review article	UDK 597.5 (262.3)
received: 2005-04-15
AN ANNOTATED CHECKLIST OF THE FAMILY GOBIIDAE
IN THE ADRIATIC SEA
Marcelo KOVACIC Natural History Museum Rijeka, HR-51000 Rijeka, Lorenzov prolaz 1, Croatia E-mail: Marcelo.Kovacic@public.srce.hr
ABSTRACT
An annotated checklist of the gobies of the Adriatic Sea is presented, including 46 species. All available data concerning the presence of gobiid species in the Adriatic Sea are compiled and critically re-examined.
Key words: check-list, Gobiidae, Adriatic Sea
LISTA AGGIORNATA DELLA FAMIGLIA GOBIIDAE IN MARE ADRIATICO
SINTESI
L'articolo presenta una lista aggiornata dei gobidi del mare Adriatico, completa di 46 specie. Vengono forniti e riesaminati con criterio critico tutti i dati disponibili inerenti la presenza delle specie di gobidi in Adriatico.
Parole chiave: lista, Gobiidae, mare Adriatico
Marcelo KOVACIC: AN ANNOTATED CHECKLIST OF THE FAMILY GOBIIDAE IN THE ADRIATIC SEA, 21-44
INTRODUCTION
The first published data on gobies of the Adriatic Sea was the record of Gobius jozo for the Split and Trogir areas (the synonym of Gobius niger Linnaeus, 1758) published in the book — Ichthyologia massiliensis» Brunnich (1765). Numerous lists of the Adriatic fish species during the following two centuries also included gobiid species (Stevcic, 1977). However, the authors of these lists were not gobiologists, in the 19th century often not even ichthyologists. These papers were rarely written by specialists on original material, and numerous lists contained just the species name, while some, in addition, contained general comments on the species (Carrara, 1846; A. Stossich, 1869; Canestrini, 1872; M. Stossich, 1880; Faber, 1883; Brusina, 1891; Sucker, 1895; Griffini, 1903; E. Ninni, 1912; Soljan, 1948, 1965; Jardas, 1985). The names of gobiid species were cited by rote from one list to another. These lists, due to unclear synonymy, also included non-valid names and names of species whose presence in the Adriatic Sea could not be proved. Only several lists were orientated just on gobies (Kolombatovic, 1891; A. P. Ninni, 1882; Damiani, 1896; E. Ninni, 1938; Cavinato, 1952; Kovacic, 1994).
The synonymy of Mediterranean gobies was cleared by Miller (1973a). Stevcic (1977), using the synonymy of Miller (1973a), listed 40 gobiid species of the Adriatic Sea. This number was later increased by the reviews of Kovacic (1994) - 42 species, and Jardas (1996a) - 44 species, due to original papers published in the meantime. Recent years have been a dynamic period for the Adriatic gobiology. Therefore, the review of Lipej & Dulcic (2004), among other fishes, listed new gobiid species found in the Adriatic Sea since Jardas (1996a), and they concluded that, with 50 species, gobies are the best represented fish family in the Adriatic Sea. However, all these authors avoided critical reconsideration of the presence of species that were previously included in the Adriatic fauna. The only exception is the replacement of Vanneaugobius pruvoti (Fage, 1907) in Jardas (1996a) with Vanneaugobius dollfusi Brownell, 1978 by Lipej & Dulcic (2004). However, this is the result of an in-between published paper on re-examined specimens by Pallaoro & Kovacic (2000). The scope of the present paper is to give complete, actual list of gobiid species in the Adriatic Sea based on critical re-examination of all available data concerning the presence of gobiid species in the Adriatic Sea.
MATERIAL AND METHODS
This review was based on scientific literature and on unpublished data on ichthyological collection of the Natural History Museum Rijeka and on ichthyological collection of the Center for Marine Research Rovinj. The gobiid species are considered to be present in the Adri-
atic Sea if the following conditions are met: Adriatic specimens of these species are deposited in the collections, or the published descriptions of Adriatic specimens contained enough morphological data for positive identification. The mentioned criteria prevent that once wrongly cited species for the Adriatic Sea would repeatedly to be listed as the part of the Adriatic fauna. The published data on species previously listed for the Adriatic Sea that should be excluded from the Adriatic fauna are critically examined. The annotation for each species contained bibliography of published records of the species in the Adriatic Sea, and of preserved specimens in the collections. The localities of the published records were listed along the Adriatic Sea in north-south direction. All records of gobiid species already known for the Adriatic Sea but difficult for identification are considered to be without positive identification, if specimens were not described, deposited, or checked in situ by fish taxonomist.
CHECKLIST
The presented checklist contains 46 gobiid species recorded in the Adriatic Sea up to the present date (Tab. 1).
Aphia minuta mediterránea De Buen, 1931
Gobius Aphya: Naccari, 1822; Martens, 1838. Gobius pellucidus: Nardo, 1827; Kolombatovic, 1891.
Brachyochirus prototypes: Nardo, 1860. Brachyochirus aphya: Trois, 1875. Latrunculus pellucidus: Giglioli, 1880; Kolombatovic, 1881, 1882; A. P. Ninni, 1882. Gobius albus: Graeffe, 1888. Aphia pellucida: D'Ancona, 1922; Zei, 1942, 1949. Brachyochirus pellucidus: Zupanovic, 1961; Mar-cuzzi, 1972.
Aphia minuta mediterranea: Jardas et al., 1996; Pal-laoro & Jardas, 1996.
Aphia minuta: Froglia & Gramitto, 1989; Ungaro et al., 1994; Nocita & Vanni, 1997;Kovacic, 1998, 2003; La Mesa, 1999; Sorice & Caputo, 1999; Ca-puto et al., 2000; Zavodnik & Kovacic, 2000; Crn-kovic, 2001.
The species was first reported in the Adriatic Sea for the Venice Lagoon under the synonym Gobius Aphya (Naccari, 1822). The species was listed under various synonyms for the Venice Lagoon, the Gulf of Triest, the Rijeka Bay, the Kvarner area, Dalmatia, the Split area, and Dubrovnik (Nardo, 1827, 1860;Martens, 1838; Trois, 1875; Giglioli, 1880; Kolombatovic, 1881, 1882, 1891; A. P. Ninni, 1882;Graeffe, 1888;D'Ancona, 1922; Zavodnik & Kovacic, 2000). The specimens from Triest, the Kvarner area, the Solta Island, and Dubrovnik
Marcelo KOVACIC: AN ANNOTATED CHECKLIST OF THE FAMILY GOBIIDAE IN THE ADRIATIC SEA, 21-44
are deposited in the collection of the Natural History Museum Rijeka, in the collection of Museo di Storia Naturale dell' Universita di Firenze, in the collection of Stazione Idrobiologica di Chioggia, and in the collection of the Institute of Oceanography and Fisheries, Split (Marcuzzi, 1972;Pallaoro & Jardas, 1996;Nocita & Vanni, 1997; Kovacic, 1998, 2003, unpublished data). Specimens were collected by the small scale fishery gear in the Kvarner area, the Kornati Islands, and the Murter Sea (Jardas et al., 1996; Crnkovic, 2001), and by the trawl in the northern Adriatic, in the channels between the mid-Dalmatian islands, in the central Adriatic, and in the Manfredonia area (Zei, 1942, 1949; Zupanovic, 1961; Froglia & Gramitto, 1989; Ungaro et al., 1994). The samples of this species were also collected at Or-tona and Ancona (central Adriatic Sea) (La Mesa, 1999; Sorice & Caputo, 1999; Caputo et al., 2000).
Buenia affinis lljin, 1930
Buenia affinis: Kovacic, 2002a.
It was considered for a long time that the species was described in the Adriatic Sea by Kolombatovic (1891). The discovery of Miller (1972a) that Kolombatovic's syntypes in the collection of the Naturhistorischen Museum, Wien belong to another species, Pomatoschistus pictus, were ignored in later reviews. In these reviews the species was treated as present in the Adriatic Sea (Tortonese, 1975; Stevcic, 1977; Kovacic, 1994; Jardas, 1996a), or as a junior synonym of other valid species (E. Ninni, 1938; Soljan, 1948; Cavinato, 1952; Bini, 1969). Kovacic (2002a) finally collected true specimens of B. affinis in 1996 and 1997 in the Kvarner area. The specimens from the Kvarner area and Seline (the Velebit Channel) are deposited in the collection of the Natural History Museum Rijeka (Kovacic, unpublished data). Visual census research recorded the species at Kostrena, the Rijeka Bay (Kovacic, 2002b).
Chromogobius quadrivittatus (Steindachner, 1863)
Gobius quadrivittatus: Steindachner, 1863; Graeffe, 1888.
Gobius planiceps: Bellotti, 1879. Gobius quadrivittatus=Gobius planiceps: Kolombatovic, 1881, 1882, 1886.
Gobius planiceps v. quadrivitatta: Kolombatovic, 1891.
Chromogobius quadrivittatus: Miller, 1971; Ahnelt, 1990; Zavodnik & Crnkovic, 1992; Kovacic, 1994, 1997, 1998, 2002b; Jardas et al., 1996, 1998; Pal-laoro & Jardas, 1996; Nocita & Vanni, 1997; Zavod-nik & Kovacic, 2000.
The species was described by Steindachner (1863)
on the specimens collected at the Hvar Island. The species was listed for Venice, Triest, the Rijeka Bay, the Hvar Island, and the Split area (Perugia, 1866; Bellotti, 1879; Kolombatovic, 1881, 1882, 1886, 1891; A. P. Ninni, 1882; Graeffe, 1888; Kovacic, 1994; Jardas et al., 1998; Zavodnik & Kovacic, 2000). The specimens from Triest, Rovinj, the Kvarner area, the Hvar Island, and the Split area are deposited in the collection of the Natural History Museum Rijeka, in the collection of the Naturhistorisches Museum Wien, and in the collection of the Institute of Oceanography and Fisheries, Split (Miller, 1971; Ahnelt, 1990; Pallaoro & Jardas, 1996; Nocita & Vanni, 1997; Kovacic, 1997, 1998, unpublished data). The species was also recorded at numerous localities in the Kvarner area (Zavodnik & Crnkovic, 1992; Kovacic, 1997, 2002b). Specimens were collected by the small scale fishery gear at the Kornati Islands and the Murter Sea (Jardas et al., 1996). Visual census research recorded the species at Kostrena, the Rijeka Bay (Kovacic, 2002b).
Chromogobius zebratus zebratus (Kolombatovic, 1891)
Gobius planiceps zebrata: Kolombatovic, 1891. Chromogobius zebratus zebratus: Miller, 1971; Kovacic, 1994.
Chromogobius zebratus: Ahnelt, 1990; Kovacic, 1997, 1998; Jardas et al., 1998; Zavodnik & Kovacic, 2000.
The species was described by Kolombatovic (1891) on the specimens collected in the Split area. The specimens from the Kvarner area (the northern Adriatic Sea), the Solta Island, the Split area, and from Mala Duba (the central Adriatic Sea) are deposited in the collection of the Natural History Museum Rijeka, and in the collection of the Naturhistorisches Museum Wien (Miller, 1971; Ahnelt, 1990; Kovacic, 1997, 1998, unpublished data; Zavodnik & Kovacic, 2000). The species was also recorded at numerous localities in the Kvarner area (Ko-vacic, 1994, 1997; Jardas et al., 1998).
Corcyrogobius liechtensteini (Kolombatovic, 1891)
Gobius liechtensteini: Kolombatovic, 1891 (part.). Corcyrogobius liechtensteini: Miller, 1972b; Kovacic, 1997, 1998; Arko Pijevac et al., 2001.
The species was described by Kolombatovic (1891) on the specimens collected in the Split area. The additional specimens at Split were collected also by Kolombatovic (1895). The specimens from the Kvarner area (the northern Adriatic Sea), from Mala Duba, and the Korcula Island (the central Adriatic Sea) are deposited in the collection of the Natural History Museum Rijeka and in the collection of the Naturhistorisches Museum Wien (Miller, 1972b; Kovacic, 1997, 1998, unpublished data).
Marcelo KOVACIC: AN ANNOTATED CHECKLIST OF THE FAMILY GOBIIDAE IN THE ADRIATIC SEA, 21-44
Benthic biocoenological research recorded the species in the Kvarner area (Arko Pijevac et al., 2001).
Crystallogobius linearis (Von Duben, 1845)
Crystallogobius Nilssonii: Kolombatovic, 1900. Crystallogobius nilssoni: Zupanovic & Grubisic, 1958; Zupanovic, 1961.
Crystallogobius Nilssoni: Jukic & Crnkovic, 1974. Crystallogobius linearis: Zupanovic & Jardas, 1989; Pall aoro & Jardas, 1996; Kovacic, 1998; La Mesa, 2001; Caputo et al., 2003.
The species was first reported in the Adriatic Sea for Dalmatia (Kolombatovic, 1900). The specimens from the central Adriatic Sea and the Split area are deposited in the collection of the Natural History Museum Rijeka and in the collection of the Institute of Oceanography and Fisheries, Split (Pallaoro & Jardas, 1996; Kovacic, 1998). Specimens were collected by the bottom trawl in the channels of the north-eastern Adriatic Sea, the Jabuka Pit, near Rogoznica, and in the channels between the mid-Dalmatian islands (Zupanovic & Grubisic, 1958; Zupanovic, 1961; Jukic & Crnkovic, 1974; Zupanovic & Jardas, 1989), without a positive identification. The samples of this species were also collected at Ortona (central Adriatic Sea) (La Mesa, 2001; Caputo et al., 2003).
Deltentosteus colonianus (Risso, 1826)
Gobius Liechtensteinii: Steindachner, 1883. Gobius colonianus: Steindachner & Kolombatovic, 1884; Kolombatovic, 1886, 1891. Deltentosteus colonianus: Jardas et al., 1996; Pallaoro & Jardas, 1996;Nocita & Vanni, 1997;Ko-vacic, 1998.
The species was first reported in the Adriatic Sea for the Solta Island by Steindachner (1883). The additional specimens from the Split area were reported by Steindachner & Kolombatovic (1884) and Kolombatovic (1886, 1891). The specimens from the Kvarner area, the Vrgada Island, the Solta Island, the Hvar Channel, and the Split area are deposited in the collection of the Natural History Museum Rijeka, in the collection of Museo di Storia Naturale dell' Universita di Firenze, and in the collection of the Institute of Oceanography and Fisheries, Split (Pallaoro & Jardas, 1996; Nocita & Vanni, 1997; Kovacic, 1998, unpubl. data). Specimens were collected by the small scale fishery gear at the Kornati Islands and the Murter Sea (Jardas et al., 1996).
Deltentosteus quadrimaculatus (Valenciennes, 1837)
Gobius marsio: Nardo, 1827, 1860.
Gobius quadrimaculatus: Perugia, 1866, 1881; Trois, 1875; Kolombatovic, 1881, 1882, 1891; A. P. Ninni, 1882; Zei, 1942, 1949; Zupanovic, 1961; Crnkovic, 1970.
Deltentosteus quadrimaculatus: Jukic & Crnkovic, 1974; Jukic, 1975, 1983; Gamulin-Brida et al., 1980; Zupanovic & Jardas, 1989; Cetinic & Pallaoro, 1990b; Jardas, 1996b; Jardas et al., 1996, 1998; Pallaoro & Jardas, 1996; Span et al., 1996; Nocita & Vanni, 1997; Kovacic, 1998; Zavodnik & Kovacic, 2000.
Deltentosteus (Gobius) quadrimaculatus: Usic, 2003.
The species was first reported in the Adriatic Sea for the Venice Lagoon under the synonym Gobius marsio (Nardo, 1827). Miller (1973a) doubted regarding identity of G. marsio in Nardo (1827). However, Nardo (1860) itself mentioned "G. quadrimaculatus, Valenc." as synonym of his G. marsio. The species was listed for the Venice Lagoon, Triest, Istria, the Kvarner area, and the Split area (Nardo, 1860;Perugia, 1866, 1881; Trois, 1875; Kolombatovic, 1881, 1882, 1891; A. P. Ninni, 1882; Zavodnik & Kovacic, 2000). The specimens from Venice, Rovinj, Istria, the Kvarner area, the Split area, the Lastovo Channel, the Kastela Bay, the Mali Ston Bay, the Murter Island, and Bari are deposited in the collection of the Center for Marine Research of the Ruder Boskovic Institute in Rovinj, in the collection of the Natural History Museum Rijeka, in the collection of Museo di Storia Naturale dell' Universita di Firenze, and in the collection of the Institute of Oceanography and Fisheries, Split (Pallaoro & Jardas, 1996; Nocita & Vanni, 1997; Kovacic, 1998, unpubl. data; Usic, 2003). Specimens were collected by the small scale fishery gear at the Kornati Islands and the Murter Sea, northern and central Dalmatia (Cetinic & Pallaoro, 1990b; Jardas et al., 1996), and by the bottom trawl in the northern Adriatic, the Kvarner area, the channels of the north-eastern Adriatic Sea, the Jabuka Pit, the central Adriatic, the channels between the mid-Dalmatian islands, the Murter Sea, and Crnogorsko primorje (Zei, 1942, 1949; Zupanovic, 1961; Crnkovic, 1970; Jukic & Crnkovic, 1974; Jukic, 1975, 1983; Zupanovic & Jardas, 1989; Jardas, 1996b; Jardas et al., 1998; Span et al., 1996). Benthic biocoenological research recorded the species at the Krk Island (Gamulin-Brida et al., 1980).
Didogobius schlieweni Miller, 1992
Didogobius schlieweni: Miller, 1992.
The species was described by Miller (1992) on the single male collected at the Unije Island, near the Cres Island, the Kvarner area on 26 June 1991. The holotype is deposited in the collection of the Zoologische Staatssammlung, München. Three additional specimens in the
Marcelo KOVACIC: AN ANNOTATED CHECKLIST OF THE FAMILY GOBIIDAE IN THE ADRIATIC SEA, 21-44
Adriatic Sea were collected at the Krk Island (the Kvarner area), at the Solta Island, and the Ugljan Island (the central Adriatic) (Pallaoro & Jardas, 1996; Kovacic, unpubl. data). These specimens are deposited in the collection of the Natural History Museum Rijeka and in the collection of the Institute of Oceanography and Fisheries, Split.
Didogobius spletchnai Ahnelt & Patzner, 1995
Didogobius spletchnai: Herler & Patzner, 2002.
Single subadult was collected for the first time in the Adriatic Sea near Pula (the Istrian Peninsula) on 3 June 2001 (Herler & Patzner, 2002). The specimen is deposited in the collection of the Naturhistorisches Museum Wien.
Gammogobius steinitzi Bath, 1971
Gammogobius steinitzi: Kovacic, 1999.
Two females were collected for the first time in the Adriatic Sea in the Vrbnik cave, at the Krk Island (the Kvarner area) on 16 October 1998 (Kovacic, 1999). The additional nine specimens were collected at the same locality on 9 and 13 September 1999 (Kovacic, unpubl. data). They all are deposited in the collection of the Natural History Museum Rijeka.
Gobius ater Bellotti, 1888
Gobius ater: Ahnelt, 2001.
Three males, collected by Kolombatovic at Split (the central Adriatic), were found in the collection of the Naturhistorisches Museum Wien and identified by Ahnelt (2001).
Gobius auratus Risso, 1810
Gobius auratus: Perugia, 1866, 1881; Trois, 1875; Giglioli, 1880; Kolombatovic, 1881, 1882, 1891; Perugia, 1881; A. P. Ninni, 1882; Faber, 1883; Vin-ciguerra, 1883; Damiani, 1896; Langhoffer, 1904; D'Ancona, 1922; Pallaoro & Jardas, 1996; Nocita & Vanni, 1997; Orepic et al., 1997; Castellarin et al., 2001; Novosel et al., 2002; Herler et al., 2005. Gobius fallax: Kovacic, 1994, 1998; Jardas et al., 1998.
Gobius xanthocephalus: Zavodnik & Kovacic, 2000; Kovacic, 2002b.
The species was first reported for the Adriatic Sea by Perugia (1866), without any notice on collection locality. The species was later listed for Venice, Triest, the
Rijeka Bay, the Pasman Island, Zadar, the Zadar Channel, the Hvar Island, the Split area, Dalmatia, the Vis Island, and the Boka Kotorska Bay, without positive identification (Trois, 1875; Giglioli, 1880; Kolombatovic, 1881, 1882, 1891; Perugia, 1881; A. P. Ninni, 1882; Faber, 1883; Vinciguerra, 1883; Damiani, 1896; Langhoffer, 1904; D'Ancona, 1922). The specimens from the Kvarner area, the central Adriatic, the Solta Island, and Split are deposited in the collection of the Institute of Oceanography and Fisheries, Split, in the collection of the Natural History Museum Rijeka, and in the collection of Museo di Storia Naturale dell' Universita di Firenze (Pallaoro & Jardas, 1996; Nocita & Vanni, 1997; Kovacic, 1998, unpubl. data). The species belongs to Gobius auratus species complex (Miller & El-Tawil, 1974; Herler et al., 2005) and specimens in the collections of Split and Firenza need re-examination because of morphological similarities between G. auratus, G. fallax and G. xanthocephalus. The specimens recorded in the Kvarner area as G. fallax in Kovacic (1994, 1998) and Jardas et al. (1998) belong to G. auratus. G. xanthocephalus reported in Zavodnik & Kovacic (2000) and Kovacic (2002b) for the Rijeka Bay is, according to Herler et al. (2005), north Adriatic color morph of G. auratus. Benthic biocoenological researches recorded the species in the Velebit Channel, and the Mljet Island (Orepic et al., 1997; Novosel et al., 2002), and the visual census researches recorded the species at Triest (Castellarin et al., 2001), without positive identification. Herler et al. (2005) collected specimens at Selce, the Krk Island, the Cres Island (the Kvarner area), and the Murter Island (the central Adriatic Sea). The part of these specimens is deposited in the collection of the Naturhistorisches Museum Wien (Herler et al., 2005).
Gobius bucchichi Steindachner, 1870
Gobius Bucchichi: Steindachner, 1870. Gobius buchichi: Perugia, 1881. Gobius Buchichii: Kolombatovic, 1881. Gobius buchichii: A. P. Ninni, 1882. Gobius bucchichi: Kolombatovic, 1891; Tortonese, 1975; Onofri, 1983; Ahnelt, 1984; Zavodnik & Zavodnik, 1986; Musin, 1989; Kraljevič & Pallaoro, 1991; Kovacic, 1994, 1998, 2002b; Jardas et al., 1996, 1998; Pallaoro & Jardas, 1996; Simonovic et al., 1996; Jaklin & Arko-Pijevac, 1997;Nocita & Vanni, 1997; Orepic et al., 1997; De Girolamo et al., 1998; Simonovic, 1999; Guidetti & Bussotti, 2000; Zavodnik & Kovacic, 2000; Novosel et al., 2002. Gobius Bucchichii: Gridelli, 1931. Gobius bucchichii: Guidetti, 2000; Castellarin et al., 2001.
The species was described by Steindachner (1870) on the specimens collected at the Hvar Island. The syn-
Marcelo KOVACIC: AN ANNOTATED CHECKLIST OF THE FAMILY GOBIIDAE IN THE ADRIATIC SEA, 21-44
types are deposited in the collection of the Museo Civico di Storia Naturale di Genova and in the collection of the Naturhistorisches Museum Wien (Tortonese, 1963; Miller, 1973a; Ahnelt, 1984). The species was listed for the Venice area, Triest, the Rijeka Bay, Zadar, and the Split area (Kolombatovic, 1881, 1891; A. P. Ninni, 1882; Perugia, 1881; Kovačic, 1994; Zavodnik & Kova-čic, 2000). The specimens from Triest, the Kvarner area, the Kornati Islands, the Biograd area, Split, the Solta Channel, the Brač Channel, the Hvar Island, the Korčula Island, and Dubrovnik area are deposited in the collection of the Natural History Museum Rijeka, in the collection of the Institute of Oceanography and Fisheries, Split, in the collection of the Natural History Museum of Split, in the collection of the Natural History Museum of the Biological Institute, Dubrovnik, in the collection of the Museo Civico di Storia Naturale di Genova, in the collection of the Museo Civico di Storia Naturale di Trieste, in the collection of the Naturhistorisches Museum Wien, and in the collection of Museo di Storia Naturale dell' Universita di Firenze (Gridelli, 1931; Tortonese, 1975; Onofri, 1983; Ahnelt, 1984; Mušin, 1989; Pal-laoro & Jardas, 1996; Nocita & Vanni, 1997; Kovačic, 1998, unpubl. data). Specimens were collected by the small scale fishery gear at the Kornati Islands and the Murter Sea (Kraljevič & Pallaoro, 1991; Jardas et al., 1996). Benthic biocoenological researches recorded the species in the Raša Bay, the Lošinj Island, the Sv. Marko Islet, the Velebit Channel, the Kornati Islands, the Murter Sea, and the Mljet Island (Zavodnik & Zavodnik, 1986; Jardas et al., 1996; Jaklin & Arko-Pijevac, 1997; Orepič et al., 1997; Novosel et al., 2002), and the visual census researches recorded the species in the Slovenian coastal waters, Triest, the Rijeka Bay, Cavtat, the Tremiti Islands, and the Boka Kotorska Bay (Simonovic et al., 1996; De Girolamo et al., 1998; Simonovic, 1999; Guidetti, 2000; Guidetti & Bussotti, 2000; Castellarin et al, 2001; Kovačic, 2002b; Jardas et al., 1998).
Gobius cobitis Pallas, 1811
Gobius capito: Perugia, 1866; Trois, 1875 Giglioli, 1880; Kolombatovic, 1881, 1891; A. P. Ninni, 1882; Graeffe, 1888;Langhoffer, 1904;E. Ninni, 1912; D'Ancona, 1922.
Gobius exanthematosus: Perugia, 1881; Vinciguerra, 1883; Usic, 2003.
Gobius cobitis: Cavinato, 1952; Marcuzzi, 1972; Tortonese, 1975; Mušin, 1989; Kovačic, 1994, 1998, 2002b; Jardas et al., 1996; 1998; Pallaoro & Jardas, 1996; Caputo et al., 1997; Nocita & Vanni, 1997; Orepic et al., 1997; Caputo, 1998; De Girolamo et al., 1998; Sorice & Caputo, 1999; Zavodnik & Kovačic, 2000; Castellarin et al., 2001; Pallaoro, 2001; Novosel et al., 2002; Turk et al., 2002; Lipej et al. 2003.
The species was first reported in the Adriatic Sea for Triest (Perugia, 1866). The species was listed under various synonyms for Venice, the Venice Lagoon, Triest, Istria, the Rijeka Bay, the Zadar Channel, the Murter Island, the Split area, the Hvar Island, the Vis Island, the Mljet Island, and Dalmatia (Trois, 1875; Giglioli, 1880; Kolombatovic, 1881, 1891; A. P. Ninni, 1882; Perugia, 1881; Vinciguerra, 1883; Graeffe, 1888; Langhoffer, 1904; E. Ninni, 1912; D'Ancona, 1922; Cavinato, 1952; Kovacic, 1994; Zavodnik & Kovacic, 2000). The specimens from Venice, Triest, the Istrian peninsula, the Kvar-ner area, the Pag Island, Dalmatia, the Split area, the central Adriatic, and the Dubrovnik area are deposited in the collection of the Center for Marine Research Rovinj, in the collection of the Natural History Museum Rijeka, in the collection of the Institute of Oceanography and Fisheries, Split, in the collection of Museo di Storia Naturale dell' Universita di Firenze, in the collection of the Museo Civico di Storia Naturale di Genova, in the collection of l'Istituto di Idrobiologica di Chioggia, and in the collection of the Natural History Museum of the Biological Institute, Dubrovnik (Marcuzzi, 1972; Tortonese, 1975; Musin, 1989; Pallaoro & Jardas, 1996; Nocita & Vanni, 1997; Kovacic, 1998, unpubl. data; Usic, 2003; Zavodnik, pers. comm.). Specimens were collected by the small scale fishery gear in the Split area (Pallaoro, 2001). Benthic biocoenological researches recorded the species in the Velebit Channel, the Kornati Islands, the Murter Sea, and the Mljet Island (Jardas et al., 1996; Orepic et al., 1997; Novosel et al., 2002), and the visual census researches recorded the species at Triest, the Slovenian coastal waters, the Rijeka Bay, the Kornati Islands, and the Murter Sea (Jardas et al., 1996; 1998; De Girolamo et al., 1998; Castellarin et al., 2001; Kovacic, 2002b; Turk et al., 2002; Lipej et al. 2003). The samples of G. cobitis were also collected at Ancona (Caputo et al., 1997; Caputo, 1998; Sorice & Caputo, 1999).
Gobius couchi Miller & El-Tawil, 1974
Gobius couchi: Kovacic, 2001a.
Fourteen females and fifteen males were collected for the first time in the Adriatic Sea at Ostro (the Kvarner area), in 1996 and 1997 (Kovacic, 2001a). The specimens are deposited in the collection of the Natural History Museum Rijeka. The additional findings in the Adriatic Sea are from Bakar, Klenovica, Kacjak (the Kvarner area), and the Solta Island (the central Adriatic) (Kovacic, unpubl. data). These specimens are also deposited in the collection of the Natural History Museum Rijeka.
Gobius cruentatus Gmelin, 1789
Gobius cruentatus: Nardo, 1827, 1860; Martens,
1838; Plucar, 1846; Perugia, 1866; Canestrini, 1872;
Marcelo KOVACIC: AN ANNOTATED CHECKLIST OF THE FAMILY GOBIIDAE IN THE ADRIATIC SEA, 21-44
Trois, 1875; Giglioli, 1880; Kolombatovic, 1881, 1891; A. P. Ninni, 1882; Vinciguerra, 1883; Graeffe, 1888; Langhoffer, 1904; D'Ancona, 1922; Zupano-vic, 1961; Marcuzzi, 1972; Tortonese, 1975; Jardas & Pallaoro, 1989; Cetinic & Pallaoro, 1990a, 1990b; Zavodnik & Crnkovic, 1992; Kovacic, 1994, 1998, 2002b, 2004; Jardas et al, 1996, 1998; Pallaoro & Jardas, 1996; Simonovic et al., 1996; Nocita & Vanni, 1997; Orepic et al., 1997; De Girolamo et al., 1998; Simonovic, 1999; Guidetti, 2000; Zavodnik & Kovacic, 2000; Arko Pijevac et al., 2001; Castellarin et al., 2001; Pallaoro, 2001; Novosel et al., 2002; Turk et al., 2002; Lipej et al. 2003; Usic, 2003.
The species was first reported in the Adriatic Sea for Venice (Nardo, 1827). The species was listed for the Venice Lagoon, Triest, the Rijeka Bay, the Ugljan Island, the Split area, the Brac Island, the Solta Island, the Las-tovo Island, the Korcula Island, the Mljet Island, and the Vis Island (Martens, 1838; Plucar, 1846; Nardo, 1860; Perugia, 1866; Canestrini, 1872; Trois, 1875; Giglioli, 1880; A. P. Ninni, 1882; Vinciguerra, 1883; Graeffe, 1888; Kolombatovic, 1881, 1891;Langhoffer, 1904; D'Ancona, 1922; Kovacic, 1994; Zavodnik & Kovacic,
2000).	The specimens from Triest, Rovinj, the Kvarner area, Zadar, the central Adriatic, Dalmatia, the Solta Island, the Split area, and Dubrovnik are deposited in the collection of the Natural History Museum Rijeka, in the collection of the Institute of Oceanography and Fisheries, Split, in the collection of l'Istituto di Idrobiologica di Chioggia, in the collection of the Museo Civico di Storia Naturale di Genova, and in the collection of Museo di Storia Naturale dell' Universita di Firenze (Marcuzzi, 1972; Tortonese, 1975; Pallaoro & Jardas, 1996; Nocita & Vanni, 1997; Kovacic, 1998; Usic, 2003). The finding by the bottom trawl in the channels between the mid-Dalmatian islands (Zupanovic, 1961) is quite surprising, considering depth and habitat preferences of the species. Specimens were collected by the small scale fishery gear at the Kornati Islands, the Murter Sea, the northern and the central Dalmatia, the Split area, the Split Channel, and the Brusnik Island (Jardas & Pallaoro, 1989; Cetinic & Pallaoro, 1990a, 1990b; Jardas et al., 1996; Pallaoro,
2001).	Benthic biocoenological researches recorded the species in the Kvarner area, the Velebit Channel, the Kornati Islands, the Murter Sea, and the Mljet Island (Zavodnik & Crnkovic, 1992; Jardas et al., 1996; Orepic et al., 1997; Arko Pijevac et al., 2001; Novosel et al.,
2002),	and the visual census researches recorded the species at Triest, the Slovenian coastal waters, the Rijeka Bay, and the Tremiti Islands (De Girolamo et al., 1998; Jardas et al., 1998; Guidetti, 2000; Castellarin et al., 2001; Kovacic, 2002b; Turk et al., 2002;Lipej et al.
2003).	The samples of this species were also collected in the Kvarner area and the Boka Kotorska Bay (Simonovic et al., 1996; Simonovic, 1999; Kovacic, 2004).
Gobius fallax Sarato, 1889
Gobius fallax: Gridelli, 1931; E. Ninni, 1938; Tortonese, 1975; Ahnelt, 1984; Cetinic & Pallaoro, 1990b; Jardas et al., 1996; Pallaoro & Jardas, 1996; De Girolamo et al., 1998; Zavodnik & Kovacic, 2000; Turk et al., 2002; Lipej et al. 2003; Herler et al., 2005.
The species was first reported in the Adriatic Sea for Triest and Šibenik (Gridelli, 1931). Miller (1973a) supposed that G. auratus v. ruginosa of Kolombatovic (1891) from the Split area is a synonym of G. fallax. The species was listed for Venice and the Korcula Island (E. Ninni, 1938; Tortonese, 1975). Single specimen, among Steindacher's syntypes of G. bucchichi from 1870, were found in the collection of the Naturhistorisches Museum Wien and identified by Ahnelt (1984). The specimens from Triest, the Goli Island in the Kvarner area, Šibenik, and the Šolta Island are deposited in the collection of the Natural History Museum Rijeka, in the collection of the Museo Civico di Storia Naturale di Trieste, and in the collection of the Institute of Oceanography and Fisheries, Split (Gridelli, 1931; Pallaoro & Jardas, 1996; Zavodnik & Kovacic, 2000; Kovacic, unpubl. data). The species belongs to Gobius auratus species complex (Miller & El-Tawil, 1974; Herler et al., 2005) and specimens in the collections in Triest and Split need reexamination considering morphological similarities between G. auratus, G. fallax and G. xanthocephalus. Specimens collected by the small scale fishery gear at the Kornati Islands and the Murter Sea, the northern and the central Dalmatia (Cetinic & Pallaoro, 1990b; Jardas et al., 1996) were not positively identified. Benthic biocoenological researches recorded the species at the Kornati Islands and the Murter Sea (Jardas et al., 1996), and the visual census researches recorded the species at Triest and the Slovenian coastal waters, (De Girolamo et al., 1998; Turk et al., 2002; Lipej et al. 2003), without positive identification. The specimens recorded in Kovacic (1994, 1998) and Jardas et al. (1998) as G. fallax, belong to another gobiid species, G. auratus. Herler et al. (2005) collected specimens at Triest, Piran, the Cres Island and the Šolta Island. The part of these specimens is deposited in the collection of Naturhistorisches Museum Wien.
Gobius geniporus Valenciennes, 1837
Gobius geniporus: Perugia, 1866; Giglioli, 1880; Kolombatovic, 1881, 1891; Faber, 1883; Vinciguerra, 1883; Marcuzzi, 1972; Tortonese, 1975; Cetinic & Pallaoro, 1990a, 1990b; Ahnelt & Elvira, 1991; Kraljevič & Pallaoro, 1991; Kovacic, 1994, 1998, 2002b; Jardas et al., 1996, 1998; Pallaoro & Jardas, 1996; Nocita & Vanni, 1997; Guidetti, 2000; Zavod-
Marcelo KOVACIC: AN ANNOTATED CHECKLIST OF THE FAMILY GOBIIDAE IN THE ADRIATIC SEA, 21-44
nik & Kovacic, 2000; Arko Pijevac et al., 2001; Pal-
laoro, 2001; Lipej et al. 2003; Usic, 2003.
The species was first reported in the Adriatic Sea for Triest (Perugia, 1866). The species was listed for Venice, Ravenna, Triest, the Rijeka Bay, Dalmatia, the Split area, the Mljet Island, the Lastovo Island, the Korcula Island, and the Boka Kotorska Bay (Giglioli, 1880; Kolombato-vic, 1881, 1891; Faber, 1883; Vinciguerra, 1883; Kovacic, 1994; Zavodnik & Kovacic, 2000). The specimens from Triest, Rovinj, the Kvarner area, the central Adriatic, Dalmatia, Dubrovnik and Kotor are deposited in the collection of the Center for Marine Research of the Ruder Boskovic Institute in Rovinj, in the collection of the Natural History Museum Rijeka, in the collection of the Institute of Oceanography and Fisheries, Split, in the collection of Museo di Storia Naturale dell' Universita di Firenze, in the collection of the Museo Civico di Storia Naturale di Genova, in the collection of l'Istituto di Idrobiologica di Chioggia, and in the collection of the Naturhistorisches Museum Wien (Marcuzzi, 1972; Tor-tonese, 1975; Ahnelt & Elvira, 1991; Pallaoro & Jardas, 1996; Nocita & Vanni, 1997; Kovacic, 1998, unpubl. data; Usic, 2003). Specimens were collected by the small scale fishery gear at the Kornati Islands, the Murter Sea, the northern and the central Dalmatia, the Split area, and the Split Channel (Cetinic & Pallaoro, 1990a, 1990b; Kraljevic & Pallaoro, 1991; Jardas et al., 1996; Pallaoro, 2001). Benthic biocoenological researches recorded the species in the Kvarner area, at the Kornati Islands, and the Murter Sea (Jardas et al., 1996; Arko Pijevac et al., 2001), and the visual census researches recorded the species in the Slovenian coastal waters, the Rijeka Bay, and the Tremiti Islands (Jardas et al., 1998; Guidetti, 2000; Kovacic, 2002b; Lipej et al. 2003).
Gobius kolombatovici Kovacic & Miller, 2000
Gobius kolombatovici: Kovacic & Miller, 2000.
The species was described by Kovacic & Miller (2000) on four females and six males collected at four closely situated localities at the Krk Island (the Kvarner area), in June and September of 1998. The holotype and paratypes are deposited in the collection of the Natural History Museum Rijeka. Single paratype was donated to the British Museum of Natural History. Single additional female in the Adriatic Sea was collected at the Cutin Island, near the Cres Island, the Kvarner area (Kovacic, unpubl. data). It is also deposited in the collection of the Natural History Museum Rijeka.
Gobius niger Linnaeus, 1758
Gobius jozo: Brunnich, 1765; Plucar, 1846; Perugia,
1866, 1881; Giglioli, 1880;Kolombatovic, 1881,
1891; A. P. Ninni, 1882; Vinciguerra, 1883; Graeffe, 1888; Langhoffer, 1904; D'Ancona, 1922; Zei, 1942, 1949; Zavodnik, 1971; Marcuzzi, 1972. Gobius Jozo: Nardo, 1827; Martens, 1838. Gobius niger: Naccari, 1822; Nardo, 1827; Martens, 1838; Plucar, 1846; Perugia, 1866, 1881; Giglioli, 1880; A. P. Ninni, 1882; Graeffe, 1888; D'Ancona, 1922; Zei, 1942, 1949; Cavinato, 1952; Zupanovic, 1961; Tortonese, 1975; Jukic & Piccinetti, 1981; Fabi & Froglia, 1983, 1984; Jukic, 1983; Onofri, 1983; Fabi & Giannetti, 1985; Zavodnik & Zavodnik, 1986; Jardas & Pallaoro, 1989; Seiwald & Patzner, 1989; Cetinic & Pallaoro, 1990b; Zavodnik & Crnkovic, 1992; Kovacic, 1994, 1998;Jardas et al., 1996, 1998; Marconato et al., 1996; Pallaoro & Jardas, 1996; Simonovic et al., 1996;Span et al., 1996; Caputo et al., 1997; McKay & Miller, 1997; Nocita & Vanni, 1997; Orepic et al., 1997; Atkinson et al., 1998; Caputo, 1998;Simonovic, 1999; Sorice & Caputo, 1999; Zavodnik & Kovacic, 2000; Pallaoro, 2001; Mazzoldi & Rasotto, 2002;Novosel et al., 2002; Rasotto & Mazzoldi, 2002; Turk et al., 2002; Lipej et al. 2003; Usic, 2003. Gobius jozo var. nigra: Nardo, 1860. Gobius jorzo: Trois, 1875.
Gobius niger jozo: Jukic & Crnkovic, 1974; Jukic, 1975.
Gobius jozzo: Usic, 2003.
The species was first reported in the Adriatic Sea for the Split and trogir areas (Brünnich, 1765). The species was listed for Venice, the Venice Lagoon, Triest, the Rijeka Bay, the Kvarner area, Ravenna, the Zadar Channel, the Split area, Dalmatia, the Hvar Island, the Korcula Island, the Lastovo Island, and the Boka Kotorska Bay (Naccari, 1822;Nardo, 1827, 1860;Martens, 1838; Plucar, 1846; Perugia, 1866, 1881; Trois, 1875; Giglioli, 1880; Kolombatovic, 1881, 1891; A. P. Ninni, 1882; Vinciguerra, 1883;Graeffe, 1888;Langhoffer, 1904; D'Ancona, 1922; Cavinato, 1952; Kovacic, 1994; Zavodnik & Kovacic, 2000). The specimens from Venice, Triest, Ravenna, Civitanova Marche, near Rovinj, the Kvarner area, the Pag Island, the Zadar Channel, the Murter Island, the Sibenik area, the Solta Island, the Hvar Island, the Hvar Channel, the Split area, the Mljet Island, the Neretva Channel, and Dubrovnik are deposited in the collection of the Center for Marine Research of the Ruder Boskovic Institute in Rovinj, in the of the Natural History Museum Rijeka, in the collection of the Institute of Oceanography and Fisheries, Split, in the collection of the Natural History Museum of Split, in the collection of the Museo Civico di Storia Naturale di Genova, in the collection of l'Istituto di Idrobiologica di Chioggia, and in the collection of Museo di Storia Naturale dell' Universita di Firenze (Marcuzzi, 1972; Tortonese, 1975; Onofri, 1983; Pallaoro & Jardas, 1996;
Marcelo KOVACIC: AN ANNOTATED CHECKLIST OF THE FAMILY GOBIIDAE IN THE ADRIATIC SEA, 21-44
Nocita & Vanni, 1997; Kovacic, 1998, unpubl. data; Usic, 2003). Specimens were collected by the small scale fishery gear at the Kornati Islands and Murter Sea, the northern and the central Dalmatia, the Split area, and the Kastela Bay (Jardas & Pallaoro, 1989; Cetinic & Pallaoro, 1990b; Jardas et al., 1996; Pallaoro, 2001), and by the bottom trawl in the northern Adriatic, the channels of the north-eastern Adriatic Sea, the central Adriatic, the channels between the mid-Dalmatian islands, the Crnogorsko primorje (Zei, 1942, 1949; Zu-panovic, 1961; Jukic & Crnkovic, 1974; Jukic, 1975; Jukic & Piccinetti, 1981; Jukic, 1983; Span et al., 1996). Benthic biocoenological researches recorded the species in the Rovinj area, the Rasa Bay, the Losinj Island, the Velebit Channel, the Kornati Islands, the Murter Sea, and the Mljet Island (Zavodnik, 1971; Zavodnik & Zavodnik, 1986; Zavodnik & Crnkovic, 1992; Jardas et al., 1996; Orepic et al., 1997; Novosel et al., 2002) and the visual census researches recorded the species in the Slovenian coastal waters and the Rijeka Bay (Jardas et al., 1998; Turk et al., 2002; Lipej et al. 2003). The samples of G. niger were also collected in the Venice Lagoon, Auris-ina, Ancona, and the Boka Kotorska Bay (Fabi & Froglia, 1983, 1984; Fabi & Giannetti, 1985; Seiwald & Patzner, 1989; Marconato et al., 1996; Simonovic et al., 1996; Caputo et al., 1997; McKay & Miller, 1997; Atkinson et al., 1998; Caputo, 1998;Simonovic, 1999;Sorice & Caputo, 1999; Mazzoldi & Rasotto, 2002;Rasotto & Mazzoldi, 2002).
Gobius paganellus Linnaeus, 1758
Gobius Paganellus: Naccari, 1822; Nardo, 1827; Martens, 1838.
Gobius paganellus: Plucar, 1846; Nardo, 1860; Perugia, 1866, 1881; Canestrini, 1872; Trois, 1875; Gi-glioli, 1880; Kolombatovic, 1881, 1891; Perugia, 1881; A. P. Ninni, 1882, Graeffe, 1888; Langhoffer, 1904; E. Ninni, 1912; D'Ancona, 1922; Zei, 1942, 1949; Cavinato, 1952; Musin, 1989; Cetinic & Pallaoro, 1990a; Jardas et al., 1996; Pallaoro & Jardas, 1996; Simonovic et al., 1996; Caputo et al., 1997; Nocita & Vanni, 1997; Orepic et al., 1997; Caputo, 1998; De Girolamo et al., 1998;Kovacic, 1998, 2002b; Simonovic, 1999; Sorice & Caputo, 1999; Zavodnik & Kovacic, 2000; Castellarin et al., 2001; Turk et al., 2002; Lipej et al. 2003; Usic, 2003.
The species was first reported in the Adriatic Sea for Venice (Naccari, 1822). The species was listed for the Venice Lagoon, Triest, Rijeka, the Ugljan Island, the Split area, and Dalmatia (Nardo, 1827; Martens, 1838; Plucar, 1846; Nardo, 1860;Perugia, 1866, 1881; Canestrini, 1872; Trois, 1875; Giglioli, 1880; Kolombatovic, 1881, 1891; A. P. Ninni, 1882; Graeffe, 1888;Langhoffer, 1904; E. Ninni, 1912; D'Ancona, 1922; Cavinato, 1952;
Zavodnik & Kovacic, 2000). The specimens from Chiog-gia, Triest, Civitanova Marche, Rovinj, the Pag Island, the Split area, the mouth of the Neretva river, and the Dubrovnik area are deposited in the collection of the Museo Zoologico di Padova, in the collection of l'Istituto di Idrobiologica di Chioggia, in the collection of the Natural History Museum Rijeka, in the collection of the Institute of Oceanography and Fisheries, Split, in the collection of the Natural History Museum of the Biological Institute, Dubrovnik, and in the collection of Museo di Storia Naturale dell' Universita di Firenze (Musin, 1989; Pallaoro & Jardas, 1996; Nocita & Vanni, 1997; Kovacic, 1998, unpubl. data; Usic, 2003). The finding by the bottom trawl in the northern Adriatic (Zei, 1942, 1949) is quite surprising, considering dept and habitat preferences of the species. Specimens were collected by the small scale fishery gear at the Kornati Islands, the Murter Sea, and the Split Channel (Cetinic & Pallaoro, 1990a; Jardas et al., 1996). Benthic biocoenological researches recorded the species at the Kornati Islands, the Murter Sea, and the Mljet Island (Jardas et al., 1996; Orepic et al., 1997) and the visual census researches recorded the species at Tri-est, the Slovenian coastal waters and Kostrena (De Girolamo et al., 1998; Castellarin et al., 2001; Kovacic, 2002b; Turk et al., 2002; Lipej et al. 2003). The samples of G. paganellus were also collected at Ancona, Buljarica and the Boka Kotorska Bay (Simonovic et al., 1996; Caputo et al., 1997; Caputo, 1998; Simonovic, 1999; Sorice & Caputo, 1999).
Gobius roulei De Buen, 1928
Gobius roulei: Kovacic, 1995, 2001b, 2002b; Jardas
et al., 1996, 1998; Pallaoro & Jardas, 1996; Zavodnik & Kovacic, 2000; Turk et al., 2002; Lipej et al.,
2003; Usic, 2003.
Single female and seven males were collected for the first time in the Adriatic Sea at four localities in the Kvarner area in 1993 and 1994 (Kovacic, 1995). The specimens are deposited in the collection of the Natural History Museum Rijeka. The additional specimens in the Kvarner area were collected by Kovacic (2001b). The species was also recorded at numerous localities in the Kvarner area (Kovacic, 1995; Zavodnik & Kovacic, 2000). The additional findings in the Adriatic Sea are from the Gulf of Triest (the northern Adriatic) (Lipej et al., 2003), the Pag Island, the Kornati Islands, and the Murter Sea (the central Adriatic) (Jardas et al., 1996; Pallaoro & Jardas, 1996). The specimens from the Pag Island and from the Rijeka Bay are deposited in the collection of the Institute of Oceanography and Fisheries, Split and in the collection of the Center for Marine Research of the Ruder Boskovic Institute in Rovinj. Specimens were collected by the small scale fishery gear at the Kornati Islands and the Murter Sea (Jardas et al.,
Marcelo KOVACIC: AN ANNOTATED CHECKLIST OF THE FAMILY GOBIIDAE IN THE ADRIATIC SEA, 21-44
1996). Benthic biocoenological researches recorded the species at the Kornati Islands and the Murter Sea (Jardas et al., 1996), and the visual census researches recorded the species in the Slovenian coastal waters and the Ri-jeka Bay (Jardas et al., 1998; Kovacic, 2002b; Turk et al., 2002; Lipej et al. 2003).
Gobius vittatus Vinciguerra, 1883
Gobius vittatus: Kolombatovic, 1886, 1891; Kovacic,
1994, 1998, 2002b; Pallaoro & Jardas, 1996; Jardas et
al., 1996; Nocita & Vanni, 1997; Jardas et al., 1998;
Zavodnik & Kovacic, 2000; Novosel et al., 2002.
Two specimens were collected for the first time in the Adriatic Sea near Split in 1884 (Kolombatovic, 1886). The additional collected specimen and details on time and locality of collecting of the first two specimens was published later (Kolombatovic, 1891). The species was reported for the Rijeka Bay (Kovacic, 1994; Zavodnik & Kovacic, 2000). The specimens from the Kvarner area and Split are deposited in the collection of the Natural History Museum Rijeka, in the collection of the Institute of Oceanography and Fisheries, Split, and in the collection of Museo di Storia Naturale dell' Universita di Firenze (Pallaoro & Jardas, 1996; Nocita & Vanni, 1997; Kovacic, 1998, unpubl. data). Specimens were collected by the small scale fishery gear at the Kornati Islands and the Murter Sea (Jardas et al., 1996). Benthic biocoe-nological researches recorded the species at the Velebit Channel, the Kornati Islands, and the Murter Sea (Jardas et al., 1996; Novosel et al., 2002) and the visual census researches recorded the species in the Rijeka Bay (Jardas et al., 1998; Kovacic, 2002b).
Knipowitschia caucasica (Kawrajsky, 1916)
Knipowitschia caucasica: Miller, 1972c; Pallaoro &
Jardas, 1996; Kovacic & Pallaoro, 2003.
Two females, collected at Zaule (near Triest), were found in the collection of the Museo Civico di Storia Naturale di Venice and identified by Miller (1972c). Additional male was collected from the Venice Lagoon by H. Bath (Miller, 1972c). Econodimis and Miller (1990) believed that Adriatic population was "a caucasica-like form, which differs from true caucasica in body proportions, and may represent a new, unnamed species". Ko-vacic & Pallaoro (2003) confirmed previously questioned presence of this species in the Adriatic Sea and provide data on morphology and ecology of the Adriatic specimens. The specimens from the northern and central Dalmatia, Croatia: the Pag Island, the river Karisnica, the Karin Sea, the Vrana Lake, Pirovac, the mouth of the river Jadro, in the Morinj Bay, the Prokljan Lake, the spring and the mouth of the river Pantan in the Kastela
Bay, and the river Cetina are deposited in the collection of the Natural History Museum Rijeka (Kovacic & Pallaoro, 2003). The specimens from the Prokljan Lake are deposited in the collection of the Institute of Oceanography and Fisheries, Split (Pallaoro & Jardas, 1996).
Knipowitschia panizzae (Verga, 1841)
Gobius Panizzae: Verga, 1841; Trois, 1875; Giglioli,
1880; E. Ninni, 1912.
Gobius panizzae: Nardo, 1860; A. P. Ninni, 1882.
Gobius Panizzai: E. Ninni, 1938.
Pomatoschistus panizzai: Bini, 1969.
Knipowitschia panizzai: Tortonese, 1975; Gandolfi &
Tongiorgi, 1976; Nocita & Vanni, 1997.
Knipowitschia panizzae: Gandolfi, 1972; Miller,
1972c; Marconato et al., 1996; Pallaoro & Jardas,
1996; McKay & Miller, 1997; Lugli & Torricelli,
1999; Marzano & Gandolfi, 2000, 2001.
The species was described by Verga (1841) on the specimens collected at the Lago di Comacchio. The species was listed for Laguna di Venice, the Sile river, the Piave river, the Livenza river, the Po river, the rivers of Veneto, Porto Tolle, Lago di Comacchio, the river Fortore, and Laguna di Lesina (Nardo, 1860; Trois, 1875; Giglioli, 1880; A. P. Ninni, 1882; E. Ninni, 1912, 1938; Bini, 1969; Gandolfi, 1972; Miller, 1972c; Gandolfi & Tongiorgi, 1976), without positive identification. The findings at the Lago di Garda and the Krk Island were considered as erroneous (Miller, 1972c). The samples of this species were also collected in the Po delta and the Venice Lagoon, without positive identification (Gandolfi, 1972; Marconato et al., 1996; McKay & Miller, 1997; Lugli & Torricelli, 1999; Marzano & Gandolfi, 2001). The specimens from the mouth of the river Po, Caorle (Veneto), the Venice Lagoon, Triest, Civitanova Marche, and the Vrana Lake are deposited in the collection of the Museo Civico di Storia Naturale di Genova, in the collection of the Institute of Oceanography and Fisheries, Split, and in the collection of Museo di Storia Naturale dell' Universita di Firenze (Tortonese, 1975; Pallaoro & Jardas, 1996; Nocita & Vanni, 1997). All these specimens should be reexamined considering morphological similarities between K. panizzae and K. caucasica (Ko-vacic & Pallaoro, 2003).
Lebetus guilleti (Le Danois, 1913)
Lebetus guilleti: Herler & Kovacic, 2002.
Eight females and five males were collected for the first time in the Adriatic Sea at Selce and Klenovica (the Kvarner area) in 1999 and 2001 (Herler & Kovacic, 2002). The specimens are deposited in the collection of the Naturhistorisches Museum Wien and in the collec-
Marcelo KOVACIC: AN ANNOTATED CHECKLIST OF THE FAMILY GOBIIDAE IN THE ADRIATIC SEA, 21-44
tion of the Natural History Museum Rijeka. The additional finding is documented photographically at the western coast of Istria, near Rovinj in May 2002 (Herler & Kovacic, 2002).
Lesueurigobius friesii (Malm, 1874)
Gobius friesii-macrolepis: Soljan, 1948; Zupanovic & Grubisic, 1958; Zupanovic, 1961. Lesueurigobius friesii: Jukic & Crnkovic, 1974; Jukic, 1975; Jardas et al, 1981, 1996, 1998; Froglia & Gramitto, 1982; Jukic, 1983; Zavodnik & Zavodnik, 1986; Zupanovic & Jardas, 1989; Jardas, 1996b; Pallaoro & Jardas, 1996; Kovacic, 1998; Zavodnik & Kovacic, 2000; Usic, 2003.
The first positive record of this species in the Adriatic Sea was based on specimens collected at the Krk Island by the bottom trawl in 1940 (Soljan, 1948). The specimens collected near Rovinj, at the Rasa Bay, the Kvarner area, the Solta Channel, NW from the Islet of Jabuka, the Brac Island and at Budva are deposited in the collection of the Center for Marine Research of the Ruder Boskovic Institute Rovinj, in the collection of the Natural History Museum Rijeka, and in the collection of the Institute of Oceanography and Fisheries, Split (Pallaoro & Jardas, 1996; Kovacic, 1998, unpubl. data; Usic, 2003). Specimens were collected by the small scale fishery gear at the Kornati Islands and the Murter Sea (Jardas et al., 1996) and by the bottom trawl in the Kvarner area, the Rijeka Bay, south of Rogoznica, at the Jabuka Pit, the Palagruza Island, the central Adriatic, the channels between the mid-Dalmatian islands and at Crnogorsko primorje (Zupanovic & Grubisic, 1958; Zupanovic, 1961; Jukic & Crnkovic, 1974; Jukic, 1975, 1983; Jardas et al., 1981, 1998; Froglia & Gramitto, 1982; Zupanovic & Jardas, 1989; Zavodnik & Kovacic, 2000). Benthic biocoenological researches recorded the species at the Rasa Bay, and in the Murter Sea (Zavodnik & Zavodnik, 1986; Jardas, 1996b).
Lesueurigobius suerii (Risso, 1810)
Gobius Lunieus: Chiereghini, 1818. Gobius lunie: Nardo, 1827. Gobius lunie: Nardo, 1860. Gobius Lesueuri: Kolombatovic, 1881. Gobius lesueuri: Kolombatovic, 1882, 1891; Zupanovic, 1961; Crnkovic, 1970; Zavodnik, 1971; Jukic & Crnkovic, 1974.
Gobius Lesueurii: Vinciguerra, 1883; Langhoffer, 1904.
Gobius lesueurii: A. P. Ninni, 1882; E. Ninni, 1912. Lesueurigobius suerii: Zavodnik & Crnkovic, 1992; Jardas et al., 1996, 1998; Usic, 2003; Ahnelt & Dor-da, 2004.
Lesueurigobius sueri: Tortonese, 1975; Nocita &
Vanni, 1997; Sorice & Caputo, 1999.
The species was recorded for the first time in the Adriatic Sea for the Venice Lagoon (Chiereghini, 1818, cited in Ninni, 1938). The species was listed under various synonyms for the Venice Lagoon, the Zadar Channel, the Split area, and the Boka Kotorska Bay (Nardo, 1827, 1860; Kolombatovic, 1881, 1882, 1891; A. P. Ninni, 1882; Vinciguerra, 1883;Langhoffer, 1904;E. Ninni, 1912). The specimens collected near Rovinj, from Dalmatia, the Sv. Andrija Island, and Bari are deposited in the collection of the Center for Marine Research of the Ruder Boskovic Institute in Rovinj, in the collection of the Naturhistorisches Museum Wien, in the collection of the Museo Civico di Storia Naturale di Genova, and in the collection of Museo di Storia Naturale dell' Universita di Firenze (Tortonese, 1975; Nocita & Vanni, 1997; Usic, 2003; Ahnelt & Dorda, 2004; Zavodnik, pers. comm.). Specimens were collected by the small scale fishery gear at the Kornati Islands and the Murter Sea (Jardas et al., 1996) and by the bottom trawl in the channels of the north-eastern Adriatic Sea, near Ro-goznica, the channels between the mid-Dalmatian islands, and Crnogorsko primorje (Zupanovic, 1961; Crnkovic, 1970; Jukic & Crnkovic, 1974). Benthic bio-coenological researches recorded the species at Rovinj and in the Kvarner area (Zavodnik, 1971; Jardas et al., 1998; Zavodnik & Crnkovic, 1992). The samples of this species were also collected at Ancona (Sorice & Caputo, 1999).
Millerigobius macrocephalus (Kolombatovic, 1891)
Gobius macrocephalus: Kolombatovic, 1891.
Millerigobius macrocephalus: Bath, 1973; Turk et al.,
2002; Lipej et al. 2003.
The species was described by Kolombatovic (1891) on single specimen collected at the Brac Island (the central Adriatic) in 1887. Bath (1973) redescribed the species on four males collected at Medulin and the Lim-ski Channel in Istria, in July 1972. The neotypes are deposited in the collection of the Senckenberg Naturmuseum, Frankfurt. The additional specimens in the Adriatic Sea were collected at the Solta Island, the central Adriatic (Kovacic, unpubl. data). They are deposited in the collection of the Natural History Museum Rijeka. Visual census research recorded the species in the Slovenian coastal waters (Turk et al., 2002; Lipej et al. 2003).
Odondebuenia balearica (Pellegrin & Fage, 1907)
Gobius liechtensteini: Kolombatovic, 1891 (part.).
Odondebuenia balearica: Miller & Tortonese, 1968;
Marcelo KOVACIC: AN ANNOTATED CHECKLIST OF THE FAMILY GOBIIDAE IN THE ADRIATIC SEA, 21-44
Ahnelt et al., 1994; Jardas et al., 1996; Pallaoro &
Jardas, 1996.
Single specimen collected at Split and four specimens collected at the Korcula Island (the central Adriatic) were found and identified by Miller & Tortonese (1968) in the collection of the Museo Zoologico, Universita di Firenze and in the collection of the Naturhistorisches Museum Wien. The additional specimens in the Adriatic Sea were collected at Rovinj (the Istrian peninsula); Urinj, Bakar, Ostro, Klenovica, the Krk Island, the Cutin Island (the Kvarner area); Split, Stobrec, the Solta Island, the Hvar Island, Mala Duba, the Bisevo Island, and the Palagruza Island (the central Adriatic); Mljet (the southern Adriatic) (Ahnelt et al., 1994; Pallaoro & Jardas, 1996; Kovacic, unpubl. data). They are deposited in the collection of the Natural History Museum Rijeka, in the collection of the Institute of Oceanography and Fisheries, Split and in the collection of the Naturhistorisches Museum Wien. The species was recorded in benthic biocoenological research at the Kornati Islands (the central Adriatic) (Jardas et al., 1996).
Pomatoschistus bathi Miller, 1982
Pomatoschistus bathi: Miller, 1982; Ahnelt et al.,
1994; Pallaoro & Jardas, 1996; Lipej et al., 2003.
Single male and twelve females from the private collection of H. Bath were identified by Miller (1982). The specimens were collected in the Bay of Kotor (the southern Adriatic) on May 10 1969. The additional Adriatic specimens were collected at the Krk Island, the Cutin Island, the Losinj Island (the Kvarner area); Seline (the Velebit Channel); the mouth of the river Zrmanja, the Prokljan Lake, the Murter Island, the Morinje Cove, the Brac Island, Blace (the central Adriatic); the Mljet Island (the southern Adriatic) (Ahnelt et al., 1994; Pallaoro & Jardas, 1996; Kovacic, unpubl. data). They are deposited in the collection of the Natural History Museum Rijeka and in the collection of the Institute of Oceanography and Fisheries, Split. Visual census researches recorded the species in the Slovenian coastal waters (the northern Adriatic) (Lipej et al., 2003).
Pomatoschistus canestrinii (Ninni, 1883)
Gobius quagga: Kolombatovic, 1881.
Gobius Canestrinii: A. P. Ninni, 1883.
Gobius Canestrini: Kolombatovic, 1888; E. Ninni,
1938.
Gobius canestrini: Kolombatovic, 1891; Cavinato,
1952.
Pomatoschistus canestrini: Bini, 1969; Tortonese,
1975; Mrakovcic et al., 1994; Lugli & Torricelli,
1999.
Pomatoschistus canestrinii: Gandolfi et al., 1982;
Pallaoro & Jardas, 1996;Nocita & Vanni, 1997;
McKay & Miller, 1997.
The species was described by A. P. Ninni (1883) on the specimens collected by Kolombatovic at the Jadro river. The species was mentioned from the type locality earlier, as a form of Gobius quagga, also by Kolombatovic (1881). The additional collected specimens were studied for the Split area by Kolombatovic (1888, 1891). The species was listed for the Venice Lagoon, Piave river and Livenza river (E. Ninni, 1938; Cavinato, 1952; Bini, 1969). The specimens from the Venice Lagoon, Triest, the Zrmanja river and its tributary Dobarnica, the Krka river, the Jadro river, the Zrnovnica river, the Cetina river, the Bacina Lakes, and the Neretva river are deposited in the collection of the Natural History Museum Ri-jeka, in the collection of the Museo Civico di Storia Naturale di Genova, in the collection of the Institute of Oceanography and Fisheries, Split, and in the collection of Museo di Storia Naturale dell' Universita di Firenze (Tortonese, 1975; Pallaoro & Jardas, 1996; Nocita & Vanni, 1997; Kovacic, unpubl. data). The samples of this species were also collected at the mouth of the Taglia-mento river, the mouth of the Stella river, the mouth of the Livenza river, the mouth of the Dese river, the Laguna del Basson, the Venice Lagoon, the mouth of river Po, and in the Zrmanja river (Gandolfi et al., 1982; Mrakovcic et al., 1994; McKay & Miller, 1997; Lugli & Torricelli, 1999).
Pomatoschistus knerii (Steindachner, 1861)
Gobius Knerii: Steindachner, 1861; Langhoffer,
1904.
Gobius Kneri: Giglioli, 1880.
Gobius knerii: Kolombatovic 1893.
Gobius steindachnerii: Kolombatovic 1900.
Pomatoschistus knerii: Jardas et al., 1996, 1998; Kovacic, 1998, 2003; Zavodnik & Kovacic, 2000.
The species was described by Steindachner (1861) on the specimens collected at the Hvar Island. The additional specimens were reported at Venice, Zaola (near Triest), the Kornati Islands, and at Seget (near Split) (Gi-glioli, 1880; Kolombatovic 1893, 1900; A. P. Ninni, 1882; Langhoffer, 1904). The specimens from the Kvarner area and the Zakan Island are deposited in the collection of the Natural History Museum Rijeka (Jardas et al., 1998; Kovacic, 1998, 2003, unpubl. data; Zavodnik & Kovacic, 2000). Specimens were collected by the small scale fishery gear at the Kornati Islands and in the Murter Sea (Jardas et al., 1996).
Marcelo KOVACIC: AN ANNOTATED CHECKLIST OF THE FAMILY GOBIIDAE IN THE ADRIATIC SEA, 21-44
Pomatoschistus marmoratus (Risso, 1810)
Gobius ferrugineus: Kolombatovic, 1891.
Gobius marmoratus: Ninni, 1938; Cavinato, 1952.
Pomatoschistus marmoratus: Bini, 1969; Pallaoro &
Jardas, 1996; Nocita & Vanni, 1997; Caputo, 1998;
Kovacic, 1998; Lugli & Torricelli, 1999; Mazzoldi &
Rasotto, 2001; Mazzoldi et al., 2002; Turk et al.,
2002; Lipej et al. 2003; Usic, 2003.
The species was first reported in the Adriatic Sea for the Split area under the synonym Gobius ferrugineus (Kolombatovic, 1891). The species was listed for Venice (E. Ninni, 1938;Cavinato, 1952; Bini, 1969). The specimens from Chioggia, the Venice Lagoon, the Triest area, the Kvarner area, the Pag Island, the Zrmanja river, the Karin Sea, the Zadar area, the Zakan Island, the Biograd area, the Sibenik area, the Trogir area, the mouth of the river Zrnovnica, Omis, the Ombla river, and Bari are deposited in the collection of the Museo Zoologico di Padova, in the collection of the Natural History Museum Rijeka, in the collection of the Institute of Oceanography and Fisheries, Split, and in the collection of Museo di Storia Naturale dell' Universita di Firenze (Pallaoro & Jardas, 1996;Nocita & Vanni, 1997; Kovacic, 1998, unpubl. data). Visual census researches recorded the species in the Slovenian coastal waters (Turk et al., 2002; Lipej et al. 2003). The samples of this species were also collected in the Po delta, in the Venice Lagoon, and at Ancona (Caputo, 1998; Lugli & Torricelli, 1999; Mazzoldi & Rasotto, 2001; Mazzoldi et al., 2002; Usic, 2003).
Pomatoschistus minutus (Pallas, 1770)
Gobius minutus: Plucar, 1846; Perugia, 1866, 1881;
Trois, 1875; Giglioli, 1880;Kolombatovic, 1881;
Graeffe, 1888; E. Ninni, 1938; Cavinato, 1952; Mar-
cuzzi, 1972; Gamulin-Brida et al., 1980.
Gobius minutus elongatus: Zupanovic, 1961.
Pomatoschistus minutus: Nocita & Vanni, 1997;
Caputo, 1998; Sorice & Caputo, 1999; Stefanni et al.,
2003; Usic, 2003.
The species was recorded for the first time in the Adriatic Sea for the Triest area (Plucar, 1846). The species was listed for Venice, Triest, and the Split area (Perugia, 1866, 1881; Trois, 1875; Giglioli, 1880; Kolombatovic, 1881, 1891; Graeffe, 1888;E. Ninni, 1938; Cavinato, 1952). The specimens from the Venice Lagoon, the Triest area, the Kvarner area, the mouth of the river Zrmanja and the mouth of the river Neretva are deposited in the collection of Museo di Storia Naturale dell' Universita di Firenze, in the collection of l'Istituto di Idrobiologica di Chioggia, and in the collection of the Natural History Museum Rijeka (Marcuzzi, 1972; Nocita
& Vanni, 1997; Kovacic, unpubl. data; Usic, 2003). Specimens were collected by the bottom trawl between the mid-Dalmatian islands (Zupanovic, 1961), without a positive identification. Benthic biocoenological research recorded the species at the Krk Island (Gamulin-Brida et al., 1980), without positive identification. The samples of this species were recently collected at Venice and Ancona (Caputo, 1998; Sorice & Caputo, 1999; Stefanni et al., 2003).
Pomatoschistus norvegicus adriaticus (Miller, 1972)
Pomatoschistus norvegicus: Stefanni, 2000.
Four females and a single male were collected for the first time in the Adriatic Sea of the littoral of Venice in March 1998 (Stefanni, 2000).
Pomatoschistus pictus (Malm, 1865)
Gobius affinis: Kolombatovic, 1891.
Pomatoschistus pictus adriaticus: Miller, 1972a;
Zander & Jelinek, 1976.
The species was recorded for the first time in the Adriatic Sea for the Venice Lagoon (E. Ninni, 1938). Miller (1972a) discovered that syntypes of Gobius affinis Kolombatovic, 1891 from the Split area in the collection of the Naturhistorischen Museum, Wien belong indeed to Pomatoschistus pictus. Miller (1972a) described the subspecies of this species, Pomatoschistus pictus adriati-cus on specimens from the Split area and on the specimens from the private collection of H. Bath collected at Triest. The samples of P. pictus were also collected at Rovinj (Zander & Jelinek, 1976).
Pomatoschistus quagga (Heckel, 1840)
Gobius quagga: Giglioli, 1880; Kolombatovic, 1881,
1882, 1891; A. P. Ninni, 1882; Vinciguerra, 1883;
Damiani, 1896; Langhoffer, 1904.
Pomatoschistus quagga: Tortonese, 1975; Jardas et
al., 1996, 1998; Pallaoro & Jardas, 1996; Nocita &
Vanni, 1997; De Girolamo et al., 1998; Zavodnik &
Kovacic, 2000; Kovacic, 2003.
The species was recorded for the first time in the Adriatic Sea in the Split area (Giglioli, 1880). The species was listed for Venice, the Rijeka Bay, the Kvarner area, the Zadar Channel, the Split area, the Hvar Island, and the Boka Kotorska Bay (Kolombatovic, 1881, 1882, 1891; A. P. Ninni, 1882; Vinciguerra, 1883; Damiani, 1896; Langhoffer, 1904; Zavodnik & Kovacic, 2000). The specimens from the Kvarner area, the Zakan Island, the central Adriatic, Split and the Boka Kotorska Bay are deposited in the collection of the Natural History Mu-
Marcelo KOVACIC: AN ANNOTATED CHECKLIST OF THE FAMILY GOBIIDAE IN THE ADRIATIC SEA, 21-44
seum Rijeka, in the collection of the Museo Civico di Storia Naturale di Genova, in the collection of the Institute of Oceanography and Fisheries, Split and in the collection of Museo di Storia Naturale dell' Universita di Firenze (Tortonese, 1975; Pallaoro & Jardas, 1996; No-cita & Vanni, 1997; Kovacic, 2003, unpubl. data). Specimens were collected by the small scale fishery gear at the Kornati Islands and in the Murter Sea (Jardas et al., 1996). Visual census researches recorded the species at Triest and the Rijeka Bay (De Girolamo et al., 1998; Jardas et al., 1998), without positive identification.
Pseudaphya ferreri (De Buen & Fage, 1908)
Gobius pusillus: Kolombatovic, 1891.
Pseudaphya ferreri: Miller, 1973b; Kovacic, 2003.
Four males and eight females, collected by Kolom-batovic as Gobius pusillus at Split (the central Adriatic), were found and identified by Miller (1973b) in the collection of the Naturhistorisches Museum Wien. The additional specimens collected at Ostro, Kacjak, and Sv. Marak (the Kvarner area) are deposited in the collection of the Natural History Museum Rijeka (Kovacic, 2003, unpubl. data).
Speleogobius trigloides Zander & Jelinek, 1976
Speleogobius trigloides: Zander & Jelinek, 1976;
Fesser, 1980; Kovacic, 1997, 2002b.
The holotype was collected at the Banjole cave, near Rovinj, the Istrian Peninsula, in June 1975. It is deposited in the collection of the Zoologisches Institut und Zoologisches Museum der Universität Hamburg. The additional findings were reported from Hvar Island, the central Adriatic, in 1974 and 1975, and Prvic Island in 1977 and 1978 by Fesser (1980). Three females and two males from the Prvic Island (the Kvarner area) are deposited in the collection of the Naturhistorisches Museum Wien. The specimens collected from Zurkovo, the Goli Island, the Prvic Island, Bakar, the Krk Island (the Kvarner area) in the period from 1997 to 2001 are deposited in the collection of the Natural History Museum Rijeka (Kovacic, 1997, 2002b, unpubl. data; Zavodnik & Kovacic, 2000).
Thorogobius ephippiatus (Lowe, 1839)
Thorogobius ephippiatus: Miller, 1969; Shultz, 1975;
Kovacic, 1994, 1997, 1998; Ahnelt & Kovacic, 1997;
Jardas et al., 1998; Zavodnik & Kovacic, 2000; Arko
Pijevac et al., 2001; Novosel et al., 2002.
The underwater photograph of this species was taken at the Prvic Island (the Kvarner area) and published in
Riedl (1966) without identification. However, Miller (1969) first identified this species in the Adriatic Sea based on underwater photograph in Riedl (1966) taken at the Prvic Island and sight-record in the Dubrovnik area on August 15 1968. The species was recorded at Banjole near Rovinj in 1962 and collected at the Hvar Island in 1969 and 1970 (Shultz, 1975). The specimens from the Kvarner area are deposited in the collection of the Natural History Museum Rijeka (Kovacic, 1998, unpubl. data). The species was also observed at numerous localities in the Kvarner area (Kovacic, 1994, 1997, unpubl. data; Ahnelt & Kovacic, 1997; Jardas et al., 1998; Zavodnik & Kovacic, 2000). Benthic biocoenological research recorded the species in the Kvarner area and in the Velebit Channel (Arko Pijevac et al., 2001; Novosel et al., 2002).
Thorogobius macrolepis (Kolombatovic, 1891)
Gobius macrolepis: Kolombatovic, 1891.
Thorogobius macrolepis: Miller, 1969; Pallaoro &
Jardas, 1996; Ahnelt & Kovacic, 1997; Kovacic,
1998; Jardas et al., 1998; Zavodnik & Kovacic, 2000;
Arko Pijevac et al., 2001; Novosel et al., 2002.
The species was described on specimens collected in the Split area by Kolombatovic (1891). Two syntypes are deposited in the collection of the Naturhistorisches Museum Wien (Miller, 1969; Ahnelt & Kovacic, 1997). Specimens from the Kvarner area are deposited in the collection of the Natural History Museum Rijeka and in the collection of the Institute of Oceanography and Fisheries, Split (Pallaoro & Jardas, 1996; Ahnelt & Kovacic, 1997; Kovacic, 1998, unpubl. data). The species was also observed at numerous localities in the Kvarner area (Ahnelt & Kovacic, 1997; Jardas et al., 1998; Zavodnik & Kovacic, 2000). Benthic biocoenological researches recorded the species in the Kvarner area and in the Velebit Channel (Arko Pijevac et al., 2001; Novosel et al., 2002). The record of Novosel et al. (2002) was without positive identification of the species. The specimens recorded in Kovacic (1994) belong to another gobiid species, Gobius roulei.
Vanneaugobius dollfusi (Brownell, 1978)
Vanneaugobius pruvoti: Jardas, 1996a.
Vanneaugobius dollfusi: Pallaoro & Kovacic, 2000;
Ahnelt & Dorda, 2004.
Single male collected near Split in 1931 and single female collected in the Drvenik Channel (the central Adriatic), in 1948, both deposited in the collection of the Institute of Oceanography and Fisheries, Split were identified by Pallaoro & Kovacic (2000). Authors also reported a single female collected near the Mljet Island
Marcelo KOVACIC: AN ANNOTATED CHECKLIST OF THE FAMILY GOBIIDAE IN THE ADRIATIC SEA, 21-44
and a single juvenile collected by D. Zavodnik near the Palagruza Island in 1998; both deposited in the collection of the Natural History Museum Rijeka. Additional specimens from the Island Palagruza, and locality between the Vis Island and the Bisevo Island were found in the collection of the Naturhistorisches Museum Wien and identified by Ahnelt & Dorda (2004).
Zebrus zebrus (Risso, 1826)
Gobius zebrus: Trois, 1875; Kolombatovic, 1881, 1891; A. P. Ninni, 1882; E. Ninni, 1912. Gobius Zebrus: Perugia, 1881. Zebrus zebrus: Tortonese, 1975; Miller, 1977; Patzner et al, 1991; Kovacic, 1994;1997, 1998, 2002b; Jardas et al., 1996, 1998; Pallaoro & Jardas, 1996; Nocita & Vanni, 1997; Zavodnik & Kovacic, 2000; Castellarin et al., 2001; Turk et al., 2002; Lipej et al. 2003.
The species was first reported for the Adriatic Sea by Trois (1875), without any notice on collection locality. The first recorded localities in the Adriatic Sea were Zaole and Servola near Triest (Perugia, 1881) and the Split area (Kolombatovic, 1881). Additional specimens were collected at Venice by A. P. Ninni (1882) and in the Split area by Kolombatovic (1891). The species was noted as common at Venice (E. Ninni, 1912). Specimens from Venice, Triest, Medulin, the Kvarner area, the mouth of the river Zrmanja, Pirovac, the Solta Island, the Kastela Bay and Mala Duba are deposited in the collection of Museo Civico di Storia Naturale di Venezia, in the collection of the Natural History Museum Rijeka, in the collection of the Museo Civico di Storia Naturale di Genova, in the collection of the Institute of Oceanography and Fisheries, Split, and in the collection of Museo di Storia Naturale dell' Universita di Firenze (Tortonese, 1975; Miller, 1977; Pallaoro & Jardas, 1996; Nocita & Vanni, 1997; Kovacic, 1998, unpubl. data). The species was also noted at numerous localities in the Kvarner area (Kovacic, 1994, 1997; Jardas et al., 1998; Zavodnik & Kovacic, 2000). Benthic biocoenological research recorded the species at the Kornati Islands and the Murter Sea (Jardas et al., 1996). Visual census researches recorded the species at Triest, the Slovenian coastal waters, and at Kostrena (Patzner et al., 1991; Castellarin et al., 2001; Kovacic, 2002b; Turk et al., 2002; Lipej et al. 2003).
Zosterisessor ophiocephalus (Pallas, 1811)
Gobius venetiarum: Nardo, 1860, Trois, 1875. Gobius lota: Canestrini, 1872; Graeffe, 1888. Gobius ophiocephalus: Giglioli, 1880; Kolombatovic, 1881, 1891; A. P. Ninni, 1882; Vinciguerra, 1883; Langhoffer, 1904; Cavinato, 1952; Marcuzzi, 1972; E. Ninni, 1912, 1938; Usic, 2003.
Zosterisessor ophiocephalus: Tortonese, 1975; Bal-estra et al., 1989; Mušin, 1989; Kraljevic & Pallaoro, 1991; Lahnsteiner et al., 1992; Zavodnik & Crnkovic, 1992; Giulianini et al., 1994; Caputo et al., 1996, 1997; Jardas et al., 1996; Marconato et al., 1996; Ota et al., 1996; Ota & Lahnsteiner, 1996; Pallaoro & Jardas, 1996; McKay & Miller, 1997; Orepic et al., 1997; Caputo, 1998;Kovacic, 1998;Ota et al., 1999; Scaggiante et al., 1999; Sorice & Caputo, 1999; Marchesan et al., 2000; Mazzoldi et al., 2000; Torricelli et al., 2000; Pallaoro, 2001; Franco et al., 2002; Malavasi et al., 2002, 2003; Usic, 2003; Dul-cic, 2004.
Zosterissessor ophiocephalus: Nocita & Vanni, 1997.
The species was recorded for the first time in the Adriatic Sea at Venice (Nardo, 1860). The species was listed under various synonyms for the Venice Lagoon, Triest, the Krk Island, the Ugljan Island, the Murter Island, the Split area, the Šolta Island, the Brac Island, the Hvar Island, the Vis Island, the Mljet Island, the Lastovo Island, the Korcula Island, and the Boka Kotorska Bay (Canestrini, 1872; Trois, 1875; Giglioli, 1880; Kolombatovic, 1881, 1891; A. P. Ninni, 1882; Vinciguerra, 1883; Graeffe, 1888; Langhoffer, 1904; E. Ninni, 1912, 1938; Cavinato, 1952). The specimens from Chioggia, the Venice Lagoon, Triest, Rovinj, the Pag Island, the Šibenik area, Dalmatia, the Split area, the Šolta Island, the Dubrovnik area, the mouth of the river Neretva, and Bari are deposited in the collection of the Museo Zoologico di Padova, in the collection of the Center for Marine Research of the Ruder Boškovic Institute in Rovinj, in the collection of the Natural History Museum Rijeka, in the collection of the Institute of Oceanography and Fisheries, Split, in the collection of the Natural History Museum of the Biological Institute, Dubrovnik, in the collection of l'Istituto di Idrobiologica di Chioggia, and in the collection of Museo di Storia Naturale dell' Universita di Firenze (Marcuzzi, 1972; Tortonese, 1975; Mušin, 1989; Pallaoro & Jardas, 1996; Nocita & Vanni, 1997; Kovacic, 1998; Usic, 2003). Specimens were collected by the small scale fishery gear at the Kornati Islands, the Murter Sea, and the Split area (Kraljevic & Pallaoro, 1991; Jardas et al., 1996;Pallaoro, 2001). Benthic biocoenological researches recorded the species at the Lošinj Island, the Kornati Islands, the Murter Sea, and the Mljet Island (Zavodnik & Crnkovic, 1992; Jardas et al., 1996; Orepic et al., 1997). Visual census research recorded the species in the Slovenian coastal waters (Turk et al., 2002). The samples of this species were also collected in the Venice Lagoon, near Grado, near Triest, at Ancona, and in the Karin Sea (Balestra et al., 1989; Lahnsteiner et al., 1992; Giulianini et al., 1994; Caputo et al., 1996, 1997; Marconato et al., 1996; Ota et al., 1996; Ota & Lahnsteiner, 1996; McKay & Miller, 1997; Caputo, 1998; Ota et al., 1999; Scaggiante et al., 1999;
Marcelo KOVACIC: AN ANNOTATED CHECKLIST OF THE FAMILY GOBIIDAE IN THE ADRIATIC SEA, 21-44
Sorice & Caputo, 1999; Marchesan et al., 2000; Maz-zoldi et al., 2000; Torricelli et al., 2000; Franco et al., 2002; Malavasi et al., 2002, 2003; Dulcic, 2004).
SPECIES EXCLUDED FROM THE CHECKLIST
The following species listed for the Adriatic Sea in the last published checklist by Jardas (1996a), should be exluded from the checklist for the Adriatic Sea:
Gobius luteus Kolombatovic, 1891
Kolombatovic (1891) described the variant of G. auratus as G. auratus v. lutea. Miller & El-Tawil (1974) raised this variant to species level as G. luteus. Heymer & Zander (1992) described new species, Gobius xantho-cephalus, from western Mediterranean. Authors agreed with Miller (1973a) that G. auratus v. ruginosa was G. fallax and concluded that the variant lutea was not a separate species, but the typical form of G. auratus (Heymer & Zander, 1992). Consequently, G. luteus is a junior synonym of G. auratus, and not a valid species.
Gobius strictus Fage, 1907
Single female collected at the Korcula Island (the central Adriatic) was found and identified by Miller (1967) as Gobius schmidti in the collection of the Museo Civico di Storia Naturale di Genova. Miller (1973a) listed Gobius schmidti by Miller (1967) as a junior synonym of Gobius strictus Fage, 1907. The later suggestion of Miller (1986) that specimens of this species could be juveniles of G. cruentatus was confirmed by Kovacic (2004). Therefore, G. strictus is a junior synonym of G. cruentatus, and not the valid species.
Gobiusculus flavescens (Fabricius, 1779)
Nardo (1860) listed Gobius ruthensparii (error for ruuthensparri) for the Venice area. Perugia (1866) listed Gobius ruthensparri for the Triest area without any data for positive identification. Gobius ruthensparii was cited by rote for the Adriatic Sea (Canestrini, 1872; Trois, 1875; Giglioli, 1880; Stossich, 1880; Perugia, 1881; Faber, 1883; Carus, 1893) with these or different spelling errors. A. P. Ninni, (1882) excluded this species from his catalogues of gobies of the Adriatic Sea. Damiani (1896) suspected that G. Ruthensparri (error for ruuthensparri) was doubtful citation by Nardo, since the species is known from the northern Atlantic. E. Ninni (1912) and Soljan (1948, 1965) excluded this synonym from their lists of the Adriatic fishes. E. Ninni (1938) was convinced in wrong identification of G. ruthensparii by Nardo. The species was listed for the first time under the valid synonym Gobiusculus flavescens (Fabricius, 1779) by Bini (1969). This was again a citation based only on
the list published by Nardo (I860). However, a new era of citations of this species for the Adriatic Sea began with Bini (1969), following the valid name (Tortonese, 1975; Stevcic, 1977;Jardas, 1985, 1996a;Kovacic, 1994). Miller (1986) listed this species just for the eastern Atlantic. According to presented data, there is no evidence for the presence of G. flavescens in the Adriatic Sea.
Pomatoschistus microps (Kroyer, 1838)
E. Ninni (1938) and Cavinato (1952) recorded this species for the Venice area. All subsequent notices of this species for the Adriatic Sea (Soljan, 1948, 1965; Bini, 1969; Vukovic i Ivanovic, 1971; Tortonese, 1975; Stevcic, 1977; Jardas, 1985, 1996a; Kovacic, 1994) were based on the original data by E. Ninni (1938) and Cavinato (1952). Miller (1972c, 1973a) suggested that descriptions by E. Ninni (1938) and Cavinato (1952) ref-ered to Knipowitschia species. Miller (1986) and Ahnelt (1991) listed this species just for the eastern Atlantic and the northwestern coast of Mediterranan. Specimens collected by the bottom trawl between the mid-Dalmatian islands were undoubtedly wrongly identified as Gobius microps laticeps (Zupanovic, 1961). According to presented data, the specimens identified as P. microps in the Venice area, belong to Knipowitschia species.
Pomatoschistus tortonesei (Miller, 1968)
The species was mentioned for the first time for the Adriatic Sea by Stevcic (1977), based on personal communication from Miller on specimens from the Boka Kotorska Bay. All later citations of this species for the Adriatic Sea (Jardas, 1985, 1996a; Kovacic, 1994) were based on data by Stevcic (1977). Miller (1982) described Pomatoschistus bathi, and redecribed P. tortonesei. The specimens from the Boka Kotorska Bay were identified as P. bathi. All specimens of P. tortonesei studied for redescription of the species (Miller, 1982) were not collected in the Adriatic Sea. Miller (1986) listed P. tortonesei just for the central Mediterranean. According to the above data, the specimens identified in the first place as P. tortonesei from the Boka Kotorska Bay (Stevcic, 1977), belong to P. bathi.
Vanneaugobius pruvoti (Fage, 1907)
Single male collected near Split in 1931 and single female collected in the Drvenik Channel (the central Adriatic) in 1948 deposited in the collection of the Institute of Oceanography and Fisheries, Split were rei-dentified as Vanneaugobius dollfusi Brownell, 1978 by Pallaoro & Kovacic (2000).
Marcelo KOVACIC: AN ANNOTATED CHECKLIST OF THE FAMILY GOBIIDAE IN THE ADRIATIC SEA, 21-44
DISCUSSION
The sources of data for this checklist of the Adriatic Sea gobies were ichthyological lists containg original collection data, papers on taxonomy and zoogeography of gobies, fisheries papers, benthic biocoenological papers, papers on fish visual census and papers on various researches that used samples of Adriatic gobies. Studies on taxonomy and zoogeography of gobies were by the far most important contributions to the checklist among all these sources of data. Eight species were added to the last published list (Jardas, 1996a): Didogobius spletch-nai, Gammogobius steinitzi, Gobius ater, Gobius cou-chi, Gobius kolombatovici, Lebetus guiletti, Pomatos-chistus norvegicus, and Vanneaugobius dollfusi (Kovacic, 1999, 2001a; Kovacic & Miller, 2000; Pallaoro & Kovacic, 2000; Stefanni, 2000; Ahnelt, 2001; Herler & Patzner, 2002; Herler & Kovacic, 2002). All these species, except of Gobius ater, were included in new fish records for the Adriatic Sea by Lipej & Dulcic (2004).
Six species were excluded from the checklist of the Adriatic gobies, based on evidence referred in the present review. Some other errors were also found in the previous checklists. The doubtful status of K. caucasica in the Adriatic Sea, questioned by Econodimis & Miller (1990), and resolved by Kovacic & Pallaoro (2003), was overlooked by checklists published in the meantime. Absence of evidence on the presence of B. affinis (Miller, 1972a) was also ignored by later reviews, until true specimens of B. affinis were found for the first time in the Adriatic Sea thirty years later (Kovacic, 2002b). The confusion on Vanneaugobius was the briefest one, it lasted for only five years (Jardas, 1996a; Pallaoro & Kovacic, 2001). The presented checklist contains 46 gobiid species recorded in the Adriatic Sea up to the present date. Four Adriatic gobies could still be considered Adriatic endemic species: Gobius kolombatovici, Kni-powitschia panizzae, Pomatoschistus canestrinii and Speleogobius trigloides. Today 59 species of Gobiidae are known to occur in the Mediterranean sensu stricto if we exlude Gobius strictus and Gobius luteus (Heymer & Zander, 1992; Kovacic, 2004) from 61 species listed for the Mediterranean (Quignard & Tomasini, 2000; Ahnelt & Dorda, 2004). The thirteen gobies recorded in the Mediterranean, and not found in the Adriatic Sea, are the Atlantic species (7 species), the Red Sea invaders (3 species), and the Mediterranean endemic species (3 species).
Three valid gobiid genera, twelve valid gobiid spe-
cies and two valid subspecies were described on the Adriatic specimens (Verga 1841; Steindachner, 1861; Steindachner, 1863; Steindachner, 1870; A. P. Ninni, 1883; Kolombatovic, 1891; Miller, 1969, 1971, 1972a, 1972b; Bath, 1973;Zander & Jelinek, 1976; Miller, 1992; Kovacic & Miller, 2000). The most fruitfull periods for Adriatic gobiology were from 1860 to 1900 with eight newly described species and ten first findings for the Adriatic Sea, and from 1968 to 2002 with three newly described species and fifteen first findings for the Adriatic Sea. The largest contributions were provided by Kolombatovic (four species description and three first findings for the Adriatic Sea), Steindachner (three species description and one first finding for the Adriatic Sea), Miller (two species description and five first findings for the Adriatic Sea), and Kovacic (one species description and six first findings for the Adriatic Sea). The locations of eight gobiid types (holotype, syntypes or neotypes) from the Adriatic Sea are known from published sources (Tortonese, 1963; Miller, 1969, 1972b, 1973a, 1992; Bath, 1973; Zander & Jelinek, 1976; Kovacic & Miller, 2000). Adriatic types of the three species are deposited in the collection of the Naturhistorisches Museum Wien, and types of the one species in each of the following collections: the collection of the Museo Civico di Storia Naturale di Genova, the collection of the Senckenberg Naturmuseum, Frankfurt, the collection of the Zoologisches Institut and Zoologisches Museum der Universität Hamburg, the collection of the Zoologische Staatssammlung, München, and the collection of the Natural History Museum Rijeka. The richest collections in Adriatic gobiid species are the collection of the Natural History Museum Rijeka (39 Adriatic species), the collection of the Institute of Oceanography and Fisheries, Split (28 Adriatic species), the collection of Museo di Storia Naturale dell' Universita di Firenze (20 Adriatic species), and the collection of the Naturhistorisches Museum Wien (13 Adriatic species).
ACKNOWLEDGEMENTS
I am grateful to D. Zavodnik for criticism and comments on this paper. I thank D. Caleta, Z. Modric, M. Vucemilovic and D. Zavodnik for help in search for literature. This paper is the second publication of the dissertation of the author, which was carried out at the University of Zagreb, Croatia under the supervision of I. Jardas in 2004.
Marcelo KOVACIC: AN ANNOTATED CHECKLIST OF THE FAMILY GOBIIDAE IN THE ADRIATIC SEA, 21-44
Tab. 1: List of gobiid species recorded in the Adriatic Sea up to the present date. Tab. 1: Seznam glavačev, do danes ugotovljenih v Jadranskem morju.
No.	Species
1	Aphia minuta mediterránea De Buen, 1931
2	Buenia affinis Iljin, 1930
3	Chromogobius quadrivittatus (Steindachner, 1863)
4	Chromogobius zebratus zebratus (Kolombatovic,1891)
5	Corcyrogobius liechtensteini (Kolombatovic, 1891)
6	Crystallogobius linearis (Von Düben, 1845)
7	Deltentosteus colonianus (Risso, 1826)
8	Deltentosteus quadrimaculatus (Valenciennes, 1837)
9	Didogobius schlieweni Miller, 1992
10	Didogobius spletchnai Ahnelt & Patzner, 1995
11	Gammogobius steinitzi Bath, 1971
12	Gobius ater Bellotti, 1888
13	Gobius auratus Risso, 1810
14	Gobius bucchichi Steindachner, 1870
15	Gobius cobitis Pallas, 1811
16	Gobius couchi Miller & El-Tawil, 1974
17	Gobius cruentatus Gmelin, 1789
18	Gobius fallax Sarato, 1889
19	Gobius geniporus Valenciennes, 1837
20	Gobius kolombatovici Kovacic & Miller, 2000
21	Gobius niger Linnaeus, 1758
22	Gobius paganellus Linnaeus, 1758
23	Gobius roulei De Buen, 1928
24	Gobius vittatus Vincijuerra, 1883
25	Knipowitschia caucasica (Kawrajsky, 1916)
26	Knipowitschia panizzae (Verja, 1841)
27	Lebetus guilleti (Le Danois, 1913)
28	Lesueurigobius friesii (Malm, 1874)
29	Lesueurigobius suerii (Risso, 1810)
30	Millerigobius macrocephalus (Kolombatovic, 1891)
31	Odondebuenia balearica (Pellejrin & Faje, 1907)
32	Pomatoschistus bathi Miller, 1982
33	Pomatoschistus canestrinii (Ninni, 1883)
34	Pomatoschistus knerii (Steindachner, 1861)
35	Pomatoschistus marmoratus (Risso, 1810)
36	Pomatoschistus minutus (Pallas, 1770)
37	Pomatoschistus norvegicus (Collett, 1903)
38	Pomatoschistus pictus adriaticus Miller, 1972
39	Pomatoschistus quagga (Heckel, 1840)
40	Pseudaphya ferreri (De Buen & Faje, 1908)
41	Speleogobius trigloides (Zander & Jelinek, 1976)
42	Thorogobius ephippiatus (Lowe, 1839)
43	Thorogobius macrolepis (Kolombatovic, 1891)
44	Vanneaugobius dollfusi (Brownell, 1978)
45	Zebrus zebrus (Risso, 1826)
46	Zosterisessor ophiocephalus (Pallas, 1811)
Marcelo KOVACIC: AN ANNOTATED CHECKLIST OF THE FAMILY GOBIIDAE IN THE ADRIATIC SEA, 21-44
SEZNAM VRST IZ DRUŽINE GOBIIDAE V JADRANSKEM MORJU
Marcelo KOVAČIČ Prirodoslovni muzej Rijeka, HR-51000 Rijeka, Lorenzov prolaz 1, Hrvaška E-mail: Marcelo.Kovacic@public.srce.hr
POVZETEK
Avtor predstavlja popoln seznam 46 vrst jadranskih glavačev. Zbrani in na novo pregledani so bili vsi obstoječi podatki o pojavljanju teh vrst v Jadranskem morju. Seznam je bil napravljen na osnovi znanstvene literature in neobjavljenih podatkov iz ihtiološke zbirke, ki jo hranijo v Prirodoslovnem muzeju na Reki. Zaradi dokazov, navedenih in pojasnjenih v tem pregledu, je bilo s prejšnjih seznamov jadranskih glavačev, na katerih je bilo odkritih tudi več drugih napak, črtanih šest vrst. Sicer pa velja, da v Jadranskem morju še vedno ni bilo odkritih trinajst vrst, doslej zabeleženih v Sredozemskem morju. Na osnovi primerkov, ujetih v Jadranskem morju, so doslej opisali tri veljavne rodove, dvanajst veljavnih vrst in dve veljavni podvrsti glavačev. Jadranska gobiologija je bila najuspešnejša v obdobju med letoma 1860 in 1900 z osmimi prvič objavljenimi vrstami in desetimi prvič najdenimi vrstami v Jadranskem morju in med letoma 1968 in 2002 s tremi prvič objavljenimi in petnajstimi prvič najdenimi vrstami jadranskih glavačev.
Ključne besede: seznam, Gobiidae, Jadransko morje
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