FOLIA BIOLOGICA ET GEOLOGICA 59/1, 5–28, LJUBLJANA 2018 A NEW ENDEMIC PLANT COMMUNITY WITH SCHOENUS NIGRICANS IN THE SOUTHEASTERN ALPS AND NORTHERN DINARIC ALPS NOVA ENDEMIČNA ZDRUŽBA Z VRSTO SCHOENUS NIGRICANS V JUGOVZHODNIH ALPAH IN SEVERNEM DELU DINARSKEGA GORSTVA Igor DAKSKOBLER1 & Andrej MARTINČIČ2 http://dx.doi.org/10.3986/fbg0037 ABSTRACT A new endemic plant community with Schoenus nigricans in the Southeastern Alps and northern Dinaric Alps We studied the phytosociology of plant communities with Schoenus nigricans, a character species of the alliance Caricion davallianae, in western Slovenia. Most of our rele- vés are classified into the new association Astrantio carnioli- cae-Schoenetum nigricantis, whose stands grow on moist rocks and in the headwater section of springs and small brooks in dolomite wooded areas, especially in the Cerkno region, in the basin of the Idrijca river. Its character species and (geographical) differential species are Schoenus nigri- cans, Molinia arundinacea, Astrantia carniolica, Palustriella commutata, Carex flacca, Valeriana saxatilis, Salix glabra, Erica carnea and Primula carniolica. In addition to the typi- cal form (subassociation -typicum) we described also the subassociation with predominating Cladium mariscus (-cla- dietosum marisci). The studied dolomite headwaters are a Natura 2000 habitat type and the sites of several vulnerable and protected species. Their role as a biotope is therefore significant and requires adequate protection. Key words: phytosociology, synsystematics, dolomite springs, Schoenus nigricans, Cladium mariscus, Primula car- niolica, Caricion davallianae, Natura 2000, Slovenia IZVLEČEK Nova endemična združba z vrsto Schoenus nigricans v Ju- govzhodnih Alpah in severnem delu Dinarskega gorstva Fitocenološko smo preučili rastlinske združbe zahodne Slovenije, v katerih uspeva vrsta Schoenus nigricans, značilnica zveze Caricion davallianae. Večino naših popisov uvrščamo v novo asociacijo Astrantio carniolicae-Schoene- tum nigricantis, katere sestoji poraščajo vlažno skalovje in povirja studencev in majhnih potokov v dolomitnih gozd- natih območjih predvsem na Cerkljanskem, v povodju Idri- jce. Njene značilnice in (geografske) razlikovalnice so vrste Schoenus nigricans, Molinia arundinacea, Astrantia carnioli- ca, Palustriella commutata, Carex flacca, Valeriana saxatilis, Salix glabra, Erica carnea in Primula carniolica. Poleg tipične oblike (subasociacija -typicum) smo opisali tudi subasociaci- jo s prevladujočo vrsto Cladium mariscus (-cladietosum marisci). Preučena dolomitna povirja sodijo med varstveno pomembne habitatne tipe v evropskem merilu (Natura 2000). So rastišča z več ranljivimi in zavarovanimi vrstami, zato imajo velik biotopski pomen in potrebujejo ustrezno varovanje. Ključne besede: fitocenologija, sinsistematika, dolomit- no povirje, Schoenus nigricans, Cladium mariscus, Primula carniolica, Caricion davallianae, Natura 2000, Slovenija 1 Institute of Biology, Scientific Research Centre of the Slovenian Academy of Sciences and Arts, Regional Unit Tolmin, Brunov drevored 13, SI-5220 Tolmin, igor.dakskobler@zrc-sazu.si 2 Zaloška 78 a, SI-1000 Ljubljana, andrej.martincic@siol.net IGOR DAKSKOBLER & ANDREJ MARTINČIČ: A NEW ENDEMIC PLANT COMMUNITY WITH SCHOENUS NIGRICANS 6 FOLIA BIOLOGICA ET GEOLOGICA 59/1 – 2018 Schoenus nigricans is a subcosmopolitan species, a character species of small-sedge rich-fen vegetation of calcareous oligotrophic flushes, soligenous mires and dune slacks in lower altitudes from the alliance Cari- cion davallianae (Aeschimann et al. 2004: 778, Mar- tinčič 2007: 803, Šilc & Čarni 2012: 128). Its distribu- tion and sites in Slovenia have been described in detail in several articles (Martinčič 1991, 1995, Kocjan et al. 2013). While mapping the flora of the Soča Valley and studying the sites and communities of the endem- ic Primula carniolica we came across, mainly in wood- ed areas, fascinating, small-area dolomite headwaters that were usually dominated by Schoenus nigricans and Molinia arundinacea, very rarely also by Cladium mariscus. The stands with the latter species were al- ready described in more detail in previous article (Dakskobler 2011: 5–6, 19–21) and we also published a phytosociological table with four relevés of the provi- sional syntaxon Astrantio carniolicae-Schoenetum nig- ricantis cladietosum marisci prov. (see also Šilc & Čarni 2012: 128). As we have collected more new rel- evés since then, mainly in the gorges and ravines of the Cerkno region, we made a phytosociological table that allows us to compare the community with Schoenus nigricans from this area with previously described communities elsewhere in Slovenia (Martinčič, ibid., Accetto 2001) and provide a more thorough synsys- tematic classification for this community. 1 INTRODUCTION 2 METHODS Communities with Schoenus nigricans were studied applying the Braun-Blanquet method (Braun- -Blanquet 1964). A total of 75 relevés (of which five had already been published: one in Dakskobler 2005, relevé 8 in Table 1, and four in Dakskobler 2011: 19– 21) were entered into the FloVegSi database (Fauna, Flora, Vegetation and Paleovegetation of Slovenia) of the Jovan Hadži Institute of Biology at ZRC SAZU (T. Seliškar, Vreš et A. Seliškar 2003). They were ar- ranged into a working table based on hierarchical clas- sification. We transformed the combined cover-abun- dance values with numerical values (1–9) according to van der Maarel (1979). Numerical comparisons were performed with the SYN-TAX 2000 program package (Podani 2001). The relevés were compared by means of “(unweighted) average linkage method” – UPGMA, using Wishart’s similarity ratio. The nomenclature source for the names of vascu- lar plants is the Mala flora Slovenije (Martinčič et al. 2007), except for the names Molinia arundinacea Schrank and Sesleria caerulea (L.) Ard. (instead of S. caerulea subsp. calcaria). Ros et al. (2007) is the no- menclature source for the names of liverworts (Mar- canthiophyta) and Ros et al. (2013) for the names of mosses. Šilc & Čarni (2012) are the nomenclature source for the names of the syntaxa, except for the name of the class Querco-Fagetea Braun-Blanquet et Vlieger in Vlieger 1937. Most of the relevés discussed in this article were made in the Cerkno and Idrija Hills at the contact of the pre-Alpine and Dinaric phytogeographical regions of Slovenia (Figure 3). The geological bedrock in the study area is mainly dolomite (Buser 2009). The stud- ied communities occur in the submontane belt (from 200 to 570 m a. s. l.) on initial soils (lithosols, rendzina or very shallow molic gleysols) – Vidic et al. (2015). The regional climate is moderately warm and humid, with mean annual precipitation of 2,000 to 2,500 mm (Zu- pančič 1998) and mean annual air temperature of 6 ºC to 8 ºC (Cegnar 1998), but the local climate is colder. 3. RESULTS AND DISCUSSION 3.1 Review of the studied syntaxa, with types of newly described communities Scheuchzerio palustris-Caricetea nigrae Tx. 1937 nom. mut. Caricetalia davalliane Br.-Bl. 1949 Caricion davallianae Klika 1934 Caricetum davallianae Dutoit 1924 Schoenetum nigricantis W. Koch 1926 Astrantio carniolicae-Schoenetum nigricantis ass. nov. hoc. loco, the nomenclature type, holotypus, is relevé 23 in Table 1. -typicum, subass. nov., the nomenclature type is the same as the nomenclature type of the association IGOR DAKSKOBLER & ANDREJ MARTINČIČ: A NEW ENDEMIC PLANT COMMUNITY WITH SCHOENUS NIGRICANS 7FOLIA BIOLOGICA ET GEOLOGICA 59/1 – 2018 -cladietosum marisci subass. nov., the nomen- clature type, holotypus, is relevé 57 in Table 1. Asplenietea trichomanis (Br.-Bl. in Meier et Br.-Bl. 1934) Oberd. 1977 Potentilletalia caulescentis Br.-Bl. in Br.-Bl. et Jenny 1926 Physoplexido comosae-Saxifragion petraeae Mu- cina et Theurillat 2015 (sin. Phyteumato-Saxifra- gion petraeae Mucina in Šilc et Čarni 2012) Primuletum carniolicae Accetto 2008 Pinguiculetum alpinae nom. prov. Elyno-Seslerietea Br.-Bl. 1948 Seslerietalia coeruleae Br.-Bl. in Br.-Bl. et Jenny 1926 Caricion austroalpinae Sutter 1962 Primulo carniolicae-Seslerietum calcariae Daks- kobler 2006 3.2. Description of the new plant community with Schoenus nigricans in western Slovenia In the first step we exported all our relevés with Schoe- nus nigricans from the FloVegSi database. We obtained a total of 78 relevés, of which three were made in forest communities, two in the stands of the association Gen- isto januensis-Pinetum sylvestris Tomažič 1940 and one in the stand of the association Rhododendro hirsuti- Ostryetum Franz ex Dakskobler 2015. These three stands, where the studied species occurred only with the rating +, were excluded from further study on ac- count of their distinctly different character. The re- maining 75 relevés were compared through hierarchi- cal classification. The result was the dendrogram in Figure 1. Figure 1: Dendrogram of relevés with Schoenus nigricans in western and northwestern Slovenia, UPGMA, 1-similarity ratio: AS – Astrantio carniolicae-Schoenetum nigricantis typicum, ASc – Astrantio carniolicae-Schoenetum nigricantis cladietosum marisci, ASs – Astrantio carniolicae-Schoenetum nigricantis var. Sesleria caerulea, PS – Primulo carniolicae-Seslerietum calcariae, PC– Primuletum carniolicae, SN – Schoenetum nigricantis, CD – Caricetum davallianae Slika 1: Dendrogram popisov sestojev z vrsto Schoenus nigricans v zahodni in sevorozahodni Sloveniji, UPGMA, 1-similarity ratio: AS – Astrantio carniolicae-Schoenetum nigricantis typicum, ASc – Astrantio carniolicae-Schoenetum nigricantis cladi- etosum marisci, ASs – Astrantio carniolicae-Schoenetum nigricantis var. Sesleria caerulea, PS – Primulo carniolicae-Seslerie- tum calcariae, PC– Primuletum carniolicae, SN – Schoenetum nigricantis, CD – Caricetum davallianae IGOR DAKSKOBLER & ANDREJ MARTINČIČ: A NEW ENDEMIC PLANT COMMUNITY WITH SCHOENUS NIGRICANS 8 FOLIA BIOLOGICA ET GEOLOGICA 59/1 – 2018 In terms of species composition the relevés made in the headwaters on glacial material, deluvium or on alluvium in Bohinj, the Završnica valley, the Vipava Valley, near Godovič and in the Bača Valley are clear- ly different and are provisionally classified into the association Schoenetum nigricantis. Also very distinct was one relevé of the dolomite headwaters along Poličnica (at the village of Police) which is temporar- ily classified into the association Caricetum davalli- anae (published already in Dakskobler 2005). These six relevés were not classified into the analytical table (Table 1) which comprises all other 69 relevés. The most different in terms of f loristic composition is the group of relevés on the left side of the dendrogram or in the right part of Table 1. Five of these relevés were made on moist rocks and are temporarily classified into the association Primuletum carniolicae s. lat., and two relevés were made on a stony grassland and are classified into the association Primulo carnioli- cae-Seslerietum calcariae. Despite a certain f loristic similarity with previously mentioned communities we nevertheless discuss two relevés from this sepa- rate group, based on their dominant species, together with the majority of relevés of the new association Astrantio carniolicae-Schoenetum nigricantis (as a provisional variant with Sesleria caerulea). In terms of species composition and sites its stands are clearly different from the stands of the association Schoene- tum nigricantis as it was documented in Slovenia (Martinčič 1991). In addition to Schoenus nigricans both communities share only the following more fre- quent species: Pinguicula alpina, Tofieldia calyculata, Parnassia palustris and Potentilla erecta, occasionally also Campylium stellatum, Cladium mariscus, Palus- triella commutata and Bryum pseudotriquetrum. However, there are a number of species that do not occur in both communities and subsequently Sørensen’s similarity coefficient (1948) is only a bit more than 20%, if we take into account also the less frequent (accidental) species. The studied stands usu- ally occur on small plots ranging between 5 m2 to not more than 30 m2 in size, on gentle to very steep, ero- sion-exposed slopes, along small springs or in moist rocks. They are in direct contact with chasmophytic communities that are frequently dominated by Pri- mula carniolica, as well as with natural grasslands with dominant Molinia arundinacea and (or) Sesleria caerulea. In part, these erosion slopes and dolomite headwaters are being overgrown, above all with wil- lows (Salix glabra, S. appendiculata, S. eleagnos) and Figure 2: Distribution of Astrantia carniolica in Slovenia Slika 2: Razširjenost vrste Astrantia carniolica v Sloveniji IGOR DAKSKOBLER & ANDREJ MARTINČIČ: A NEW ENDEMIC PLANT COMMUNITY WITH SCHOENUS NIGRICANS 9FOLIA BIOLOGICA ET GEOLOGICA 59/1 – 2018 with individual specimens of Picea abies, Frangula alnus and Ostrya carpinifolia. The headwaters occur within wooded areas dominated by communities of Scots pine (Pinus sylvestris), hop hornbeam (Ostrya carpinifolia) and beech (Fagus sylvatica), stands of the associations Genisto januensis-Pinetum sylvestris, Rhododendro hirsuti-Ostryetum, Rhododendro hirsu- ti-Fagetum and Ostryo-Fagetum. The diagnostic spe- cies that very clearly differentiate the new association from all of the previously described, similar fen and headwater communities are: Schoenus nigricans and Molinia arundinacea (dominant species), Astrantia carniolica and Palustriella commutata (with high constancy; they are characteristic for rocky headwa- ter sites), Carex flacca, Erica carnea (frequent in sur- rounding forests and grasslands), Salix glabra (indi- cates both moist sites as well as cold gorges between the Julian Alps and the Trnovski Gozd Plateau) as well as Valeriana saxatilis and Primula carniolica (character species of chasmophytic communities on moist rocks). The latter is an endemic and as such also a phytogeographical differential species that jointly with Astrantia carniolica points to an endemic headwater community of the Southeastern Alps and northern part of the Dinaric Alps. The new association was named after Astrantia carniolica, a southeastern-Alpine Illyrian species that usually occurs on more or less moist and stony lime- stone and dolomite sites with shallow soils (lithosol, rendzina) in numerous forest and shrub communi- ties, primarily of the montane, altimontane and sub- alpine belts (Anemono trifoliae-Fagetum, Homogyno sylvestris-Fagetum, Polysticho lonchitis-Fagetum, Sax- ifrago cuneifolii-Fagetum, Rhododendro hirsuti-Fage- tum, Arunco-Fagetum, Adenostylo glabrae-Piceetum, Petasiti paradoxi-Piceetum, Rhodothamno-Laricetum, Rhododendro hirsuti-Ostryetum, Hemerocallido-Os- tryetum, Lamio orvalae-Alnetum incanae, Alno inca- nae-Pinetum sylvestris, Lamio orvalae-Salicetum elea- gni, Rhodothamno-Pinetum mugo, Rhododendro hir- suti-Alnetum viridis, Rhododendro hirsuti-Salicetum glabrae), in stony grasslands (Primulo carniolicae- Seslerietum calcariae, Ranunculo hybridi-Caricetum sempervirentis, Caricetum ferrugineae), in scree com- munities (Astrantio carniolicae-Adenostyletum glabrae), and most abundantly in headwater basins and in chasmophytic communities of moist rocks with Primula carniolica, Paederota lutea and (or) Pin- guicula alpina. Its distribution in Slovenia (according to the FloVegSi database) is shown in Figure 2. Its site Figure 3: Aproximate localities of stands of the association Astrantio carniolicae-Schonetum nigricantis on the map of Slovenia Slika 3: Približna nahajališča sestojev asociacije Astrantio carniolicae-Schonetum nigricantis na zemljevidu Slovenije IGOR DAKSKOBLER & ANDREJ MARTINČIČ: A NEW ENDEMIC PLANT COMMUNITY WITH SCHOENUS NIGRICANS 10 FOLIA BIOLOGICA ET GEOLOGICA 59/1 – 2018 requirements indicate that it is a very good character species of the newly described association, both in terms of ecology and chorology. The existing locali- ties of the stands of the new association are mainly in the Cerkno and Idrija regions: gorge Kazarska (Buko- vska) Grapa with Beli Potok creek, Orehovska Grapa gorge, Poličnica, Dabrška Grapa gorge, the Idrijca valley at Stopnik – all of them still in the pre-Alpine phytogeographical region, the Sjavnica (Sevnica) val- ley with its tributary the Kopačnica, the area of Do- lenja Trebuša with Hotenja, the area of Gorenja Trebuša with the Pršjak, the Kanomlja valley with Govška Grapa gorge – already in the Dinaric phyto- geographical region. We came across slightly less typical stands of this association in the Breginjski Kot and in the Bača Valley (above the ravine of the Korit- nica) – both in the Alpine phytogeographical region, and in the river basin of the Poljanska Sora (above the Kopačnica valley before Hotavlje) – also in the pre- Alpine phytogeographical region. The existing distri- bution of the described community on the map of Slovenia is presented in Figure 3. Most relevés in Table 1 are classified into the typi- cal subassociation (Astrantio carniolicae-Schoenetum nigricantis typicum); in addition to the already men- tioned two relevés (var. Sesleria caserulea prov.) there are five more relevés that stand out with the dominant Cladium mariscus. These relevés were made in the gorges of Pršjak (Gorenja Trebuša) and Vidršek (below Ravne at Cerkno). We discussed these stands in the past already (Dakskobler 2011) and also justified why they could not be classified neither into the association Mariscetum serrati (Cladietum marisci) nor into the as- sociation Euphorbio villosae-Cladietum marisci (Accetto 2001), the alliance Magnocaricion elatae and class Phragmito-Magnocaricetea. In this article we typ- ify these stands, which differ from others mainly with considerable coverage of Cladium mariscus, as a new subassociation Astrantio carniolicae-Schoenetum nigri- cantis cladietosum marisci. Its only differential species is Cladium mariscus, which is dominant in the highest stand layer. According to this criterion these stands could also be classified at the rank of the new associa- tion Schoeno nigricantis-Cladietum marisci. 4 CONCLUSIONS Dolomite headwaters are recognised as vulnerable and threatened habitat types (calcareous fens) also at the Eu- ropean level (Natura 2000) – Jogan et al. (2004). In the study area they were located in rather remote areas where no major human interventions have been made recently and where their development is mainly subject to the natural dynamics, i.e. succession processes and increasing forest cover. These are also the sites and lo- calities of several red listed vulnerable species of Slove- nia (Anon. 2002): Schoenus nigricans, Cladium mari- scus, Carex hostiana, C. davalliana, Epipactis palustris, Eriophorum latifolium, protected species Pinguicula al- pina, Dactylorhiza fuchsii, Gymnadenia conopsea, Lis- tera ovata and Platanthera bifolia (Anon. 2004) as well as of the endemic and Natura 2000 species Primula car- niolica. Foresters should pay special attention to envis- aged interventions (especially when planning chutes or heavy logging) in these wooded areas. As the studied headwaters are very small they may easily be over- looked. As such, they required accurate mapping and should be indicated in large-scale maps. We advise na- ture conservation services to encourage research into similar dolomite headwaters with Schoenus nigricans also in other parts of Slovenia, with the financial sup- port of the European Union. They have been reported also in the Notranjska region (Polhov Gradec Hills) and in the Dolenjska region (the headwaters of the Besnica and the vicinity of Lipoglav) – Kocjan et al. (2013: 154). 5 POVZETEK Po srednjeevropski metodi (Braun-Blanquet 1964) smo fitocenološko preučili malopovršinska dolomitna povirja, v katerih v zahodni Sloveniji raste črnikasti si- tovec (Schoenus nigricans). Našli smo jih predvsem v Cerkljanskem in Idrijskem hribovju: Kazarska (Buko- vska) grapa z Belim potokom, Orehovska grapa, Polič- nica, Dabrška grapa, dolina Idrijce pri Stopniku, v do- lini Sjavnice (Sevnice) s pritokom Kopačnico, v Dolenji Trebuši s Hotenjo, v Gorenji Trebuši s Pršjakom, v do- lini Kanomlje z Govško grapo, nekaj sestojev tudi v Breginjskem kotu, v Baški dolini (nad grapo Koritnice) in v povodju Poljanske Sore (nad dolino Kopačnice pred Hotavljami); torej v alpskem, predalpskem in di- narskem fitogeografskem območju. Od skupno 78 po- IGOR DAKSKOBLER & ANDREJ MARTINČIČ: A NEW ENDEMIC PLANT COMMUNITY WITH SCHOENUS NIGRICANS 11FOLIA BIOLOGICA ET GEOLOGICA 59/1 – 2018 pisov smo izločili tri, ki smo jih naredili v gozdnih se- stojih (dva v rdečeborovju in enega v črnogabrovju z dlakavim slečem). Na podlagi primerjave 75 popisov (slika 1) smo ugotovili, da so se ločeno združevali popi- si naslednjih asociacij: Schoenetum nigricantis, Carice- tum davallianae, Primuletum carniolicae s. lat., Primulo carniolicae-Seslerietum calcariae. Večina popisov pa je bila od njih različnih. Njihova vrstna sestava je v pri- merjavi s sestoji asociacije Schoenetum nigricantis (Martinčič 1991) precej drugačna, podobno velja za rastiščne razmere, zato jih uvrščamo v novo asociacijo Astrantio carniolicae-Schoenetum nigricantis. Obema združbama so poleg vrste Schoenus nigricans skupne le naslednje pogostejše vrste Pinguicula alpina, Tofieldia calyculata, Parnassia palustris in Potentilla erecta, delo- ma tudi Campylium stellatum, Cladium mariscus, Pa- lustriella commutata in Bryum pseudotriquetrum. Pre- cej vrst pa obema združbama ni skupnih in posledično je floristična podobnost po Sørensen-u (1948), če upo- števamo tudi manj pogoste (slučajne) vrste, le malo več kot 20 %. Preučeni sestoji uspevajo navadno na majh- nih površinah, 5 m2 do največ 30 m2, na položnih do zelo strmih erozijskih pobočjih, ob majhnih studencih ali v vlažnem skalovju. So v neposrednem stiku z združbami skalnih razpok, v katerih pogosto prevla- duje vrsta Primula carniolica, pa tudi z naravnimi tra- višči z dominantima vrstama Molinia arundinacea in (ali) Sesleria caerulea. Deloma se ta erozijska pobočja in dolomitna povirja zaraščajo, predvsem z vrbami (Salix glabra, S. appendiculata, S. eleagnos), posamično tudi s smreko, navadno krhliko in črnim gabrom. Po- virja so v gozdnem prostoru, kjer prevladujejo združbe rdečega bora, črnega gabra in bukve (sestoji asociacij Genisto januensis-Pinetum sylvestris, Rhododendro hir- suti-Ostryetum, Rhododendro hirsuti-Fagetum in Ost- ryo-Fagetum). Diagnostične vrste, ki novo asociacijo zelo jasno razlikujejo od vseh dosedaj opisanih podob- nih združb nizkih barij in povirij so: Schoenus nigri- cans in Molinia arundinacea (dominantni vrsti), Astrantia carniolica in Palustriella commutata (z veliko stalnostjo, zelo značilni za povirna skalnata rastišča), Carex flacca, Erica carnea (pogosti vrsti okoliških goz- dov in travišč), Salix glabra (označuje ne samo vlažna rastišča, temveč tudi hladne grape med Julijskimi Al- pami in Trnovskim gozdom) ter Valeriana saxatilis in Primula carniolica (značilnici združb vlažnih skalnih razpok). Druga je kot endemit tudi fitogeografska raz- likovalnica in kaže skupaj s kranjskim zalim kobulčk- om (Astrantia carniolica) na endemično povirno združbo Jugovzhodnih Alp in severnega dela Dinar- skega gorstva. Večino popisov v preglednici 1 uvršča- mo v tipično subasociacijo Astrantio carniolicae-Schoe- netum nigricantis typicum. Izstopata dva popisa, ki kažeta na podobnost s sestoji asociacije Primulo carni- olicae-Seslerietum calcariae (začasno ju vrednotimo kot var. Sesleria caerulea prov.) in pet popisov, v katerih je dominantna navadna rezika (Cladium mariscus). O njih smo že pisali (Dakskobler 2011) in jih zdaj tipizi- ramo kot novo subasociacijo Astrantio carniolicae- -Schoenetum nigricantis cladietosum marisci. Njena edina razlikovalnica je vrsta Cladium mariscus. Po me- rilih dominantne vrste v najvišji sestojni plasti bi bila mogoča tudi uvrstitev teh sestojev v samostojno asoci- acijo Schoeno nigricantis-Cladietum marisci. Dolomitna povirja sodijo med ranljive in ogrožene habitatne tipe (karbonatna nizka barja) tudi v evrop- skem okviru (Natura 2000) – Jogan et al. (2004). V preučenem območju smo jih večinoma našli v precej odmaknjenih območjih, kjer v zadnjem času ni večjih človekovih posegov in na njihov razvoj najbolj vpliva naravna dinamika, torej sukcesijski procesi in večanje gozdnatosti. V sestojih opisane združbe uspeva tudi več v Sloveniji ranljivih vrst iz rdečega seznama (Anon. 2002): Schoenus nigricans, Cladium mariscus, Carex hostiana, C. davalliana, Epipactis palustris, Eriopho- rum latifolium, nekaj zavarovanih vrst: Pinguicula al- pina, Dactylorhiza fuchsii, Gymnadenia conopsea, Li- stera ovata in Platanthera bifolia (Anon. 2004) in ende- mit in Natura 2000 vrsta kranjski jeglič (Primula car- niolica). Gozdarji morajo biti pozorni pri morebitnih načrtovanih posegih (predvsem gradnji vlak in moč- nejših sečnjah) v ta gozdnata območja. Ker se preučena povirja pojavljajo na zelo majhnih površinah, jih nepo- učeni prav lahka spregledajo, zato bi bilo potrebno nji- hovo natančno kartiranje in vnos v zemljevide po- drobnega merila. Službam za varstvo narave priporo- čamo, da z denarno podporo Evropske skupnosti to spodbudijo, prav tako raziskave podobnih dolomitnih povirij s črnikastim sitovcem v drugih delih Slovenije. Poznamo jih tudi na Notranjskem (Polhograjsko hri- bovje) in na Dolenjskem (v povirju Besnice in v okolici Lipoglava) – Kocjan et al. (2013: 154). IGOR DAKSKOBLER & ANDREJ MARTINČIČ: A NEW ENDEMIC PLANT COMMUNITY WITH SCHOENUS NIGRICANS 12 FOLIA BIOLOGICA ET GEOLOGICA 59/1 – 2018 ACKNOWLEDGEMENTS and corrections. We also acknowledge the financial support from the Slovenian Research Agency (research core funding No. P1-0236). English translation by An- dreja Šalamon Verbič. The authors owe special thanks to Dr. Branko Vreš, Mag. Andrej Seliškar and Brane Anderle, co-authors of Figure 2. Academician Dr. Mitja Zupančič and Prof. Dr. Franc Batič helped us with valuable improvements REFERENCES – LITERATURA Accetto, M., 2001: Nova spoznanja o rastlinstvu Kočevske in Bele krajine. Gozdarski vestnik (Ljubljana) 59 (5–6): 248–259. Aeschimann, D., K. Lauber, D. M. Moser & J.-P.Theurillat, 2004: Flora alpina. Bd. 2: Gentianaceae−Orchi- daceae. Haupt Verlag, Bern, Stuttgart, Wien. 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Dakskobler IGOR DAKSKOBLER & ANDREJ MARTINČIČ: A NEW ENDEMIC PLANT COMMUNITY WITH SCHOENUS NIGRICANS 14 FOLIA BIOLOGICA ET GEOLOGICA 59/1 – 2018 Figure 4: Typical dolomite area with small springs and stands with Schoenus nigricans (Beli potok, Kazarska grapa, Bukovo) Slika 4: Značilno dolomitno območje, kjer so povirja s črnikastim sitovcem (Schoenus nigricans) – Beli potok (Kazarska grapa, Bukovo) Figure 5: Stands of the association Genisto januensis-Pinetum sylvestris are typical for dolomite areas with stands of Schoenus nigricans (Pršjak, Gorenja Trebuša) Slika 5: Sestoji asociacije Genisto januensis-Pinetum sylvestris so značilni za območja z dolomitnimi povirji s črnikastim sitovcem (Schoenus nigricans), Pršjak, Gorenja Trebuša IGOR DAKSKOBLER & ANDREJ MARTINČIČ: A NEW ENDEMIC PLANT COMMUNITY WITH SCHOENUS NIGRICANS 15FOLIA BIOLOGICA ET GEOLOGICA 59/1 – 2018 Figure 7: Also in the stands of the association Primulo carniolicae-Seslerietum calcariae in spots thrive Schoenus nigricans Slika 7: Tudi v sestojih asociacije Primulo carniolicae-Seslerietum calcariae ponekod uspeva črnikasti sitovec (Schoenus nigri- cans) Figure 6: Stands of the association Rhododendro hirsuti-Ostryetum are also characteristic for dolomite areas with stands of Schoenus nigricans, Poličnica (Police) Slika 6: Sestoji asociacije Rhododendro hirsuti-Ostryetum so prav tako značilni za dolomitna območja s sestoji črnikastega sitovca (Schoenus nigricans), Poličnica (Police) IGOR DAKSKOBLER & ANDREJ MARTINČIČ: A NEW ENDEMIC PLANT COMMUNITY WITH SCHOENUS NIGRICANS 16 FOLIA BIOLOGICA ET GEOLOGICA 59/1 – 2018 Figure 8: Stand of the association Primuletum carniolicae s. lat. with Schoenus nigricans, Beli potok (Bukovo) Slika 8: Sestoj asociacije Primuletum carniolicae s. lat. s črnikastim sitovcem (Schoenus nigricans), Beli potok (Bukovo) Figure 9: Stand of the association Astrantio carniol- icae-Schoenetum nigricantis, detail Slika 9: Sestoj asociacije Astrantio carniolicae- Schoenetum nigricantis, detajl IGOR DAKSKOBLER & ANDREJ MARTINČIČ: A NEW ENDEMIC PLANT COMMUNITY WITH SCHOENUS NIGRICANS 17FOLIA BIOLOGICA ET GEOLOGICA 59/1 – 2018 Figure 11: Late spring aspect of the association Astrantio carniolicae-Schoenetum nigricantis, Kopačnica (Šebrelje) Slika 12: Pozno pomladanski videz asociacije Astrantio carniolicae-Schoenetum nigricantis, Kopačnica (Šebrelje) Figure 10: Early spring aspect of the association Astrantio carniolicae-Schoenetum nigricantis, Beli potok (Bukovo) Slika 10: Zgodnje spomladanski videz asociacije Astrantio carniolicae-Schoenetum nigricantis, Beli potok (Bukovo) IGOR DAKSKOBLER & ANDREJ MARTINČIČ: A NEW ENDEMIC PLANT COMMUNITY WITH SCHOENUS NIGRICANS 18 FOLIA BIOLOGICA ET GEOLOGICA 59/1 – 2018 Figure 12: Astrantia carniolica, typical plant of stony and moist sites on dolomite bedrock Slika 12: Kranjski zali kobulček (Astrantia carniolica), značilna vrsta kamnitih in vlažnih rastišč na dolomitu Figure 13: Primula carniolica, Slovenian endemic and one of the diagnostic species of the association Astrantio carniolicae- Schoenetum nigricantis Slika 13: Kranjski jeglič (Primula carniolica), slovenski endemit in ena izmed diagnostičnih vrst asociacije Astrantio carnioli- cae-Schoenetum nigricantis IGOR DAKSKOBLER & ANDREJ MARTINČIČ: A NEW ENDEMIC PLANT COMMUNITY WITH SCHOENUS NIGRICANS 19FOLIA BIOLOGICA ET GEOLOGICA 59/1 – 2018 Figure 14: Stand of the subassociation Astrantio carniolicae-Schoenetum nigricantis cladietosum marisci, Pršjak (Gorenja Trebuša) Slika 14: Sestoj subasociacije Astrantio carniolicae-Schoenetum nigricantis cladietosum marisci, Pršjak (Gorenja Trebuša) IGOR DAKSKOBLER & ANDREJ MARTINČIČ: A NEW ENDEMIC PLANT COMMUNITY WITH SCHOENUS NIGRICANS 20 FOLIA BIOLOGICA ET GEOLOGICA 59/1 – 2018 Table 1: Communities with Schoenus nigricans in western Slovenia Preglednica 1: Združbe z vrsto Schoenus nigricans v zahodni Sloveniji Number of relevé (Zaporedna številka popisa) 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 31 32 Database number of relevé (Delovna številka popisa) 21 02 85 24 87 95 24 87 99 24 88 01 21 22 24 26 93 20 24 49 45 26 83 07 26 87 73 23 78 53 26 91 34 26 31 64 26 91 37 26 92 02 26 94 21 26 91 99 26 37 52 26 92 48 26 92 54 26 94 03 26 61 40 26 31 70 26 92 00 26 67 09 26 67 15 26 67 17 24 49 47 26 31 66 26 67 07 26 85 04 23 78 54 25 85 86 Elevation in m (Nadmorska višina v m) 480340 350400360300225240 357 480 350440400 470 400440390 515 560400253 475 475 530 565440202440530 480500440 Aspect (Lega) SW N EE N E N E SE N E S SE SW S E W S E N N E N S E N E S N SW SW SW S S N E S N E SE N E S SW Slope in degrees (Nagib v stopinjah) 10 20 15 45 50 30 45 20 35 15 30 60 40 35 30 35 10 50 45 70 40 50 50 70 70 45 30 30 80 25 10 10 Parent material (Matična podlaga) D D D D DC D D D D D D D D D D D D D D D D D D D D D D D D D D D Soil (Tla) Li Hy Hy Hy Li Hy Hy Hy Hy Hy Hy Hy Hy Hy Hy Hy Li Hy Hy Hy Li Hy Hy Li Li Li Hy Hy Li Re Hy Hy Cover of shrub layer in % (Zastiranje grmovne plasti v %) E2 . . 10 10 10 5 20 . . . . . . . . 5 . . . . . . 5 . . . . . . . . 20 Cover of herb layer in % (Zastiranje zeliščne plasti v %) E1 100 70 80 70 70 80 80 80 95 70 80 80 60 70 80 80 80 70 70 60 90 80 80 80 80 80 80 90 70 80 70 80 Cover of moss layer in % (Zastiranje mahovne plasti v %) E0 20 20 30 20 10 20 10 20 10 20 40 30 30 10 30 10 30 30 20 30 20 20 5 20 10 20 30 30 20 20 20 Number of species (Število vrst) 19 22 24 24 15 16 13 14 15 17 18 16 15 20 12 23 14 18 18 15 13 14 16 14 17 25 17 13 19 22 17 21 Relevé area (Velikost popisne ploskve) m 2 5 10 15 15 10 20 20 20 10 10 10 10 10 10 10 10 10 10 10 20 20 10 10 20 20 20 10 10 10 5 10 10 Date of taking relevé (Datum popisa) 5/ 30 /2 00 5 6/ 23 /2 01 3 6/ 23 /2 01 3 6/ 23 /2 01 3 6/ 5/ 20 06 5/ 18 /2 01 7 5/ 29 /2 01 2 8/ 13 /2 01 7 10 /8 /2 01 7 6/ 8/ 20 10 5/ 30 /2 01 7 9/ 12 /2 01 6 5/ 30 /2 01 7 4/ 7/ 20 17 3/ 31 /2 01 7 4/ 7/ 20 17 6/ 24 /2 01 6 5/ 25 /2 01 7 5/ 25 /2 01 7 5/ 26 /2 10 7 4/ 9/ 20 17 9/ 12 /2 01 6 4/ 7/ 20 17 4/ 13 /2 01 7 4/ 13 /2 01 7 4/ 13 /2 01 7 5/ 29 /2 01 2 9/ 12 /2 01 6 4/ 13 /2 01 7 9/ 22 /2 01 7 6/ 8/ 20 10 9/ 29 /2 01 5 Locality (Nahajališče) Po lic e- M lin i R ek a- Sv . I va n R ek a- Sv . I va n R ek a- Sv .Iv an K or it ni ca St op ni k- D ab rč ek St op ni k- V M lin u G or en ja T re bu ša -B el e vo de H ot en ja R av ne -V id rš ek Sj av ni ca -K op ač ni ca O re ho vs ka g ra pa Sj av ni ca -K op ač ni ca O re ho vs ka g ra pa K az ar sk a gr ap a- Se ls ka v od a O re ho vs ka g ra pa K az ar sk a gr ap a Po lic e- Po lič ni ca Po lic e- Po lič ni ca K az ar sk a gr ap a- Se ls ka v od a G ač ni k O re ho vs ka g ra pa O re ho vs ka g ra pa K az ar sk a gr ap a- Be li po to k K az ar sk a gr ap a- Be li po to k K az ar sk a gr ap a- Be li po to k St op ni k- K re pa ka O re ho vs ka g ra pa K az ar sk a gr ap a- Be li po to k O re ho vs ka g ra pa -O re he k R av ne -V id rš ek K an om lja -G ov šk ar ic a Quadrant (Kvadrant) 98 49 /3 98 49 /4 98 49 /4 98 49 /4 98 49 /1 98 49 /3 99 49 /1 99 49 /1 99 49 /1 98 49 /4 99 49 /2 98 49 /4 99 49 /2 98 49 /4 98 49 /3 98 49 /4 98 49 /3 98 49 /3 98 49 /3 98 49 /3 99 49 /1 98 49 /4 98 49 /4 98 49 /3 98 49 /3 98 49 /3 98 49 /3 98 49 /4 98 49 /3 98 49 /4 98 49 /4 99 49 /4 Coordinate GK Y (D-48) m 41 58 20 41 69 19 41 68 89 41 68 42 41 36 88 41 50 28 41 40 33 40 97 88 41 33 37 41 87 41 41 72 02 41 81 39 41 70 88 41 81 35 41 58 92 41 81 53 41 60 76 41 56 30 41 53 07 41 58 85 41 04 19 41 80 04 41 81 48 41 55 06 41 53 65 41 57 25 41 32 83 41 81 22 41 54 65 41 82 35 41 87 33 42 22 29 Coordinate GK X (D-48) m 51 09 99 1 51 07 72 9 51 07 61 7 51 07 58 4 51 15 97 7 51 08 81 3 51 06 34 4 51 03 40 2 51 03 89 7 51 08 23 2 51 06 36 4 51 10 89 9 51 06 25 3 51 10 91 7 51 11 32 4 51 10 89 7 51 11 04 7 51 10 15 8 51 10 52 8 51 11 33 9 51 04 09 1 51 11 05 2 51 10 96 6 51 10 73 2 51 10 74 4 51 10 63 2 51 06 88 3 51 10 87 8 51 10 71 6 51 11 15 5 51 08 28 4 50 99 66 6 Diagnostic species of the associations (Diagnostične vrste asociacije) CD Schoenus nigricans E1 3 4 3 4 4 2 3 4 3 4 3 4 3 3 3 4 3 4 4 4 4 3 3 4 4 4 4 3 3 3 4 3 EP Molinia arundinacea E1 3 4 4 3 2 4 4 4 3 3 4 3 3 4 4 3 2 3 3 2 4 4 4 2 3 3 3 4 3 3 2 4 TR Astrantia carniolica E1 1 1 2 1 2 3 1 2 2 1 2 1 1 1 + 1 4 1 2 2 1 1 1 1 1 1 1 1 1 1 . . MCPalustriella commutata E0 . 2 2 1 1 2 1 1 + 2 2 1 2 2 4 2 2 2 1 2 1 2 1 + 1 + 1 1 2 3 2 2 EP Erica carnea E1 + + + + . + + + + 1 + + . 1 + 1 + + + + + 1 + + + 2 . + + 1 1 2 QP Carex flacca E1 . + + + + 1 . . . . + . + . + . . . + + . + + 1 2 2 + + . . . 1 AT Valeriana saxatilis E1 1 1 + + . . . . . . . . + + . + + 1 1 1 + + + + . 1 . . + . . . BA Salix glabra E2a + . + + . . . . . . . r . . + + + 1 + + . . + + + + . . + + . 1 AT Primula carniolica E1 + . 1 2 . . . . . . . + . 1 . . . . + . . 1 1 2 + + . + 2 2 . . Diferential species of lower units (Razlikovalna vrsta subasociacije) PM Cladium mariscus E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . IGOR DAKSKOBLER & ANDREJ MARTINČIČ: A NEW ENDEMIC PLANT COMMUNITY WITH SCHOENUS NIGRICANS 21FOLIA BIOLOGICA ET GEOLOGICA 59/1 – 2018 33 34 35 36 37 38 39 40 41 42 43 44 45 46 47 48 49 50 51 52 53 54 55 56 57 58 59 60 61 62 63 64 65 66 67 68 69 26 92 49 24 84 03 26 32 91 26 32 92 26 32 93 25 86 04 26 79 64 26 79 70 26 32 94 21 78 73 26 67 20 25 63 58 26 92 51 26 13 55 26 62 18 22 71 22 24 20 75 23 78 88 25 58 47 26 37 50 26 94 04 26 91 86 26 31 79 21 80 59 21 80 60 23 78 52 23 50 43 23 50 44 21 02 82 21 02 87 21 06 31 21 78 70 26 31 63 26 91 87 26 92 01 26 31 65 26 87 72 525 310 475475480570484498445260440240544206203205390210360390370465490342342450430430490480220270440475470440357 N E SW S S SS W SW S S SS W N E S N W SW N W N SW SE E N E SW S E SE SW W SW SS W S SE SE SE SW W N N N E N SW N EE SE SW 10 20 30 35 35 40 20 50 5 70 90 40 80 70 60 35 45 15 25 10 5 45 25 50 45 25 25 15 35 25 30 15 90 90 80 80 80 D D D D D D D D D D D D D D D D D D D Tu D D Gr D D D D D D D D D D D D Tu D Hy Hy Hy Hy Hy Hy Hy Hy Hy Li Li Hy Hy Li Li Hy Hy Hy Hy Hy Hy Hy Li Li Re Hy Hy Hy Li Li Li Li Li Li Li Li Li . 10 . . . 5 . . . . . . . . . . . 20 10 . . . . 5 5 20 10 10 5 5 . 5 . . . . . 80 70 100100100 85 80 80 90 40 60 75 50 60 60 95 50 80 80 90 90 100 80 90 90 90 80 70 60 80 70 80 60 40 30 50 60 20 15 20 20 20 30 30 20 10 30 20 30 30 20 10 20 10 10 10 10 20 40 40 30 20 5 11 19 16 12 6 18 14 14 10 11 10 10 11 14 12 12 9 35 34 14 15 18 12 11 13 18 26 20 24 25 22 21 15 14 10 19 9 10 30 10 10 10 50 10 10 10 10 20 5 10 10 20 10 10 20 50 10 20 10 10 10 20 10 20 10 10 5 4 10 10 10 5 10 10 5/ 25 /2 01 7 5/ 19 /2 01 3 10 /7 /2 01 6 10 /7 /2 01 6 10 /7 /2 01 6 9/ 29 /2 01 5 7/ 18 /2 01 7 7/ 18 /2 01 7 10 /7 /2 01 6 4/ 19 /2 00 7 4/ 13 /2 01 7 4/ 22 /2 00 2 5/ 25 /2 01 7 5/ 8/ 20 16 4/ 9/ 20 17 9/ 17 /2 00 9 7/ 21 /2 01 1 6/ 3/ 20 10 5/ 25 /2 00 0 6/ 24 /2 01 6 5/ 26 /2 10 7 4/ 7/ 20 17 9/ 12 /2 01 6 6/ 5/ 20 07 6/ 5/ 20 07 10 /8 /2 01 0 4/ 9/ 20 10 4/ 9/ 20 10 5/ 30 /2 00 5 5/ 30 /2 00 5 4/ 22 /2 00 2 4/ 19 /2 00 7 9/ 12 /2 01 6 4/ 7/ 20 17 4/ 7/ 20 17 9/ 12 /2 01 6 10 /8 /2 01 7 Po lic e- Po lič ni ca St op ni k- V M lin u H ot av lje -S re dn je B rd o H ot av lje -S re dn je B rd o H ot av lje -S re dn je B rd o K an om lja -G ov šk ar ic a Br eg in js ki k ot -B or ja na Br eg in js ki k ot - St an ov iš če H ot av lje -S re dn je B rd o G or en ja T re bu ša -P rš ja k K az ar sk a gr ap a- Be li po to k Pr ve jk -T em ni ka r Po lic e- Po lič ni ca D ol en ja T re bu ša D ol en ja T re bu ša St op ni k- K re pa ka K or it ni ca -R ut D ol en ja T re bu ša -P ol og G or en ja T re bu ša -P rš ja k K az ar sk a gr ap a K az ar sk a gr ap a- Se ls ka v od a O re ho vs ka g ra pa O re ho vs ka g ra pa Pr šj ak -S op ot a Pr šj ak -S op ot a R av ne -V id rš ek R av ne -V id rš ek R av ne -V id rš ek Po lic e- M lin i Po lic e. M lin i D ol en ja T re bu ša -P rv ej k G or en ja T re bu ša -P rš ja k O re ho vs ka g ra pa O re ho vs ka g ra pa O re ho vs ka g ra pa O re ho vs ka g ra pa H ot en ja 98 49 /3 99 49 /1 98 50 /4 98 50 /4 98 50 /4 99 49 /4 97 46 /2 97 46 /2 98 50 /4 99 49 /1 98 49 /3 99 48 /2 98 49 /3 99 49 /1 99 49 /1 98 49 /3 98 49 /1 99 48 /2 99 49 /1 98 49 /3 98 49 /3 98 49 /4 98 49 /4 99 49 /1 99 49 /1 98 49 /4 98 49 /4 98 49 /4 98 49 /3 98 49 /3 99 48 /2 99 49 /1 98 49 /4 98 49 /4 98 49 /4 98 49 /4 99 49 /1 41 55 48 41 42 15 43 10 98 43 11 18 43 10 60 42 16 43 38 19 99 38 12 01 43 10 39 40 98 28 41 56 90 40 97 44 41 54 10 41 00 99 41 00 77 41 32 57 41 34 82 40 97 70 41 04 55 41 60 76 41 60 74 41 81 52 41 79 54 41 03 79 41 03 63 41 87 51 41 87 83 41 87 62 41 57 95 41 58 40 40 97 10 40 98 43 41 81 36 41 81 48 41 81 35 41 81 25 41 33 23 51 10 27 3 51 06 29 7 51 09 20 3 51 09 20 9 51 09 27 6 50 99 69 9 51 24 53 8 51 24 68 2 51 09 20 8 51 01 93 6 51 10 65 9 51 06 46 4 51 10 47 0 51 06 10 1 51 06 09 0 51 06 89 2 51 15 68 0 51 05 12 5 51 01 76 2 51 11 06 5 51 11 06 6 51 10 93 8 51 11 17 8 51 01 83 4 51 01 84 7 51 08 19 8 51 08 14 3 51 08 21 3 51 10 04 0 51 09 94 2 51 06 51 1 51 01 89 5 51 10 90 0 51 10 93 4 51 10 90 3 51 10 90 5 51 03 90 1 Pr . 1 -5 5 Fr . 1 -5 5 Pr . 5 6- 60 Fr . 5 6- 60 Pr . 6 1- 69 Fr . 6 1- 69 2 4 4 5 5 3 4 3 4 3 3 4 3 3 4 5 4 3 3 2 2 1 3 2 3 1 3 + 3 1 r r + + + 1 1 55 100 5 100 9 100 4 3 4 4 3 4 3 1 3 + + 1 1 . . 2 1 3 2 4 3 4 1 1 1 + + + + + + + . + . . . 53 96 5 100 5 56 2 1 . . . . . . . 1 r + + 1 + 2 1 1 1 3 3 1 . . 1 . . + 1 + 1 1 1 . 1 2 . 45 82 2 40 7 78 1 1 . 2 . 2 2 2 2 1 . . . 2 1 . . . 2 1 2 . . . 1 . + + 1 . + + 2 2 2 2 . 44 80 3 60 7 78 . . . . . . . + . . . + . . . . . 1 + + + 1 1 . . . . . . . . . . . . . . 37 67 0 0 0 0 1 + . . . + . . . . . 1 . . . . . 1 1 . + 1 . . . . + + + . . + . . . . . 26 47 2 40 2 22 2 . . . . . . . . 1 + + + + r . . . 1 . . + . . . . . . 1 2 1 1 + + . . . 26 47 0 0 6 67 1 . . . . 1 . . . . . . + . . . . . + . . + . . . . . . + + . . . . . . . 22 40 0 0 2 22 . . . . . . . . . 1 1 . . . . . . . . . . 1 . . . . . . 2 3 1 1 2 3 1 + . 17 31 0 0 8 89 . . . . . . . . . . . . . . . . . . . . . . . 4 4 5 4 4 . . . . . . . . . 0 0 5 100 0 0 IGOR DAKSKOBLER & ANDREJ MARTINČIČ: A NEW ENDEMIC PLANT COMMUNITY WITH SCHOENUS NIGRICANS 22 FOLIA BIOLOGICA ET GEOLOGICA 59/1 – 2018 CD Caricetalia davallianae Pinguicula alpina E1 2 1 1 1 1 + . . + 1 1 1 1 2 1 1 1 2 2 2 2 1 1 2 1 1 + . 3 1 2 1 Tofieldia calyculata E1 1 + + + . . . . . + + + + 1 + + + . + 1 . . . + + . 1 1 1 1 1 1 Parnassia palustris E1 . + . . . . . + 1 + + + . . . . + + . . . . . . . . . + + + . . Carex lepidocarpa E1 + + + . + 1 + . . . + . . . . . . . . . . . . . . . + . . . . 1 Campylium stellatum E0 . . + . . . . . . . . . . + . . . 1 . . . . . . . 1 . . . . . . Carex flava s.str. E1 . 1 + . + . . . . . . . + . . . . . . . . . . . . . . . . . . . Eriophorum latifolium E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Carex hostiana E1 + . . . . . . . . . . . . . . . . + . + . . . . . . . . . . . . Carex davalliana E1 + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Juncus articulatus E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + MCMontio-Cardaminetea Hymenostylium recurvirostre E0 . + . . . . . . . . . 1 1 . . 1 . . + . . . . . . . . . 2 . . . Bryum pseudotriquetrum E0 . . . . . . . . . . . . + . . 1 . . . . . . + . . . . . . 1 . . AT Asplenietea trichomanis Phyteuma scheuchzeri subsp. columnae E1 . . . . . . . . . . . . . . . + . . . . . . . . . . . . + . . . Hieracium bifidum E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Paederota lutea E1 . . . . . . . . . . . . . . . . . . . . + . . . . . . . . . . . Potentilla caulescens E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . TR Thlaspietea rotundifolii Petasites paradoxus E1 . . . . + . 1 . . . . . . . . . . . . . . . . . . . 1 . . . . + Campanula cespitosa E1 . . . . . . . . . . . + . . . . . . . . . + . . . . . . . . . . Adenostyles glabra E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 Achnatherum calamagrostis E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Aquilegia einseleana E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Peucedanum verticillare E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Equisetum ramosissimum E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . Equisetum variegatum E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . ES Elyno-Seslerietea Sesleria caerulea E1 + . 1 + . . + . . 1 1 + 1 + + + + + . + + . . . . + + . 1 1 + . Carex mucronata E1 . . . . . . . . . . . . . + . + . + . . + . + + + + . . + + . . Phyteuma orbiculare E1 . + . + . . . . . . . . . . . . . . . . + 1 . + + + . . + . . . Globularia cordifolia E1 . . . . . . . r . . . . . . . . . . + . . . . . + + . + . . . . Aster bellidiastrum E1 1 . . . . . . . . . + . r . . . . . . + . . . . . . . . . + . . Betonica alopecuros E1 . . + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Carex ferruginea E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . CU Calluno-Ulicetea Potentilla erecta E1 . + + + . . + 2 1 + + + + + . + + + 1 + . + . . . . r . . + + 1 FB Festuco-Brometea Gymnadenia conopsea E1 . . . . . . . r r . . . . . . . . . . . . . . . . . . . . . . . Brachypodium rupestre E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Peucedanum oreoselinum E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Teucrium chamaedrys E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + Euphorbia cyparissias E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Euphorbia verrucosa E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Galium verum E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Gentianella ciliata E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Linum catharticum E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Briza media E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Dorycnium herbaceum E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Prunella grandiflora E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Bromopsis erecta E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Carex humilis E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . MAMolinion, Molinio-Arrhenatheretea Mo Cirsium oleraceum E1 . . . . + . . . . . . . . . + . . . . . . . . . . . . . . . . . Mo Epipactis palustris E1 . . . . . . . + . . . . . . . . . . . . . . . . . . . . . . . . MuAVeratrum album E1 . . . . . . . . . . . . . . . . . . . . 1 . . . . . . . . . . . Mo Succisa pratensis E1 . . . . . . . + + . . . . . . . . . . . . . . . . . . . . . . . Mo Crepis paludosa E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Number of relevé (Zaporedna številka popisa) 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 31 32 IGOR DAKSKOBLER & ANDREJ MARTINČIČ: A NEW ENDEMIC PLANT COMMUNITY WITH SCHOENUS NIGRICANS 23FOLIA BIOLOGICA ET GEOLOGICA 59/1 – 2018 . . . . . 1 1 . . 2 3 2 2 2 3 2 2 . 1 . . . . . + . . + 2 1 1 1 1 2 3 2 3 40 73 2 40 9 100 . . . . . + . 1 . . + . 1 + + 2 1 1 1 + . . . + . + . . 1 1 + . + + + + 1 33 60 2 40 8 89 . . + + . 1 1 1 2 . . . + . . + . . . . . . . . . + . . . . . . . . . + + 19 35 1 20 2 22 + + . . . 1 1 + . . . . . . . . . 1 1 . . . . . . . . . + . . . . . . . . 16 29 0 0 1 11 . 1 . + . . . . . . . . . . . . . . 1 . . . . . + . . . . . . . . . . + . 7 13 1 20 1 11 . + . . . . . . . . . . . . . + . . . . . . . . . . . . . . . . . . . . . 6 11 0 0 0 0 . . . . . . . . . . . . . . . . . 1 2 1 3 . . . . . . . . . . . . . . . . 4 7 0 0 0 0 . . . . . . . . . . . . . . . . . + . . . . . . . . . . + 1 . . . . . . . 4 7 0 0 2 22 . . . . . . . . . . . . . . . . . . . 1 3 . . . . . . . . . . . . . . . . 3 5 0 0 0 0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 2 0 0 0 0 . . . . . + . . . . 1 . . . . . . . . . . . . . . . . . . . . . 3 3 2 1 1 8 15 0 0 5 56 . . . + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . 5 9 0 0 1 11 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 2 4 0 0 0 0 . . . . . . . + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 2 0 0 0 0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . . 1 2 0 0 1 11 . . . . . . . . . . r . . . . . . . . . . . . . . . . . . . . . . + . . . 1 2 0 0 1 11 . . . . . + + + . . . . . + . . + . . . . . . . . . . + . . . . . . . . . 9 16 1 20 0 0 . . . . . + . + . . . . . + + . + . . . . . + . . . . . . . . . 1 1 . . + 8 15 0 0 3 33 . . . . . r . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 2 4 0 0 0 0 . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . 1 2 0 0 0 0 . . . . . . + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 2 0 0 0 0 . . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . . . . . . 1 2 0 0 0 0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 2 0 0 0 0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 2 0 0 0 0 . 1 . . . . . . . r . 1 + 1 + . r . . . + 1 1 . . . . + 2 1 + 2 + 1 1 + . 30 55 1 20 8 89 . . . . . . . . . 1 1 . . + . . . . + . . . . . . . . . + 1 + 1 + 1 + . 2 14 25 0 0 8 89 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 . . . . . . 8 15 0 0 1 11 . . . . . . . r . . . . . . . . . . . . . r . . . . . . + + . . . . . . . 7 13 0 0 2 22 . . . . . . . . . . . . . 1 . . . + . . . . . . . . . . + 1 . + . . . . . 7 13 0 0 3 33 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 2 0 0 0 0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 . . . . . . 0 0 0 0 1 11 + 1 1 + + + 1 1 . . . . . . . + . 1 1 . 1 . . + + 1 + + + . . . . . . . . 33 60 5 100 1 11 . . . . . r . . . . . . . . . . . + . . . . . . . . . . . . . . . . . . . 4 7 0 0 0 0 . . . . . . . . . . . . . . . . . . . + + . . . . . . . . . . . . . . . . 2 4 0 0 0 0 + . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . . . . . . 2 4 0 0 0 0 . . + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 2 4 0 0 0 0 . . + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 2 0 0 0 0 . . + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 2 0 0 0 0 . . + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 2 0 0 0 0 . . . . . + . . . . . . . . . . . . . . . . . . . . r . . . . . . . . . . 1 2 1 20 0 0 . . . . . . . . . . . . . . . 1 . . . . . . . . . . . . . . . . . . . . . 1 2 0 0 0 0 . . . . . . . . . . . . . . . . . r . . . . . . . . . . . . . . . . . . . 1 2 0 0 0 0 . . + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 2 0 0 0 0 . . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . . . . . . 1 2 0 0 0 0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 0 0 0 0 0 0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . 0 0 0 0 1 11 . . . . . . 1 . . . . . . . . + . + . . . . . . . . . . . . . . . . . . . 5 9 0 0 0 0 . + . . . . . . 1 . . . . . . . . + . . . . . . . . . . . . . . . . . . . 4 7 0 0 0 0 . . . . . . . . . . . + . . + . . + . . . . . . . . . . . . r . . . . . . 4 7 0 0 1 11 . . . . . . . . . . . . . . . . . 1 . . . . . . . . . . . . . . . . . . . 3 5 0 0 0 0 . . . . . + . . . . . . . . . . . + . . . . . . . . . . . . . . . . . . . 2 4 0 0 0 0 33 34 35 36 37 38 39 40 41 42 43 44 45 46 47 48 49 50 51 52 53 54 55 56 57 58 59 60 61 62 63 64 65 66 67 68 69 Pr . 1 -5 5 Fr . 1 -5 5 Pr . 5 6- 60 Fr . 5 6- 60 Pr . 6 1- 69 Fr . 6 1- 69 IGOR DAKSKOBLER & ANDREJ MARTINČIČ: A NEW ENDEMIC PLANT COMMUNITY WITH SCHOENUS NIGRICANS 24 FOLIA BIOLOGICA ET GEOLOGICA 59/1 – 2018 MALotus corniculatus E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Mo Carex distans E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Mo Angelica sylvestris E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . MARanunculus nemorosus E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . TG Trifolio-Geranietea Geranium sanguineum E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Salvia pratensis subsp. saccardiana E1 . . + . . . . . . . . . . . . . . . . . . . . . . r . . . . . . Vincetoxicum hirundinaria E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Viola hirta E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Laserpitium latifolium E1 . . . . . . + . . . . . . . . . . . . . . . . . . . . . . . . . Anthericum ramosum E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Silene nutans E1 . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . BA Betulo-Alnetea Salix appendiculata E2a . . + + + + . + + + + . . . . + . + . . . . . . . . . . . . . + FP Filpendulo-Petasition Lysimachia vulgaris E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Bidens frondosa E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . PM Phhragmiti-Magnocaricetea Carex paniculata E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Mentha aquatica E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . RP Rhamno-Prunetea Juniperus communis E2a . . . . . . . . . . . . . . . . . . r . . . . . . . . . . + . . Ligustrum vulgare E2a . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Rhamnus catharticus E2a . . + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Berberis vulgaris E2a . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Rubus fruticosus agg. E2a . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Viburnum lantana E2 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . EA Epilobietea angustifolii Eupatorium cannabinum E1 . + + . + + . . . 1 + . + + . . . . . . . . . . . . + + . . 1 . Tussilago farfara E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Crepis sp. E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . EP Erico-Pinetea Buphthalmum salicifolium E1 . . . . . . . . . r . . . . . + . . . . . + 1 . + . + . . + + . Polygala chamaebuxus E1 + . . . . . . . . . . . . . . . . . + . . . . . . . . . + + . . Calamagrostis varia E1 . . . . . . . . + . . . . . . . . . . . . . . . . . . . . . . . Allium ericetorum E1 . . . . . . . + . . . . . . . . . . . . . . . . + + . . . + . . Aquilegia nigricans E1 . + . . . . . . . . . . . + . . . . . . . . . . . . + . . . . . Pinus sylvestris E1 . . . + . . . . . . . . . . . . . . . . . + . . . . . . . . . . Cirsium erisithales E1 . . . . . . . . . . . . . . . + . . . . . . + . . . . . . . . . Pinus sylvestris E2a . . . . . . . . + . . . . . . . . . . . . . . . . . . . . . . r Leontodon incanus E1 . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . . . Amelanchier ovalis E2a . . . r . . . . . . . . . . . . . . . . . . . . + . . . . . . . Chamaecytisus hirsutus E1 . . . . . . . . . r . . . . . . . . . . . . . . . . . . . . . + Rhododendron hirsutum E2a . . . r . . . . . . + . . . . . . . . . . . . . . . . . . . . . Gymnadenia odoratissima E1 . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . . Asperula aristata E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Chamaecytisus purpureus E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . Crepis slovenica E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . VP Vaccinio-Piceetea Picea abies E2a + r . + . . . . + . . . . + . + . . + . . . + . . + . . . . . + Gentiana asclepiadea E1 + . + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . AI Alnion incanae Frangula alnus E2 . . + . . + + . + . . . . . . . . . . . . . . . . . . . . . + . Salix eleagnos E2 . . + + + . 2 + . . . . . . . . . + . . . . . . . . . . . . + . Equisetum arvense E1 . + . . . . . . . . . . . . . . . . . . . . . . . . . . . . + + Listera ovata E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Alnus incana E2a . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Viburnum opulus E2a . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Equisetum hyemale E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Number of relevé (Zaporedna številka popisa) 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 31 32 IGOR DAKSKOBLER & ANDREJ MARTINČIČ: A NEW ENDEMIC PLANT COMMUNITY WITH SCHOENUS NIGRICANS 25FOLIA BIOLOGICA ET GEOLOGICA 59/1 – 2018 . . . . . . . . . . . . . . . . . + + . . . . . . . . . . . . . . . . . . 2 4 0 0 0 0 1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 2 0 0 0 0 . . . . . . . . . . . . . . . . . . . + . . . . . + . . . . . . . . . . . 1 2 1 20 0 0 . . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . . . . . . 1 2 0 0 0 0 . . + + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 2 4 0 0 0 0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 2 4 0 0 0 0 . . + + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 2 4 0 0 0 0 . . . . . . . . . . . . . . . . . . + . . . + . . + + . . . . . . . . . . 2 4 2 40 0 0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 2 0 0 0 0 . . . . . . . . . . . . . . . . . . + . . . . . . . . . . + . . . . . . . 1 2 0 0 1 11 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 2 0 0 0 0 . . . . . . . . . . . . + . + . . . . . . . . . + . + . . . . . . . . . + 13 24 2 40 1 11 . + . . . . . . 1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . 2 4 0 0 0 0 . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . . . . . . . . 1 2 0 0 0 0 . . . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . . . . . 1 2 0 0 0 0 . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . . . 0 0 1 20 0 0 . . . . . . . . . . . . . . . . . + + . . + . . . . . + . + . + . . . . . 5 9 1 20 2 22 . + . . . . . . . . . . . . . . . . . . . . . . . . + + . . . . . . . . . 1 2 2 40 0 0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 2 0 0 0 0 . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . . . . 0 0 1 20 0 0 . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . . . 0 0 1 20 0 0 . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . . 0 0 1 20 0 0 . . . . . . . . + . . . . . . . . . + . . . . . . . 1 1 . . . . . . . r . 13 24 2 40 1 11 . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . . . . . . . . 1 2 0 0 0 0 . . . . + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 2 0 0 0 0 . . . . . . . + + . . . . . . . . . + . . 1 + . . . . . . . . . . . . . . 13 24 0 0 0 0 . . + + . . . . . . . . . . + . . + + . . . . . . . . . . . . . . . . . . 9 16 0 0 0 0 . . . . . + 1 1 + . . . . + . . 1 . . . . . . . . . . . . . . . . . . . . 7 13 0 0 0 0 . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . 5 9 0 0 0 0 . . . + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 4 7 0 0 0 0 . . . . . . . . + . . . . . . . . . . . . . + . . + . . . . . . . . . . . 4 7 1 20 0 0 . . . . . . . . . . . . . . . . . . . . . 1 . . . + . . . . r 2 . . . . . 3 5 1 20 2 22 . . . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . . . . . 3 5 0 0 0 0 . . . . . . . . . . . . . . . . . . . . . . 1 . . . . . . . . . . . . . . 2 4 0 0 0 0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . r . . . . . 2 4 0 0 1 11 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . 2 4 0 0 1 11 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . 2 4 0 0 1 11 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . 1 2 0 0 1 11 . . . . . . . . . . . . . . . . . . . . . . + . . . . . r . . . . . . . . 1 2 0 0 1 11 . . . . . . . . . . . . . . . . . . . . . . . + . . . . + + . . . . . . . 1 2 1 20 2 22 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . 0 0 0 0 1 11 . . . . . . . . . . . . . . . . . . + + . + . + . . . . . . . . . . . . . 13 24 1 20 0 0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 2 4 0 0 0 0 . + 1 + + . . . . . . . . . . . . + + . + . . + 1 + + + + + . . . . . . . 12 22 5 100 2 22 . . . . . . . . . r . . . + . . . + + . . . . . . . . . . . . . . . . . . 11 20 0 0 0 0 . + . . . . + . . . . . . . . . . + 1 . . . . . . . . . . . . . . . . . . 7 13 0 0 0 0 . . . . . . . . . . . . . . . . . . + . . . . . . . + . . . . . . . . . . 1 2 1 20 0 0 . . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . . . . . . 1 2 0 0 0 0 . . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . . . . . . 1 2 0 0 0 0 . 1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 2 0 0 0 0 33 34 35 36 37 38 39 40 41 42 43 44 45 46 47 48 49 50 51 52 53 54 55 56 57 58 59 60 61 62 63 64 65 66 67 68 69 Pr . 1 -5 5 Fr . 1 -5 5 Pr . 5 6- 60 Fr . 5 6- 60 Pr . 6 1- 69 Fr . 6 1- 69 IGOR DAKSKOBLER & ANDREJ MARTINČIČ: A NEW ENDEMIC PLANT COMMUNITY WITH SCHOENUS NIGRICANS 26 FOLIA BIOLOGICA ET GEOLOGICA 59/1 – 2018 AF Aremonio-Fagion Cyclamen purpurascens E1 . . . . . . . . . . . . . + . + . . . . . . . . . . . . . + . . Anemone trifolia E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Knautia drymeia E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Hemerocallis lilioasphodelus E1 . . . + . . . . . . . . . . . . . . . . . . . . . . . . . . . . Omphalodes verna E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Euphorbia carniolica E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Potentilla carniolica E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . TA Tilio-Acerion Acer pseudoplatanus E1 . . . + . + . . . + . . . . . + . + . . . . . . . . . . . . + . Acer pseudoplatanus E2 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Tilia platyphyllos E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Ulmus glabra E2a . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . FS Fagetalia sylvaticae Fagus sylvatica E2 + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Fraxinus excelsior E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Salvia glutinosa E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Fraxinus excelsior E2 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Melica nutans E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . QP Quercetalia pubescenti-petraeae Ostrya carpinifolia E2a . . . 1 + + + . . + + . . . . + . . . . . . . . . + . . . . . r Sorbus aria E2a . . . . . . . . . + . . . . . . . . . . . . . . . . . . . . . . Fraxinus ornus E2 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Mercurialis ovata E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . Clematis recta E1 . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . Tamus communis E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . QR Quercetalia roboris Serratula tinctoria E1 . . . . . . . . . . . . . . . . . . . . . . . . . . r . . . . + Pteridium aquilinum E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Rubus hirtus E2a . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . QF Querco-Fagetea Corylus avellana E2a . r . . . r . . . . . . . . . . . . . . . . . . . . . . . . . . Anemone nemorosa E1 . . . . . . . . . . . . . + . . . . . . . . . . . . . . . . . . Carex digitata E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Carex umbrosa E1 . . . . . . . . . . . . . . + . . . . . . . . . . . . . . . . . Dactylorhiza fuchsii E1 . . . . . r . . . . . . . . . . . . . . . . . . . . . . . . . . Platanthera bifolia E1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Rosa arvensis E2a . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . ML Mosses (Mahovi) Orthothecium rufescens E0 . . . + . . . . . . . . . . . . . + 1 1 1 . . + 1 r . . 1 . . . Fissidens adianthoides E0 . . . . . . . . . . . . . + . . . . . . . . . . . r . 1 . . . . Neckera crispa E0 . . . . . . . . . . . . . . . . . . . . . . + . . . . + . . . . Ctenidium molluscum E0 . . . . . . . . . . . . . + . . . . . . . . . . . + . . . . . . Scleropodium purum E0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Tortella tortuosa E0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . Preissia quadrata E0 . . . . . . . . . . . + . . . . . . . . . . . . . . . . . . . . Jungermannia atrovirens E0 . . . . . . . . . . . + . . . . . . . . . . . . . . . . . . . . Campylidium calcareum E0 . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . . Riccardia chamaedryfolia E0 . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . . Heterocladium heteropterum E0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Leiocolea collaris E0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Neckera complanata E0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Orthothecium intricatum E0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Legend-Legenda 1-55 Astrantio carniolicae-Schoenetum nigricantis typicum 56-60: Astrantio carniolicae-Schoenetum nigricantis cladietosum marisci 61-62: Astrantio carniolicae-Schoenetum nigricantis var. Sesleria caerulea prov. 63-65: Primulo carniolicae-Seslerietum calcariae var. Schoenus nigricans 66-68 Primuletum carniolicae s. lat. var. Schoenus nigricans Number of relevé (Zaporedna številka popisa) 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 31 32 IGOR DAKSKOBLER & ANDREJ MARTINČIČ: A NEW ENDEMIC PLANT COMMUNITY WITH SCHOENUS NIGRICANS 27FOLIA BIOLOGICA ET GEOLOGICA 59/1 – 2018 . . . . . . . . . . . . . . . . . . . . . r . . . . . . . . . . . . . . . 4 7 0 0 0 0 . . . . . . . . . . . . . . . . . + . + + . . . . . . . . . + + . . . . . 3 5 0 0 2 22 . . . . . . . . . . . . . . . . . + . + . . . . . + . . . . . + . . . . . 2 4 1 20 1 11 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 2 0 0 0 0 . . + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 2 0 0 0 0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . r . . . . 0 0 0 0 2 22 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . 0 0 0 0 1 11 . . + . . . + . . r . . . . . . . + + . + . . . . + 1 . . . . . . . . . . 12 22 2 40 0 0 . . . . . . . . . . . . . . . . . + . . . . . . . . + r . . . . . . . . . 1 2 2 40 0 0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . r . . . . . . 0 0 0 0 1 11 . . . . . . . . . . . . . . . . . . . . . . . . . . r . . . . . . . . . . 0 0 1 20 0 0 . . . . . . . . . . . . . . . . . . . . . . . . . . + . . r . . . . . . . 1 2 1 20 1 11 . . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . . . . . . 1 2 0 0 0 0 . . . . + . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . . . 1 2 1 20 0 0 . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . . . 0 0 1 20 0 0 . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . . . 0 0 1 20 0 0 . . . . . . . . . . . . . . . . . . + . . . . + + + + . + + . . . . . . . 10 18 4 80 2 22 . . + . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . . . . . 3 5 0 0 0 0 . 1 . . . . . . . . . . . . . . . . + . . . . + . + + r . . . . . . . . . 2 4 4 80 0 0 . . . + . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 2 4 0 0 0 0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 2 0 0 0 0 . r . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 2 0 0 0 0 . + . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . . . . . . 4 7 0 0 0 0 . . . . . . . . . . . . . . . . . . + . . . . . . . . r . . . . . . . . . 1 2 1 20 0 0 . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . . . . 0 0 1 20 0 0 . . . . . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . . . 3 5 0 0 0 0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 2 0 0 0 0 . . . . . . . . . . . . . . . . . . . . . 1 . . . . . . . . . + . . . . . 1 2 0 0 1 11 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 2 0 0 0 0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 2 0 0 0 0 . . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . . 0 0 1 20 0 0 . . . . . . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . . . 0 0 1 20 0 0 . . . . . . . . . . . 2 1 1 1 . . . . . . . . . . . . . . . + . . 1 1 . . 13 24 0 0 3 33 . . . . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . . + . 4 7 0 0 1 11 . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . . 3 5 0 0 0 0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . + + + 1 . . . + . 2 4 0 0 5 56 . . . . . . . . . . . . . . . . . . . . . + . . . . . . . . . . . . . . . 1 2 0 0 0 0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . + . . . . . 1 2 0 0 2 22 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 . . . . + . 1 2 0 0 2 22 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 2 0 0 1 11 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 2 0 0 0 0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 2 0 0 0 0 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . r . . . . 0 0 0 0 1 11 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . 0 0 0 0 1 11 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . 0 0 0 0 1 11 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . + . 0 0 0 0 1 11 33 34 35 36 37 38 39 40 41 42 43 44 45 46 47 48 49 50 51 52 53 54 55 56 57 58 59 60 61 62 63 64 65 66 67 68 69 Pr . 1 -5 5 Fr . 1 -5 5 Pr . 5 6- 60 Fr . 5 6- 60 Pr . 6 1- 69 Fr . 6 1- 69 IGOR DAKSKOBLER & ANDREJ MARTINČIČ: A NEW ENDEMIC PLANT COMMUNITY WITH SCHOENUS NIGRICANS 28 FOLIA BIOLOGICA ET GEOLOGICA 59/1 – 2018 69 Pinguicula alpina community prov. Pr. Presence (number of relevés in which the species is presented) - število popisov, v katerih se pojavlja vrsta Fr. Frequency in % - frekvenca v % A Limestone - apnenec D Dolomite - dolomit C Chert - roženec Tu Tufa - lehnjak Gr Gravel - grušč Hy Molic Gleysols - organsko-mineralna tla Li Lithosol - kamnišče Re Rendzina - rendzina