Otiorhynchus porcatus v Sloveniji. Figure 9: Distribution of Otiorhynchus porcatus in Slovenia. Slika 11: Razširjenost Otiorhynchus remotegranulatus v Sloveniji. Figure 11: Distribution of Otiorhynchus remotegranulatus in Slovenia. Slika 10: Razširjenost Otiorhynchus primigenius v Sloveniji. Figure 10: Distribution of Otiorhynchus primigenius in Slovenia. Slika 12: Razširjenost Otiorhynchus rugifrons v Sloveniji. Figure 12: Distribution of Otiorhynchus rugifrons in Slovenia. Slika 14: Razširjenost Otiorhynchus tenebricosus v Sloveniji. Figure 14: Distribution of Otiorhynchus tenebricosus in Slovenia. Slika 13: Razširjenost Otiorhynchus subdentatus v Sloveniji. Figure 13: Distribution of Otiorhynchus subdentatus in Slovenia. Slika 15: Razširjenost Otiorhynchus truncatus v Sloveniji. Figure 15: Distribution of Otiorhynchus truncatus in Slovenia. Slika 16: Razširjenost Otiorhynchus turca v Sloveniji. Figure 16: Distribution of Otiorhynchus turca in Slovenia. Slika 17: Razširjenost Otiorhynchus uncinatus v Sloveniji. Figure 17: Distribution of Otiorhynchus uncinatus in Slovenia. Sl. 18: Osebki hroščev rilčkarjev iz zbirke Biološkega inštituta ZRC SAZU (Ljubljana): A – Dodecastichus aurosignatus, B – Do­decastichus brevipes, C – Dodecastichus heydeni, D – Otiorhynchus uncinatus, E – Otiorhynchus subdentatus, F – Otiorhyn­chus alutaceus, G – Otiorhynchus turca in H – Otiorhynchus pantherinus. Merilo = 5 mm. Foto: B. Vreš Fig. 18: Specimens from the weevil beetles collection of the Institut of biology ZRC SAZU (Ljubljana): A – Dodecastichus auro­signatus, B – Dodecastichus brevipes, C – Dodecastichus heydeni, D – Otiorhynchus uncinatus, E – Otiorhynchus subdentatus, F – Otiorhynchus alutaceus, G – Otiorhynchus turca in H – Otiorhynchus pantherinus. Scale bar = 5 mm. Photo: B. Vreš NAVODILA AVTORJEM Folia biologica et geologica so znanstvena revija IV. razreda SAZU za naravoslovne vede. Objavljajo naravoslovne znanstvene razprave in pregledne članke, ki se nanašajo predvsem na raziskave v našem etničnem območju Slovenije, pa tudi raziskave na območju Evrope in širše, ki so pomembne, potrebne ali primerljive za naša preučevanja. 1. ZNANSTVENA RAZPRAVA Znanstvena razprava zajema celovit opis izvirne raziskave, ki vključuje teoretični pregled tematike, po­drobno predstavlja rezultate z razpravo in zaključki ali sklepi in pregled citiranih avtorjev. V izjemnih primerih so namesto literaturnega pregleda dovoljeni viri, če to zahteva vsebina razprave. Razprava naj ima klasično razčlenitev (uvod, ma­terial in metode, rezultati, diskusija z zaključki, zahvale, literatura idr.). Dolžina razprave, vključno s tabelami, grafikoni, tablami, slikami ipd., praviloma ne sme presegati 2 avtor­skih pol oziroma 30 strani tipkopisa. Zaželene so razpra­ve v obsegu ene avtorske pole oziroma do dvajset strani tipkopisa. Razpravo ocenjujeta recenzenta, od katerih je eden praviloma član SAZU, drugi pa ustrezni tuji strokovnjak. Recenzente na predlog uredniškega odbora revije Folia biologica et geologica potrdi IV. razred SAZU. Razprava gre v tisk, ko jo na predlog uredniškega odbora na seji sprejmeta IV. razred in predsedstvo SAZU. 2. PREGLEDNI ČLANEK Pregledni članek objavljamo po posvetu uredniške­ga odbora z avtorjem. Na predlog uredniškega odbora ga sprejmeta IV. razred in predsedstvo SAZU. Članek naj praviloma obsega največ 3 avtorske pole (tj. do 50 tipka­nih strani). 3. NOVOSTI Revija objavlja krajše znanstveno zanimive in aktu­alne prispevke do 7000 znakov. 4. IZVIRNOST PRISPEVKA Razprava oziroma članek, objavljen v reviji Folia biologica et geologica, ne sme biti predhodno objavljen v drugih revijah ali knjigah. 5. JEZIK Razprava ali članek sta lahko pisana v slovenščini ali katerem od svetovnih jezikov. V slovenščini zlasti te­daj, če je tematika lokalnega značaja. Prevod iz svetovnih jezikov in jezikovno lektorira­nje oskrbi avtor prispevka, če ni v uredniškem odboru dogovorjeno drugače. 6. POVZETEK Za razprave ali članke, pisane v slovenščini, mora biti povzetek v angleščini, za razprave ali članke v tujem jeziku ustrezen slovenski povzetek. Povzetek mora biti do­volj obširen, da je tematika jasno prikazana in razumljiva domačemu in tujemu bralcu. Dati mora informacijo o na­menu, metodi, rezultatu in zaključkih. Okvirno naj pov­zetek zajema 10 do 20 % obsega razprave oziroma članka. 7. IZVLEČEK Izvleček mora podati jedrnato informacijo o na­menu in zaključkih razprave ali članka. Napisan mora biti v slovenskem in angleškem jeziku. 8. KLJUČNE BESEDE Število ključnih besed naj ne presega 10 besed. Pred­staviti morajo področje raziskave, podane v razpravi ali članku. Napisane morajo biti v slovenskem in angleškem jeziku. 9. NASLOV RAZPRAVE ALI ČLANKA Naslov razprave ali članka naj bo kratek in razum­ljiv. Za naslovom sledi ime/imena avtorja/avtorjev (ime in priimek). 10. NASLOV AVTORJA/AVTORJEV Pod ključnimi besedami spodaj je naslov avtorja/avtorjev, in sicer akademski naslov, ime, priimek, ustano­va, mesto z oznako države in poštno številko, država, ali elektronski poštni naslov. 11. UVOD Uvod se mora nanašati le na vsebino razprave ali članka. 12. ZAKLJUČKI ALI SKLEPI Zaključki ali sklepi morajo vsebovati sintezo glavnih ugotovitev glede na zastavljena vprašanja in razrešujejo ali nakazujejo problem raziskave. 13. TABELE, TABLE, GRAFIKONI, SLIKE IPD. Tabele, table, grafikoni, slike ipd. v razpravi ali član­ku naj bodo jasne, njihovo mesto mora biti nedvoumno označeno, njihovo število naj racionalno ustreza vsebini. Tabele, table, slike, ilustracije, grafikoni ipd. skupaj z na­slovi naj bodo priloženi na posebnih listih. Če so slike v digitalni obliki, morajo biti pripravljene u zapisu .tiff v barvni skali CMYK in resoluciji vsaj 300 DPI/inch. Risa­ne slike pa v zapisu .eps. Pri fitocenoloških tabelah se tam, kjer ni zastopana rastlinska vrsta, natisne pika. 14. LITERATURA IN VIRI Uporabljeno literaturo citiramo med besedilom. Citirane avtorje pišemo v kapitelkah. Enega avtorja piše­mo » (Priimek leto)« ali »(Priimek leto: strani)« ali »Pri­imek leto« [npr. (Bukry 1974) ali (Oberdorfer 1979: 218) ali ... Poldini (1991) ...]. Če citiramo več del istega avtorja, objavljenih v istem letu, posamezno delo ozna­čimo po abecednem redu »Priimek leto mala črka« [npr. ...Horvatić (1963 a)... ali (Horvatić 1963 b)]. Avtor­jem z enakim priimkom dodamo pred priimkom prvo črko imena (npr. R. Tuxen ali J. Tuxen). Več avtorjev istega dela citiramo po naslednjih načelih: delo do treh avtorjev »Priimek, Priimek & Priimek leto: strani« [npr. (Shearer, Papike & Simon 1984) ali Pearce & Cann (1973: 290-300)...]. Če so več kot trije avtorji, citiramo »Priimek prvega avtorja et al. leto: strani« ali »Priimek prvega avtorja s sodelavci leto« [npr. Noll et al. 1996: 590 ali ...Meusel s sodelavci (1965)]. Literaturo uredimo po abecednem redu. Imena av­torjev pišemo v kapitelkah: – Razprava ali članek: Dakskobler, L, 1997: Geografske variante asoci­acije Seslerio autumnalis-Fagetum (Ht.) M. Wraber ex Borhidi 1963. Razprave IV razreda SAZU (Ljubljana) 38 (8): 165–255. Kajfež, L. & A. Hočevar, 1984: Klima. Tlatvorni činitelji. V D. Stepančič: Komentar k listu Murska Sobota. Osnovna pedološka karta SFRJ. Pedološka karta Slovenije 1:50.000 (Ljubljana): 7–9. Le Loeuff, J., E. Buffeaut, M. Martin & H. Tong, 1993: Decouverte d’Hadrosauridae (Dinosauria, Orni­thischia) dans le Maastrichtien des Corbieres (Aude, France). C. R. Acad. Sci. Paris, t. 316, Ser. II: 1023–1029. – Knjiga: GORTANI, L. & M. GORTANI, 1905: Flora Friuliana. Udine. Če sta različna kraja založbe in tiskarne, se navaja kraj založbe. – Elaborat ali poročilo: PRUS, T., 1999: Tla severne Istre. Biotehniška fakulteta. Univerza v Ljubljani. Center za pedologijo in varstvo okolja. Oddelek za agronomijo. Ljubljana. (Elabo­rat, 10 str.). – Atlasi, karte, načrti ipd.: KLIMATOGRAFIJA Slovenije 1988: Prvi zvezek: Temperatura zraka 1951–1980. Hidrometeorološki zavod SR Slovenije. Ljubljana. LETNO poročilo meteorološke službe za leto 1957. Hidrometeorološki zavod SR Slovenije. Ljubljana. Za vire veljajo enaka pravila kot za literaturo. 15. LATINSKA IMENA TAKSONOV Latinska imena rodov, vrst in infraspecifičnih tak­sonov se pišejo kurzivno. V fitocenoloških razpravah ali člankih se vsi sintaksoni pišejo kurzivno. 16. FORMAT IN OBLIKA RAZPRAVE ALI ČLAN­KA Članek naj bo pisan v formatu RTF z medvrstičnim razmikom 1,5 na A4 (DIN) formatu. Uredniku je treba oddati izvirnik in kopijo ter zapis na disketi 3,5 ali na CD-ROM-u. Tabele in slike so posebej priložene tekstu. Slike so lahko priložene kot datoteke na CD-ROM-u, za podrobnosti se vpraša uredništvo. INSTRUCTIONS FOR AUTHORS Folia biologica et geologica is a scientific periodical of the Classis IV: Natural history that publishes natural sci­entific proceedings and review articles referring mainly to researches in ethnic region of ours, and also in Europe and elsewhere being of importance, necessity and comparison to our researches. 1. SCIENTIFIC TREATISE It is the entire description of novel research includ­ing the theoretical review of the subjects, presenting in detail the results, conclusions, and the survey of litera­ture of the authors cited. In exceptional cases the survey of literature may be replaced by sources, if the purport requires it. It should be composed in classic manner: introduc­tion, material and methods, results, discussion with con­clusions, acknowledgments, literature, etc. The treatise should not be longer than 30 pages, in­cluding tables, graphs, figures and others. Much desired are treatises of 20 pages. The treatises are reviewed by two reviewers, one of them being member of SASA as a rule, the other one a foreign expert. The reviewers are confirmed by the Classis IV SASA upon the proposal of the editorial board of Folia biologica et geologica. The treatise shall be printed when adopted upon the proposal of the editorial board by Classis IV and the Presidency SASA. 2. REVIEW ARTICLE On consultation with the editorial board and the author, the review article shall be published. Classis IV and the Presidency SASA upon the proposal of the edito­rial board adopt it. It should not be longer than 50 pages. 3. NEWS The periodical publishes short, scientificaly relevant and topical articles up to 7000 characters in lenght. 4. NOVELTY OF THE CONTRIBUTION The treatise or article ought not to be published previously in other periodicals or books. 5. LANGUAGE The treatise or article may be written in one of world language and in Slovenian language especially when the subjects are of local character. The author of the treatise or article provides the translation into slovenian language and corresponding editing, unless otherwise agreed by the editorial board. 6. SUMMARY When the treatise or article is written in Slovenian, the summary should be in English. When they are in foreign language, the summary should be in Slovenian. It should be so extensive that the subjects are clear and understandable to domestic and foreign reader. It should give the information about the intention, method, result, and conclusions of the treatise or article. It should not be longer than 10 to 20% of the treatise or article itself. 7. ABSTRACT It should give concise information about the inten­tion and conclusions of the treatise or article. It must be written in English and Slovenian. 8. KEY WORDS The number of key words should not exceed 10 words. They must present the topic of the research in the treatise or article and written in English and Slovenian. 9. TITLE OF TREATISE OR ARTICLE It should be short and understandable. It is fol­lowed by the name/names of the author/authors (name and surname). 10. ADDRESS OF AUTHOR/AUTHORS The address of author/authors should be at the bot­tom of the page: academic title, name, surname, institu­tion, town and state mark, post number, state, or e-mail of the author/authors. 11. INTRODUCTION Its contents should refer to the purports of the trea­tise or article only. 12. CONCLUSIONS Conclusions ought to include the synthesis of the main statements resolving or indicating the problems of the research. 13. TABLES, GRAPHS, FIGURES, ETC. They should be clear, their place should be marked unambiguously, and the number of them must ration­ally respond to the purport itself. Tables, figures, illus­trations, graphs, etc. should be added within separated sheets. In case that pictures in digital form, TIFF format and CMYK colour scale with 300 DPI/inch resolution should be used. For drawn pictures, EPS format should be used. In cases, when certan plant species are not represented, a dot should be always printed in phytocenologic tables. 14. LITERATURE AND SOURCES The literature used is to be cited within the text. The citation of the authors is to be marked in capitals. One writes the single author as follows: “(Surname year)” or “(Surname year: pages)” or “Surname year” [(Bukry 1974) or (Oberdorfer 1979: 218) or ... Poldini (1991)...]. The works of the same author are to be cited in alphabeti­cal order: “Surname year small letter” [...Horvatić (1963 a)... or (Horvatić (1963 b)]. The first letter of the au­thor’s name is to be added when the surname of several authors is the same (R. Tuxen or J. Tuxen). When there are two or three authors, the citation is to be as follows: “Surname, Surname & Surname year: pages” [(Shearer, Papike & Simon 1984) or Pearce & Cann (1973: 290-300)...]. When there are more than three authors, the ci­tation is to be as follows: “Surname of the first one et al. year: pages” or “Surname of the first one with collabora­tors year” [Noll et al. 1996: 590 or Meusel with collabo­rators (1965)]. The literature is to be cited in alphabetical order. The author’s name is written in capitals as follows: – Treatise or article: Dakskobler, L, 1997: Geografske variante asoci­acije Seslerio autumnalis-Fagetum (Ht.) M. Wraber ex Borhidi 1963. Razprave IV. Razreda SAZU (Ljubljana) 38 (8): 165-255. Kajfež, L. & A. Hočevar, 1984: Klima. Tlatvorni činitelji. V D. Stepančič: Komentar k listu Murska Sobota. Osnovna pedološka karta SFRJ. Pedološka karta Slovenije 1:50.000 (Ljubljana): 7–9. Le Loeuff, J., E. Buffeaut, M. Martin & H. Tong, 1993: Déecouverte d’Hadrosauridae (Dinosauria, Ornithis­chia) dans le Maastrichtien des Corbieres (Aude, France). C. R. Acad. Sci. Paris, t. 316, Ser. II: 1023-1029. – Book: GORTANI, L. & M. GORTANI, 1905: Flora Friuliana. Udine. In case that the location of publishing and printing are different, the location of publishing is quoted. – Elaborate or report: PRUS, T., 1999: Tla severne Istre. Biotehniška fakulteta. Univerza v Ljubljani. Center za pedologijo in varstvo okolja. Oddelek za agronomijo. Ljubljana. (Elabo­rat, 10 str.). – Atlases, maps, plans, etc.: KLIMATOGRAFIJA Slovenije 1988: Prvi zvezek: Temperatura zraka 1951-1980. Hidrometeorološki zavod SR Slovenije. Ljubljana. LETN0 poročilo meteorološke službe za leto 1957. Hidrometeorološki zavod SR Slovenije. Ljubljana. The same rules hold for sources. 15. LATIN NAMES OF TAXA Latin names for order, series, and infraspecific taxa are to be written in italics. All syntaxa written in phyto­coenological treatises or articles are to be in italics. 16. SIZE AND FORM OF THE TREATISE OR AR­TICLE The contribution should be written in RTF format, spacing lines 1.5 on A4 (DIN) size. The original and copy ought to be sent to the editor on diskette 3.5 or on CD-Rom. Tables and figures are to be added separately. Figures may be added as files on CD-Rom. The editorial board is to your disposal giving you detailed information. 17. THE TERM OF DELIVERY The latest term to deliver your contribution is May 31. FOLIA BIOLOGICA ET GEOLOGICA 56/1 - 2015 Slovenska akademija znanosti in umetnosti v Ljubljani Grafična priprava za tisk Medija grafično oblikovanje, d.o.o. Tisk Collegium graphicum, d.o.o. Ljubljana 2015 FOLIA BIOLOGICA ET GEOLOGICA 56/1 – 2015 FOLIA BIOLOGICA ET GEOLOGICA = Ex RAZPRAVE IV. RAZREDA SAZU issn 1855-7996 · Letnik / Volume 56 · Številka / Number 1 · 2015 ISSN 1855-7996 VSEBINA / CONTENTS RAZPRAVE / ESSAYS Igor Dakskobler Phytosociological analysis of montane beech forests on steep shady slopes on mixed geological bedrock in western Slovenia Fitocenološka analiza montanskih bukovih gozdov na strmih osojnih pobočjih na mešani geološki podlagi v zahodni Sloveniji Igor Dakskobler, Branko Dolinar, Branko Zupan, Florijan Poljšak & Peter Strgar Fitocenološka oznaka rastišč vrste Nigritella archiducis-joannis v Dolini Triglavskih jezer (Julijske Alpe, Slovenija) Phytosociological characteristics of the sites of Nigritella archiducis-joannis in the Triglav Lakes Valley (The Julian Alps, Slovenia) Igor Paušič Confirmation of the Austrian vanilla orchid, Nigritella austriaca (Teppner & E. Klein) P. Delforge (Orchidaceae) a new species in the Slovenian flora Potrditev avstrijske murke, Nigritella austriaca (Teppner & E. Klein) P. Delforge (Orchidaceae), nove vrste v flori Slovenije Mitja Zupančič & Jože Skumavec Flora and vegetation in Pokljuka Gorge (Julian Alps, NW Slovenia) Flora in vegetacija Pokljuške soteske (Julijske Alpe, SZ Slovenija) Božidar Drovenik & Branko Vreš Novosti v favni rodu Otiorhynchus s. lat. (Coleoptera: Curculionidea) v Sloveniji Novelties in the fauna of Otiorhynchus s. lat. (Coleoptera: Curculionidea) in Slovenia 56/1 · 2015