155 Plant communities of moist rock crevices with endemic Primula carniolica in the (sub)montane belt of western Slovenia Abstract Based on our analysis of a large number of relevés of communities of moist rock crevices in western Slovenia (southern Julian Alps, northern part of the T rnovski Gozd Plateau) we described several new syntaxa with the endemic Primula carniolica. The most unique in terms of ecology is a hygrophytic chasmophytic community classified into the association Astrantio carniolicae- Primuletum carniolicae, which is, together with similar hygrophytic associations Astrantio carniolicae-Pinguiculetum alpinae and Campanulo cespitosae-Saxifragetum aizoidis, associations Phyteumato columnae-Primuletum carniolicae, Primulo carniolicae-Potentilletum clusianae and the subassociation Primuletum carniolicae violetosum biflorae, temporarily classified into the alliance Cystopteridion and order Potentilletalia caulescentis. Only the new association Paederoto luteae-Potentilletum caulescentis is classified into the alliance Physoplexido comosae-Saxifragion petraeae. The elevational range of chasmophytic communities with Primula carniolica is 200 to 1460 m a.s.l. Species that most commonly accompany Primula carniolica in rock crevices include Paederota lutea, Phyteuma scheuchzeri subsp. columnae, Sesleria caerulea, Aster bellidiastrum and moss Orthothecium rufescens. Iz vleček Z obdelavo obsežnega števila fitocenoloških popisov združb vlažnih skalnih razpok v zahodni Sloveniji (južne Julijske Alpe, severni del T rnovskega gozda) smo opisali nekatere nove sintaksone, v katerih uspeva endemit Primula carniolica. Ekološko najbolj posebna je vlagoljubna združba skalnih razpok, ki jo uvrščamo v asociacijo Astrantio carniolicae-Primuletum carniolicae. Skupaj s podobnima vlagoljubnima asociacijama Astrantio carniolicae-Pinguiculetum alpinae in Campanulo cespitosae-Saxifragetum aizoidis ter tudi asociacijama Phyteumato columnae-Primuletum carniolicae in Primulo carniolicae-Potentilletum clusianae ter subasociacijo Primuletum carniolicae violetosum biflorae jih za zdaj uvrščamo v zvezo Cystopteridion in red Potentilletalia caulescentis. Le novo asociacijo Paederoto luteae-Potentilletum caulescentis uvrščamo v zvezo Physoplexido comosae-Saxifragion petraeae. Razpon nahajališč združb skalnih razpok z vrsto Primula carniolica je od 200 m do 1460 m nm. v. Vrste, ki v skalnih razpokah najbolj pogosto rastejo skupaj z njo pa so Paederota lutea, Phyteuma scheuchzeri subsp. columnae, Sesleria caerulea, Aster bellidiastrum in mah Orthothecium rufescens. Key words: phytosociology, synsystematics, calcareous rock faces, Astrantia carniolica, Pinguicula alpina, Primula carniolica, Saxifraga aizoides, Natura 2000, Julian Alps, Dinaric Alps, Slovenia. Ključne besede: fitocenologija, sinsistematika, karbonatno skalovje, Astrantia carniolica, Pinguicula alpina, Primula carniolica, Saxifraga aizoides, Natura 2000, Julijske Alpe, Dinarsko gorstvo, Slovenija. Received: 6. 2. 2020 Revision received: 30. 3. 2020 Accepted: 6. 4. 2020 1 Scientific Research Centre of the Slovenian Academy of Sciences and Arts, Institute of Biology, Regional Unit Tolmin, Brunov drevored 13, 5220 Tolmin, Slovenia. E-mail: igor.dakskobler@zrc-sazu.si 2 Zaloška 78 a, SI-1000 Ljubljana, Slovenia. E-mail: andrej.martincic@siol.net Igor Dakskobler1 & Andrej Martinčič2 DOI: 10.2478/hacq-2020-0005 19/2 • 2020, 155–231 19/2 • 2020, 155–231 156 Igor Dakskobler & Andrej Martinčič Plant communities of moist rock crevices with endemic Primula carniolica in the (sub)montane belt of western Slovenia & Martinčič 2018) and in two shrub communities of rocky sites, namely Heliospermo pusillae-Rhododendretum hirsuti and Rhododendro hirsuti-Salicetum appendiculatae (Dakskobler & Surina 2017b). In most of the listed forest and grassland communities Primula carniolica occurs with a lower frequency and cover than in chasmophytic (rock-face) communities, where its optimal sites are. Only two such communities have been described to date: Primulo carniolicae-Potentilletum caules- centis (Dakskobler 1998, Accetto 2008) and Primuletum carniolicae (Accetto 2008, 2015, Dolinar et al. 2013, 2015, 2017). Most of our relevés of chasmophytic communities with Primula carniolica have not yet been processed and published. In this article we provide a brief description of the identified communities and try to find an appropriate syntaxonomic classification for some of them. Introduction Primula carniolica is endemic to Slovenia and has a rela- tively small distribution area in the pre-Alpine and Di- naric phytogeographical region of Slovenia, which spans the foothills of the southern Julian Alps and NW Dinaric Alps. Its northernmost localities are in the northeastern edge of the Šentviška Gora Plateau at the village Bukovski Vrh, in the gorge Selska Voda under the village of Bukovo and in Orehovska Grapa gorge under the hamlet of Nemci at the foothills of Mt. Kojca. The southeastern- most locality is under Pšeničev Vrh above Sodražica and the ridge of Velika Gora above Ribnica (Figure 1, see also Dakskobler et al. 2004, Dolinar et al. 2013, Dakskobler & Vreš 2014). There are many localities in its distribu- tion area and they frequently feature many specimens. The Carniolan primrose occurs in different plant com - munities, but mainly on dolomite or dolomite-limestone bedrock, most often on rocky and at least slightly moist sites, in narrow, hard-to-access ravines, more rarely on karstified Dinaric plateaus, at elevations spanning 200 to 1460 m. Numerous plant communities featuring this species have already been supported by phytosociological tables. It has been identified (by the authors of this study and others) in stands of the following forest syntaxa: Om- phalodo-Fagetum rhododendretosum hirsuti (Dakskobler et al. 2000, Surina & Dakskobler 2013), Rhododendro hir- suti-Fagetum (Dakskobler 2003, Accetto 2015), Arunco- Fagetum (Dakskobler 2015a), Polysticho lonchitis-Fagetum (Dakskobler & Rozman 2020, in preparation), Lamio or- valae-Aceretum pseudoplatani (Dakskobler 2007), Lamio orvalae-Salicetum eleagni caricetosum albae (Dakskobler 2010), Fraxino orni-Ostryetum phyteumatetosum colum- nae (Dakskobler 2015b), Rhododendro hirsuti-Ostryetum (Dakskobler 2015b, Accetto 2015), Ostryo carpinifoliae- Piceetum (Accetto (2013), Fraxino orni-Pinetum nigrae var. geogr. Primula carniolica (Dakskobler 1998, 1999, Accetto 2015), Primulo carniolicae-Pinetum nigrae (Ac- cetto 2008, 2015), Genisto januensis-Pinetum sylvestris campanuletosum cespitosae (Rozman et al. 2020). It has been recorded in several grassland communities as well: Primulo carniolicae-Caricetum firmae, Primulo carniolicae- -Seslerietum calcariae (Surina & Dakskobler 2005, Dak- skobler 2006), Saxifrago squarrosae-Caricetum mucronatae var. geogr. Primula carniolica (Dakskobler 2006, Daksko- bler & Surina 2017a), Primulo carniolicae-Caricetum sem- pervirentis (Accetto 2010), in a frost pocket community of moist rocky hollows Drepanoclado uncinati-Heliosper- metum pusillae (Surina & Vreš 2009), in a community of moist screes Astrantio carniolicae-Adenostyletum glabrae (Dakskobler 2008), community of dolomite headwaters Astrantio carniolicae-Schoenetum nigricantis (Dakskobler Figure 1: Distribution of Primula carniolica in Slovenia (FloVegSi database). Slika 1: Razširjenost vrste Primula carniolica v Sloveniji (podatkovna baza FloVegSi). Methods Our analysis was based on the phytosociological rele- vés recorded on the sites of Primula carniolica using the Central-European (Braun-Blanquet 1964) approach. The relevés were made in the northwestern part of its distri- bution area, between Čepovan, Slap ob Idrijci, Bukovski Vrh, Bukovo, Orehek in the northwest and north, and the Golaki range, Črni Vrh nad Idrijo and the Žirovnica valley near Žiri in the south and southeast. The plot size was determined based on site homogeneity. Relevés were entered into the FloVegSi database (Fauna, Flora, Vegeta- tion and Paleovegetation of Slovenia) of the Jovan Hadži Institute of Biology at ZRC SAZU (Seliškar et al. 2003). In further analyses we focused only on the relevés of chas- mophytic communities in this area (altogether about 320 – Figure 2) and ordered them using hierarchical classifica-                                                                       19/2 • 2020, 155–231 157 Igor Dakskobler & Andrej Martinčič Plant communities of moist rock crevices with endemic Primula carniolica in the (sub)montane belt of western Slovenia tion, unweighted average linkage method – UPGMA and Wishart’s similarity ratio, using SYN-TAX 2000 program package (Podani 2001). We transformed the combined cover-abundance values with numerical values (1–9) ac- cording to van der Maarel (1979). The UPGMA method was used because it better recognises the relevés that stand out from others. The results of the UPGMA correspond to our experience in the field, as the clusters comprise the relevés (stands) that are in fact ecologically similar. The relevés where Primula carniolica is mostly accom- panied by distinctly hygrophytic species Pinguicula al- pina, Astrantia carniolica, Hymenostylium recurvirostre and Palustriella commutata (the smaller group) clustered separately from all other relevés (the bigger group), which either comprised only some of the above-listed species, usually with low mean coverage, or none of these spe- cies (Dendrogram, Figure 14 in Appendix). In the next step, these two, distinctly different groups of relevés were processed separately, but without taking into account all similar relevés in the bigger group, in order to avoid the tables being too extensive. Figure 2: Localities of rock crevice communities with Primula carniolica in western Slovenia. Slika 2: Nahajališča združb skalnih razpok z vrsto Primula carniolica v zahodni Sloveniji. 19/2 • 2020, 155–231 158 Igor Dakskobler & Andrej Martinčič Plant communities of moist rock crevices with endemic Primula carniolica in the (sub)montane belt of western Slovenia For the sake of comparison our analysis included also several other communities of moist rock crevices in the submontane-montane belt of the southern Julian Alps and the northern part of the Dinaric Alps, which are dominated by Pinguicula alpina and (or) Saxifraga aizoides, have a rich moss layer, but do not comprise Primula carniolica (these communities are partly dis- tributed outside its distribution area). These relevés were arranged into tables in the same manner, through hier- archical classification, the same as the columns in the synoptic table. Numerical comparisons were performed with the SYN-TAX 2000 program package (Podani 2001). When describing new associations of plant com- munities with poor species composition and few domi- nant species, we focused on the latter and their mean coverage, rather than only on floristic similarity between the compared syntaxa. The nomenclatural source for the names of vascu- lar plants was the Mala flora Slovenije (Martinčič et al. 2007), except for the name Molinia arundinacea Schrank. The nomenclature of Flora alpina (Aeschimann et al. 2004) – Sesleria caerulea was used for the taxon Sesleria caerulea subsp. calcaria (MFS). Ros et al. (2007) was the nomenclatural source for the names of liverworts (March- anthiophyta) and Ros et al. (2013) for the names of moss- es. Šilc & Čarni (2012), Accetto (2015), Mucina et al. (2016), Dakskobler & Surina (2017a, b) and Dakskobler & Martinčič (2018) served as nomenclatural sources for the names of the syntaxa. The data source for geological bedrock was Buser (2009). The geographic coordinates of the relevés were determined according to the Slovenian geographic coordinate system D 48 (Zone 5) on the Bes- sel ellipsoid and with Gauss-Krüger projection. Results and discussion Arrangement of relevés into tables Our analysis of 348 relevés of rock crevices with Primula carniolica (from its entire distribution area), showed that even though they comprise about 300 species of vascular plants and mosses, the species with constancy higher than 50% are (in addition to Primula carniolica) only Paed- erota lutea, Phyteuma scheuchzeri subsp. columnae, Sesleria caerulea, Aster bellidiastrum and moss Orthothecium rufe- scens. Seven species have the constancy of 40% or higher: Astrantia carniolica, Asplenium ruta-muraria, Valeriana saxatilis, Asplenium trichomanes, Calamagrostis varia and mosses Neckera crispa and Ctenidium molluscum. The en- tire species composition of these communities features less than 10% of the species with constancy higher than 20% and only 42 species (35 vascular plants and 7 mosses) with constancy more than 10% (the list of the most com- mon species in chasmophytic communities with Primula carniolica is in Table 11). This clearly demonstrates that the decisive factor when classifying communities with the average number of only 15 to 20 species per relevé into a syntaxonomic system are in the first place the species with constancy of at least 40% or higher. Some species with the constancy of 10 to 35%, but high mean coverage can be in the first place differential species or species that indicate entirely different sites. The relevés of distinctly moist calcareous rock crevic- es, with occasional tufa formations, were arranged into two tables. Table 1 comprises hygrophilous communities where Primula carniolica has similar constancy and mean coverage as Pinguicula alpina and Astrantia carniolica, while Hymenostylium recurvirostre in Palustriella commu- tata stand out in the moss layer, both with their frequency and mean coverage. With hierarchical clustering we arranged the stands that were the most similar to these in terms of site moisture (stands with dominating Pinguicula alpina and Astrantia carniolica, with some stands comprising also Saxifraga aizoides) in Table 2 and Table 3 (stands with dominating Saxifraga aizoides). Table 5 comprises the relevés with three dominant spe- cies alongside Primula carniolica: Orthothecium rufescens, Paederota lutea and Phyteuma scheuchzeri subsp. colum- nae. The latter subspecies is not included in the phyto- sociological table of the original description of the as- sociation Primuletum carniolicae (Accetto 2008), because its distribution area (Figure 3) does not overlap with the entire distribution area of Primula carniolica, but only with its western part, primarily the Idrijca river basin with the Trnovski Gozd Plateau. Ecology of Phyteuma columnae is slightly different from the ecology of Primula carniolica. As it requires less moisture it is more com- mon in very dry rock crevices. It is a character species of the association Phyteumato-Potentilletum caulescentis, whose stands dominate on the southern rocky edge of the Trnovski Gozd and Nanos plateaus (Poldini 1978), more rarely also on their northern side, and of the as- sociation Phyteumato-Moehringietum villosae from the foothills of the southern Julian Alps (Dakskobler 2000). In stands with Primula carniolica it is therefore above all a phytogeographical differential species. The entire species composition of the community in Table 5 is not considerably different from the floristic composition of the community in Table 1, but there are obvious differ- ences in terms of constancy and mean coverage of the most frequent species, and in the percentage of groups of diagnostic species (Table 10). 19/2 • 2020, 155–231 159 Igor Dakskobler & Andrej Martinčič Plant communities of moist rock crevices with endemic Primula carniolica in the (sub)montane belt of western Slovenia Figure 3: Distribution of Phyteuma scheuchzeri subsp. columnae in Slovenia (FloVegSi database). Slika 3: Razširjenost taksona Phyteuma scheuchzeri subsp. columnae v Sloveniji (podatkovna baza FloVegSi). While the relevés arranged in Table 6 are similar to the relevés in Table 5, many of them were made at higher el- evations and have a higher frequency of more hygrophytic species such as Astrantia carniolica and Viola biflora. In terms of ecological conditions these relevés indicate a cer- tain similarity also with the community in Table 1 (com- pare structure by groups of diagnostic species, Table 10). Table 7 comprises the relevés that grouped separately in the comparison of the relevés of the association Primuletum carniolicae s. lat. and have until now been classified into the association Primulo carniolicae-Potentilletum caulescentis. Table 8 comprises six relevés from Govci (rock faces un- der the top of Zeleni Rob and Poldanovec on the north- ern edge of the Trnovski Gozd Plateau above the Trebuša valley), characterised by Potentilla clusiana, Carex firma, Rhodothamnus chamaecistus and Campanula cochleariifolia. The relevés in T able 4 grouped separately and were main- ly made on the fringe of the distribution area of Primula carniolica, in the gorge Raskovec and the Železnica valley (municipality of Idrija, although historically this territory belongs to Žiri and Gorenjska regions). Viola biflora and Pinguicula alpina are frequent here, but Paederota lutea and Phyteuma scheuchzeri subsp. columnae are very rare. Syntaxonomic classification of of recorded communities based on the synoptic table Table 9 was decisive in our assessment and synsystematic classification of the studied communities. The synoptic table was used because the syntaxa were described with more than one relevé. As no single relevé is fully rep- resentative of a syntaxon it is only in a synthetic form (a cluster of relevés) that the author can convey how he understands a certain syntaxon. Individual stands (com- munities) that occur in nature and are depicted by our relevés can never fully correspond to what the author recognised as a (specific) abstract syntaxon. We therefore compared the syntaxa described in this article, which were obtained with the cluster analysis of individual relevés (see the dendrogram, Figure 14 in the Appendix), with other, already published syntaxa, which were con- sidered in a synthetic form. In addition to most of the communities in the listed tables (except the community in Table 3, which is obviously different from other chas- mophytic communities with dominant Pinguicula alpina and/or Primula carniolica) it comprises, in synoptic form, also the following, already published communities: four tables of the association Primuletum carniolicae s. lat. (Dolinar et al. 2013, 2015, 2017) and two published ta- bles of the same association (Accetto 2008, 2015). These six synoptic columns present chasmophytic communities in the central and southeastern part of the distribution area of Primula carniolica (the above-mentioned tables have no relevés from this area). We also added two other forms of the association Primulo carniolicae-Potentille- tum caulescentis (Dakskobler 1998, Accetto 2008) to the synoptic table. The table comprises 15 syntaxa. By com- paring them we obtained the dendrogram in Figure 4, in which we took into account the species’ constancy, and not only their presence or absence. This decision is based on the already mentioned observation that the species that play the decisive role in these communities, which are distributed across small areas and have few recorded species, are in the first place the species with higher con- stancy and mean coverage. Some researchers assess mean coverage also with a specific cover index (for communi- ties in the territory of Slovenia comp. Surina 2005). Based on the synoptic table and hierarchical classifi- cation we can describe two new associations, Astrantio carniolicae-Primuletum carniolicae (Table 1) and Astrantio carniolicae-Pinguiculetum alpinae (Table 2) whose stands occur on apparently moister sites than those of the com- munities from previously described associations Primu- letum carniolicae and Primulo carniolicae-Potentilletum caulescentis. They are similar also in terms of constancy of diagnostic species, except that the stands of the first asso- ciation differ so significantly from the stands of the other in terms of certain community-defining species (see also T able 10) that we find the classification of the community in T able 1 into the subassociation Astrantio-Pinguiculetum alpinae primuletosum carniolicae less appropriate. In terms of entire species composition the stands of the association Astrantio-Pinguiculetum alpinae are slightly similar to the stands of the association Hymenostylio recur-                                                                       19/2 • 2020, 155–231 160 Igor Dakskobler & Andrej Martinčič Plant communities of moist rock crevices with endemic Primula carniolica in the (sub)montane belt of western Slovenia virostri-Pinguiculetum poldinii from the Venetian Prealps, which its authors Giovagnoli & T assinazzo (2012) classify into the alliance Adiantion and class Adiantetea. Given the presence of certain bryophytes, character species of this class (Hymenostylium recurvirostre, Preissia quadrata, Pel- lia endiviifolia and Eucladium verticillatum) in the stands of the associations Astrantio-Pinguiculetum alpinae and Astrantio-Primuletum carniolicae, these associations could be classified in the same way. Nevertheless, the fact that among the listed moss species only Hymenostylium re- curvirostre has higher constancy and mean coverage, the absence of Adiantum capillus-veneris from our relevés as well as the entire species composition speak against such classification. (Sub)montane stands of moist rock crevices with Saxifraga aizoides, which according to the hierarchical classification do not belong to the association Astrantio carniolicae-Pinguiculetum alpinae, are a curiosity in the Southeastern Alps, and we therefore classify them into the new association Campanulo cespitosae-Saxifragetum aizoidis. The second group with the highest degree of similarity features two forms of the association Primuletum carni- olicae s. lat. with the taxon Phyteuma scheuchzeri subsp. columnae and two forms of the association Primulo carn- iolicae-Potentilletum caulescentis. This means that similar stands from the northwestern part of Primula carniolica’s distribution area are grouped together. These commu- nities could be treated as a special geographical variant of the association Primuletum carniolicae. According to our criteria the rank of geographical variant (a small or regional association) is higher than the rank of subasso- ciation. Since the applicable Code of Phytosociological Nomenclature (Weber et al. 2000) does not deal with this rank, we have two options for a valid typification: • to reduce the rank of geographical variant to the rank of subassociation, or • to bring the rank of geographical variant to the rank of association. The results of hierarchical classification speak in favour of the latter option (see also Tables 9 and 10), as they point to certain floristic differences between the studied stands and the stands described by Accetto (2008, 2015) and Dolinar et al. (2013, 2015, 2017) as the association Primuletum carniolicae, also in terms of the updated se- lection of diagnostic species (in both cases the common species are Primula carniolica, Orthothecium rufescens and Valeriana tripteris). Based on these criteria it makes sense to classify our stands into the new association Phyteumato columnae-Primuletum carniolicae. Only in this case can we take into account the results of hierarchical classifica- tion (Figure 4), which demonstrate considerable similar- Figure 4: Dendrogram of rock crevices communities with endemic Primula carniolica, UPGMA, 1-similarity ratio. Slika 4: Dendrogram združb skalnih razpok z endemitom Primula carniolica (UPGMA, 1-similarity ratio). Legend / Legenda: AcPa Astrantio carniolicae-Pinguiculetum alpinae, this article, Table 2. AcPc Astrantio carniolicae-Primuletum carniolicae, this article, Table 1. PhcPc-ac Phyteumato columnae-Primuletum carniolicae astran- tietosum carniolicae, this article, Table 6. Phc-Pc-ty Phyteumato carniolicae-Primuletum carniolicae typi- cum, this article, Table 5. PcPcl-CT Phyteumato columnae-Primuletum carniolicae poten- tilletosum caulescentis, this article, Table 7 (CT – Cerkno and T rebuša regions). PcPcl-TG Phyteumato columnae-Primuletum carniolicae poten- tilletosum caulescentis (Primulo carniolicae-Potentilletum cau- lescentis), Dakskobler (1998, Table 1, pp. 286–287) – TG (T rnovski Gozd). Pcty Primuletum carniolicae, Accetto (2008, T able 1, pp. 42–44). Pccb Primuletum carniolicae caricetosum brachystachyos, Accetto (2015, Table 27, pp. 98–99). PcPcl-As Paederoto luteae-Potentilletum caulescentis (Primulo carniolicae-Potentilletum caulescentis var. A spleni um seelosii), Accetto (2008, Table 5, pp. 49–50). Pcvb Primuletum carniolicae violetosum biflorae, this article, Table 4. Pc-No Primuletum carniolicae s. lat., Dolinar et al. (2017, Table 1, pp. 30–32) – No – Notranjska region Pc-Do Primuletum carniolicae s. lat., Dolinar et al. (2013, Table 1, pp. 13–16) – Do – Dolenjska region. Pc-Npr Primuletum carniolicae s. lat., Dolinar et al. (2015, Table 1, pp. 40–43) – Notranjski Park Cerknica. Pc-Iv Primuletum carniolicae s. lat., Dolinar et al. (2013: Table 2, pp. 17–18) – Iv – Iški Vintgar. PcPcl Primulo carniolicae-Potentilletum clusianae, this article, Table 8.                                19/2 • 2020, 155–231 161 Igor Dakskobler & Andrej Martinčič Plant communities of moist rock crevices with endemic Primula carniolica in the (sub)montane belt of western Slovenia ity of the stands of the association Primulo carniolicae- Potentilletum caulescentis from the northwestern part of the distribution area of Primula carniolica (Dakskobler 1998, 2000) with presently described communities from the same area. With these findings in mind, this associa- tion should be reduced to the rank of the subassociation Phyteumato columnae-Primuletum carniolicae potentilleto- sum caulescentis. Based on floristic similarity (Figure 4), however, the relevés from the rock faces of Govci under Zeleni Rob and Poldanovec, where Potentilla clusiana occurs or even dominates, cannot be classified into the association Phy- teumato-Primuletum carniolicae. In the dendrogram in Figure 4 they grouped separately from other relevés and are therefore classified into the new association Primulo carniolicae-Potentilletum clusianae. Based on the results of hierarchical classification (Figure 4, Table 9) it would make sense to rename the subassociation Primuletum carniolicae caricetosum brach- ystachyos (Accetto 2015) into Primuletum carniolicae caricetosum mucronatae. The nomenclatural type is the same, but Carex mucronata (frequency 100%) defines these stands significantly more than C. brachystachys (fre- quency 50%). However, the Code (Weber et al. 2000) does not allow such renaming, because the valid name caricetosum brachystachyos is not based on a taxonomical error (Arts. 29a and 43). Justification of the possible new suballiance (alliance) of (sub)mon- tane chasmophytic communities Table 9 serves also as the basis for classification of the described communities into higher syntaxonomic units. So far, these communities have been classified either into the alliance Physoplexido comosae-Saxifragion petraeae or into the alliance Cystopteridion (Šilc & Čarni 2012, Accetto 2015, Dolinar et al. 2015, 2017). A more recent review of the vegetation of Europe at the rank of classes, orders and alliances (Mucina et al. 2016) indicates that the alliance Violo biflorae-Cystopteridion alpinae (syntax. syn. Cystopteridion) belongs to the order Violo biflorae- Cystopteridetalia alpinae and class Polypodietea. This clas- sification is different from that of Theurillat (2004), and E. and S. Pignatti (2014, 2016), who classify the alliance Cystopteridions into the order Potentilletalia caulescentis and class Asplenietea trichomanis. The studied communi- ties with their entire species composition also undoubt- edly belong to this order and class, and are characterised by higher occurrence of hygrophytic species. According to Mucina et al. (2016) they cannot be classified into the alliance Violo biflorae-Cystopteridion (syn. Cystopte- ridion). According to Theurillat (in litt.) the classifica - tion of the alliance Cystopteridion into the order Violo biflorae-Cystopteridetalia alpinae is problematic. In the original diagnosis this order corresponds to communities that are intermediate between the classes Asplenietea tri- chomanis and Thlaspietea rotundifoliae , and indifferent to the substrate. Therefore, the alliance Cystopteridion Rich- ard 1972 is not the same syntaxon as the alliance Violo biflorae-Cystopteridion alpinae Fernández Casas 1970, as described in Mucina & al. (2016). With this in mind, we provisionally classify most of the communities described in this article into the alli- ance Cystopteridion and order Potentilletalia caulescentis. We also propose another possible alliance, an alliance of hygrophytic rock crevice (chasmophytic) communi- ties of the Southeastern Alps and northern Dinaric Alps (Astrantio carniolicae-Paederotion luteae nom. prov.) an ecological vicariant of the alliance Physoplexido comosae- -Saxifragion petraeae. Based on Table 9, Paederota lutea and Astrantia carniolica seem to be appropriate name- giving species for such an alliance. Paederota lutea is an eastern-Alpine-Illyrian species and a character species of chasmophytic communities (Aeschimann et al. 2004). In Slovenia it is distributed in chasmophytic commu- nities, on screes, stony subalpine-alpine grasslands and stony shrub communities and forests from the colline to the alpine belt. In the colline, submontane and mon- tane belts it frequently occurs on shady moist rocks and is one of the most constant companions of the endemic Primula carniolica. Surina (2005) described its associa- tion Ranunculo traunfellneri-Paederotetum luteae in the Krn Mts. and classified it into the alliance Cystopter- idion. Paederota lutea is also the name-giving species of the association Paederoto luteae-Minuartietum rupestris (Wraber 1986, Surina 2005). Both described associations occur in the subalpine or alpine belt. A detailed ecologi- cal, phytogeographical and phytosociological description of Astrantia carniolica was published recently (Daksko- bler & Martinčič 2018), and is therefore not repeated herein. Although it occurs also in different communities, in dolomite headwaters, on moist screes and in stony for- ests, Astrantia carniolica nevertheless meets the criteria (together with Paederota lutea, which has a very similar, but slightly larger distribution area) for naming south- eastern-Alpine‒northern-Dinaric communities of moist rock crevices. However, to ensure that the description of the new alliance is valid, we should take into account also similar syntaxa from other parts of the Southeast- ern Alps and northern Dinaric Alps, and perform more extensive numerical comparisons (opinion of two anony- mous reviewers, with which the authors agree). 19/2 • 2020, 155–231 162 Igor Dakskobler & Andrej Martinčič Plant communities of moist rock crevices with endemic Primula carniolica in the (sub)montane belt of western Slovenia An overview of newly described syntaxa with their nomenclatural types and diagnostic species Asplenietea trichomanis (Br.-Bl. in Meier et Br.-Bl. 1934) Oberd. 1977 Potentilletalia caulescentis Br.-Bl. in Br.-Bl. et Jenny 1926 Cystopteridion fragilis Richard 1972 (non. Violo biflorae-Cystopteridion alpinae sensu Mucina et al. 2016) (Astrantio carniolicae-Paederotion luteae nom. prov.) In the context of this article the diagnostic spe- cies of this alliance include the species expected to be diagnostic also for the suggested possible new alliance: Astrantia carniolica, Paederota lutea, Pri- mula carniolica, Orthothecium rufescens, Aster bel- lidiastrum, Pingiucula alpina, Carex brachystachys, Asplenium viride, Valeriana saxatilis, V. tripteris, Viola biflora, Saxifraga aizoides, Tofieldia calycula- ta, Cystopteris fragilis, Hymenostylium recurvirostre, Palustriella commutata. Astrantio carniolicae-Primuletum carniolicae ass. nov. hoc loco Nomenclatural type, holotypus, is relevé 35 in Table 1. Diagnostic species: Primula carniolica, Astran- tia carniolica, Pinguicula alpina, Hymenosty- lium recurvirostre, Palustriella commutata (char- acter species) and Phyteuma scheuchzeri subsp. columnae, Molinia arundinacea, Valeriana saxa- tilis and Carex mucronata (differential species). • asteretosum bellidiastri subass. nov. hoc loco (relevés 4–26 in Table 1), nomen- clatural type, holotypus, is relevé 15 in Table 1. Differential species: Aster bellidiastrum, Paederota lutea. • hymenostylietosum recurvirostris subass. nov. hoc loco (relevés 27–56 in Table 1), nomenclatural type, holotypus, is relevé 35 in Table 1. Differential species: Hymenostylium recur- virostre, Pinguicula alpina (because of high- er medium cover value than in the stands of the former subassociation). Relevés 1–3 in T able 1 are classified only at the rank of the association. Astrantio carniolicae-Pinguiculetum alpinae ass. nov. hoc loco Nomenclatural type, holotypus, is relevé 11 in Table 2. Diagnostic species: Pinguicula alpina, Astran- tia carniolica, Hymenostylium recurvirostre, Palustriella comutata, Petasites paradoxus. var. Molinia arundinacea (relevés 13–19 in Table 2) Campanulo cespitosae-Saxifragetum aizoidis ass. nov. hoc loco Nomenclatural type, holotypus, is relevé 6 in Table 3. Diagnostic species: Saxifraga aizoides, Campa- nula cespitosa, Molinia arundinacea, Palustriella commutata, Calamagrostis varia, Preissia qu- adrata. Variants: var. Pinguicula alpina (relevés 1–2 in Ta- ble 3): differential species Pinguicula al- pina, Carex flacca var. Caltha palustris (relevés 3–4 in Ta- ble 3): differential species Caltha palustris, Gymnostomum aeruginosum. var. Hymenostylium recurvirostre (relevés 5–9 in Table 3): differential species Hyme- nostylium recurvirostre, Tofieldia calyculata. var. Aster bellidiastrum (relevés 10–14 in Table 3): differential species Aster bellidi- astrum, Orthothecium rufescens, Veronica urticifolia, Saxifraga cuneifolia. Phyteumato columnae-Primuletum carniolicae ass. nov. hoc loco Nomenclatural type, holotypus, is relevé 28 in Table 5. Diagnostic species: Primula carniolica, Phy- teuma scheuchzeri subsp. columnae, Paederota lutea, Orthothecium rufescens, Valeriana tript- eris, Carex bracyhstachys. • typicum subass. nov. hoc loco (nomen- clatural type is the same as the nomen- clatural type of the association, relevé 28 in T able 5) var. Viola biflora (relevés 1–3 in Table 5), differential species Viola biflora, Cystopteris fragilis. • astrantietosum carniolicae subass. nov. hoc loco Nomenclatural type, holotypus, is relevé 17 in T able 6. Differential species: Astrantia carniolica, Viola biflora, Valeriana saxatilis. • potentilletosum caulescentis (Dakskobler (1998) 2000) subass. nov. stat. nov. (basionym Primulo carniolicae-Potentille- tum caulescentis Dakskobler (1998) 2000, 19/2 • 2020, 155–231 163 Igor Dakskobler & Andrej Martinčič Plant communities of moist rock crevices with endemic Primula carniolica in the (sub)montane belt of western Slovenia see Dakskobler 1998: 272–273, 286–287; 2000: 56) Nomenclatural type, lectotypus, is relevé 1 in Table 1 (Dakskobler 1998: 286–287). Differential species: Potentilla caulescens, Erica carnea, Polygala chamaebuxus, Prim- ula auricula, P . × venusta. Primulo carniolicae-Potentilletum clusianae ass. nov. hoc loco Nomenclatural type, holotypus, is relevé 4 in Table 8. Diagnostic species: Potentilla clusiana, Primula carniolica, Carex firma, Campanula cochleariifo- lia, Rhodothamnus chamaecistus, Hladnikia pas- tinacifolia, Hieracium pillosum, Aquilegia iulia. Primuletum carniolicae Accetto 2008 violeto- sum biflorae subass. nov. hoc loco Nomenclatural type, holotypus, is relevé 11 in T able 4. Differential species: Viola biflora, Pinguic- ula alpina, Veronica urticifolia. Physoplexido comosae-Saxifragion petraeae Mucina et Theurillat 2015 Paederoto luteae-Potentilletum caulescentis Accetto ex Dakskobler et Martinčič ass. nov. hoc loco (basionym Primulo carniolicae-Potentilletum caulescentis Dakskobler (1998) 2000 var. Asp- lenium seelosii Accetto 2008) Nomenclatural type, lectotypus, is relevé 6 in Table 5 (Accetto 2008: 49–50). Diagnostic species: Potentilla caulescens, Paederota lutea, Primula carniolica, Asple- nium seelosii. Note: Based on the synoptic table (Table 9) and Figure 4 we reduced the rank of the as- sociation Primulo carniolicae-Potentilletum caulescentis Dakskobler (1998) 2000 to the rank of the subassociation Phyteuma- to-Primuletum carniolicae potentilletosum caulescentis. As this subassociation clearly does not comprise the syntaxon Primulo carniolicae-Potentilletum caulescentis var. As- plenium seelosii Accetto 2008 (see Figure 4), we gave it a new name and described a new association, whose author is M. Accetto. The stands of the association Primuletum carniolicae s. lat., which were studied in the central and partly in the southwestern part of the distribution area of Primula carniolica by Dolinar et al. (2013, 2015, 2017), differ from the stands of the association Primuletum carniolicae described by Accetto (2008, 2015) mainly because they comprise Aruncus dioicus, Cystopteris fragilis, Moehringia muscosa, Pinguicula alpina, Hieracium bifidum, Polysti- chum aculeatum and Scopolia carniolica, which indicates slightly moister sites, and in percentages of the groups of diagnostic species (a higher percentage of the species of the class Vaccinio-Piceetea, order Fagetalia sylvaticae and alliances Aremonio-Fagion and Tilio-Acerion – Table 10). For the time being, however, we do not describe them at the level of subassociation. Ecology of the studied communities Astrantio carniolicae-Primuletum carniolicae Distribution area of the stands of this association is shown in Figure 5. They are located in Orehovska Grapa gorge (under Orehek), in Kazarska Grapa (in particular in Beli Potok) under Bukovo and Bukovski Vrh, and in Poličnica at Police (all of them in the Cerkno region, in the pre-Alpine phytogeographical region), and were re- corded also in the Trebuša valley, along the Trebušica, Gačnik, Srna Grapa and Makčeva Grapa gorges, in gorges under Bukov Vrh, Poldanovec and Stanov Rob (Govci), and in Govškarca in the Kanomlja valley. The localities occur at elevations ranging from 230 to 760 m, with a little more than a half of the relevés on shady, and the other on sunny aspects. The parent material is mainly dolomite, with occasional tufa formations, and the aver- age number of species per relevé is 16 (in the range of 7 to 29 species).                                                                       Figure 5: Distribution area of the stands of the association Astrantio carniolicae-Primuletum carniolicae on the map of Slovenia. Slika 5: Območje razširjenosti sestojev asociacije Astrantio carniolicae- -Primuletum carniolicae na zemljevidu Slovenije. 19/2 • 2020, 155–231 164 Igor Dakskobler & Andrej Martinčič Plant communities of moist rock crevices with endemic Primula carniolica in the (sub)montane belt of western Slovenia Astrantio carniolicae-Pinguiculetum alpinae The distribution area of the stands of this association is shown in Figure 6. They are located in Tolmin Gorge, in the Soča ravine under Magozd (at Otona pool and Globoščak gorge), in Velike Luti (Kneža valley), in the Bača ravine at Klonte and Humar (between Koritnica and Hudajužna), at Brinta waterfall at Selce, in Dabrček gorge under the Šentviška Gora Plateau, in Orehovska Grapa, in the Hotenja valley, the T rebuša valley (at the T rebušica and Gačnik streams). The stands of the association were recorded at elevations ranging from 205 to 590 m, with half of the relevés on shady, and the other half on sunny aspects. The parent material is dolomite, limestone, brec- cia, in places admixture of marlstone and chert, and in some spots tufa formations. The average number of spe- cies per relevé is 12 (in the range of 6 to 30 species). cies). The Arctic-Alpine species Saxifraga aizoides is char- acteristic primarily for moist stony sites in the subalpine and alpine belt; it is distributed only in the Alpine phy- togeographical region, with several localities also in the pre-Alpine phytogeographical region: Črni vrh above the stream of Batava, Porezen, Prodarjeva Grapa gorge and the Bača valley between Humar and Klonte (Figure 7). Its localities in the valleys of the Godiča, Volarja and Bača and along the Soča under Magozd are among the lowest- lying in Slovenia (comp. Dakskobler et al. 2017b). In Slovenia, the eastern-Alpine species Campanula cespitosa is distributed also beyond the Alps, in the pre-Alpine and Dinaric phytogeographical regions (Figure 8) and is not rare on rocky and gravelly areas in the submontane belt.                                                                                                                                                                                                                   Figure 6: Distribution area of the stands of the association Astrantio carniolicae-Pinguiculetum alpinae on the map of Slovenia. Slika 6: Območje razširjenosti sestojev asociacije Astrantio carniolicae- -Pinguiculetum alpinae na zemljevidu Slovenije. Campanulo cespitosae-Saxifragetum aizoidis The distribution area of the stands of this association is shown in Figure 7. They are located in the upper course of the Volarja river under Gregorčič waterfall, at the stream Godiča at Beri waterfall and downstream toward Poljubinj, in Prodarjeva Grapa gorge under the village of Porezen, in gorges Linderpoh (under Šprickovbl waterfall) and Kacenpoh at Bača near Podbrdo, in the Zadlaščica river valley at Perbla and in the Tolminka valley (at the Zala gorge). It was recorded at elevations ranging from 255 to 990 m, most often on southeastern and north- eastern aspects. The predominant geological bedrock is limestone with an admixture of marlstone and chert, oc- casionally also dolomite with chert. The average number of species per relevé is 17 (in the range of 10 to 26 spe- Figure 7: Distribution of Saxifraga aizoides (FloVegSi database) and of the stands of the association Campanulo cespitosae-Saxifragetum aizoidis in Slovenia. Slika 7: Razširjenost vrste Saxifraga aizoides (podatkovna baza FloVegSi) in sestojev asociacije Campanulo cespitosae-Saxifragetum aizoidis na zemljevidu Slovenije. Figure 8: Distribution of Campanula cespitosa in Slovenia (FloVegSi database). Slika 8: Razširjenost vrste Campanula cespitosa v Sloveniji (podatkovna baza FloVegSi). 19/2 • 2020, 155–231 165 Igor Dakskobler & Andrej Martinčič Plant communities of moist rock crevices with endemic Primula carniolica in the (sub)montane belt of western Slovenia Phyteumato columnae-Primuletum carniolicae typicum The distribution area of the stands of this subassociation is shown in Figure 9. They are located in the rock faces of Maslenica above the saddle of Oblakov Vrh, in the T rebuša valley (Gačnik, Pršjak, Srna Grapa), under Skopica above the Idrijca valley, under Vrše, Drnulk and Kobilica above the T rebuša valley, under Drnulk and the Stador-Kobilica ridge above the Čepovan valley, in the Sjavnica (Sevnica) gorge under the village of Jagršče, under the hill of De- garnik above Beli Potok gorge and in gorges of Kazar- ska Grapa and Orehovska Grapa, in Govškarca and V Studencu above the Kanomlja valley, under Sončni Rob above Hudo Polje at Vojsko, under Mali Golak. It was recorded at elevations ranging from 215 to 1460 m (Mali Golak) on predominantly shady aspects. This explains the large elevation range of the localities – in the T rebuša val- ley at around 200 m a.s.l., for example, Primula carniolica can be accompanied also by the subalpine species Rho- dodendron hirsutum and Rhodothamnus chamaecistus, and on similar localities and sites Carex ferruginea also occurs at very low elevations. Site conditions in moist and shady gorges of the Trebušica, Gačnik and some other streams with a cold local climate are similar to those in the alti- montane belt. The parent material is almost exclusively dolomite. The average number of species per relevé is 17 (in the range of 6 to 34 species). Gačnik and Makčeva Grapa gorges in the Trebuša valley, under the hill of Hudournik above the Hotenja valley. It was recorded at elevations ranging from 215 to 1440 m, with most of the relevés at 750 m a.s.l., predominantly on shady aspects. The parent material is dolomite, rarely dolomite limestone. The average number of species per relevé is 18 (in the range of 11 to 24 species).                                                                                                                                                                                                                   Figure 9: Distribution area of the stands of the subassociation Phyteu- mato columnae-Primuletum carniolicae typicum on the map of Slovenia. Slika 9: Območje razširjenosti sestojev subasociacije Phyteumato colum- nae-Primuletum carniolicae typicum na zemljevidu Slovenije. Phyteumato columnae-Primuletum carniolicae astrantietosum carniolicae The distribution area of the stands of this subassocia- tion is shown in Figure 10: on the Trnovski Gozd Pla- teau: Sončni Rob above Hudo Polje, under Mts. Mali and Srednji Golak, under Bukov Vrh and Mali Govci; in Figure 10: Distribution area of the stands of the subassociation Phyteu- mato columnae-Primuletum carniolicae astrantietosum carniolicae on the map of Slovenia. Slika 10: Območje razširjenosti sestojev subasociacije Phyteumato co- lumnae-Primuletum carniolicae astrantietosum carniolicae na zemljevidu Slovenije. Phyteumato columnae-Primuletum carniolicae potentilletosum caulescentis The distribution area of the stands of this subassocia- tion is shown in Figure 11: in Orehovska Grapa gorge, in the Idrijca valley under Dolenji Potoki (Straža) and under Sv. Ivan (Šebrelje), under Skopica, Hudournik Figure 11: Distribution area of the stands of the subassociation Phyteu- mato columnae-Primuletum carniolicae potentilletosum caulescentis on the map of Slovenia. Slika 11: Območje razširjenosti sestojev subasociacije Phyteumato columnae-Primuletum carniolicae potentilletosum caulescentis na zemlje- vidu Slovenije. 19/2 • 2020, 155–231 166 Igor Dakskobler & Andrej Martinčič Plant communities of moist rock crevices with endemic Primula carniolica in the (sub)montane belt of western Slovenia and Poldanovec, under Drnulk and Kobilica above the T rebuša valley. Its localities are situated at elevations rang- ing from 240 to 1050 m, with most of them above 550 m a.s.l. Aspects are shady, the parent material is dolomite. The average number of species per relevé is 19 (in the range of 12 to 31 species). Primulo carniolicae-Potentilletum clusianae The localities of the stands of this association are under Zeleni Rob and Poldanovec on the northern edge of the Trnovski Gozd Plateau (Figure 12). Elevations range be- tween 1170 and 1330 m, the parent material is dolomite, aspect shady. The average number of species per relevé is 17 (in the range of 13 to 23 species). In terms of species composition the stands of this association show consider- able similarity and in places contact with the stands of the association Primulo carniolicae-Caricetum firmae (Da- kskobler 2006), but are substantially different in terms of mean coverage and constancy of certain edifier species. In the stands of the association Primulo-Caricetum firmae the species Carex firma and Sesleria caerulea occur at similar constancy to that in the stands of the association Primulo- Potentilletum clusianae, but with a visibly higher mean coverage, and Potentilla clusiana occurs with a substan- tially lower constancy. There are obvious differences also in the constancy and mean coverage of Aster bellidiastrum, Campanula cespitosa (higher in the first compared associa- tion), C. cochleariifolia, Hladnikia pastinacifolia, Hieraci- um pilosum (higher in the second compared association), in the average number of species per relevé (higher in the first compared association) and in mean coverage (cover) of the herb layer (visibly higher in the first compared as- sociation) – comp. Dakskobler (2006, Table 1, relevés 1–23, pp. 90–99). Into the association Primulo carnioli- cae-Potentilletum clusianae we classify stands of moist rock crevices (as demonstrated also by percentages of groups of diagnostic species, Table 10), whereas the stands of stony subalpine grasslands are classified into the association Primulo carniolicae-Caricetum firmae. Primuletum carniolicae violetosum biflorae The distribution area of the stands of this subassociation is shown in Figure 13. Most of the relevés were made in the Raskovec gorge in the Žirovnica valley at Žiri (in Idrija municipality), several relevés also in the gorge of Gačnik and the Govškarca above the Kanomlja valley. It was recorded at elevations ranging from 430 to 850 m on predominantly shady aspects. The parent material is dolomite. The average number of species per relevé is 20 (in the range of 7 to 28 species).                                                                       Figure 12: Localities of the stands of the association Primulo carnioli- cae-Potentilletum clusianae on the northern edge of the T rnovski Gozd Plateau. Slika 12: Nahajališča sestojev asociacije Primulo carniolicae-Potentille- tum clusianae na severnem robu T rnovskega gozda. Figure 13: Distribution area of the stands of the subassociation Primu- letum carniolicae violetosum biflorae on the map of Slovenia. Slika 13: Območje razširjenosti sestojev subasociacije Primuletum carniolicae violetosum biflorae na zemljevidu Slovenije. 19/2 • 2020, 155–231 167 Igor Dakskobler & Andrej Martinčič Plant communities of moist rock crevices with endemic Primula carniolica in the (sub)montane belt of western Slovenia Nature conservation status Primula carniolica is listed in Appendix 2 of the Coun- cil Directive 92/43/EEC on the Conservation of Natural Habitats and of Wild Fauna and Flora – 1992 (Daksko- bler et al. 2004). In the study area, i.e. in the northwest- ern part of its distribution area, this species occurs the most abundantly in shady gorges in the Cerkno region (Kazarska Grapa, Orehovska Grapa, Poličnica, Sjavnica, on the northeastern edge of the Šentviška Gora Plateau under Degarnik and on the pasture at the homestead Sel- jak, in rock faces under Sv. Ivan near Šebrelje above the Idrijca valley) and in the T rebuša valley (Gačnik, Makčeva Grapa, Srna Grapa, Pršjak, Mali Govci, also along the Trebušica and the gorges that flow into this river on the left bank), and in the rock faces of Govci from Bukov Vrh across Zeleni Rob, Poldanovec and Stanov Rob to Stador, Drnulk and Skopica). The population is abun- dant and vital, and has not yet been affected by the hu- man impact, except on isolated localities. Also abundant is the population in the Raskovec gorge in the Železnica valley in the Žiri region (Dakskobler et al. 2017a, Do- linar et al. 2017), in Govškarca, Klavžarica and V Stu- dencu above the Kanomlja valley. Another two species of European conservation concern sporadically occur in the studied communities of moist rock crevices, namely Aquilegia iulia and Hladnikia pastinacifolia (Čušin et al. 2004, Podobnik et al. 2013) as well as a number of protected species (Anon. 2004): Cyclamen purpurascens, Gentiana clusii, Gymnadenia odoratissima, Hemerocallis lilioasphodelus, Leontopodium alpinum, Pinguicula alpina, Primula auricula, Primula × venusta, and T axus baccata as well as several other Red List species (Anon. 2002): Scho- enus nigricans, Veratrum nigrum. Unlike flowering plants, bryophytes do not comprise rare or phytogeographically important species, nor do they comprise species from the Slovenian (Martinčič 2016) or European Red List (Hodgetts et al. 2019). The studied communities belong to Natura 2000 type 8210 Calcareous rocky slopes with chasmophytic vegetation, and the stands of associations Astrantio-Primuletum and Astrantio-Pinguiculetum also to Natura 2000 habitat type 7220 Petrifying springs with tufa formation (Cratoneurion) – Jogan et al. (2004). Conclusions A large number of the relevés made in the moist rocks in the submontane and montane belt (mainly in the area of Illyrian beech forests from the alliance Aremonio-Fagion) in western Slovenia between the southern Julian Alps and the northern Dinaric Alps can be arranged in tables in different ways. Our classification followed certain field findings where it was established that as a rule only a few dominant species occur with high mean coverage in these special site conditions. In the predominantly dolo- mite bedrock it is the moisture conditions that determine which species will dominate. In the southern Julian Alps, where the bedrock comes also as platy limestones with an admixture of marlstone and chert, this depends on other factors as well. With this in mind, it makes sense to classi- fy the stands with Primula carniolica on the moistest rocks into the association Astrantio carniolicae-Primuletum car- niolicae, because Pinguicula alpina does not have as high constancy and mean coverage here as in the stands of a similar association, Astrantio-Pinguiculetum alpinae, in which Primula carniolica does not occur, as these stands are, at least in part, outside its distribution area. All other relevés of rock crevices with Primula carniolica could be classified into the association Primuletum car- niolicae (in the broadest sense), which was first recorded in Slovenia and named by T . Wraber (he found it at Divje Jezero / Wild Lake at Idrija already on 16/8/1973, and later in the ravine of Kadice at Sodražica on 25/4/1982). Unfortunately, Wraber never processed or published his relevés and we only came across them by chance in 2016 and 2017 when we studied his legacy, which is kept at Wraber’s Library at the University of Ljubljana’s Botani- cal Garden. Tone Wraber was without a doubt a pioneer in the phytosociological investigation of chasmophytic communities with Primula carniolica, but his relevés can- not have been known to Accetto (2008), who was the first to validly publish the description of the associa- tion Primuletum carniolicae. The relevés of the associa- tion Primuletum carniolicae in the northwestern part of the distribution area are nevertheless slightly different from the relevés of this association in the central and southwestern part of its distribution area. The most ap- propriate syntaxonomic solution would be to describe a new geographical variant with the subspecies Phyteuma scheuchzeri subsp. columnae. However, given the results of hierarchical classification we could also describe a new association, Phyteumato-Primuletum carniolicae, which would allow us to evaluate the established variability of individual stands at the rank of subassociations and not only at the rank of variants. Wraber’s notebook comprises a relevé dated 3 May 1982, locality Strug, rocks on the right side of the road from Fežnar toward Divje Jezero lake, which comprises most of the diagnostic species of the subassociation Phyteumato-Primuletum carniolicae as- trantietosum carniolicae. The decision for an independent association Phyteumato-Primuletum carniolicae, on the other hand, means reducing the rank of the association Primulo carniolicae-Potentilletum caulescentis to the rank 19/2 • 2020, 155–231 168 Igor Dakskobler & Andrej Martinčič Plant communities of moist rock crevices with endemic Primula carniolica in the (sub)montane belt of western Slovenia of subassociation (which could not happen if we insisted on the association Primuletum carniolicae in the broadest sense – comp. Accetto 2008). Because Primula carniolica characterises very hygrophytic chasmophytic communi- ties it is rightly considered a character species of the al- liance Cystopteridion (not Violo biflorae-Cystopteridion sensu Mucina et al. 2016), into which we provisionally classify most of the described new syntaxa (except for as- sociation Paederoto luteae-Potentilletum caulescentis, which is classified into the alliance Physoplexido comosae-Saxifra- gion petraeae). The article also proposes a new possible alliance of hygrophytic rock crevice (chasmophytic) com- munities of the Southeastern Alps and northern Dinaric Alps, an ecological vicariant of the alliance Physoplexido comosae-Saxifragion petraeae. The name-giving species for such an alliance could be Paederota lutea and Astrantia carniolica (Astrantio carniolicae-Paederotion luteae nom. prov.). However, a valid description of a new alliance re- quires further research that takes into account also similar syntaxa of hygrophytic chasmophytic communities from other parts of the Southeastern Alps and northern Di- naric Alps, and more extensive numerical comparisons. Povzetek Združbe vlažnih skalnih razpok z endemitom Primula carniolica v (sub)montanskem pasu zahodne Slovenije Primula carniolica, evropsko varstveno pomembna vrsta (Natura 2000), je v severozahodnem delu svojega areala (v zahodni Sloveniji med južnimi Julijskimi Alpami in severnim delom Dinarskega gorstva) najbolj množično prisotna v senčnih grapah na Cerkljanskem (Kazarska grapa, Orehovska grapa, Poličnica, Sjavnica, v ostenjih pod sv. Ivanom nad dolino Idrijce pod Šebreljami, tudi na severovzhodnem robu Šentviške planote: pod Degar- nikom in na pašniku pri domačiji Seljak v Bukovskem Vrhu, že na Tolminskem) in v dolini Trebuše (Gačnik, Makčeva grapa, Srna grapa, Pršjak, Mali Govci, tudi ob sami Trebušici in grapah, ki se vanjo izlivajo na levem bregu) ter v ostenju Govcev od Bukovega vrha preko Zelenega roba, Poldanovca in Stanovega roba vse do Sta- dorja, Drnulka in Skopice). Populacija je bogata, vitalna in za zdaj človek nanjo razen na nekaterih nahajališčih nima večjega vpliva. Bogata je tudi populacija v grapi Raskovec v dolini Železnice na Žirovskem in v Govškarci, Klavžarici in V Studencu nad dolino Kanomlje. Veliko število fitocenoloških popisov, ki smo jih nar- edili v vlažnem skalovju v submontanskem in montan- skem pasu (večinoma v območju ilirskih bukovih goz- dov iz zveze Aremonio-Fagion) v raziskovanem območju je mogoče v tabele razvrščati na več načinov. Naša ure- ditev je sledila nekaterim spoznanjem na terenu, kjer je očitno, da je v teh posebnih rastiščnih razmerah navadno le malo prevladujočih vrst z večjim srednjim zastiranjem. Katere so, pa je v prevladujoči dolomitni podlagi pred- vsem odvisno od vlažnostnih razmer. V južnih Julijskih Alpah, kjer so podlaga tudi ploščasti apnenci s primesjo laporovca in roženca, lahko nanjo vplivajo tudi drugi de- javniki. Ob tej podmeni je uvrstitev sestojev z vrsto Pri- mula carniolica na najbolj vlažnem skalovju v asociacijo Astrantio carniolicae-Primuletum carniolicae smiselna, saj v njej vrsta Pinguicula alpina nima tako velike stalnosti in srednjega zastiranja kot v sestojih podobne asociacije Astrantio-Pinguiculetum alpinae, v kateri pa vrsta Primula carniolica ne uspeva in so ti sestoji vsaj deloma tudi zunaj njenega areala. Vse ostale popise skalnih razpok, kjer uspeva vrsta Pri- mula carniolica, bi lahko uvrstili v široko zajeto asociacijo Primuletum carniolicae, ki jo je v Sloveniji prvi popisal in tako imenoval T. Wraber (pri Divjem jezeru pri Idriji že 16. 8. 1973, v soteski Kadice pri Sodražici 25. 4. 1982). Žal popisov ni nikoli obdelal ali objavil in smo nekatere od njih slučajno opazili v letih 2016 in 2017 v njegovi zapuščini, ki jo hrani Wraberjeva knjižnica v Botaničnem vrtu Univerze v Ljubljani. Vsekakor je bil Tone Wraber pionir pri fitocenološkem raziskovanju združb skalnih razpok z vrsto Primula carniolica, a njegovih popisov Ac- cetto (2008), ki je prvi veljavno objavil opis asociacije Primuletum carniolicae ni mogel poznati. Popisi asociacije Primuletum carniolicae v severozahodnem delu areala se vendarle nekoliko razlikujejo od popisov te asociacije v osrednjem in jugozahodnem delu areala. Najprimernejša sintaksonomska rešitev bi bil opis nove geografske vari- ante s podvrsto Phyteuma scheuchzeri subsp. columnae. Toda rezultati hierarhične klasifikacije dopuščajo tudi opis nove asociacije Phyteumato-Primuletum carniolicae, ki omogoča, da tudi ugotovljeno variabilnost posameznih sestojev vrednotimo na rangu subasociacij in ne zgolj na rangu variant. V Wraberjevi beležnici, datum popisa je 3. 5. 1982, nahajališče pa Strug, skalovje na desni strani ceste od Fežnarja proti Divjemu jezeru, je popis, ki vs- ebuje večino diagnostičnih vrst subasociacije Phyteu- mato-Primuletum carniolicae astrantietosum carniolicae. Odločitev za samostojno asociacijo Phyteumato-Primule- tum carniolicae pa ima za posledico tudi znižanje ranga asociacije Primulo carniolicae-Potentilletum caulescentis na rang subasociacije (kar bi, če bi ostali pri široko zajeti asociaciji Primuletum carniolicae, ne mogli storiti – prim. Accetto 2008). Ker vrsta Primula carniolica označuje precej vlagoljubne združbe skalnih razpok, jo upravičeno štejemo za značilnico zveze Cystopteridion (ne Violo biflo- rae-Cystopteridion v smislu Mucina et al. 2016), v katero 19/2 • 2020, 155–231 169 Igor Dakskobler & Andrej Martinčič Plant communities of moist rock crevices with endemic Primula carniolica in the (sub)montane belt of western Slovenia za zdaj uvrščamo večino novo opisanih sintaksonov (ra- zen asociacije Paederoto luteae-Potentilletum caulescentis, ki jo uvrščamo v zvezo Physoplexido comosae-Saxifragion petraeae). V članku razmišljamo tudi o novi zvezi združb vlažnih skalnih razpok v Jugovzhodnih Alpah in severnem delu Dinarskega gorstva, ki bi bila ekološka vikarianta zvezi Physoplexido comosae-Saxifragion petraeae. Primerni vrsti za poimenovanje take zveze sta vrsti Paederota lutea in Astrantia carniolica (Astrantio carniolicae-Paederotion luteae nom. prov.). Za veljaven opis takšne zveze bi bile potrebne nadaljnje raziskave, predvsem pa upoštevanje podobnih sintaksonov iz drugih delov Jugovzhodnih Alp in Dinarskega gorstva in bolj obsežna primerjava z numeričnimi metodami. V preučenih združbah vlažnih skalnih razpok pone- kod uspevata še dve evropsko varstveni pomembni vrsti Aquilegia iulia in Hladnikia pastinacifolia in večje število zavarovanih vrst: Cyclamen purpurascens, Gentiana clusii, Gymnadenia odoratissima, Hemerocallis lilioasphodelus, Le- ontopodium alpinum, Pinguicula alpina, Primula auricula, Primula × venusta in Taxus baccata in nekaterih drugih vrst iz rdečega seznama: Schoenus nigricans, Veratrum nig- rum. Preučene združbe sodijo v Natura 2000 habitatni tip 8210 Karbonatna skalnata pobočja z vegetacijo skalnih razpok, sestoji asociacij Astrantio-Primuletum in Astran- tio-Pinguiculetum tudi v Natura 2000 habitatni tip 7220 Lehnjakotvorni izviri (Cratoneurion). Acknowledgements The authors owe special thanks to Dr. Branko Vreš, Mag. Andrej Seliškar, Brane Anderle, Janez Mihael Kocjan and Branko Dolinar, co-authors of Figures 1, 3, 8 and 9. Iz- tok Sajko prepared Figures 1–13 and Prof. Dr. Andrej Rozman Figure 14 (Appendix) for print. We would like to thank the heirs of late Tone Wraber for giving his manu- scripts and professional literature to the safekeeping of the Botanical Garden of the University of Ljubljana, and to its director, Dr. Jože Bavcon, who allowed us to examine professor’s legacy. Sincere thanks to Prof. Dr. Jean-Paul Theurillat for his valuable data and advice about the rank- ing of new syntaxa. Three anonymous reviewers helped us with valuable improvements and corrections. We also acknowledge the financial support from the Slovenian Research Agency (research core funding No. P1-0236). English translation by Andreja Šalamon Verbič. 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Biološki vestnik 34 (1): 115–124. Appendix Figure 14: Dendrogram of relevés of rock crevice communities with Primula carniolica in the northwestern part of its ditribu- tion area. Slika 14: Dendrogram popisov združb skalnih razpok z vrsto Primula carniolica v severozahodnem delu njenega areala.        19/2 • 2020, 155–231 172 Igor Dakskobler & Andrej Martinčič Plant communities of moist rock crevices with endemic Primula carniolica in the (sub)montane belt of western Slovenia N umber of relevé (Zaporedna štev . popisa) 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 31 32 33 34 35 36 37 38 39 40 41 42 43 Database number of relevé (Delovna številka popisa) 269367 263162 269183 269213 274405 274329 278101 234544 234553 274475 278088 269264 241571 241573 271067 274320 273974 275147 275162 258062 268309 271065 260958 266149 273716 258590 269215 269184 263158 269178 269256 263160 263180 269180 263163 269187 269193 266721 269201 269253 263171 266719 266711 Elevation in m (Nadmorska višina v m) 450 435 435 410 600 480 435 590 700 455 540 590 760 750 310 380 755 430 430 230 278 270 230 255 240 480 408 435 430 425 570 430 465 430 440 475 520 440 470 560 475 440 370 Aspect (Lega) SSE NE N N N N SW W N SW SW S NE N S NNE N SSW NNE N E W W NE NW SE S E SW SE S N N NE N SW NE S NEE NE NE S N Slope in degrees (Nagib v stopinjah) 90 90 90 80 80 85 80 80 95 90 90 70 80-100 80 90 80 85 90 90 95 80 85-95 95 80 95 80 85 90 80 85 85 80 80 85 90 90 90 85 80 80 90 80 90 Parent material (Matična podlaga) D D D D D D D D D D D D D D D D D D D D D D D D D D D D Tu D D D D D D D D Tu D D D D D Soil (Tla) Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Stoniness in % (Kamnitost v %) 100 100 100 100 100 100 100 100 100 100 100 100 60 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 Cover of herb layer in % (Zastiranje zeliščne plasti v %): E1 20 20 30 30 20 30 30 30 20 30 30 30 40 50 30 40 30 30 20 30 30 30 30 30 25 30 30 60 40 30 35 30 40 30 60 40 20 40 30 30 30 40 30 Cover of moss layer in % (Zastiranje mahovne plasti v %) E0 20 20 20 40 20 20 10 20 20 20 30 20 30 30 10 50 20 20 30 30 20 20 20 40 30 30 40 30 80 80 60 70 60 80 40 40 30 70 30 50 40 39 40 Number of species (Število vrst) 15 20 13 14 18 26 10 18 16 12 11 11 21 14 21 22 16 27 24 32 17 17 26 12 27 15 21 23 10 8 13 13 14 14 15 14 14 11 10 15 13 19 12 Relevé area (Velikost popisne ploskve) m 2 5 10 5 5 10 10 15 10 10 10 20 5 10 10 10 10 10 15 15 10 10 10 10 10 10 5 10 5 10 10 10 10 10 5 10 10 10 20 5 5 10 10 10 Date of taking relevé (Datum popisa) 3/31/2017 9/12/2016 4/7/2017 4/13/2017 4/23/2018 5/8/2018 12,6.2019 8/26/2003 8/26/2003 6/14/2018 12,6.2019 5/25/2017 5/5/2011 5/5/2011 4/29/2018 5/8/2018 7/2/2018 4/22/2019 4/22/2019 8/30/2015 13,8,2017 4/29/2018 4/17/2016 4/9/2017 9/6/2018 9/29/2015 4/13/2017 4/7/2017 9/12/2016 4/7/2017 5/25/2017 9/12/2016 9/12/2016 4/7/2017 9/12/2016 4/7/2017 4/7/2017 4/13/2017 4/7/2017 5/25/2017 9/12/2016 4/13/2017 4/13/2017 Locality (Nahajališče) Kazarska grapa-Žabže Orehovska grapa Orehovska grapa Kazarska grapa-Beli potok Kazarska grapa Gorenja T rebuša Srna grapa Gačnik-Statarska grape Poldanovec - Govčenk Stanov rob - Govška grapa Gorenja T rebuša Makčeva grapa Gačnik Police-Poličnica Bukov vrh Brusova grapa Bukov vrh-Suha grapa Gorenja T rebuša Srna grapa Gorenja T rebuša Srna grapa Gačnik Gačnik Gačnik-Statarska grape Gorenja T rebuša - Žveplenica Gorenja T rebuša - T rebušica Gorenja T rebuša - T rebušica Gorenja T rebuša - Žveplenica Gačnik Gorenja T rebuša - Žrčin Govškarca-Kanomlja Kazarska grapa - Bei potok Orehovska grapa Orehovska grapa Orehovska grapa Police-Poličnica Orehovska grapa Orehovska grapa Orehovska grapa Orehovska grapa Orehovska grapa Orehovska grapa Kazarska grapa-Beli ptok Orehovska grapa Police-Poličnica Orehovska grapa Kazarska grapa - Beli potok Kazarska grapa - Beli potok Quadrant (Kvadrant) 9849/4 9849/4 9849/4 9849/3 9849/3 9949/3 9949/1 9948/4 9948/4 9949/3 9949/1 9849/3 9949/3 9949/3 9949/3 9949/3 9949/1 9949/1 9949/1 9948/2 9948/4 9948/4 9948/2 9949/1 9948/2 9949/4 9849/3 9849/4 9849/4 9849/4 9849/3 9849/4 9849/4 9849/4 9849/4 9849/4 9849/4 9849/3 9849/4 9849/3 9849/4 9849/3 9849/3 Coordinate GK Y (D-48) m 416348 418168 418167 415883 415406 410696 411018 409507 409086 410176 411215 415172 413871 413929 410038 410195 412054 410952 411029 409693 409690 409679 409629 410415 409699 422008 415865 418170 418202 418217 415246 418199 418005 418186 418136 418148 418033 415685 418135 415303 418003 415705 415587 Coordinate GK X (D-48) m 5110970 5110871 5110855 5110618 5111232 5100026 5103400 5097980 5098562 5100701 5103015 5110614 5095748 5095734 5100136 5100069 5101908 5103425 5103378 5103390 5100882 5100840 5103317 5104084 5102978 5099627 5110646 5110868 5110828 5110864 5110577 5110837 5111065 5110842 5110900 5110934 5111054 5110658 5110903 5110488 5111044 5110657 5110716 Diagnostic species of the association (Diagnostične vrste asociacije) AP Primula carniolica E1 1 3 3 2 2 2 1 r + 2 + 2 1 2 2 + 1 1 1 1 1 1 2 1 + 3 3 3 + 1 + 2 2 3 2 3 2 2 1 2 2 3 2 AP Palustriella commutata E0 + + . 1 1 + . 1 1 1 2 1 1 1 1 1 1 2 2 + + . 1 1 1 2 3 1 4 4 3 3 2 2 2 2 1 2 2 3 1 2 1 AP Astrantia carniolica E1 . 1 + 1 + . + 2 2 2 2 1 1 1 1 1 + + + 1 + 1 + . + . 1 + 3 1 3 2 2 + 1 . 1 2 1 + 2 2 2 AP Pinguicula alpina E1 1 . + . + . . 1 + . 1 . + + + + . + + 2 2 1 . + 1 + + 1 1 + 3 + 1 + 1 2 2 2 3 3 + 2 3 AP Hymenostylium recurvirostre E0 + 1 1 . . + + 3 . + 1 . . . + . . 1 . + . + + . 1 2 1 1 4 4 4 4 3 4 3 3 3 3 2 2 2 1 3 PsSp Phyteuma scheuchzeri subsp. columnae E1 + + + + + + + r . . . + + . + + + + . + + + + . . + r + . . . . . + . . . . . r . r . EP Molinia arundinacea E1 . . . . . . . + + 1 1 + . . . . . + . . . + . + . . . . + . 1 . + . + + + . . + 1 + . AP Valeriana saxatilis E1 . . . . . . + + + . + . + + . . + + + 1 1 1 1 3 2 . . . . . . . . . + + . . . . . . . ES Carex mucronata E1 . . . . . . . r . . . + . . . . . . . 1 . + + . . . . . . . . . . . + 1 + + + . + + + Diagnostic species of the subassociation (Diagnostične vrste subasociacij) AP Paederota lutea E1 . . . . + 1 + 2 1 + 1 . 2 1 2 2 1 1 2 1 + 1 2 1 1 + . . . . . . . . . . . . . . . . . AP Aster bellidiastrum E1 . . . 1 + + + 1 + 1 1 2 1 + 1 1 1 + + 2 2 + 1 1 + + . . . . . . . . . . . . . . . . . Table 1 (Tabela 1): Astrantio carniolicae-Primuletum carniolicae. Relevé number 1–43. 19/2 • 2020, 155–231 173 Igor Dakskobler & Andrej Martinčič Plant communities of moist rock crevices with endemic Primula carniolica in the (sub)montane belt of western Slovenia N umber of relevé (Zaporedna štev . popisa) 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 31 32 33 34 35 36 37 38 39 40 41 42 43 Database number of relevé (Delovna številka popisa) 269367 263162 269183 269213 274405 274329 278101 234544 234553 274475 278088 269264 241571 241573 271067 274320 273974 275147 275162 258062 268309 271065 260958 266149 273716 258590 269215 269184 263158 269178 269256 263160 263180 269180 263163 269187 269193 266721 269201 269253 263171 266719 266711 Elevation in m (Nadmorska višina v m) 450 435 435 410 600 480 435 590 700 455 540 590 760 750 310 380 755 430 430 230 278 270 230 255 240 480 408 435 430 425 570 430 465 430 440 475 520 440 470 560 475 440 370 Aspect (Lega) SSE NE N N N N SW W N SW SW S NE N S NNE N SSW NNE N E W W NE NW SE S E SW SE S N N NE N SW NE S NEE NE NE S N Slope in degrees (Nagib v stopinjah) 90 90 90 80 80 85 80 80 95 90 90 70 80-100 80 90 80 85 90 90 95 80 85-95 95 80 95 80 85 90 80 85 85 80 80 85 90 90 90 85 80 80 90 80 90 Parent material (Matična podlaga) D D D D D D D D D D D D D D D D D D D D D D D D D D D D Tu D D D D D D D D Tu D D D D D Soil (Tla) Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Stoniness in % (Kamnitost v %) 100 100 100 100 100 100 100 100 100 100 100 100 60 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 Cover of herb layer in % (Zastiranje zeliščne plasti v %): E1 20 20 30 30 20 30 30 30 20 30 30 30 40 50 30 40 30 30 20 30 30 30 30 30 25 30 30 60 40 30 35 30 40 30 60 40 20 40 30 30 30 40 30 Cover of moss layer in % (Zastiranje mahovne plasti v %) E0 20 20 20 40 20 20 10 20 20 20 30 20 30 30 10 50 20 20 30 30 20 20 20 40 30 30 40 30 80 80 60 70 60 80 40 40 30 70 30 50 40 39 40 Number of species (Število vrst) 15 20 13 14 18 26 10 18 16 12 11 11 21 14 21 22 16 27 24 32 17 17 26 12 27 15 21 23 10 8 13 13 14 14 15 14 14 11 10 15 13 19 12 Relevé area (Velikost popisne ploskve) m 2 5 10 5 5 10 10 15 10 10 10 20 5 10 10 10 10 10 15 15 10 10 10 10 10 10 5 10 5 10 10 10 10 10 5 10 10 10 20 5 5 10 10 10 Date of taking relevé (Datum popisa) 3/31/2017 9/12/2016 4/7/2017 4/13/2017 4/23/2018 5/8/2018 12,6.2019 8/26/2003 8/26/2003 6/14/2018 12,6.2019 5/25/2017 5/5/2011 5/5/2011 4/29/2018 5/8/2018 7/2/2018 4/22/2019 4/22/2019 8/30/2015 13,8,2017 4/29/2018 4/17/2016 4/9/2017 9/6/2018 9/29/2015 4/13/2017 4/7/2017 9/12/2016 4/7/2017 5/25/2017 9/12/2016 9/12/2016 4/7/2017 9/12/2016 4/7/2017 4/7/2017 4/13/2017 4/7/2017 5/25/2017 9/12/2016 4/13/2017 4/13/2017 Locality (Nahajališče) Kazarska grapa-Žabže Orehovska grapa Orehovska grapa Kazarska grapa-Beli potok Kazarska grapa Gorenja T rebuša Srna grapa Gačnik-Statarska grape Poldanovec - Govčenk Stanov rob - Govška grapa Gorenja T rebuša Makčeva grapa Gačnik Police-Poličnica Bukov vrh Brusova grapa Bukov vrh-Suha grapa Gorenja T rebuša Srna grapa Gorenja T rebuša Srna grapa Gačnik Gačnik Gačnik-Statarska grape Gorenja T rebuša - Žveplenica Gorenja T rebuša - T rebušica Gorenja T rebuša - T rebušica Gorenja T rebuša - Žveplenica Gačnik Gorenja T rebuša - Žrčin Govškarca-Kanomlja Kazarska grapa - Bei potok Orehovska grapa Orehovska grapa Orehovska grapa Police-Poličnica Orehovska grapa Orehovska grapa Orehovska grapa Orehovska grapa Orehovska grapa Orehovska grapa Kazarska grapa-Beli ptok Orehovska grapa Police-Poličnica Orehovska grapa Kazarska grapa - Beli potok Kazarska grapa - Beli potok Quadrant (Kvadrant) 9849/4 9849/4 9849/4 9849/3 9849/3 9949/3 9949/1 9948/4 9948/4 9949/3 9949/1 9849/3 9949/3 9949/3 9949/3 9949/3 9949/1 9949/1 9949/1 9948/2 9948/4 9948/4 9948/2 9949/1 9948/2 9949/4 9849/3 9849/4 9849/4 9849/4 9849/3 9849/4 9849/4 9849/4 9849/4 9849/4 9849/4 9849/3 9849/4 9849/3 9849/4 9849/3 9849/3 Coordinate GK Y (D-48) m 416348 418168 418167 415883 415406 410696 411018 409507 409086 410176 411215 415172 413871 413929 410038 410195 412054 410952 411029 409693 409690 409679 409629 410415 409699 422008 415865 418170 418202 418217 415246 418199 418005 418186 418136 418148 418033 415685 418135 415303 418003 415705 415587 Coordinate GK X (D-48) m 5110970 5110871 5110855 5110618 5111232 5100026 5103400 5097980 5098562 5100701 5103015 5110614 5095748 5095734 5100136 5100069 5101908 5103425 5103378 5103390 5100882 5100840 5103317 5104084 5102978 5099627 5110646 5110868 5110828 5110864 5110577 5110837 5111065 5110842 5110900 5110934 5111054 5110658 5110903 5110488 5111044 5110657 5110716 Diagnostic species of the association (Diagnostične vrste asociacije) AP Primula carniolica E1 1 3 3 2 2 2 1 r + 2 + 2 1 2 2 + 1 1 1 1 1 1 2 1 + 3 3 3 + 1 + 2 2 3 2 3 2 2 1 2 2 3 2 AP Palustriella commutata E0 + + . 1 1 + . 1 1 1 2 1 1 1 1 1 1 2 2 + + . 1 1 1 2 3 1 4 4 3 3 2 2 2 2 1 2 2 3 1 2 1 AP Astrantia carniolica E1 . 1 + 1 + . + 2 2 2 2 1 1 1 1 1 + + + 1 + 1 + . + . 1 + 3 1 3 2 2 + 1 . 1 2 1 + 2 2 2 AP Pinguicula alpina E1 1 . + . + . . 1 + . 1 . + + + + . + + 2 2 1 . + 1 + + 1 1 + 3 + 1 + 1 2 2 2 3 3 + 2 3 AP Hymenostylium recurvirostre E0 + 1 1 . . + + 3 . + 1 . . . + . . 1 . + . + + . 1 2 1 1 4 4 4 4 3 4 3 3 3 3 2 2 2 1 3 PsSp Phyteuma scheuchzeri subsp. columnae E1 + + + + + + + r . . . + + . + + + + . + + + + . . + r + . . . . . + . . . . . r . r . EP Molinia arundinacea E1 . . . . . . . + + 1 1 + . . . . . + . . . + . + . . . . + . 1 . + . + + + . . + 1 + . AP Valeriana saxatilis E1 . . . . . . + + + . + . + + . . + + + 1 1 1 1 3 2 . . . . . . . . . + + . . . . . . . ES Carex mucronata E1 . . . . . . . r . . . + . . . . . . . 1 . + + . . . . . . . . . . . + 1 + + + . + + + Diagnostic species of the subassociation (Diagnostične vrste subasociacij) AP Paederota lutea E1 . . . . + 1 + 2 1 + 1 . 2 1 2 2 1 1 2 1 + 1 2 1 1 + . . . . . . . . . . . . . . . . . AP Aster bellidiastrum E1 . . . 1 + + + 1 + 1 1 2 1 + 1 1 1 + + 2 2 + 1 1 + + . . . . . . . . . . . . . . . . . 19/2 • 2020, 155–231 174 Igor Dakskobler & Andrej Martinčič Plant communities of moist rock crevices with endemic Primula carniolica in the (sub)montane belt of western Slovenia N umber of relevé (Zaporedna štev . popisa) 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 31 32 33 34 35 36 37 38 39 40 41 42 43 AP Cystopteridion s. lat. (Astrantio-Paederotion luteae nom. prov.) Orthothecium rufescens E0 1 + 1 2 1 + . + + + . + 2 1 + + 1 1 2 1 + . 1 1 1 2 1 2 . . 1 + + 1 . 1 1 . 1 1 + . 3 Tofieldia calyculata E1 . r . . . . . . . . . . 1 . + . . + + + + + + + 1 . + + . . . + 1 + + + + 1 + + + + . Aster bellidiastrum E1 . . . 1 + + + 1 + 1 1 2 1 + 1 1 1 + + 2 2 + 1 1 + + . . . . . . . . . . . . . . . . . Valeriana saxatilis E1 . . . . . . + + + . + . + + . . + + + 1 1 1 1 3 2 . . . . . . . . . + + . . . . . . . Carex brachystachys E1 . . . . 1 r + 2 + . . . + . + + . r . . 1 . . . + 1 + . . . . . . . . . . . . . . . + Preissia quadrata E0 . 1 . . 1 . . . . . . . . . . . . . . + . . . . + . + + . . + . + . . . + . . + + . . Asplenium viride E1 . + . . . + . . . . . . . . + + + + + . . . . . . . . + . . . 1 . . . . . . . . . . . Valeriana tripteris E1 . . . . . + . . . r . . . . + + . . 1 . . + . . . . . . . . . . . . . . . . . . . . . Fissidens dubius E0 . + . . . + . . . . . . . . . . . . . + . . . . . . . . . . . . . . . . . . . . . . . Barbula crocea E0 . . . . . . . . . . . . . . . . . . . . . . + . 1 . . 1 . . . . . . . . . . . . . . . Viola biflora E1 . . . . . . . + 1 . . . . . . . + . + . . . . . . . . . . . . . . . . . . . . . . . . Eucladium verticillatum E0 + . . . . . . . . . . . . . . . . . . . . . . . . . + + . . . + . . . . . . . . . . . Pellia endiviifolia E0 . . . . . + . . . . . . . . . . . . . . . . . . 1 . . . + . . . . . . . . . . . . . . Soldanella minima E1 . . . . . . . . . . . . . 1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Moehringia muscosa E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . PcSp Physoplexido comosae-Saxifragion petraeae Campanula cespitosa E1 . . . . . . . . . . . . . . . . . . . . . . . . . + r . . . 1 . . . 1 1 + . . + + + . Hieracium pospichalii E1 . . . . . . + . . r . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . PC Potentilletalia caulescentis Potentilla caulescens E1 . . + . . . . . r . . . . . . . . r . + . . . . . . . . . . . r . . . + + r . . r r . Kernera saxatilis E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . + . . . . . . . . . . . . AT Asplenietea trichomanis Asplenium trichomanes E1 + . + . + + . . r r . . . . . . . . . . . . . . + . . . . . . . . . . . . . . . . . . Asplenium ruta-muraria E1 . . . . + . . . . . . . . . + . . + . r . . . . . . + . . . . . . . . . . . . . . . . TR Thlaspietea rotundifolii Hieracium bifidum E1 . + + . . . . . . . . . . + . . . . . + . . r . + r . . . . . . . . . . + . . . . . . Adenostyles glabra E1 . . . . . . . + . . . . r + . . + + + . . . . . . + . . . . . . . . . . . . . . . . . Gymnocarpium robertianum E1 . . . . + . . . . . . . . . . + . . r . . . . . r . . . . . . . . . . . . . . . . . . Achnatherum calamagrostis E1 . . . . . . . . . . . . . . . . . . . r . . . . . . . . . . . . . . . . . . . . . . . Petasites paradoxus E1 . . . . . . . . . . . . . . . . . . . r . . . . . . . . . . . . . . . . . . . . . . . Hieracium porrifolium E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . CD Caricetalia davallianae Parnassia palustris E1 . . . . . . . . r . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + Schoenus nigricans E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + + . . + . . . . MC Montio-Cardaminetea Conocephalum conicum E0 + + 2 1 1 + . . . 1 . . . . + 2 . . . + . . + + 1 . . 1 . . . + 1 1 . . . . . . . + . Campylium stellatum E0 . + . . . . . . . . . . . . . . . . . . . . + . . . . . . . + . . . . . . . . . . . . Bryum pseudotriquetrum E0 . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . + . . . . . . . . . . . . Cratoneuron filicinum E0 + . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . . . . . . . Distichium capillaceum E0 . . . . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . . . . . . . . . . Campylium calcareum E0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Campylium elodes E0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Ditrichum flexicaule E0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . ES Elyno-Seslerietea Sesleria caerulea E1 . . . . . . 1 r . . + + + 1 . . . + + + . . + 1 r . + . + . . . 1 . + 1 1 1 1 + . + 1 Phyteuma orbiculare E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . Carex ferruginea E1 . . . . . . . . + . . . . . . . r . . r . . + . . . . . . . . . . . . . . . . . . . . Carex firma E1 . . . . . . . . . . . . 2 2 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Laserpitium peucedanoides E1 . . . . . . . . . . . . r . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Betonica alopecuros E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . r . . . . . . . . . . . . . . . 19/2 • 2020, 155–231 175 Igor Dakskobler & Andrej Martinčič Plant communities of moist rock crevices with endemic Primula carniolica in the (sub)montane belt of western Slovenia N umber of relevé (Zaporedna štev . popisa) 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 31 32 33 34 35 36 37 38 39 40 41 42 43 AP Cystopteridion s. lat. (Astrantio-Paederotion luteae nom. prov.) Orthothecium rufescens E0 1 + 1 2 1 + . + + + . + 2 1 + + 1 1 2 1 + . 1 1 1 2 1 2 . . 1 + + 1 . 1 1 . 1 1 + . 3 Tofieldia calyculata E1 . r . . . . . . . . . . 1 . + . . + + + + + + + 1 . + + . . . + 1 + + + + 1 + + + + . Aster bellidiastrum E1 . . . 1 + + + 1 + 1 1 2 1 + 1 1 1 + + 2 2 + 1 1 + + . . . . . . . . . . . . . . . . . Valeriana saxatilis E1 . . . . . . + + + . + . + + . . + + + 1 1 1 1 3 2 . . . . . . . . . + + . . . . . . . Carex brachystachys E1 . . . . 1 r + 2 + . . . + . + + . r . . 1 . . . + 1 + . . . . . . . . . . . . . . . + Preissia quadrata E0 . 1 . . 1 . . . . . . . . . . . . . . + . . . . + . + + . . + . + . . . + . . + + . . Asplenium viride E1 . + . . . + . . . . . . . . + + + + + . . . . . . . . + . . . 1 . . . . . . . . . . . Valeriana tripteris E1 . . . . . + . . . r . . . . + + . . 1 . . + . . . . . . . . . . . . . . . . . . . . . Fissidens dubius E0 . + . . . + . . . . . . . . . . . . . + . . . . . . . . . . . . . . . . . . . . . . . Barbula crocea E0 . . . . . . . . . . . . . . . . . . . . . . + . 1 . . 1 . . . . . . . . . . . . . . . Viola biflora E1 . . . . . . . + 1 . . . . . . . + . + . . . . . . . . . . . . . . . . . . . . . . . . Eucladium verticillatum E0 + . . . . . . . . . . . . . . . . . . . . . . . . . + + . . . + . . . . . . . . . . . Pellia endiviifolia E0 . . . . . + . . . . . . . . . . . . . . . . . . 1 . . . + . . . . . . . . . . . . . . Soldanella minima E1 . . . . . . . . . . . . . 1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Moehringia muscosa E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . PcSp Physoplexido comosae-Saxifragion petraeae Campanula cespitosa E1 . . . . . . . . . . . . . . . . . . . . . . . . . + r . . . 1 . . . 1 1 + . . + + + . Hieracium pospichalii E1 . . . . . . + . . r . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . PC Potentilletalia caulescentis Potentilla caulescens E1 . . + . . . . . r . . . . . . . . r . + . . . . . . . . . . . r . . . + + r . . r r . Kernera saxatilis E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . + . . . . . . . . . . . . AT Asplenietea trichomanis Asplenium trichomanes E1 + . + . + + . . r r . . . . . . . . . . . . . . + . . . . . . . . . . . . . . . . . . Asplenium ruta-muraria E1 . . . . + . . . . . . . . . + . . + . r . . . . . . + . . . . . . . . . . . . . . . . TR Thlaspietea rotundifolii Hieracium bifidum E1 . + + . . . . . . . . . . + . . . . . + . . r . + r . . . . . . . . . . + . . . . . . Adenostyles glabra E1 . . . . . . . + . . . . r + . . + + + . . . . . . + . . . . . . . . . . . . . . . . . Gymnocarpium robertianum E1 . . . . + . . . . . . . . . . + . . r . . . . . r . . . . . . . . . . . . . . . . . . Achnatherum calamagrostis E1 . . . . . . . . . . . . . . . . . . . r . . . . . . . . . . . . . . . . . . . . . . . Petasites paradoxus E1 . . . . . . . . . . . . . . . . . . . r . . . . . . . . . . . . . . . . . . . . . . . Hieracium porrifolium E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . CD Caricetalia davallianae Parnassia palustris E1 . . . . . . . . r . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + Schoenus nigricans E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + + . . + . . . . MC Montio-Cardaminetea Conocephalum conicum E0 + + 2 1 1 + . . . 1 . . . . + 2 . . . + . . + + 1 . . 1 . . . + 1 1 . . . . . . . + . Campylium stellatum E0 . + . . . . . . . . . . . . . . . . . . . . + . . . . . . . + . . . . . . . . . . . . Bryum pseudotriquetrum E0 . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . + . . . . . . . . . . . . Cratoneuron filicinum E0 + . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . . . . . . . Distichium capillaceum E0 . . . . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . . . . . . . . . . Campylium calcareum E0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Campylium elodes E0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Ditrichum flexicaule E0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . ES Elyno-Seslerietea Sesleria caerulea E1 . . . . . . 1 r . . + + + 1 . . . + + + . . + 1 r . + . + . . . 1 . + 1 1 1 1 + . + 1 Phyteuma orbiculare E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . Carex ferruginea E1 . . . . . . . . + . . . . . . . r . . r . . + . . . . . . . . . . . . . . . . . . . . Carex firma E1 . . . . . . . . . . . . 2 2 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Laserpitium peucedanoides E1 . . . . . . . . . . . . r . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Betonica alopecuros E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . r . . . . . . . . . . . . . . . 19/2 • 2020, 155–231 176 Igor Dakskobler & Andrej Martinčič Plant communities of moist rock crevices with endemic Primula carniolica in the (sub)montane belt of western Slovenia N umber of relevé (Zaporedna štev . popisa) 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 31 32 33 34 35 36 37 38 39 40 41 42 43 FB Festuco-Brometea Buphthalmum salicifolium E1 . . . . . . . . . . . . . . . . . . . . . . . . r . . + . . . . . . . . . . . . . . . Galium lucidum E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . CA Calthion Caltha palustris E1 . . . . . . . . . . . . . . . r . . . . . . . . . . . . . . . . . . . . . . . . . . . Cirsium oleraceum E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . BA Betulo-Alnetea Salix appendiculata E2a . . . . . . . . . . . . + . . r . . . + . . + . r . . . . . . . . . . . . . . . . . . Salix glabra E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . + . . . MuA Mulgedio-Aconitetea Chaerophyllum hirsutum E1 . . . . . . . . . . r . . . r . . . . . . . . . . . . . . . . . . . . . . . . . . . . Crepis paludosa E1 . . . . . . . . . . . . . . . . . . . . . . . . r . . . . . . . . . . . . . . . . . . EA Epilobietea angustifolii Eupatorium cannabinum E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . TG Trifolio-Geranietea Laserpitium siler E1 . . . . . . . . r . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Laserpitium latifolium E1 . . . . . . . . . . . r . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . EP Erico-Pinetea Calamagrostis varia E1 . . . . . . . . . . . + + . + . + + + . + + + . . + . . . . . + . + . . . . . . . + + Erica carnea E1 . . . r . . . . . . . . r . . . . . . . . . r . . . . + . . . . + . . + . . . . . . . Rhododendron hirsutum E1 . . . . . . . . . . . . + . . . + + . . . . . . + + . . . . . . . . . . . . . . . . . Rhodothamnus chamaecistus E1 . . . . . . . . . . . . 1 + . . . . . r . . . . . . . . . . . . . . . . . . . . . . . Carex ornithopoda E1 . . . + . . . . . . . . . . . . . . . r . . . . . . . . . . . . . . . . . . . r . . . Aquilegia nigricans E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . Polygala chamaebuxus E1 . . . . . . . . . . . . r . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Cirsium erisithales E1 . . . . . . . . . . . . . . . . . r . . . . . . . . . . . . . . . . . . . . . . . . . VP Vaccinio-Piceetea Veronica urticifolia E1 + + . . + + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Solidago virgaurea E1 . . . . . . . . . . . . . . . . . . . . + + . . . . . . . . . . . . . . . . . . . . . Aposeris foetida E1 . . . . . . . . . . . . . . . . . . . r . . . . . . . . . . . . . . . . . . . . . . . Homogyne sylvestris E1 . . . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . . . . . . . . . . . Oxalis acetosella E1 . . . . . . . . . . . . . . . . . . r . . . . . . . . . . . . . . . . . . . . . . . . Picea abies E2a . . . . . . . . . . . . . . . . . . . . . . . . . . . . . r . . . . . . . . . . . . . Gentiana asclepiadea E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . AF Aremonio-Fagion Cyclamen purpurascens E1 . . . . . r . . . . . . . . . . . . + . . . . + . . r . . . . . . . . . . . . . . . . Euphorbia carniolica E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . r . . . . + r r . Primula vulgaris E1 . . . . . . . . . . . . . . . . . r . . r . . . . . . . . . . . . . . . . . . . . . . Knautia drymeia E1 . . . r . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . r . Cardamine trifolia E1 . . . . . . . . . . . . . . . . . r . . . . . . . . . . . . . . . . . . . . . . . . . Helleborus niger E1 . . . . . . . . . . . . . . . . . . . r . . . . . . r . . . . . . . . . . . . . . . . Potentilla carniolica E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . . + . . . . . . . . Anemone trifolia E1 . . . . . . . . . . . . . . . r . . . . . . . . . . . . . . . . . . . . . . . . . . . Hemerocallis lilioasphodelus E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . Lamium orvala E1 . . . . . + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . TA Tilio-Acerion Aruncus dioicus E1 r . . . . . . . . . . . . . . . . . . . . . . . r . . . . . . . . . . . . . . . . . . Phyllitis scolopendrium E1 r . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . . Polystichum aculeatum E1 . . . . . . . . . . . . . . . r . . . . . . . . . . . . . . . . . . . . . . . . . . . Acer pseudoplatanus E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . r . . . . . . . . . . . . . 19/2 • 2020, 155–231 177 Igor Dakskobler & Andrej Martinčič Plant communities of moist rock crevices with endemic Primula carniolica in the (sub)montane belt of western Slovenia N umber of relevé (Zaporedna štev . popisa) 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 31 32 33 34 35 36 37 38 39 40 41 42 43 FB Festuco-Brometea Buphthalmum salicifolium E1 . . . . . . . . . . . . . . . . . . . . . . . . r . . + . . . . . . . . . . . . . . . Galium lucidum E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . CA Calthion Caltha palustris E1 . . . . . . . . . . . . . . . r . . . . . . . . . . . . . . . . . . . . . . . . . . . Cirsium oleraceum E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . BA Betulo-Alnetea Salix appendiculata E2a . . . . . . . . . . . . + . . r . . . + . . + . r . . . . . . . . . . . . . . . . . . Salix glabra E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . + . . . MuA Mulgedio-Aconitetea Chaerophyllum hirsutum E1 . . . . . . . . . . r . . . r . . . . . . . . . . . . . . . . . . . . . . . . . . . . Crepis paludosa E1 . . . . . . . . . . . . . . . . . . . . . . . . r . . . . . . . . . . . . . . . . . . EA Epilobietea angustifolii Eupatorium cannabinum E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . TG Trifolio-Geranietea Laserpitium siler E1 . . . . . . . . r . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Laserpitium latifolium E1 . . . . . . . . . . . r . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . EP Erico-Pinetea Calamagrostis varia E1 . . . . . . . . . . . + + . + . + + + . + + + . . + . . . . . + . + . . . . . . . + + Erica carnea E1 . . . r . . . . . . . . r . . . . . . . . . r . . . . + . . . . + . . + . . . . . . . Rhododendron hirsutum E1 . . . . . . . . . . . . + . . . + + . . . . . . + + . . . . . . . . . . . . . . . . . Rhodothamnus chamaecistus E1 . . . . . . . . . . . . 1 + . . . . . r . . . . . . . . . . . . . . . . . . . . . . . Carex ornithopoda E1 . . . + . . . . . . . . . . . . . . . r . . . . . . . . . . . . . . . . . . . r . . . Aquilegia nigricans E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . Polygala chamaebuxus E1 . . . . . . . . . . . . r . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Cirsium erisithales E1 . . . . . . . . . . . . . . . . . r . . . . . . . . . . . . . . . . . . . . . . . . . VP Vaccinio-Piceetea Veronica urticifolia E1 + + . . + + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Solidago virgaurea E1 . . . . . . . . . . . . . . . . . . . . + + . . . . . . . . . . . . . . . . . . . . . Aposeris foetida E1 . . . . . . . . . . . . . . . . . . . r . . . . . . . . . . . . . . . . . . . . . . . Homogyne sylvestris E1 . . . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . . . . . . . . . . . Oxalis acetosella E1 . . . . . . . . . . . . . . . . . . r . . . . . . . . . . . . . . . . . . . . . . . . Picea abies E2a . . . . . . . . . . . . . . . . . . . . . . . . . . . . . r . . . . . . . . . . . . . Gentiana asclepiadea E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . AF Aremonio-Fagion Cyclamen purpurascens E1 . . . . . r . . . . . . . . . . . . + . . . . + . . r . . . . . . . . . . . . . . . . Euphorbia carniolica E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . r . . . . + r r . Primula vulgaris E1 . . . . . . . . . . . . . . . . . r . . r . . . . . . . . . . . . . . . . . . . . . . Knautia drymeia E1 . . . r . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . r . Cardamine trifolia E1 . . . . . . . . . . . . . . . . . r . . . . . . . . . . . . . . . . . . . . . . . . . Helleborus niger E1 . . . . . . . . . . . . . . . . . . . r . . . . . . r . . . . . . . . . . . . . . . . Potentilla carniolica E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . . + . . . . . . . . Anemone trifolia E1 . . . . . . . . . . . . . . . r . . . . . . . . . . . . . . . . . . . . . . . . . . . Hemerocallis lilioasphodelus E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . Lamium orvala E1 . . . . . + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . TA Tilio-Acerion Aruncus dioicus E1 r . . . . . . . . . . . . . . . . . . . . . . . r . . . . . . . . . . . . . . . . . . Phyllitis scolopendrium E1 r . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . . Polystichum aculeatum E1 . . . . . . . . . . . . . . . r . . . . . . . . . . . . . . . . . . . . . . . . . . . Acer pseudoplatanus E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . r . . . . . . . . . . . . . 19/2 • 2020, 155–231 178 Igor Dakskobler & Andrej Martinčič Plant communities of moist rock crevices with endemic Primula carniolica in the (sub)montane belt of western Slovenia N umber of relevé (Zaporedna štev . popisa) 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 31 32 33 34 35 36 37 38 39 40 41 42 43 FS Fagetalia sylvaticae Mycelis muralis E1 . r + . . + . r . . . . . . . . . . + . . . . . . + + + + . . . . + . . . . . . . . . Salvia glutinosa E1 r . . . + . . . . . . . . . . . . . . . . . . . . . + + . . . . . . . . . . . . . . . Carex umbrosa E1 . . . . . . . . . . . . . . + + . . . . . + . . . . . . . . . . . . . . . . . . . . . Galeobdolon flavidum E1 . . . + . . . . . . . . . . . + . + . . . . . . . . . . . . . . . . . . . . . . . . . Galium laevigatum E1 . . . . . r . . . . . . . . . . . . r . . . . . . . . . . . . . . . . . . . . . . . . Brachypodium sylvaticum E1 . . . . . . . . . . . . . . . . . . . . . . . . r . . + . . . . . . . . . . . . . . . Melica nutans E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + Prenanthes purpurea E1 . r . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Asarum europaeum subsp. caucasicum E1 . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . . . . . . . . . . . . . . Cardamine pentaphyllos E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . QP Quercetalia pubescenti-petraeae Carex flacca E1 . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . + . . . . . . . . . . . . Fraxinus ornus E1 . . . r . r . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . . Ostrya carpinifolia E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . QF Querco-Fagetea Carex digitata E1 + + + + + . . . . . . . . . + r . . + r . . r . . . . + . . . . . + . . . . . . . . . Hedera helix E1 . + . . . . . . . . . . . . + . . . . . + . . . . . r . . . . . . + . . . . . . . . . Clematis vitalba E1 . . . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . . . . . . . . . + . Hepatica nobilis E1 . . . . . + . . . . . . . . . . + + . . . . . . . . . . . . . . . . . . . . . . . . . T axus baccata E1 . . . . . . . . . . . . . . . . . . . r . . . . . . . . . . . . . . . . . . . . . . . Frangula alnus E2a . . . . . . . . . . . . . . . . . . . . . . r . . . . . . . . . . . . . . . . . . . . ML Mosses and lichens (Mahovi in lišaji) Neckera crispa E0 . 1 1 + 1 1 . . . . . . . . . . . + 1 . . . . . 1 . + . . . . + 1 . . . . . . . . . . Ctenidium molluscum E0 . . . . . + . . . . . . . . + . 1 + . + . + . . . . . . . . . . . . . . . . . . . . . Tortella tortuosa E0 . . . . . + . . . . . . . . . 1 . . . . + . . . + . . . . . . . . . . . . . . . . . . Heterocladium heteropterum E0 . + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . r . . . . . . . . Marchantia polymorpha E0 . . . . . . . + . . . . . . . . . . . . + . . . . . . . . . . . . . . . . . . . . . . Mnium thomsonii E0 . . . . . + . . . . . . . . . . . . . + . . . . . . . . . . . . . . . . . . . . . . . Lophozia sp. E0 . . . . . + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Brachythecium rutabulum E0 . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . . . . . . . Dichodontium pellucidum E0 . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . . . . . . . Bryum sp. E0 . . . . . . . . . . . . . . . . . . . . . . . . . . 1 . . . . . . . . . . . . . . . . Encalypta vulgaris E0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Mnium marginatum E0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Rhynchostegium murale E0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 19/2 • 2020, 155–231 179 Igor Dakskobler & Andrej Martinčič Plant communities of moist rock crevices with endemic Primula carniolica in the (sub)montane belt of western Slovenia N umber of relevé (Zaporedna štev . popisa) 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 31 32 33 34 35 36 37 38 39 40 41 42 43 FS Fagetalia sylvaticae Mycelis muralis E1 . r + . . + . r . . . . . . . . . . + . . . . . . + + + + . . . . + . . . . . . . . . Salvia glutinosa E1 r . . . + . . . . . . . . . . . . . . . . . . . . . + + . . . . . . . . . . . . . . . Carex umbrosa E1 . . . . . . . . . . . . . . + + . . . . . + . . . . . . . . . . . . . . . . . . . . . Galeobdolon flavidum E1 . . . + . . . . . . . . . . . + . + . . . . . . . . . . . . . . . . . . . . . . . . . Galium laevigatum E1 . . . . . r . . . . . . . . . . . . r . . . . . . . . . . . . . . . . . . . . . . . . Brachypodium sylvaticum E1 . . . . . . . . . . . . . . . . . . . . . . . . r . . + . . . . . . . . . . . . . . . Melica nutans E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + Prenanthes purpurea E1 . r . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Asarum europaeum subsp. caucasicum E1 . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . . . . . . . . . . . . . . Cardamine pentaphyllos E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . QP Quercetalia pubescenti-petraeae Carex flacca E1 . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . + . . . . . . . . . . . . Fraxinus ornus E1 . . . r . r . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . . Ostrya carpinifolia E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . QF Querco-Fagetea Carex digitata E1 + + + + + . . . . . . . . . + r . . + r . . r . . . . + . . . . . + . . . . . . . . . Hedera helix E1 . + . . . . . . . . . . . . + . . . . . + . . . . . r . . . . . . + . . . . . . . . . Clematis vitalba E1 . . . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . . . . . . . . . + . Hepatica nobilis E1 . . . . . + . . . . . . . . . . + + . . . . . . . . . . . . . . . . . . . . . . . . . T axus baccata E1 . . . . . . . . . . . . . . . . . . . r . . . . . . . . . . . . . . . . . . . . . . . Frangula alnus E2a . . . . . . . . . . . . . . . . . . . . . . r . . . . . . . . . . . . . . . . . . . . ML Mosses and lichens (Mahovi in lišaji) Neckera crispa E0 . 1 1 + 1 1 . . . . . . . . . . . + 1 . . . . . 1 . + . . . . + 1 . . . . . . . . . . Ctenidium molluscum E0 . . . . . + . . . . . . . . + . 1 + . + . + . . . . . . . . . . . . . . . . . . . . . Tortella tortuosa E0 . . . . . + . . . . . . . . . 1 . . . . + . . . + . . . . . . . . . . . . . . . . . . Heterocladium heteropterum E0 . + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . r . . . . . . . . Marchantia polymorpha E0 . . . . . . . + . . . . . . . . . . . . + . . . . . . . . . . . . . . . . . . . . . . Mnium thomsonii E0 . . . . . + . . . . . . . . . . . . . + . . . . . . . . . . . . . . . . . . . . . . . Lophozia sp. E0 . . . . . + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Brachythecium rutabulum E0 . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . . . . . . . Dichodontium pellucidum E0 . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . . . . . . . Bryum sp. E0 . . . . . . . . . . . . . . . . . . . . . . . . . . 1 . . . . . . . . . . . . . . . . Encalypta vulgaris E0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Mnium marginatum E0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Rhynchostegium murale E0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 19/2 • 2020, 155–231 180 Igor Dakskobler & Andrej Martinčič Plant communities of moist rock crevices with endemic Primula carniolica in the (sub)montane belt of western Slovenia Number of relevé (Zaporedna štev. popisa) 44 45 46 47 48 49 50 51 52 53 54 55 56 Pr. Fr. Database number of relevé (Delovna številka popisa) 269257 269260 269218 269258 269185 269195 269216 266151 269188 266708 269214 269194 269259 Elevation in m (Nadmorska višina v m) 575 580 410 580 464 520 410 260 460 525 410 520 580 Aspect (Lega) NE SE N NE E SSW S N SE N N SW S Slope in degrees (Nagib v stopinjah) 90 90 80 85 70 80 80 90 90 70 65 95 80 Parent material (Matična podlaga) D D D D D D D D D D D D D Soil (Tla) Li Li Li Li Li Li Li Li Li Li Li Li Li Stoniness in % (Kamnitost v %) 100 100 100 100 70 100 100 100 100 100 100 100 100 Cover of herb layer in % (Zastiranje zeliščne plasti v %): E1 25 30 40 40 60 40 50 40 30 70 40 20 30 Cover of moss layer in % (Zastiranje mahovne plasti v %) E0 65 30 50 40 30 60 80 30 40 30 50 20 30 Number of species (Število vrst) 12 16 17 16 15 23 20 7 7 23 22 17 13 Relevé area (Velikost popisne ploskve) m 2 10 10 10 20 5 5 10 10 5 10 10 5 5 Date of taking relevé (Datum popisa) 5/25/2017 5/25/2017 4/13/2017 5/25/2017 4/7/2017 4/7/2017 4/13/2017 4/9/2017 4/7/2017 4/13/2017 4/13/2017 4/7/2017 5/25/2017 Locality (Nahajališče) Police-Poličnica Police-Poličnica Kazarska grapa-Beli potok Police-Poličnica Orehovska grapa Orehovska grapa Kazarska grapa-Beli potok Gačnik Orehovska grapa Kazarska grapa-Beli potok Kazarska grapa-Beli potok Orehovska grapa Police-Poličnica Quadrant (Kvadrant) 9849/3 9849/3 9849/3 9849/3 9849/4 9849/4 9849/3 9949/1 9849/4 9849/3 9849/3 9849/4 9849/3 Coordinate GK Y (D-48) m 415224 415228 415848 415222 418156 418009 415836 410394 418144 415493 415892 418037 415225 Coordinate GK X (D-48) m 5110523 5110540 5110618 5110537 5110906 5111059 5110650 5104079 5110929 5110727 5110602 5111081 5110585 Diagnostic species of the association (Diagnostične vrste asociacije) Pr. Fr. AP Primula carniolica E1 + 1 1 3 1 2 + 1 2 2 1 2 1 56 100 AP Palustriella commutata E0 4 1 3 3 2 2 4 2 4 . 1 . . 50 89 AP Astrantia carniolica E1 1 1 2 2 1 1 3 . + + 1 . 1 49 88 AP Pinguicula alpina E1 2 2 2 1 2 2 1 3 3 1 1 2 2 48 86 AP Hymenostylium recurvirostre E0 3 1 + 2 + 3 1 . . 1 1 1 1 43 77 PsSp Phyteuma scheuchzeri subsp. columnae E1 . + . . . . + . + . . . . 27 48 EP Molinia arundinacea E1 . + . 1 1 + . . . 1 + . + 24 43 AP Valeriana saxatilis E1 . . + 1 . . . + . + + + + 24 43 ES Carex mucronata E1 . 1 + + . + . . . + 1 1 2 21 38 Diagnostic species of the subassociation (Diagnostične vrste subasociacij) AP Paederota lutea E1 . . . . . . . + . . . . . 22 39 AP Aster bellidiastrum E1 . . . . . . . . . . 1 . . 24 43 Table 1 (Tabela 1): Astrantio carniolicae-Primuletum carniolicae. Relevé number 44–56. 19/2 • 2020, 155–231 181 Igor Dakskobler & Andrej Martinčič Plant communities of moist rock crevices with endemic Primula carniolica in the (sub)montane belt of western Slovenia Number of relevé (Zaporedna štev. popisa) 44 45 46 47 48 49 50 51 52 53 54 55 56 Pr. Fr. AP Cystopteridion s. lat. (Astrantio-Paederotion luteae nom. prov.) Orthothecium rufescens E0 3 2 2 2 1 + 1 1 . 1 1 + 1 47 84 Tofieldia calyculata E1 + + + . + + . . . 1 . . . 30 54 Aster bellidiastrum E1 . . . . . . . . . . 1 . . 24 43 Valeriana saxatilis E1 . . + 1 . . . + . + + + + 24 43 Carex brachystachys E1 . + . 1 . . + . . + . . . 18 32 Preissia quadrata E0 . + + . . . + . . + 1 . . 16 29 Asplenium viride E1 . . r . . . . . . + + . . 12 21 Valeriana tripteris E1 . . . . . . . . . . . . . 6 11 Fissidens dubius E0 . . . . . . . . . . + + . 5 9 Barbula crocea E0 . . . . . + . . . . . 1 . 5 9 Viola biflora E1 . . . . . . . . . . . . . 4 7 Eucladium verticillatum E0 . . . . . . . . . . . . . 4 7 Pellia endiviifolia E0 . . . . . . . . . . . . . 3 5 Soldanella minima E1 . . . . . . . . . . . . . 1 2 Moehringia muscosa E1 . . . . . . . . . . . . . 1 2 PcSp Physoplexido comosae-Saxifragion petraeae Campanula cespitosa E1 + + + + . . + . . . . . 1 15 27 Hieracium pospichalii E1 . . . . . . . . . . . . . 2 4 PC Potentilletalia caulescentis Potentilla caulescens E1 . . . . . . . . . . . + r 12 21 Kernera saxatilis E1 . . . . . . . . . . . . . 2 4 AT Asplenietea trichomanis Asplenium trichomanes E1 . . . . . . . . . . . + . 8 14 Asplenium ruta-muraria E1 . . . . . + . . . . . . . 6 11 TR Thlaspietea rotundifolii Hieracium bifidum E1 r . . + . . . . . . . . . 10 18 Adenostyles glabra E1 . . . . . . . . . . . . . 7 13 Gymnocarpium robertianum E1 . . . . . . + . . . . . . 5 9 Achnatherum calamagrostis E1 . . . . . . . . . . . . . 1 2 Petasites paradoxus E1 . . . . . . . . . . . . . 1 2 Hieracium porrifolium E1 . . . . . . + . . . . . . 1 2 CD Caricetalia davallianae Parnassia palustris E1 + . . . . . . . . . . . . 3 5 Schoenus nigricans E1 . . . . . . . . . . . . . 3 5 MC Montio-Cardaminetea Conocephalum conicum E0 . . . . . . . . . . . . . 18 32 Campylium stellatum E0 . . . . . . . . . . 1 . . 4 7 Bryum pseudotriquetrum E0 . . . . . . . . . . . . . 2 4 Cratoneuron filicinum E0 . . . . . . . . . . . . . 2 4 Distichium capillaceum E0 . . . . . . . . . . . . . 1 2 Campylium calcareum E0 . . . . . . . . . . 1 . . 1 2 Campylium elodes E0 . . . . . . . . . . . + . 1 2 Ditrichum flexicaule E0 . . . . . . . . . . . + . 1 2 ES Elyno-Seslerietea Sesleria caerulea E1 . + + 1 1 1 . + 1 1 + + 1 34 61 Phyteuma orbiculare E1 . . + . 1 + . . . + . . . 5 9 Carex ferruginea E1 . . . . . . . . . . . . . 4 7 Carex firma E1 . . . . . . . . . . . . . 2 4 Laserpitium peucedanoides E1 . . . . . . . . . . . . . 1 2 Betonica alopecuros E1 . . . . . . . . . . . . . 1 2 19/2 • 2020, 155–231 182 Igor Dakskobler & Andrej Martinčič Plant communities of moist rock crevices with endemic Primula carniolica in the (sub)montane belt of western Slovenia Number of relevé (Zaporedna štev. popisa) 44 45 46 47 48 49 50 51 52 53 54 55 56 Pr. Fr. FB Festuco-Brometea Buphthalmum salicifolium E1 . . . . . . . . . . . . . 2 4 Galium lucidum E1 . . . . . . . . . + . . . 1 2 CA Calthion Caltha palustris E1 . . . . . . . . . . . . . 1 2 Cirsium oleraceum E1 . . . . . . . . . . . . . 1 2 BA Betulo-Alnetea Salix appendiculata E2a . . . . . . . . + . + . . 7 13 Salix glabra E1 . . r + . . . . . + . . r 6 11 MuA Mulgedio-Aconitetea Chaerophyllum hirsutum E1 . . . . . . . . . . . . . 2 4 Crepis paludosa E1 . . . . . . . . . . . . . 1 2 EA Epilobietea angustifolii Eupatorium cannabinum E1 r . . + . + . . . . . . . 3 5 TG Trifolio-Geranietea Laserpitium siler E1 . . . r . . . . . . . . . 2 4 Laserpitium latifolium E1 . . . . . . . . . . . . . 1 2 EP Erico-Pinetea Calamagrostis varia E1 + + . . . . + . . . . + . 18 32 Erica carnea E1 . . + . 1 + . . . + + + . 12 21 Rhododendron hirsutum E1 . . . . . . . . . + . . . 6 11 Rhodothamnus chamaecistus E1 . . . . . . . . . + . . . 4 7 Carex ornithopoda E1 . . . . . . . . . . . . . 3 5 Aquilegia nigricans E1 . . . . + . . . . . . . . 2 4 Polygala chamaebuxus E1 . . . . . . . . . . . . . 1 2 Cirsium erisithales E1 . . . . . . . . . . . . . 1 2 VP Vaccinio-Piceetea Veronica urticifolia E1 . . . . . . . . . . . . . 4 7 Solidago virgaurea E1 . . . . . . . . . . . . . 2 4 Aposeris foetida E1 . . . . . . . . . . . . . 1 2 Homogyne sylvestris E1 . . . . . . . . . . . . . 1 2 Oxalis acetosella E1 . . . . . . . . . . . . . 1 2 Picea abies E2a . . . . . . . . . . . . . 1 2 Gentiana asclepiadea E1 . . . . . . + . . . . . . 1 2 AF Aremonio-Fagion Cyclamen purpurascens E1 . . . . + + . . . + . . . 7 13 Euphorbia carniolica E1 . . . . . . . . . . . . . 5 9 Primula vulgaris E1 . . . . . . + . . . . . . 3 5 Knautia drymeia E1 . . . . . . . . . . . . . 2 4 Cardamine trifolia E1 . . . . . . + . . . . . . 2 4 Helleborus niger E1 . . . . . . . . . . . . . 2 4 Potentilla carniolica E1 . . . . . . . . . . . . . 2 4 Anemone trifolia E1 . . . . . . . . . . . . . 1 2 Hemerocallis lilioasphodelus E1 . . . . . . . . . . . . . 1 2 Lamium orvala E1 . . . . . . . . . . . . . 1 2 TA Tilio-Acerion Aruncus dioicus E1 . . . . . . . . . . . . . 2 4 Phyllitis scolopendrium E1 . . . . . . . . . . . . . 2 4 Polystichum aculeatum E1 . . . . . . . . . . . . . 1 2 Acer pseudoplatanus E1 . . . . . . . . . . . . . 1 2 19/2 • 2020, 155–231 183 Igor Dakskobler & Andrej Martinčič Plant communities of moist rock crevices with endemic Primula carniolica in the (sub)montane belt of western Slovenia Number of relevé (Zaporedna štev. popisa) 44 45 46 47 48 49 50 51 52 53 54 55 56 Pr. Fr. FS Fagetalia sylvaticae Mycelis muralis E1 . . . . . . + . . . . . . 11 20 Salvia glutinosa E1 . . . . . . . . . . + . . 5 9 Carex umbrosa E1 . . . . . . . . . . . . . 3 5 Galeobdolon flavidum E1 . . . . . . . . . . . . . 3 5 Galium laevigatum E1 . . . . . . . . . . . . . 2 4 Brachypodium sylvaticum E1 . . . . . . . . . . . . . 2 4 Melica nutans E1 . . . . . . + . . . . . . 2 4 Prenanthes purpurea E1 . . . . . . . . . . . . . 1 2 Asarum europaeum subsp. caucasicum E1 . . . . . . . . . . . . . 1 2 Cardamine pentaphyllos E1 . . . . . . . . . . . . + 1 2 QP Quercetalia pubescenti-petraeae Carex flacca E1 . . + . . . . . . . . . . 3 5 Fraxinus ornus E1 . . . . . . . . . . . . . 3 5 Ostrya carpinifolia E1 . . . . . . . . . + . . . 1 2 QF Querco-Fagetea Carex digitata E1 . + . . + + + . . + + . . 18 32 Hedera helix E1 . . . . . . . . . . . . . 5 9 Clematis vitalba E1 . . . . . . + . . . + . . 4 7 Hepatica nobilis E1 . . . . . . . . . . . . . 3 5 T axus baccata E1 . . . . . . . . . . . . . 1 2 Frangula alnus E2a . . . . . . . . . . . . . 1 2 ML Mosses and lichens (Mahovi in lišaji) Neckera crispa E0 . . . . 1 1 . . . 1 + . . 15 27 Ctenidium molluscum E0 . . . . . + . . . . . + . 8 14 Tortella tortuosa E0 . . . . . + . . . . . 1 . 6 11 Heterocladium heteropterum E0 . . . . . . . . . . . . . 2 4 Marchantia polymorpha E0 . . . . . . . . . . . . . 2 4 Mnium thomsonii E0 . . . . . . . . . . . . . 2 4 Lophozia sp. E0 . . . . . . . . . . . . . 1 2 Brachythecium rutabulum E0 . . . . . . . . . . . . . 1 2 Dichodontium pellucidum E0 . . . . . . . . . . . . . 1 2 Bryum sp. E0 . . . . . . . . . . . . . 1 2 Encalypta vulgaris E0 . . . . . + . . . . . . . 1 2 Mnium marginatum E0 . . . . . + . . . . . . . 1 2 Rhynchostegium murale E0 . . . . . + . . . . . . . 1 2 Legend / Legenda D Dolomite – dolomit T u T ufa – lehnjak Li Lithosol – kamnišče Pr. Presence (number of relevés in which the species is presented) – število popisov, v katerih se pojavlja vrsta Fr. Frequency in % – frekvenca v % 19/2 • 2020, 155–231 184 Igor Dakskobler & Andrej Martinčič Plant communities of moist rock crevices with endemic Primula carniolica in the (sub)montane belt of western Slovenia Table 2 (Tabela 2): Astrantio carniolicae-Pinguiculetum alpinae. Number of relevé (Zaporedna številka popisa) 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 Pr. Fr. Database number of relevé (Delovna številka popisa) 218422 274938 277751 277752 269176 273916 275187 275846 230488 273681 269333 271028 272241 275019 275441 275440 275163 275020 273679 Elevation in m (Nadmorska višina v m) 205 233 240 240 420 590 250 470 350 265 380 338 340 340 240 240 460 330 270 Aspect (Lega) E NNE S SW E SW NW SW NW NW SW SW NW SW NW NW SWW SW SE Slope in degrees (Nagib v stopinjah) 80 90 85 85 85 100 90 95 80 95 80 90 80 90 80 85 90 85 80 Parent material (Matična podlaga) D D L L D D Br D DC D D L DC LCM D D Tu LCM Do Soil (Tla) Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Stoniness in % (Kamnitost v %) 80 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 Cover of herb layer in % (Zastiranje zeliščne plasti v %): E1 30 30 15 20 30 30 50 30 30 30 30 40 30 40 60 60 35 70 40 Co ver of moss layer in % (Zastiranje maho vne plasti v %) E0 30 30 15 20 20 30 20 290 40 20 20 50 40 40 30 40 40 20 40 Number of species (Število vrst) 9 7 10 14 12 10 10 10 33 19 12 12 10 8 10 10 14 11 11 Relevé area (Velikost popisne ploskve) m 2 10 5 10 10 10 20 10 10 10 20 10 10 10 10 10 10 15 10 10 Date of taking relevé (Datum popisa) 3/23/2008 5/20/2018 7/9/2019 7/9/2019 4/7/2017 7/25/2018 4/19/2019 6/17/2019 6/2/2009 9/4/2018 5/18/2017 4/17/2018 8/23/2018 7/1/2018 5/7/2019 5/7/2019 4/22/2019 7/1/2018 9/4/2018 Locality (Nahajališče) Tolminska korita Tolminka-Korita Magozd-Soča-Globoščak Magozd-Soča-Globoščak Orehovska grapa Gačnik-Medved Magozd-Soča-Otona Hotenja- Riževec Humar-Bača Gorenja T rebuša- T rebušica Dabrček Malenšček-Brinta Klonte-Bača Kneža-Velike luti Magozd-Soča-Otona Magozd-Soča-Otona Gačnik-Statarska grapa Kneža-Velike luti Gorenja T rebuša- T rebušica Quadrant (Kvadrant) 9848/1 9748/3 9747/1 9747/1 9849/4 9949/1 9747/1 9949/1 9849/1 9948/2 9849/3 9747/4 9849/1 9849/1 9747/1 9747/1 9949/1 9849/1 9948/4 Coordinate GK Y (D-48) m 403250 403232 390441 390448 418247 411191 390505 413890 415394 409638 414615 396405 414991 410855 390493 390492 411100 410935 409708 Coordinate GK X (D-48) m 5118042 5118066 5126604 5126600 5110810 5102833 5126397 5103353 5113963 5101452 5109345 5120770 5113776 5116774 5126440 5126430 5103416 5116669 5101244 19/2 • 2020, 155–231 185 Igor Dakskobler & Andrej Martinčič Plant communities of moist rock crevices with endemic Primula carniolica in the (sub)montane belt of western Slovenia Number of relevé (Zaporedna številka popisa) 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 Pr. Fr. Diagnostic species of the association (Diagnostične vrste asociacije) Pr. Fr. AP Pinguicula alpina E1 2 2 2 2 3 3 3 2 2 3 2 3 + 3 3 + 3 3 3 19 100 AP Hymenostylium recurvirostre E0 1 . 1 1 2 2 1 1 2 3 + 3 1 2 1 3 2 . . 16 84 AP Palustriella commutata E0 2 2 . 1 . . + . 2 . 1 3 2 3 2 3 3 + 3 14 74 AP Astrantia carniolica E1 . 1 . . . + 1 . . + 1 . 2 2 2 3 2 3 . 11 58 TR Petasites paradoxus E1 . . + . . . . . r + . + . + . . . + + 7 37 Differential species of the variant (Razlikovalnica variante) EP Molinia arundinacea E1 . . . . . . . . r + + + 1 2 2 3 1 2 3 11 58 AT Cystopteridion s. lat. (Astrantio-Paederotion luteae nom. prov.) Tofieldia calyculata E1 2 2 + + 1 . . . 1 1 . . + + . . + . + 11 58 Aster bellidiastrum E1 + + 1 1 . + . . 2 1 . . + . . . 2 1 1 11 58 Orthothecium rufescens E0 . . . . 2 + + + 1 1 + . . . . . 1 . 1 9 47 Paederota lutea E1 . . . . . + . 1 . + 2 . . . . . + . . 5 26 Saxifraga aizoides E1 . . . . . . . . 2 . . . + . + + . . . 4 21 Carex brachystachys E1 . . . + . + . . + . . . . . . . . . . 3 16 Pellia endiviifolia E0 . . . . . . + . . . . . + . . . . . + 3 16 Barbula crocea E0 . . . . . . . . 2 1 . . . . . . . 1 . 3 16 Preissia quadrata E0 . . . . . . . . + 2 + . . . . . . . . 3 16 Valeriana saxatilis E1 . . . . . + . . . . . . . . . . . . . 1 5 Selaginella helvetica E1 . . . . . . . . + . . . . . . . . . . 1 5 Eucladium verticillatum E0 . . . . . . . . . . . . . . + . . . . 1 5 Fissidens dubius E0 . . . . . . . . . . . . . . . + . . . 1 5 PCSp Physoplexido comosae-Saxifragion petraeae Campanula cespitosa E1 + . + + + . + + + . . + . . . . . . + 9 47 Phyteuma scheuchzeri subsp. columnae E1 . . . . 1 . . . . + . . . . . . . . . 2 11 Micromeria thymifolia E1 . . . . . . . . . . . . . . . . . + . 1 5 PcSp Potentilletalia caulescentis Potentilla caulescens E1 . . . + r . . . . 2 . . . . . . . . . 3 16 AT Asplenietea trichomanis Asplenium trichomanes E1 . . . . + . . + . . . . . . . . . . . 2 11 TR Thlaspietea rotundifolii Achnatherum calamagrostis E1 . . . . r . . r . . . . . . . . . . . 2 11 19/2 • 2020, 155–231 186 Igor Dakskobler & Andrej Martinčič Plant communities of moist rock crevices with endemic Primula carniolica in the (sub)montane belt of western Slovenia Number of relevé (Zaporedna številka popisa) 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 Pr. Fr. Adenostyles glabra E1 . . . . . r . . r . . . . . . . . . . 2 11 Hieracium bifidum E1 . . . . . . . . r . . + . . . . . . . 2 11 Hieracium porrifolium E1 . . + . . . . . . . . . . . . . . . . 1 5 CD Caricetalia davallianae Carex lepidocarpa E1 . . . . . . . . . + . . 1 . + 1 . . . 4 21 MC Montio-Cardaminetea Conocephalum conicum E0 . 1 . . . . . . 1 . . . . . . . . . . 2 11 Cratoneuron filicinum E0 . . . 1 . . . . + . . . . . . . . . . 2 11 Bryum pseudotriquetrum E0 . . . . . . + . + . . . . . . . . . . 2 11 ES Elyno-Seslerietea Sesleria caerulea E1 1 1 + . + + . . + . 1 1 . 1 1 + 1 + . 13 68 Carex mucronata E1 1 . . . . . . . . . + . . . . . . . . 2 11 MA Molinio-Arrhenatheretea Angelica sylvestris E1 . . . . . . . . + . . . . . . . . . . 1 5 Pimpinella major E1 . . . . . . . . r . . . . . . . . . . 1 5 Crepis paludosa E1 . . . . . . . . . . . . . . . . + . . 1 5 BA Betulo-Alnetea Salix appendiculata E1 . . . . . . + . r + . + . . . . . . . 4 21 NuA Mulgedio-Aconitetea Petasites hybridus E1 . . . . . . . . r . . . . . . . . . . 1 5 Chaerophyllum hirsutum E1 . . . . . . . . . . . . . . . . r . . 1 5 EA Epilobietea angustifolii Eupatorium cannabinum E1 . . . + . . . . . . . . . . . . . . . 1 5 EP Erico-Pinetea Calamagrostis varia E1 . . + + . . + . 2 + . 1 . . . . + . . 7 37 Erica carnea E1 . . . . . . . . r . + + . . . . . . . 3 16 Leontodon incanus E1 r . . . . . . . . + . . . . . . . . . 2 11 Carex ornithopoda E1 . . . + . . . . . . . . . . . . r . . 2 11 Gymnadenia odoratissima E1 . . . . . . . . . . . . . . . . . r . 1 5 VP Vaccinio-Piceetea Picea abies E1 . . . . . . . r . . . . . . . . . . . 1 5 Veronica urticifolia E1 . . . . . . . . 1 . . . . . . . . . . 1 5 FS Fagetalia sylvaticae Galium laevigatum E1 . . . + . . . . r . . . . . . . . . . 2 11 19/2 • 2020, 155–231 187 Igor Dakskobler & Andrej Martinčič Plant communities of moist rock crevices with endemic Primula carniolica in the (sub)montane belt of western Slovenia Legend / Legenda Br Breccia – breča C Chert – roženec D Dolomite – dolomit L Limestone – apnenec M Marlstone – laporovec Li Lithosol – kamnišče Pr. Presence (number of relevés in which the species is presented) – število popisov, v katerih se pojavlja vrsta Fr. Frequency in % – frekvenca v % Number of relevé (Zaporedna številka popisa) 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 Pr. Fr. Salvia glutinosa E1 . . . . + . . . . . . . . . . . . . . 1 5 Fagus sylvatica E1 . . . . . . . r . . . . . . . . . . . 1 5 Laburnum alpinum E2a . . . . . . . . r . . . . . . . . . . 1 5 QP Quercetalia pubescenti-petraeae Carex flacca E1 . . . . . . . . . . . 1 . . + 1 . 1 . 4 21 Ostrya carpinifolia E1 . . . . + . . r . . . . . . . . . . . 2 11 QF Querco-Fagetea Potentilla erecta E1 . . . . . . . . . . . . . . . . . . + 1 5 SP Salicetea purpureae Salix eleagnos E1 . . + + . . . . . + . . . . . . . . + 4 21 ML Mosses and lichens (Mahovi in lišaji) Brachythecium rutabulum E0 . . . . . . . . + . . . . . . . . . . 1 5 Ctenidium molluscum E0 . . . . . . . . 1 . . . . . . . . . . 1 5 Leiocolea collaris E0 . . . . . . . . + . . . . . . . . . . 1 5 Reboulia hemisphaerica E0 . . . . . . . . + . . . . . . . . . . 1 5 Tortella tortuosa E0 . . . . . . . . . + . . . . . . . . . 1 5 Neckera crispa E0 . . . . . . . . . . + . . . . . . . . 1 5 19/2 • 2020, 155–231 188 Igor Dakskobler & Andrej Martinčič Plant communities of moist rock crevices with endemic Primula carniolica in the (sub)montane belt of western Slovenia Number of relevé (Zaporedna štev. popisa) 1 2 3 4 5 6 7 8 9 10 11 12 13 14 Pr. Fr. Database number of relevé (Delovna številka popisa) 271077 271078 271075 272101 272102 272105 272126 272127 272106 256295 269090 257491 271257 273342 Elevation in m (Nadmorska višina v m) 400 400 392 414 265 260 255 255 260 327 990 755 420 460 Aspect (Lega) SSE SE SW SW SE SE SE SSE SE SE NW NEE NE E Slope in degrees (Nagib v stopinjah) 70 70 85 80 85 90 85 85 80 100 80 80 85 80 Parent material (Matična podlaga) LMC LMC LMC LMC D DC DC DC DC A AC A LMC LMC Soil (Tla) Li Li Li Li Li Li Li Li Li Li Li Li Li Li Stoniness in % (Kamnitost v %) 80 80 80 80 80 90 80 80 70 100 80 80 70 70 Cover of herb layer in % (Zastiranje zeliščne plasti v %): E1 40 40 30 40 30 20 40 40 50 15 30 30 50 50 Cover of moss layer in % (Zastiranje mahovne plasti v %) E0 40 10 40 40 30 25 39 30 20 10 20 20 50 30 Number of species (Število vrst) 15 10 12 25 18 18 15 13 20 11 11 18 25 26 Relevé area (Velikost popisne ploskve) m 2 10 10 10 10 10 10 10 10 10 5 10 10 10 10 Date of taking relevé (Datum popisa) 4/26/2018 4/26/2018 4/26/2018 6/2/2018 6/2/2018 6/2/2018 8/15/2018 8/15/2018 6/2/2018 4/19/2015 5/17/2017 5/12/2015 5/20/2018 8/10/2018 Locality (Nahajališče) Volarja-Gregorčičev slap Volarja-Gregorčičev slap Volarja-Gregorčičev slap Godiča-Beri Godiča Godiča Godiča Godiča Godiča Zadlaščica-Perbla Kacenpoh-Kobla Linderpoh-Šprickovbl Tolminka Valley - Zala Prodarjeva grapa Quadrant (Kvadrant) 9747/4 9747/4 9747/4 9848/2 9848/2 9848/2 9848/2 9848/2 9848/2 9748/4 9749/4 9749/4 9748/3 9849/2 Coordinate GK Y (D-48) m 396956 396957 397001 405021 404708 404660 404631 404636 404636 404710 420118 419818 402551 418767 Coordinate GK X (D-48) m 5120880 5120873 5120880 5116236 5116076 5116070 5116045 5116052 5116052 5119380 5121314 5120042 5119318 5117456 Diagnostic species of the association (Diagnostične vrste asociacije) Pr. Fr. AP Saxifraga aizoides E1 2 2 1 + 3 1 + 1 1 + + 1 3 + 14 100 AP Palustriella commutata E0 2 1 3 2 . 1 1 . . . 1 1 1 2 10 71 PcSp Campanula cespitosa E1 . . + + + + 1 1 1 + 1 . + . 10 71 EP Molinia arundinacea E1 1 . 1 2 + 1 3 2 1 . . + . 1 10 71 EP Calamagrostis varia E1 1 . . + . + . . + + + + 1 + 9 64 AP Preissia quadrata E0 + . . . + + + + + . . + + + 9 64 AP Cystopteridion s. lat. (Astrantio-Paederotion luteae nom. prov.) Hymenostylium recurvirostre E0 . + . . 2 2 3 2 + . . + . . 7 50 Barbula crocea E0 . . . . + + . . + . . . 1 + 5 36 Pinguicula alpina E1 2 1 . . . . . . . . r r . . 4 29 Carex brachystachys E1 + . . . . + . . . . . . + 1 4 29 Tofieldia calyculata E1 . . . . . 1 1 1 2 . . . . . 4 29 Table 3 (Tabela 3): Campanulo cespitosae-Saxifragetum aizoidis. 19/2 • 2020, 155–231 189 Igor Dakskobler & Andrej Martinčič Plant communities of moist rock crevices with endemic Primula carniolica in the (sub)montane belt of western Slovenia Number of relevé (Zaporedna štev. popisa) 1 2 3 4 5 6 7 8 9 10 11 12 13 14 Pr. Fr. Orthothecium rufescens E0 . . . . . . . . . 2 1 . 1 1 4 29 Aster bellidiastrum E1 . . . . . . . . . 1 1 + . 1 4 29 Pellia endiviifolia E0 + . + 2 . . . . . . . . + 1 5 36 Jungermannia atrovirens E0 . . . . . . . . + . . . + + 3 21 Valeriana tripteris E1 . . . . . . . . . . . . + 1 2 14 Fissidens dubius E0 . . . . . . . . . . . . + + 2 14 Paederota lutea E1 . . . . . . . . . 1 . . . . 1 7 Valeriana saxatilis E1 . . . . . . . . . + . . . . 1 7 Asplenium viride E1 . . . . . . . . . . . . 1 . 1 7 Viola biflora E1 . . . . . . . . . . . . . 1 1 7 PcSp Physoplexido comosae-Saxifragion petraeae Campanula carnica E1 . . . + . . . . . + . . . + 3 21 Hieracium pospichalii E1 . . . . . . . . . . . . + . 1 7 PC Potentilletalia caulescentis Potentilla caulescens E1 . . . . + + . . + . . + . . 4 29 Primula auricula E1 . . . . . . . . . . . 1 . . 1 7 Rhamnus pumilus E1 . . . . . . . . . . . + . . 1 7 AT Asplenietea trichomanis Asplenium ruta-muraria E1 . . . . . . . . . + . . + . 2 14 TR Thlaspietea rotundifolii Hieracium porrifolium E1 . . . + + + + 1 + . . . . . 6 43 Petasites paradoxus E1 + . + . + . + . . . . . . . 4 29 Hieracium bifidum E1 . . . + . + + . . . . . 1 . 4 29 Adenostyles glabra E1 . . . . . . . . . . + . r 1 3 21 Leontodon hispidus subsp. hyoseroides E1 + + . . . . . . . . . . . . 2 14 Trisetum argenteum E1 . . 1 . . . . . . . . + . . 2 14 Achnatherum calamagrostis E1 . . . . + . . . . . . . . . 1 7 CD Caricetalia davallianae Carex flava agg. E1 . 1 . . . . . . . . . . . . 1 7 Epilobium alsinifolium E1 . . + . . . . . . . . . . . 1 7 Pinguicula vulgaris E1 . . . . . . . . . . . . . 1 1 7 MC Montio-Cardaminetea Gymnostomum aeruginosum E0 2 . 2 1 . . . . . . . . . . 3 21 Campylium stellatum E0 + . . . . . . . . . . . + + 3 21 Conocephalum conicum E0 . . . . . . . . . . . 1 . . 1 7 Bryum pseudotriquetrum E0 . . . . . . . . . . . + . . 1 7 Oxyrrhynchium schleicheri E0 . . . . . . . . . . . + . . 1 7 Cololejeunea calcarea E0 . . . . . . . . . . . . + . 1 7 Barbilophozia sp. E0 . . . . . . . . . . . . . + 1 7 ES Elyno-Seslerietea Sesleria caerulea E1 . . . + 1 . 1 + + + . 2 1 2 9 64 Carex mucronata E1 . . . . . r + . . . + . . . 3 21 Carex ornithopodoides E1 . . . . . . . . r . . . . . 1 7 Carex ferruginea E1 . . . . . . . . . . 1 . . . 1 7 FB Festuco-Brometea Buphthalmum salicifolium E1 . . . . + + . 1 + . . . . . 4 29 MA Molinio-Arrhenatheretea Caltha palustris E1 . . 2 + . . . . . . . . . . 2 14 Angelica sylvestris E1 . . . . . . . . . . . . . + 1 7 BA Betulo-Alnetea Salix appendiculata E1 . . . . + . + + + . . . . + 5 36 19/2 • 2020, 155–231 190 Igor Dakskobler & Andrej Martinčič Plant communities of moist rock crevices with endemic Primula carniolica in the (sub)montane belt of western Slovenia Number of relevé (Zaporedna štev. popisa) 1 2 3 4 5 6 7 8 9 10 11 12 13 14 Pr. Fr. MuA Mulgedio-Aconitetea Chaerophyllum hirsutum E1 . . . + . . . . . . . . . . 1 7 Petasites hybridus E1 . . r . . . . . . . . . . . 1 7 EA Epilobietea angustifolii Eupatorium cannabinum E1 . . . 1 + + . . . . . . . . 3 21 Tussilago farfara E1 + . . . . . . . . . . . . . 1 7 EP Erico-Pinetea Erica carnea E1 . . . . + + 1 + + . . . . . 5 36 Leontodon incanus E1 . . . + + . . . + . . . . . 3 21 Peucedanum austriacum E1 . . . . . + . . + . . . . . 2 14 Carex ornithopoda E1 . . . . . . . . + . . . . . 1 7 VP Vaccinio-Piceetea Veronica urticifolia E1 . . . + . . . . . + . + + 1 5 36 Saxifraga cuneifolia E1 . . . . . . . . . . . . + + 2 14 AF Aremonio-Fagion Lamium orvala E1 . . . + . . . . . . . . . . 1 7 Primula vulgaris E1 . . . + . . . . . . . . . . 1 7 TA Tilio-Acerion Acer pseudoplatanus E1 . + . . . . . . . . . . . . 1 7 Tilia platyphyllos E2a . r . . . . . . . . . . . . 1 7 Aruncus dioicus E1 . . . . r . . . . . . . . . 1 7 FS Fagetalia sylvaticae Petasites albus E1 . . . + . . . . . . . . + . 2 14 Galium laevigatum E1 . . . + . . . . . . . . . + 2 14 Brachypodium sylvaticum E1 . . . + . . . . . . . . . . 1 7 Salvia glutinosa E1 . . . . . . . . . . . . . + 1 7 QP Quercetalia pubescenti-petraeae Carex flacca E1 2 2 + . . . . . . . . . . . 3 21 Ostrya carpinifolia E2a . . . + . . . + . . . . + . 3 21 Fraxinus ornus E2a r . . . . . . . . . . . . . 1 7 QF Querco-Fagetea Hedera helix E1 . . . + . . . . . . . . . . 1 7 Carex digitata E1 . . . . . . . . . . + . . . 1 7 SP Salicetea purpureae Salix purpurea E2a . r . . . . . . . . . . . . 1 7 ML Mosses and lichens (Mahovi in lišaji) Tortella tortuosa E0 . . . . + . 1 + + . . . . . 4 29 Brachythecium rutabulum E0 . . . + . . . . . . . . . . 1 7 Rhynchostegium riparioides E0 . . . + . . . . . . . . . . 1 7 Ctenidium molluscum E0 . . . . . . . . . . . + . . 1 7 Encalypta streptocarpa E0 . . . . . . . . . . . . 1 . 1 7 Legend / Legenda C Chert – roženec D Dolomite – dolomit L Limestone – apnenec M Marlstone – laporovec Li Lithosol – kamnišče Pr. Presence (number of relevés in which the species is presented) – število popisov, v katerih se pojavlja vrsta Fr. Frequency in % – frekvenca v % 19/2 • 2020, 155–231 191 Igor Dakskobler & Andrej Martinčič Plant communities of moist rock crevices with endemic Primula carniolica in the (sub)montane belt of western Slovenia Number of relevé (Zaporedna številka popisa) 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 Pr. Fr. Database number of relevé (Delovna štev. popisa) 258588 260931 260936 260938 260929 261055 260932 268210 260915 260940 260930 275149 260916 260917 261056 Elevation in m (Nadmorska višina v m) 460 560 550 550 570 600 560 850 550 550 560 430 570 580 600 Aspect (Lega) N W W NWW SE NW NW NE NE NW NW N NW N NE Slope in degrees (Nagib v stopinjah) 90 95 85 90 90 70 95 90 80 80 80 80 80 90 95 Parent material (Matična podlaga) D D D D D D D D D D D D D D D Soil (Tla) Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Stoniness in % (Kamnitost v %) 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 Cover of herb layer in % (Zastiranje zeliščne plasti v %) E1 20 15 30 20 20 30 20 20 40 60 20 20 30 50 30 Cover of moss layer in % (Zastiranje mahovne plasti v %) E0 10 20 20 10 30 30 20 20 60 80 20 40 20 30 10 Number of species (Število vrst) 18 15 28 28 23 26 14 14 17 26 20 17 22 22 7 Relevé area (Velikost popisne ploskve) m 2 10 4 4 10 5 5 5 5 5 10 10 15 10 5 5 Date of taking relevé (Datum popisa) 9/29/2015 4/15/2016 4/15/2016 4/15/2016 4/15/2016 4/20/2016 4/15/2016 8/30/2017 4/15/2016 4/15/2016 4/15/2016 4/22/2019 4/15/2016 4/15/2016 4/20/2016 Locality (Nahajališče) Govškarca - Kanomlja Žirovnica - Raskovec Žirovnica - Raskovec Žirovnica - Raskovec Žirovnica - Raskovec Žirovnica - Raskovec Žirovnica - Raskovec Gačnik - Brdo Žirovnica - Raskovec Žirovnica - Raskovec Žirovnica - Raskovec Gačnik Žirovnica - Raskovec Žirovnica - Raskovec Žirovnica - Raskovec Quadrant (Kvadrant) 9949/4 9950/4 9950/4 9950/4 9950/4 9950/4 9950/4 9949/3 9950/4 9950/4 9950/4 9949/1 9950/4 9950/4 9950/4 Coordinate GK Y (D-48) m 422149 430440 430488 430430 430388 430318 430464 412447 430396 430418 430422 410984 430391 430373 430310 Coordinate GK X (D-48) m 5099632 5096271 5096326 5096266 5096235 5096162 5096296 5100926 5096265 5096248 5096252 5103382 5096208 5096190 5096174 Diagnostic species of the association (Diagnostične vrste asociacije) Pr. Fr. AP Primula carniolica E1 2 1 2 2 2 1 2 2 1 1 2 1 2 2 1 15 100 AP Orthothecium rufescens E0 + 1 + 1 2 1 1 + 3 2 2 2 1 2 1 15 100 AP Valeriana tripteris E1 + + + . r + . . + . + . + . . 8 53 Differential species of the subassociation (Razlikovalnice asociacije) AP Viola biflora E1 + + + + 1 + + 1 2 3 1 + + + 2 15 100 VP Veronica urticifolia E1 + + + + + 1 + + + + + . . . . 11 73 AP Pinguicula alpina E1 . + . + + . . . + . + + 1 1 + 9 60 Cystopteridion s. lat. (Astrantio-Paederotion luteae nom. prov.) Aster bellidiastrum E1 1 + + 1 1 1 + . 1 + 1 1 1 1 + 14 93 Asplenium viride E1 + 1 + + 1 1 . . + 1 . . + . . 9 60 Cystopteris fragilis E1 . . + + + . 1 + . 1 1 . + . . 8 53 Carex brachystachys E1 + . + 1 . . . + . . 1 . + + . 7 47 Paederota lutea E1 1 . . . . . . + . . . 1 . . . 3 20 Table 4 (Tabela 4): Primuletum carniolicae Accetto 2008 violetosum biflorae. 19/2 • 2020, 155–231 192 Igor Dakskobler & Andrej Martinčič Plant communities of moist rock crevices with endemic Primula carniolica in the (sub)montane belt of western Slovenia Number of relevé (Zaporedna številka popisa) 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 Pr. Fr. Hymenostylium recurvirostre E0 . . + . . + . . . . . . . + . 3 20 Tofieldia calyculata E1 . . . + . . . . . . . 1 . + . 3 20 Palustriella commutata E0 + . . . . . . . . . . . . . . 1 7 Moehringia muscosa E1 . . . r . . . . . . . . . . . 1 7 Astrantia carniolica E1 . . . . . . . + . . . . . . . 1 7 Fissidens dubius E0 . . . . . . . 1 . . . . . . . 1 7 Valeriana saxatilis E1 . . . . . . . . . . . . + . . 1 7 PcSpPhysoplexido comosae-Saxifragion petraeae Campanula cespitosa E1 . . . . . . . . . . . . + 1 + 3 20 Phyteuma scheuchzeri subsp. columnae E1 . . . . . . . + . . . . . . . 1 7 AT Asplenietea trichomanis Asplenium ruta-muraria E1 + r + + + + + + . + . . r + . 11 73 Asplenium trichomanes E1 + + + 1 + . 1 1 . + + . . . . 9 60 TR Thlapietea rotundifolii Hieracium bifidum E1 r . . + . + + . . . + + . . . 6 40 Gymnocarpium robertianum E1 . . . r . . . . . . . . . . . 1 7 MC Montio-Cardaminetea Conocephalum conicum E0 1 1 + + + 1 . + 2 2 1 2 . + . 12 80 Oxyrrhynchium hians E0 + . + . . + . . . . . . . . . 3 20 Campylium stellatum E0 . . . . + + . . . . . . . . . 2 13 Oxyrrhynchium schleicheri E0 . . . . + . . . . . . . . + . 2 13 Cirriphyllum cirrhosum E0 . . . . + . . . . . . . . . . 1 7 Distichium capillaceum E0 . . . . . . . . . . . . . + . 1 7 ES Elyno-Seslerietea Sesleria caerulea E1 . . . . . + . . . . . . 2 1 . 3 20 Carex mucronata E1 . . r . . . . . . . . . . . . 1 7 FB Festuco-Brometea Buphthalmum salicifolium E1 . . . . . . . . . . . . r + . 2 13 BA Betulo-Alnetea Salix appendiculata E1 . . . . . r . . . . . . + . . 2 13 MuA Mulgedio-Aconitetea Petasites hybridus E1 . . . . . r . . . . . . . . . 1 7 EP Erico-Pinetea Calamagrostis varia E1 + + + + . . . . + + + + + 2 . 10 67 Rhododendron hirsutum E1 . . . . . . . . . . . . + 1 . 2 13 Rubus saxatilis E1 . . r . . . . . . . . . . . . 1 7 Molinia arundinacea E1 . . . . . . . . . r . . . . . 1 7 Erica carnea E1 . . . . . . . . . . . . . + . 1 7 VP Vaccinio-Piceetea Veronica urticifolia E1 + + + + + 1 + + + + + . . . . 11 73 Homogyne sylvestris E1 . . . + . r . . . . . . . . . 2 13 Oxalis acetosella E1 . . . . . . . . . 1 . + . . . 2 13 Clematis alpina E1 . . . r . . . . . . . . . . . 1 7 Rosa pendulina E1 . . . r . . . . . . . . . . . 1 7 Aposeris foetida E1 . . . . . . . . + . . . . . . 1 7 AF Aremonio-Fagion Scopolia carniolica E1 . . r . . . . . . . . . r . . 2 13 Cyclamen purpurascens E1 . . . r . . . . . . . + . . . 2 13 Helleborus niger E1 . . . . r . . . . . . . . . . 1 7 19/2 • 2020, 155–231 193 Igor Dakskobler & Andrej Martinčič Plant communities of moist rock crevices with endemic Primula carniolica in the (sub)montane belt of western Slovenia Number of relevé (Zaporedna številka popisa) 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 Pr. Fr. Cardamine enneaphyllos E1 . . . . . . . . . r . . . . . 1 7 Euphorbia carniolica E1 . . . . . . . . . r . . . . . 1 7 Cardamine trifolia E1 . . . . . . . . . r . . . . . 1 7 TA Tilio-Acerion Aruncus dioicus E1 . r . . . + . . + + + . . . . 5 33 Thalictrum aquilegiifolium E1 . . . . . . . . . + r + r . . 4 27 Polystichum aculeatum E1 . . + . . . . . . . r . . . . 2 13 Ulmus glabra E2a . . . . . . . . + r . . . . . 2 13 Acer pseudoplatanus E1 . . r . . . . . . . . . . . . 1 7 Phyllitis scolopendrium E1 . . r . . . . . . . . . . . . 1 7 FS Fagetalia sylvaticae Galeobdolon flavidum E1 r . + + r + . . . . + . + . . 7 47 Cardamine pentaphyllos E1 . r . . . . . . . r r . . r . 4 27 Campanula trachelium E1 . + r . . . . . . + . . . . . 3 20 Lathyrus vernus E1 . . r . . . . . + . . . . . . 2 13 Galium laevigatum E1 . . . . . + r . . . . . . . . 2 13 Salvia glutinosa E1 r . . . . . . . . . . . . . . 1 7 Daphne mezereum E1 . . . r . . . . . . . . . . . 1 7 Mercurialis perennis E1 . . . r . . . . . . . . . . . 1 7 Mycelis muralis E1 . . . . . + . . . . . . . . . 1 7 Fagus sylvatica E1 . . . . . . . . . r . . . . . 1 7 Prenanthes purpurea E1 . . . . . . . . . r . . . . . 1 7 QF Querco-Fagetea Carex digitata E1 . . . r . + + . + r r + + + . 9 60 Hepatica nobilis E1 . . r r . . . . + + . + . . . 5 33 T axus baccata E2b . . . . . . . . . . . . . . + 1 7 ML Mosses and lichens (Mahovi in lišaji) Neckera crispa E0 . . + + 2 1 + . . . . + . . . 6 40 Ctenidium molluscum E0 . . . . . . . + . + 2 1 1 . . 5 33 Mnium thomsonii E0 . . + + + + . . . . . . . . . 4 27 Tortella tortuosa E0 . . + . . . + . 1 . . . . . . 3 20 Pedinophyllum interruptum E0 . . . . 1 + . . . . . . . 1 . 3 20 Mnium marginatum E0 . . . . + + . . . . . . . + . 3 20 Brachythecium starkei E0 . . . . + . . . . . . . . . . 1 7 Plagiomnium rostratum E0 . . . . + . . . . . . . . . . 1 7 Marchantia polymorpha E0 . . . . . . + . . . . . . . . 1 7 Legend / Legenda D Dolomite – dolomit Li Lithosol – kamnišče Pr. Presence (number of relevés in which the species is presented) – število popisov, v katerih se pojavlja vrsta Fr. Frequency in % – frekvenca v % 19/2 • 2020, 155–231 194 Igor Dakskobler & Andrej Martinčič Plant communities of moist rock crevices with endemic Primula carniolica in the (sub)montane belt of western Slovenia Table 5 (Tabela 5): Phyteumato columnae-Primuletum carniolicae. Relevé number 1–37. Number of relevé (Zaporedna številka popisa) 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 31 32 33 34 35 36 37 Database number of relevé (Delovna številka popisa) 259019 259020 273980 217874 258592 275224 269300 278013 278010 226622 266154 269299 269303 269301 273965 230232 246867 274036 230234 268195 241497 259048 255777 258581 273979 258597 275218 269305 274329 269366 269369 230610 258593 275599 241495 275601 246308 Elevation in m (Nadmorska višina v m) 975 975 745 260 485 230 475 700 660 215 250 475 570 475 780 350 1250 895 325 890 800 1460 450 390 745 500 650 470 480 460 420 830 495 775 780 760 440 Aspect (Lega) NE NE NE SW S E NW N N NW N NE SEE E NW NE N SW N E NE NW E N SW NE NNE NE N SSW E NWW SW NW NE NE NW Slope in degrees (Nagib v stopinjah) 80 90 90 90 80 95 70 90 80 90 90 90 90 80 85 90 80 90 90 85 80 80 80 90 90 90 80 80 85 70 80 95 90 85 80 85 90 Parent material (Matična podlaga) D D D D D D D D D D D D D D D D D D D D D L D D D D D D D D D D D D D D D Soil (Tla) Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Stoniness in % (Kamnitost v %) 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 Cover of herb layer in % (Zastiranje zeliščne plasti v %) E1 30 20 30 30 40 40 35 20 30 20 25 30 25 30 15 30 35 20 20 30 30 30 30 20 30 30 30 40 30 30 20 30 30 20 20 25 20 Cover of moss layer in % (Zastiranje mahovne plasti v %) E0 30 20 30 10 20 10 50 10 5 10 20 20 10 30 20 10 20 30 30 20 10 20 20 10 10 20 30 40 20 20 20 5 10 10 5 10 20 Number of species (Število vrst) 24 24 16 17 13 10 20 27 20 11 10 14 13 17 15 29 32 14 18 15 24 15 25 12 10 18 15 26 26 15 15 17 11 14 16 15 20 Relevé area (Velikost popisne ploskve) m 2 10 10 10 10 3 10 10 15 10 5 10 10 10 10 10 10 10 10 10 5 10 4 5 4 10 4 10 10 10 5 5 10 2 10 10 10 10 Date of taking relevé (Datum popisa) 6/25/2015 6/25/2015 7/2/2018 4/19/2007 9/29/2015 4/18/2019 6/20/2017 4/29/2019 4/29/2019 5/8/2009 4/9/2017 6/20/2017 6/20/2017 6/20/2017 7/2/2018 4/14/2009 8/13/2003 7/16/2018 4/14/2009 8/30/2017 4/12/2011 6/22/2015 5/14/2002 9/29/2015 7/2/2018 9/29/2015 4/18/2019 6/20/2017 5/8/2018 3/31/2017 3/31/2017 5/13/2005 9/29/2015 5/16/2019 4/12/2011 5/16/2019 6/17/2011 Locality (Nahajališče) Oblakov vrh-Maslinca Oblakov vrh-Maslinca Gačnik Pršjak Govškarca-Kanomlja Gorenja trebuša - Ščura Jagršče-Sjavnica Gorenja T rebuša-Drnulk Gorenja T rebuša-Drnulk Stopnik-Idrijca Gačnik Jagršče-Sjavnica Jagršče-Sjavnica Jagršče-Sjavnica Gačnik Divje jezero Sončni rob-Hudo polje Gačnik Divje jezero Gačnik Skopica Mali Golak Skopica-Vučja grapa Govškarca-Kanomlja Gačnik Govškarca-Kanomlja Drnulk-Kobilica Jagršče-Sjavnica Gorenja T rebuša-Srna grapa Kazarska grapa-Žabže Kazarska grapa-Žabže Kanomlja-V Studencu Govškarca-Kanomlja Gorenja T rebuša-Drnulk Skopica Gorenja T rebuša-Drnulk Šebrelje-Kanačnk Quadrant (Kvadrant) 9949/1 9949/1 9949/1 9949/1 9949/4 9948/2 9949/2 9948/2 9948/2 9849/3 9949/1 9949/2 9949/2 9949/2 9949/1 0050/1 0049/1 9949/3 0050/1 9949/3 9948/2 0049/1 9848/4 9950/3 9949/1 9949/4 9948/2 9949/2 9949/3 9849/4 9849/3 9949/3 9949/4 9948/2 9948/2 9948/2 9849/3 Coordinate GK Y (D-48) m 414439 414436 412009 409858 421985 409627 418040 408491 408574 415408 410338 418059 418052 418044 412186 425057 414658 412655 425017 412456 408404 412475 408583 422739 411972 421919 408536 418048 410696 416359 416285 415822 421955 408234 408398 408264 415931 Coordinate GK X (D-48) m 5101703 5101709 5101929 5101922 5099617 5103347 5105396 5102623 5102681 5106880 5104116 5105394 5105339 5105356 5101817 5093568 5094216 5100796 5093614 5100959 5106342 5093446 5106940 5099403 5101954 5099660 5102698 5105306 5100026 5110990 5110887 5100016 5099625 5103552 5106348 5103490 5108041 Diagnostic species of the association (Diagnostične vrste asociacije) AP Primula carniolica E1 r 1 + 1 3 3 2 2 2 + + 1 1 1 1 2 2 1 2 2 1 + 1 + 1 2 2 1 2 2 2 2 2 1 1 + 2 PcSp Phyteuma scheuchzeri subsp. columnae E1 + + 1 + + + + + r + + 1 + 1 r + 1 + + + 1 1 1 + + + 1 1 + + + + 1 1 + 1 1 AP Paederota lutea E1 1 2 2 2 1 1 2 2 2 + 1 2 1 1 . 1 2 2 1 1 2 1 1 + 1 + 2 1 1 . . 2 2 2 1 2 1 AP Orthothecium rufescens E0 + + + + 2 1 1 + 1 + 1 1 + 1 1 1 + . . + . . + + + 1 3 2 + 1 1 . + + . . . AP Valeriana tripteris E1 1 + 1 . . . 1 r . . . . + + . + + . r . . . 1 + + + + 1 + + . + . + + + 2 AP Carex brachystachys E1 + r 2 + 1 1 . . + . . + + . . + + + . r . . . . + + r + r . r r + + . + + AP Cystopteridion s. lat. (Astrantio-Paederotion luteae nom. prov.) Aster bellidiastrum E1 + 1 1 1 1 . . . . 1 1 + + 1 + + + 1 1 1 1 1 + 1 1 + . 1 + . . + . . . . . Asplenium viride E1 . r + . . . + r . . . . . . + 1 + + . . . + + + + + + 1 + . . + . r . . . Fissidens dubius E0 + + . . . . 1 + . . . . . 1 . . . . . . . + . + . . . 1 + . . . . + . . . Valeriana saxatilis E1 . . . . . 1 . . 1 . 2 1 1 1 + + r 1 + . . . r . . . . . . . . + . . . . . Tofieldia calyculata E1 . . . 1 . + + 1 1 . + . . . . + . . . . . . . + . . . . . . . . . . . . . Viola biflora E1 2 + + . . . . . . . . . . . . . + 1 . + . . . + . + . . . . . + . . . . . Barbula crocea E0 . . 2 . . . . . . . . . . . 1 . + . . . . . . . . . . . . . . . . . . . . Cystopteris fragilis E1 1 1 + . . . . . . . . . . . . . . + . . r . . . . . . . . . . . . . . . . Preissia quadrata E0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . 19/2 • 2020, 155–231 195 Igor Dakskobler & Andrej Martinčič Plant communities of moist rock crevices with endemic Primula carniolica in the (sub)montane belt of western Slovenia Number of relevé (Zaporedna številka popisa) 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 31 32 33 34 35 36 37 Database number of relevé (Delovna številka popisa) 259019 259020 273980 217874 258592 275224 269300 278013 278010 226622 266154 269299 269303 269301 273965 230232 246867 274036 230234 268195 241497 259048 255777 258581 273979 258597 275218 269305 274329 269366 269369 230610 258593 275599 241495 275601 246308 Elevation in m (Nadmorska višina v m) 975 975 745 260 485 230 475 700 660 215 250 475 570 475 780 350 1250 895 325 890 800 1460 450 390 745 500 650 470 480 460 420 830 495 775 780 760 440 Aspect (Lega) NE NE NE SW S E NW N N NW N NE SEE E NW NE N SW N E NE NW E N SW NE NNE NE N SSW E NWW SW NW NE NE NW Slope in degrees (Nagib v stopinjah) 80 90 90 90 80 95 70 90 80 90 90 90 90 80 85 90 80 90 90 85 80 80 80 90 90 90 80 80 85 70 80 95 90 85 80 85 90 Parent material (Matična podlaga) D D D D D D D D D D D D D D D D D D D D D L D D D D D D D D D D D D D D D Soil (Tla) Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Stoniness in % (Kamnitost v %) 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 Cover of herb layer in % (Zastiranje zeliščne plasti v %) E1 30 20 30 30 40 40 35 20 30 20 25 30 25 30 15 30 35 20 20 30 30 30 30 20 30 30 30 40 30 30 20 30 30 20 20 25 20 Cover of moss layer in % (Zastiranje mahovne plasti v %) E0 30 20 30 10 20 10 50 10 5 10 20 20 10 30 20 10 20 30 30 20 10 20 20 10 10 20 30 40 20 20 20 5 10 10 5 10 20 Number of species (Število vrst) 24 24 16 17 13 10 20 27 20 11 10 14 13 17 15 29 32 14 18 15 24 15 25 12 10 18 15 26 26 15 15 17 11 14 16 15 20 Relevé area (Velikost popisne ploskve) m 2 10 10 10 10 3 10 10 15 10 5 10 10 10 10 10 10 10 10 10 5 10 4 5 4 10 4 10 10 10 5 5 10 2 10 10 10 10 Date of taking relevé (Datum popisa) 6/25/2015 6/25/2015 7/2/2018 4/19/2007 9/29/2015 4/18/2019 6/20/2017 4/29/2019 4/29/2019 5/8/2009 4/9/2017 6/20/2017 6/20/2017 6/20/2017 7/2/2018 4/14/2009 8/13/2003 7/16/2018 4/14/2009 8/30/2017 4/12/2011 6/22/2015 5/14/2002 9/29/2015 7/2/2018 9/29/2015 4/18/2019 6/20/2017 5/8/2018 3/31/2017 3/31/2017 5/13/2005 9/29/2015 5/16/2019 4/12/2011 5/16/2019 6/17/2011 Locality (Nahajališče) Oblakov vrh-Maslinca Oblakov vrh-Maslinca Gačnik Pršjak Govškarca-Kanomlja Gorenja trebuša - Ščura Jagršče-Sjavnica Gorenja T rebuša-Drnulk Gorenja T rebuša-Drnulk Stopnik-Idrijca Gačnik Jagršče-Sjavnica Jagršče-Sjavnica Jagršče-Sjavnica Gačnik Divje jezero Sončni rob-Hudo polje Gačnik Divje jezero Gačnik Skopica Mali Golak Skopica-Vučja grapa Govškarca-Kanomlja Gačnik Govškarca-Kanomlja Drnulk-Kobilica Jagršče-Sjavnica Gorenja T rebuša-Srna grapa Kazarska grapa-Žabže Kazarska grapa-Žabže Kanomlja-V Studencu Govškarca-Kanomlja Gorenja T rebuša-Drnulk Skopica Gorenja T rebuša-Drnulk Šebrelje-Kanačnk Quadrant (Kvadrant) 9949/1 9949/1 9949/1 9949/1 9949/4 9948/2 9949/2 9948/2 9948/2 9849/3 9949/1 9949/2 9949/2 9949/2 9949/1 0050/1 0049/1 9949/3 0050/1 9949/3 9948/2 0049/1 9848/4 9950/3 9949/1 9949/4 9948/2 9949/2 9949/3 9849/4 9849/3 9949/3 9949/4 9948/2 9948/2 9948/2 9849/3 Coordinate GK Y (D-48) m 414439 414436 412009 409858 421985 409627 418040 408491 408574 415408 410338 418059 418052 418044 412186 425057 414658 412655 425017 412456 408404 412475 408583 422739 411972 421919 408536 418048 410696 416359 416285 415822 421955 408234 408398 408264 415931 Coordinate GK X (D-48) m 5101703 5101709 5101929 5101922 5099617 5103347 5105396 5102623 5102681 5106880 5104116 5105394 5105339 5105356 5101817 5093568 5094216 5100796 5093614 5100959 5106342 5093446 5106940 5099403 5101954 5099660 5102698 5105306 5100026 5110990 5110887 5100016 5099625 5103552 5106348 5103490 5108041 Diagnostic species of the association (Diagnostične vrste asociacije) AP Primula carniolica E1 r 1 + 1 3 3 2 2 2 + + 1 1 1 1 2 2 1 2 2 1 + 1 + 1 2 2 1 2 2 2 2 2 1 1 + 2 PcSp Phyteuma scheuchzeri subsp. columnae E1 + + 1 + + + + + r + + 1 + 1 r + 1 + + + 1 1 1 + + + 1 1 + + + + 1 1 + 1 1 AP Paederota lutea E1 1 2 2 2 1 1 2 2 2 + 1 2 1 1 . 1 2 2 1 1 2 1 1 + 1 + 2 1 1 . . 2 2 2 1 2 1 AP Orthothecium rufescens E0 + + + + 2 1 1 + 1 + 1 1 + 1 1 1 + . . + . . + + + 1 3 2 + 1 1 . + + . . . AP Valeriana tripteris E1 1 + 1 . . . 1 r . . . . + + . + + . r . . . 1 + + + + 1 + + . + . + + + 2 AP Carex brachystachys E1 + r 2 + 1 1 . . + . . + + . . + + + . r . . . . + + r + r . r r + + . + + AP Cystopteridion s. lat. (Astrantio-Paederotion luteae nom. prov.) Aster bellidiastrum E1 + 1 1 1 1 . . . . 1 1 + + 1 + + + 1 1 1 1 1 + 1 1 + . 1 + . . + . . . . . Asplenium viride E1 . r + . . . + r . . . . . . + 1 + + . . . + + + + + + 1 + . . + . r . . . Fissidens dubius E0 + + . . . . 1 + . . . . . 1 . . . . . . . + . + . . . 1 + . . . . + . . . Valeriana saxatilis E1 . . . . . 1 . . 1 . 2 1 1 1 + + r 1 + . . . r . . . . . . . . + . . . . . Tofieldia calyculata E1 . . . 1 . + + 1 1 . + . . . . + . . . . . . . + . . . . . . . . . . . . . Viola biflora E1 2 + + . . . . . . . . . . . . . + 1 . + . . . + . + . . . . . + . . . . . Barbula crocea E0 . . 2 . . . . . . . . . . . 1 . + . . . . . . . . . . . . . . . . . . . . Cystopteris fragilis E1 1 1 + . . . . . . . . . . . . . . + . . r . . . . . . . . . . . . . . . . Preissia quadrata E0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . 19/2 • 2020, 155–231 196 Igor Dakskobler & Andrej Martinčič Plant communities of moist rock crevices with endemic Primula carniolica in the (sub)montane belt of western Slovenia Number of relevé (Zaporedna številka popisa) 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 31 32 33 34 35 36 37 Hymenostylium recurvirostre E0 + + . . . . . . + . . . . . . . + . . . . . . . . . . . + . . . . . . . . Moehringia muscosa E1 + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Primula x venusta E1 . . . . . . . . . . . . . . . . . . . . r . . . . . . . . . . . . . . . . Pellia endiviifolia E0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . Selaginella helvetica E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Palustriella commutata E0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . Heliosperma veselskyi subsp. veselskyi E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . r . PcSp Physoplexido comosae-Saxifragion petraeae Campanula cespitosa E1 . . . + + . . + + . . . . . . . . . . . . . . . . . . . . . . . . . . . . Athamanta turbith E1 . . . . . . . . r . . . . . . . . . . . . . . . . . . . . . . . . . . . . Hieracium porrifolium E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . r . . PC Potentilletalia caulescentis Potentilla caulescens E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + + Kernera saxatilis E1 . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . . . . . . . Primula auricula E1 . . . . . . . . . . . . . . . . . . . . 1 . . . . . . . . . . . . . . . . AT Asplenietea trichomanis Asplenium ruta-muraria E1 . . . + + . . r . + . . . . + + 1 1 . + + + + . . + + . . + . + r + + . 1 Asplenium trichomanes E1 1 + . . . . . r . . + . . . + + . + . . . . 1 . . . + . + 1 . + 1 1 + + 1 Hieracium glaucum E1 . . . . . . . + 1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . TR Thlaspietea rotundifolii Hieracium bifidum E1 . . . + + . + + + . . . . + . . . . . . + r . . . . + + . . + + r . + + + Gymnocarpium robertianum E1 . . . . . . . . . . . . . . r . . . . . . . . . . . . + . . . . . r . . . Adenostyles glabra E1 . . . . . . . . . . . . . . . . . . . . . . . . . r . . . . . . . . . . . Aquilegia iulia E1 . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . . . . . . . Dryopteris villarii E1 . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . . MC Montio-Cardaminetea Conocephalum conicum E0 . . . . . . . . . . . 1 . . 1 . . . 1 + . . + 1 . 1 + 1 + . + . . . . . . Distichium capillaceum E0 + + . + . . . . . . . . . . . . 1 . . . . . . . . . . . . . . . . . . . . Ditrichum flexicaule E0 . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . . . . . . . Campylium stellatum E0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Cololejeunea calcarea E0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . ES Elyno-Seslerietea Sesleria caerulea E1 . . . . . . + 1 1 1 + 1 1 2 + 1 1 . + . 1 r . . . . . + . . + . . . + + . Carex mucronata E1 . . . . . . . . + . . . . . . + . . . . + . . . . . . . . . . . . . + . . Betonica alopecuros E1 . . . . . . . . . . . + . . . . . . . . . . . . . . . . . . . . . . . . . Carduus crassifolius E1 . . . . . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . r . . Carex ferruginea E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Gentiana clusii E1 . . . . . . . . + . . . . . . . . . . . . . . . . . . . . . . . . . . . . Laserpitium peucedanoides E1 . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . . . . . . . FB Festuco-Brometea Carex humilis E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Buphthalmum salicifolium E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . r . . Galium lucidum E1 . . . . . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . . . Brachypodium rupestre E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . r . . . . . . MA Molinio-Arrhenatheretea Angelica sylvestris E1 . . . . . . . . . . . . . . . r . . r . . . . . . . . + . . . . . . . . . Caltha palustris E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . r . . . . . . T araxacum x Ruderalia E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . BA Betulo-Alnetea Salix appendiculata E1 . . . + . . . + . . . + . . . . . . . . + . . . . . . . . . . . . . . . . Salix glabra E1 . . . . . . . + + . . . . . . . . . . . . . . . . . . . . . . . . . . . . MuA Mulgedio-Aconitetea Senecio ovatus E1 . . . . . . . . . . . . . . . . . . . . . . r . . . . . . . . . . . . . r Ranunculus platanifolius E1 . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . . Aconitum degenii subsp. paniculatum E1 . . . . . . . . . . . . . . . . . . . . . . r . . . . . . . . . . . . . . Saxifraga rotundifolia E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . TG Trifolio-Geranietea Laserpitium siler E1 . . . . . . . . . . . . + + . . . . . . . . . . . . . . . . . . . . . . . Viola hirta E1 . . . . . . . . . . . . . . . . . . . . r . . . . . . . . . . . . . . . . 19/2 • 2020, 155–231 197 Igor Dakskobler & Andrej Martinčič Plant communities of moist rock crevices with endemic Primula carniolica in the (sub)montane belt of western Slovenia Number of relevé (Zaporedna številka popisa) 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 31 32 33 34 35 36 37 Hymenostylium recurvirostre E0 + + . . . . . . + . . . . . . . + . . . . . . . . . . . + . . . . . . . . Moehringia muscosa E1 + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Primula x venusta E1 . . . . . . . . . . . . . . . . . . . . r . . . . . . . . . . . . . . . . Pellia endiviifolia E0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . Selaginella helvetica E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Palustriella commutata E0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . Heliosperma veselskyi subsp. veselskyi E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . r . PcSp Physoplexido comosae-Saxifragion petraeae Campanula cespitosa E1 . . . + + . . + + . . . . . . . . . . . . . . . . . . . . . . . . . . . . Athamanta turbith E1 . . . . . . . . r . . . . . . . . . . . . . . . . . . . . . . . . . . . . Hieracium porrifolium E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . r . . PC Potentilletalia caulescentis Potentilla caulescens E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + + Kernera saxatilis E1 . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . . . . . . . Primula auricula E1 . . . . . . . . . . . . . . . . . . . . 1 . . . . . . . . . . . . . . . . AT Asplenietea trichomanis Asplenium ruta-muraria E1 . . . + + . . r . + . . . . + + 1 1 . + + + + . . + + . . + . + r + + . 1 Asplenium trichomanes E1 1 + . . . . . r . . + . . . + + . + . . . . 1 . . . + . + 1 . + 1 1 + + 1 Hieracium glaucum E1 . . . . . . . + 1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . TR Thlaspietea rotundifolii Hieracium bifidum E1 . . . + + . + + + . . . . + . . . . . . + r . . . . + + . . + + r . + + + Gymnocarpium robertianum E1 . . . . . . . . . . . . . . r . . . . . . . . . . . . + . . . . . r . . . Adenostyles glabra E1 . . . . . . . . . . . . . . . . . . . . . . . . . r . . . . . . . . . . . Aquilegia iulia E1 . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . . . . . . . Dryopteris villarii E1 . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . . MC Montio-Cardaminetea Conocephalum conicum E0 . . . . . . . . . . . 1 . . 1 . . . 1 + . . + 1 . 1 + 1 + . + . . . . . . Distichium capillaceum E0 + + . + . . . . . . . . . . . . 1 . . . . . . . . . . . . . . . . . . . . Ditrichum flexicaule E0 . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . . . . . . . Campylium stellatum E0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Cololejeunea calcarea E0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . ES Elyno-Seslerietea Sesleria caerulea E1 . . . . . . + 1 1 1 + 1 1 2 + 1 1 . + . 1 r . . . . . + . . + . . . + + . Carex mucronata E1 . . . . . . . . + . . . . . . + . . . . + . . . . . . . . . . . . . + . . Betonica alopecuros E1 . . . . . . . . . . . + . . . . . . . . . . . . . . . . . . . . . . . . . Carduus crassifolius E1 . . . . . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . r . . Carex ferruginea E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Gentiana clusii E1 . . . . . . . . + . . . . . . . . . . . . . . . . . . . . . . . . . . . . Laserpitium peucedanoides E1 . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . . . . . . . FB Festuco-Brometea Carex humilis E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Buphthalmum salicifolium E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . r . . Galium lucidum E1 . . . . . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . . . Brachypodium rupestre E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . r . . . . . . MA Molinio-Arrhenatheretea Angelica sylvestris E1 . . . . . . . . . . . . . . . r . . r . . . . . . . . + . . . . . . . . . Caltha palustris E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . r . . . . . . T araxacum x Ruderalia E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . BA Betulo-Alnetea Salix appendiculata E1 . . . + . . . + . . . + . . . . . . . . + . . . . . . . . . . . . . . . . Salix glabra E1 . . . . . . . + + . . . . . . . . . . . . . . . . . . . . . . . . . . . . MuA Mulgedio-Aconitetea Senecio ovatus E1 . . . . . . . . . . . . . . . . . . . . . . r . . . . . . . . . . . . . r Ranunculus platanifolius E1 . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . . Aconitum degenii subsp. paniculatum E1 . . . . . . . . . . . . . . . . . . . . . . r . . . . . . . . . . . . . . Saxifraga rotundifolia E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . TG Trifolio-Geranietea Laserpitium siler E1 . . . . . . . . . . . . + + . . . . . . . . . . . . . . . . . . . . . . . Viola hirta E1 . . . . . . . . . . . . . . . . . . . . r . . . . . . . . . . . . . . . . 19/2 • 2020, 155–231 198 Igor Dakskobler & Andrej Martinčič Plant communities of moist rock crevices with endemic Primula carniolica in the (sub)montane belt of western Slovenia Number of relevé (Zaporedna številka popisa) 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 31 32 33 34 35 36 37 Anthericum ramosum E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Laserpitium latifolium E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Thalictrum minus E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . EA Epilobietea angustifolii Rubus idaeus E1 . . . . . . . . . . . . . . . . r . . . . . . . . . . . . . . . . . . . . EP Erico-Pinetea Calamagrostis varia E1 1 . . + + + + + . + . . . + . . . . + + + r . . . . . . . . . + . . . . . Rhododendron hirsutum E1 . . . . . . + + + . . + . + + . + . . . . . . . . . . . . . . . . . . + . Erica carnea E1 . . . + . + . + + . . . . . . . . . . . . . . . . . . . . . . . . . . . . Polygala chamaebuxus E1 . . . + . . + + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Carex ornithopoda E1 . . . . + . . . . . . . . . . . . . + . . . r . . . . . . . . . . . r . . Rubus saxatilis E1 . . . . . . . . . . . . + . . . . . . . . . . . . . . . . . . r . . . . . Carex alba E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Cirsium erisithales E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Leontodon incanus E1 . . . 1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Molinia arundinacea E1 . . . . . + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Amelanchier ovalis E1 . . . . . . . + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Aster amellus E1 . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . . . . . . . . Rhodothamnus chamaecistus E1 . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . . . . . . . Aquilegia nigricans E1 . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . . . . Pinus sylvestris E2a . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . VP Vaccinio-Pinetea Veronica urticifolia E1 . . . . . . . . . . . . + . . . . . . . . . r . . + . . + + r . . . . . . Homogyne sylvestris E1 . . . . . . . . . . . . . . + . . . . + . . . . . . r + . . . . . . . r . Solidago virgaurea E1 . . . . . . . r . . . . . . . . . . . . . . . . . . . r . . . r . . . . . Hieracium murorum E1 . . . . . . . . . . . . . . . + . . + . . . r . . . . . . . . . . . . . . Clematis alpina E1 . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . . r . . . . . Gentiana asclepiadea E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Oxalis acetosella E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Picea abies E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Polystichum lonchitis E1 . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . . Aposeris foetida E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Rosa pendulina E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . AF Aremonio-Fagion Cyclamen purpurascens E1 . . + . . . . . . . . . . . . . . . . . . . r . . . . . r . . . . . . . . Helleborus niger E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Hemerocallis lilioasphodelus E1 . . . . . . . . . + . . . . . . . . . . . . + . . . . . . . . . . . . . . Potentilla carniolica E1 . . . . . . . . . . . . . . . + . . . . . . . . . . . + . . . . . . . . . Cardamine trifolia E1 . . . . . . . . . . . . . . . r . . . . . . . . . . . . . . . . . . . . . Primula vulgaris E1 . . . . . . . . . . . . . . . . . . . . r . . . . . . . . . . . . . . . . Anemone trifolia E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Euphorbia carniolica E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . r . . . . . . . . . Lamium orvala E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . Rhamnus fallax E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Laserpitium krapfii E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Omphalodes verna E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Knautia drymeia E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . TA Tilio-Acerion Aruncus dioicus E1 . . . . . . . . . . . . . + . . . . . r . . + . . . . + . + . . . . . . . Polystichum aculeatum E1 r . . . . . . . . . . . . . . . . . . . . . . . . r . . . . . . . . . . + Phyllitis scolopendrium E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . Acer pseudoplatanus E1 r . . . . . . . . . . . . . . . . . r . + . r . . . . . . . . . . . . . . Geranium robertianum E1 + + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . T ephroseris pseudocrispa E1 . . . . . . . . . . . . . + . . . . . . . . . . . . . . . . r . . . . . . Thalictrum aquilegiifolium E1 . . . . . . . . . . . . . . . r . . . . . . . . . . . . . . . . . . . . . Ulmus glabra E2a . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 Acer platanoides E2a . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + 19/2 • 2020, 155–231 199 Igor Dakskobler & Andrej Martinčič Plant communities of moist rock crevices with endemic Primula carniolica in the (sub)montane belt of western Slovenia Number of relevé (Zaporedna številka popisa) 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 31 32 33 34 35 36 37 Anthericum ramosum E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Laserpitium latifolium E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Thalictrum minus E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . EA Epilobietea angustifolii Rubus idaeus E1 . . . . . . . . . . . . . . . . r . . . . . . . . . . . . . . . . . . . . EP Erico-Pinetea Calamagrostis varia E1 1 . . + + + + + . + . . . + . . . . + + + r . . . . . . . . . + . . . . . Rhododendron hirsutum E1 . . . . . . + + + . . + . + + . + . . . . . . . . . . . . . . . . . . + . Erica carnea E1 . . . + . + . + + . . . . . . . . . . . . . . . . . . . . . . . . . . . . Polygala chamaebuxus E1 . . . + . . + + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Carex ornithopoda E1 . . . . + . . . . . . . . . . . . . + . . . r . . . . . . . . . . . r . . Rubus saxatilis E1 . . . . . . . . . . . . + . . . . . . . . . . . . . . . . . . r . . . . . Carex alba E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Cirsium erisithales E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Leontodon incanus E1 . . . 1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Molinia arundinacea E1 . . . . . + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Amelanchier ovalis E1 . . . . . . . + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Aster amellus E1 . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . . . . . . . . Rhodothamnus chamaecistus E1 . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . . . . . . . Aquilegia nigricans E1 . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . . . . Pinus sylvestris E2a . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . VP Vaccinio-Pinetea Veronica urticifolia E1 . . . . . . . . . . . . + . . . . . . . . . r . . + . . + + r . . . . . . Homogyne sylvestris E1 . . . . . . . . . . . . . . + . . . . + . . . . . . r + . . . . . . . r . Solidago virgaurea E1 . . . . . . . r . . . . . . . . . . . . . . . . . . . r . . . r . . . . . Hieracium murorum E1 . . . . . . . . . . . . . . . + . . + . . . r . . . . . . . . . . . . . . Clematis alpina E1 . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . . r . . . . . Gentiana asclepiadea E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Oxalis acetosella E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Picea abies E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Polystichum lonchitis E1 . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . . Aposeris foetida E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Rosa pendulina E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . AF Aremonio-Fagion Cyclamen purpurascens E1 . . + . . . . . . . . . . . . . . . . . . . r . . . . . r . . . . . . . . Helleborus niger E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Hemerocallis lilioasphodelus E1 . . . . . . . . . + . . . . . . . . . . . . + . . . . . . . . . . . . . . Potentilla carniolica E1 . . . . . . . . . . . . . . . + . . . . . . . . . . . + . . . . . . . . . Cardamine trifolia E1 . . . . . . . . . . . . . . . r . . . . . . . . . . . . . . . . . . . . . Primula vulgaris E1 . . . . . . . . . . . . . . . . . . . . r . . . . . . . . . . . . . . . . Anemone trifolia E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Euphorbia carniolica E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . r . . . . . . . . . Lamium orvala E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . Rhamnus fallax E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Laserpitium krapfii E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Omphalodes verna E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Knautia drymeia E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . TA Tilio-Acerion Aruncus dioicus E1 . . . . . . . . . . . . . + . . . . . r . . + . . . . + . + . . . . . . . Polystichum aculeatum E1 r . . . . . . . . . . . . . . . . . . . . . . . . r . . . . . . . . . . + Phyllitis scolopendrium E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . Acer pseudoplatanus E1 r . . . . . . . . . . . . . . . . . r . + . r . . . . . . . . . . . . . . Geranium robertianum E1 + + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . T ephroseris pseudocrispa E1 . . . . . . . . . . . . . + . . . . . . . . . . . . . . . . r . . . . . . Thalictrum aquilegiifolium E1 . . . . . . . . . . . . . . . r . . . . . . . . . . . . . . . . . . . . . Ulmus glabra E2a . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 Acer platanoides E2a . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + 19/2 • 2020, 155–231 200 Igor Dakskobler & Andrej Martinčič Plant communities of moist rock crevices with endemic Primula carniolica in the (sub)montane belt of western Slovenia Number of relevé (Zaporedna številka popisa) 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 31 32 33 34 35 36 37 FS Fagetalia sylvaticae Salvia glutinosa E1 . . . . . . . . . . . . . . . . . . . . . . r . . . . . . . . . . . + . + Galeobdolon flavidum E1 . r + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Mercurialis perennis E1 . . . . . . . . . . . . . . . . . . . . r . . . . . . . . . . . r . . . . Mycelis muralis E1 r + . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . Galium laevigatum E1 . . . . . . . . . . . . . + . r . . . . . . . . . . . + r . . . . . . . . Fagus sylvatica E1 . . . . . . . . . . . . . . . r . . . . . . . . . . . . . . . . . . . . . Melica nutans E1 . . . . . . . . . . . + . . . . . . . . . . . . . . . . . . . . . . . . . Symphytum tuberosum E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Campanula trachelium E1 . . . . . . . . . . . . . . . r . . . . . . . . . . . . . . . . . . . . . Cardamine pentaphyllos E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Asarum europaeum subsp. caucasicum E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Petasites albus E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Sambucus nigra E2a . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Lathyrus vernus E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . QP Quercetalia pubescenti-petraeae Fraxinus ornus E1 . . . . . . . . . . . . . . . . . . . . + . r . . . . + r . . . . . r + + Ostrya carpinifolia E1 . . . . . . . + + . . . . . . . . . . . . . . . . . . . . . r . . . . r . Mercurialis ovata E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Clematis recta E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + Euonymus verrucosa E2a . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + QF Querco-Fagetea Carex digitata E1 . . . . . . + + . . . . . . . . . . + + r . . . . + + + . 1 + . r . . . . Hepatica nobilis E1 . . + . . . . . . . . . . . . r . + r . r . . . . r . . + . . . . r . . . Hedera helix E1 . . . . . . . . . + . . . . . . . . . . . . . . . . . . . + + . . . . . + Veratrum nigrum E1 . . . . . . . . . . . . . . . . . . . . r . . . . . . . . . . . . . . . . Acer campestre E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Clematis vitalba E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . Quercus petraea E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Cardamine impatiens E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Lonicera xylosteum E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . ML Mosses and lichens (Mahovi in lišaji) Neckera crispa E0 1 1 1 . + . . + . . . . . . . 1 1 . . . . . . . . 1 + 1 1 1 + + 1 1 + + 2 Ctenidium molluscum E0 1 1 . . . . + + . . . . . . . + 1 1 2 1 1 1 . . + . 1 1 + 1 . . . . . . . Tortella tortuosa E0 . . 1 + 1 . 2 1 + 1 + 1 1 2 1 + + + + + . . . + . . . 1 + . . . . . . . . Mnium thomsonii E0 . . . . . . 1 . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . Schistidium apocarpum E0 . . . . . . . . . . . . . . . + + . + . . . . . . . . . . . . . . . . . . Plagiochila porreloides E0 + + . . . . . . . . . . . . . . + . . . . . . . . . . . . . . . . . . . . Encalypta streptocarpa E0 + + . . . . . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . Leiocolea collaris E0 . . . . . . + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Marchantia polymorpha E0 . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . . . . . . . . Bryum capillare E0 . . . . . . . . . . . . . . . . + . . . . 1 . . . . . . . . . . . . . . . Pedinophyllum interruptum E0 . . . . . . + . . . . . . . . . + . . . . . . . . . . . . . . . . . . . . Mnium stellare E0 . . . . . . . . . . . . . . . . . . . . . . + . . . . + . . . . . . . . . Isothecium alopecuroides E0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + Lejeunea cavifolia E0 . + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Myurella sibirica E0 . + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Neckera complanata E0 . + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Didymodon ferrugineus E0 . . . . . . + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Trichostomum brachydontium E0 . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . . . . . . . Myurella julacea E0 . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . . . . . . . Hypnum cupressiforme var. resupinatum E0 . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . . . . . . . Lichenes div. E0 . . . . . . . . . . . . . . . . . . . . . . 1 . . . . . . . . . . . . . . Lophozia sp. E0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . Dichodontium pellucidum subsp. flavescens E0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Brachythecium rutabulum E0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Plagiomnium rostratum E0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Mnium marginatum E0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Plagiobryum zierii E0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 19/2 • 2020, 155–231 201 Igor Dakskobler & Andrej Martinčič Plant communities of moist rock crevices with endemic Primula carniolica in the (sub)montane belt of western Slovenia Number of relevé (Zaporedna številka popisa) 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 31 32 33 34 35 36 37 FS Fagetalia sylvaticae Salvia glutinosa E1 . . . . . . . . . . . . . . . . . . . . . . r . . . . . . . . . . . + . + Galeobdolon flavidum E1 . r + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Mercurialis perennis E1 . . . . . . . . . . . . . . . . . . . . r . . . . . . . . . . . r . . . . Mycelis muralis E1 r + . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . Galium laevigatum E1 . . . . . . . . . . . . . + . r . . . . . . . . . . . + r . . . . . . . . Fagus sylvatica E1 . . . . . . . . . . . . . . . r . . . . . . . . . . . . . . . . . . . . . Melica nutans E1 . . . . . . . . . . . + . . . . . . . . . . . . . . . . . . . . . . . . . Symphytum tuberosum E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Campanula trachelium E1 . . . . . . . . . . . . . . . r . . . . . . . . . . . . . . . . . . . . . Cardamine pentaphyllos E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Asarum europaeum subsp. caucasicum E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Petasites albus E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Sambucus nigra E2a . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Lathyrus vernus E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . QP Quercetalia pubescenti-petraeae Fraxinus ornus E1 . . . . . . . . . . . . . . . . . . . . + . r . . . . + r . . . . . r + + Ostrya carpinifolia E1 . . . . . . . + + . . . . . . . . . . . . . . . . . . . . . r . . . . r . Mercurialis ovata E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Clematis recta E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + Euonymus verrucosa E2a . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + QF Querco-Fagetea Carex digitata E1 . . . . . . + + . . . . . . . . . . + + r . . . . + + + . 1 + . r . . . . Hepatica nobilis E1 . . + . . . . . . . . . . . . r . + r . r . . . . r . . + . . . . r . . . Hedera helix E1 . . . . . . . . . + . . . . . . . . . . . . . . . . . . . + + . . . . . + Veratrum nigrum E1 . . . . . . . . . . . . . . . . . . . . r . . . . . . . . . . . . . . . . Acer campestre E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Clematis vitalba E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . Quercus petraea E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Cardamine impatiens E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Lonicera xylosteum E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . ML Mosses and lichens (Mahovi in lišaji) Neckera crispa E0 1 1 1 . + . . + . . . . . . . 1 1 . . . . . . . . 1 + 1 1 1 + + 1 1 + + 2 Ctenidium molluscum E0 1 1 . . . . + + . . . . . . . + 1 1 2 1 1 1 . . + . 1 1 + 1 . . . . . . . Tortella tortuosa E0 . . 1 + 1 . 2 1 + 1 + 1 1 2 1 + + + + + . . . + . . . 1 + . . . . . . . . Mnium thomsonii E0 . . . . . . 1 . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . Schistidium apocarpum E0 . . . . . . . . . . . . . . . + + . + . . . . . . . . . . . . . . . . . . Plagiochila porreloides E0 + + . . . . . . . . . . . . . . + . . . . . . . . . . . . . . . . . . . . Encalypta streptocarpa E0 + + . . . . . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . Leiocolea collaris E0 . . . . . . + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Marchantia polymorpha E0 . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . . . . . . . . Bryum capillare E0 . . . . . . . . . . . . . . . . + . . . . 1 . . . . . . . . . . . . . . . Pedinophyllum interruptum E0 . . . . . . + . . . . . . . . . + . . . . . . . . . . . . . . . . . . . . Mnium stellare E0 . . . . . . . . . . . . . . . . . . . . . . + . . . . + . . . . . . . . . Isothecium alopecuroides E0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + Lejeunea cavifolia E0 . + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Myurella sibirica E0 . + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Neckera complanata E0 . + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Didymodon ferrugineus E0 . . . . . . + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Trichostomum brachydontium E0 . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . . . . . . . Myurella julacea E0 . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . . . . . . . Hypnum cupressiforme var. resupinatum E0 . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . . . . . . . Lichenes div. E0 . . . . . . . . . . . . . . . . . . . . . . 1 . . . . . . . . . . . . . . Lophozia sp. E0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . Dichodontium pellucidum subsp. flavescens E0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Brachythecium rutabulum E0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Plagiomnium rostratum E0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Mnium marginatum E0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Plagiobryum zierii E0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 19/2 • 2020, 155–231 202 Igor Dakskobler & Andrej Martinčič Plant communities of moist rock crevices with endemic Primula carniolica in the (sub)montane belt of western Slovenia Table 5 (Tabela 5): Phyteumato columnae-Primuletum carniolicae. Relevé number 38–73. Number of relevé (Zaporedna številka popisa) 38 39 40 41 42 43 44 45 46 47 48 49 50 51 52 53 54 55 56 57 58 59 60 61 62 63 64 65 66 67 68 69 70 71 72 73 Pr. Fr. Database number of relevé (Delovna številka popisa) 230615 278030 278031 242921 274414 278019 242923 275598 273969 274020 242931 258598 269424 269425 274402 269198 278005 278008 269430 269297 246397 266158 263169 246395 268193 259018 269394 274407 275254 275255 263172 269177 269181 269182 263753 269396 Elevation in m (Nadmorska višina v m) 1050 910 925 830 505 750 840 770 775 785 890 500 770 770 535 540 680 670 780 450 330 250 450 330 870 970 654 560 690 730 420 425 430 435 380 660 Aspect (Lega) NW NW NW NE W NE N NNE S SSW NE W SEE N N N NNE NNE N NE N NW N NE NW NE NNE W N NE NE NE NE NE NW N Slope in degrees (Nagib v stopinjah) 90 90 70 90 80 90 90 90 85 90 85 80 85 80 70 70 80 80 80 85 90 100 90 80 85 90 80 80 85 90 80 80 60 80 80 90 Parent material (Matična podlaga) D D D D D D D D D D D D D D D D D D D D D D D D D D D D D D D D D D D D Soil (Tla) Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Stoniness in % (Kamnitost v %) 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 Cover of herb layer in % (Zastiranje zeliščne plasti v %) E1 20 20 20 20 30 25 15 20 20 20 30 40 30 30 30 30 35 40 30 30 30 25 20 30 30 25 30 30 30 30 30 40 30 30 30 15 Cover of moss layer in % (Zastiranje mahovne plasti v %) E0 10 10 30 20 40 10 20 20 20 10 20 40 40 40 30 50 10 20 30 30 30 25 19 20 30 20 10 30 30 30 30 60 5 20 20 10 Number of species (Število vrst) 15 18 14 14 20 16 11 14 11 11 10 18 10 17 21 15 19 23 13 23 17 12 16 34 20 24 22 24 21 29 16 24 15 18 13 6 Relevé area (Velikost popisne ploskve) m 2 10 20 10 10 10 15 10 5 10 10 10 2 5 10 10 5 15 15 5 10 50 10 10 100 5 10 5 10 15 15 10 10 5 5 10 2 Date of taking relevé (Datum popisa) 7/14/2004 4/16/2019 4/16/2019 4/18/2012 4/23/2018 4/29/2019 4/18/2012 5/16/2019 7/2/2018 7/16/2018 4/18/2012 9/29/2015 4/20/2017 4/20/2017 4/23/2018 4/7/2017 4/29/2019 4/29/2019 4/20/2017 6/20/2017 4/22/2011 4/9/2017 9/12/2016 4/22/2011 8/30/2017 6/25/2015 5/26/2107 4/23/2018 4/27/2019 5/16/2019 9/12/2016 4/7/2017 4/7/2017 4/7/2017 6/24/2016 5/26/2017 Locality (Nahajališče) Vojsko-V Studencu Stador-Kobilica Stador-Kobilica Skopica Kazarska grapa Gorenja T rebuša-Drnulk Skopica Gorenja T rebuša-Drnulk Gačnik Gačnik Velike Vrše Govškarca-Kanomlja Bukovski Vrh-Degarnik Bukovski Vrh-Degarnik Kazarska grapa Orehovska grapa-Nemci Goirenja T rebuša-Drnulk Goirenja T rebuša-Drnulk Bukovski Vrh-Degarnik Jagršče-Sjavnica Jagršče-Sjavnica Gačnik Orehovska grapa Jagršče-Sjavnica Gačnik Oblakov vrh-Maslinca Kazarska grapa-Degarnik Kazarska grapa Čepovan-Drnulk Čepovan-Drnulk Orehovska grapa Orehovska grapa Orehovska grapa Orehovska grapa Kazarska grapa Kazarska grapa-Degarnik Quadrant (Kvadrant) 9949/3 9948/2 9948/2 9948/2 9849/3 9948/2 9948/2 9948/2 9949/1 9949/1 9948/2 9949/4 9849/3 9849/3 9849/3 9849/4 9948/2 9948/2 9849/3 9949/2 9949/2 9949/1 9849/4 9849/4 9949/3 9949/1 9849/3 9849/3 9948/2 9948/2 9849/4 9849/4 9849/4 9849/4 9849/3 9849/3 Coordinate GK Y (D-48) m 415736 408043 408060 408327 415320 408334 408354 408230 412117 412115 407993 421925 415139 415127 415532 417834 408621 408411 415240 418011 417202 410303 418076 417201 412447 414448 415147 415307 407582 407804 418220 418224 418177 418172 416035 415385 Coordinate GK X (D-48) m 5099687 5101657 5101677 5106389 5111461 5102809 5106389 5103541 5101925 5101933 5105332 5099650 5110930 5110939 5111307 5111257 5103083 5103022 5110954 5105487 5106621 5104150 5110995 5106770 5100954 5101710 5111240 5111290 5102792 5103192 5110806 5110830 5110838 5110848 5110995 5111208 Diagnostic species of the association (Diagnostične vrste asociacije) Pr. Fr. AP Primula carniolica E1 1 + r + + 1 + + 1 1 2 2 2 1 1 2 2 3 1 1 1 1 1 1 1 1 1 + 2 2 1 1 3 2 + 1 73 100 PcSp Phyteuma scheuchzeri subsp. columnae E1 + + + + 1 + + + + + + + + + 1 2 1 1 r + + + + + + 1 2 1 + + + + + + + + 73 100 AP Paederota lutea E1 1 1 1 + 1 1 + 1 2 1 . + . . 1 . 2 3 . 2 1 1 . 1 1 1 . . . . . . . . . . 54 74 AP Orthothecium rufescens E0 + + + . + + . . . . . . . . . + . + . . . + + . . . . + . 1 . . . + . . 41 56 AP Valeriana tripteris E1 . . + + . . . . . . . . . r + . . + . . 1 + . 1 + + + + . 1 . . . . . . 36 49 AP Carex brachystachys E1 r . . . 1 + . . . . . . . . . . . . . + . . . r . 1 1 . + + . . . . . . 33 45 AP Cystopteridion s. lat. (Astrantio-Paederotion luteae nom. prov.) Aster bellidiastrum E1 . . . . . . . . 1 1 . + . . . . . . . + + 1 + . . . . . . + . . . . . . 33 45 Asplenium viride E1 . . . + + . + . . . . + . . . + . . . + . . . + + . . + + + . . . . . . 29 40 Fissidens dubius E0 + + . . . + . . . . . . . . . + . . . . . . . + + . . + . . + + . . 1 . 20 27 Valeriana saxatilis E1 . . . . . + . r + . . . . . r + + . . 1 . . . . . . . . . . . + + + . . 23 32 Tofieldia calyculata E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . + + . . r + . . 12 16 Viola biflora E1 + . . . . . . . . . . . . . . . . . . . . . . . + + . . . . . . . . . . 12 16 Barbula crocea E0 . . + . . . . . . . . . . . . . . + . + . . . . . . . . . . . + . . . . 7 10 Cystopteris fragilis E1 . r . . . . . . . . . . . . . . . . . . . . . . + + . + . . . . . . . . 9 12 Preissia quadrata E0 . . . . 2 1 . . . . . . . . . + + + . . . . . . . . . 1 1 + . . . . . . 9 12 19/2 • 2020, 155–231 203 Igor Dakskobler & Andrej Martinčič Plant communities of moist rock crevices with endemic Primula carniolica in the (sub)montane belt of western Slovenia Number of relevé (Zaporedna številka popisa) 38 39 40 41 42 43 44 45 46 47 48 49 50 51 52 53 54 55 56 57 58 59 60 61 62 63 64 65 66 67 68 69 70 71 72 73 Pr. Fr. Database number of relevé (Delovna številka popisa) 230615 278030 278031 242921 274414 278019 242923 275598 273969 274020 242931 258598 269424 269425 274402 269198 278005 278008 269430 269297 246397 266158 263169 246395 268193 259018 269394 274407 275254 275255 263172 269177 269181 269182 263753 269396 Elevation in m (Nadmorska višina v m) 1050 910 925 830 505 750 840 770 775 785 890 500 770 770 535 540 680 670 780 450 330 250 450 330 870 970 654 560 690 730 420 425 430 435 380 660 Aspect (Lega) NW NW NW NE W NE N NNE S SSW NE W SEE N N N NNE NNE N NE N NW N NE NW NE NNE W N NE NE NE NE NE NW N Slope in degrees (Nagib v stopinjah) 90 90 70 90 80 90 90 90 85 90 85 80 85 80 70 70 80 80 80 85 90 100 90 80 85 90 80 80 85 90 80 80 60 80 80 90 Parent material (Matična podlaga) D D D D D D D D D D D D D D D D D D D D D D D D D D D D D D D D D D D D Soil (Tla) Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Stoniness in % (Kamnitost v %) 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 Cover of herb layer in % (Zastiranje zeliščne plasti v %) E1 20 20 20 20 30 25 15 20 20 20 30 40 30 30 30 30 35 40 30 30 30 25 20 30 30 25 30 30 30 30 30 40 30 30 30 15 Cover of moss layer in % (Zastiranje mahovne plasti v %) E0 10 10 30 20 40 10 20 20 20 10 20 40 40 40 30 50 10 20 30 30 30 25 19 20 30 20 10 30 30 30 30 60 5 20 20 10 Number of species (Število vrst) 15 18 14 14 20 16 11 14 11 11 10 18 10 17 21 15 19 23 13 23 17 12 16 34 20 24 22 24 21 29 16 24 15 18 13 6 Relevé area (Velikost popisne ploskve) m 2 10 20 10 10 10 15 10 5 10 10 10 2 5 10 10 5 15 15 5 10 50 10 10 100 5 10 5 10 15 15 10 10 5 5 10 2 Date of taking relevé (Datum popisa) 7/14/2004 4/16/2019 4/16/2019 4/18/2012 4/23/2018 4/29/2019 4/18/2012 5/16/2019 7/2/2018 7/16/2018 4/18/2012 9/29/2015 4/20/2017 4/20/2017 4/23/2018 4/7/2017 4/29/2019 4/29/2019 4/20/2017 6/20/2017 4/22/2011 4/9/2017 9/12/2016 4/22/2011 8/30/2017 6/25/2015 5/26/2107 4/23/2018 4/27/2019 5/16/2019 9/12/2016 4/7/2017 4/7/2017 4/7/2017 6/24/2016 5/26/2017 Locality (Nahajališče) Vojsko-V Studencu Stador-Kobilica Stador-Kobilica Skopica Kazarska grapa Gorenja T rebuša-Drnulk Skopica Gorenja T rebuša-Drnulk Gačnik Gačnik Velike Vrše Govškarca-Kanomlja Bukovski Vrh-Degarnik Bukovski Vrh-Degarnik Kazarska grapa Orehovska grapa-Nemci Goirenja T rebuša-Drnulk Goirenja T rebuša-Drnulk Bukovski Vrh-Degarnik Jagršče-Sjavnica Jagršče-Sjavnica Gačnik Orehovska grapa Jagršče-Sjavnica Gačnik Oblakov vrh-Maslinca Kazarska grapa-Degarnik Kazarska grapa Čepovan-Drnulk Čepovan-Drnulk Orehovska grapa Orehovska grapa Orehovska grapa Orehovska grapa Kazarska grapa Kazarska grapa-Degarnik Quadrant (Kvadrant) 9949/3 9948/2 9948/2 9948/2 9849/3 9948/2 9948/2 9948/2 9949/1 9949/1 9948/2 9949/4 9849/3 9849/3 9849/3 9849/4 9948/2 9948/2 9849/3 9949/2 9949/2 9949/1 9849/4 9849/4 9949/3 9949/1 9849/3 9849/3 9948/2 9948/2 9849/4 9849/4 9849/4 9849/4 9849/3 9849/3 Coordinate GK Y (D-48) m 415736 408043 408060 408327 415320 408334 408354 408230 412117 412115 407993 421925 415139 415127 415532 417834 408621 408411 415240 418011 417202 410303 418076 417201 412447 414448 415147 415307 407582 407804 418220 418224 418177 418172 416035 415385 Coordinate GK X (D-48) m 5099687 5101657 5101677 5106389 5111461 5102809 5106389 5103541 5101925 5101933 5105332 5099650 5110930 5110939 5111307 5111257 5103083 5103022 5110954 5105487 5106621 5104150 5110995 5106770 5100954 5101710 5111240 5111290 5102792 5103192 5110806 5110830 5110838 5110848 5110995 5111208 Diagnostic species of the association (Diagnostične vrste asociacije) Pr. Fr. AP Primula carniolica E1 1 + r + + 1 + + 1 1 2 2 2 1 1 2 2 3 1 1 1 1 1 1 1 1 1 + 2 2 1 1 3 2 + 1 73 100 PcSp Phyteuma scheuchzeri subsp. columnae E1 + + + + 1 + + + + + + + + + 1 2 1 1 r + + + + + + 1 2 1 + + + + + + + + 73 100 AP Paederota lutea E1 1 1 1 + 1 1 + 1 2 1 . + . . 1 . 2 3 . 2 1 1 . 1 1 1 . . . . . . . . . . 54 74 AP Orthothecium rufescens E0 + + + . + + . . . . . . . . . + . + . . . + + . . . . + . 1 . . . + . . 41 56 AP Valeriana tripteris E1 . . + + . . . . . . . . . r + . . + . . 1 + . 1 + + + + . 1 . . . . . . 36 49 AP Carex brachystachys E1 r . . . 1 + . . . . . . . . . . . . . + . . . r . 1 1 . + + . . . . . . 33 45 AP Cystopteridion s. lat. (Astrantio-Paederotion luteae nom. prov.) Aster bellidiastrum E1 . . . . . . . . 1 1 . + . . . . . . . + + 1 + . . . . . . + . . . . . . 33 45 Asplenium viride E1 . . . + + . + . . . . + . . . + . . . + . . . + + . . + + + . . . . . . 29 40 Fissidens dubius E0 + + . . . + . . . . . . . . . + . . . . . . . + + . . + . . + + . . 1 . 20 27 Valeriana saxatilis E1 . . . . . + . r + . . . . . r + + . . 1 . . . . . . . . . . . + + + . . 23 32 Tofieldia calyculata E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . + + . . r + . . 12 16 Viola biflora E1 + . . . . . . . . . . . . . . . . . . . . . . . + + . . . . . . . . . . 12 16 Barbula crocea E0 . . + . . . . . . . . . . . . . . + . + . . . . . . . . . . . + . . . . 7 10 Cystopteris fragilis E1 . r . . . . . . . . . . . . . . . . . . . . . . + + . + . . . . . . . . 9 12 Preissia quadrata E0 . . . . 2 1 . . . . . . . . . + + + . . . . . . . . . 1 1 + . . . . . . 9 12 19/2 • 2020, 155–231 204 Igor Dakskobler & Andrej Martinčič Plant communities of moist rock crevices with endemic Primula carniolica in the (sub)montane belt of western Slovenia Number of relevé (Zaporedna številka popisa) 38 39 40 41 42 43 44 45 46 47 48 49 50 51 52 53 54 55 56 57 58 59 60 61 62 63 64 65 66 67 68 69 70 71 72 73 Pr. Fr. Hymenostylium recurvirostre E0 . . . . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . . . 6 8 Moehringia muscosa E1 + . . . . . . . + . . . . . . . . . . . . . . . . + . . . . + + . . . . 6 8 Primula x venusta E1 . r . . . . . r . . . . . . . . . . . . . . . . . . . . . . . . . . . . 3 4 Pellia endiviifolia E0 . . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . . . . . 2 3 Selaginella helvetica E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . + + . . . . . . 2 3 Palustriella commutata E0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 1 Heliosperma veselskyi subsp. veselskyi E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 1 PcSp Physoplexido comosae-Saxifragion petraeae Campanula cespitosa E1 . . . . . . . . . . . . . . . . + . . . . . + . . . . . + + + + + . . . 11 15 Athamanta turbith E1 . . . . . . . r . . . . . . . . r . + . . . . . . . . . . . . . . . . . 4 5 Hieracium porrifolium E1 . . . . . . . . . . . . . . . . . . . . . . + . r . . . + . . . . . . . 4 5 PC Potentilletalia caulescentis Potentilla caulescens E1 . r . . . . . r . . 1 . . . . . . . . . . . . . . . . . + . . . . + . . 7 10 Kernera saxatilis E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + + 1 + . . 5 7 Primula auricula E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 1 AT Asplenietea trichomanis Asplenium ruta-muraria E1 + + + 1 1 + + + + + 1 + + + + + r . . + + 1 1 . . + + 1 . . + + . + 1 + 49 67 Asplenium trichomanes E1 1 + + 1 1 1 1 . + + 1 + . + + . . + . . . . . 1 + 1 1 1 + + . . . 1 1 + 41 56 Hieracium glaucum E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 2 3 TR Thlaspietea rotundifolii Hieracium bifidum E1 . + + . + + . . . . . . + 1 + + r + . . . . + . + 1 + r . + . + + + . . 35 48 Gymnocarpium robertianum E1 . . . . + . . . . . . r . . + . . . . . . + . . . . . . . r r . . . . . 9 12 Adenostyles glabra E1 . . . . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . . . 2 3 Aquilegia iulia E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 1 Dryopteris villarii E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 1 MC Montio-Cardaminetea Conocephalum conicum E0 . . + . . . . . . . . . . . . . . . . . . . . 1 + . . . . . 1 + . + + . 18 25 Distichium capillaceum E0 + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 5 7 Ditrichum flexicaule E0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 1 Campylium stellatum E0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . 1 1 Cololejeunea calcarea E0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . 1 1 ES Elyno-Seslerietea Sesleria caerulea E1 . r . . . . + + 1 + 1 . 1 + r . 1 1 + 1 2 1 + . . . . + . . 1 + 1 1 . . 39 53 Carex mucronata E1 . . . + . . + . . . . . . . . + + . + . . . . . . . . . + . . . . . . . 10 14 Betonica alopecuros E1 . . . . . . . . . . . . . . . . . . r . . . . . . . . . + . . . . + . . 4 5 Carduus crassifolius E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 2 3 Carex ferruginea E1 . . . . . . . . . . . . . . . . r r . . . . . . . . . . . . . . . . . . 2 3 Gentiana clusii E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 1 Laserpitium peucedanoides E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 1 FB Festuco-Brometea Carex humilis E1 . . . . . . . r . . . . + . . . . . . . . . . . . . . . . . . . . . . + 3 4 Buphthalmum salicifolium E1 . r . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 2 3 Galium lucidum E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 1 Brachypodium rupestre E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 1 MA Molinio-Arrhenatheretea Angelica sylvestris E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 3 4 Caltha palustris E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 1 T araxacum x Ruderalia E1 . . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . . . . . 1 1 BA Betulo-Alnetea Salix appendiculata E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . r . . . . . . 5 7 Salix glabra E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 2 3 MuA Mulgedio-Aconitetea Senecio ovatus E1 . . . . . . . . . . . . . . . . . . . . . . . r . . . + . . . . . . . . 4 5 Ranunculus platanifolius E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 1 Aconitum degenii subsp. paniculatum E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 1 Saxifraga rotundifolia E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 1 TG Trifolio-Geranietea Laserpitium siler E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 2 3 Viola hirta E1 . . . . . . . . . . . . . . . . r . . . . . . . . . . . . . . . . . . . 2 3 19/2 • 2020, 155–231 205 Igor Dakskobler & Andrej Martinčič Plant communities of moist rock crevices with endemic Primula carniolica in the (sub)montane belt of western Slovenia Number of relevé (Zaporedna številka popisa) 38 39 40 41 42 43 44 45 46 47 48 49 50 51 52 53 54 55 56 57 58 59 60 61 62 63 64 65 66 67 68 69 70 71 72 73 Pr. Fr. Hymenostylium recurvirostre E0 . . . . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . . . 6 8 Moehringia muscosa E1 + . . . . . . . + . . . . . . . . . . . . . . . . + . . . . + + . . . . 6 8 Primula x venusta E1 . r . . . . . r . . . . . . . . . . . . . . . . . . . . . . . . . . . . 3 4 Pellia endiviifolia E0 . . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . . . . . 2 3 Selaginella helvetica E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . + + . . . . . . 2 3 Palustriella commutata E0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 1 Heliosperma veselskyi subsp. veselskyi E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 1 PcSp Physoplexido comosae-Saxifragion petraeae Campanula cespitosa E1 . . . . . . . . . . . . . . . . + . . . . . + . . . . . + + + + + . . . 11 15 Athamanta turbith E1 . . . . . . . r . . . . . . . . r . + . . . . . . . . . . . . . . . . . 4 5 Hieracium porrifolium E1 . . . . . . . . . . . . . . . . . . . . . . + . r . . . + . . . . . . . 4 5 PC Potentilletalia caulescentis Potentilla caulescens E1 . r . . . . . r . . 1 . . . . . . . . . . . . . . . . . + . . . . + . . 7 10 Kernera saxatilis E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + + 1 + . . 5 7 Primula auricula E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 1 AT Asplenietea trichomanis Asplenium ruta-muraria E1 + + + 1 1 + + + + + 1 + + + + + r . . + + 1 1 . . + + 1 . . + + . + 1 + 49 67 Asplenium trichomanes E1 1 + + 1 1 1 1 . + + 1 + . + + . . + . . . . . 1 + 1 1 1 + + . . . 1 1 + 41 56 Hieracium glaucum E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 2 3 TR Thlaspietea rotundifolii Hieracium bifidum E1 . + + . + + . . . . . . + 1 + + r + . . . . + . + 1 + r . + . + + + . . 35 48 Gymnocarpium robertianum E1 . . . . + . . . . . . r . . + . . . . . . + . . . . . . . r r . . . . . 9 12 Adenostyles glabra E1 . . . . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . . . 2 3 Aquilegia iulia E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 1 Dryopteris villarii E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 1 MC Montio-Cardaminetea Conocephalum conicum E0 . . + . . . . . . . . . . . . . . . . . . . . 1 + . . . . . 1 + . + + . 18 25 Distichium capillaceum E0 + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 5 7 Ditrichum flexicaule E0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 1 Campylium stellatum E0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . 1 1 Cololejeunea calcarea E0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . 1 1 ES Elyno-Seslerietea Sesleria caerulea E1 . r . . . . + + 1 + 1 . 1 + r . 1 1 + 1 2 1 + . . . . + . . 1 + 1 1 . . 39 53 Carex mucronata E1 . . . + . . + . . . . . . . . + + . + . . . . . . . . . + . . . . . . . 10 14 Betonica alopecuros E1 . . . . . . . . . . . . . . . . . . r . . . . . . . . . + . . . . + . . 4 5 Carduus crassifolius E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 2 3 Carex ferruginea E1 . . . . . . . . . . . . . . . . r r . . . . . . . . . . . . . . . . . . 2 3 Gentiana clusii E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 1 Laserpitium peucedanoides E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 1 FB Festuco-Brometea Carex humilis E1 . . . . . . . r . . . . + . . . . . . . . . . . . . . . . . . . . . . + 3 4 Buphthalmum salicifolium E1 . r . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 2 3 Galium lucidum E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 1 Brachypodium rupestre E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 1 MA Molinio-Arrhenatheretea Angelica sylvestris E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 3 4 Caltha palustris E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 1 T araxacum x Ruderalia E1 . . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . . . . . 1 1 BA Betulo-Alnetea Salix appendiculata E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . r . . . . . . 5 7 Salix glabra E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 2 3 MuA Mulgedio-Aconitetea Senecio ovatus E1 . . . . . . . . . . . . . . . . . . . . . . . r . . . + . . . . . . . . 4 5 Ranunculus platanifolius E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 1 Aconitum degenii subsp. paniculatum E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 1 Saxifraga rotundifolia E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 1 TG Trifolio-Geranietea Laserpitium siler E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 2 3 Viola hirta E1 . . . . . . . . . . . . . . . . r . . . . . . . . . . . . . . . . . . . 2 3 19/2 • 2020, 155–231 206 Igor Dakskobler & Andrej Martinčič Plant communities of moist rock crevices with endemic Primula carniolica in the (sub)montane belt of western Slovenia Number of relevé (Zaporedna številka popisa) 38 39 40 41 42 43 44 45 46 47 48 49 50 51 52 53 54 55 56 57 58 59 60 61 62 63 64 65 66 67 68 69 70 71 72 73 Pr. Fr. Anthericum ramosum E1 . . . . . . . . . . . . . . . . . . r . . . . . . . . . . . . . . . . . 1 1 Laserpitium latifolium E1 . . . . . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . . 1 1 Thalictrum minus E1 . . . . . . . . . . . . . . . . . . . . . . . . . r . . . . . . . . . . 1 1 EA Epilobietea angustifolii Rubus idaeus E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 1 EP Erico-Pinetea Calamagrostis varia E1 + + . . . . . . . . . + . . . . . + . . . . . . + + . + + + . + . . . . 23 32 Rhododendron hirsutum E1 . + r r . 1 . . . . r . . r . . . + + + 1 . . . . . . . . + . . . . . . 19 26 Erica carnea E1 . . . . . . . . . . . r . . . . . . r . + . + . . . . . + r . . . . . . 10 14 Polygala chamaebuxus E1 . . . . . . . . . . . . . . . . . . . . r . . . . . . . . . . . . . . . 4 5 Carex ornithopoda E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 4 5 Rubus saxatilis E1 . . . . . . . . . . . . . . + . . . . . . . . . . . . . . r . . . . . . 4 5 Carex alba E1 . . . . . . . . . . . . . r + + . . . . . . . . . . . . . . . . . . . . 3 4 Cirsium erisithales E1 . . . . . . . . . . . . . . . + . + . . . . . . . . . . . . . . . . . . 2 3 Leontodon incanus E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 1 Molinia arundinacea E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 1 Amelanchier ovalis E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 1 Aster amellus E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 1 Rhodothamnus chamaecistus E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 1 Aquilegia nigricans E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 1 Pinus sylvestris E2a . . . . . . . . . . . . . . . . . . . . . . . . . . . . r . . . . . . . 1 1 VP Vaccinio-Pinetea Veronica urticifolia E1 + . . . . . . . . . . . . + . . . . . . . . . r . . 1 + . + . . . . + . 13 18 Homogyne sylvestris E1 . . . + . r . . . . . . . . . . . r . + . . . + . . . . . + . . . . . . 11 15 Solidago virgaurea E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 3 4 Hieracium murorum E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 3 4 Clematis alpina E1 . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . . . 3 4 Gentiana asclepiadea E1 . . . . + . . . . . . . . . . . . . . . . . . r . . . . . . . . . . . . 2 3 Oxalis acetosella E1 . . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . . . r . 2 3 Picea abies E1 . . . . . . . . . . . . . . . . . . . . . . . . . . + . . r . . . . . . 2 3 Polystichum lonchitis E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 1 Aposeris foetida E1 . . . . . . . . . . . r . . . . . . . . . . . . . . . . . . . . . . . . 1 1 Rosa pendulina E1 . . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . . . . . 1 1 AF Aremonio-Fagion Cyclamen purpurascens E1 . . . . . . . . . . . + . r . + . . . . + + + + . r . r r + . + + + . . 17 23 Helleborus niger E1 . . . . . . . . . . . . . . . . + . . + . . . . . . + . r r . . . . . . 5 7 Hemerocallis lilioasphodelus E1 . . . . . . . . . . . . . . . . r . . . . . . . . . . . . . . . . . . . 3 4 Potentilla carniolica E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . 3 4 Cardamine trifolia E1 . . . . . . . . . . . . . . . . . . . . . . . r . . . . . . . . . . r . 3 4 Primula vulgaris E1 . . . . + . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . 3 4 Anemone trifolia E1 . . . . . . . . . . . . . . . . . . . + . . . r . . . . + . . . . . . . 3 4 Euphorbia carniolica E1 . . . . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . . . 2 3 Lamium orvala E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 1 Rhamnus fallax E1 . . . . . . . . . . . . . . . . . r . . . . . . . . . . . . . . . . . . 1 1 Laserpitium krapfii E1 . . . . . . . . . . . . . . . . . . . . . . . . r . . . . . . . . . . . 1 1 Omphalodes verna E1 . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . . 1 1 Knautia drymeia E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . 1 1 TA Tilio-Acerion Aruncus dioicus E1 . . . . . . . . . . . r . . . . . . . . . . . + . . . . . r . . . . . . 8 11 Polystichum aculeatum E1 . . . . . . . . . . . . . . . . . . . . . . . + . r + . . . . . . . . . 6 8 Phyllitis scolopendrium E1 . . . . . . . . . . . . . . . . . . . . . . . r . r . . . . + + . + . . 6 8 Acer pseudoplatanus E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . 5 7 Geranium robertianum E1 . . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . + . . . . 4 5 T ephroseris pseudocrispa E1 . . . . . . . . . . . . . . . . . . . . . . . r . . . . . . . . . . . . 3 4 Thalictrum aquilegiifolium E1 . . . . . . . . . . . . . . . . . . . . . . . . . . r r . . . . . . . . 3 4 Ulmus glabra E2a . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 1 Acer platanoides E2a . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 1 19/2 • 2020, 155–231 207 Igor Dakskobler & Andrej Martinčič Plant communities of moist rock crevices with endemic Primula carniolica in the (sub)montane belt of western Slovenia Number of relevé (Zaporedna številka popisa) 38 39 40 41 42 43 44 45 46 47 48 49 50 51 52 53 54 55 56 57 58 59 60 61 62 63 64 65 66 67 68 69 70 71 72 73 Pr. Fr. Anthericum ramosum E1 . . . . . . . . . . . . . . . . . . r . . . . . . . . . . . . . . . . . 1 1 Laserpitium latifolium E1 . . . . . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . . 1 1 Thalictrum minus E1 . . . . . . . . . . . . . . . . . . . . . . . . . r . . . . . . . . . . 1 1 EA Epilobietea angustifolii Rubus idaeus E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 1 EP Erico-Pinetea Calamagrostis varia E1 + + . . . . . . . . . + . . . . . + . . . . . . + + . + + + . + . . . . 23 32 Rhododendron hirsutum E1 . + r r . 1 . . . . r . . r . . . + + + 1 . . . . . . . . + . . . . . . 19 26 Erica carnea E1 . . . . . . . . . . . r . . . . . . r . + . + . . . . . + r . . . . . . 10 14 Polygala chamaebuxus E1 . . . . . . . . . . . . . . . . . . . . r . . . . . . . . . . . . . . . 4 5 Carex ornithopoda E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 4 5 Rubus saxatilis E1 . . . . . . . . . . . . . . + . . . . . . . . . . . . . . r . . . . . . 4 5 Carex alba E1 . . . . . . . . . . . . . r + + . . . . . . . . . . . . . . . . . . . . 3 4 Cirsium erisithales E1 . . . . . . . . . . . . . . . + . + . . . . . . . . . . . . . . . . . . 2 3 Leontodon incanus E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 1 Molinia arundinacea E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 1 Amelanchier ovalis E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 1 Aster amellus E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 1 Rhodothamnus chamaecistus E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 1 Aquilegia nigricans E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 1 Pinus sylvestris E2a . . . . . . . . . . . . . . . . . . . . . . . . . . . . r . . . . . . . 1 1 VP Vaccinio-Pinetea Veronica urticifolia E1 + . . . . . . . . . . . . + . . . . . . . . . r . . 1 + . + . . . . + . 13 18 Homogyne sylvestris E1 . . . + . r . . . . . . . . . . . r . + . . . + . . . . . + . . . . . . 11 15 Solidago virgaurea E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 3 4 Hieracium murorum E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 3 4 Clematis alpina E1 . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . . . 3 4 Gentiana asclepiadea E1 . . . . + . . . . . . . . . . . . . . . . . . r . . . . . . . . . . . . 2 3 Oxalis acetosella E1 . . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . . . r . 2 3 Picea abies E1 . . . . . . . . . . . . . . . . . . . . . . . . . . + . . r . . . . . . 2 3 Polystichum lonchitis E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 1 Aposeris foetida E1 . . . . . . . . . . . r . . . . . . . . . . . . . . . . . . . . . . . . 1 1 Rosa pendulina E1 . . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . . . . . 1 1 AF Aremonio-Fagion Cyclamen purpurascens E1 . . . . . . . . . . . + . r . + . . . . + + + + . r . r r + . + + + . . 17 23 Helleborus niger E1 . . . . . . . . . . . . . . . . + . . + . . . . . . + . r r . . . . . . 5 7 Hemerocallis lilioasphodelus E1 . . . . . . . . . . . . . . . . r . . . . . . . . . . . . . . . . . . . 3 4 Potentilla carniolica E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . 3 4 Cardamine trifolia E1 . . . . . . . . . . . . . . . . . . . . . . . r . . . . . . . . . . r . 3 4 Primula vulgaris E1 . . . . + . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . 3 4 Anemone trifolia E1 . . . . . . . . . . . . . . . . . . . + . . . r . . . . + . . . . . . . 3 4 Euphorbia carniolica E1 . . . . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . . . 2 3 Lamium orvala E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 1 Rhamnus fallax E1 . . . . . . . . . . . . . . . . . r . . . . . . . . . . . . . . . . . . 1 1 Laserpitium krapfii E1 . . . . . . . . . . . . . . . . . . . . . . . . r . . . . . . . . . . . 1 1 Omphalodes verna E1 . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . . 1 1 Knautia drymeia E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . 1 1 TA Tilio-Acerion Aruncus dioicus E1 . . . . . . . . . . . r . . . . . . . . . . . + . . . . . r . . . . . . 8 11 Polystichum aculeatum E1 . . . . . . . . . . . . . . . . . . . . . . . + . r + . . . . . . . . . 6 8 Phyllitis scolopendrium E1 . . . . . . . . . . . . . . . . . . . . . . . r . r . . . . + + . + . . 6 8 Acer pseudoplatanus E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . 5 7 Geranium robertianum E1 . . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . + . . . . 4 5 T ephroseris pseudocrispa E1 . . . . . . . . . . . . . . . . . . . . . . . r . . . . . . . . . . . . 3 4 Thalictrum aquilegiifolium E1 . . . . . . . . . . . . . . . . . . . . . . . . . . r r . . . . . . . . 3 4 Ulmus glabra E2a . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 1 Acer platanoides E2a . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 1 19/2 • 2020, 155–231 208 Igor Dakskobler & Andrej Martinčič Plant communities of moist rock crevices with endemic Primula carniolica in the (sub)montane belt of western Slovenia Number of relevé (Zaporedna številka popisa) 38 39 40 41 42 43 44 45 46 47 48 49 50 51 52 53 54 55 56 57 58 59 60 61 62 63 64 65 66 67 68 69 70 71 72 73 Pr. Fr. FS Fagetalia sylvaticae Salvia glutinosa E1 . . . . + . . . . . . . . r . . . r . . . . r . . r + + . . . . . . . . 10 14 Galeobdolon flavidum E1 . . . . + . . . . . . r . . . . . . . . + . . + . . . . . . r . . . r . 8 11 Mercurialis perennis E1 . . . . . . . . . . . . . r + . . . . . + . . + . r . . . . . . . . . . 7 10 Mycelis muralis E1 . . + . . . . . . . . . . . . . . . . . . . . . . . . . . . r . + . . . 6 8 Galium laevigatum E1 . . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . . . . . 5 7 Fagus sylvatica E1 . . . . . . . . . . . . . . + . . . . . . . . . . . + . . + . . . + . . 5 7 Melica nutans E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . r . 2 3 Symphytum tuberosum E1 . . . . + . . . . . . . . . . . . . . . . . . . . . . r . . . . . . . . 2 3 Campanula trachelium E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 1 Cardamine pentaphyllos E1 . . . . + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 1 Asarum europaeum subsp. caucasicum E1 . . . . . . . . . . . . . . . . . . . . . . . r . . . . . . . . . . . . 1 1 Petasites albus E1 . . . . . . . . . . . . . . . . . . . . . . . r . . . . . . . . . . . . 1 1 Sambucus nigra E2a . . . . . . . . . . . . . . . . . . . . . . . r . . . . . . . . . . . . 1 1 Lathyrus vernus E1 . . . . . . . . . . . . . . . . . . . . . . . . . r . . . . . . . . . . 1 1 QP Quercetalia pubescenti-petraeae Fraxinus ornus E1 . . . . . . . . . . r . . . + . . + . . . . + . . . . . . . . . . . . . 11 15 Ostrya carpinifolia E1 . . . . . . . . . . . . r . + . . . . . . . . . . . + . . . . . . . . . 7 10 Mercurialis ovata E1 . . . . . . . r . . . . . . . . + . . . . . . . . . . . . . + + + . . . 5 7 Clematis recta E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 1 Euonymus verrucosa E2a . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 1 QF Querco-Fagetea Carex digitata E1 . . . . . + . . . . . + + + + . . + + . + + . + + . + + . . . + r 1 + . 28 38 Hepatica nobilis E1 . r . . . + . . . . . r . + + . r . . . . . . . . + + . . . . . . . . . 16 22 Hedera helix E1 . . . . . . . . . . . . . . r . . . . . . . . + . . . . . . . . . . . . 6 8 Veratrum nigrum E1 . . . . . . . . . . . . . . . . . . . . + . . . . . r r . . . . . . . . 4 5 Acer campestre E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . 1 1 Clematis vitalba E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 1 Quercus petraea E1 . . . . . . . . . . . . . . . . . . . . . . r . . . . . . . . . . . . . 1 1 Cardamine impatiens E1 . . . . . . . . . . . . . . . . . . . . . . . r . . . . . . . . . . . . 1 1 Lonicera xylosteum E1 . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . . 1 1 ML Mosses and lichens (Mahovi in lišaji) Neckera crispa E0 1 + + + 2 . 1 1 1 1 1 2 3 3 2 3 1 2 2 2 2 + 1 2 + 1 1 . 2 + 2 3 + 2 2 1 53 73 Ctenidium molluscum E0 . + 1 + + . 1 1 1 1 2 2 1 2 2 1 2 2 + + . . . 2 1 2 1 2 1 2 . + . . . . 42 58 Tortella tortuosa E0 + . . + + + . . . 1 . . . . . . . . . . . . . . 1 + . . . 2 + . . . . . 29 40 Mnium thomsonii E0 . . . . . . + + . + . . . . . . . . . . . . . . . . . + . . . . . . . . 6 8 Schistidium apocarpum E0 . . . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . . . . 4 5 Plagiochila porreloides E0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 3 4 Encalypta streptocarpa E0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 3 4 Leiocolea collaris E0 . . . . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . . . 2 3 Marchantia polymorpha E0 . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . 2 3 Bryum capillare E0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 2 3 Pedinophyllum interruptum E0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 2 3 Mnium stellare E0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 2 3 Isothecium alopecuroides E0 . . . 1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 2 3 Lejeunea cavifolia E0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 1 Myurella sibirica E0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 1 Neckera complanata E0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 1 Didymodon ferrugineus E0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 1 Trichostomum brachydontium E0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 1 Myurella julacea E0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 1 Hypnum cupressiforme var. resupinatum E0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 1 Lichenes div. E0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 1 Lophozia sp. E0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 1 Dichodontium pellucidum subsp. flavescens E0 . . . . . . . . . . . . . . . . . . . 1 . . . . . . . . . . . . . . . . 1 1 Brachythecium rutabulum E0 . . . . . . . . . . . . . . . . . . . 1 . . . . . . . . . . . . . . . . 1 1 Plagiomnium rostratum E0 . . . . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . . . 1 1 Mnium marginatum E0 . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . . . . 1 1 Plagiobryum zierii E0 . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . . . . 1 1 Legend / Legenda: D Dolomite – dolomit L Limestone – apnenec Li Lithosol – kamnišče Fr. Frequency in % – frekvenca v % 19/2 • 2020, 155–231 209 Igor Dakskobler & Andrej Martinčič Plant communities of moist rock crevices with endemic Primula carniolica in the (sub)montane belt of western Slovenia Pr. Presence (number of relevés in which the species is presented) – število popisov, v katerih se pojavlja vrsta Number of relevé (Zaporedna številka popisa) 38 39 40 41 42 43 44 45 46 47 48 49 50 51 52 53 54 55 56 57 58 59 60 61 62 63 64 65 66 67 68 69 70 71 72 73 Pr. Fr. FS Fagetalia sylvaticae Salvia glutinosa E1 . . . . + . . . . . . . . r . . . r . . . . r . . r + + . . . . . . . . 10 14 Galeobdolon flavidum E1 . . . . + . . . . . . r . . . . . . . . + . . + . . . . . . r . . . r . 8 11 Mercurialis perennis E1 . . . . . . . . . . . . . r + . . . . . + . . + . r . . . . . . . . . . 7 10 Mycelis muralis E1 . . + . . . . . . . . . . . . . . . . . . . . . . . . . . . r . + . . . 6 8 Galium laevigatum E1 . . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . . . . . 5 7 Fagus sylvatica E1 . . . . . . . . . . . . . . + . . . . . . . . . . . + . . + . . . + . . 5 7 Melica nutans E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . r . 2 3 Symphytum tuberosum E1 . . . . + . . . . . . . . . . . . . . . . . . . . . . r . . . . . . . . 2 3 Campanula trachelium E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 1 Cardamine pentaphyllos E1 . . . . + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 1 Asarum europaeum subsp. caucasicum E1 . . . . . . . . . . . . . . . . . . . . . . . r . . . . . . . . . . . . 1 1 Petasites albus E1 . . . . . . . . . . . . . . . . . . . . . . . r . . . . . . . . . . . . 1 1 Sambucus nigra E2a . . . . . . . . . . . . . . . . . . . . . . . r . . . . . . . . . . . . 1 1 Lathyrus vernus E1 . . . . . . . . . . . . . . . . . . . . . . . . . r . . . . . . . . . . 1 1 QP Quercetalia pubescenti-petraeae Fraxinus ornus E1 . . . . . . . . . . r . . . + . . + . . . . + . . . . . . . . . . . . . 11 15 Ostrya carpinifolia E1 . . . . . . . . . . . . r . + . . . . . . . . . . . + . . . . . . . . . 7 10 Mercurialis ovata E1 . . . . . . . r . . . . . . . . + . . . . . . . . . . . . . + + + . . . 5 7 Clematis recta E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 1 Euonymus verrucosa E2a . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 1 QF Querco-Fagetea Carex digitata E1 . . . . . + . . . . . + + + + . . + + . + + . + + . + + . . . + r 1 + . 28 38 Hepatica nobilis E1 . r . . . + . . . . . r . + + . r . . . . . . . . + + . . . . . . . . . 16 22 Hedera helix E1 . . . . . . . . . . . . . . r . . . . . . . . + . . . . . . . . . . . . 6 8 Veratrum nigrum E1 . . . . . . . . . . . . . . . . . . . . + . . . . . r r . . . . . . . . 4 5 Acer campestre E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . 1 1 Clematis vitalba E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 1 Quercus petraea E1 . . . . . . . . . . . . . . . . . . . . . . r . . . . . . . . . . . . . 1 1 Cardamine impatiens E1 . . . . . . . . . . . . . . . . . . . . . . . r . . . . . . . . . . . . 1 1 Lonicera xylosteum E1 . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . . 1 1 ML Mosses and lichens (Mahovi in lišaji) Neckera crispa E0 1 + + + 2 . 1 1 1 1 1 2 3 3 2 3 1 2 2 2 2 + 1 2 + 1 1 . 2 + 2 3 + 2 2 1 53 73 Ctenidium molluscum E0 . + 1 + + . 1 1 1 1 2 2 1 2 2 1 2 2 + + . . . 2 1 2 1 2 1 2 . + . . . . 42 58 Tortella tortuosa E0 + . . + + + . . . 1 . . . . . . . . . . . . . . 1 + . . . 2 + . . . . . 29 40 Mnium thomsonii E0 . . . . . . + + . + . . . . . . . . . . . . . . . . . + . . . . . . . . 6 8 Schistidium apocarpum E0 . . . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . . . . 4 5 Plagiochila porreloides E0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 3 4 Encalypta streptocarpa E0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 3 4 Leiocolea collaris E0 . . . . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . . . 2 3 Marchantia polymorpha E0 . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . 2 3 Bryum capillare E0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 2 3 Pedinophyllum interruptum E0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 2 3 Mnium stellare E0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 2 3 Isothecium alopecuroides E0 . . . 1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 2 3 Lejeunea cavifolia E0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 1 Myurella sibirica E0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 1 Neckera complanata E0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 1 Didymodon ferrugineus E0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 1 Trichostomum brachydontium E0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 1 Myurella julacea E0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 1 Hypnum cupressiforme var. resupinatum E0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 1 Lichenes div. E0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 1 Lophozia sp. E0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 1 Dichodontium pellucidum subsp. flavescens E0 . . . . . . . . . . . . . . . . . . . 1 . . . . . . . . . . . . . . . . 1 1 Brachythecium rutabulum E0 . . . . . . . . . . . . . . . . . . . 1 . . . . . . . . . . . . . . . . 1 1 Plagiomnium rostratum E0 . . . . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . . . 1 1 Mnium marginatum E0 . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . . . . 1 1 Plagiobryum zierii E0 . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . . . . 1 1 19/2 • 2020, 155–231 210 Igor Dakskobler & Andrej Martinčič Plant communities of moist rock crevices with endemic Primula carniolica in the (sub)montane belt of western Slovenia Table 6 (Tabela 6): Phyteumato-Primuletum carniolicae astrantietosum carniolicae. Number of relevé (Zaporedna številka popisa) 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 Pr. Fr. Database number of relevé (Delovna številka popisa) 230633 241567 241568 234480 235012 246868 259052 259053 242177 234555 271153 271155 241563 246867 268197 268209 273971 268206 268207 268204 273912 273973 268212 275148 278093 273964 278100 Elevation in m (Nadmorska višina v m) 1190 810 810 1110 215 1250 1430 1440 1380 810 420 430 800 1250 890 845 775 845 890 850 590 755 840 430 470 780 460 Aspect (Lega) NW SE NW W NE NE N NW NE NE N NW NE N E SE N E NW W W NE NW SE SW NE N Slope in degrees (Nagib v stopinjah) 90 90 90 95 90 80 90 80 80 80 90 95 80 80 95 80 85 80 80 90 95 95 85 90 90 80 80 Parent material (Matična podlaga) DL D D D D D L L L D D D D D D D D D D D D D D D D D D Soil (Tla) Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Stoniness in % (Kamnitost v %) 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 Cover of herb layer in % (Zastiranje zeliščne plasti v %): E1 20 20 30 20 30 30 30 20 15 40 20 35 30 35 20 30 30 40 60 30 15 20 35 30 30 60 60 Cover of moss layer in % (Zastiranje mahovne plasti v %) E0 5 5 5 5 30 20 20 10 5 10 10 10 10 20 10 10 20 30 30 20 5 20 20 10 30 30 70 Number of species (Število vrst) 16 17 15 15 15 23 24 18 11 22 18 16 24 22 16 18 17 18 24 21 18 17 22 14 17 17 20 Relevé area (Velikost popisne ploskve) m 2 10 10 10 10 5 10 5 5 5 5 10 10 10 10 5 5 20 5 5 5 20 10 5 15 15 10 15 Date of taking relevé (Datum popisa) 7/3/2000 5/5/2011 5/5/2011 7/20/2004 4/25/2005 8/13/2003 6/22/2015 6/22/2015 8/4/1998 5/19/2004 5/3/2018 5/3/2018 5/5/2011 8/13/2003 8/30/2017 8/30/2017 7/2/2018 8/30/2017 8/30/2017 8/30/2017 7/25/2018 7/2/2018 8/30/2017 4/22/2019 12,6.2019 7/2/2018 12,6.2019 Locality (Nahajališče) Bukov vrh Bukov vrh - Brusova grapa Bukov vrh - Brusova grapa Hudournik Gačnik Sončni rob Mali Golak Mali Golak Srednji Golak Gorenja T rebuša - Lazna - Špičast rob Gorenja T rebuša - Makčeva grapa Gorenja T rebuša - Makčeva grapa Mali Govci Sončni rob Gačnik Gačnik - Brdo Gačnik Gačnik Gačnik Gačnik Gačnik - Medved Gačnik Gačnik-Brdo Gačnik Gačnik Gačnik Gačnik Quadrant (Kvadrant) 0049/1 9949/3 9949/3 9949/1 9949/1 0049/1 0049/1 0049/1 0049/1 9948/4 9949/3 9949/3 9949/3 0049/1 9949/3 9949/3 9949/1 9949/1 9949/3 9949/1 9949/1 9949/1 9949/3 9949/1 9949/1 9949/1 9949/1 Coordinate GK Y (D-48) m 413635 413790 413770 413757 410107 414653 412426 412435 413074 408748 410066 410105 413679 414658 412494 412459 412108 412495 412577 412503 411222 412050 412457 410992 411150 412218 411121 Coordinate GK X (D-48) m 5094650 5095679 5095671 5102207 5104236 5094330 5093455 5093412 5093352 5099263 5100660 5100659 5095879 5094216 5101188 5100916 5101912 5101431 5100830 5101371 5102794 5101938 5100950 5103416 5103138 5101768 5103184 Diagnostic species of the association (Diagnostične vrste asociacije) Pr. Fr. AP Primula carniolica E1 + 1 1 1 1 1 1 1 + 2 1 2 + 2 2 2 1 1 1 1 + 1 1 2 1 + 1 27 100 AP Paederota lutea E1 1 2 1 1 1 2 2 2 1 1 1 2 2 2 2 2 2 1 2 1 1 1 1 1 1 2 1 27 100 PcSp Phyteuma scheuchzeri subsp. columnae E1 + + + . + + + + . + + + 1 1 + + + + 1 + . + + . + . + 22 81 AP Orthothecium rufescens E0 . . . + 2 2 2 1 1 1 . 1 + . . 1 1 + 1 1 1 + + . . 1 1 19 70 AP Carex brachystachys E1 . . . . . . + . . + . . . + + + + + 1 + + + . . . + + 13 48 AP Valeriana tripteris E1 . . . . . . + . . + + + + + + . . . . . . + + . . . + 10 37 Differential species of the subassociation (Razlikovalnice subasociacije) AP Viola biflora E1 + + + + . + 1 + + 1 + + 1 + + 1 + 1 + r + . + r + + + 25 93 AP Valeriana saxatilis E1 + + + + 1 + 1 1 + . . + r r + 1 + 1 1 1 1 + . . 1 1 . 22 81 AP Astrantia carniolica E1 r . . . 1 + . . . . + + + . + r 1 + 2 r . + 2 1 1 3 1 18 67 AP Cystopteridion s. lat. (Astrantio-Paederotion luteae nom. prov.) Aster bellidiastrum E1 + + + . 1 1 + + + + r . + + + 1 1 + . . + + + 1 + 1 1 23 85 Asplenium viride E1 . . . . . . + + . + 1 1 + + + r + . + 1 + . 1 + + . . 16 59 Pinguicula alpina E1 r + + r + + + + r . . . . . . . . . + + . . . . . . . 11 41 Tofieldia calyculata E1 . + + . 1 + + . . . . . . . . . . + + . . . . + . + + 10 37 Cystopteris fragilis E1 + . . + . . . . + + . . + . . . . . . + . . + . . . . 7 26 Fissidens dubius E0 . + . . . . + . . . . . . . . . . . 1 . . . + 1 + . . 6 22 19/2 • 2020, 155–231 211 Igor Dakskobler & Andrej Martinčič Plant communities of moist rock crevices with endemic Primula carniolica in the (sub)montane belt of western Slovenia Number of relevé (Zaporedna številka popisa) 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 Pr. Fr. Database number of relevé (Delovna številka popisa) 230633 241567 241568 234480 235012 246868 259052 259053 242177 234555 271153 271155 241563 246867 268197 268209 273971 268206 268207 268204 273912 273973 268212 275148 278093 273964 278100 Elevation in m (Nadmorska višina v m) 1190 810 810 1110 215 1250 1430 1440 1380 810 420 430 800 1250 890 845 775 845 890 850 590 755 840 430 470 780 460 Aspect (Lega) NW SE NW W NE NE N NW NE NE N NW NE N E SE N E NW W W NE NW SE SW NE N Slope in degrees (Nagib v stopinjah) 90 90 90 95 90 80 90 80 80 80 90 95 80 80 95 80 85 80 80 90 95 95 85 90 90 80 80 Parent material (Matična podlaga) DL D D D D D L L L D D D D D D D D D D D D D D D D D D Soil (Tla) Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Stoniness in % (Kamnitost v %) 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 Cover of herb layer in % (Zastiranje zeliščne plasti v %): E1 20 20 30 20 30 30 30 20 15 40 20 35 30 35 20 30 30 40 60 30 15 20 35 30 30 60 60 Cover of moss layer in % (Zastiranje mahovne plasti v %) E0 5 5 5 5 30 20 20 10 5 10 10 10 10 20 10 10 20 30 30 20 5 20 20 10 30 30 70 Number of species (Število vrst) 16 17 15 15 15 23 24 18 11 22 18 16 24 22 16 18 17 18 24 21 18 17 22 14 17 17 20 Relevé area (Velikost popisne ploskve) m 2 10 10 10 10 5 10 5 5 5 5 10 10 10 10 5 5 20 5 5 5 20 10 5 15 15 10 15 Date of taking relevé (Datum popisa) 7/3/2000 5/5/2011 5/5/2011 7/20/2004 4/25/2005 8/13/2003 6/22/2015 6/22/2015 8/4/1998 5/19/2004 5/3/2018 5/3/2018 5/5/2011 8/13/2003 8/30/2017 8/30/2017 7/2/2018 8/30/2017 8/30/2017 8/30/2017 7/25/2018 7/2/2018 8/30/2017 4/22/2019 12,6.2019 7/2/2018 12,6.2019 Locality (Nahajališče) Bukov vrh Bukov vrh - Brusova grapa Bukov vrh - Brusova grapa Hudournik Gačnik Sončni rob Mali Golak Mali Golak Srednji Golak Gorenja T rebuša - Lazna - Špičast rob Gorenja T rebuša - Makčeva grapa Gorenja T rebuša - Makčeva grapa Mali Govci Sončni rob Gačnik Gačnik - Brdo Gačnik Gačnik Gačnik Gačnik Gačnik - Medved Gačnik Gačnik-Brdo Gačnik Gačnik Gačnik Gačnik Quadrant (Kvadrant) 0049/1 9949/3 9949/3 9949/1 9949/1 0049/1 0049/1 0049/1 0049/1 9948/4 9949/3 9949/3 9949/3 0049/1 9949/3 9949/3 9949/1 9949/1 9949/3 9949/1 9949/1 9949/1 9949/3 9949/1 9949/1 9949/1 9949/1 Coordinate GK Y (D-48) m 413635 413790 413770 413757 410107 414653 412426 412435 413074 408748 410066 410105 413679 414658 412494 412459 412108 412495 412577 412503 411222 412050 412457 410992 411150 412218 411121 Coordinate GK X (D-48) m 5094650 5095679 5095671 5102207 5104236 5094330 5093455 5093412 5093352 5099263 5100660 5100659 5095879 5094216 5101188 5100916 5101912 5101431 5100830 5101371 5102794 5101938 5100950 5103416 5103138 5101768 5103184 Diagnostic species of the association (Diagnostične vrste asociacije) Pr. Fr. AP Primula carniolica E1 + 1 1 1 1 1 1 1 + 2 1 2 + 2 2 2 1 1 1 1 + 1 1 2 1 + 1 27 100 AP Paederota lutea E1 1 2 1 1 1 2 2 2 1 1 1 2 2 2 2 2 2 1 2 1 1 1 1 1 1 2 1 27 100 PcSp Phyteuma scheuchzeri subsp. columnae E1 + + + . + + + + . + + + 1 1 + + + + 1 + . + + . + . + 22 81 AP Orthothecium rufescens E0 . . . + 2 2 2 1 1 1 . 1 + . . 1 1 + 1 1 1 + + . . 1 1 19 70 AP Carex brachystachys E1 . . . . . . + . . + . . . + + + + + 1 + + + . . . + + 13 48 AP Valeriana tripteris E1 . . . . . . + . . + + + + + + . . . . . . + + . . . + 10 37 Differential species of the subassociation (Razlikovalnice subasociacije) AP Viola biflora E1 + + + + . + 1 + + 1 + + 1 + + 1 + 1 + r + . + r + + + 25 93 AP Valeriana saxatilis E1 + + + + 1 + 1 1 + . . + r r + 1 + 1 1 1 1 + . . 1 1 . 22 81 AP Astrantia carniolica E1 r . . . 1 + . . . . + + + . + r 1 + 2 r . + 2 1 1 3 1 18 67 AP Cystopteridion s. lat. (Astrantio-Paederotion luteae nom. prov.) Aster bellidiastrum E1 + + + . 1 1 + + + + r . + + + 1 1 + . . + + + 1 + 1 1 23 85 Asplenium viride E1 . . . . . . + + . + 1 1 + + + r + . + 1 + . 1 + + . . 16 59 Pinguicula alpina E1 r + + r + + + + r . . . . . . . . . + + . . . . . . . 11 41 Tofieldia calyculata E1 . + + . 1 + + . . . . . . . . . . + + . . . . + . + + 10 37 Cystopteris fragilis E1 + . . + . . . . + + . . + . . . . . . + . . + . . . . 7 26 Fissidens dubius E0 . + . . . . + . . . . . . . . . . . 1 . . . + 1 + . . 6 22 19/2 • 2020, 155–231 212 Igor Dakskobler & Andrej Martinčič Plant communities of moist rock crevices with endemic Primula carniolica in the (sub)montane belt of western Slovenia Number of relevé (Zaporedna številka popisa) 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 Pr. Fr. Soldanella minima E1 . . . + . + + . . . . . . . . . . . . . . . . . . . . 3 11 Cystopteris regia E1 . . . + . . . + . . . . . . . . . . . r . . . . . . . 3 11 Hymenostylium recurvirostre E0 . . . + . 1 . . . + . . . . . . . . . . . . . . . . . 3 11 Pellia endiviifolia E0 . . . . . . . . . . . . . . . . . . . . + . . . 1 1 . 3 11 Barbula crocea E0 . . . . . . . . . . . . . . . . . . . . . 1 . . . 1 . 2 7 Solorina saccata E0 . . . . . + . . . . . . . . . . . . . . . . . . . . . 1 4 Heliosperma pusillum E1 . . . . . . + . . . . . . . . . . . . . . . . . . . . 1 4 Preissia quadrata E0 . . . . . . . . . . . + . . . . . . . . . . . . . . . 1 4 Palustriella commutata E0 . . . . . . . . . . . . . . . . . . . . + . . . . . . 1 4 PcSp Physoplexido comosae-Saxifragion petraeae Campanula cespitosa E1 . . . . . + . . . . . . . . . + . . . . . . . . . . . 2 7 PC Potentilletum caulescentis Potentilla caulescens E1 r . . + . r . . . . . . . . . . . r . . . . . . . . . 4 15 Kernera saxatilis E1 . . . . . . . . . . . . . + . . . . . . . . . . . . . 1 4 AT Asplenietea trichomanis Asplenium ruta-muraria E1 r + . . . . . + . . . . + 1 + . . + + + . + + . . . . 11 41 Asplenium trichomanes E1 . + r + . . . . . + + + + . . . . . . + + 1 . . . . . 10 37 TR Thlapietea rotundifolii Adenostyles glabra E1 . . . . r r + . . . . . . . + r + + . . . . . . . r + 9 33 Hieracium bifidum E1 . + . . + . . . . r + + + . . . . + . . . . . . . . . 7 26 Dryopteris villarii E1 . . . . . . + + . . . . . . . . . . . . . . . . . . . 2 7 Aquilegia iulia E1 . . . . . . . . . . . . . + . . . . . . . . . . . . . 1 4 Gymnocarpium robertianum E1 . . . . . . . . . . . . . . . . + . . . . . . . . . . 1 4 CD Caricetalia davallianae Parnassia palustris E1 . . . . . + . . . . . . . . . . . . . . . . . . . . . 1 4 MC Montio-Cardaminetea Conocephalum conicum E0 . . . . . . + . + 1 1 + . . . . . . + 1 + . + 1 1 1 2 13 48 Campylium stellatum E0 . . . . . . . . + . . . . . . . . . . . . . . . . . . 1 4 Distichium capillaceum E0 . . . . . . . . . . . . . 1 . . . . . . . . . . . . . 1 4 ES Elyno-Seslerietea Sesleria caerulea E1 + + + + + + r . . . . . . 1 + . 1 + + . + + . . + . 1 16 59 Laserpitium peucedanoides E1 . . r . . . + + . . . . . + . . . . . . . . . . . . . 4 15 Carex ferruginea E1 + . . . . . . . . . . . + . . . . . . . . . . . . . + 3 11 Carex mucronata E1 . + r + . . . . . . . . . . . . . . . . . . . . . . . 3 11 Carex firma E1 . . r . . 1 . . . . . . . . . . . . . . . . . . . . . 2 7 Campanula witasekiana E1 . . . . . . + . . . . . . . . . . . . . . . . . . . . 1 4 Phyteuma orbiculare E1 . . . . . . . . . . . . . . . . . . . . . . . . . . r 1 4 TG Trifolio-Geranietea Digitalis grandiflora E1 . . . . . . . . . . . . r . . . . . . . . . . . . . . 1 4 BA Betulo-Alnetea Salix appendiculata E1 . . . . . . . r . . . . . . . . . . + . . . . . . . . 2 7 MuA Mulgedio-Aconitetea Chaerophyllum hirsutum E1 . . . . . . . . . . . . . . . . . . . . . . . r + . . 2 7 Veratrum album E1 . . . . . . r . . . . . . . . . . . . . . . . . . . . 1 4 Crepis paludosa E1 . . . . . . . . . . . . . . . . . . . . . . . r . . . 1 4 EA Epilobietea angustifolii Rubus idaeus E1 . . . . . . . . . . . . . r . . . . . . . . . . . . . 1 4 EP Erico-Pinetea Calamagrostis varia E1 . . . . . + . . . + . . + . . + + . . + + . + . + . . 9 33 Rhododendron hirsutum E1 . r r . . . + + . . . . . + . . . + + + . . . . . r . 9 33 Rhodothamnus chamaecistus E1 + + + . . r . . . . . . . + . . . . . . . . . . . . . 5 19 19/2 • 2020, 155–231 213 Igor Dakskobler & Andrej Martinčič Plant communities of moist rock crevices with endemic Primula carniolica in the (sub)montane belt of western Slovenia Number of relevé (Zaporedna številka popisa) 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 Pr. Fr. Soldanella minima E1 . . . + . + + . . . . . . . . . . . . . . . . . . . . 3 11 Cystopteris regia E1 . . . + . . . + . . . . . . . . . . . r . . . . . . . 3 11 Hymenostylium recurvirostre E0 . . . + . 1 . . . + . . . . . . . . . . . . . . . . . 3 11 Pellia endiviifolia E0 . . . . . . . . . . . . . . . . . . . . + . . . 1 1 . 3 11 Barbula crocea E0 . . . . . . . . . . . . . . . . . . . . . 1 . . . 1 . 2 7 Solorina saccata E0 . . . . . + . . . . . . . . . . . . . . . . . . . . . 1 4 Heliosperma pusillum E1 . . . . . . + . . . . . . . . . . . . . . . . . . . . 1 4 Preissia quadrata E0 . . . . . . . . . . . + . . . . . . . . . . . . . . . 1 4 Palustriella commutata E0 . . . . . . . . . . . . . . . . . . . . + . . . . . . 1 4 PcSp Physoplexido comosae-Saxifragion petraeae Campanula cespitosa E1 . . . . . + . . . . . . . . . + . . . . . . . . . . . 2 7 PC Potentilletum caulescentis Potentilla caulescens E1 r . . + . r . . . . . . . . . . . r . . . . . . . . . 4 15 Kernera saxatilis E1 . . . . . . . . . . . . . + . . . . . . . . . . . . . 1 4 AT Asplenietea trichomanis Asplenium ruta-muraria E1 r + . . . . . + . . . . + 1 + . . + + + . + + . . . . 11 41 Asplenium trichomanes E1 . + r + . . . . . + + + + . . . . . . + + 1 . . . . . 10 37 TR Thlapietea rotundifolii Adenostyles glabra E1 . . . . r r + . . . . . . . + r + + . . . . . . . r + 9 33 Hieracium bifidum E1 . + . . + . . . . r + + + . . . . + . . . . . . . . . 7 26 Dryopteris villarii E1 . . . . . . + + . . . . . . . . . . . . . . . . . . . 2 7 Aquilegia iulia E1 . . . . . . . . . . . . . + . . . . . . . . . . . . . 1 4 Gymnocarpium robertianum E1 . . . . . . . . . . . . . . . . + . . . . . . . . . . 1 4 CD Caricetalia davallianae Parnassia palustris E1 . . . . . + . . . . . . . . . . . . . . . . . . . . . 1 4 MC Montio-Cardaminetea Conocephalum conicum E0 . . . . . . + . + 1 1 + . . . . . . + 1 + . + 1 1 1 2 13 48 Campylium stellatum E0 . . . . . . . . + . . . . . . . . . . . . . . . . . . 1 4 Distichium capillaceum E0 . . . . . . . . . . . . . 1 . . . . . . . . . . . . . 1 4 ES Elyno-Seslerietea Sesleria caerulea E1 + + + + + + r . . . . . . 1 + . 1 + + . + + . . + . 1 16 59 Laserpitium peucedanoides E1 . . r . . . + + . . . . . + . . . . . . . . . . . . . 4 15 Carex ferruginea E1 + . . . . . . . . . . . + . . . . . . . . . . . . . + 3 11 Carex mucronata E1 . + r + . . . . . . . . . . . . . . . . . . . . . . . 3 11 Carex firma E1 . . r . . 1 . . . . . . . . . . . . . . . . . . . . . 2 7 Campanula witasekiana E1 . . . . . . + . . . . . . . . . . . . . . . . . . . . 1 4 Phyteuma orbiculare E1 . . . . . . . . . . . . . . . . . . . . . . . . . . r 1 4 TG Trifolio-Geranietea Digitalis grandiflora E1 . . . . . . . . . . . . r . . . . . . . . . . . . . . 1 4 BA Betulo-Alnetea Salix appendiculata E1 . . . . . . . r . . . . . . . . . . + . . . . . . . . 2 7 MuA Mulgedio-Aconitetea Chaerophyllum hirsutum E1 . . . . . . . . . . . . . . . . . . . . . . . r + . . 2 7 Veratrum album E1 . . . . . . r . . . . . . . . . . . . . . . . . . . . 1 4 Crepis paludosa E1 . . . . . . . . . . . . . . . . . . . . . . . r . . . 1 4 EA Epilobietea angustifolii Rubus idaeus E1 . . . . . . . . . . . . . r . . . . . . . . . . . . . 1 4 EP Erico-Pinetea Calamagrostis varia E1 . . . . . + . . . + . . + . . + + . . + + . + . + . . 9 33 Rhododendron hirsutum E1 . r r . . . + + . . . . . + . . . + + + . . . . . r . 9 33 Rhodothamnus chamaecistus E1 + + + . . r . . . . . . . + . . . . . . . . . . . . . 5 19 19/2 • 2020, 155–231 214 Igor Dakskobler & Andrej Martinčič Plant communities of moist rock crevices with endemic Primula carniolica in the (sub)montane belt of western Slovenia Number of relevé (Zaporedna številka popisa) 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 Pr. Fr. Pinus nigra E1 . r . . . . . . . . . . . . . . . . . . . . . . . . . 1 4 Polygala chamaebuxus E1 . . . . r . . . . . . . . . . . . . . . . . . . . . . 1 4 Erica carnea E1 . . . . r . . . . . . . . . . . . . . . . . . . . . . 1 4 Cirsium erisithales E1 . . . . . . . . . + . . . . . . . . . . . . . . . . . 1 4 Aquilegia nigricans E1 . . . . . . . . . . . . . . . r . . . . . . . . . . . 1 4 Molinia arundinacea E1 . . . . . . . . . . . . . . . . . . . . . . . . . . 1 1 4 VP Vaccinio-Piceetea Homogyne sylvestris E1 . . . . . . . . . + r . . . . . + . r . . . . . . . . 4 15 Clematis alpina E1 . . . . . . . . . . . . . . . . . . + . . + 2 . . + . 4 15 Veronica urticifolia E1 . . . . . . . . . . . . . . . . . . . . . . . . . . 1 1 4 AF Aremonio-Fagion Anemone trifolia E1 . . . . . . . . . + + . . . . . . . . . . . . . . . . 2 7 Cyclamen purpurascens E1 . . . . . . . . . . . . . . + . . . . . . + . . . . . 2 7 Cardamine enneaphyllos E1 . . . . . . . . . . . . r . . . . . . . . . . . . . . 1 4 Cardamine trifolia E1 . . . . . . . . . . . . . . . . . . . . . . . + . . . 1 4 TA Tilio-Acerion Acer pseudoplatanus E1 . . . . . r . . . . . . . . . . . . r . . . . . . . . 2 7 Aruncus dioicus E1 . . . . . . . . . . . . . . . . . . . . . . + . . . . 1 4 Thalictrum aquilegiifolium E1 . . . . . . . . . . . . . . . . . . . . . . . . . . + 1 4 FS Fagetalia sylvaticae Galeobdolon flavidum E1 . . . . . . . . . . + + . . . . . . . + . . . . . . . 3 11 Fagus sylvatica E1 . . . . . . . . . r . . . . . . . . . . . . + . . . . 2 7 Galium laevigatum E1 . . . . . . . . . . . . . . . . + . . . . . . . . . + 2 7 Mycelis muralis E1 . . . . . . . . . . . . . . . . . . . + + . . . . . . 2 7 Salvia glutinosa E1 . . . . . . . . . . . . r . . . . . . . . . . . . . . 1 4 QP Quercetalia pubescentis Fraxinus ornus E1 . . . . . . . . . . + . + . . . . . . . . . . . . . . 2 7 Ostrya carpinifolia E2a . . . . . . . . . . . . + . . . . . . . . . . . . . . 1 4 QF Querco-Fagetea Carex digitata E1 . . . . . . . . . 1 r + . . . r . . r + . + + . + . . 9 33 Hepatica nobilis E1 . . . . . . . . . . . . . . . . . . . . . . 2 + . . . 2 7 Carex umbrosa E1 . . . . r . . . . . . . . . . . . . . . . . . . . . . 1 4 ML Mosses and lichens (Mahovi in lišaji) Ctenidium molluscum E0 + . . . . + . . . + + . 1 1 + 1 1 . 1 . + . + 1 1 . . 14 52 Neckera crispa E0 . . . + . . . . . 1 + 1 1 1 + 1 . 1 + . . . 1 . . . . 11 41 Tortella tortuosa E0 . . . . . . . . + . . . . + . 1 . + 2 + . + 1 . . 1 . 9 33 Brachythecium rutabulum E0 . . . . . . . . . . . . . . . . . . . . . . . . 1 1 2 3 11 Plagiopus oederi E0 + . . . . . . + . . . . . . . . . . . . . . . . . . . 2 7 Bryum sp. E0 . . . . . . 1 . . . . . . . . . . . . . . . . . . . . 1 4 Lescuraea plicata E0 . . . . . . . + . . . . . . . . . . . . . . . . . . . 1 4 Dicranum scoparium E0 . . . . . . . + . . . . . . . . . . . . . . . . . . . 1 4 Marchantia polymorpha E0 . . . . . . . . . . . . + . . . . . . . . . . . . . . 1 4 Schistidium apocarpum E0 . . . . . . . . . . . . . + . . . . . . . . . . . . . 1 4 Pedinophyllum interruptum E0 . . . . . . . . . . . . . . . . . . . . + . . . . . . 1 4 Plagiochila porelloides E0 . . . . . . . . . . . . . . . . . . . . + . . . . . . 1 4 Legend / Legenda D Dolomite – dolomit L Limestone – apnenec Li Lithosol – kamnišče Pr. Presence (number of relevés in which the species is presented) – število popisov, v katerih se pojavlja vrsta Fr. Frequency in % – frekvenca v % 19/2 • 2020, 155–231 215 Igor Dakskobler & Andrej Martinčič Plant communities of moist rock crevices with endemic Primula carniolica in the (sub)montane belt of western Slovenia Number of relevé (Zaporedna številka popisa) 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 Pr. Fr. Pinus nigra E1 . r . . . . . . . . . . . . . . . . . . . . . . . . . 1 4 Polygala chamaebuxus E1 . . . . r . . . . . . . . . . . . . . . . . . . . . . 1 4 Erica carnea E1 . . . . r . . . . . . . . . . . . . . . . . . . . . . 1 4 Cirsium erisithales E1 . . . . . . . . . + . . . . . . . . . . . . . . . . . 1 4 Aquilegia nigricans E1 . . . . . . . . . . . . . . . r . . . . . . . . . . . 1 4 Molinia arundinacea E1 . . . . . . . . . . . . . . . . . . . . . . . . . . 1 1 4 VP Vaccinio-Piceetea Homogyne sylvestris E1 . . . . . . . . . + r . . . . . + . r . . . . . . . . 4 15 Clematis alpina E1 . . . . . . . . . . . . . . . . . . + . . + 2 . . + . 4 15 Veronica urticifolia E1 . . . . . . . . . . . . . . . . . . . . . . . . . . 1 1 4 AF Aremonio-Fagion Anemone trifolia E1 . . . . . . . . . + + . . . . . . . . . . . . . . . . 2 7 Cyclamen purpurascens E1 . . . . . . . . . . . . . . + . . . . . . + . . . . . 2 7 Cardamine enneaphyllos E1 . . . . . . . . . . . . r . . . . . . . . . . . . . . 1 4 Cardamine trifolia E1 . . . . . . . . . . . . . . . . . . . . . . . + . . . 1 4 TA Tilio-Acerion Acer pseudoplatanus E1 . . . . . r . . . . . . . . . . . . r . . . . . . . . 2 7 Aruncus dioicus E1 . . . . . . . . . . . . . . . . . . . . . . + . . . . 1 4 Thalictrum aquilegiifolium E1 . . . . . . . . . . . . . . . . . . . . . . . . . . + 1 4 FS Fagetalia sylvaticae Galeobdolon flavidum E1 . . . . . . . . . . + + . . . . . . . + . . . . . . . 3 11 Fagus sylvatica E1 . . . . . . . . . r . . . . . . . . . . . . + . . . . 2 7 Galium laevigatum E1 . . . . . . . . . . . . . . . . + . . . . . . . . . + 2 7 Mycelis muralis E1 . . . . . . . . . . . . . . . . . . . + + . . . . . . 2 7 Salvia glutinosa E1 . . . . . . . . . . . . r . . . . . . . . . . . . . . 1 4 QP Quercetalia pubescentis Fraxinus ornus E1 . . . . . . . . . . + . + . . . . . . . . . . . . . . 2 7 Ostrya carpinifolia E2a . . . . . . . . . . . . + . . . . . . . . . . . . . . 1 4 QF Querco-Fagetea Carex digitata E1 . . . . . . . . . 1 r + . . . r . . r + . + + . + . . 9 33 Hepatica nobilis E1 . . . . . . . . . . . . . . . . . . . . . . 2 + . . . 2 7 Carex umbrosa E1 . . . . r . . . . . . . . . . . . . . . . . . . . . . 1 4 ML Mosses and lichens (Mahovi in lišaji) Ctenidium molluscum E0 + . . . . + . . . + + . 1 1 + 1 1 . 1 . + . + 1 1 . . 14 52 Neckera crispa E0 . . . + . . . . . 1 + 1 1 1 + 1 . 1 + . . . 1 . . . . 11 41 Tortella tortuosa E0 . . . . . . . . + . . . . + . 1 . + 2 + . + 1 . . 1 . 9 33 Brachythecium rutabulum E0 . . . . . . . . . . . . . . . . . . . . . . . . 1 1 2 3 11 Plagiopus oederi E0 + . . . . . . + . . . . . . . . . . . . . . . . . . . 2 7 Bryum sp. E0 . . . . . . 1 . . . . . . . . . . . . . . . . . . . . 1 4 Lescuraea plicata E0 . . . . . . . + . . . . . . . . . . . . . . . . . . . 1 4 Dicranum scoparium E0 . . . . . . . + . . . . . . . . . . . . . . . . . . . 1 4 Marchantia polymorpha E0 . . . . . . . . . . . . + . . . . . . . . . . . . . . 1 4 Schistidium apocarpum E0 . . . . . . . . . . . . . + . . . . . . . . . . . . . 1 4 Pedinophyllum interruptum E0 . . . . . . . . . . . . . . . . . . . . + . . . . . . 1 4 Plagiochila porelloides E0 . . . . . . . . . . . . . . . . . . . . + . . . . . . 1 4 19/2 • 2020, 155–231 216 Igor Dakskobler & Andrej Martinčič Plant communities of moist rock crevices with endemic Primula carniolica in the (sub)montane belt of western Slovenia Table 7 (Tabela 7): Phyteumat o-Primuletum carniolicae potentilletosum caulescentis. Number of relevé (Zaporedna številka popisa) 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 Pr. Fr. Database number of relevé (Delovna številka popisa) 228717 244377 246923 247018 275606 246607 269206 269310 255778 278001 278002 278004 275371 277987 278020 278018 Elevation in m (Nadmorska višina v m) 240 950 1050 1050 670 470 543 485 510 670 670 665 660 710 730 720 Aspect (Lega) N NE NE NE SEE NE NE NEE N NE NE NNE NNE NEE N N Slope in degrees (Nagib v stopinjah) 100 80 80 80 80 80 70 90 80 85 80 80 90 90 90 95 Parent material (Matična podlaga) D D D D D D D D D D D D D D D D Soil (Tla) Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Li Stoniness in % (Kamnitost v %) 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 Cover of herb layer in % (Zastiranje zeliščne plasti v %): E1 30 25 20 20 30 30 30 30 35 35 25 30 30 20 25 20 Co ver of moss layer in % (Zastiranje maho vne plasti v %) E0 30 5 5 5 30 5 10 10 10 20 5 10 10 5 5 5 Number of species (Število vrst) 27 12 13 15 21 19 19 17 23 29 26 31 23 18 15 13 Relevé area (Velikost popisne ploskve) m 2 10 5 12 12 15 10 10 5 5 15 15 15 15 10 15 15 Date of taking relevé (Datum popisa) 4/22/2004 8/10/2012 7/1/1998 7/1/1998 5/16/2019 5/4/2011 4/7/2017 11/18/2017 5/14/2002 4/29/2019 4/29/2019 4/29/2019 4/29/2019 7/12/2019 4/29/2019 4/29/2019 Locality (Nahajališče) Straža (Idrijca) Poldanovec Hudournik Hudournik Gorenja T rebuša - Drnulk Šebrelje - Sv. Ivan Orehovska grapa - Nemci Šebrelje - Sv. Ivan Skopica - Vučja grapa Gorenja T rebuša - Drnulk Gorenja T rebuša - Drnulk Gorenja T rebuša - Drnulk Drnulk - Kobilica Gorenja T rebuša - Drnulk Gorenja T rebuša - Drnulk Gorenja T rebuša - Drnulk Quadrant (Kvadrant) 9849/4 9949/3 9949/1 9949/1 9948/2 9849/4 9849/4 9849/4 9848/4 9948/2 9948/2 9948/2 9948/2 9948/2 9948/2 9948/2 Coordinate GK Y (D-48) m 418442 409813 413662 413728 408332 416374 417696 416791 408523 408643 408646 408635 408562 408453 408424 408320 Coordinate GK X (D-48) m 5106926 5097119 5102365 5102274 5103605 5108067 5111204 5107974 5106876 5103053 5103051 5103073 5102692 5102624 5102700 5103002 19/2 • 2020, 155–231 217 Igor Dakskobler & Andrej Martinčič Plant communities of moist rock crevices with endemic Primula carniolica in the (sub)montane belt of western Slovenia Number of relevé (Zaporedna številka popisa) 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 Pr. Fr. Diagnostic species of the association (Diagnostične vrste asociacije) Pr. Fr. AP Primula carniolica E1 + + + + + 1 1 + 1 1 1 1 + 1 1 1 16 100 PcSp Phyteuma scheuchzeri subsp. columnae E1 + + + + 1 + + + + + 1 1 + 1 + + 16 100 AP Paederota lutea E1 1 2 1 1 r + . + 1 1 1 1 1 1 + 1 15 94 AP Orthothecium rufescens E0 + . . + . + 1 + + + + + . . . . 9 56 AP Valeriana saxatilis E1 . . + + . . + . 1 1 + + + . . + 9 56 AP Carex brachystachys E1 . + . + + . . . . r . r . + + 1 8 50 Differential species of the subassociation (Razlikovalnice subasociacije) PC Potentilla caulescens E1 . . + r . + + 1 + + . + + 1 1 1 12 75 EP Erica carnea E1 . . . . r + + + + + + + + + + . 11 69 EP Polygala chamaebuxus E1 + . . . . . + + + + + + 2 . . . 8 50 PC Primula auricula E1 . . . . 1 . . . . + + + . . . . 4 25 AP Primula x venusta E1 . . . . . . . . . + + r . . . . 3 19 AP Cystopteridion s. lat. (Astrantio-Paederotion luteae nom. prov.) Fissidens dubius E0 + . . . + . . + . . . + . . . . 4 25 Asplenium viride E1 . . . . . . . . . . . . r . . . 1 6 Tofieldia calyculata E1 . . . . . . . . . . . . 1 . . . 1 6 Preissia quadrata E0 . . . . . . . . . + . . . + + . 3 19 Hymenostylium recurvirostre E0 1 . . . . . . . . . . . . . . . 1 6 Pellia endiviifolia E0 + . . . . . . . . . . . . . . . 1 6 Eucladium verticillatum E0 + . . . . . . . . . . . . . . . 1 6 Barbula crocea E0 . . . . . . . . . . . . + . . . 1 6 PcSp Physoplexido comosae-Saxifragion petraeae Campanula cespitosa E1 . . . . . . + . r . . . + . r . 4 25 Athamanta turbith E1 . . . . + . . . . . . . . . . . 1 6 PC Potentilletalia caulescentis Potentilla caulescens E1 . . + r . + + 1 + + . + + 1 1 1 12 75 Primula auricula E1 . . . . 1 . . . . + + + . . . . 4 25 Kernera saxatilis E1 . . + + . . + . . . . . . . . . 3 19 Rhamnus pumilus E1 . . . . . . . . . + . . . . . . 1 6 AT Asplenieta trichomanis Asplenium trichomanes E1 + . + + + . . . r . . + + + + r 10 63 Asplenium ruta-muraria E1 + . + + + . . r r + . + . . . . 8 50 Hieracium glaucum E1 . . . . . . . . . . . . 1 . + . 2 13 19/2 • 2020, 155–231 218 Igor Dakskobler & Andrej Martinčič Plant communities of moist rock crevices with endemic Primula carniolica in the (sub)montane belt of western Slovenia Number of relevé (Zaporedna številka popisa) 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 Pr. Fr. TR Thlaspietea rotundifolii Hieracium bifidum E1 . r . . 1 . . . . . . . + + + + 6 38 Gymnocarpium robertianum E1 . . . . + + . . . . + . . . . . 3 19 Aquilegia iulia E1 . r . . . . . . . . . . . . . . 1 6 Campanula cochleariifolia E1 . 1 . . . . . . . . . . . . . . 1 6 Hieracium porrifolium E1 . . . . . . . . . . . . . 1 . . 1 6 MC Montio-Cardaminetea Campylium stellatum E0 + . . . . . . . . . . . + . . . 2 13 Bryum pseudotriquetrum E0 1 . . . . . . . . . . . . . . . 1 6 Oxyrrhynchium hians E0 1 . . . . . . . . . . . . . . . 1 6 Cratoneuron filicinum E0 + . . . . . . . . . . . . . . . 1 6 Cololejeunea calcarea E0 . . . . . . . + . . . . . . . . 1 6 ES Elyno-Seslerietea Sesleria caerulea E1 1 r 1 1 1 + 2 + 1 2 1 1 1 + . + 15 94 Carex mucronata E1 . . + . . 1 1 1 1 + 1 + . . . + 9 56 Betonica alopecuros E1 . . . . . . . . r . r + . + . . 4 25 Aster bellidiastrum E1 1 + . . . + . . . . . . . . . . 3 19 Gentiana clusii E1 . . . . . . . . . + r . 1 . . . 3 19 Phyteuma orbiculare E1 + . . . . + . . + . . . . . . . 3 19 Hieracium villosum E1 . . + + . . . . . . . . . . . . 2 13 Carex firma E1 . + . . . . . . . . . . . . . . 1 6 Globularia cordifolia E1 . . . . . . . . . r . . . . . . 1 6 FB Festuco-Brometea Carex humilis E1 . . . . + . . . . . . + . . . . 2 13 Buphthalmum salicifolium E1 . . . . . . . . . . . + . . . . 1 6 BA Betulo-Alnetea Salix appendiculata E1 + . . . . + . . r . + . + . + 2 7 44 Salix glabra E1 . . . . . . + . . r . . . . . . 2 13 RP Rhamno-Prunetea Juniperus communis E2a . . . . . . . . . . r . . . . . 1 6 TG Trifolio-Geranietea Laserpitium siler E1 . . . . . . . . + r + + . . . . 4 25 Thesium bavarum E1 . . . . . r . . . . . . . . . . 1 6 Viola hirta E1 . . . . . . . . . . . + . . . . 1 6 19/2 • 2020, 155–231 219 Igor Dakskobler & Andrej Martinčič Plant communities of moist rock crevices with endemic Primula carniolica in the (sub)montane belt of western Slovenia Number of relevé (Zaporedna številka popisa) 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 Pr. Fr. EP Erico-Pinetea Erica carnea E1 . . . . r + + + + + + + + + + . 11 69 Rhododendron hirsutum E1 + . . . . + . . + + + + + + + . 9 56 Calamagrostis varia E1 1 r + + + . + . . + . . . + . . 8 50 Polygala chamaebuxus E1 + . . . . . + + + + + + 2 . . . 8 50 Asperula aristata E1 . . . . . . . . . . . . + 1 + . 3 19 Leontodon incanus E1 . . . . . . . . . r r . + . . . 3 19 Rhodothamnus chamaecistus E1 . . . . . . . . . + + . . . . . 2 13 Amelanchier ovalis E1 . . . . . . . . . r . . . . . . 1 6 Aquilegia nigricans E1 . . . . . + . . . . . . . . . . 1 6 Genista januensis E1 . . . . . . . . . r . . . . . . 1 6 Molinia arundinacea E1 . . . . . . . . . r . . . . . . 1 6 Pinus nigra E1 . . . . r . . . . . . . . . . . 1 6 VP Vaccinio-Piceetea Clematis alpina E1 . . + r . . . . . . . . . . . . 2 13 Hieracium murorum E1 + . . . . . . . r . . . . . . . 2 13 Veronica urticifolia E1 + . . . . . . . . . . . . . . . 1 6 AF Aremonio-Fagion Cyclamen purpurascens E1 . . . . . . + . . . + + . . . . 3 19 Helleborus niger E1 . . . . . . . . . . . + . . . . 1 6 Primula vulgaris E1 . . . . . . + . . . . . . . . . 1 6 FS Fagetalia sylvaticae Fagus sylvatica E1 . . . . . . . . r . . r . . . . 2 13 Mycelis muralis E1 . . . . r . . . . . . . . . . . 1 6 QP Quercetalia pubescenti-petraeae Ostrya carpinifolia E1 + . . . r + + . . . r + + . . + 8 50 Fraxinus ornus E1 . . . . r . . . + r r r . + . . 6 38 Mercurialis ovata E1 . . . . + . . + . . + + . . . . 4 25 Melittis melissophyllum E1 . . . . . . . . . . . r . . . . 1 6 QF Quereco-Fagetea Carex digitata E1 . . . . + + . . . . . + . . . . 3 19 Hedera helix E1 . . . . . . . . . . . . . . r . 1 6 Hepatica nobilis E1 . . . . . . . . . . . + . . . . 1 6 19/2 • 2020, 155–231 220 Igor Dakskobler & Andrej Martinčič Plant communities of moist rock crevices with endemic Primula carniolica in the (sub)montane belt of western Slovenia Number of relevé (Zaporedna številka popisa) 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 Pr. Fr. ML Mosses and lichens (Mahovi in lišaji) Ctenidium molluscum E0 1 . . . . + + + . + + . 1 . . + 8 50 Tortella tortuosa E0 . + . + . . + . + . . . + + . . 6 38 Neckera crispa E0 . . . . . + 1 1 . . + . . . . . 4 25 Pedinophyllum interruptum E0 1 . . . . . . . . . . . . . . . 1 6 Trichostomum crispulum E0 1 . . . . . . . . . . . . . . . 1 6 Anomodon attenuatus E0 1 . . . . . . . . . . . . . . . 1 6 Plagiomnium rostratum E0 + . . . . . . . . . . . . . . . 1 6 Encalypta streptocarpa E0 + . . . . . . . . . . . . . . . 1 6 Anomodon viticulosus E0 + . . . . . . . . . . . . . . . 1 6 Homalothecium sericeum E0 . . . . . . . + . . . . . . . . 1 6 Neckera complanata E0 . . . . . . . + . . . . . . . . 1 6 Lichenes div. E0 . . . . . . . . 2 . . . . . . . 1 6 Dicranodontium denudatum E0 . . . . . . . . . . . . + . . . 1 6 Schistidium apocarpum E0 . . . . . . . . . . . . . + . . 1 6 Legend / Legenda D Dolomite – dolomit Li Lithosol – kamnišče Pr. Presence (number of relevés in which the species is presented) – število popisov, v katerih se pojavlja vrsta Fr. Frequency in % – frekvenca v % 19/2 • 2020, 155–231 221 Igor Dakskobler & Andrej Martinčič Plant communities of moist rock crevices with endemic Primula carniolica in the (sub)montane belt of western Slovenia Table 8 (Tabela 8): Primulo carniolicae-Potentilletum clusianae. Number of relevé (Zaporedna številka popisa) 1 2 3 4 5 6 Pr. Fr. Database number of relevé (Delovna številka popisa) 236342 234471 246510 236341 236351 236350 Elevation in m (Nadmorska višina v m) 1320 1280 1170 1330 1325 1320 Aspect (Lega) NW NW NE NW NW NE Slope in degrees (Nagib v stopinjah) 90 90 85 90 90 90 Parent material (Matična podlaga) D D D D D D Soil (Tla) Li Li Li Li Li Li Stoniness in % (Kamnitost v %) 100 100 100 70 70 100 Cover of herb layer in % (Zastiranje zeliščne plasti v %): E1 30 30 30 30 30 30 Co ver of moss layer in % (Zastiranje maho vne plasti v %) E0 5 5 5 5 5 5 Number of species (Število vrst) 13 21 23 18 21 22 Relevé area (Velikost popisne ploskve) m 2 2 10 10 5 3 10 Date of taking relevé (Datum popisa) 7/31/2010 8/27/2004 5/11/2011 8/4/2010 8/4/2010 8/4/2010 Locality (Nahajališče) Zeleni rob Zeleni rob Poldan- ovec Zeleni rob Zeleni rob Zeleni rob Quadrant (Kvadrant) 9949/3 9949/3 9949/3 9949/3 9949/3 9949/3 Coordinate GK Y (D-48) m 411517 411773 410299 411442 411445 411512 Coordinate GK X (D-48) m 5095832 5095846 5096834 5095838 5095850 5095836 Diagnostic species of the association (Diagnostične vrste asociacije) Pr. Fr. PC Potentilla clusiana E1 + 1 1 2 1 + 6 100 AP Primula carniolica E1 + + + 1 1 1 6 100 ES Carex firma E1 + 1 1 + + + 6 100 EP Rhodothamnus chamaecistus E1 + 1 1 + + + 6 100 PC Campanula cochleariifolia E1 + r . + + 1 5 83 TR Hladnikia pastinacifolia E1 . + + r . . 3 50 TR Aquilegia iulia E1 . . + . r + 3 50 ES Hieracium pilosum E1 . . . + + + 3 50 AP Cystopteridion s. lat. (Astrantio-Paederotion luteae nom. prov.) Paederota lutea E1 1 + . + 1 2 5 83 Valeriana saxatilis E1 . r + + + + 5 83 Astrantia carniolica E1 r + . . + + 4 67 Viola biflora E1 + . . r + 1 4 67 Tofieldia calyculata E1 . r . . + . 2 33 Carex brachystachys E1 . . . r . + 2 33 Orthothecium rufescens E0 . . . . + 1 2 33 Pinguicula alpina E1 . + . . . . 1 17 Ranunculus traunfellneri E1 1 . . . . . 1 17 Soldanella minima E1 + . . . . . 1 17 Primula x venusta E1 . . r . . . 1 17 Cystopteris fragilis E1 . . . . . 1 1 17 19/2 • 2020, 155–231 222 Igor Dakskobler & Andrej Martinčič Plant communities of moist rock crevices with endemic Primula carniolica in the (sub)montane belt of western Slovenia Legend / Legenda D Dolomite – dolomit Li Lithosol – kamnišče Pr. Presence (number of relevés in which the species is presented) – število popisov, v katerih se pojavlja vrsta Fr. Frequency in % – frekvenca v % Number of relevé (Zaporedna številka popisa) 1 2 3 4 5 6 Pr. Fr. PcSp Physoplexido comosae-Saxifragion petraeae Phyteuma scheuchzeri subsp. columnae E1 . . . + + 1 3 50 Saxifraga squarrosa E1 . + + . . . 2 33 PC Potentilletalia caulescentis Bupleurum petraeum E1 . . . 1 1 . 2 33 Primula auricula E1 . . 1 . . . 1 17 Rhamnus pumilus E1 . . + . . . 1 17 AT Asplenieta trichomanis Valeriana tripteris E1 . . . . . + 1 17 TR Thlaspietea rotundifolii Adenostyles glabra E1 . . . . . r 1 17 Hieracium bifidum E1 . . . . . r 1 17 MC Montio-Cardaminetea Distichium capillaceum E0 . + . . . . 1 17 ES Elyno-Seslerietea Sesleria caerulea E1 r 1 1 2 1 1 6 100 Aster bellidiastrum E1 + . . . + + 3 50 Carex mucronata E1 . . + + + . 3 50 Laserpitium peucedanoides E1 . . r + + . 3 50 Hieracium pilosum E1 . . . + + + 3 50 Hieracium villosum E1 . r . . . . 1 17 Leontopodium alpinum E1 . . + . . . 1 17 Dryas octopetala E1 . . + . . . 1 17 Gentiana clusii E1 . . r . . . 1 17 BA Betulo-Alnetea Salix appendiculata E1 . r r . . . 2 33 Salix glabra E1 . + + . . . 2 33 EP Erico-Pinetea Rhododendron hirsutum E1 . + + . + + 4 67 Pinus nigra E1 . . r . . . 1 17 ML Mosses (Mahovi) Neckera crispa E0 . + + 1 . . 3 50 Tortella tortuosa E0 . + + + + 1 5 83 Ctenidium molluscum E0 + + . . . . 2 83 19/2 • 2020, 155–231 223 Igor Dakskobler & Andrej Martinčič Plant communities of moist rock crevices with endemic Primula carniolica in the (sub)montane belt of western Slovenia Table 9: Synoptic table of rock crevice communities with Primula carniolica. Tabela 9: Sintezna tabela združb skalnih razpok z vrsto Primula carniolica. Successive number (Zaporedna številka) 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 Sign for syntaxa (Oznaka sintaksonov) AcPa AcPc PhcPc-ac PhcPc-ty PcPcl-CT PcPcl-TG Pcty Pccb Pcvb Pc-No Pc-Do PC-Nrp Pc-Iv PcPcl-As PhPc-pc Author (Avtor) IDAM IDAM IDAM IDAM IDAM ID MA MA ID IDBD IDBDBV BDBV BDID MA ID Number of relevés (Število popisov) 19 56 27 73 16 8 28 10 15 6 14 5 7 9 6 Cystopteridion s. lat. (Astrantio carniolicae-Paederotion luteae nom. prov.) Pinguicula alpina E1 100 86 41 . . 50 7 . 60 33 14 40 71 . 17 Hymenostylium recurvirostre E0 84 77 11 8 6 . . . 20 . 7 . . . . Palustriella commutata E0 74 89 4 1 . . . . 7 . . . . . . Aster bellidiastrum E1 58 43 85 45 19 88 7 20 93 83 43 60 100 . 50 Tofieldia calyculata E1 58 54 37 16 6 38 4 40 20 17 14 60 43 . 33 Astrantia carniolica E1 58 88 67 . . 38 . . 7 . . . . . 67 Orthothecium rufescens E0 47 84 70 56 56 38 50 40 100 50 29 . . . 33 Paederota lutea E1 26 39 100 74 94 100 100 100 20 67 29 80 86 100 83 Saxifraga aizoides E1 21 . . . . . . . . . . . . . . Carex brachystachys E1 16 32 48 45 50 . 4 50 47 33 7 20 43 . 33 Preissia quadrata E0 16 29 4 12 19 . . . . . . . . . . Barbula crocea E0 16 9 7 10 6 . . . . . . . . . . Pellia endiviifolia E0 16 5 11 3 6 . . . . . 7 . . . . Valeriana saxatilis E1 5 43 81 32 56 75 29 70 7 . . . . 33 83 Fissidens dubius E0 5 9 22 27 25 38 29 10 7 . 36 20 . . . Eucladium verticillatum E0 5 7 . . 6 . . . . . . . . . . Selaginella helvetica E1 5 . . 3 . . . . . . . . . . . Primula carniolica E1 . 100 100 100 100 100 100 100 100 100 100 100 100 100 100 Asplenium viride E1 . 21 59 40 6 13 11 70 60 50 36 80 29 . . Valeriana tripteris E1 . 11 37 49 . . 18 30 53 83 64 80 71 . 17 Viola biflora E1 . 7 93 16 . . . . 100 . . . 14 . 67 Soldanella minima E1 . 2 11 . . . . . . . . . . . 17 Moehringia muscosa E1 . 2 . 8 . . . . 7 33 36 20 29 . . Cystopteris fragilis E1 . . 26 12 . . . . 53 33 14 60 . . 17 Cystopteris regia E1 . . 11 . . . . . . . . . . . . Heliosperma pusillum E1 . . 4 . . . . . . . . . . . . Solorina saccata E0 . . 4 . . . 7 . . . . . . . . Primula x venusta E1 . . . 4 19 . . . . . . . . . 17 Heliosperma veselskyi subsp. veselskyi E1 . . . 1 . . . . . . . . . . . Collema fuscovirens E0 . . . . . . 21 . . . . . . 56 . Ranunculus traunfellneri E1 . . . . . . . . . . . . . . 17 Physoplexido comosae-Saxifragion petraeae Campanula cespitosa E1 47 27 7 15 25 . . 10 20 . 29 20 . . . Phyteuma scheuchzeri subsp. columnae E1 11 48 81 100 100 88 . . 7 . . . . . 50 Micromeria thymifolia E1 5 . . . . . . . . . . . . . . Hieracium pospichalii E1 . 4 . . . . . . . . . 20 . . . Athamanta turbith E1 . . . 5 6 38 . . . . . . . . . Saxifraga squarrosa E1 . . . . . 25 . . . . . . . . 33 Silene hayekiana E1 . . . . . . . . . . . 20 . . . 19/2 • 2020, 155–231 224 Igor Dakskobler & Andrej Martinčič Plant communities of moist rock crevices with endemic Primula carniolica in the (sub)montane belt of western Slovenia Successive number (Zaporedna številka) 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 Potentilletalia caulescentis Potentilla caulescens E1 16 21 15 10 75 88 14 . . . . 100 . 100 . Kernera saxatilis E1 . 4 4 7 19 . . . . . . . . 22 . Primula auricula E1 . . . 1 25 . . . . . . . . . 17 Rhamnus pumilus E1 . . . 6 13 . 10 . . 7 . 29 78 17 Campanula cochleariifolia E1 . . . 6 25 . . . . . . . . 83 Asplenium seelosii E1 . . . . . . . . . . . . . 33 . Potentilla clusiana E1 . . . . . . . . . . . . . . 100 Bupleurum petraeum E1 . . . . . . . . . . . . . . 33 Asplenietea trichomanis Asplenium trichomanes E1 11 14 37 56 63 25 25 50 60 83 64 20 57 22 . Asplenium ruta-muraria E1 . 11 41 67 50 38 25 80 73 100 71 60 71 56 . Hieracium glaucum E1 . . . 3 13 . . . . . . . . . . Erysimum sylvestre E1 . . . . . . . . . . 7 . . . . Thlaspietea rotundifolii Petasites paradoxus E1 37 2 . . . . . . . . . . . . . Achnatherum calamagrostis E1 11 2 . . . . . . . . . . . . . Adenostyles glabra E1 11 13 33 3 . 25 4 10 . . . . . . 17 Hieracium bifidum E1 11 18 26 48 38 50 . 10 40 33 57 60 57 . 17 Hieracium porrifolium E1 5 2 . 5 6 25 . . . . 7 . . . . Gymnocarpium robertianum E1 . 9 4 12 19 . . . 7 33 14 40 . . . Aquilegia iulia E1 . . 4 1 6 13 . . . . . . . . 50 Dryopteris villarii E1 . . 7 1 . . . . . . . . . . . Saxifraga caesia E1 . . . . . 13 . . . . . . . . . Hladnikia pastinacifolia E1 . . . . . . . . . . . . . . 50 Caricetalia davallianae Carex lepidocarpa E1 21 . . . . . . . . . . . . . . Parnassia palustris E1 . 5 4 . . . . . . . . . . . . Schoenus nigricans E1 . 5 . . . . . . . . . . . . . Montio-Cardaminetea Conocephalum conicum E0 11 32 48 25 . 13 . . 80 67 14 . . . . Bryum pseudotriquetrum E0 11 4 . . 6 . . . . . . . . . . Cratoneuron filicinum E0 5 4 . . 6 . . . . . . . . . . Campylium stellatum E0 . 7 4 1 13 . . . 13 . . . . . . Distichium capillaceum E0 . 2 4 7 . . . . . . . . . . 17 Ditrichum flexicaule E0 . 2 . 1 . . . . . . . . . . . Campylium calcareum E0 . 2 . . . . . . . . . . . . . Campylium elodes E0 . 2 . . . . . . . . . . . . . Cololejeunea calcarea E0 . . . 1 6 . . . . . . . . . . Oxyrrhynchium hians E0 . . . . 6 . . . 20 . . . . . . Oxyrrhynchium schleicheri E0 . . . . . . . . 13 . . . . . . Elyno-Seslerietea Sesleria caerulea E1 68 61 59 53 94 100 75 100 20 33 29 40 43 89 100 Carex mucronata E1 11 38 11 14 56 75 11 100 . . . 20 . 100 50 Phyteuma orbiculare E1 . 9 4 19 . 4 10 . 17 . . 14 . . Carex ferruginea E1 . 7 11 3 . 13 . . . . . . . . . Carex firma E1 . 4 7 6 38 . . . . . . . . 100 Betonica alopecuros E1 . 2 . 5 25 . 7 10 . . . . 14 . . Laserpitium peucedanoides E1 . 2 15 1 . 38 11 10 . . . . 29 . 50 Campanula witasekiana E1 . . 4 . . . . . . . . . . . . 19/2 • 2020, 155–231 225 Igor Dakskobler & Andrej Martinčič Plant communities of moist rock crevices with endemic Primula carniolica in the (sub)montane belt of western Slovenia Successive number (Zaporedna številka) 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 Carduus crassifolius E1 . . . 3 . . . . . . . . . . . Gentiana clusii E1 . . . 1 19 . . . . . . . . . 17 Hieracium villosum E1 . . . 13 . . . . 17 . . . . 17 Leontopodium alpinum E1 . . . . . 13 . . . . . . . . 17 Dryas octopetala E1 . . . . . 13 . . . . . . . . 17 Ranunculus carinthiacus ? E1 . . . . . 13 . . . . . . . . . Hieracium pillosum E1 . . . . . . . . . . . . . . 50 Festuco-Brometea Buphthalmum salicifolium E1 . 4 . 3 6 . . 10 13 . 7 20 29 11 . Galium lucidum E1 . 2 . 1 . . . 10 . . 14 . . . . Carex humilis E1 . . . 4 13 . 4 10 . . . . . . . Brachypodium rupestre E1 . . . 1 . . . . . . . . . . . Linum catharticum E1 . . . . . 13 . . . . . . . . . Globularia punctata E1 . . . . . . 4 . . . . . . . . Campanula rotundifolia E1 . . . . . . . 10 . . . . . . . Sanguisorba minor E1 . . . . . . . . . . . 20 . . . Molinio-Arrhenatheretea, Calthion Angelica sylvestris E1 5 . . 4 . . . . . . . . . . . Pimpinella major E1 5 . . . . . . . . . . . . . . Caltha palustris E1 . 2 . 1 . . . . . . . . . . . Cirsium oleraceum E1 . 2 . . . . . . . . . . 14 . . T araxacum x Ruderalia E1 . . . 1 . . . . . . . . . . . Euphorbia villosa E1 . . . . . . . . . . . . 14 . . Trifolio-Geranietea Laserpitium siler E1 . 4 . 3 25 . . . . . 7 . 14 11 . Laserpitium latifolium E1 . 2 . 1 . . . . . . 7 20 14 . . Digitalis grandiflora E1 . . 4 . . . . . . . . . 14 . . Viola hirta E1 . . . 3 6 . . 10 . 17 . . . . . Anthericum ramosum E1 . . . 1 . . . . . . 7 . . . . Thalictrum minus E1 . . . 1 . . . . . . . . . . . Thesium bavarum E1 . . . 6 . . . . . . . . . . Vincetoxicum hirundinaria E1 . . . . . . . . . . 14 . . 11 . Betulo-Alnetea Salix appendiculata E1 21 13 7 7 44 25 4 10 13 17 21 40 43 . 33 Salix glabra E1 . 11 . 3 13 63 . . . . . . . . 33 Mulgedio-Aconitetea Chaerophyllum hirsutum E1 5 4 7 . . . . . . . . . . . Crepis paludosa E1 5 2 4 . . . . . . . 14 . . . . Petasites hybridus E1 5 . . . . . . . 7 . . . 14 . . Veratrum album E1 . . 4 . . . . . . . . 20 . . . Senecio ovatus E1 . . . 5 . . . . . . 7 40 . . . Aconitum degenii subsp. paniculatum E1 . . . 1 . . . . . 17 7 . . . . Ranunculus platanifolius E1 . . . 1 . . . . . . . . . . . Saxifraga rotundifolia E1 . . . 1 . . . . . . . . . . . Athyrium filix-femina E1 . . . . . . . . . 17 . 20 . . . Centaurea montana E1 . . . . . . . . . . . 20 . . . Epilobietea angustifolii Eupatorium cannabinum E1 5 5 . . . . . . . . . . 14 . . Urtica dioica E1 . . . . . . . . . 17 . . . . . 19/2 • 2020, 155–231 226 Igor Dakskobler & Andrej Martinčič Plant communities of moist rock crevices with endemic Primula carniolica in the (sub)montane belt of western Slovenia Successive number (Zaporedna številka) 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 Sambuco-Salicion capreae, Rhamno-Prunetea Rubus idaeus E1 . . 4 1 . . . . . . . . . . . Juniperus communis (inc. J. intermedia) E2a . . . 6 . . 20 . . 14 . . . . Salix caprea E1 . . . . . . . . . . . 20 . . . Sorbus aucuparia E2a . . . . . . . . . . . 20 . . . Viburnum lantana E2a . . . . . . . . . . . . 14 . . Erico-Pinetea Molinia arundinacea E1 58 43 4 1 6 . 4 . 7 . . . 29 . . Calamagrostis varia E1 37 32 33 32 50 13 . . 67 50 50 . 29 . . Erica carnea E1 16 21 4 14 69 25 36 70 7 17 36 60 . 11 . Carex ornithopoda E1 11 5 . 5 . . 11 20 . . 29 . . 11 . Leontodon incanus E1 11 . . 1 19 . . . . . 7 . 14 . . Gymnadenia odoratissima E1 5 . . . . 13 . . . . . . . . . Rhododendron hirsutum E1 . 11 33 26 56 38 21 40 13 . 14 20 43 . 67 Rhodothamnus chamaecistus E1 . 7 19 1 13 75 . . . . . . . . 100 Aquilegia nigricans E1 . 4 4 1 6 . . . . . 7 20 . . . Cirsium erisithales E1 . 2 4 3 . . . . . . . . 14 . . Polygala chamaebuxus E1 . 2 4 5 50 13 7 30 . . . . . 22 . Pinus nigra E1 . . 4 6 . . 10 . . . . . . 17 Rubus saxatilis E1 . . . 5 . . . . . . . . 29 . . Carex alba E1 . . . 4 . . 7 . . 17 29 . . . . Amelanchier ovalis E1 . . . 1 6 25 . 40 . . . 20 . 11 . Pinus sylvestris E2a . . . 1 . . . . . . . . 14 . . Aster amellus E1 . . . 1 . . . . . . . . . . . Asperula aristata E1 . . . 19 13 . . . . . . . . . Globularia cordifolia E1 . . . 6 13 4 10 . 17 . . . 11 . Genista januensis E1 . . . 6 . . . . . . . . . . Allium ericetorum E1 . . . . . . . 10 . . . . . . . Cotoneaster tomentosus E1 . . . . . . . 10 . . . . . . . Chamaecytisus hirsutus E1 . . . . . . . . . . 7 . . . . Euphrasia cuspidata ? E1 . . . . . . . . . . . . . 22 . Vaccinio-Piceetea Picea abies E1 5 2 . 3 . 13 18 . . 17 21 40 71 . . Veronica urticifolia E1 5 7 4 18 6 . . . 73 50 . . . . . Solidago virgaurea E1 . 4 . 4 . . . . . . . . . . . Aposeris foetida E1 . 2 . 1 . . . . 7 . . 20 . . . Gentiana asclepiadea E1 . 2 . 3 . . 4 . . . . . . . . Homogyne sylvestris E1 . 2 15 15 . . 21 40 13 17 14 60 . . . Oxalis acetosella E1 . 2 . 3 . . . . 13 17 . . . . . Clematis alpina E1 . . 15 4 13 . . . 7 17 . . . . . Hieracium murorum E1 . . . 4 13 . . 10 . 50 . 20 . . . Polystichum lonchitis E1 . . . 1 . . . . . . . . . . . Rosa pendulina E1 . . . 1 . . 4 10 . . 7 . . . . Abies alba E1 . . . . . . 4 30 . . 14 20 . . . Rhytidiadelphus triquetrus E0 . . . . . . . . . . . 40 . . . Aremonio-Fagion Cyclamen purpurascens E1 . 13 7 23 19 . 14 60 13 33 7 20 . . . Euphorbia carniolica E1 . 9 . 3 . . . . 7 . . 20 . . . Primula vulgaris E1 . 5 . 4 6 . 7 . . . . . . . . 19/2 • 2020, 155–231 227 Igor Dakskobler & Andrej Martinčič Plant communities of moist rock crevices with endemic Primula carniolica in the (sub)montane belt of western Slovenia Successive number (Zaporedna številka) 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 Cardamine trifolia E1 . 4 4 4 . . . . 7 . . 20 . . . Helleborus niger E1 . 4 . 7 6 13 14 40 7 33 21 . 14 . . Knautia drymeia E1 . 4 . 1 . . . . . . . . 14 . . Potentilla carniolica E1 . 4 . 4 . . . . . . . . . . . Anemone trifolia E1 . 2 7 4 . 13 . . . . . . . . . Hemerocallis lilioasphodelus E1 . 2 . 4 . 25 . . . . 7 . . . . Lamium orvala E1 . 2 . 1 . . . . . . . . . . . Dentaria enneaphyllos E1 . . 4 . . . . 7 33 . . . . . Laserpitium krapfii E1 . . . 1 . . . 10 . . . . . . . Rhamnus fallax E1 . . . 1 . . . 10 . . . . . . . Omphalodes verna E1 . . . 1 . . . . . 17 . . 14 . . Scopolia carniolica E1 . . . . . . . . 13 33 7 20 . . . Tilio-Acerion Aruncus dioicus E1 . 4 4 11 . . . . 33 33 14 20 14 . . Phyllitis scolopendrium E1 . 4 . 8 . . . . . 33 . . . . . Acer pseudoplatanus E1 . 2 7 7 . . 11 . 7 . 14 20 . . . Polystichum aculeatum E1 . 2 . 8 . . . . 13 33 7 40 . . . Thalictrum aquilegiifolium E1 . . 4 4 . . 7 20 27 . 21 40 . . . Geranium robertianum E1 . . . 5 . . . . . . 21 20 29 . . T ephroseris pseudocrispa E1 . . . 4 . . . . . . . . . . . Acer platanoides E2a . . . 1 . . . . . . . . . . . Ulmus glabra E2a . . . 1 . . . . 13 . . . . . . Fagetalia sylvaticae Galium laevigatum E1 11 4 7 7 . . 7 . 13 . 14 20 . . . Fagus sylvatica E1 5 . 7 7 13 13 4 20 7 17 14 60 29 . . Laburnum alpinum E2a 5 . . . . 25 . . . . . . . . . Salvia glutinosa E1 5 9 4 14 . . 7 . . . . 20 . . . Mycelis muralis E1 . 20 7 8 6 . 4 . 7 50 14 . . . . Galeobdolon flavidum E1 . 5 11 11 . 13 . . 47 . . 20 . . . Brachypodium sylvaticum E1 . 4 . . . . . . . . . . . . . Melica nutans E1 . 4 . 3 . . . . . . 29 20 . . . Asarum europaeum subsp. caucasicum E1 . 2 . 1 . . . . . . . . . . . Dentaria pentaphyllos E1 . 2 . 1 . . . . 27 50 . . . . . Prenanthes purpurea E1 . 2 . . . . . . 7 . 7 40 . . . Mercurialis perennis E1 . . . 10 . . 7 30 7 . 21 20 14 . . Symphytum tuberosum E1 . . . 3 . . . . . . . . . . . Campanula trachelium E1 . . . 1 . . . . 20 67 7 . . . . Lathyrus vernus E1 . . . 1 . . . . 13 . . . . . . Petasites albus E1 . . . 1 . . . . . . . 20 . . . Sambucus nigra E2a . . . 1 . . . . . . . . . . . Viola reichenbachiana E1 . . . . . . 4 . . . . . . . . Daphne mezereum E2a . . . . . . . 10 7 . . 20 . . . Lonicera alpigena E2a . . . . . . . . . 17 . . . . . Myosotis sylvatica agg. E1 . . . . . . . . . 33 . . . . . Heracleum sphondylium E1 . . . . . . . . . . 7 . . . . Poa nemoralis E1 . . . . . . . . . . . 20 . . . Polygonatum multiflorum E1 . . . . . . . . . . . 20 . . . Quercetalia pubescenti-petraeae Carex flacca E1 21 5 . . . . . . . . . 14 . . Ostrya carpinifolia E1 11 2 4 10 50 13 21 60 . . 14 . 29 33 . 19/2 • 2020, 155–231 228 Igor Dakskobler & Andrej Martinčič Plant communities of moist rock crevices with endemic Primula carniolica in the (sub)montane belt of western Slovenia Successive number (Zaporedna številka) 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 Fraxinus ornus E1 . 5 7 15 38 38 14 20 . . 21 20 . 18 . Mercurialis ovata E1 . . . 7 25 . . . . . . . . . . Clematis recta E1 . . . 1 . . . . . . . . . . . Euonymus verrucosa E2a . . . 1 . . . . . . . . . . . Melittis melissophyllum E1 . . . 6 . . . . . . . . . . Sorbus aria E2a . . . . . . . 30 . . 7 . 29 . . Querco-Fagetea Potentilla erecta E1 5 . . . . . . . . . . . 14 . . Carex digitata E1 . 32 33 38 19 13 . . 60 33 21 40 . . . Hedera helix E1 . 9 . 8 6 . . . . . 7 . . . . Clematis vitalba E1 . 7 . 1 . . . . . . . . . . . Carex umbrosa E1 . 5 4 . . . . . . . . . . . . Hepatica nobilis E1 . 5 7 22 6 . 11 10 33 33 7 20 . . . Frangula alnus E2a . 2 . . . . . . . . . . . . . T axus baccata E1 . 2 . . . . . . 7 . 7 . . . . Veratrum nigrum E1 . . . 5 . . . . . . . . . . . Acer campestre E1 . . . 1 . . . . . . . . . . . Cardamine impatiens E1 . . . 1 . . . . . . . . . . . Lonicera xylosteum E1 . . . 1 . . . . . . . . . . . Quercus petraea E1 . . . 1 . . . . . . . . . . . Anemone nemorosa E1 . . . . . . . . . 17 7 20 . . . Corylus avellana E1 . . . . . . . . . . . 20 . . . Salicetea purpureae Salix eleagnos E2a 21 . . . . . . . . . . . . . . Mosses and lichens (Mahovi in lišaji) Brachythecium rutabulum E0 5 2 11 1 . . . . . . . . . . . Ctenidium molluscum E0 5 14 52 58 50 25 21 60 33 83 64 40 . . 33 Neckera crispa E0 5 27 41 73 25 38 54 70 40 100 71 100 . 22 50 Tortella tortuosa E0 5 11 33 40 38 25 57 50 20 33 7 . . . 83 Leiocolea collaris E0 5 . . 3 . . . . . . . . . . . Reboulia hemisphaerica E0 5 . . . . . . . . . . . . . . Marchantia polymorpha E0 . 4 4 3 . . 4 . 7 . . 40 . . . Heterocladium heteropterum E0 . 4 . . . . . . . . . . . . . Mnium thomsonii E0 . 4 . 8 . . . . 27 . . . . . . Bryum sp. E0 . 2 4 . . . . . . . . . . . . Mnium marginatum E0 . 2 . 1 . . . . 20 . . . . . . Dichodontium pellucidum E0 . 2 . . . . . . . . . . . . . Encalypta vulgaris E0 . 2 . . . . . . . . . . . . . Lophozia sp. E0 . 2 . 1 . . . . . . . . . . . Rhynchostegium murale E0 . 2 . . . . . . . . . . . . . Schistidium apocarpum E0 . . 4 5 6 . . 30 . . . . . 33 . Pedinophyllum interruptum E0 . . 4 3 6 . . . 20 . . . . . . Plagiochila porelloides E0 . . 4 4 . . . . . . . . . . . Plagiopus oederi E0 . . 7 . . . . . . . . . . . . Lescuraea plicata E0 . . 4 . . . . . . . . . . . . Dicranum scoparium E0 . . 4 . . . . . . . . . . . . Encalypta streptocarpa E0 . . . 4 6 . 7 . . . . . . . . Isothecium alopecuroides E0 . . . 3 . . . . . . . . . . . Mnium stellare E0 . . . 3 . . . . . . 7 . . . . Bryum capillare E0 . . . 3 . . . . . . . . . . . 19/2 • 2020, 155–231 229 Igor Dakskobler & Andrej Martinčič Plant communities of moist rock crevices with endemic Primula carniolica in the (sub)montane belt of western Slovenia Successive number (Zaporedna številka) 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 Plagiomnium rostratum E0 . . . 1 6 . . . . . . . . . . Dichodontium pellucidum subsp. flavescens E0 . . . 1 . . . . . . . . . . . Didymodon ferrugineus E0 . . . 1 . . . . . . . . . . . Plagiobryum zierii E0 . . . 1 . . . . . . . . . . . Lejeunea cavifolia E0 . . . 1 . . . . . . . . . . . Myurella sibirica E0 . . . 1 . . . . . . . . . . . Neckera complanata E0 . . . 1 6 . . . . . . . . . . Trichostomum brachydontium E0 . . . 1 . . . . . . . . . . . Myurella julacea E0 . . . 1 . . . . . . . . . . . Lichenes div. E0 . . . 1 6 13 . . . . . . . . . Homalothecium sericeum E0 . . . . 6 . . . . . 7 . . 33 . Dicranodontium denudatum E0 . . . . 6 . . . . . . . . . . Trichostomum crispulum E0 . . . . 6 . . . . . . . . . . Anomodon attenuatus E0 . . . . 6 . . . . . . . . . . Anomodon viticulosus E0 . . . . 6 . . . . . . . . . . Musci sp. E0 . . . . . 13 . . . . . . . . . Lepraria crassissima E0 . . . . . . 32 70 . . . . . 22 . Collema polycarpon E0 . . . . . . 18 . . . . . . . . Trentepohlia aurea E0 . . . . . . 4 10 . . . . . . . Brachythecium sp. E0 . . . . . . . 10 . . . . . . . Brachythecium starkei E0 . . . . . . . . 7 . . . . . . Cirriphyllum cirrhosum E0 . . . . . . . . 7 . . . . . . Plagiomnium undulatum E0 . . . . . . . . . 17 . 60 . . . Tortella sp. E0 . . . . . . . . . . . . 33 . Caloplaca sp. E0 . . . . . . . . . . . . . 22 . Legend (Legenda) 1 AcPa Astrantio carniolicae-Pinguiculetum alpinae, this article, Table 2. 2 AcPc Astrantio carniolicae-Primuletum carniolicae, this article, Table 1. 3 PhcPc-ac Phyteumato columnae-Primuletum carniolicae astrantietosum carniolicae, this article, Table 6. 4 PhcPc-ty Phyteumato columnae-Primuletum carniolicae typicum, this article, Table 5. 5 PcPcl-CT Phyteumato columnae-Primuletum carniolicae potentilletosum caulescentis, this article, Table 7. 6 PcPcl-TG Phyteumato columnae-Primuletum carniolicae potentilletosum carniolicae (Primulo carniolicae-Potentilletum caulescentis), Dakskobler (1998, Table 1, pp. 286–287). 7 Pcty Primuletum carniolicae, Accetto (2008, Table 1, pp. 42–44). 8 Pccb Primuletum carniolicae caricetosum brachystachyos, Accetto (2015, Table 27, pp. 98–99). 9 Pcvb Primuletum carniolicae violetosum biflorae, this article, Table 4. 10 Pc-No Primuletum carniolicae s. lat., Dolinar et al. (2017, Table 1, pp. 30–32). 11 Pc-Do Primuletum carniolicae s. lat., Dolinar et al. (2013, Table 1, pp. 13–16). 12 Pc-Npr Primuletum carniolicae s. lat., Dolinar et al. (2015, Table 1, pp. 40–43). 13 Pc-Iv Primuletum carniolicae s. lat., Dolinar et al. (2013: Table 2, pp. 17–18). 14 PcPcl-As Paederoto luteae-Potentilletum caulescentis (Primulo carniolicae-Potentilletum caulescentis var. Asplenium seelosii), Accetto (2008, Table 5, pp. 49–50). 15 PcPcl Primulo carniolicae-Potentilletum clusianae, this article, Table 8. ID Igor Dakskobler BV Branko Vreš AM Andrej Martinčič MA Marko Accetto BD Branko Dolinar 19/2 • 2020, 155–231 230 Igor Dakskobler & Andrej Martinčič Plant communities of moist rock crevices with endemic Primula carniolica in the (sub)montane belt of western Slovenia Succesive number (Zaporedna številka) 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 Sign for syntaxa (Oznaka sintaksonov) AcPa AcPc PhcPc-ac PhcPc-ty PcPcl-CT PcPcl-TG Pcty Pccb Pcvb Pc-No Pc-Do PC-Nrp Pc-Iv PcPcl-As PhPc-pc Author (Avtor) ID ID ID ID ID ID MA MA ID IDBD IDBDBV BDBV BDID MA ID Number of relevés (Število popisov) 19 56 27 73 16 8 28 10 15 6 14 5 7 9 6 Cystopteridion s. lat. (Astrantio carniolicae-Paederotion luteae) 50.2 50.1 50.2 32.1 22.7 29.2 37.0 27.0 39.1 27.5 27.8 25.4 38.0 25.7 33.1 Physoplexido comosae-Saxifragion petraeae 5.32 4.76 4.79 7.03 6.78 7.62 0 0.51 1.4 0 1.85 2.46 0 0 4.22 Potentilletalia caulescentis 1.35 1.51 1.03 1.04 6.78 6.36 1.34 0.51 0 0 0.45 4.1 1.88 20.7 12.7 Asplenietea trichomanis 0.93 1.51 4.25 7.35 6.52 3.18 4.78 6.63 6.89 8.64 9.06 3.28 8.31 6.93 0 Thlaspietea rotundifolii 6.33 2.77 4.03 4.15 3.57 6.36 0.38 1.02 2.44 3.12 4.97 4.1 3.7 0 6.81 Montio-Cardaminetea, Caricetalia davallianae 3.63 3.8 3.05 1.6 0.31 0.66 0 0 5.18 3.16 0.89 0 0 0 0 Elyno-Seslerietea 6.67 7.41 6.04 4.71 12 15.3 10.3 11.73 1.04 3.16 1.85 2.46 6.49 16.8 21.2 Festuco-Brometea 0.00 0.36 0 0.56 0.98 0.66 0.77 2.04 0.67 0 1.34 1.64 1.88 0.98 0 Molinio-Arrhenatheretea, Calthion 0.84 0.24 0 0.4 0 0 0 0 0 0 0 0 1.82 0 0 Trifolio-Geranietea 0 0.36 0.22 0.56 1.91 0 0 0.51 0 0.8 2.23 0.82 2.73 1.95 0 Betulo-Alnetea 1.77 1.45 0.38 0.56 2.95 4.44 0.38 0.51 0.67 0.8 1.34 1.64 2.79 0 3.35 Mulgedio-Aconitetea 1.27 0.36 0.82 0.56 0 0 0 0 0.36 1.61 1.79 4.1 0.91 0 0 Sambuco-Salicion capreae, Rhamno-Prunetea, Epilobietea 0.42 0.3 0.22 0.08 0.31 0 0 1.02 0 0.8 0.89 1.64 1.82 0 0 Erico-Pinetea 11.7 7.66 5.93 6.07 16.1 11.5 8.61 12.2 4.87 4.77 11.4 4.92 11.2 7.82 9.34 Vaccinio-Piceetea 0.84 1.27 1.85 3.35 1.66 0.66 4.88 4.59 5.85 7.94 3.57 8.2 4.61 0 0 Aremonio-Fagion 0 2.95 1.2 3.51 1.6 2.57 3.35 6.12 2.8 7.04 2.68 3.28 2.73 0 0 Tilio-Acerion 0 0.72 0.82 2.96 0 0 1.72 1.02 4.82 4.68 4.91 5.74 2.79 0 0 Fagetalia sylvaticae 2.2 3.13 1.96 4.07 0.98 2.57 3.16 3.06 8.03 11.05 7.21 11.5 2.79 0 0 Quercetalia pubescenti-petraeae 2.7 0.72 0.6 2 6.16 2.57 3.35 5.61 0 0 2.68 0.82 4.67 4.53 0 Querco-Fagetea, Salicetea purpureae 2.2 3.74 2.4 4.71 1.6 0.66 1.05 0.51 5.18 3.92 3.13 4.1 0.91 0 0 Mosses and lichens (Mahovi in lišaji) 1.69 4.88 10.2 12.6 7.09 5.75 18.9 15.3 10.7 11 9.95 9.84 0 14.7 9.29 Total (Skupaj) 100 100 100 100 100 100 100 100 100 100 100 100 100 100 100 Table 10: Phytosociological groups in the researched rock crevice communities in the western and central Slovenia (relative frequencies). Tabela 10: Skupine diagnostičnih vrst v preučenih združbah skalnih razpok v zahodni in osrednji Sloveniji (relativne frekvence). Legend – see Table 9 (Legenda – glej tabelo 9) 19/2 • 2020, 155–231 231 Igor Dakskobler & Andrej Martinčič Plant communities of moist rock crevices with endemic Primula carniolica in the (sub)montane belt of western Slovenia Number of relevés Frequency Vascular plants (Cevnice) Primula carniolica 348 100 Paederota lutea 209 60 Phyteuma scheuchzeri subsp. columnae 196 56 Sesleria caerulea 187 54 Aster bellidiastrum 178 51 Astrantia carniolica 147 42 Asplenium ruta-muraria 147 42 Valeriana saxatilis 142 41 Asplenium trichomanes 141 41 Calamagrostis varia 139 40 Carex brachystachys 132 38 Carex digitata 126 36 Pinguicula alpina 120 34 Valeriana tripteris 118 34 Asplenium viride 114 33 Tofieldia calyculata 109 31 Hieracium bifidum 105 30 Carex mucronata 77 22 Rhododendron hirsutum 76 22 Erica carnea 75 22 Cyclamen purpurascens 63 18 Viola biflora 60 17 Campanula cespitosa 55 16 Veronica urticifolia 55 16 Potentilla caulescens 52 15 Salix appendiculata 47 14 Molinia arundinacea 45 13 Galeobdolon flavidum 44 13 Hepatica nobilis 44 13 Ostrya carpinifolia 43 12 Cystopteris fragilis 41 12 Fraxinus ornus 39 11 Aruncus dioicus 37 11 Salvia glutinosa 35 10 Mosses (Mahovi) Orthothecium rufescens 221 64 Neckera crispa 166 48 Ctenidium molluscum 143 41 Conocephalum conicum 110 32 Tortella tortuosa 101 29 Palustriella commutata 62 18 Fissidens dubius 56 16 Hymenostylium recurvirostre 48 14 Table 11: List of species in rock crevice communities with Primula carniolica, stored in the FLoVegSi database, with frequency more than 10%. Tabela 11: Seznam vrst v združbah skalnih razpok z vrsto Primula carniolica s frekvenco več kot 10 %.