CONTRIBUTION TO THE KNOWLEDGE OF NEUROPTERIDA  
(RAPHIDIOPTERA, NEUROPTERA) OF SERBIA COLLECTED  
IN THE PERIOD 2015-2016 
 
Dušan DEVETAK1,2, Predrag JAKŠIĆ3, Ana NAHIRNIĆ-BESHKOVA4,  
Franc JANŽEKOVIČ1, Tina KLENOVŠEK1, Jan PODLESNIK1 and Vesna KLOKOČOVNIK1 
 
1Department of Biology, Faculty of Natural Sciences and Mathematics,  
University of Maribor, Koroška cesta 160, 2000 Maribor, Slovenia 
2e-mail: dusan.devetak@guest.arnes.si 
3Čingrijina 14/25, Zvezdara, 11000 Beograd, Serbia 
4National Museum of Natural History, Bulgarian Academy of Sciences,  
Tsar Osvoboditel Blvd 1, 1000 Sofia, Bulgaria 
 
Abstract – As a result of field studies in 2015–2016 in Serbia, mainly during two 
field collecting trips, four species of Raphidioptera and 65 species of Neuroptera 
were collected. The following species are reported from Serbia for the first time: 
Ornatoraphidia flavilabris (A. Costa, 1855), Phaeostigma pilicollis (Stein, 1863), 
Nothochrysa fulviceps (Stephens, 1836), Nineta pallida (Schneider, 1846), 
Apertochrysa ventralis (Curtis, 1834), Chrysoperla mediterranea (Hölzel, 1972), 
Hemerobius nitidulus Fabricius, 1777, Coniopteryx tineiformis Curtis, 1834 and 
Mantispilla perla (Pallas, 1772).  
 
KEY WORDS: Lacewings, faunistics, Balkan Peninsula 
 
Izvleček – PRISPEVEK K POZNAVANJU MREŽEKRILCEV - NEUROPTERI-
DA (RAPHIDIOPTERA, NEUROPTERA) SRBIJE, ZBRANIH V OBDOBJU OD 
2015 DO 2016 
Med terenskim delom v Srbiji v obdobju 2015–2016, predvsem na dveh odpra-
vah, smo nabrali štiri vrste kamelovratnic (Raphidioptera) in 65 vrst pravih mreže-
krilcev (Neuroptera). Naslednje vrste so nove za Srbijo: Ornatoraphidia flavilabris 
(A. Costa, 1855), Phaeostigma pilicollis (Stein, 1863), Nothochrysa fulviceps 
(Stephens, 1836), Nineta pallida (Schneider, 1846), Apertochrysa ventralis (Curtis, 
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ACTA ENTOMOLOGICA SLOVENICA
LJUBLJANA, JUNIJ 2023                  Vol. 31, øt. 1: 77–100
1834), Chrysoperla mediterranea (Hölzel, 1972), Hemerobius nitidulus Fabricius, 
1777, Coniopteryx tineiformis Curtis, 1834 and Mantispilla perla (Pallas, 1772). 
KLJUČNE BESEDE: Mrežekrilci, favnistika, Balkanski polotok 
 
Introduction 
 
The data on Neuropterida in Serbia are fragmentary and scattered in individual 
papers. To date, there is no comprehensive work on the superorder in the country.  
 
In a period 2011–2019, zoologists from the Department of Biology, Faculty of 
Natural Sciences and Mathematics, University of Maribor, Slovenia organized nine 
neuropterological expeditions to the Balkan Peninsula with the aim of improving the 
knowledge of the less investigated Balkan countries. Two field trips in the periods of 
21-28 June 2015 and 3-10 July 2016 were devoted to Serbia (Devetak 2021). The 
aim of the collecting trips was to emend the species lists of Neuropterida for the 
country. In addition to Slovenian biologists, local researchers also collected data in 
the field.  
 
In the present study, a survey of Neuropterida collected in Serbia in 2015 and 
2016 is given. 
 
Material and methods 
 
Serbia is divided into five administrative units – statistical regions. Field trips 
took place in four of them. Collection sites included in the checklist are detailed in 
Table 1. For each species, a list of localities is given, based on collected material. 
The locality-codes include the three-letter abbreviations of the regions and a numer-
ical code (Table 1). 
 
Insects were collected using a sweep net and portable light traps. Specimens were 
preserved in 70% ethanol and deposited in the first author’s collection. We followed 
the nomenclature and taxonomy proposed by Aspöck et al. (1980, 2001), Rausch and 
Weißmar (2007), Lacewing Digital Library (Oswald 2017) and Duelli and Henry 
(2022). In our paper the understanding of the taxonomic status of owlflies 
(Ascalaphidae) doesn’t follow Oswald (2017); ascalaphids are here treated at the 
family level. We list only those species of Apertochrysa Tjeder, 1966 whose taxo-
nomic status and terminology are clear (Duelli and Henry, 2022). 
 
Results 
 
In Serbia in the period 2015-2016 four species of Raphidioptera and 65 species 
of Neuroptera were collected.  
Acta entomologica slovenica, 31 (1), 2023
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Table 1: List of collection sites in Serbia in 2015-2016.
Region Code Collection site
Vojvodina
VOJ 1
South Banat District: Deliblatska peščara, center; steppe; 146 m; 
44°53’06.7”N 021°04’45.5”E; 8. VII. 2016; Klenovšek, Janžekovič, 
Jakšić & Devetak leg.
VOJ 2
South Banat District: Deliblatska peščara, center; dry meadow; 152 m; 
44°53’59.4”N 021°06’11.6”E; 09.VII.2016; Klenovšek, Janžekovič, 
Jakšić & Devetak leg.
VOJ 3
South Banat District: Deliblatska peščara, Šumarak; grassland-sand 
dunes; 100 m; 44°48’32.1”N 021°10’25.8”E; 09.VII.2016; Klenovšek, 
Janžekovič, Jakšić & Devetak leg.
VOJ 4
South Banat District: Deliblatska peščara, north of Šušara; grassland; 
180 m; 44°56’56”N 021°08’05”E; 8.VII.2016; Klenovšek, Janžekovič, 
Jakšić & Devetak leg.
VOJ 5
South Banat District: Deliblatska peščara, near Deliblato; sands/steppe; 
135 m; 44°51’51.0”N 021°03’35.6”E; 8.VII.2016; Klenovšek, 
Janžekovič, Jakšić & Devetak leg.
VOJ 6
South Banat District: Deliblatska peščara, west of Grebenac; mixed 
forest (Robina, Pinus); 125 m; 44°54’19”N 021°12’48”E; 09.VII.2016; 
Klenovšek, Janžekovič, Jakšić & Devetak leg.
VOJ 7
South Banat District: Deliblatska peščara, south east; forest house; light 
trap; 86 m; 44°48’44.4”N 021°13’41.9”E; 10.VII.2016; Klenovšek, 
Janžekovič, Jakšić & Devetak leg.
Belgrade
BEL 1
Beograd, Zvezdara; 185 m; 44°47’53’’N 20°30’18’’E; 25.VII.2015; 
05.VIII.2015; 14.VIII.2015; 1.IX.2015; 22.-23.II.2016; 01.IV.2016; 
29.VIII.2016; 21.IX.2016; P.Jakšić leg.
BEL 2
Beograd, Kumodraž, 230 m, 44°43’22’’N 20°30’43’’E; 3.VI.2015; 
P.Jakšić leg.
Šumadija and 
Western Serbia
ŠWS 1
Rasina District: river Zapadna Morava, 7 km NNE of Kruševac; 140 m; 
43° 38.375’ N 21° 22.664’ E; 25.VI.2015; Klokočovnik, Klenovšek, 
Podlesnik, Janžekovič, Devetak leg.
ŠWS 2
Zlatibor District: river Drina, lake Perućačko jezero, near Perućac; 300-
315 m; 43° 57.563’ N 19° 23.572’ E; 25.VI.2015; Klokočovnik, 
Klenovšek, Podlesnik, Janžekovič, Devetak leg.
ŠWS 3
Zlatibor District: river Drina, lake Perućačko jezero, Perućac; Hotel 
Jezero; 43°57’44”N 19°24’29”E; 27.VI.2015; Devetak leg.
Acta entomologica slovenica, 31 (1), 2023
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ŠWS 4
Zlatibor District: Tara National Park: Gorge of the Derventa river; 430 
m; 43° 57.087’ N 19° 21.451’ E; 25.VI.2015; Klokočovnik, Klenovšek, 
Podlesnik, Janžekovič, Devetak leg.
ŠWS 5
Zlatibor District: Tara National Park: Gorge of the Derventa river, 
Predov Krst; 610 m; 43° 56.518’ N 19° 20.723’ E; 25.VI.2015; 
Klokočovnik, Klenovšek, Podlesnik, Janžekovič, Devetak leg.
ŠWS 6
Zlatibor District: Tara National Park: Križevac near Predov Krst; 
Quercus forest; 724 m; 43° 56.415’ N 19° 20.163’ E; 27.VI.2015; 
Klokočovnik, Klenovšek, Podlesnik, Janžekovič, Devetak leg.
ŠWS 7
Zlatibor District: Tara National Park: Predov Krst; meadow with 
Juniperus, Pinus, Picea, Acer, Salix; 997 m; 43° 56.460’ N 19° 18.899’ 
E; 27.VI.2015; Klokočovnik, Klenovšek, Podlesnik, Janžekovič, 
Devetak leg.
ŠWS 8
Zlatibor District: Tara National Park: Križevac near Predov Krst; mixed 
forest; 755 m; 43° 56.521’ N 19° 20.165’ E; 27.VI.2015; Klokočovnik, 
Klenovšek, Podlesnik, Janžekovič, Devetak leg.
ŠWS 9
Zlatibor District: Tara National Park: Jezero Zaovine; Pinus, Picea; 935 
m; 43°52.633’N 19°24.972’E; 27.VI.2015; Klokočovnik, Klenovšek, 
Podlesnik, Janžekovič, Devetak leg.
ŠWS 10
Zlatibor District: Tara National Park: Jezero Zaovine: Bjeluša; brook; 
885 m; 43° 52.988’ N 19° 21.707’ E; 27. VI. 2015; Klokočovnik, 
Klenovšek, Podlesnik, Janžekovič, Devetak leg.
ŠWS 11
Zlatibor District: Tara National Park: Vasilići-Filipovići; meadow with 
Juniperus, Pinus, Picea; 1105 m; 43° 51.514’ N 19° 22.910’ E; 27. VI. 
2015; Klokočovnik, Klenovšek, Podlesnik, Janžekovič, Devetak leg.
ŠWS 12
Zlatibor District: Javor Mt., Vasilin vrh peak; 1489 m; 43°26’28’’N 
20°03’06’’E; 12.VII.2015; A. Nahirnić-Beshkova & S. Beshkov leg.
ŠWS 13
Zlatibor District: Prijepolje. Mileševka river; 685 m; 07.VI.2015, A. 
Nahirnić-Beshkova leg.; 
ŠWS 14
Zlatibor District: Prijepolje: Jadovnik Mt., Sopotnica: Šćepanica, 
8.VII.2015; 26.VI.2016; P. Jakšić leg.
ŠWS 15
Zlatibor District: Ćetanica Mt. plateau; near Karaula village; 1355 m; 
43°19’13’’N 19°49’47’’E; 04.VIII.2015; A. Nahirnić-Beshkova & S. 
Beshkov leg.
ŠWS 16
Zlatibor District: Krnjača Mt., near main road, between Bare and 
Busara; 43°25’25’’N 19°23’53’’E; 26.VII.2015; A. Nahirnić-Beshkova 
leg.
ŠWS 17
Zlatibor District: Zlatar Mt., Suvi Čečar; 1118 m; 43°26’31’’N 
19°49’45’’E; 23.VII.2015; A. Nahirnić-Beshkova leg.
ŠWS 18
Zlatibor District: Kitonja Mt., above Vujovići village; 1100 m; 
43°29’54’’N 19°41’49’’E; 27.VII.2015; A. Nahirnić-Beshkova leg.
D. Devetak, P. Jakøiå, A. Nahirniå-Beshkova, F. Janæekoviœ, T. Klenovøek, J. Podlesnik, V. Klokoœovnik: Contribution to the
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ŠWS 19
Zlatibor district: Prijepolje: Jadovnik Mt., Milošev do; 43°18’46”N 
19°48’14”E ; 28.VI.2014; P. Jakšić leg.
ŠWS 20
Kolubara District: Maljen Mt., Divčibare; 850 m; 11.VII.2015; 
19.VII.2015; P. Jakšić leg.
ŠWS 21
Zlatibor district: Javor Mt., between Vasilin vrh and Jankov vrh peaks, 
1397 m, 43°26’14’’N, 20°02’56’’E, 04.VIII.2015; leg. A. Nahirnić & 
S. Beshkov
ŠWS 22
Zlatibor district: Pobijenik Mt., Pribojska Goleša, Đurovići village, 
43°30’54’’N, 19°29’54’’E, 1187 m, 25.VII.2015, leg. A. Nahirnić
ŠWS 23
Zlatibor district: Pobijenik Mt., Borak summit, 1420 m, 43°28’45’’N, 
19°32’47’’, 25.VII.2015; leg. A. Nahirnić
Southern and 
Eastern Serbia
SES 1
Braničevo District: National Park Đerdap: Brnjička, Brnjica; near river 
Danube; 75 m; 44°39’21.8”N 021°45’54.2”E; 07.VII.2016; Devetak 
leg.
SES 2 Braničevo District: Golubac; 250 m; 12.VI.2015, P. Jakšić leg.
SES 3
Braničevo District: National Park Đerdap: Golubac: Ridan, close to the 
river Donava; 90 m; 44°39’36.5”N 021°41’20.4”E; 08.VII.2016; 
Devetak leg.
SES 4
Pirot district: Kanjon Jerme: Vlasi; 491 m; 42° 59.672’ N 22° 38.230’ 
E; 22.VI.2015; Klokočovnik, Klenovšek, Podlesnik, Janžekovič, 
Devetak leg.
SES 5
Pirot district: Zvonačka Banja; 690 m; 42° 55.740’ N 22° 35.558’ E; 
22.VI.2015; Klokočovnik, Klenovšek, Podlesnik, Janžekovič, Devetak 
leg.
SES 6
Pirot district: Dimitrovgrad: Mt Vidlič, south slope; 630 m; 43° 01.806’ 
N 22° 47.917’ E; 22.VI.2015; Klokočovnik, Klenovšek, Podlesnik, 
Janžekovič, Devetak leg. 
SES 7
Pirot district: Dimitrovgrad: Mt Vidlič, south slope; planted forest with 
Pinus nigra; 650 m; 43°02’31.6”N 22°49’16.3”E; 22.VI.2015; 
Klokočovnik, Klenovšek, Podlesnik, Janžekovič, Devetak leg. 
SES 8
Pirot district: Mt. Vidlič, Crni Vrh, 1046-1115 m; 43°11’13”N 
22°39’03”E; 29.V.-15.VI.2015; A.Nahirnić-Beshkova leg. & 
S.Beshkov leg. 
SES 9
Pirot District: Vidlič Mt.: Crni vrh; meadows/ light forest; 1115 m; 
43°11’12.1”N 022°39’02”E; 06. VII. 2016; Jakšić, Klenovšek, 
Janžekovič & Devetak leg.
Acta entomologica slovenica, 31 (1), 2023
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SES 10
Pirot district: Mt. Vidlič, Srednja Glama near Koprivštica; 995 m; 
43°14’05”N 22°36’37”E; 14.VI.2015; A.Nahirnić-Beshkova leg.
SES 11
Pirot district: Mt. Vidlič, road between Pirot and Rsavci below Skrča; 
943 m; 43°11’07’’N 22°43’21’’E; 31.V.2015; A.Nahirnić-Beshkova 
leg.
SES 12
Pirot District: Mt. Vidlič, Basarski Kamen, 991 m; 43°09’09’’N 
22°41’06’’E; 30.V.2015; A.Nahirnić-Beshkova leg.
SES 13
Pirot District: Mt. Vidlič, 1.5 km NWW from Basara village, 885 m; 
43°10’11’’N 22°40’17’’E; 30.V.2015; A.Nahirnić-Beshkova leg.
SES 14
Pirot District: Vidlič Mt.; uvala; 985 m; 43°10’37”N 022°40’07”E; 
06.VII.2016; Jakšić, Klenovšek, Janžekovič & Devetak leg.
SES 15
Pirot District: Vidlič Mt.: north east of Pirot; Pinus nigra planted forest; 
725 m; 43°10’19”N 022°38’35”E = 43.171944, 22.643056; 
06.VII.2016; Jakšić, Klenovšek, Janžekovič & Devetak leg.
SES 16
Pirot District: Vidlič Mt.: Hotel Stara; 1052 m; light trap; 43°10’49.5”N 
022°41’19.7”E; 04.-07.VII.2016; Jakšić, Klenovšek, Janžekovič & 
Devetak leg.
SES 17
Pirot District: Vidlič Mt.; oak forest (Fagus, Acer, Crataegus, Corylus); 
980 m; 43°11’22.2”N 022°39’12.7”E; 06.VII.2016; Jakšić, Klenovšek, 
Janžekovič & Devetak leg.
SES 18
Pirot District: Vidlič Mt.; dry rocky grassland (sporadic Ulmus, 
Carpinus orientalis, Crataegus pentagyna); 970 m; 43°11’22.7”N 
022°38’35.7”E; 06.VII.2016; Jakšić, Klenovšek, Janžekovič & Devetak 
leg.
SES 19
Pirot District: Vidlič Mt.; meadow/ forest edge; sporadic Ulmus, 
Carpinus orientalis, Crataegus; 970 m; 43°11’16”N 022°38’44”E; 
06.VII.2016; Jakšić, Klenovšek, Janžekovič & Devetak leg.
SES 20
Pirot District: Visočka Ržana; 1030 m; 43°11’04”N 022°42’31”E; 05. 
VII. 2016; Jakšić, Klenovšek, Janžekovič & Devetak leg.  
SES 21
Pirot District: between Visočka Ržana and Motel Tigar; mixed forest 
(Pinus, Crataegus, Fagus); 955 m; 43°10’54”N 022°43’28”E; 
05.VII.2016; Jakšić, Klenovšek, Janžekovič & Devetak leg.
SES 22
Pirot District: Balkan (Stara planina), Dojkinci; along river Dojkinačka 
reka; 958 m; 43°15’19”N 022°46’34”E; 05.VII.2016; Jakšić, 
Klenovšek, Janžekovič & Devetak leg.
SES 23
Pirot district: Balkan (Stara planina), Dojkinci; along river Dojkinačka 
reka; 920 m; 43°14’45”N 022°46’35”E; 05.VII.2016; Jakšić, 
Klenovšek, Janžekovič & Devetak leg.
SES 24
Pirot District: south of Dojkinci; planted forest with Pinus; 770 m; 
43°11’36”N 022°48’54”E; 05.VII.2016; Jakšić, Klenovšek, Janžekovič 
& Devetak leg.
D. Devetak, P. Jakøiå, A. Nahirniå-Beshkova, F. Janæekoviœ, T. Klenovøek, J. Podlesnik, V. Klokoœovnik: Contribution to the
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SES 25
Pirot District: Dojkinci; Ponor; 1550 m; 43°15’23.5”N 022°48’31.2”E; 
05.VII.2016; Jakšić leg.
SES 26
Pirot District: Rsovci: Manastir Sv. Ilija; 725 m; 43°10’09.0”N 
022°45’20”E; 05.VII.2016; Jakšić, Klenovšek, Janžekovič & Devetak 
leg.
SES 27
Nišava district: Jelašnička klisura, Jelašnica, 23.VI.2015; Klokočovnik, 
Klenovšek, Podlesnik, Janžekovič, Devetak leg.
SES 28
Nišava district: Niš, east, Nišava river, 23. VI. 2015; Klokočovnik, 
Klenovšek, Podlesnik, Janžekovič, Devetak leg.
SES 29
Nišava district: Svrljiške planine: Gradinje, Padina; 961 m; 
43°19’13’’N 22°20’09’’E; 07.VII.2015; A. Nahirnić-Beshkova leg.
SES 30
Nišava District: Suva planina; Bojanine vode; forest Acer, Fagus, Tilia; 
860 m; 43°13’16.7”N 022°06’46.3”E; 04.VII.2016; Jakšić, Klenovšek, 
Janžekovič & Devetak leg.
SES 31
Nišava District: Suva planina; forest; 840 m; 43°12’37.2”N 
022°07’53.7”E; 04.VII.2016; T. Klenovšek leg.
SES 32
Nišava District: Suva planina; meadow; 1005 m; 43°12’55.5”N 
022°07’25.7”E; 04.VII.2016; Jakšić, Klenovšek, Janžekovič & Devetak 
leg.
SES 33
Nišava District: Suva planina; Devojački grob; mountain crest, 
meadows with sporadic bushes (Sorbus aria, Juniperus); 1305-1320 m; 
43°11’52.6”N 022°08’35.2”E; 04.VII.2016; Jakšić, Klenovšek, 
Janžekovič & Devetak leg.
SES 34
Nišava District: Suva planina; near Gornja Studena; 743 m; 
43°12’34.3”N 022°08’00.0”E; 04.VII.2016; Jakšić, Klenovšek, 
Janžekovič & Devetak leg.
SES 35
Pčinja District: Preševo town, 2 km W Trnava village; 696 m; 
42°16’33”N 21°36’57”E; 27.VIII.2015; 18.IX.2015; 10.XI.2015; light 
trap; A. Nahirnić-Beshkova & S. Beshkov leg.
SES 36
Pčinja District: Preševo, Reljan, 714 m; 42°19’34’’N 021°47’13’’E; 
07.V.2015; P. Jakšić leg. 
SES 37
Pčinja District: Starac Mt., Turski grob; 780 m; 42°20’38’’N 
21°53’03’’E ; 4.VII.2015; 19.IX.2015; 11.XI.2015; A. Nahirnić-
Beshkova & S.Beshkov leg.
SES 38
Pčinja District: Starac Mt., between Budovija and Staračka Kula peak; 
823 m; 42°20’11”N 021°52’27”E; 05.VIII.2016; light trap; A.Nahirnić-
Beshkova, S.Beshkov leg.
SES 39
Pčinja District: Pčinja river Valley: Trgovište, Vražji kamen; 663 m; 
42°23’06’’N 22°03’06’’E; 21.IX.2015; A. Nahirnić-Beshkova & S. 
Beshkov leg.
 
 
RAPHIDIOPTERA 
 
RAPHIDIIDAE  
 
Dichrostigma flavipes (Stein, 1863)  
Literature records for Serbia: Guelmino (1996); Sziráki (2014).  
Material examined: Southern and Eastern Serbia: SES 24.  
 
Ornatoraphidia flavilabris (Costa, 1855) 
Material examined: Southern and Eastern Serbia: SES 41. 
First record in Serbia. 
 
Phaeostigma pilicollis (Stein, 1863) 
Material examined: Southern and Eastern Serbia: SES 36. 
First record in Serbia. 
Acta entomologica slovenica, 31 (1), 2023
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SES 40
Pčinja District: Preševo, 2 km W from Trnava  village; serpentine; 696 
m; 42°16’33’’N 21°36’57’’E; 10.XI.2015; A. Nahirnić-Beshkova & S. 
Beshkov leg.
SES 41
Pčinja District: Mt. Kozjak, Delinovica village; 774 m; 42°19’02’’N 
021°54’36’’E; 03.VII.2015; A.Nahirnić-Beshkova leg.
SES 42
Nišava district: Mt. Tresibaba; Tresibaba pass, 720 m, 43°29’16’’N 
22°13’25’’E; 22.V.2016; A.Nahirnić-Beshkova leg.
SES 43
Pirot district: Mt. Vidlič, 1 km N of Basara village, 985 m; 
43°09’59’’N 22°40’53’’E; 2016.07.22; A.Nahirnić-Beshkova leg.
SES 44
Pirot district: Šljivovički Vis Mt., Stranje, 912 m, 43°09’02’’N, 
22°22’00’’E, 19.V.2016; A.Nahirnić-Beshkova leg.
SES 45
Pčinja district: Kostin Čukar Mt., Goli Čukar, 750 m, 42°21’03’’N, 
21°54’01’’E, 02.VII.2015, leg. A. Nahirnić
SES 46
Pčinja district: Kostin Čukar Mt., Bejavica, 733 m, 42°21’01’N, 
21°53’19’’E, 02.VII.2015; leg. A. Nahirnić
SES 47
Pčinja district: Kozjak Mt., Virovi peak, 1200-1285 m, 42°18’36’’N, 
21°55’42’’E, 03.VII.2015; leg. A. Nahirnić.
SES 48
Starac Mt., Vrh above Gornji Starac village, 794 m, 42°19’32’’N, 
21°52’26’’E, 04.VII.2015; leg. A. Nahirnić
Phaeostigma notatum (Fabricius, 1781) 
Literature records for Serbia: Aspöck et al. (2001). 
Material examined: Šumadija and Western Serbia: ŠWS 7.  
 
 
NEUROPTERA 
 
Suborder Nevrorthiformia 
 
NEVRORTHIDAE 
 
Nevrorthus apatelios H. Aspöck, U. Aspöck & Hölzel, 1977 
Literature records for Serbia: Aspöck et al. (1977), Malicky (1984), Devetak and 
Jakšić (2003), Jones and Devetak (2009), Marković V. et al. (2016).  
Material examined: Southern and Eastern Serbia: SES 5, SES 22. 
A few specimens have been found: two along a stram near Zvonačka Banja and 
ten individuals on riparian vegetation along the Dojkinačka reka river (Fig. 1).  
D. Devetak, P. Jakøiå, A. Nahirniå-Beshkova, F. Janæekoviœ, T. Klenovøek, J. Podlesnik, V. Klokoœovnik: Contribution to the
85
Fig. 1: Riparian vegetation along the Dojkinačka reka river, habitat of Nevrorthus 
apatelios. Photo: D. Devetak. 
Suborder Hemerobiiformia 
 
OSMYLIDAE 
 
Osmylus fulvicephalus (Scopoli, 1763) 
Literature records for Serbia: Živojinović (1950), Živić et al. (2001, 2006), 
Devetak and Jakšić 2003), Đuknić et al. (2010), Devetak et al. (2019b).  
Material examined: Southern and Eastern Serbia: SES 5, SES 28. Šumadija and 
Western Serbia: ŠWS 4, ŠWS 10, ŠWS 13, ŠWS 14 (8.VII.2015), ŠWS 19.  
 
CHRYSOPIDAE 
 
Nothochrysa fulviceps (Stephens, 1836) 
 
Material examined: Southern and Eastern Serbia: SES 35 (18.IX.2015).  
First record in Serbia. 
 
Hypochrysa elegans (Burmeister, 1839) 
Literature records for Serbia: Devetak et al. (2019b).  
Material examined: Southern and Eastern Serbia: SES 8. Šumadija and Western 
Serbia: ŠWS 8. 
 
Nineta flava (Scopoli, 1763) 
Literature records for Serbia: Mocsáry (1899), Devetak and Jakšić (2003).  
Material examined: Southern and Eastern Serbia: SES 32. 
Nineta pallida (Schneider, 1846) 
Material examined: Šumadija and Western Serbia: ŠWS 12.  
First record in Serbia. 
 
Nineta principiae Monserrat, 1981   
Literature record for Serbia: Devetak et al. (2019b). 
Material examined: Southern and Eastern Serbia: SES 39. Šumadija and Western 
Serbia: ŠWS 6.  
 
Chrysopidia ciliata (Wesmael, 1841) 
Literature record for Serbia: Devetak and Jakšić (2003), Guelmino (1996).  
Material examined: Southern and Eastern Serbia: SES 22.  
 
Chrysopa perla (Linnaeus, 1758) 
Literature records for Serbia: Živojinović (1950), Petrik (1958), Stančić (1962), 
Devetak (1992), Devetak and Jakšić (2003), Devetak et al. (2019b). 
Acta entomologica slovenica, 31 (1), 2023
86
Material examined: Belgrade: BEL 2. Southern and Eastern Serbia: SES 23, SES 
26, SES 29, SES 38. Šumadija and Western Serbia: ŠWS 7, ŠWS 11,  
 
Chrysopa walkeri McLachlan, 1893 
Literature records for Serbia: Mocsáry (1899), Devetak (1992). 
Material examined: Belgrade: BEL 2. Southern and Eastern Serbia: SES 6, SES 
10, SES 11, SES 12, SES 24, SES 29, SES 32, SES 38. Šumadija and Western 
Serbia: ŠWS 20 (11.VII.2015), ŠWS 14 (8.VII.2015).  
 
Chrysopa dorsalis Burmeister, 1839 
Literature records for Serbia: Devetak and Jakšić (2003), Devetak et al. (2019b).  
Material examined: Southern and Eastern Serbia: SES 6, SES 7 (on Pinus nigra), 
SES 24. Šumadija and Western Serbia: ŠWS 8, ŠWS 9.  
 
Chrysopa formosa Brauer, 1851 
Literature records for Serbia: Frivaldszky (1877), Mocsáry (1899), Petrik (1958), 
Devetak and Jakšić (2003).  
Material examined: Southern and Eastern Serbia: SES 35 (27.VIII.2015).  
 
Chrysopa pallens (Rambur, 1838) 
Literature records for Serbia: Devetak (1992), Devetak and Jakšić (2003), 
Devetak et al. (2015).  
Material examined: Belgrade: BEL 1 (05.VIII.2015). Šumadija and Western 
Serbia: ŠWS 20 (11.VII.2015).  
 
Chrysopa gibeauxi (Leraut, 1989) 
Literature records for Serbia: Canard and Thierry (2017), Devetak et al. (2019b).  
Material examined: Šumadija and Western Serbia: ŠWS 9.  
In the Tara National Park, close to the lake a male was found in coniferous forest 
with Pinus and Picea at the altitude 935 m. 
 
Apertochrysa flavifrons (Brauer, 1851) 
Literature record for Serbia: Devetak and Jakšić (2003), Devetak et al. (2019a).  
Material examined: Vojvodina: VOJ 7. Southern and Eastern Serbia: SES 4, SES 
7, SES 17, SES 23, SES 30, SES 38.  
 
Apertochrysa prasina s.str. (Burmeister, 1839) 
(sensu Duelli and Henry 2022) 
This species has only recently been redefined (Duelli and Henry 2022). 
Material examined: Southern and Eastern Serbia: SES 8, SES 14, SES 16, SES 
19, SES 21. Šumadija and Western Serbia: ŠWS 8.
D. Devetak, P. Jakøiå, A. Nahirniå-Beshkova, F. Janæekoviœ, T. Klenovøek, J. Podlesnik, V. Klokoœovnik: Contribution to the
87
Apertochrysa ventralis (Curtis, 1834)  
Material examined: Belgrade: BEL 1. Southern and Eastern Serbia: SES 9, SES 
30, SES 31.  
First record in Serbia.  
 
Cunctochrysa albolineata (Killington, 1935) 
Literature records for Serbia: Aspöck et al. (1980), Devetak and Jakšić (2003). 
Material examined: Southern and Eastern Serbia: SES 14, SES 31.  
 
Chrysoperla carnea (Stephens, 1836) sensu stricto 
Literature records for Serbia: Devetak et al. (2019a, 2019b). 
Material examined: Vojvodina: VOJ 7. Belgrade: BEL 1 (29.VIII.2016). 
Southern and Eastern Serbia: SES 1, SES 3, SES 9, SES 17, SES 19, SES 23, SES 
30, SES 31, SES 32, SES 37 (11.XI.2015), SES 40.  
 
Chrysoperla lucasina (Lacroix, 1912) 
Literature records for Serbia: Devetak and Jakšić (2003), Devetak et al. (2019a, 
2019b). 
Material examined: Vojvodina: VOJ 7. Belgrade: BEL 1 (14.VIII.2015, 
1.IX.2015, 29.VIII.2016). Southern and Eastern Serbia: SES 4, SES 5, SES 7, SES 
8, SES 9, SES 14, SES 15, SES 17, SES 19, SES 23, SES 30, SES 34, SES 39. 
Šumadija and Western Serbia: ŠWS 1, ŠWS 7, ŠWS 8, ŠWS 9, ŠWS 10, ŠWS 11, 
ŠWS 15, ŠWS 16, ŠWS 17, ŠWS 18, ŠWS 19.  
C. lucasina is one of the most widespread lacewing species in the country.  
 
Chrysoperla pallida Henry, Brooks, Duelli and Johnson, 2002 
Literature records for Serbia: Devetak et al. (2019a).  
Material examined: Vojvodina: VOJ 7. Belgrade: BEL 1 (25.VII.2015, 
1.IX.2015, 22.-23.II.2016, 01.IV.2016). Southern and Eastern Serbia: SES 14, SES 
17, SES 23, SES 28, SES 32, SES 40.  
 
Chrysoperla agilis Henry, Brooks, Duelli and Johnson, 2003 
Literature records for Serbia: Devetak et al. (2019a).  
Material examined: Vojvodina: VOJ 7. Belgrade: BEL 1 (14.VIII.2015). 
Southern and Eastern Serbia: SES 1, SES 14, SES 17, SES 23, SES 25, SES 30, SES 
40. 
 
Chrysoperla mediterranea (Hölzel, 1972) 
Material examined: Southern and Eastern Serbia: SES 23.  
A male and two females were collected from the vegetation close to the 
Dojkinačka reka river.  
First record in Serbia. 
Acta entomologica slovenica, 31 (1), 2023
88
HEMEROBIIDAE 
 
Hemerobius humulinus Linnaeus, 1758 
Literature records for Serbia: Guelmino (1996), Devetak and Jakšić (2003), 
Devetak et al. (2019a, 2019b), Podlesnik et al. (2019). 
Material examined: Vojvodina: VOJ 6. Belgrade: BEL 1 (25.VII.2015). 
Southern and Eastern Serbia: SES 4, SES 5, SES 14, SES 20, SES 23, SES 24, SES 
26. Šumadija and Western Serbia: ŠWS 2, ŠWS 3, ŠWS 4, ŠWS 5, ŠWS 6, ŠWS 7, 
ŠWS 8, ŠWS 11.   
 
Hemerobius simulans Walker, 1853 
Literature records for Serbia: Devetak and Jakšić (2003), Podlesnik et al. (2019). 
Material examined: Southern and Eastern Serbia: SES 8.  
 
Hemerobius pini Stephens, 1836 
Literature records for Serbia: Devetak et al. (2019b), Podlesnik et al. (2019). 
Material examined: Southern and Eastern Serbia: SES 20. Šumadija and Western 
Serbia: ŠWS 7. 
Hemerobius pini was collected in the Tara National Park in an alpine meadow 
with sporadic Juniperus, Pinus, Picea, Acer and Salix trees, at the altitude 997 m 
(Fig. 2).   
 
Hemerobius contumax Tjeder, 1932 
Literature records for Serbia: Podlesnik et al. (2019). 
Material examined: Šumadija and Western Serbia: ŠWS 12.  
 
Hemerobius nitidulus Fabricius, 1777 
Material examined: Southern and Eastern Serbia: SES 22.  
First record in Serbia. 
 
Hemerobius handschini Tjeder, 1957 
Literature records for Serbia: Devetak and Jakšić (2003), Devetak et al. (2019b), 
Podlesnik et al. (2019).  
Material examined: Southern and Eastern Serbia: SES 6, SES 7, SES 9, SES 14, 
SES 15, SES 24, SES 34, SES 39. Šumadija and Western Serbia: ŠWS 2, ŠWS 3, 
ŠWS 5, ŠWS 8, ŠWS 9, ŠWS 11.  
 
Hemerobius micans Olivier, 1793 
Literature records for Serbia: Devetak (1992), Devetak and Jakšić (2003), 
Devetak et al. (2019b), Podlesnik et al. (2019). 
D. Devetak, P. Jakøiå, A. Nahirniå-Beshkova, F. Janæekoviœ, T. Klenovøek, J. Podlesnik, V. Klokoœovnik: Contribution to the
89
Material examined: Southern and Eastern Serbia: SES 4, SES 5, SES 9, SES 14, 
SES 17, SES 20, SES 21, SES 23, SES 24, SES 30, SES 31, SES 32, SES 39. 
Šumadija and Western Serbia: ŠWS 2, ŠWS 3, ŠWS 4, ŠWS 5, ŠWS 6, ŠWS 7, ŠWS 
8, ŠWS 9, ŠWS 10, ŠWS 15.  
 
Hemerobius gilvus Stein, 1863 
Literature records for Serbia: Devetak and Jakšić (2003), Podlesnik et al. (2019). 
Material examined: Southern and Eastern Serbia: SES 5, SES 39.  
 
Hemerobius marginatus Stephens, 1836 
Literature record for Serbia: Devetak and Jakšić (2003).  
Material examined: Southern and Eastern Serbia: SES 17, SES 20, SES 30.  
Hemerobius marginatus was collected in oak and beech forests with Fagus, Acer, 
Crataegus, Tilia and Corylus at altitudes 860-980 m.  
 
Wesmaelius quadrifasciatus (Reuter, 1894) 
Literature records for Serbia: Podlesnik et al. (2019).  
Material examined: Šumadija and Western Serbia: ŠWS 12.  
Acta entomologica slovenica, 31 (1), 2023
90
Fig. 2: Predov Krst in the Tara National Park - alpine meadow with sporadic 
Juniperus, Pinus, Picea, Acer and Salix trees. Photo: D. Devetak.
Sympherobius pygmaeus (Rambur, 1842) 
Literature record for Serbia: Devetak and Jakšić (2003), Podlesnik et al. (2019).  
Material examined: Southern and Eastern Serbia: SES 35 (18.IX.2015), SES 39. 
Šumadija and Western Serbia: ŠWS 9, ŠWS 15.  
 
Sympherobius fuscescens (Wallengren, 1863) 
Literature record for Serbia: Devetak et al. (2019b), Podlesnik et al. (2019). 
Material examined: Šumadija and Western Serbia: ŠWS 9.  
 
Megalomus tortricoides Rambur, 1842 
Literature records for Serbia: Devetak and Jakšić (2003), Devetak et al. (2019b), 
Podlesnik et al. (2019). 
Material examined: Southern and Eastern Serbia: SES 8 (02.-15.VI.2015), SES 
10, SES 35 (18.IX.2015). Šumadija and Western Serbia: ŠWS 7, ŠWS 12.  
 
Megalomus tineoides Rambur, 1842 
Literature records for Serbia: Podlesnik et al. (2019), Devetak et al. (2020).  
Material examined: Southern and Eastern Serbia: SES 35 (18.IX.2015).  
 
Micromus variegatus (Fabricius, 1793) 
Literature records for Serbia: Devetak (1992), Guelmino (1996), Devetak and 
Jakšić (2003), Podlesnik et al. (2019). 
Material examined: Southern and Eastern Serbia: SES 8, SES 28, SES 35 
(18.IX.2015). Šumadija and Western Serbia: ŠWS 12.  
 
Micromus angulatus (Stephens, 1836) 
Literature records for Serbia: Devetak and Jakšić (2003), Devetak et al. (2019b), 
Podlesnik et al. (2019). 
Material examined: Belgrade: BEL 1 (21.IX.2016). Šumadija and Western 
Serbia: ŠWS 2, ŠWS 8, ŠWS 12, ŠWS 15.  
 
Micromus lanosus (Zelený, 1962) 
Literature records for Serbia: Devetak et al. (2019b), Podlesnik et al. (2019). 
Material examined: Southern and Eastern Serbia: SES 4, SES 17, SES 19, SES 
30, SES 31, SES 35 (18.IX.2015). Šumadija and Western Serbia: ŠWS 2, ŠWS 7, 
ŠWS 8, ŠWS 12. 
 
SISYRIDAE 
 
Sisyra nigra (Retzius, 1783) 
Literature records for Serbia: Guelmino (1996), Podlesnik et al. (2017), Devetak 
et al. (2019b). 
D. Devetak, P. Jakøiå, A. Nahirniå-Beshkova, F. Janæekoviœ, T. Klenovøek, J. Podlesnik, V. Klokoœovnik: Contribution to the
91
Material examined: Šumadija and Western Serbia: ŠWS 2.  
 
Sisyra terminalis Curtis, 1854 
Literature records for Serbia: Podlesnik et al. (2017).  
Material examined: Southern and Eastern Serbia: SES 1 (Fig. 3), SES 3. Šumadija 
and Western Serbia: ŠWS 1.  
Riparian vegetation is a typical habitat for Spongillaflies (along rivers: the 
Zapadna Morava river;  the Danube – Fig. 3).    
 
CONIOPTERYGIDAE 
 
Aleuropteryx loewii Klapálek, 1894 
Literature records for Serbia: Devetak and Jakšić (2003), Devetak et al. (2019b).  
Material examined: Southern and Eastern Serbia: SES 6. Šumadija and Western 
Serbia: ŠWS 9.  
Helicoconis lutea (Wallengren, 1871) 
Literature records for Serbia: Devetak et al. (2019b). 
Material examined: Šumadija and Western Serbia: ŠWS 9, ŠWS 11.  
Acta entomologica slovenica, 31 (1), 2023
92
Fig. 3: Riparian vegetation along the Danube is habitat of Sisyra terminalis 
adults. Photo: D. Devetak.
Helicoconis pseudolutea Ohm, 1965 
Literature records for Serbia: Devetak and Jakšić (2003), Devetak et al. (2019b). 
Material examined: Šumadija and Western Serbia: ŠWS 5, ŠWS 8.  
 
Coniopteryx borealis Tjeder, 1930 
Literature records for Serbia: Devetak and Jakšić (2003).  
Material examined: Šumadija and Western Serbia: ŠWS 5.  
 
Coniopteryx pygmaea Enderlein, 1906  
(syn.: Coniopteryx (C.) parthenia (Navás and Marcet, 1910)) 
Literature records for Serbia: Devetak and Jakšić (2003).  
Material examined: Southern and Eastern Serbia: SES 15, SES 24, SES 34.  
 
Coniopteryx tineiformis Curtis, 1834 
Material examined: Southern and Eastern Serbia: SES 17.  
First record in Serbia.  
 
Coniopteryx arcuata Kis, 1965 
Literature records for Serbia: Devetak et al. (2019b).  
Material examined: Šumadija and Western Serbia: ŠWS 4.  
Coniopteryx (Metaconiopteryx) esbenpeterseni Tjeder, 1930 
Literature records for Serbia: Devetak and Jakšić (2003), Devetak et al. (2019b).  
Material examined: Southern and Eastern Serbia: SES 15, SES 34. Šumadija and 
Western Serbia: ŠWS 5, ŠWS 8.  
 
Coniopteryx (Metaconiopteryx) lentiae H. Aspöck and U. Aspöck, 1964 
Literature records for Serbia: Devetak et al. (2019b).  
Material examined: Šumadija and Western Serbia: ŠWS 9.  
 
Conwentzia pineticola Enderlein, 1905 
Literature records for Serbia: Devetak and Jakšić (2003), Devetak et al. (2019b). 
Material examined: Šumadija and Western Serbia: ŠWS 11.  
 
Semidalis aleyrodiformis (Stephens, 1836) 
Literature records for Serbia: Guelmino (1996), Devetak and Jakšić (2003), 
Devetak et al. (2019b). 
Material examined: Šumadija and Western Serbia: ŠWS 2, ŠWS 5, ŠWS 6, ŠWS 
8, ŠWS 10, ŠWS 11. 
 
MANTISPIDAE 
 
Mantispa styriaca (Poda, 1761) 
Literature records for Serbia: Devetak and Jakšić (2003). 
Material examined: Southern and Eastern Serbia: SES 6, SES 39. 
D. Devetak, P. Jakøiå, A. Nahirniå-Beshkova, F. Janæekoviœ, T. Klenovøek, J. Podlesnik, V. Klokoœovnik: Contribution to the
93
Mantispa aphavexelte U. Aspöck and H. Aspöck, 1994 
Literature records for Serbia: Devetak and Jakšić (2003). 
Material examined: Southern and Eastern Serbia: SES 35 (18.IX.2015), SES 37 
(19.IX.2015), SES 38, SES 39.  
 
Mantispilla perla (Pallas, 1772) 
Material examined: Southern and Eastern Serbia: SES 18 
On Vidlič Mountain, the habitat of this species was dry rocky grassland with spo-
radic Ulmus, Carpinus orientalis and Crataegus pentagyna trees or bushes (Fig. 4). 
On one isolated Ulmus tree more than 50 individuals of  M. perla were observed.  
First record in Serbia.  
 
Suborder Myrmeleontiformia 
 
MYRMELEONTIDAE 
 
Myrmecaelurus trigrammus (Pallas, 1771) 
Literature records for Serbia: Frivaldszky (1877), Biró (1885), Mocsáry (1899), 
Grozdanić and Stevanovič (1969), Mitić and Gradojević (1983), Stevanović and 
Bjelić (1985), Devetak (1992), Devetak et al. (2019a), Ivajnšič and Devetak (2020). 
Acta entomologica slovenica, 31 (1), 2023
94
Fig. 4. A dry rocky graslans with sporadic Ulmus, Carpinus orientalis and 
Crataegus pentagyna trees, Vidlič Mountain. Photo: D. Devetak.
Material examined: Vojvodina: VOJ 1, VOJ 2, VOJ 3, VOJ 4, VOJ 5. Southern 
and Eastern Serbia: SES 37 (04.VII.2015), SES 45.  
In Deliblato Sands (=Deliblatska peščara), M. trigrammus adults appeared en 
masse in fragments of dry meadows (Fig. 5).  
 
Myrmeleon formicarius Linnaeus, 1767 
Literature records for Serbia: Petrik (1958), Grozdanić and Stevanović (1969), 
Stevanović and Bjelić (1985), Devetak (1992), Devetak and Jakšić (2003), Devetak 
et al. (2019a, 2019b). Petrik (1958) listed this species in Deliblato sands, but it was 
impossible to identify larval antlions at the specific level in his time.  
Material examined: Vojvodina: VOJ 5. Southern and Eastern Serbia: SES 5, SES 
8, SES 24, SES 42. Šumadija and Western Serbia: ŠWS 8, ŠWS 14 (8.VII.2015), 
ŠWS 20 (11.VII.2015).  
 
Myrmeleon inconspicuus Rambur, 1842 
Literature records for Serbia: Grozdanić and Stevanovič (1969), Stevanović and 
Bjelić (1985),  
Devetak (1992), Devetak and Jakšić (2003), Devetak et al. (2019a), Ivajnšič and 
Devetak (2020).   
D. Devetak, P. Jakøiå, A. Nahirniå-Beshkova, F. Janæekoviœ, T. Klenovøek, J. Podlesnik, V. Klokoœovnik: Contribution to the
95
Fig. 5: Dry meadows in Deliblato sands. Photo: D. Devetak.
Material examined: Vojvodina: VOJ 2.  
In Deliblato sands M. inconspicuus larvae occurred in fragments of dry meadows 
(Fig. 5).  
 
Euroleon nostras (Geoffroy in Fourcroy, 1785) 
Literature records for Serbia: Živojinović (1950), Grozdanić and Stevanović 
(1969), Stevanović and Bjelić (1985), Devetak (1992), Devetak and Jakšić (2003), 
Devetak et al. (2019a, 2019b), Ivajnšič and Devetak (2020).  
Material examined: Southern and Eastern Serbia: SES 6. Šumadija and Western 
Serbia: ŠWS 4, ŠWS 5, ŠWS 8, ŠWS 11.   
 
Neuroleon microstenus (McLachlan, 1898) 
Literature records for Serbia: Devetak 1992: Serbia. 
Material examined: Southern and Eastern Serbia: SES 38.  
A female of N. microstenus was attracted by light trap. It is a rare species in 
Serbia. 
Distoleon tetragrammicus (Fabricius, 1798) 
Literature records for Serbia: Frivaldszky (1877), Biró (1885), Devetak (1992), 
Guelmino (1996), Devetak and Jakšić (2003), Ivajnšič and Devetak (2020).  
Material examined: Southern and Eastern Serbia: SES 14, SES 33, SES 43.  
 
Creoleon plumbeus (Olivier, 1811) 
Literature records for Serbia: Frivaldszky (1877), Biró (1885), Mocsáry (1899), 
Grozdanić and Stevanovič (1969), Stevanović and Bjelić (1985), Devetak et al. 
(2019a), Ivajnšič and Devetak (2020). 
Material examined: Vojvodina: VOJ 2.  
 
ASCALAPHIDAE 
 
Libelloides lacteus (Brullé, 1832) 
Literature records for Serbia: Devetak and Jakšić (2003), Petrović (2013), Parenta 
et al. (2022).  
Material examined: Southern and Eastern Serbia: SES 8 (29. V. 2015; Fig. 6), 
SES 13, SES 44.  
 
Libelloides macaronius (Scopoli, 1763) 
Literature records for Serbia: Devetak (1992), Devetak and Jakšić (2003), Parenta 
et al. (2022). 
Material examined: Southern and Eastern Serbia: SES 2, SES 6, SES 8, SES 10, 
SES 19, SES 20, SES 26, SES 27, SES 29, SES 37 (04.VII.2015), SES 46, SES 47, 
SES 48. Šumadija and Western Serbia: ŠWS 14 (26.VI.2016), ŠWS 16, ŠWS 20, 
ŠWS 21, ŠWS 22, ŠWS 23.  
Acta entomologica slovenica, 31 (1), 2023
96
 
 
Discussion 
 
Here we report four snakefly and 65 lacewing species collected in Serbia in the 
period 2015-2016. Nine species are reported from the country for the first time.  
 
Some older reports on the occurrence of a few species in Serbia are doubtful. 
Antlions are an example. Some authors, e.g., Petrik (1958) listed only one species 
(M. formicarius) for the Deliblato sands, but in his time a review literature on antlion 
larval identification did not exist.  
 
Another example is green lacewing genus Apertochrysa. Due to the lack of the 
knowledge of taxonomy of the genus in the past, listing some species for the country 
was incorrect (e.g., Devetak et al. 2019b). Some specimens of the newely recognized 
D. Devetak, P. Jakøiå, A. Nahirniå-Beshkova, F. Janæekoviœ, T. Klenovøek, J. Podlesnik, V. Klokoœovnik: Contribution to the
97
Fig. 6. Limestone dry clearing surrounded by Fagetum submontanum carpineto-
sum betuli and Fago – Aceri intermedii – Coryletum colurnae at Crni Vrh 1046-1115 
m, Vidlič Mountain. Photo: A. Nahirnić Beshkova. 
species (Ap1, Ap2, Ap3) are impossible to identify without live coloration (for 
explanation, see Duelli and Henry 2022).  
 
 
Acknowledgements 
We are grateful to Davide Badano who confirmed our identification of Neuroleon 
microstenus. Insects were collected in protected nature areas with the permissions of 
the Ministry of Agriculture, Forestry and Water Economy of the Republic of Serbia. 
Two anonymous reviewers are gratefully acknowledged for thoroughly reviewing 
and improving the manuscript.  
 
 
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Received / Prejeto: 9.4. 2023
Acta entomologica slovenica, 31 (1), 2023
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